Reservation Status of Tasmanian Native Higher Plants By Naomi Lawrence - David Storey - Jennie Whinam Biodiversity Conservation Branch February 2008 Depar tment of Pr imar y Industr ies and Water Reservation Status of Tasmanian Native Higher Plants 1 Authors: N. Lawrence, D. Storey and J. Whinam This book is copyright. Copyright © 2008 Crown in right of State of Tasmania. Drawings: Richard Hale Apart from fair dealing for the purposes of private study, research, criticism or review as permitted under the Copyright Act 1968, no part may be reproduced by any means without the permission of the publisher. Design: Graphic Services, Information & Land Services Division The Crown does not accept legal responsibility for the accuracy, completeness, or relevance to the user’s purpose of the information herein and those using it for whatever purpose are advised to verify it and to obtain any appropriate professional advice. ISBN-13: 978 0 7246 6431 3 (book) ISBN-13: 978 0 7246 6432 0 (pdf) Published by: Department of Primary Industries and Water, GPO Box 44 Hobart, Tasmania 7001. Cite as: Lawrence N., Storey D. & Whinam J. (2008). Reservation Status of Tasmanian Native Higher Plants, Biodiversity Conservation Report 07/1. Department of Primary Industries and Water, Tasmania, Australia. All of the data used in this assessment can be made available electronically by the Department of Primary Industries and Water (DPIW) upon request. An extract of the data in this report will be made available on the Natural Values Atlas (NVA) that is maintained by DPIW. Additional data can also be made available on request, including the full data set ordered by reserves (providing a list of all species that have been recorded in each reserve) and ordered by species (providing a list of all reserves in which each species occurs). This will enable users to undertake analyses of the data that are tailored to their own requirements. For data requests contact the Vegetation Section of DPIW Phone: 03 62336556 E-mail: [email protected] CONTENTS Acknowledgments 1 Executive summary 2 Background 3 Methods 4 Results 5 Summary 5 Look up table and explanations 16 Breakdown for each taxon 19 Appendices 329 References 339 ACKNOWLEDGEMENTS The authors are greatly indebted to the staff of the Tasmanian Herbarium for their help with distributional information and taxonomic changes and all those who have contributed data to the Natural Values Atlas (previously GTSPOT) over the years. We would like to thank the many people who have assisted in the development of the data set used to undertake the assessment. In particular, we would like to thank Mick Brown, Jamie Kirkpatrick, Fred Duncan and Louise Gilfedder for their assistance and Penny Wells, Mick Brown, Brooke Craven and Stephen Harris for their advice and guidance. We also acknowledge the invaluable technical assistance provided by Kristy Goddard, Felicity Faulkner, Wieslawa Misiak and Adam Smith. We gratefully acknowledge the skills of Gina Donnelly and Andrew Gibson, who designed the cover and layout of the book, and Richard Hale for the botanical illustrations. Funding for the research for this report and the review of the database was largely provided by the Tasmanian Wilderness World Heritage Area Program. Reservation Status of Tasmanian Native Higher Plants 1 EXECUTIVE SUMMARY This report provides a range of information on the reservation status of vascular plant species in Tasmania, including records of individual species within the reserve system and summary analyses of various trends in the data. The report aims to support conservation planners, biological consultants, land managers and land use decision-makers by providing them with detailed information on the reservation status of individual species and groups of species. The information underpins various reporting requirements including the Tasmanian Reserve System and the National Reserve System, and is intended to be freely available. The full report includes detailed information on the reservation of each species at the State and bioregional level, as well as summary information relating to a series of indicators. These indicators include: »» the extent of reservation, including whether the species is fully reserved (reserved at all of its locations), reserved in all bioregions, or reserved in more or less than half of the bioregions in which it occurs; »» information on the size classes of reserves in which the species occurs (eg occurs in one or more reserves that are >500ha, or >1,000 hectares); »» information on the type of reserves in which the species occurs (eg informal reserves, private reserves and dedicated formal reserves including the World Heritage Area); and »» stochastic risk (measured by whether the species has a restricted distribution either in a single reserve or in adjoining reserves, meaning that it is at potential risk of a stochastic event affecting the entire population for the species). The introductory section of the report provides summary information relating to the above indicators, which has been analysed at the family level and supra-group level (ie dicotyledons, gymnosperms, monocotyledons and pteridophytes). It includes summary data on the total number, and percentage, of species that are unreserved, partially reserved, and fully reserved, as well as lists of species that were unreserved, only recorded from one reserve, or only recorded from one reserve class as at June 2006. A list of reserves for which flora data was not available at the time of this analysis is also provided. The assessment is based on the Tasmanian CAR Reserves Spatial Layer as at June 2006, and reserve type classifications follow the Tasmanian Regional Forest Agreement 1997. Species location data is based on the Natural Values Atlas as at June 2005 which also incorporates data from the Tasmanian Herbarium1. The NVA is the most comprehensive data set of species locations currently in Tasmania. However it does not, as yet, capture all of the survey data that exists for Tasmanian species and therefore there are still some gaps in the information, which this project has helped to identify. For subsequent assessments all efforts will be made to identify any data gaps and to locate and incorporate this data into the NVA. The methodology is explained in detail in the Background Section to the report and appendix 2. 1 All of the data used in this assessment can be made available electronically by the Department of Primary Industries and Water (DPIW) upon request. An extract of the data in this report will be made available on the Natural Values Atlas (NVA) that is maintained by DPIW. Additional data can also be made available on request, including the full data set ordered by reserves (providing a list of all species that have been recorded in each reserve) and ordered by species (providing a list of all reserves in which each species occurs). This will enable users to undertake analyses of the data that are tailored to their own requirements. B ACKGROUND This report provides information on the reservation status of the native vascular flora in Tasmania. The Tasmanian reserve system forms an integral part of the Australian reserve system and represents the combined efforts of public and private land-holders to achieve a state wide system of terrestrial protected areas. The state’s reserve system is a major contributor to the conservation of native biodiversity at the state and national level. Within this system, reserves that have perpetual protection have the greatest capacity to protect our native biodiversity in the long term and are thus a major and important component of the system. The aim of the reserve system is to protect examples of all regional ecosystems, along with all the elements that make up these ecosystems and their associated conservation values, including individual native vascular plant species. As a part of reviewing the effectiveness of the reserve system in achieving this aim, it is important to know which species occur within the reserve system, which do not and which are under represented. To this end, periodic assessments of the reservation status of the biodiversity within the reserves needs to be undertaken, using new information about reserves, species and species assemblages. Botanical surveys undertaken in the 1970s revealed that a large proportion of species were either unreserved or poorly reserved at that time, particularly species within dry sclerophyll vegetation. The earliest assessment of the reservation and conservation status of native vascular flora in Tasmania was an analysis of unreserved species, undertaken by Brown et al. (1977). The list of unreserved higher plant species was revised and a list of poorly reserved species was compiled and published by Duncan (1985). As the pool of information on species locations expanded, and the requirement for information to help identify areas to be incorporated into the reserve system emerged, the need for an updated assessment became evident. This was Reservation Status of Tasmanian Native Higher Plants undertaken by Kirkpatrick et al. (1991a, 1991b) and remains the most recent published assessment of the reservation status of Tasmanian native vascular plant species prior to this report. There were further upgrades to the published data as part of the studies leading to the Tasmanian Regional Forest Agreement 1997 (R. Barnes and J. Sheridan 1996) however this data was not widely circulated. In addition, North et al. (1998) carried out a large number of surveys in reserves and analysed the contribution of the resulting flora records to the reservation of the State’s flora. Since that time a significant amount of additional data has been collected, the reserve system has been expanded, and plant taxonomy has undergone revision. Advances in technology, particularly desk-top GIS and spatial data sets, have also enabled a more complex and comprehensive analyses of species reservation status to be undertaken. The 1991 assessments of Kirkpatrick et al. predated the establishment of the National Reserves System (NRS) Program and the Regional Forest Agreement (RFA). There is a need to align the assessment of species reservation status with the requirements of both the NRS and RFA for comprehensiveness, adequacy and representativeness (see definitions, Appendix 1). These assessments also predated the development of bioregional boundaries under the Interim Bioregionalisation of Australia (IBRA) system. Bioregions and subregions are now used as a key mechanism for assessing gaps in the current reserve system and identifying priorities for improved representation of ecosystems and their components. be undertaken in the near future to review other special attributes of native vascular species. Rather than providing a simple categorisation of the reservation status of each species, such as “well, moderately or poorly reserved”, this assessment provides both the full data set and a more detailed analysis against a range of indicators that contribute to the adequacy, comprehensiveness and representativeness of each species in the reserve system. These reservation indicators are designed to form a useful tool to assist those involved in land-use planning, management and decision making. Further indicators could potentially be developed for different users as needs arise. Unlike the assessments undertaken in 1991 that also examined conservation status, the current assessment focuses only on the reservation status of Tasmanian native vascular plants. The more complex task of undertaking a review of the conservation status of native plants is now predominantly a process undertaken under the auspices of the Tasmanian Threatened Species Protection Act 1995 and the Commonwealth Environment Protection and Biodiversity Conservation Act 1999. However, subsequent work will 3 METHODS Geo-referenced point data for native vascular plant species was extracted from the Natural Values Atlas in 2005 and intersected with various spatial layers including CAR reserves, bioregions, 10km grid and coasts and islands. The data extracted was then exported into other database programs and queried to generate the results in accordance with the indicators listed below in table 1. The methods used to undertake the analysis are described in detail in appendix 2. It is important to note that not all plant survey data for the state has been captured within the NVA and thus there will be some gaps in the data. Consequently the reservation status of some species is likely to be underestimated. This assessment will serve to help identify some of these gaps and these will become priorities for incorporation into the NVA for future assessments. Aligning the analysis with the Reserve System requirements for CAR All previous assessments of the reservation status of Tasmanian vascular flora predate the National Reserve System Program and the Tasmanian Regional Forest Agreement 1997 (RFA). Under the RFA the Tasmanian and Australian Governments agreed to establish a comprehensive, adequate and representative (CAR) reserve system to ensure the longterm conservation and protection of Tasmania’s biodiversity, old growth forest and wilderness values. The current assessment aligns with CAR principles (Appendix 1) and the reserve categories defined under the RFA. The indicators that this assessment reports against were developed to help determine the degree to which the CAR principles are being met for native vascular plant species. The CAR reserve system established under the RFA comprised existing and new reserves on both public and private land. The reserve classes used for this assessment are: Indicators »» Formal Reserves. Reserves equivalent to IUCN Protected Area Management Categories I, II, III, IV, or VI as defined by the IUCN Commission for National Parks and Protected Areas (1994). The status of Formal Reserves is secure, requiring action by the Tasmanian Parliament for revocation. Some Formal Reserves are also Dedicated Reserves, which, in Tasmania, comprise the following reserves: National Parks, State Reserves, Game Reserves, Nature Reserves, Historic Sites and Forest Reserves not subject to the Mineral Resources Development Act 1995 (Tas)2 . The indicators reported against in this assessment are those which would best assist in the determination of the degree to which the CAR requirements are currently met for a species by the current reserve system. There are 13 indicators that were reported against in this assessment, (Table 1). »» Informal Reserves. Reserves, other than a Forest Reserve, on State Forest comprising an area identified as Protection Zone under the Management Decision Classification System, or other administrative reserve on public land, which is managed to protect CAR values. »» Private CAR Reserves. Private land that is managed in the long term for the protection of CAR values under secure arrangements, including proclamation under legislation, contractual agreements such as management agreements and covenants, and reserves set aside under independently certified forest management systems. »» Values managed by prescription. These include those areas protected by management prescription under the Forest Practices Code or in Forest Management Plans. This assessment includes only formal, informal and private CAR reserves, as values managed by prescription are not spatially defined across their full extent and, as a consequence, are not available within the CAR reserves spatial layer. The definitions for CAR detailed in appendix 1 were used as the basis for developing useful indicators that would enable assessments of reservation status to be undertaken by others. 2 The descriptions for Formal and Dedicated Reserves are based on the definitions in the Tasmanian Regional Forest Agreement, November 1997. Table 1. The 13 assessment indicators No. Indicator Explanation 1 Fully Reserved All records occur with in the CAR reserve system. 2 Partially reserved 1 Examples reserved in all bioregions in which the species occurs. 3a Partially reserved 2 Reserved in half or more of the bioregions in which the species occurs. Indicates Comprehensiveness Representativeness Comprehensiveness Representativeness Comprehensiveness Representativeness 3b Partially reserved 3 Reserved in less than half the bio-regions in which the species occurs Comprehensiveness Representativeness 4 Not reserved Was not recorded with in the CAR reserve system. Comprehensiveness Representativeness 5 No data There were no observations for the species recorded in the NVA as at 2005. 6 Not in a reserve >1000 Ha Was not recorded in any CAR reserve greater than 1000 hectares. Adequacy 7 Not in a reserve > 500 Ha Was not recorded in any CAR reserve greater than 500 hectares. Adequacy 8 Not in a dedicated formal reserve There are no records occurring in any dedicated formal reserve. Adequacy 9 Reserved only in Private reserves All the records in reserves occur only in private reserves. Adequacy 10 Reserved only in informal reserves All the records in reserves occur only in informal reserves. Adequacy 11 Reserved only in the WHA All records in reserves occur only within the Tasmanian Wilderness World Heritage Area Adequacy 12 Potential stochastic risk Potentially has restricted distribution within a single or adjoining reserve/s Adequacy *Note: The analysis was based on NVA records as at June 2005. Because the data used in the assessment is point data, it is not possible to indicate what proportion of the species population is incorporated into the reserve system. Therefore it is currently not possible to have an effective indicator for this component of the adequacy requirement. The indicators relating to reserve size and reserve number have only been reported on for the dedicated formal and formal reserves in this assessment. This is because in the current CAR reserve layer informal reserves on State forest (which are, in many places, interconnected linear reserves in the landscape) have been coded with the same identification tag and grouped as the same reserve. The same is also true of some groups of private reserves. This makes it difficult to undertake a meaningful analysis on these reserves for indicators 6 and 7 in particular. Reservation Status of Tasmanian Native Higher Plants 5 RESULTS Table 2. Comparison of results for 1991–2007 Reservation Status Summary This section provides an overall summary of the results of this assessment. The Macquarie Island species were assessed separately and the results are presented at the end of this section. The results for each of the species assessed are given in the next section. No. of species % 1991 2007 1991 2007 Unreserved 256 74 14.3 4.1 Partially or fully reserved 1535 1693 85.7 93.7 100 100 No data 40 Total 1791 1807 2.2 *Note: Results based on NVA records as at June 2005. There were 1807 taxa assessed, of these 474 are endemic and 459 (25%) are listed under the Tasmanian Threatened Species Protection Act 1995 and/or the Commonwealth Environment Protection and Biodiversity Conservation Act 1999. The data in table 2 shows that the number of species not reserved has dropped considerably from 14.3% of the native vascular plant species assessed in 1991 to 4.0% of species assessed in 2007. The increase in the number of species assessed is due to taxonomic changes and the discovery of new species over the 16 years since the last assessment was published. Table 2 presents a comparison of the results of the 1991 and the 2007 assessments. It was not possible to do an exact comparison in accordance with the criteria used in the 1991 assessment for two reasons: »» the exact identity of many of the species assessed in 1991 could not always be determined due to taxonomic changes. »» the original data set used to undertake the analysis could not be located in electronic form and it was not possible to efficiently recreate the data from the hardcopy of the 1991 report. Table 3. Breakdown of results against indicators for supra groups Indicators 1 2 3a 3b 4 5 6 7 8 9 10 11 12 No. taxa in group Fully res. Res. in all bio-regions occurs Res. in >50% bioregions Res. in <50% bioregions Not reserved. No data Not in res. >1000 ha Not in res. >500 ha Not in dedic. formal res. Private reserves only Informal res’s only All res’d in WHA Stochastic risk ? Dicotyledonae 1073 65 506 473 26 41 27 34 22 75 1 5 14 29 Gymnospermae 11 0 7 3 1 0 0 0 0 0 0 0 0 0 Monocotyledonae 622 39 244 305 29 31 13 32 17 57 9 11 6 29 Supra Group Pteridophyta 101 4 59 37 3 2 0 3 1 5 0 0 2 3 Totals 1807 108 816 818 59 74 40 69 40 137 10 16 22 61 *Note: Results based on NVA records as at June 2005. Table 3 shows a breakdown of the results by supra group. Note that some species fall into more than one indicator category. For example, a species that is ‘fully reserved’ will also be ‘reserved in all bioregions in which it occurs’. There is a relatively low proportion of the species (6%) within the dicotyledons that are fully reserved, a little less than half (47%) are reserved in all the bio-regions in which they occur, less than half (46%) of the dicotyledons are partially reserved and 4% are not reserved. Most of the dicotyledons that are reserved occur within either dedicated formal or formal reserves that are larger than 500 hectares. For the monocotyledons, relatively few are fully reserved (6.5%) or not reserved (5%). Close to half of all the monocotyledons are reserved in all the bioregions in which they occur. There are relatively few monocotyledons not reserved in a reserve greater than 1000 hectares (5%) or which do not occur in a dedicated formal reserve (9%). All the native conifer species (Gymnospermae) are reserved with none fully reserved, with more than half reserved in all the bioregions in which they occur. All gymnosperms are within reserves of greater than 1000 hectares in the formal reserve system and all occur in at least one, dedicated formal reserve. Nearly all of the ferns and fern allies (Pteridophyta) are reserved, with only 2 species unreserved. There are very few Pteridophytes reserved in less than half the bioregions in which they occur (3%) or which do not occur in a dedicated formal reserve or in a reserve greater than 1000 hectares. Overall there are relatively few species (6%) that are fully reserved, or not reserved (4%) or for which there is no data (2%). Overall more than half (54%) of all the native vascular plant species in the state are reserved in all the bioregions in which they occur. The species for which there is currently no data are either monocotyledons or dicotyledons, with dicotyledons making up over half (67%) of these. The data for species that are reserved only in one reserve class are given in table 4. Less than 2% of all species are reserved only in private or informal reserves. This is a low percentage for the private and informal reserve system and reflects that much of the survey data for these reserves was not in the NVA at the time this assessment was undertaken. Reservation Status of Tasmanian Native Higher Plants Thus this result is not an accurate indication of the extent to which private and informal reserves are increasing representativeness of species within the reserve system. A larger percentage of species occur only in dedicated formal and formal reserves. While this would be expected, given that the largest number of species is represented within them, it may also be influenced by the limited data for private and informal reserves available at the time of this analysis. Of the dedicated formal reserves, the WHA has 26% of the species that only occur in this reserve class. A list of species which only occur in each of the reserve classes (including WHA) is given in appendix 3. Table 5 shows the number of reserves for which there is currently no vascular flora species data recorded in the data set within each of the reserve classes. A list of these reserves is provided in appendix 4. There are in total 305 reserves with no vascular flora species data, which is 30% of all the reserves (excluding informal). Table 5. No. of reserves with no vascular flora species data recorded Reserve Class Reserve type Dedicated Formal Forest Reserve Table 4. No. of species which only occur in one reserve class Reserve Class No. of species % of vasc flora 84 4.6 Dedicated formal reserves WHA Component 22 1.2 Formal reserves 67 3.7 Private reserves 10 0.5 Informal reserves 16 0.9 *Note: Results based on NVA records as at June 2005. 2 Game Reserve 5 Historic Site 17 National Park 1 Nature Reserve 26 State Reserve 14 Total Formal No 78 Conservation Area 31 Forest Reserve subject to MRDA 13 Nature Recreation Area Total 3 34 Private Nature Reserve* Private Sanctuary* Total Informal Total 6 187 193 Data not available N/A 305 * Note – where a Private Nature Reserve or a Private Sanctuary also has a conservation covenant registered on title, the Private Nature Reserve or Private Sanctuary is given precedence in reporting (ie it will not also be counted in the conservation covenant category). Results based on NVA records as at June 2005. A breakdown of the data by family group was undertaken to determine if there were families that stood out as relatively less well reserved based on the indicator data. The indicators provide an overall indication of how well CAR principles are being met for each family group. (Table 6). 7 Table 6. Breakdown of assessment results by family INDICATOR FAMILY No of taxa 1 2 3a 3b 4 5 6 7 8 9 Fully res’d Reserved in all bio-reg’s occurs Res’d in >0.5 bioreg’s occurs Res’d in <0.5 bioreg’s occurs Un-res’d No data Not in res >1000 ha in any bio-reg Not res >500 ha in any bio-reg Not in dedic. formal res. Res’d priv. res. only 10 11 Res’d inform Res’d only in res. only WHA 12 Stochastic risk? Adiantaceae 7 0 2 4 1 0 0 0 0 0 0 0 0 1 Aizoaceae 4 0 2 1 1 0 0 0 0 0 0 0 0 0 Alismataceae 1 0 0 1 0 0 0 0 0 1 0 0 0 1 Amaranthaceae 2 0 1 1 0 0 0 0 0 1 0 0 0 0 Apiaceae 39 2 14 24 0 1 0 0 0 4 0 1 2 3 Apocynaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Araliaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Asclepiadaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 9 0 6 3 0 0 0 1 1 1 0 0 0 0 Asteraceae Aspleniaceae 200 19 94 80 6 11 9 8 5 11 0 0 2 3 Athyriaceae 2 0 0 2 0 0 0 0 0 0 0 0 0 0 Azollaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Bignoniaceae 1 0 1 0 0 0 0 0 0 1 0 0 0 0 Blechnaceae 11 0 8 3 0 0 0 0 0 1 0 0 0 0 Boraginaceae 5 0 2 2 0 1 0 0 0 0 0 0 0 0 Brassicaceae 22 4 6 11 1 3 1 2 2 3 0 0 0 3 Brunoniaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Burmanniaceae 1 0 1 0 0 0 0 0 0 0 0 0 0 0 Callitrichaceae 3 0 0 1 1 1 0 0 0 1 0 0 0 1 Campanulaceae 17 0 7 10 0 0 0 1 1 1 0 0 0 1 Caprifoliaceae 1 0 1 0 0 0 0 0 0 0 0 0 0 0 Caryophyllaceae 16 2 8 7 0 0 1 1 1 2 0 0 2 0 Casuarinaceae 7 1 5 2 0 0 0 0 0 0 0 0 0 0 Centrolepidaceae 14 4 7 6 1 0 0 0 0 1 0 0 1 0 Chenopodiaceae 15 0 5 7 0 1 2 1 1 0 0 0 0 0 Clusiaceae 2 0 2 0 0 0 0 0 0 0 0 0 0 0 Convolvulaceae 7 0 4 3 0 0 0 2 2 2 1 0 0 0 Crassulaceae 7 0 3 3 0 1 0 0 0 0 0 0 0 0 INDICATOR FAMILY No of taxa 1 2 3a 3b 4 5 6 7 8 9 Fully res’d Reserved in all bio-reg’s occurs Res’d in >0.5 bioreg’s occurs Res’d in <0.5 bioreg’s occurs Un-res’d No data Not in res >1000 ha in any bio-reg Not res >500 ha in any bio-reg Not in dedic. formal res. Res’d priv. res. only 10 11 Res’d inform Res’d only in res. only WHA 12 Stochastic risk? Cucurbitaceae 1 1 1 0 0 0 0 0 0 1 0 0 0 0 Cunoniaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Cupressaceae 6 0 4 1 1 0 0 0 0 0 0 0 0 0 Cuscutaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Cyatheaceae 3 0 1 2 0 0 0 0 0 0 0 0 0 0 Cyperaceae 118 6 57 53 4 4 0 2 1 5 1 0 2 5 Dennstaedtiaceae 7 0 6 1 0 0 0 1 0 0 0 0 0 0 Dicksoniaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Dilleniaceae 14 0 7 4 0 1 2 0 0 1 0 0 0 0 Donatiaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Droseraceae 8 1 4 4 0 0 0 0 0 0 0 0 0 0 Dryopteridaceae 4 0 4 0 0 0 0 0 0 0 0 0 0 0 Elaeocarpaceae 2 0 0 2 0 0 0 0 0 0 0 0 0 0 Elatinaceae 1 0 0 0 1 0 0 0 0 0 0 0 0 0 Epacridaceae 94 6 52 40 1 0 1 1 1 5 0 0 2 3 Ericaceae 4 0 1 3 0 0 0 0 0 0 0 0 0 0 Escalloniaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Eucryphiaceae 3 1 1 2 0 0 0 0 0 0 0 0 0 0 Euphorbiaceae 13 1 10 3 0 0 0 0 0 2 0 0 1 1 Fabaceae 58 0 28 26 1 2 1 1 1 5 0 0 0 1 Fagaceae 2 0 2 0 0 0 0 0 0 0 0 0 0 0 Frankeniaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Fumariaceae 1 0 1 0 0 0 0 0 0 0 0 0 0 0 Gentianaceae 14 3 5 6 1 0 2 0 0 1 0 1 1 3 Geraniaceae 7 0 4 3 0 0 0 0 0 1 0 0 0 1 Gleicheniaceae 7 0 5 2 0 0 0 0 0 0 0 0 1 0 Goodeniaceae 15 1 6 6 0 0 3 0 0 0 0 0 0 0 Grammitidaceae 7 2 5 2 0 0 0 0 0 0 0 0 1 0 Gunneraceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Gyrostemonaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Haemodoraceae 1 1 1 0 0 0 0 0 0 0 0 0 0 0 Haloragaceae 20 0 4 13 2 0 1 1 0 4 0 0 0 0 Hydatellaceae 2 0 0 2 0 0 0 0 0 0 0 0 0 0 Hydrocharitaceae 2 0 1 1 0 0 0 0 0 1 0 0 0 0 Reservation Status of Tasmanian Native Higher Plants 9 INDICATOR FAMILY Hymenophyllaceae 1 2 3a 3b 4 5 6 7 8 9 No of taxa Fully res’d Reserved in all bio-reg’s occurs Res’d in >0.5 bioreg’s occurs Res’d in <0.5 bioreg’s occurs Un-res’d No data Not in res >1000 ha in any bio-reg Not res >500 ha in any bio-reg Not in dedic. formal res. Res’d priv. res. only 8 0 7 1 0 0 0 0 0 0 0 10 11 Res’d inform Res’d only in res. only WHA 0 0 12 Stochastic risk? 0 Iridaceae 6 0 2 4 0 0 0 0 0 0 0 0 0 0 Isoetaceae 5 0 1 3 0 1 0 0 0 0 0 0 0 0 Juncaceae 34 1 13 19 1 1 0 0 0 1 0 0 0 0 Juncaginaceae 7 0 0 5 1 0 1 0 0 0 0 0 0 1 Lamiaceae 15 0 7 7 0 1 0 0 0 2 0 0 0 1 Lauraceae 4 0 2 2 0 0 0 0 0 1 0 0 0 1 Lemnaceae 3 0 1 1 1 0 0 0 0 2 0 0 0 0 Lentibulariaceae 6 2 3 3 0 0 0 0 0 1 0 0 0 0 Liliaceae 40 3 13 23 2 0 2 2 2 3 2 0 1 0 Linaceae 2 0 0 2 0 0 0 0 0 0 0 0 0 0 Lindsaeaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Loganiaceae 6 0 2 4 0 0 0 0 0 0 0 0 0 0 Lycopodiaceae 9 1 5 4 0 0 0 0 0 0 0 0 0 0 Lythraceae 2 0 0 1 1 0 0 0 0 1 0 0 0 0 Malvaceae 4 0 0 3 1 0 0 0 0 0 0 0 0 0 Marsileaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Menyanthaceae 4 0 1 3 0 0 0 0 0 0 0 0 0 0 Mimosaceae 23 0 11 10 0 2 0 1 0 2 0 0 0 0 Monimiaceae 2 0 2 0 0 0 0 1 0 1 0 1 0 0 Myoporaceae 2 0 2 0 0 0 0 0 0 1 0 0 0 0 Myrtaceae 56 0 28 27 1 0 0 4 3 1 0 0 0 0 0 Oleaceae 1 0 1 0 0 0 0 0 0 0 0 0 0 Onagraceae 11 0 3 7 0 1 0 0 0 0 0 0 0 0 Ophioglossaceae 3 0 1 0 1 1 0 0 0 1 0 0 0 0 15 211 15 74 100 13 18 6 25 12 28 4 7 0 Osmundaceae Orchidaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Oxalidaceae 3 0 2 1 0 0 0 0 0 0 0 0 1 0 Pittosporaceae 11 0 6 3 0 2 0 0 0 0 0 0 0 0 Plantaginaceae 13 1 5 7 0 1 0 0 0 0 0 0 0 0 Plumbaginaceae 2 0 0 1 0 1 0 0 0 1 0 0 0 0 137 7 58 67 3 6 3 3 2 9 2 1 2 5 Poaceae INDICATOR FAMILY No of taxa 1 2 3a 3b 4 5 6 7 8 9 Fully res’d Reserved in all bio-reg’s occurs Res’d in >0.5 bioreg’s occurs Res’d in <0.5 bioreg’s occurs Un-res’d No data Not in res >1000 ha in any bio-reg Not res >500 ha in any bio-reg Not in dedic. formal res. Res’d priv. res. only 10 11 Res’d inform Res’d only in res. only WHA 12 Stochastic risk? Podocarpaceae 5 0 3 2 0 0 0 0 0 0 0 0 0 0 Polygalaceae 5 0 2 3 0 0 0 0 0 0 0 0 0 0 Polygonaceae 10 0 0 8 2 0 0 1 1 3 0 0 0 1 Polypodiaceae 1 0 1 0 0 0 0 0 0 0 0 0 0 0 Portulacaceae 6 0 1 3 0 2 0 0 0 1 0 0 0 0 Posidoniaceae 1 0 1 0 0 0 0 0 0 0 0 0 0 0 Potamogetonaceae 6 0 1 5 0 0 0 0 0 1 0 0 0 0 Primulaceae 1 0 0 1 0 0 0 0 0 0 0 0 0 0 Proteaceae 40 5 25 14 0 1 0 1 1 1 0 0 1 0 Psilotaceae 3 1 3 0 0 0 0 0 0 0 0 0 0 1 Pteridaceae 2 0 0 2 0 0 0 0 0 0 0 0 0 0 Ranunculaceae 25 0 9 13 0 2 1 0 0 0 0 0 0 0 Restionaceae 14 1 5 9 0 0 0 0 0 0 0 0 0 0 Rhamnaceae 28 0 11 15 1 1 0 0 0 2 0 0 0 0 Rosaceae 12 1 8 2 1 0 1 1 1 1 0 0 0 1 Rubiaceae 18 0 10 6 2 0 0 1 0 1 0 0 0 1 Ruppiaceae 3 0 0 1 1 1 0 0 0 0 0 0 0 0 Rutaceae 28 3 18 9 0 1 0 1 0 1 0 0 0 0 Santalaceae 8 0 4 3 0 1 0 0 0 0 0 0 0 0 Sapindaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Schizaeaceae 3 0 1 1 1 0 0 0 0 1 0 0 0 1 Scrophulariaceae 39 8 21 16 1 1 0 1 0 2 0 0 2 2 Selaginellaceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Solanaceae 4 0 1 2 0 1 0 0 0 0 0 0 0 0 Stackhousiaceae 5 0 4 1 0 0 0 1 1 2 0 0 0 0 Sterculiaceae 4 0 2 2 0 0 0 1 0 0 0 0 0 1 Stylidiaceae 9 1 5 3 0 1 0 0 0 0 0 0 0 0 Thelypteridaceae 1 0 0 1 0 0 0 1 0 1 0 0 0 0 Thymelaeaceae 21 0 10 9 0 0 2 1 0 2 0 1 0 1 Tremandraceae 5 0 3 2 0 0 0 0 0 1 0 0 0 0 Typhaceae 3 0 0 0 2 1 0 0 0 2 0 1 0 1 Urticaceae 5 1 4 1 0 0 0 1 1 0 0 0 0 0 Violaceae 9 1 5 4 0 0 0 0 0 1 0 0 0 0 Winteraceae 1 0 1 0 0 0 0 0 0 0 0 0 0 0 Reservation Status of Tasmanian Native Higher Plants 11 INDICATOR FAMILY Xanthorrhoeaceae 1 2 3a 3b 4 5 6 7 8 9 No of taxa Fully res’d Reserved in all bio-reg’s occurs Res’d in >0.5 bioreg’s occurs Res’d in <0.5 bioreg’s occurs Un-res’d No data Not in res >1000 ha in any bio-reg Not res >500 ha in any bio-reg Not in dedic. formal res. Res’d priv. res. only 5 0 1 4 0 0 0 0 0 0 0 10 11 Res’d inform Res’d only in res. only WHA 0 0 12 Stochastic risk? 0 Xyridaceae 4 0 3 1 0 0 0 0 0 0 0 0 0 0 Zannichelliaceae 7 1 4 2 0 0 1 0 0 3 0 2 0 1 Zosteraceae 2 0 1 1 0 0 0 0 0 0 0 0 0 0 Zygophyllaceae Totals 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1807 108 816 818 59 74 40 69 40 137 10 16 22 61 *Note: Results based on NVA records as at June 2005. There are two large families that do stand out, which together comprise almost one-quarter of all the species assessed: the Orchidaceae (12%) and Asteraceae (11%). The nature of the life history and ecology of species provides an insight as to why many species within these families are apparently less frequent in the reserve system. In the case of the Orchidaceae, some of the most extensively reserved vegetation types are also those with a low level of orchid diversity. Low light levels generally preclude orchids from being a significant component of rainforest, swamp forest or wet sclerophyll forests. Buttongrass moorlands are too acidic and alpine environments too harsh for many orchids to thrive. By far the greatest diversity of orchids is in dry eucalypt forests and woodlands. These forests are subject to frequent fires and other kinds of disturbance that favour orchid diversity and abundance. These forests and woodlands coincide with extensive tracts of non-reserved tenures. It is also probable that the orchids are under surveyed as many are virtually impossible to identify without flowers and flowering is often sporadic and may not occur for several seasons in a row. The distribution patterns of orchids is another factor possibly contributing to this situation. Orchids, in general, tend to occur in small, isolated, widely distributed populations, an attribute that would make them subject to a low level of chance of being picked up in surveys. This in turn might lead to orchid records giving a poor indication of their real distribution in reserves. The Asteraceae, like orchids, generally require disturbance for recruitment. Many species in the Asteraceae are annuals or short-lived perennials. As a result of their life histories and ecology, members of these families may be under-surveyed in the reserve system. In addition, both these families have a relatively high proportion of species, (33% of species in the Orchidaceae and 20% of species in the Asteraceae), listed as threatened under State and Commonwealth legislation. Another big group, the Cyperaceae, also has a large proportion of species for which the CAR principles are potentially not well met. This may reflect the fact that wetlands are not well reserved in many regions of the State and/or are not well surveyed, with 18% of species in this family also listed as threatened. Other groups with a relatively high proportion of species listed as threatened in Tasmania include the Fabaceae (32%), Epacridaceae (20%), and Rhamnaceae (46%). This data provides valuable information in regard to species or groups that may not be effectively conserved by reservation alone and may need different strategies to ensure they are effectively protected. It also indicates groups where additional survey data is required within the reserve system to gain a more accurate assessment for those species. To gain an indication of how well the current reserve system incorporates the full range of native vascular plants, a breakdown of the number of species not reserved in one or more of the bioregions in which they occur was undertaken (Table 7). Table 7 shows that there are many species, including common species (see results next section), that are not reserved in all of the bioregions they occur in. A further review of these species is required to determine if this adversely impacts on the effective conservation of them. Table 7. No. of species occurring in a bioregion that are not reserved in that bioregion Bioregion No. of species Figure 1. Reserve distribution by bioregion in Tasmania Total number and area of reserves Dedicated Formal Formal Private No. reserves Area ha No. reserves Area ha No. reserves Informal Area ha No. reserves Area ha. King 144 26 19200 34 51800 21 6000 - 14500 Northern Slopes 239 28 26500 77 48200 65 11100 - 51600 Flinders 163 30 51400 76 65000 24 2400 - 11800 West 45 13 801400 37 472500 3 1500 - 98700 Central Highlands 97 12 219400 41 207600 9 19200 - 30800 Northern Midlands 323 8 1700 20 8200 40 16500 - 1500 Ben Lomond 186 12 26300 57 67600 38 6300 - 38300 Southern Ranges 99 20 294500 37 20300 26 3600 - 71900 South East 170 82 80900 85 46800 114 18300 - 53500 Total 948 231 1521300 464 988000 340 84900 - 372600 Total number of all reserves 1037 Total area of all reserves Ha 2956800 ha *Note: Reserves with cross-bioregional boundaries will be counted in each of the bioregions in which they occur. A species may be unreserved in more than one bioregion, therefore the total number of species will not equal the combined total for each bioregion. Results based on NVA records as at June 2005. In many cases the result may be due to deficiencies in the data itself. For example, there are cases where a species has a single point recorded in a bioregion and where it has not been possible to determine what that record actually represents. It could potentially be a mis-identification; it may represent a single plant or a large population. It could, for example, be a single plant on a roadside in which case reservation of the species in that bioregion may not be possible or desirable. Any further review of these species would serve to increase the accuracy of the data in the Natural Values Atlas and to refine future analyses. The data in Table 7 and Figure 1 shows, as might be expected, that bioregions with relatively large numbers of species not represented in the reserve system are those that either have fewer reserves, in terms of number and/or area reserved, (for example the Northern Midlands), and/or are regions with large areas subject to intensive land uses. For example the Northern Slopes, Flinders and South East bioregions are major agricultural regions. In such bioregions the capacity Reservation Status of Tasmanian Native Higher Plants to expand the reserve system may be limited and other strategies for the conservation of unreserved species and other species for which the CAR requirements are not well met, may need to be considered. Table 8 shows the distribution of the reserves by class and type according to size class and the number of species represented within each reserve category. This data shows that over half of the dedicated formal reserves (66%) and other formal reserves (67%) and nearly all the private reserves (95%) are less than 500 ha in size. The proportion of reserves in the size classes less than and greater than 500 ha is very similar for dedicated formal reserves and other formal reserves. When looking at the number of native vascular flora species that are currently recorded as represented within these different reserve classes the highest representation (80%) is within the 19 National Parks. Although National Parks represent a small proportion in terms of number of reserves within the reserve system (2%) they are all substantially larger than 1000 ha and in area make up a large and significant component of the reserve system. Almost equal in terms of species represented within them are the Conservation Areas. There are, however, 185 of these Conservation Areas compared to 19 National Parks with a similar distribution of reserve size classes. Within the private reserves, Private Property Conservation Covenants have the highest number of species represented (35%) within them. Overall the total number of species in the State that are represented in dedicated formal (86%) and other formal (85%) reserves is very similar. The informal reserves currently hold 69% and the private reserves 49% of the entire native vascular flora according to the data currently held. These results however are probably an underestimate given that data for private reserves is not yet fully captured in the NVA and the same may be the case for informal reserves. 13 Table 8. Reserve size class by reserve class and type and number of species within each category Reserve size class Reserve class/type Dedicated formal reserves 6 6 25 5000-<10000 2000-<5000 1 3 20 4 44 1 2 2 1 12 1 2 3 8 1 345 1440 3 Nature Reserve 727 54 21 2 2 3 1 State Reserve 977 32 16 3 5 5 2 Wellington Park 490 10000<20000 20000<50000 50000<100000 100000 or > 1 29 1 3 63 50 7 13 32 11 Conservation Area 1428 87 52 10 11 13 7 Forest Reserves subject to Mineral Resources Act 967 19 72 36 20 20 4 Nature Recreation Area 902 7 9 2 1 1 2 1 1 Regional Reserve 757 1 2 3 5 3 4 2 1532 114 Conservation Agreement 0 3 Private land in WHA 60 Private Nature Reserve 172 Private Property Conservation Covenant 645 11 1 123 1 19 84 1556 Unable to separate 1 1 0 Total no. of reserves 1000-<2000 Historic Site Private Sanctuary Total all 500-<1000 National Park Total Total 10 478 406 Total Informal 100-<500 Forest Reserve Public authority land in WHA Private reserves <100 Game Reserve Total Other formal reserves No sp. 1 1 3 252 2 1 2 185 171 24 1 21 1 134 50 35 39 16 5 5 2 2 402 3 1 1 2 10 4 1 15 255 76 4 5 340 335 15 5 2 895 283 86 8 5 1 1 0 0 0 0 0 383 23 + + + + + + + + + + + + 525 271 66 54 52 23 17 6 3 5 1037 1250 *Note: species will occur in more than one of the reserve classes, so the total number of species that occur in the reserve classes (dedicated formal etc.) will not equal the combined total of the reserve types. Results based on NVA records as at June 2005. In conclusion, a substantially larger percentage of Tasmanian native vascular plant species has been recorded as occurring within the Tasmanian reserve system compared with the previous assessments of Kirkpatrick et al. in 1991. There are still a significant number of species (over 100) and some families for which the CAR principles are potentially not being well met (based on the data set as at 2005) and these need further assessment. There are a large number of species that do not have their full distributional range represented in the reserve system and further assessment of these species should be undertaken to determine if they are priorities for reservation in particular bioregions. Crassula moschata is the only species occurring on Macquarie Island that is listed as threatened (rare) under the Tasmanian Threatened Species Protection Act 1995. All species on Macquarie Island that do not occur in the rest of Tasmania, have been assessed as fully reserved. Where species also occur on the Tasmanian mainland the relevant indicator is given in the results. When looking at the number of native vascular flora species currently recorded as represented within these different reserve classes the highest representation is currently within the formal reserves, with National Parks alone having 80% of the native vascular flora represented within them. There are currently 74 species (a list of these is provided in appendix 5) that are not known to be reserved, and therefore should be specifically targeted for inclusion in the reserve system, where this is possible. There are also 137 species that are not represented within dedicated formal reserves, and 61 species (a list of these is provided in appendix 6) that are potentially subject to stochastic risk due to their distribution in the reserve system. There are 40 species for which there is currently no data and therefore are priorities for survey to establish how well reserved they are. Macquarie Island Given the substantial geographic separation of Macquarie Island from the rest of Tasmania and its sub-antarctic latitude (the rest of Tasmania is within temperate latitudes) a separate assessment has been undertaken for Macquarie Island vascular flora. A total of 40 native vascular plant species are recorded on Macquarie Island, these are listed in Buchanan (2005). Of these only eight species also occur in Tasmania (3 dicotyledons, 2 monocotyledons and 3 pteridophytes). Reservation Status of Tasmanian Native Higher Plants 15 Lookup Tables And Explanations This section explains how to use and interpret the results of the assessment undertaken. The tabulated results are presented in the breakdown of the results. The results are presented for each taxon assessed. Where the assessment was undertaken at species level and incorporates all data for all subspecies and varieties of that species this is indicated by # directly after the species name. Species names are listed in alphabetical order. An explanation regarding this is given in the methods in appendix 2. Table 9 and Table 10 provide definitions for each symbol used in the results breakdown for each taxon provided in the following section. Table 9. Key to Codes Code Is endemic to Tasmania t In Australia only occurs in Tasmania (occurs beyond Australia) r Rare in Tasmania v Vulnerable in Tasmania e Endangered in Tasmania x Extinct in Tasmania x? Presumed to be extinct in Tasmania VU Nationally Vulnerable EN Nationally Endangered CR Nationally Critically Endangered EX Nationally Extinct EW Nationally Extinct in the Wild Assessment undertaken at the species level. In this case all data for the species, including all subspecies and varieties of that species, was combined and used for the assessment. In all other cases the assessment was undertaken for each taxon. Field Name Other Formal Res. Table 10. Explanation of fields in the assessment table Field Name Explanation Species name, family For each species, the full name is given as it appears in Buchanan 2005, as well as its current common name as it appears in Wapstra et al. 2005. The family to which the species belongs is also listed. Endem. Taxa that are considered endemic at state and/ or national level are indicated. See Table 9 for code meanings. Listed State/Nat Species listed in the EPBCA 1999 (codes in capital letters) and/or the TSPA 1995 (codes in lower case letters) are indicated. See Table 9 for code meanings. Bioreg. A list of the number and name of each bioregion in which the species occurs. Records The total number of species records currently recorded within each bioregion Reserves The number of species records currently recorded within reserves in each bioregion No. of Reserves The total number of reserves in which the species occurs within each bioregion listed. Dedic. Formal Res. The number of dedicated formal reserves, as defined on page 4 in which the species occurs for each bioregion listed and how many of these reserves are of a size greater than 1000 ha. Explanation Y # There are 16 columns of information and data provided in the assessment results. Table 10 explains how to interpret the information presented in these columns. No. Ha »»Is the total number of reserves in the bioregion listed that are greater than 1000 ha. The number of other formal reserves, as defined on page 4 in which the species occurs for each bioregion listed and how many of these reserves are of a size greater than 1000 ha. There are two sub fields under this field which provide this information. »»Is the total number of reserves for the bioregion listed. No. Ha »»Is the total number of reserves in the bioregion listed that are greater than 1000 ha. Informal Res. The number of informal reserves, as defined on page 4 in which the species occurs for each bioregion listed and how many of these reserves are of a size greater than 1000 hectares. There are two sub fields under this field which provide this information. »»Is the total number of reserves for the bioregion listed. No. Ha »» Is the total number of reserves in the bioregion listed that are greater than 1000 ha. Private Reserves The number of private reserves, as defined on page 4, in which the species occurs for each bioregion listed and how many of these reserves are of a size greater than 1000 ha. There are two sub fields under this field which provide this information. »»Is the total number of reserves for the bioregion listed. There are two sub fields under this field which provide this information. »»Is the total number of reserves for the bioregion listed. Explanation No. Ha »»Is the total number of reserves in the bioregion listed that are greater than 1000 ha % of Records in Reserves The percentage of the total number of records for a species occurring within reserves in each of the bioregions listed. Indicator The 13 indicators numbered from 1-12 as defined in the table below (Table 11) Table 11. Explanation for indicator numbers in the indicator columns Indicator Explanation 1 Fully Reserved: 2 Partially reserved 1. 3a Partially reserved 2. 3b Partially reserved 3. 4 Not reserved: 5 No data 6 Not in a reserve > 1000 ha 7 Not in a reserve > 500 ha. 8 Not in a dedicated formal reserve 9 Reserves only in private reserves 10 Reserved only in informal reserves 11 Reserved only in the WHA 12 Potential stochastic risk Note. For expanded explanations refer to table 1. Reservation Status of Tasmanian Native Higher Plants 17
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