Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) Big Idea 1: Evolution a.Structural and functional evidence supports the relatedness of all domains (homeostatic mechanisms). Organisms are linked by lines of descent from common ancestry. (1.B.1.) Lecture Presentations for • Early taxonomists - Two Kingdoms: plant & animal • Late 1960’s - Five Kingdoms: Monera, Protista, Plantae, Fungi, Animalia. • Modern hypothesis - Eukarya Bacteria Biology Archaea • The earliest organisms Eighth Edition Neil Campbell and Jane Reece Pearson Education, Inc., publishing as Person Benjamin Cummings College Board, AP Biology Curriculum Framework 2012-2013 form a “ring of life” through horizontal gene transfer. 25.1, 25.3 Fig. 26-23 Copyright © Rebecca Rehder Wingerden Bozeman Biology: Homeostatic Evolution (12:00 min.) Copyright © 2012 Rebecca Rehder Wingerden Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) http://www.bozemanscience.com/021-homeostatic-evolution Fig. 26-21 Land plants Green algae transferred from one genome to another through mechanisms such as exchange of transposable elements and plasmids, viral infection and perhaps fusion of organisms. • 3 Copyright © 2012 Rebecca Rehder Wingerden Amoebas Cellular slime molds • Most recent common • Bacteria 2 3 1 Eukarya Archaea 4 3 2 1 Billions of years ago 0 Archaea consist of a diverse group of prokaryotic organisms that inhabit a wide variety of environments. Fungi Sulfolobus Nuclear envelope Green nonsulfur bacteria Thermophiles Halophiles (Mitochondrion) Spirochetes Chlamydia COMMON ANCESTOR OF ALL LIFE Methanobacterium Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) Euglena Trypanosomes Leishmania Animals Fig. 26-22 The role of horizontal gene transfer in the history of life. This tree show two major episodes of horizontal gene transfer, the dates of which are uncertain. It is known that many more such events occurred. 1. DNA and RNA are carriers of genetic information through transcription, translation and replication. Dinoflagellates Forams Diatoms Ciliates Red algae 1 2 Eukarya consists of all the organisms that have cells containing true nuclei, singlecelled as well as multicellular plants, fungi, and animals EUKARYA • Horizontal gene transfer - process in which genes are ancestor of all living things. Gene transfer between mitochondrial ancestor and ancestor of eukaryotes Gene transfer between chloroplast ancestor and ancestor of green plants. Three Domains: - Bacteria - Archaea - Eukarya ARCHAEA Copyright © 2012 Rebecca Rehder Wingerden Green sulfur bacteria Bacteria contains most of the currently known prokaryotes, including the bacteria closely related to chloroplast and mitochondria BACTERIA Cyanobacteria (Plastids, including chloroplasts) Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) 2. Major features of the genetic code are shared by all modern living systems. Expression of genes from different species. Because diverse forms of life share a common genetic code, one species can be programmed to produce proteins characteristic of a second species by introducing DNA from a second species into the first. DNA TRANSCRIPTION Pre-mRNA RNA PROCESSING mRNA TRANSCRIPTION DNA TRANSLATION mRNA Ribosome Polypeptide (a) Bacterial cell. In a bacterial cell, which lacks a nucleus, mRNA produced by transcription is immediately translated without additional processing. TRANSLATION Ribosome Polypeptide (b) Eukaryotic cell. The nucleus provides a separate compartment for transcription. The original RNA transcript, called pre-mRNA, is processed in various ways before leaving the nucleus as mRNA (a) Tobacco plant expressing a firefly gene Fig. 17-3a-2 Copyright © 2012 Rebecca Rehder Wingerden Copyright © 2012 Rebecca Rehder Wingerden (b) Pig expressing a jellyfish gene Fig. 17-6 Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) 3. Metabolic pathways are conserved across all currently recognized domains. b. Structural evidence supports the relatedness of all eukaryotes. Glucose • Glycolysis is the most widespread metabolic pathway among Earth’s organisms suggests that it evolved very early in the history of life. 1. Cytoskeleton (a network of structural proteins that facilitate cell movement, morphological integrity and organelle transport). Glycolysis CYTOSOL Microtubule Pyruvate No O2 present: Fermentation Fig. 6-20 O2 present: Aerobic cellular respiration - is common to fermentation MITOCHONDRION (anaerobic) and cellular respiration (aerobic) Ethanol or lactate Acetyl CoA Citric acid cycle - puruvate represents a fork in the catabolic pathways of glucose oxidation Fig. 9-19 Bozeman Biology: Life Requires Free Energy (13:00 min.) http://www.bozemanscience.com/012-life-requires-free-energy 0.25 µm Microfilaments The cytoskeleton. In this TEM, prepared by a method known as deep-etching, the thicker, hollow microtubules and thinner, solid microfilaments are visible. A third component of the cytoskeleton, intermediate filaments, is not evident here. Copyright © 2012 Rebecca Rehder Wingerden Copyright © 2012 Rebecca Rehder Wingerden Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) 2. Membrane-bound organelles: energy transformation • chloroplasts: photosynthesis, convert solar energy to chemical • mitochondria: cellular respiration, generates ATP by extracting energy form sugars, fats, and other fuels with the help of oxygen Intermembrane space Outer membrane energy by absorbing light and using it to drive a synthesis of organic compound such as sugars from carbon dioxide and water Ribosomes Stroma Inner and outer membranes Free ribosomes in the mitochondrial matrix Granum Inner membrane 1 µm Thylakoid Fig. 6-18 • Mitochondria and chloroplast are semiautonomous organelles Cristae Matrix Fig. 6-17 0.1 µm Copyright © 2012 Rebecca Rehder Wingerden that: - grow and reproduce within the cell, and - contain a small amount of DNA that programs the synthesis of proteins made on the organelle’s ribosomes. Copyright © 2012 Rebecca Rehder Wingerden Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) 3. Linear chromosomes: • contained in the cell nucleus, • made up of chromatin, a complex or protein and DNA, and • each eukaryotic species has a characteristic number of chromosomes. Organisms share many conserved core processes and features that evolved and are widely distributed among organisms today. (1.B.1) 4. Endomembrane systems, including the nuclear envelope • synthesis of proteins and their transport into membranes and organelles or out of the cell, • metabolism, • movement of lipids, and • detoxification of poisons. Smooth ER Nuclear envelope Rough ER ER lumen Cisternae Transitional ER Ribosomes Transport vesicle Smooth ER Rough ER 200 nm Bozeman Biology: Essential Characteristics of Life (10:00 min.) http://www.bozemanscience.com/005-essential-characteristics-of-life Copyright © 2012 Rebecca Rehder Wingerden Copyright © 2012 Rebecca Rehder Wingerden Fig. 6-12
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