Cent re for
Ecology & Hydrology
N AT U RA L EN V I RO N M E NT RES EA RC H C O U N C I L
t
á
10 0 0 (T 4 - 2 0 0 1
PLAN TATT
Attributes of British and Irish Plants:
Status, Size, Life History,
Geography and Habitats
for use in connection with the
New atlas of the British and Irish fl ora
M . 0 . Hill, C. D. Preston & D . B . Roy
Biological Records Centre
NERC Centre for Ecology and Hydrology
Ce ntre tor Et c totry entl Hydrology
Bus h Es te:e
Penicuik
Midlottna t
6 326 OAS
2004
@ N ERC Copyright 2004
Print ed in 2004 by Raven Marketing Group, Cambridgeshire
ISBN I 870393 74 0
The Cent re for Ecology and Hydrology (CEH) is one of t he Cent res and Surveys of t he
Nat ural Enviro nment Research Council (NERC). Established in 1994, CEH is a mult idisciplinary, environmental research organisat ion wit h some 600 staff and well-equipped
laboratories and fi eld facilities at nine sites t hroughout t he Unit ed Kingdom. CEH's
mission is to "advance t he science of ecology, environmental microbiology and hydrology
t hrough high quality and internationally recognised research leading to bett er
understanding and quant ifi cat ion of the physical, chemical and biological processes relating
to land and freshwat er and living organisms wit hin t hese environments".
The Biological Records Cent re is operated by CEH, based at CEH Monks Wood. BRC is
joint ly funded by CEH and t he Joint Nature Conservation Committ ee (www.j ncc.gov.uk),
t he latter act ing on behalf of the statutory nat ure conservation agencies in England,
Scot land,Wales and Nort hern Ireland. CEH and JNCC support BRC as an import ant
component of t he developing National Biodiversity Network. BRC seeks to help
nat uralists and research biologists to co-ordinate t heir eff ort s in studying the occurrence
of plant s and animals in Britain and Ireland, and to make t he results of t hese st udies
available to ot hers.
For fur t her informat ion, visit www.ceh.ac.uk.
Cover photograph shows the endemic plant Coincya w right ii on the cliffs of Lundy (courtesy of
Roger Key, English Nature).
Published by
Cent re for Ecology & Hydrology
Monks W ood
A bbot s Ripton
Hunt ingdon
Cambridgeshire
PE28 2LS
Copies can be ordered by writing to t he above address or by telephoning
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CON TE N TS
Int roduct ion
Species, fam ilies, stat us and change
3
Species and families
.3
Native/afi en status
.3
Conservation status . .
.3
Rarity status
.3
Change Index .
4
Size and life history at tributes . . .
Height and length .
Perennation
.5
Life form . . . .
.5
Woodiness
Clonality
Geographic att ributes
European and wider distributions, and continentality
.
. 9
Native distribution of species alien in Britain and Ireland . .
. 10
Counts of 10-km squares in the New Atlas
. 10
Climatic means
11
H abitat att ributes . . . .
. 12
Coastal species
. 12
Broad Habitats
. 12
Ellenberg's indicator values
. 14
Light values (L)
. 14
Moisture values (F)
. 15
Reaction (R)
. 16
..
. 16
Sak tolerance (S) .
. 17
Nitrogen (N)
List ing of species with t heir att ribut es
18
A ppendix
61
Sources for plant height .
. 61
Defi nition of life forms
. 61
Examples of how fi fe-form criteria are applied .
. 62
Links between life forms and plant height
. 63
Clonality
. 64
Broad Habitats
. 64
Initial calculation of species' preferences for Broad Habitats
. 68
Extension and validation of species' preferences
. 69
A cknowledgem ent s
70
Bibliography
71
IN TR OD UCTI ON
In the course of our research on geographical distributions and climate change, we have
frequent ly wanted t o characterize the plants t hat const itute the w ild fl ora of Britain and Ireland.
Several sources of information were available, including t he Electronic Comparative Plant Ecology
(Hodgson et al., 1994), t he Czech clonal plant database Clopla I (http.//www.butbn.cas.ce kfi mes/
clopla I .htm), t he BSBI database (http://www.bsbi.org.uk/html/database.html), t he Ecoflora database
(Fitter & Peat, 1994), and the indicator values of Ellenberg (Ellenberg et S., 199 1). Inevitably, these
sources were not suffi cient for our purposes, and we have gradually assembled our own set of
att ribute data, linked to t he database of the Biological Records Centre (BRC).
The establishment of the National Biodiversity Network (N BN) provided an additional st imulus
to the assembling of att ribute data, off ering a suitable platfor m for online searches for species
att ributes. Some of the infor mation published here is also available fro m the N BN gateway
(www.searchnbn.net). In 2002, the publicat ion of the New Atlas of the British and Irish Flora (Preston,
Pearman & Dines, 2002), toget her with results of a companion project to analyse change
(Preston,Telfer et aL, 2002), allowed us to complete the dataset that is presented here.
The dataset is partly old and partly new (Table 1). Biogeographic elements and Ellenberg values
are taken with few modifi cat ions from earlier publications (Preston & Hill, 1997, 1999; Hill et
1999). Information that depends directly on t he New Atlas is either new or (Change Index) is
taken from the text of the New Atlas. Data on t he Broad Habitats of plants are mostly drawn
from an unpublished report (Preston et al., 2003). Infor mation on plant height, perennat ion, life
form and clonal growt h was assembled from a variety of fl oras and monographs, with some
personal observation; it is a new compilat ion. Furt her details of t hese att ributes are provided in
t he following pages. The attribute categories are tabulated (Tables 2- 13), with the numbe of taxa
in each category listed in t he column 'N ='.
Table I . A t t ributes, codes and names listed as colum ns in PLA N TAT T
Colum n nam e
A bbreviat ion
Source or ot her com ment
(a) Status and taxonomic
Name in curr ent flora (Stace, 1997)
Taxon name
Family
Fam
Stace ( 1997)
Nat ive status
NS
New Atlas (Prest on et al., 2002)
Conservat ion status
CS
Cheffi ngs (2004)
Rarity stat us
RS
Calculat ed from New Atlas data
Change Index
Chg
New Atias (Prest on et al., 2002)
Height (terrest rial)
Hght
Several sources, list ed below
Length (aquat ic)
Len
Several sources, list ed below
Perennat ion
PI , P2
Mainly Clapham,Tut in & Warburg ( 1962)
Life form
LF1, LF2
Clapham et al. ( 1962), heavily revised
(b) Size and life form
Mainly Clapham et al. ( 1962)
W oodiness
Clonel ,
Clone2
Sources are listed below; clone2 is mainly blank
Major Biome (European
dist ribut ion)
EI
Part of biogeographic element
accor ding t o Preston & Hill ( 1997)
East ern limit code
E2
Part of biogeographic element
according t o Preston & Hill ( 1997)
Clonal spread
(c) Geography & climate
Prest on & Hill ( 1997) plus additions
Cont inentality in Europe
O rigin of alien taxa
O rigin
New Atlas (Prest on et al., 2002)
Number of 10-km squares in
Britain (including Isle of Man)
GB
New Atlas (Prest on et al., 2002)
Number. of 10-km squares in
Ireland
IR
New Atlas (Prest on et al., 2002)
Number of 10-km squares in
Channel Islands
CI
New Atlas (Prest on et al., 2002)
January mean terhperature
Tjan
Calculat ed from New Atlas plus climat ic data
July mean t emperat ure
Tjul
Calculat ed from New Atlas plus climat ic data
Annual precipitat ion
Prec
Calculat ed from New Atlas plus climat ic data
Coastal
Co
Calculat ed from New Atlas plus Ellenberg salt value
Broad Habitats
Br Habitats
Preston et al. (2003), plus previously unreport ed data
for aliens
Ellenberg indicator value
L, F, R, N, S
Hill et aL ( 1999)
(d) Habitat
SPE CI E S, FA M ILIE S, S TA T US A ND CHA N GE
Species and families
The list of 1885 taxa includes all nat ive British and Irish species, as well as natives of the Channel
Islands not known from Britain or Ireland. There are no species restricted to the Isle of Man. All
archaeophyt e species are included, plus a number of well-established neophytes.A small number
of subspecies, hybrids and aggregate species are also included. The number of taxa has been
slightly increased from the 1791 t hat were listed for Ellenberg indicator values (Hill et al., 1999).
Most of t he addit ions are of neophytes, w hich, following the New Atlas (Preston et al., 2002), we
now t hink t o be suffi cient ly common o r potent ially interest ing to be included.
T he names of families have been abbreviated to four letters and follow Stace ( 1997).
N at ive/alien stat us
Species of uncert ain stat us are classified as 'nat ive or alien'. Int roduced species have been
classifi ed as archaeophytes, neophytes and casuals (Preston et al., 2002). Bot h archaeophytes and
neophytes are introduced species w hich are present in t he wild as naturalized populations, that is
t hey are spreading vegetat ively or reproducing eff ectively by seed. An archaeophyte is a plant
t hat became naturalized before AD 1500. A neophyte is one that was first int roduced after 1500,
or was only present as a casual before 1500 and is naturalized now only because it was
reint roduced subsequent ly. In contrast to archaeophytes and neophytes, a casual is a plant that is
present only as populat ions w hich fail to persist in t he w ild for periods of more t han
approximately fi ve years. Such a species is dependent on constant reint roduction (Macpherson et
at , 1996).
Hybrids between two alien parents which were introduced to the w ild as hybrids (e.g. Crocosmia
x crocosmiifl ora, Euphorbia x pseudovirgata) are classified as neophytes. O nly hybrids between
w hich have been formed spontaneously in the w ild between two alien parents are classified as
'AX'.
Conservat ion stat us
The conservat ion status of plant s is coded according to the system proposed by Hodgetts,
Palmer & W igginton ( 1996); see also Palmer et al. ( 1997). This status is an assessment of threat
rather t han rarity, alt hough closely related to rarity in that only species known at the time of t he
assessment in less t han 10 1 10-km squares have been allocated t o one of the categories listed.
The stat us list ed is t hat given by Cheffi ngs (2004). This replaces t he stat uses provided in the Red
Data Book (W igginton, 1999) and Scarce plants in Britain (Stewart , Pearman & Preston, 1994).
Cheffi ngs assigns a conservat ion status to nat ive species and archaeophytes, with only a few
exceptions. PLA NTATT cannot be used to provide a complete list of t hreatened taxa as many
subspecies listed by Cheffi ngs are not t reated separately by us.
Rarity stat us
Rarity status is based on counts of the number of 10-km squares in Britain and t he Isle of Man in
w hich the plant was recorded as a nat ive in t he period 1987- 1999. Rare plants are those
recorded in 1- 15 10-km squares during t his period; scarce plants are recorded in 16- 100.The
t hresholds were those that were used in defining rare and scarce species for the Red Data Book
and Scarce plants. Rarity status, however, is solely a measure of rarity rat her than threat, and is
based on data from t he New Atlas. Cheffi ngs assigns a conservat ion status to nat ive species and
archaeophyt es, w ith only a few except ions. PLA NTATT cannot be used to provide a complete
list of t hreat ened taxa as many subspecies listed by Cheffi ngs are not t reated separately by us.
A lien species have by defi nition no nat ive records, and therefore are not given a rarity status.
Table 2. N at ive stat us, Conservation stat us and Rar ity stat us
A t t ribute and codes
N = Explanation
(a) Native status
AC
12 Alien casual; many are crop plants
AN
259 Neophyte, alien int roduced after 1500
AR
15 1 A rchaeophyte, alien int roduced before 1500
AX
1 Spontaneous hybrid between two alien parent s
1362 Nat ive, not endemic
NA
46 Nat ive or alien (nat ive status doubtful)
NE
47 Nat ive endemic
NH
7 Spontaneous hybrid bet ween two nat ive parent s
(b) Conservation status
CR
DD
EN
EW
EX
VU
20 Cr itically endangered
I Data defi cient
34 Endangered
4 Ext inct in t he wild
10 Ext inct
119 Vulnerable
(c) Rarity status
933 Present, not rare or scarce
234 Rare ( 1- 15 10-km squares in Britain, 1987- 1999)
254 Scarce ( 16- 100 10-km squares in Britain, 1987- 1999)
27
Absent from Britain and Isle of Man as a nat ive, but nat ive in
Ireland or t he Channel Islands
12 Apparent ly ext inct (not recorded since 1986)
2 Insuffi cient data available to assess rarity
(blank)
423 Alien taxa
(d) Change Index
(values)
Change between 1930- 1960 and 1987- 1999
Change Index
The Change Index (Telfer et al., 2002) measures the relat ive magnitude of change, compar ing the
period 1930- 1960 with 1987- 1999. It is explained in more detail in t he Appendix. It ranges from
-4.78 for the critically endangered Gahum tricornutum to 4.70 for t he much more frequent ly
recorded Prunus laurocerasus. The Change Index has not been calculated for species not included
in the fi rst Atlas of the British Flora (Perr ing & Walters, 1962). Consequent ly, several aliens such as
Amsinckia micrantha and Crassula helmsii, w hich have increased spectacularly in the intervening
period, lack a value for the Change Index.
SIZ E A N D L I FE H IS TOR Y A TTRIB UTES
H eight and length
Heights, measured in cent imet res, are given for terrestrial (or emergent) plants and lengths (cm)
for submerged aquatic plants. A few amphibious or emergent plants such as Persicaria amphibia
are scored for both height and length.
These values are int ended as a general indication of the size of t he plant. For procumbent plants,
heights are an indicat ion of the height of the less procumbent shoots. For scapose plants and
ferns, heights are not stem lengths but leaf lengths. Likewise, lengths for isoetids (i.e. linear-leaved
ro sette-for ming root ed aquat ics) are in fact leaf lengths. For example Lobelia dortmanna has
length 4 cm, w hich is certainly not the length of t he scape.
Height and length values were drawn from a variety of sources, which are detailed in the
Appendix.
Pe rennat ion
The categories are annual, biennial and perennial. Biennial is used as a short hand to denote also
monocarpic perennials. For example Car/Ma vulgaris and Heracleum mantegazzianum normally
take several years to reach maturity but are categor ized as biennials. The sources of data for
perennation are most ly t he same as t hose used for height and length. In a few cases (e.g. Linum
catharticum), we have relied on personal communication and scientifi c papers rat her than fl oras.
A relat ively small number of species fall into more t han one of these categories. The secondary
category is intended to be either less frequent or equally frequent. Thus Poe annua has primary
perennation category PI =annual and secondary category P2 = perennial. Several fl oras suggest
t hat Cardamine flexuosa and Montia fontana can be annual, biennial or perennial. These have been
coded as PI = p, P2 = a and PI = a, P2 = p respectively.
Life form
T he Raunkiaer system of life forms was set out by Clapham,Tutin & Warburg ( 1962), who listed
life forms for most species. Their assignments provide an excellent start ing point but contain
numerous inconsistencies. Life forms have been fully revised for PLANTATT. Following German
and Sw iss aut hors (e.g. Lindacher, 1995), the category of helophytes or marsh plants has been
omitt ed. The major ity of t hese can grow in places that are not permanently fl ooded in winter.
T he position of the overwintering buds in such situations is taken as a guide to t heir life form.
A n additional life form of annual water plants has been added, so t hat t he categor y t herophytes
are all terrest rial.
As wit h perennat ion, a secondary life for m is given for some species. Details of how life forms
were assigned and the relation of life form to plant height are given in t he Appendix.
Woodiness
W oodiness is an attr ibut e that applies to some chamaephytes, nanophanerophytes and
phanerophytes. All ot her life forms are t reated as herbaceous. Three categories are recognized,
woody, semi-woody and herbaceous. Although we have been guided in part by Clapham,Tutin &
W arburg ( 1962), we have not hesitated to express our own opinions where these diff er.
Table 3. Size and life hist ory at t ribut es ot her t han clonality; count s of primar y
att ributes are in t he column N 1=, co unt s of secondary at tribut es are in N 2=
A t t r ibut e or code
N 1=
(a) Height (cm)
Hght
(b) Length (cm)
Len
(c) Perennation
PI
a
455
b
96
p
1334
(d) Life form
LFl
N 2= Ex planat io n
Height (leaf lengt h for scapose plants and ferns)
Length (aquat ic plant s; leaf length for isoet ids)
P2 Primary and secondary type of perennation
14 Annual
14 Biennial, including monocarpic perennials
42 Perennial
LF2 Primary and secondary fi fe form (see Table Al )
Ch
148
Gb
32
Gn
134
11 Non-bulbous geophyte (rhizome, corm or t uber)
hc
8 11
50 Hemicryptophyte
Hy
124
34 Perennial hydrophyt e (perennial water plant )
Hz
25
Ph
108
Pn
73
Th
430
24 Chamaephyte
Bulbous geophyte
1 Annual hydrophyte (aquat ic therophyte)
15 Mega-, meso- and microphanerophyte
2 1 Nanophanerophyte
14 Therophyte (annual land plant )
(e) Woodiness
sw
Vt
1651
Herbaceous
27
Semi-woody
207
Woody
Clonality
Clonal growt h is defi ned here as vegetative reproduct ion combined with lateral spread. Like
perennat ion, clonality may have more t han one value for a given species, refl ecting the various
ways in which clonal growth may occur. In some species t here are varieties with clonal growt h
(Arrhenatherum elatius van bulbosum, Caltha palustris var. radicans) while the normal for m is not
markedly clonal. In these two species, the variety with clonal growth is less frequent than the
type variety, so that both plants are given Clone] = 0 (not spreading clonally) as the primary
state, w hereas Clone2 = D Rg and St o12 respectively. Two species with proliferous
infl orescences really ought to have been assigned to three categories, namely Juncus bulbosus
(N ode2, Irreg and omitted D Ri) and Butomus umbellatus (Rhiz2, D Rg and omitted D Ri). It did
not seem wort h having a third column just for these two, so the inflorescence character has
been omitt ed.
Table 4. C at egor ies of clonality; count s of primar y at t ributes are in t he colum n
N 1=, count s of secondary at t ributes are in N 2=
A t t ribute or code
(I) Clonafi ty
N 1=
Gone
0
1228
N 2= Explan at io n
Clone2
Litt le or no vegetative spread
Ogr
25
Tussock-forming graminoid, may slowly spread
Ot b
26
Tuberous or bulbous, slowing cloning by off set s
D Ra
20
10 Detaching ramet s above ground (oft en axillary)
D Rg
4
D Ri
16 Detaching ramet s at or below ground
11 Detaching ramet s on infl orescence
D R1
1 Detaching rarnets on leaves (Hammarbya)
D Rp
1 Detaching ramet s on prot hallus (Trichomanes)
Frag
7
Ir reg
38
Leaf
Fragmenting as part of normal growth
17 Irregularly fragment ing (mainly wat er plants)
1 Plant let s for med on leaves (Cardamine pratensis)
N ode l
19
N ode2
76
Rhiz l
152
Rhiz2
209
Root
37
St ol l
14
St o12
26
5 Far-creeping by st olons in illuminated medium
T ip
5
Tip root ing (the stems oft en t urn downwards)
20 Short ly creeping and root ing at nodes
9 Ext ensively creeping and rooting at nodes
8 Rhizome short ly creeping
15 Rhizome far-creeping
3 Clones formed by suckering from root s
Short ly creeping, stolons in illuminat ed medium
The following comments may be helpful in distinguishing the clonal categories.
I. T he category Ogr is rather poor ly defi ned. If a perennial graminoid (grass, sedge or rush) is
repeatedly mown or heavily grazed, it may gradually form clones, even though this is not the
normal condition.
2.
T he category Ot b assumes that the ground is litt le disturbed. If there is regular disturbance,
t he off sets w ill be dispersed to new sites and t he plant would then be indicated as D Rg
(detaching ramets at or below ground).
3.
T he small category Frag applies to duckweeds and Azola
4.
T he categor ies N ode (creeping and root ing at nodes), Rhiz (rhizomatous or with
subterranean stolons) and St ol (stoloniferous in the illuminated medium, eit her air or
water) are divided into two, according to whether they spread a short distance in a season
or are far-creeping. For rhizomatous plants, a rule of t humb is that a species whose new
shoots arise at a distance less t han a quart er of t he height of the plant is short ly creeping,
w hile those that spread further are deemed to be far-creeping. Analogous distinct ions are
made for N ode and Stol.
5.
The distinction between N ode and St ol is that in N ode the horizontal or decumbent
stem extends with indeterminate growth, rooting at the leafy nodes and not forming a new
stem axis at each rooted position. In St ol, a new stem axis normally arises at the point of
root ing, or the stolon is determinate, bending upwards and not progressing wit h
indeterminat e growt h.
6.
In the t ip-root ing categor y, T ip, leafy stems that are not obviously stolons turn down at the
apex and root there.
GE OGR A PH I C A TTRIB UTES
Euro ean and wider distribut ions and cont inent alit
The categorizat ion of taxa to biogeographic elements and their designat ion as 'continental' or
ot herwise (Table 5) follow Preston & Hill ( 1997), with minor revisions and t he addit ion of the
hyperoceanic category from a subsequent paper (Preston & Hill, 1999). Element s are provided
for nat ive species and some archaeophytes; for archaeophyt es the classifi cation describes the
archaeophyt ic range. In addition, some but not all alien species originat ing in Europe have also
been assigned to biogeographic elements for PLA NTATT and designated as continental.
Geographic att ribut es are not provided for some hybrids.
Table 5. European and wide r dist ribut ion, and continentality
A tt r ibut e
and codes
N = Ex planat io n
N ative dist ribution o s ecies alien in Britain and Ireland
The native dist ribution of aliens (Table 6) is taken from the New Atlas. This information is
provided for neophytes and some archaeophytes. In Table 6, taxa present in t wo areas are
included in bot h totals but taxa are excluded from the totals if they are only doubtfully native in
the relevant area.
Table 6. N at ive dist ribut ion of species alien in Brit ain and Ireland
Code
N=
Am
24
N ort h America
Am4
22
W est ern Nort h America
Am6
5
Eastern Nort h America
As
10
Asia east of 60°E
As I
24
Asia bet ween 60°E and I 20°E
As2
9
Asia E of I 20°E
Aus
3
Aust ralia
Crop
15
Eur
149
Gard
13
N Hem
6
N Hemisphere (Europe,Asia and Nort h America)
NZ
4
N ew Zealand
SAf
5
Southern Afr ica
SAm
20
South America and/or Cent ral America
Unk
10
Unknown
Ex planat ion
Crop plant , does not have a nat ive range
Europe
Garden or igin, does not have a nat ive range
Counts o I O-km s uares in the New Atlas
For each taxon, the number of 10-km squares in Britain, Ireland and the Channel Islands is.
enumerated (Table 7). For most natives, only t he native distribution has been counted (blue dots
in the New Atlas). For the small number of nat ive species in w hich it proved impossible even to
attempt to delimit the nat ive range in the New Atlas, the count is for all squares. For alien taxa,
all squares with records have been counted. Squares have been counted without regard to date,
so that Otanthus maritimus, ext inct in Britain, is recorded from 23 squares t here.
10
Table 7. Counts of squares and climat ic m eans
Attribute
Min
(a) Counts
Max Explanat ion
Counts of 10-km squares
GB
0
IR
0
CI
0
(b) Climatic means
2805 Great Britain and Isle of Man
985 Ireland
14 Channel Islands
Mean values for 10-km squares
Tian
- 1.3
Tjul
10.4
Prec
553
7.0 January mean t emperat ure (°C)
17.0 July mean t emperature (°C)
32 18 Annual precipitation (mm)
Climatic means
Climat ic values for plants were calculated as t he mean climate of t he 10-km squares where they
occur in Britain, Ireland and the Channel Islands, averaging over t he squares enumerated for the
counts. Climate data for 10-km squares were taken from baseline climate summaries of t he UK
Climate Impacts Programme (HuIme & Jenkins, 1998). These baseline summaries were
const ructed by interpolat ion of daily weat her measurements from individual met stat ions,
averaged over the 30-year period 1961- 1990 (Barrow et al., 1993).
HA BI TA T A TTRIB UTES
Coastal species
Species are deemed to be coastal if 80% of occupied squares are on the coast and if they depend
on coastal habitat. Most coastal species have Ellenberg salt values (S values, defi ned below)
greater than 0. Those with Ellenberg S = 0 were scrutinized carefully to ascer tain whether their
habitat was indeed coastal.
Broad Habitats
The preferences of species for Broad Habitats of the UK BAP are listed in the column labelled Br
Habitats. Species' main habitat (s) are_list ed. No species is deemed to have a preference for
more t han four habitats. Minor habitats are ignored. For more infor mat ion about Broad
Habitats and species' preferences, refer to the Appendix.
Table 8. C oast al species and preferences for Broad H abit ats
A tt r ibut e
and codes
N=
Explanat io n
(a) Co
Co
Coastal species
145
(b) Br Nabs
At least 80% of occupied squares contain sea at high t ide
Broad Habitats
3 10
Broadleaved, mixed and yew woodland
2
32
3
543
Boundary and linear feat ures (eg hedges, roadsides, walls)
4
198
A rable and hor t icult ural (includes orchards, excludes domest ic gardens)
5
22
6
163
Neut ral grassland (includes coarse Arrhenatherum grassland)
7
2 18
Calcareous grassland (includes lowland and montane types)
8
89
Acid grassland (includes non-calcareous sandy grassland)
9
10
Bracken
10
83
Dw arf shrub heat h (cover of dwarf shrubs at least 25%)
II
254
12
41
13
174
St anding wat er and canals
14
149
Rivers and st reams
IS
103
Montane habit at s (acid grassland and heat h with montane species)
16
292
Inland rock (het erogeneous - includes quarries, limest one pavement , cliff s, screes
and skeletal soils over rock)
17
23 1
Built-up areas and gardens
18
74
Supralitt oral rock (does not include marit ime grassland)
19
135
Supralitt oral sediment (st randlines, shingle, coastal dunes)
21
65
Litt oral sediment (includes saltmarsh and salt marsh pools)
23
1
Coniferous woodland
Improved grassland
Fen, marsh and swamp (not wooded; includes flushes, rush-pastures, springs and
mud communit ies)
Bog (on deep peat ; includes bog pools as well as acid lowland valley mires on
slight ly shallower peat )
Inshore sublitt oral sediment (only Zostera marina)
Ellenbe s indicator values
Ellenberg defi ned seven major scales, of which if ve are presented here. The two that are
omitted, T (temperat ure) and K (cont inentality), correspond quite closely to the major biome
and east ern limit categor ies defi ned for European dist ribut ions by Preston & Hill ( 1997). N either
T nor K values are satisfactory in an oceanic climate such as that of Britain; t hose for K are
par ticularly unreliable, especially as Ellenberg's defi nition was geographical rather than climat ic.
The fi ve remaining scales have values defi ned in the tables that follow. The values are based on
those of Ellenberg et at ( 1991). They are mostly reproduced fro m a previous publication (Hill et
al., 1999), w ith some additions because of the ext ra species included here. A few example
species are given for each value, by way of explanation.
Light values (L)
The full range of Ellenberg values for light (Table 9) is not represent ed in the British fl ora. For
canopy t rees, light values refer to the tolerance of t he sapling stage of t he life cycle.
Table 9. Ellenberg values for light (L)
Code
N=
Ex planat io n
0
Plant in deep shade (no examples for Britain or Ireland)
2
4
Between 1 and 3 (Epipogium aphyllum, Neottia nidus-avis, Trichomenes speciosum)
3
20
Shade plant, most ly less t han 5% relat ive illuminat ion, seldom more t han 30%
illuminat ion when t rees are in full leaf (Gahurn odoraturn, Listera cordate, Mercurialis
perennis)
4
70
Between 3 and 5 (Circaea lutetiono, Lamiastrum galeobdolum, Poa nérnoralis)
5
120
Semi-shade plant , rarely in full light , but generally w ith more than 10% relat ive
illuminat ion when t rees are in leaf (Carex pendula, Hyacinthoides non-scripta, Primula
vulgaris)
6
2 13
Bet ween 5 and 7 (Anthriscus sylvestris, Digitalis purpurea, Teucriurn scorodonia)
7
680
Plant generally in well lit places, but also occurring in part ial shade (Arrhenatherurn
elatius, Carex flacco, Poo trivialis, Vicia cracca)
8
576
Light-loving plant rarely found where relat ive illuminat ion in summer is less t han 40%
(Cardarnine hirsute, Orchis morio,Thymus polytrichus,Vacciniurn oxycoccus)
9
202
Plant in full light , found most ly in full sun (Aster tripolium, Mel/lotus albus, Poa compressa,
Primula farinosa)
14
M oisture values (F)
Unlike the ot her Ellenberg values, moisture is on a scale of I to 12 (Table 10). We use the
abbreviation F from t he German Feuchtigkeit.
Table 10. Ellenberg values for moist ure ( F)
C ode
N=
Explanat ion
Indicat or of ext reme dryness, rest ricted to soils t hat oft en dry out for some t ime
(Corynephorus canescens, Helianthemum apenninum, Koeleria vallesiana)
1
3
2
28
3
168
Dry-sit e indicat or, more oft en found on dry ground than in moist places (Asplenium
trichomanes, Centaurea scabiosa, Spergularia rubra)
4
378
Between 3 and 5 (Arctium minus, Hefi ctotrichon pratense, Iris foetidissima, Thymus
polytrichus)
5
492
Moist-sit e indicator, mainly on fresh soils of average dampness (Anthriscus sylvestris,
Euphorbia amygdaloides, Hyacinthoides non-scripta, Solarium nigrum)
6
226
Bet ween 5 and 6 (Agrostis stolonifera, Empetrum nigrum, Rumex crispus)
7
14 1
Dampness indicator, mainly on const ant ly moist or damp, but not on wet soils (Carex
Dactylorhiza maculata, Pulicaria dysenterica, Ranunculus repens)
8
170
Between 7 and 9 (Cardamine pratensis, Equisetum telmateia, Phalaris arundinacea, Schoenus
nigricans)
9
126
W et-sit e indicator, oft en on wat er-sat urat ed, badly aerated soils (Drosera rotundifofi a,
Myosotis scorpioides,Vaccinium oxK occus, Viola palustris)
10
64
Indicat or of shallow-water sites t hat may lack standing water for ext ensive periods
(Afi sma plantago-aquatica, Carex limosa, Ranunculus fi ngua, Typha latifofi a)
II
34
Plant root ing under wat er, but at least for a t ime exposed above, or plant fl oat ing on
t he surface (Lemna minor, Nuphar lutea, Sagitt aria sagittifolia, Schoenoplectus lacustris)
12
55
Submerged plant, permanent ly or almost constant ly under wat er (Isoetes lacustris,
Potamogeton crispus, Ranunculus circinatus, Zostera marina)
Bet ween 1 and 3 (Clinopodium acinos, Saxifraga tridactyfi tes, Sedum acre)
15
React ion (R)
Reaction (Table I I) refers to environmental acidity, which would ordinarily be measured by pH.
Except for water plants, R values refl ect preferences for soil acidity.
Table I I . Ellenberg values for react ion (R)
Code
Ex planat io n
N=
Indicator of ext reme acidity, never found on weakly acid or basic soils (Andromeda
polifolia, Lycopodium clavatum, Rubus chamaemorus, Ulex minor)
Between I and 3 (Agrostis curtisii, Calluna vulgaris, Drosera rotundifofi a, Polygala serpyllifolia)
Acidity indicat or, mainly on acid soils, but except ionally also on nearly neut ral ones
(Agrostis vinealis, Dactylorhiza M a de r a , Galium saxatile, Pteridium aquilinum)
Between 3 and 5 (Agrostis capillaris, Carex panicea,Juncus eff usus,Teucrium scorodonia)
Indicator of moderat ely acid soils, only occasionally found on very acid or on neut ral t o
basic soils (Cardamine pratensis, Cirsium palustre, Rubus idaeus, Ulex eumpaeus)
Bet ween 5 and 7 (Ammophila arenaria, Carex sylvatica, Lofi um perenne, Ranunculus ficaria)
Indicat or of weakly acid t o weakly basic condit ions; never found on very acid soils
(Agrimonia eupatoria,Atriplex prostrata, Nuphar lutea, Phleum pretense)
Between 7 and 9 (Artemisia vulgaris, Carduus nutans, Iris foetidissima, Viola hirsuta)
Indicat or of basic react ion, always found on calcareous or ot her high-pH soils (Bunium
bulbocastanum, Clinopodium calamintha, Dryopteris submontana, Primula farMosa)
Nitrogen (N)
Nitrogen values (Table 12) are in fact a general indication of preference for soil fert ility. They are
closely correlated with t he st ress values of Grime ( 1979, 200 1), low N values corresponding to
plants with high st ress tolerance (Grime et al., 1997) and vice-versa.
Table 12. Ellenberg values for nitrogen (N)
Ex planat io n
Cod e
N=
1
84
2
323
Between 1 and 3 (Aira praecox, Carex panicea, Linum catharticum, Scabiosa columbaria)
3
286
Indicat or of more or less infert ile sites (Centaured scabiosa, Golium saxatile, Pimpinella
saxifraga,Teucrium scorodonia)
4
245
Between 3 and 5 (Agrostis capillaris, Cirsium palustre, Plantago lanceolata, Primula vulgaris)
5
342
Indicat or of sites of int ermediat e fert ility (Angelica sylvestris, Digitalis purpurea, Iris
foetidissima, Trifolium pratense)
6
3 19
Between 5 and 7 (Cirsium arvense, Glyceria fluitans, Poa trivialis, Rumex crisp s)
7
223
Plant oft en found in richly fertile places (Atriplex prostrate, Epilobium hirsutum, Stellaria
media,Typha latifolia)
8
55
9
8
Indicat or of ext remely infert ile sites (Agrostis curtisii, Cli nopodium acinos, Drosera
rotundifolia, Rubus chamaemorus)
Between 7 and 9 (Beta vulgaris, Galium aparine, Lamium album, Urtica dioica)
Indicator of ext remely rich situat ions, such as cat t le rest ing places or near pollut ed
rivers (Arctium laPpa,Artemisia absinthium, Hyoscyamus niger, Rumex obtusifolius)
Salt tolerance (5)
Values for salt tolerance (Table 13) start at zero, corresponding to no tolerance of salt.
Table 13. Ellenberg values for salt t olerance (S)
Explanat ion
Absent from saline sites; if in coastal situat ions, only accidental and non-persist ent if
subjected to saline spray o r wat er (85% of t he fl ora)
Slightly salt-tolerant species, rare to occasional on saline soils but capable of persist ing
in t he present of salt (includes dune and dune-slack species where t he ground wat er is
fresh but w here some input s of salt spray are likely) (Calystegia sepium, Chenopodium
album, Oenanthe crocata, Sedum angfi cum)
Species occurring in both saline and non-saline situat ions, for which saline habitat s are
not st ro ngly predominant (Atriplex prostrata, Elytrigia repens, Phragmites australis, Rumex
crispus)
Species most common in coastal sites but regularly present in freshwat er or on nonsaline soils inland (includes st rictly coastal species occurring in sites such as cliff crevices
and sand dunes t hat are not obviously salt-aff ect ed) (Cakile maritima, Cochlearia officinalis,
Juncus gerardii, Spergularia rupicola)
Species of salt meadows and upper saltmarsh, subject t o at most only very occasional
t idal inundat ion (includes species of brackish condit ions, i.e. of consist ent but low
salinity) (Atriplex littoralis, Elytrigia atherica, Glaux maritima,Triglochin maritimum)
Species of the upper edge of saltmarsh, where not inundat ed by all t ides (includes
obligat e halophytes of cliff s receiving regular salt spray) (Aster tripofi um, Crithmum
maritimum, Puccinellia maritima, Suaeda vera)
Species of mid-level saltmarsh (Atriplex portulacoides, Cochlearia anglica, Limonium vulgare)
Species of lower saltmarsh (Spartina anglica, Suaeda maritima)
Species more or less permanent ly inundat ed in sea wat er (Zostera spp.)
Species of extremely saline condit ions, in sites where sea water evaporat es,
precipit at ing salt (Safi cornia europaea agg.; t hese could equally well be t reat ed as
species of t he lower marsh)
L IS TIN G OF SPE CIES WI TH THEIR A T TRI B UTES
The listing of species with their attributes has 32 columns. Codes used in these columns are set
out in the foregoing tables. The short and full names of the columns are given here (Table 14), as
are the pages on which the tables of codes appear
Table 14. Plant at t ributes cross-referenced t o ta bles and page numbers where t hese
are described more fully
Table N o
Page
Short nam e
A t tr ibute
Taxon name
Name in N ew At las
1
2
Fam
Family (4-let ter abbreviation)
1
2
NS
Nat ive status
2
4
CS
Conservat ion status
2
4
RS
Rarit y stat us
2
4
Chg
Change Index
2
Hght
Height in cm (terrest rial)
3
6
Len
Length in cm (aquat ic)
3
6
PI
Perennat ion - primary
3
6
P2
Perennat ion - secondary
3
6
LF 1
Life for m - primary
3
6
LF2
Life for m - secondary
•
Woodiness
3
6
3
6
Clone 1
Clonal spread .- primary
4
7
Clone 2
Clonal spread - secondary
4
7
EI
Major Biome (European dist ribut ion)
5
9
E2
East ern limit code
5
9
C'
Continentality in Europe
5
9
O rigin
Origin of alien taxa
6
10
GB
Number of 10-km squares in Britain (including Isle of Man)
7
1I
IR
Number of I 0-km squares in Ireland
7
II
CI
Number of 10-km squares in Channel Islands
7
II
Tjan
January mean t emperat ure
7
II
Tjul
July mean t emperat ure
7
II
Prec
Annual precipitat ion
7
Co
Coastal
8
13
Br Habitats
Broad Habitats
8
13
Ellenberg indicator value - light
9
14
Ellenberg indicat or value - moist ure
10
15
Ellenli erg indicat or value - reaction
II
16
Ellenberg indicator value - nit rogen
12
16
Ellenberg indicator value - salt t olerance
13
17
ft
18
T a xo n n a m e
A cae na no vae - se lan dia e
Ac er cam p est re
A c e r p lata noi de s
A ce r p seu dop latanus
A ce ras an thm pop hor um
A chi llea mille folium
A ch illea p tarm ica
A con itu m nap ell us
A con itur n nap ellus se ns . la t.
A cor us cal am us
A cta ea spi ca ta
A dia ntum cap ill us - ven eris
A do nis an nua
Ad ox a m o s eh a te ll in a
A egop odi um
da graria
A es cul us hip po ca sta nu m
A ethu sa cynap ium
A grimo nia eup atoria
A grimo nia proc ere
A gros tem ma githa go
A gros tis can ine
A gros tis can ine sen s .lat .
A gros tis cap illa ris
A gro stis cur tisii
A gros tis giga ntea
A gros tis stoloni fera
A grost is vi n ea li s
Aire ca ryop hy llea
Aire p raeco x
Aj ug a cha maep itys
Aj ug a py ramid alis
Aj ug a rept an s
A lch em illa ac utilo ba
Alch em illa alp ina
Alch em illa filica ulis
Alch em illa gla bra
Alch em illa gla uce scen s
A lch em illa glo merula ns
A lch em illa mica ns
.
A lch em illa min ima
A lch em illa mo llis
A lch em illa mo ntico la
A lch em illa sub crena ta
A lch em illa vul garis agg .
A lch em ill a wichu rae
A lch em ill a xa nth oc hlo ra
Fa m
Rosa
Acer
Acer
A cer
O rc h
A st e
A st e
Ranu
Ranu
A ra c
Ranu
A d ia
Ranu
Adox
A p ia
H ip p
A p ia
Ro s a
Ro s a
Cary
Po a c
Poac
Poet
Poac
Poac
Poac
Poac
P oa c
Po a c
La m i
La rn i
La m i
Ro s a
Ro s a
Ro s a
Ro s a
Ro s a
Ro s a
Ro s a
Ro sa
R o sa
R o sa
R o sa
R o sa
R o sa
R o sa
NS
AN
N
AN
AN
N
N
N
NA
NA
AN
N
N
AR
N
AR
AN
NA
N
N
AR
N
N
N
N
AR
N
N
N
N
NA
N
N
N
N
N
N
N
N
N
NE
AN
N
N
N
N
N
EN
VU
VU
VU
EW
VU
CS
r
r
n
s
n
n
n
n
n
r
5
n
r
n
n
n
s
s
r
r
n
n
n
n
n
n
n
n
s
5
s
n
n
5
s
n
RS
-0 .0 1
-0 .6 1
-0 .5 2
- 0 . 19
-0 .6 2
-0 .3 4
-0 .5 6
1.3 2
1.2 8
-0 .26
1 .3 9
3 .66
1.4 2
0 .6 9
- 0 .4 4
0 .5 4
- 2 . 19
-0 .0 5
-0 .4 5
1.0 8
-0 .4 1
-0 .8 9
-0 .3 8
-0 .7 5
-0 .4 0
-0 .76
0 .2 9
-0 .6 5
0 .3 5
C hg
H g ht L e n
11
15 00
3 0 00
3 0 00
40
45
60
10 0
10 0
1 12
60
30
40
12
10 0
32 0 0
10 0
60
10 0
10 0
60
60
62
60
80
45
60
25
10
20
30
30
60
15
30
40
15
30
40
5
60
35
35
35
20
40 .
1 P2 L F 1 L F2 W
Ch
Ph
Ph
Ph
Gn
Ch
h
hc
h
Gn
h
Gn
h
Hy
h
Gn
h
hc
Th
Gn
h
hc
h
Ph
Th
hc
hc
Th
hc
hc
hc
he
hc
hc
he
Th
Th
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
to 12
iz 2
t o 12
iz 2
t o 12
iz2
iz 2
h iz l
h iz 2
h iz 2
h iz l
h iz t
h iz l
h iz 2
h iz l
lo ne l
ode2
St o 12
5
5
5
4
7
3
3
3
3
3
6
6
4
1
5
6
3
3
2
3
3
3
3
3
3
3
3
3
3
1
3
4
3
7
8
7
5
5
5
8
8
6
7
8
8
8
4
7
5
1
4
5
5
2
5
4
6
1
1
6
4
3
3
3
1
5
5
3
3
4
9
9
5
7
7
7
7
9
5
5
7
7
O r i in
A us , N Z
c
c
c
c
c
Eur
Unk
Eur, A s 1
Eur
U nk
c Eur
c Eur
C lo ne 2 E l E 2 C
GB
82
13 8 9
14 19
2 59 9
10 9
2 77 4
2 38 2
10 4
10 4
5 15
33
38
2 34
17 2 0
2533
2 186
164 0
18 5 9
8 19
8 15
16 0 9
19 37
2 7 58
207
16 13
2 77 6
1 12 6
2046
2450
43
10 6
2439
15
3 84
14 0 7
12 7 1
20
57
4
3
805
9
2
19 3 5
65
10 6 0
9
0
43
942
0
968
492
0
0
17
0
29
3
2
8 19
557
0
532
16 1
96
36 1
44 3
9 22
0
10 9
977
15 0
6 56
6 59
0
8
7 17
0
4
308
223
3
0
0
0
17
0
0
4 96
0
24 0
IR
1
1
1
1
1
I
1
1
CI
r an r ul
3 .8 15 .2
3 .6 15 .7
3 .4 15 .2
3 .6 14 .6
3 .6 16 .3
3 .5 14 .5
3 .3 14 .3
4 .0 15 .8
4 .0 15 .8
3 .6 15 .8
2 .5 14 .3
5 .2 15 .2
3 .8 16 .0
3 .3 14 .9
3 .6 14 .6
3 .6 14 .8
3 .6 15 .5
3 .8 15 . 1
3 .9 15 .1
3 .9 15 .6
3 .5 14 .5
3 .4 14 .3
3 .5 14 .5
4 .9 15 .8
3 .6 15 .3
3 .6 14 .5
3 .3 14 .0
3 .7 14 .6
3 .5 14 .3
3 .8 16 .5
3 .3 12 .6
3 .4 14 .6
1 .6 13 .2
1 .6 12 . 1
2 .9 14 . 1
2 .6 13 .4
2 .0 13 .3
0 .4 1 1.5
2 .0 13 .5
1.7 13 .5
3 .5 15 .0
1.3 13 .0
0 .9 12 .6
3 . 1 14 .1
0 .9 12 .0
2 .8 13 .9
P re c
Co
83 1
839
903
10 8 3
690
1 10 3
1 14 6
964
964
7 70
10 54
1 10 1
745
10 0 5
10 6 4
10 14
863
953
10 2 6
852
1 12 8
1 14 5
1 10 4
10 8 2
889
1 10 1
12 0 0
10 8 7
1 14 9
7 13
14 2 5
10 9 5
969
18 4 3
1 18 5
13 3 8
14 0 8
19 7 8
10 56
16 6 7
10 09
10 84
1 12 5
1 17 9
19 9 2
1 17 4
B r H a b it at s
3 , 10 , 18 , 19
1, 3
1 , 3 , 17
1 , 3 , 17
7
6
11
1, 3 . 14 , 17
1, 3 , 14 , 17
13 , 14
16
16
4
1
3 , 17
3 , 17
3 , 4 , 17
6
1, 3 , 6
4
1 1, 13
8 , 11
8
8 , 10
3, 4
4, 6
8 , 10
10 , 16
8 , 16
4, 7
10 , 16
1
6
7 , 15 , 16
7 , 15 . 16
6 , 15 . 16
7
8 , 15
7
7
3 , 17
6
6
6 , 7 , 15 , 16
7
6, 7
L F R N S
8 3 6 3
5 5 7 6
4 5 7 7'
4 5 6 6
71 4 8 3
7 5 6 4
7 7 5 3
5 7 7 6
5 7 7 6
8 10 7 7 1
3 5 8 6
4 7 8 3
7 4 7 4
4 5 6 5
6 5 6
71
5 5 7 7
6 4
7 6
7 4 7 4
5 6 7 5
7 5 6 5
7 7 3 3
7 6 3 3
6 5 4 4
7 6 2
1
7 6 6 71
7 6 7 6
7 6 3 2
8 2 5 2
8 2 4 2
8 2
71 4
7' 5 5 2
5 7 5 5
7 4 6 5
7 5 4 3
8 6
6 3
7 6
6 4
T 5 7 5
7 5 5 4
7 5 7 5
8 3
71 6
6 5 7 6
7 4 6 4
7 4 6 5
T 5 6 4
71 5 5 3
6 5 6 4
Tax on na me
Fam
NS
CS
RS Ch g. Hght
Le n P 1 P2 LF 1 LF2 W
Clona l
Clo ne2 E l E2
Taxon name
Anthoxanthum aristatum
Anthoxanthum odoratum
Anthriscus caucalis
Anthriscus sylvestris
Anthylris vulneraria
Anfi rrhinum maj us
Apera interrupta
Apera spica-venti
Aphanes arvensis
Aphanes arvensis agg.
Aphanes australis
Apium grayeolens
Apium inundatum
Apium nodiflorum
Apium repens
Aquilegia vulgaris
Arabidopsis thaliana
Arabis alpina
Arabis glabra
Arabis hirsuta
Arabis petraea
Arabis scabra
Arbutus unedo
Arctium lappa
Arctium minus
Arctostaphybos alpinus
Arctostaphylos uva-ursi
Arenaria ciliata
Arenaria norvegica
Arenaria serpyllifolia
Armeria arenaria
Armada maritima
Armoracia rusticana
Arnoseris minima
Arrhenatherurn elatius
Artemisia absinthium
Art emisia campestris
Artemisia norvegica
Artemisia vulgaris
Arum italicum
Arum maculatum
Asparagus off icinalis
Asparagus off icinalis
subsp.officinalis
Asparagus offi cinalis
subsp.prostratus
AR
N
Lili
Uh
VU
VU
EN
VU
CR
CS
N
N
N
N
N
N
N
AR
A R EX
N
AR
N EN
N VU
AR
N
N
N
N
NS
AN
N
N
N
N
AN
AN
AR
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
AR
Fam
Poac
Poac
Apia
Apia
Faba
Scro
Poac
Poac
Rosa
Rosa
Rosa
Apia
Apia
Apia
Apia
Ranu
Bras
Bras
Bras
Bras
Bras
Bras
Eric
Aste
Aste
Eric
Eric
Cary
Cary
Cary
Plum
Plum
Bras
Aste
Poac
Aste
Aste
Aste
Aste
Arac
Arae
Lili
r
30
150
RS Chg Hght Len
-2.65
40
n 0.90
50
n -0.16
70
n -0.19
100
n 0.45
60
2.84
30
0.80
40
-0.2 1
100
n
10
n -0.32
10
n
10
n -0.63
80
n -0.54
30 so
n -0.3 1
60 100
r
15
n
1.70
100
n
1.21
30
r
15
s -1.16
100
n -1.02
40
s -0.64
25
r
20
o
1.18 500
0.51
150
n -0.4 1
150
s -0.22
20
n -0.75
20
o
6
r
0.21
6
n -0.76
30
o
20
n -0. 14
15
0.05
150
-3.72
30
n 0.37
150
-0.46
90
r -0,42
60
r
8
-0.20
150
s
2.09
60
n -0.28
50
n
1.78
150
Gn
Gn
1 P2 LF1 LF2
Th
hc
Th
hc
he
Ch
Th
Th
Th
Th
Th
hc
Hy
Hy
hc
hc
Th
Ch
hc
p he
Ch
Ch
Ph
hc
hc
Ch
Ch
Ch
a Ch Th
Th
Ch
Ch
hc
Th
hc
Ch
Ch
hc
hc
Gn
Gn
Gn
0
h 0
7
7
1
4
W Clone l Clone2 El E2 C
Origin
0
Eur
0
6 4
0
7 3
0
5 5
0
5 3
0
Eur
0
8 4
Eur, As l
0
5 4
0
7 3
0
7 3
0
7 3
0
8 4
Irreg
7 2
Irreg
8 4
Node2
7 3
0
7 3
0
7 4
0
1 4
7 4
0
0
5 6
0
1 5
0
9 3
w O
9 1
h 0
7 4
h 0
7 5
w Node2
1 6
w Node2
4 6
0
1 3
0
1 3
0
8 4
0
8 2
0
3 6
Rhiz2
Crop
0
7 3
0
DRg
7 3
sw 0
7 4
sw Rhizl
7 4 c
h 0
1 3
h 0
7 4
h Rhizl
9 1
h Rhizl
7 3
h 0
7 1
171
5871
GB
93
2782
659
2480
1798
1043
104
326
1608
2302
1549
519
9271
1661
3
1504
2218
1
151
1042
78
1
0
971
2424
134
473
0
16
2 147
0
1137
1532
83
2678
1005
9
3
2109
39
1604
17
7
7
I
7
4
16.2
Tj ul
16.0
14.4
15.71
14.6
14.6
15.7'
15.9
16.0
15.0
14.7
14.6
15.8
14.8
15.2
16.4
15.0
14.8
11.8
16.0
14.6
11.9
16.5
14.4
16.0
14.7
11.6
12.3
12.9
12.3
14.9
16.9
14.1
15.6
16.1
14.5
15.6
16. 1
10.9
14.9
16.2
15.3
15.9
5.9 15.9
3.8
CI Tja n
0 I
3.9
965 14
3.5
471 12
3.9
805 5 3.6
465 9
3.7
84 8
3.9
0 0
3.4
1 3 3.7
432 4
3.7
659 14
3.6
230 14
3.5
82 8 4.3
351 6 3.8
855 12 4.0
0 0 3.6
197 5 3.6
509 13 3.5
0 0 2.5
0 1 3.5
143 4
3.3
2 0
1.7
0 0
4.4
8 0
4.7
4 1 3.8
846 14
3.7
0 0
1.6
37 0 2.1
1 0 2.6
1 0 2.5
522 13 3.6
0 5 6.1
319 14 3.9
111 10 3.8
0 0
3.6
958 13 3.6
41 6
3.8
0 0
3.3
0 0 0.9
489 12
3.7
0 8 5.5
650 11 3.9
7 7 5.4
IR
950
713
18, 19
3
Prec Co Br Habitats
775
4, 17
1106
6
747
3, 4, 8
1039
3
1030
7
837
3, 17
671
3, 4, 16
726
3, 4
956
4, 16
1038
4, 8, 16
1070
8, 16
849
13
1023
11, 13
961
14
651
13
986
1, 3, 16
1034
16, 17
3218
16
686
8
1104
7, 16
1966
15, 16
844
16
1335
1, 16
769
3
1051
3
1750
15
16 15
10, 15
1454
15
1904
16
986
16
843 Co 18, 19
1265 Co 18, 2 1
850
3
685
4
1089
3, 6
861
3, 16, 17
620
3, 8
1982
15
984
3, 17
962
1, 3
931
1
953
3, 18, 19
8
4
7j 5
6
6
3
5,
L F R N S
7 4 4 5
7 6 4 3
71 5 6 5
6 5 7 T
8 4 7 2
8 3 7' 5
9 5 8 6
7i 4 5 5
8 4 6 4
7 4 6 4
7 4 5 4
8 8 7 7
71 10 6 4
7 10 7 7'
9 9 7 7
6 4 6 5
8 3 6 2
7 5 71 3
7 3 8 5
7 5 8 3
9 3 8 1
7 3 8 2
6 5 71 2
9 5 7 9
6 4 7 5
7 6 2 2
7 5 2 2
9 5 8 2
9 3 8 2
8 3 T 5
8 3 6 2
8 7 5 5
8 5 7 Tj
7 4 3 3
7 5 7 7
7 4 7 9
8 3 6 5
9 4 4 1
7 4 8 7
4 5 6 6
4 5 7 7'
7 5 6 5
NJ
1‘ )
Taxon name
Asperula cynanchica
Asp leniurn adianturn-nigrum
Asp lenium adianturn-nigrum
sens.lat
Asp lenium marinum
Asp leniurn obovatum
Asp lenium onopteris
Asp leniurn ruta-muraria
Asp leniurn septentrionale
Aspleniurn trichomanes
Asplenium viride
Aster (alien N American taxa)
Aster lanceolatus
Aster lanceolatus x novi-belgii
(A. x salignus)
Aster linosyris
Aster novae-angliae
Aster novi-belgii
Aster tripolium
Astragalus alpinus
Astragalus danicus
Astragalus glycyphyllos •
A thyrium distentifolium
Athyrium filix-femina
Athyriurn flexile
Atriplex glabri uscula
Atriplex laciniata
Atriplex littoralis
Atriplex longipes
Atrip/ex patula
Atriplex pedunculata
Atriplex portulacoides
Atriplex praecox
Atriplex prostrata
Atrop a belladonna
Avena fatua
A vena sativa
Avena strigosa
Azoll a filicuboides
.
Baldellia ranunculoides
Ball ota nigra
Barbarea intermedia
Barbarea stricta
Barbarea verna
Barbarea vulgaris
Bartsia alpina
n
r
n
-0.44
n
r
n
n
s
n
r
n
n
n
s
n
r
n
s
n
n
-3.0 1
2.76
-1.08
-0.37
1.92
0.50
1.34
-0.02
-0.10
1.10
-0.33
1.17
0.06
-0.34
-0.93
0.38
1.59
-0.88
-0.36
0.38
0.25
-0.10
r
20
100
60
100
90
90
22
150
50
200
150
100
30
30
100
30
90
20
33
30
100
90
87
30
80
10
100
150
150
150
120
1
20
AN
N
AN
AN
N
N VU
N
N
N
N
NE V U
N
N
N
N
N
N CR
N
N
N
N
AR
AC
AC
AN
N
AR
AN
AN
AN
N
N
Aste
Aste
Aste
Aste
Aste
Faba
Faba
Faba
Wood
Wood
Wood
Chen
Chen
Chen
Chen
Chen
Chen
Chen
Chen
Chen
Sola
Poac
Poac
Poac
Azol
Alis
Lami
Bras
Bras
Bras
Bras
Scro
0.15
-0.08
0.07
-0.2 1
45
35
30
45
12
15
35
20
150
120
n
n
s
o
n
s
n
n
N
N
N
N
N
N
N
N
AN
AN
Asp
Asp
Asp
Asp
Asp
Asp
Asp
Asp
Aste
Aste
0.35
0.02
-0.18
RS Chg li ght Len
n -0.47
50
n
45
Fam NS CS
Rubi N
Asp
N
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
Th
Th
Th
Th
Th
Th
Pn
Th
Th
hc
Th
Th
Th
Hy
Hy
hc
hc
p hc
b Th
p hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
hc
Rhizl
0
Rhizl
Rhir l
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
Frag
0
Rhiz2
0
0
0
0
Rhizl
0
0
0
0
0
0
0
0
Rhizl
Rhizl
Sto12
4
5
4
3
2
3
7
1
c
Eur
Eur
Crop
Am4, SAm
Am6
Am6
Gard
Am
Am6
Origin
Eur
c Eur, Asl
Eur
5
6
6 c
3
6
6
1
2
1
6
3
4
4
1
3
4
3
3
3
2
1
1
6
3
6
6
8
8
7
1
7
7
1
5
1
5
7
7
4
6
7
9
4
6
7
7
8
8
9
9
7
7
8
4
1 P2 LF1 LF2 W Clonel Clone2 El E2 C
hc
Rhiz2
7 3
hc
0
7 3
373
9
83
557
703
4
242
357
98
2574
16
780
418
362
39
2318
20
322
62
1847
414
1556
728
270
660
536
1365
735
104
514
1869
18
2146
530
135
0
2184
55
2294
435
1211
211
9
0
0
12
267 '
0
3
0
0
849
0
159
92
36
0
787
0
68
0
349
0
170
131
72
21
344
70
207
1
28
615
.".0
708
172
19
34
898
2
911
43
53
5
0
0
1
3
3
0
0
0
0
12
0
10
13
3
0
12
0
10
0
5
0
5
4
6
9
6
12
7
0
7
3
0
14
12
11
0
11
0
12
0
5
0
3.8
4.9
3.9
3.7
4.3
-0.9
3.3
3.4
0.3
3.5
-0.9
4.2
4.5
4.2
4.2
3.7
3.9
4.6
3.6
3.9
3.6
3.71
3.9
4.0
4.0
4. 1
3.8
3.7
3.7
4, 1
3.7
0.4
3.7
4.4
5.3
4.6
3.7
2.5
3.6
2.0
3.7
3.9
15.6
15.8
16.1
15.2
14,8
11.0
14.8
15.7
11.2
14.4
10.7
14.4
14.8
15.4
15.6
14.8
16.4
15.7
12.8
15.1
16.1
15A
15A
14.6
16.0
15.1
15.7
15.0
15.9
15.6
15.0
11.9
14.6
14.0
15.4
14.5
14.7
13.2
14.5
12.8
15.3
15.8
890
1006
806
936
1109
1275
745
711
2098
1132
1753
1163
979
822
982
1013
603
869
1503
955
717
849
903
1042
774
974
797
975
705
943
943
2069
3, 17
Co 18
3
3, 17
Co 21
7, 15
7
1, 3, / 6
15
1, 16
15, 16
Co 19
Co 19
Co 19
Co 21
3, 4
Co 21
Co 21
Co 19
3, 4, 19, 21
1, 3
3, 4
4
4
13, 14
13, 14
3
4
11, 13, 14
3
3, 14
7, 11, 15
1110
3
1261 Co 18
1197
16
1200
1, 3, 16
1074
3, 16, 17
1825
16
1121
3, 16, 17
1701
15, 16
934
3, 17, 2 1
844
3, 17
GB
IR CI Tjan Tj ul Prec Co Br Habitats
329 47 3 4.0 15.9
842
7
2146 708 14
3.7 14.6 1110
3, 16
7
8
7
7
9
9
8
6
6
5
7
9
9
9
9
7
91
9
9
8
5
71
7
7
7 11
8 10
71 4
8 5
8 7
8 5
7 6
5 8
6
9
5
5
7
8
5
4
7
7
L F
7
6
7
8
7'
7
7
6
8
7
3
5
3
7
7
7
7
7
7
8
7
7
6
7
7
5
8
6
8
6
7
6
7
71
5
5
4
7
7
2
8
8
7
7
6
1
6
6
6
2
2
3
4
6
4
8
7
6
8
7
8
6
6
7
6
7
7
5
8
2
6
7
8
6
8
2
5
5
3
3
2
2
2
3
6
6
R N S
8 2
5 5
Taxon name
Bellis pe ren nis
Berberis vulgaris
Baru la erecta
Beta vulgaris
Betula na na
Betula p endula
Betula pubescens
Bidens cernua
Bide ns tripartita
Blacks ton ia perfoliata
Blechnum spicant
Blysmus compressus
Blysmus rufus
Bo lboscho e nus maritimus
Bo trychium lunaria
Brachy podium pinnatum
Brachy podium sylvaticum
Brassica napus
Brassica nigra
Brassica oleracea
Brassica rapa
Briza maxima
Briza media
Briza minor
Bromopsis benekenii
Bromopsis erecta
Bromopsis inermis
Bromopsis ramosa
Bromus commutatus
Bromus hordeaceus
Bromus racemosus
Bromus secalinus
Bryon ia dioica
Bu ddleja davidii
Bu nium bulbo ca s tan um
Bupleururn baldense
Bupleurum falcatum
Bupleurum rotun difolium
Bupleurum tenuissimum
Butomus umbellatus
Buxus sempervirens
Cakile maritima
Calamagrostis canescens
Calamagrostis epigej os
Calamagrostis purp urea
Calamagrostis scotica
Fam
Aste
Serb
A pia
Chen
Betu
Betu
Betu
Aste
Aste
Gent
Blec
Cype
Cype
Cype
Ophi
Poac
Poac
Bras
Bras
Bras
Bras
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Cucu
Budd
Apia
Apia
Apia
Apia
Apia
Buto
Buxa
Bras
Poac
Poac
Poac
Poac
NS
N
NA
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
AN
NA
NA
AR
AN
N
AR
N
N
AN
N
N
N
N
AR
N
AN
N
N
AN
AR
N
N
N
N
N
N
N
NE
VU
EW
EN
CS
s
n
r
n
n
n
r
r
r
r
n
n
n
n
n
s
n
n
n
s
RS
n
n
n
n
s
n
n
n
n
n
n
n
n
n
n
n
n
Chg Hght Len
8
0.89
250
-0 61
100
-0.02
150
1.23
100
-0.09
-0.23 2500
0.40 2000
67
-0.54
67
-0.43
45
0.12
50
-0.39
37
-1.28
37'
-0.53
100
0.00
15
-0.43
90
0.15
95
-0.17
130
2.88
150
-0.02
130
0.90
100
0.74
62
62
0.75
55
0.28
105
0.25
110
-0.01
150
1.71
170
-0.18
95
1.07
80
-0.37
95
0.74
90
-1.15
400
-0.50
500
3.73
50
0. 14
10
100
30
-4.58
50
-0.97
150 150
-0.04
500
2.54
30
-0.38
120
-0.33
200
0.47
150
100
Origin
1 P2 LF1 LF2 W Clone l Clone2 El E2 C
7 3
h Rhizl
hc
7 3
w Rhizl
Pn
7 3
h Irreg
Hy
b he
9 1
h 0
w Rhizl
2 6
Pn
5 4
w o
Ph
5 4
Ph
w 0
7 6
Th
0
Th
7 5
o
9 2
0
Th
7 3
o
hc
7 3 c
Rhiz2
hc
4 3
Rhiz2
hc
8 4
DRg
Rhizl
Hy
5 6
0
Gn
7 4
Rhizl
he
he
7 3
0
Gard
0
b Th hc
7 3
Th
0
sw 0
8 2
Pn
7 4
0
b Th hc
Eur
Th
0 3
0
7 3
Rhizl
he
9 1
0
Th
7 3 c
0
hc
7 3
hc
0
Eur, Asl
7 4
hc
Rhiz2
hc
7 3
0
7 3
0
Th
8 3
o
Th
7 3
0
Th
Unk
0
Th
0
Gn
9 2
w 0
As
Ph
8 2
0
Gn
9 1
0
Th
Eur, As
8 5
hc
0
Eur?
Th
0
8 3
0
Th
7 4
Rhiz2 DRg
Hy
w O
Ph
9 2
6 3
h 0
Th
5 4 c
h Rhiz2
hc
5 5
h Rhiz2
hc
4 4
h Rhiz2
hc
4 1
h Rhizl
hc
GB
2797
974
1111
607
125
2293
2399
874
1055
7871
2159
384
367
766
1109
6 12
2310
1758
1080
98
1407
239
1853
92
64
798
263
1887
675
2406
516
403
1004
1434
13
3
8
287
161
685
2
576
293
936
10
1
IR CI Tja n Tj ul Prec
3,5 14.5 1105
984 14
892
26 1 3.4 15.2
843
265 1 3.8 15.5
952
205 14 4.6 15.4
0 0 0.71 11.5 1585
390' 0 3.3 14.6 1073
779 0 3.4 14.4 1123
883
281 2 3.9 15.5
884
3.9 15.5
222 5
8 12
198 2 4.0 15.8
3.5 14. 1 1198
831 9
840
0 3.2 15.2
3.8 13.6 1371
0
7
7 4.3 15.1 1020
21
1 3.0 13.8 1259
13
745
3.6 15.9
0
2
3.7 14.7 1062
78 12
9 14
3.6 15.1
7
14
822
5 11 4.0 15.8
922
1 4.9 15.6
993
7 3.9 15.0
63
932
11 4.6 15.8
971
3.5 14.9
2
58
9 19
5.2 16. 1
8
896
3.0 15.3
0
764
3.7 15.9
3
4
757
3.6 15.9
0
995
3.5 14.9
0
45
772
3.9 16. 1
0
3
3.71 14.7 1051
83 14
852
0 4.0 15.8
7
8 16
3.9 15.8
4
1
726
3.6 16.0
4
930
3.9 15.4
26 14
633
3.3 16.3
0
808
8 6.1 16.5
776
3.4 16.0
0
753
1 3.8 16.0
684
0 4.2 16.4
715
0 3.71 16.0
716
3.7 16.6
0
131 13 4.4 14.6 1011
723
0 0 3.4 157
812
9 4 3.7 15.7
1.6 13.1 1346
0 0
888
0 0 3.6 12.8
Co
Co
Co
Co
Co
Co
Br Habitats
6
1, 3
11
18, 19
10, 12, 15
1
1
13, 14
11, 13
7
1, 2, 10, 16
11
21
21
7, 16
7
1
3, 4, 17
3, 4
17, 18
4
3, 17, 19
7
4
1
7
3
1
3
4, 6
3, 6
4
3
3, 17
4
18, 19
3
4
3, 6
13, 14
1, 3
19
11
3, 11
1, 11
11
L F R N S
8 5 6 4
7 4 8 3
7 10 7 7
9 5 7 8
7 8 1 1
7 5 4 4
7 7 4 4
8 9 7 7
8 8 7 7
8 5 8 2
5 6 3 3
8 8 8 3
8 8 7 4
8 10 8 7 '
8 4 6 2
7 3 8 3
6 5 6 5
7 4 7 7
8 5 7 6
8 4 71 8
7 5 7 6
7 3 4 2
8 5 7 3
7 4 5 5
5 5 7 5
7 4 5 3
8 4 8 5
4 6 7 7
7 4 8 6
8 4 71 4
6 6 7 8
6 4 5 4
7 5 7 7
7 5 7 5
71 4 9 4
9 3 8 2
6 3 9 3
8 3 9 4
9 7 8 4
7i 11 71 71
4 4 8 5
9 6 7 7
7 9 7 5
7 7 7 6
71 8 6 3
8 8 6 4
Taxon name
Calamagrostis stricta
Calendula officinalis
Callitriche brutia
CaIlitriche hamulata
Callitriche hamulata sens.lat
Callitriche hermap hm ditica
Callitriche obtusangula
Callitriche platycarpa
Callitriche stagnalis
Ca& riche stagnalis sens.lat
Callitriche truncata
Fam
Poac
Aste
Cal
Cal
Cal
Cal
Cal
Cal
Cal
Cal
Cal
Eric
NS CS
N
AN
N
N
N
N
N
N
N
N
N
N
RS Chg_ Hght Len
100
r -0.74
50
25
80
25 80
1.12
50
0.21
60
1.35
15 100
15 60
15 80
1.51
n
20
0.47
s
60
n -0.64
P1 P2 LF1
hc
Th
a
Hz
a
a p Hz
a p Hz
Hy
p
Hy
p Hz
p Hz
p Hz
Hz
Ch
LF2 W Clonel
Rhiz1
0
Node2
Node2
Hy
Node2
Hy
Irreg
Irreg
Node2
Hy
Node2
Hy
Node2
Hy
0
0
Pn
lrreg
Irreg
Irreg
Irreg
Nodel
Irreg
lrreg
9
5
6
4
8
7
7
7
9
5
1
2
3
6
2
3
3
3
1
3
Clone2 E1 E2 C
2 6
Unk
Origin
GB
22
764
213
1758
1758
392
757
1006
1488
2647
52
2434
CI Tjan Tjul Prec Co Br Habitats
11
928
6 0 2.9 14.2
3, 17
822
24 10 4.0 15.7
13
63 3 4.1 14.9 1087
13, 14
3.3 14.2 1162
282 6
11, 13, 14
282 6 3.3 14.2 1162
13
115 0 3.1 13.7 1120
13, 14
840
157 4 4.1 15.7
11, 13
947
173 5 3.7 15.1
13
346 5 3.71 14.6 1083
11, 13, 14
769 11 3.5 14.4 1105
13, 14
719
1 2 4. 1 16.1
10, 12
905 11 3.5 14.3 1157
IR
L
9
8
8
7
7
7
7
6
7
6
7
7
F R N S
9 4 2
5 7 7
10 5 5
11 6 5
10 5
12 7 5
11 7 6 1
10 7 7 0
10 6 6 1
10 6 6
12 7 7
6 2 2
NJ
in
Taxon name
Carex binervis
Carex buxbaumii
Carex capillaris
Carex cary ophyllea
Carex chordorrhiza
Carex curia
Carex davalliana
Carex depauperata
Carex diandra
Carex digitate
Carex dioica
Carex distans
Carex disticha
Carex divisa
Carex divulsa
Carex echinata
Carex elate
Carex elongate
Carex erk etorum
Carex extensa
Carex filiformis
Carex /lam a
Carex if ava
Carex hirta
Carex hostiana
Carex humifi s
Carex lachenalii
Carex laevigata
Carex lasiocarpa
Carex if mosa
Carex magellanica
Carex marifi ma
Carex microglochin
Carex montane
Carex muricata
Carex nigra
Carex norvegica
Carex ornithopoda
Carex otrubae
Carex Ova rls
Carex pallescens
Carex panicea
Carex paniculata
Carex pauciflora
Carex pendula
Carex pil ulifera
Fan/
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
Cype
NS CS
N
N VU
N
N
N VU
N
N EX
N CR
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N VU
N
N
N
N
N
N
N
N
N
N VU
N
N
N
N VU
N
N
N
N
N
N
N
N
N
Chg Hght Len
-0.17
90
70
-0.35
40
-0.20
15
40
0.17
50
25
60
0.22
60'
0.04
15
-0.35
30
-0.47
72
-0.03
90
-0.35
70
82
-0.75
40
-0.32
95
0.06
80
-0.46
17
-0.23
40
0.23
50
0.53
50
70
0.17
70
-0.05
57
-0.0 1
10
-0.22
20
-0.0 1
120
0.73
120
n
0.14
40
s -0.02
40
s -1.34
18
r
12
s
0.68
35
n
85
n -0.0 1
70
r
30
r
0.28
15
n -0. 14
100
n -0.2 1
90
n -0.5 1
60
n -0.3 1
50
n -0.11
150
n -0.59
25'
n
1.30
150
n -0.04
35
RS
n
r
s
n
r
n
x
r
n
s
n
n
n
s
n
n
n
s
s
n
r
n
r
n
n
s
r
n
n
1 P2 LF1 LF2 W Clonel Clone2 El E2 C
hc
0
7 /
hc
Rhizl
4 4 c
hc
Ogr
2 6
hc
Rhizl
7 4
hc
Rhiz2
2 6 c
hc
Rhizl
4 6
hc
0
7 3 c
hc
Rhiz l
9 2
hc
Rhizl
5 6
hc
0
5 3 c
hc
Rhizl
2 6
hc
0
8 3
hc
Rhiz2
7 4
hc
Rhizl
9 2
hc
0
8 4
hc
0
5 3
Hy hc
0
7 5
hc
0
5 4 c
hc
Rhizl
5 4 c
hc
Ogr
8 3
hc
Rhiz2
7 4 c
hc
Rhiz2
8 3
hc
0
5 3
hc
Rhiz2
7 3
hc
0
7 3
hc
Ogr
7 5 c
hc
Rhizl
1 6
hc
0
7 1
Hy hc
Rhiz2
4 6
Hy hc
Rhiz2
4 6
hc
Rhiz l
4 6
hc
Rhiz2
1 6
hc
Rhiz2
1 3
hc
Rhizl
7 3 c
he
0
8 4
hc
Rhiz2
5 4
hc
0
1 6
hc
0
4 3
hc
0
8 4
hc
0
5 4
hc
0
5 4
hc
Rhiz2
5 3
hc Hy
0
7 3
hc
Rhizl
4 6
hc
0
8 3
hc
0
7 3
Origin
GB
IR
1927 647
3
1
120
0
1866 437
4
0
1190 219
1
0
9
1
378 294
39
0
1100 204
814 213
12271 48 1
162
780 24
2100 78
294 22
72
1
33
440 19
13
2706 95
1
1958 57
1578 42
30
9
1012 29
461 16
423 19
131
83
1
48
912 6
2582 87
6
15
1636 40
2422 74
1596 18
2526 89
1516 50
377
1409 30
2111 44
CI Tjan
3 3.3
0 3.3
0
1.2
11 3.4
0
1.4
0
2.7
0 4.0
0 4. 1
0
3.6
0
3.3
0 2.7
9 4.3
3 3.6
1 4.4
4.2
3.4
3.8
3.6
2.9
4.5
3.5
3.5
3.2
3.7
3.2
3.9
-1.1
3.6
3.1
3.3
1.7
3.6
-0.7
4.0
1
3.8
3.5
-0.8
0
2.1
9 4.0
7
3.4
1 3.0
6 3.5
5 3.7
0
1.7
10
3.9
8 3.3
Tjul Prec
14.0 1243
13.5 1410
11.9 1636
14.7 1079
11.9 1176
13.6 1303
16.3
833
15.8
823
14.4 1042
15.5
859
13.2 1404
15.0 1039
15.0
9 10
16.4
727
15.8
844
14.1 1207
15.2
903
15.5
899
15.3
801
14.5 1228
16.3
708
14.5 1104
14.7i 1585
15.0
963
13.8 1290
16. 1
845
10.4 1948
14.4 1240
13.6 1383
13.4 1431
12.8 1808
12.9
964
11.0 1855
15.5 1193
15.4
939
14.3 1131
10.8 1901
13.8 1272
15.3
939
14.3 1138
14.2 1236
14.3 1139
14.9 1036
12.1 1789
15.3
928
14.1 1197
1, 11
12
1, 14
8
11
Br Habitats
8, 10, 16
11
7
7
11
11
11
3
11
1, 16
11
Co 21
11
6
3, 6, 7
11, 12, 14
11
1
7
Co 21
6, 7
7, 11
1, 11
6
11
7
11, 15
1, 16
11
11, 12
12
Co 19
11
6, 8
3, 7, 8
11
15
7, 16
11
3, 10
1
Co
L
7
8
9
7
9
8
9
5
8
5
8
8
7
8
7'
8
7
5
6
8
7
7
7i
T
8
8
8
5
8
8
9
9
9
7
T
7
8
8
6
7
6
8
6
8
5
7
F
6
8
6
4
9
9
9
4
9
5
9
6
8
7
4
8
10
8
4
7
7
5
9
7
9
3
7
8
10
10
9
8
9
6
4
8
7
3
8
7
6
8
9
9
8
5
R N S
3 2
7 2
8 2
7 2
4 3
3 2
8 2
71 4
5 3
8 4
6 2
7 5
6 4
7 6
7 6
3 2
7 5
6 6
7 1
7 5
8 5
6 2
8 2
7 6
6 2
8 2
4 1
5 4
6 3
4 1
2 1
7 2
8 2
4 1
6 4
4 2
7 2
9 3
7 7
5 4
5 4
4 2
6 6,
1 1
7 6
3 2
odel
hizl
ode2
ode2
1
3
1
2
3
2
4
3
3
4
0
1
3
3
3
3
3
5
9
9
7
7
7
8
7
a
8
9
7
7
7
1
1
2
3
4 c
4
3
5
5
7
7
7
8
3
E2 C
4
2
2
6
1
3
4
6
6
6
3
2
5
6
6
6
7
one l Clone2 El
7
hizl
7
8
1
1
hiiz2
h
4
7
7
hiz2
5
hiz2
1
hIzz21
h
1
7
7
7
2
hiz2
5
hizl
5
Eur
Eur
Eur
SAf
Eur?, Asl ?
Origin
.
Co
Co
Co
Co
Co
14
11, 19
13
7
1
18, 19
3, 6, 17
11, 14
1
13, 14
18
7, 17
3
3, 4, 17
6, 7
6, 7
7, 19
19, 21
21
10
18
3, 16, 17, 18
1
1
1
15, 16
15, 16
1194
1278
731
947
895
962
868
1578
950
936
1027
878
744
836
1093
632
980
1063
841
1170
825
896
763
1166
797
2003
2178
14.2
13.9
16.4
15.3
15.4
15.9
15.0
13.9
15.2
15.0
15.1
15.5
16.1
15.4
14.5
15.6
15.1
14.3
16.1
15.4
16.5
15.5
16.2
14.7
16.2
11.4
11.3
3.4
3.8
3.9
3.9
3.6
5.7
3.6
3.4
3.6
3.8
4.8
4.1
4.0
3.8
3.6
3.8
4.0
3.8
4.4
5.8
5.5
4.0
3.7
3.7
3.8
0.4
0.8
6
5
0
10
9
10
2
1
12
3
14
10
3
5
11
2
14
0
8
0
2
14
0
0
0
0
0
761
179
0
243
52
8
30
42
140
304
172
494
1
56
975
125
710
3
17
0
0
357
0
31
0
0
0
2203
477
24
1131
1488
79
303
296
1693
955
463
1216
153
884
2658
1239
1811
111
457
3
3
1361
233
131
10
77
46
11,14
1200
3.1 13.9
Tj ul Pree Co Br Habitats
11
754
16.0
11, 16
13.9 1239
15.4 1125 Co 18
15
10.6 1541
11
936
13.1
1, 14
14.8 1063
11, 14
814
15.7
11
14.0 1193
15, 16
11.5 1699
15
11.3 2262
3, 6, 7
823
15.7
1, 14
841
15.7
1
14.8 1050
15
11.3 1904
11
14.6 1072
11, 14, 19
14.2 1169
2
IR CI Tjan
72 3 3.8
656 4 3.3
42 7 5.3
0 0 -1.2
0 0 3.1
3.7
675 5
160 41 3.9
3.3
731 0
0 0 0.7
0 0 0.3
48 1 3.6
3.8
102 0
3.6
608 2
0 0 -0.1
230 1 3.3
855 9 3.5
1168 434
GB
672
1874
56
17
4
1879
1186
1924
31
71
1043
417
1899
83
913
2337
4
9
7
s
7
7
71
8
8
9
8
9
8
8
4
a
2
3
6
2
s 6
4 5
7 6
4 2
s 5
7 71
7 3
7 2
7 3
6 5
6 5
8 3
6 3
a 3
8 3
5 2
7
8
7
8
8
7
8
4
9'
8
7
5
6
9
2
8
4
R N
6 6
5 2
7 3
3 2
7 5
6 6
7 7
4 2
7 2
7 3
6 4
7 6
6 5
6 3
5 4
6 2
a,
4
71
8 1
8
8
7
3
4
7'
8 1
8
8
L F
7
8,
9
8
0
0
1
0
0
0
3
1
0
0
1
1
t•J
Taxon name
Fam
Cerastium arvense
Cary
Co a st/UM brachypetalum
Cary
Cerastiurn cera stoides
Cary
Cerastiurn diffusum
Cary
Cerastium fontanum
Cary
Cerastium glomeratum
Cary
Cerastium nigrescens
Cary
Cerastiurn pumilum
Cary
Cerastium sernidecandrum
Cary
Cerastium tomentosurn
Cary
Cera tocapnos claviculata
Furna
Ceratochloa carinata
Poac
Ceratoch/oa cathartica
Poac
Ceratophyllum demersum
Gera
Ceratophyllum submersum
Cera
Ceterach off icinarum
AspI
Chaenorhinum minus
Scro
Chaerophyllum temulum
Apia
Chamaecyparis lawsoniana
Cupr
Chamaemelum nobile
Aste
Charnerion angustifolium
Onag
Chelidonium rnaj us
Papa
Chenopodium album
Chen
Chenopodium album agg.
Chen
Chenopodiurn bonus-henricus Chen
Chenopodium
chenopodioides
Chen
Chenopodiurn ficifoliurn
Chen
Chenopodiurn glaucurn
Chen
Chenopodiurn hybridum
Chen
Chenopodiurn murale
Chen
Chenopodiurn polyspermum Chen
Chenopodiurn rubrum
Chen
Chenopodiurn urbicurn
Chen
Chenopodiurn
Chen
Chrysanthemum segetum
Aste
Chrysospleniurn alternifolium Saxi
Chrysosplenium
oppositifolium
Saxi
Cicendia filiformis
Gent
Cicerbita alpina
Aste
Cicerbita rnacrophylla
Aste
Cichorium intybus
Aste
Cicuta virosa
Apia
Circaea alpina
Onag
. -1.27'
s
0.55
5.
-0.36
-0.70
15
10
130
175
110
150
30
p
a
p
p
p
p
p
Ch
Th
hc
hc
hc
Hy
Gn
hc
n
s
r
N
N
N VU
AN
AR
N
N
Th
Th
Th
Th
Th
Th
Th
Th
Th
Th
he
-0.17
1.90
-1.32
-0.32
-1.63
0.62
n' 1.00
1 -4.57
-2.60
-1.80
n
0.62
a
a
a
a
a
a
a
a
a
a
p
s
N
AR
AR
AR
AR
AR
N
AR
AR V U
AR
N
30
90
50
100
100
50
70
100
35
60
20
RS Chg Hght Len P1 P2 LF1 LF2
n -1.05
30
p
Ch
30
a
Th
s -0.05
12
p
Ch
n 0.38
23
a
Th
n
1.40
30
p
Ch
n
1.44
30
a
Th
r
5
p
Ch
s -0.171
11
a
Th
n
0.50
14
a
Th
2.97
30
p
Ch
n 0.57
75
a
Th
2.09
80
p
hc
0.63
100
p
hc
n 0.87
100 p
Hy
n
0.39
100 p
Hy
n -0.30
20
p
he
-0.63
25
a
Th
n -0.64
100
b
hc
4 100
p
Ph
n -0.92
30
p
hc
n -0.0 1
150
p
Gn hc
-0.72
90
p
he
n
100
a
Th
n -0.73
100
a
Th
-1.79
50
p
hc
NS CS
N
AN
N
N
N
N
NE VU
N
N
AN
N
AN
AN
N
N
N
AR
N
AN
N
N
AR
N
N
AR
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
Node2
0
Rhiz2
Rhiz2
0
0
Rhiz2
0
0
0
0
0
0
0
0
0
0
Sto12
W Clonal Clone2
Rhiz2
0
Nodal
0
Rhizl
Nodal
0
0
0
0
Rhiz2 Node2
'0
0
0
Irreg
Irreg
0
0
0
w 0
h Node2
h Root
h 0
h 0
h 0
h 0
8
5
4
7
9
4
7
7
7
7
8
7
7
7
8
8
3
4
5 c
6
2
2
3
4
4
6
6
4
4
4
4
4
3
6
Eur
El E2 C
Origin
5 6
8 3
Eur
1 3
7 3
5 4
8 3
4 1
7 3
7 3
Eur
7 1
Am4
Am, SAm
6 6
7 4
9 2
7 3
7 3
Arn4
8 2
5 6
7 5
6 5
6 5
Eur
IR
39
0
0
247
985
823
0
0
61
140
20
1
0
62
3
787
170
0
46
115
650
220
782
782
81
2067 688 6
66
371 5
4
0 0
724 42 0
1312 60 10
139 139 0
40
0 0
3.4
5.4
-1.3
3.3
3.8
3.5
2.2
4.2
3.9
4.0
3.7
4.2
3.9
3.8
4.0
4. 1
3.8
2.8
CI Tja n
0 3.3
0
3,4
0 -0.4
12 4.0
14 3.5
14 3.6
0 3.0
0
4,1
12
3.7
8
3.7
2
3.3
2 3,8
9 4.2
1 3,7
5 3,9
9 4.0
0
3.6
5 3.6
4
3.6
13 4.6
10
3.4
9 3.7
14 3.7
14
3.7
1 3,4
39
0 2
745
11 7
157
2 2
285
0 1
412
8 13
998
8 12
1267 142 8
239
1 0
180
0 6
1682 471 10
790
0 0
GB
8 14
2
29
1174
2805
263 1
2
87
1117
1303
1122
183
184
927
208
1093
1468
1786
827
308
2603
1662
2340
2340
1363
14.3
15.6
10.5
14.9
15.5
14.8
13.0
16.7
16. 1
16. 1
16.2
16.0
16.0
15.7
16.1
16.1
14.8
14.4
1168
1063
1375
920
840
926
1836
1, 14
10
15, 16
3, 17
3
1/ , 13, 14
1
6 12 Co 6, 2 1
745
4
724
17
695
4
797
4
794
4
800
4, 11
762
4
752 Co 15, 19
1022
4
1042
11, 14
Tjul Prec Co Br Habitats
15.3
773
8
16.3
657
3, 7
10.8 1994
15
14.6 1062 Co 18, 19
14.4 1106
6
14.5 1079
3, 5
11.5 1256
16
16.2
806
7
15.3
846
8, 18, 19
15.2
898
3, 17, 19
14.6 1111
1, 9
16. 1
744
3, 4
16.1
796
3, 4
15.8
770
13
16.3
705
13
15.0 1061
3, 16
15.3
899
3, 4, 17
15.4
890
3
15.3
965
17
15.7
990
8
14.5 1075
3, 17
15.3
905
3
14.8 1018
4
14.8/ 1018
4
15.2
883
3
5
9,
7
7
5
7
4
8
7
8
7
8
7
7
71
7•
7
5
L
8
9
8
9
7
7
9
8
8
8
5
8
7i
7
7
7
8
6
5
8
6
6
7
7
8
9
8
6
5
4
10
71
7
6
6,
4
6
6
7
5
4
5
8
F
4
3
8
4
5
5
3
2
3
3
5
5
4
12
12
3
4
5
5
7
5
5
5
5
5
5
3
6
6
7
7
5;
7
6
7i
7
6
7
7
71
7
6
6
6
5
6
8
71
7
7
5
2
6
6
5
5
5
8
7
9
7
7
8
8
7t
9
5
6
4
0
3
0
0
0
1
4
5
5
7
71 1
7 1
8 0
71 7i
R N S
5. 3 0
8 2 0
5 4 0
61 3 1
5 4 0
6 5 0
5 1 0
8 1 0
6 3 0
7 5 1
4 5 0
6 6 0
5, 5 0
7 7
8 8
8 1
7 4
t.. )
CO
Taxon name
Circaea alpina x lutetiana (C
x intermedia)
Circaea lutetiana
Cirsium acaule
Cirsium arvense
Cirsium dissectum
Cirsium eriophorum
Cirsium heterophyllum
Cirsium palustre
Cirsium tuberosurn
Cirsium vulgare
Cladium mariscus
Claytonia perfoliata
Claytonia sibirica
Clematis vitalba
Clinopodium acinos
Clinopodium ascendens
Clinopodium calamintha
Clinopodium menthifolium
Clinopodium vulgare
Cochlearra anglica
Cochlearia atlantica
Cochlearia danica
Cochlearia micacea
Cochlearia officinalis
Cochlear la officinalis sens.lat
Cochleada pyrenaica
Coeloglossum viride
Coincya monensis
Coincya wrightii
Colchicum autumnale
Colutea arborescens
Conium maculatum
Conopodium maj us
Consolida aj acis
Convallaria maj alis
Convolvulus arvensis
Conyza canadensis
Corallorhiza trifida
Coriandrum sativum
Cornus sanguinea
Cornus sericea
Cornus suecica
Coronopus didymus
Coronopus squamatus
Corrigiola litoralis
Onag
Onag
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Cype
Port
Port
Ranu
Lami
Lami
Lami
Lami
Lami
Bras
Bras
Bras
Bras
Bras
Bras
Bras
Orch
Bras
Bras
Lili
Faba
Apia
Apia
Ranu
Lili
Cony
Aste
Orch
Apia
Corn
Corn
Corn
Bras
Bras
Cary
NH
N
N
N
N
N
N
N
N
N
N
AN
AN
N
N
N
N
N
N
N
NE
N
NE
N
N
N
N
N
NE
N
AN
AR
N
AN
N
N
AN
N
AN
N
AN
N
AN
AR
N
CR
VU
DD
EN
VU
Fam NS CS
r
n
n
s
n
n
n
n
n
n
s
r
n
n
r
n
s
n
n
s
n
s
r
n
n
n
n
n
n
n
n
n
r
n
n
a
P
P
a
p
a
P
p
p
a
a
a
a
a
a
p
a
P
P
p
P
b
P
a
b
P
b
b
P
b
P
p
p
b
Hght Len P
0.48
45
-0.38
60
-0.52
15
0.47
120
-0. 14
60
-0.08
150
-0.44
120
0.15
175
0.41
80
0.80
150
0.11 200
0.50
30
1.28
40
0.00 3000
-1.59
22
0.04
60
-0.3 1
60
60
-0.67
77
0.02
40
20
3.31
25
10
30
-0.18
30
30
-1.34
22
0.43
50
90
-0. 14
35
400
-0.02
250
-0.19
40
60
0,25
25
-0.70
100
1.12
100
0.61
22
50
-0.06
400
300
-0.42
20
1.77
18
0.33
25
-0.96
25
RS Ch
p
p
p
p
Gn
Gn
hc
Gn
hc
hc
he
hc
hc
hc
Hy Gn
Th
Th hc
Ph
Th
hc
hc
hc
hc
hc
he
Th
hc
hc
hc
hc
Gn
hc
Ch hc
Gn
Ph
hc
Gn
Th
Gn
Gn
Th
Gn
Th
Ph
Pn Ph
hc
Th
Th
Th
0
Rhir l
0
0
0
0
Rhizl
Rhiz2
Rhizl
Rhizl
0
0
0
0
0
0
0
0
0
0
Dtb
0
0
0
0
Rhiz2
Rhiz2
0
0
0
Root
0
Rhiz2
0
0
0
o
Rhiz2
Rhiz2
Rhial
Root
Rhizl
0
Rhizl
0
P2 LF1 LF2 W Clonal
Nodal
3
3
2
8
8
3
6
2
7
4
3
4
4
1
8
7
5
8
3
3
3
2
3
6
1
1
1
1
6
6
3
6
2
1
3
3
3
3
5
1
3
4
4
2
4
4
7
7
7
9
7
7
7
4
7
1
3
3
1
4
8
7
7
7
7
7
7
7
7
4
5
8
7
8
c
Clone2 El E2 C
Unk
Am
Eur?
Am
Eur
Eur
Am4
As2, Am4
Origin
570
2053
734
2736
539
447
761
2697
17
2789
232
704
1166
954
551
657
129
1
1317
297
2
588
32
1051
1245
124
964
61
1
301
166
1847
2520
360
439
1841
1048
102
201
1179
421
218
1284
1290
2
GB
2.7
3.7
3.7
3.6
4.0
3.6
2.0
3.5
3.9
3.5
4.1
3.6
3.2
4.0
3.6
4.2
3.7
4.8
3.5
4.5
4.4
4.5
0.2
3.9
3.7
1.5
3.2
4.2
5.8
3.6
3.7
3.9
3.5
3.8
3.3
3.8
3.8
2.5
3.9
3.7
3.5
0.9
4.1
4.0
6.2
CI T an
49 0
748 8
0 3
968 14
553 2
0 0
1 0
95 12
0
98 14
25
1
10
2
6
0
0
6
2
4
0
2
9
0
0
13 14
0
27
2
30
3
0
21
0
3
0
1
0
0
47 13
75
7
7
0
43 14
11 10
0 0
2 3
52 1
111 0
0 0
315 14
149 9
0 0
IR
13.4
14,9
16.1
14.5
15.0
16.0
12.9
14.5
16.1
14.5
14.7
15.6
14.4
16.0
15.8
15.9
16.4
16.3
15.4
15.6
12.8
14.9
11.4
14.0
13.9
12.9
14.1
14.8
15.7
15.8
16.3
15.2
14.5
16.2
15.5
15.2
16.0
13.5
16.0
15.8
15.0
11.6
15.5
15.7
16.0
T'ul
1477
1041
742
1092
1043
745
1476
1105
882
1103
1085
768
1092
815
778
854
641
844
880
917
1830
1056
2221
1246
1287
1567
1148
1037
986
809
671
915
1102
703
831
929
766
956
763
810
936
1853
921
821
1052
Prec
Co
Co
Co
Co
Co
Co
Co
1
1
7
3, 4, 6
11
7
6, 16
11, 14
7
3, 5, 6, 7
11
4, 17, / 9
1, 3
3
16
3, 16
3
1
7
21
18
3, 18
15
18, 21
18
11, 16
7
3, 17, 19
18
6
3, 17
3
1, 6
3, 4, 17
1, 7
3, 4
3, 4, 17, 19
1, 2
3, 17
1, 3
1, 3, 17
10, 15
4, 17
3, 4
19
Br Habitats
L
5
4
5
4
4
5
4
5
7
6
5
5
7
6
7
7
4
4
4
6
4
6
6
4
6
8
4
6
8
6
5
10
6
7'
4
2
5
3
5
4
8
6
6
F
N
6
6
3
6
2
5
5
4
3
6
4
5
6
5
1
6
3
5
• 4
6
5
5
2
5
i 4
3
2
3
3
4
3
8
5
4
5
6
6
4
5
6
6
1 2
6 7
7 7
5 5
R
S
Taxon name
Corylus avellana
Cory nephorus ca nes cens
Cotoneaster bullatus
Cotoneaster cambricus
Cotoneaster horiz ontalis
Cotoneaster integrifolius
Cotoneaster microphyllus
agg.
Cotoneaster simonsii
Crambe maritima
Crassula aquatica
Crassula helmsii
Crassula tillaea
Crataegus laevigata
Crataegus m on ogy na
Crepis bien nis
Crepis capillaris
Crepis foetida
Crepis morn s
Crepis paludosa
Crepis praemorsa
Crepis vesicaria
Crithmurn maritimum
Crocosmia aurea x poli sh (C.
x crocosmiiflora)
Cruciata laevipes
Cryptogramma crispa
Cuscuta epithyrnum
Cuscuta europaea
Cymbalaria muralis
Cynodon dactylon
Cynoglossum ge rma nicurn
Cynoglossum offi cinale
Cynosurus cristatus
Cyperus fuscus
Cyperus longus
Cypripedium calceolus
Cystopteris dickie an a
Cystopteris fragilis
Cystopteris montan a
Cytisus seopar/us
Dab oe cia ca ntabrica
Dactylis glomerata
Dacty/orhiza fuchsii
Dactylorhiza incarnata
Dactylorhiza lapponica
N
N
N
AR EN
N
N
N EN
AN
N
AN
N
N
N
N
AN
NA
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
Irid
Rubi
Adia
Cuse
Cusc
Scro
Poac
Bora
Bora
Poac
Cype
Cype
Orch
Wood
Wood
Wood
Faba
Eric
Poac
Orch
Orch
Orch
CR
VU
VU
VU
VU
AN
AN
N
NA VU
AN
N
Rosa
Rosa
Bras
Cras
Cras
Cras
Rosa
Rosa
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Apia
N
NS CS
N
N
AN
AN EN
AN
AN
Fam
Beth
Poac
Rosa
Rosa
Rosa
Rosa
-1.20
-0.27
0.60
0.23
$
n
n
n
n
r
n
n
r
r
n
n
r
s
r
r
n
r
n
o
n
n
n
s
-0.69
-0.25
0.00
0.12
-0.06
0.33
-0.33
3.11
-0.77
-0.63
-1.28
0.04
-0.10
-0.10
-0.52
-1.09
0.02
-0.32
2.22
0.86
0.32
-0.76
-0.02
-0.17
n
n
n
n
s
1.54
3.55
0.29
60
60
15
1
2
8,
30
75
75
75
20
100
30
20
20
15
200
50
120
50
45
21
100
300
62
8
10
5
1000
1000
120
75
60
60
80
60
80
45
a
a
a
a
a
30 p
a
Gn
hc
hc
Th
Th
Ch
hc
hc
hc
hc
Th
he
Gn
hc
hc
hc
Pn
Ch Pn
he
Gn
Gn
Gn
Pn
Pn Ph
hc
Hz
Hy Ch
Th
Ph
Ph
hc
b Th hc
a hc Th
hc
hc
hc
hc
hc
w
w
h
h
h
h
w
w
h
h
h
h
w
w
Nodel
Node2
Rhizl
ORg
Rhiz2
0
Sto12
Sto12
Node2
Rh1z2 Node2
0
0
0
0
Rhizl
Rhizl
0
0
Rhiz2
0
0
0
0
0
0
0
0
0
0
Irreg
0
0
0
0
0
0
0
0
0
0
0
8
7
7
7
8
8
5
?
3
4
7
8
8
7
5
4
7
4
8
7
9
9
7
7
7
7
8
7
5
7
7
7
c
5
3 c
4
3
4
3
6 c
?
6
6
3
1
4
4
4
3
4
3
4
4
1
2
3
3
3
3
4
3 c
3
4 c
3
6
Eur
Gard
Eur
Aus, NZ
Asl
Asl
,
RS Chg Hght Len P1 P2 LF1 LF2 W Clone l Clone2 El E2 C
Origin
n -0.54
600
p
Ph
w 0
7 3
hc
r
0.01
32
P
h 0
8 3
400
p
Ph
w 0
As l
150
p
Pn
vv 0
Eur
7 1
100
p
Pn
w 0
As 1
100
p
Pn
w 0
As l
1446
1475
466
494
126
2059
2
50
7 18
2745
11
33
22
5
1118
22
2288
0
2707
2214
1189
15
576
973
25 1
2
604
104
597
2496
288
2525
33
75
1162
1
1227
301
731
0
23
27
0
619
0
0
42
962
0
0
0
0
191
0
553
33
981
774
331
0
135
122
46
0
8
0
0
946
th
907
0
0
357
0
302
138
1
1
1
1
1
1
1
12
3
5
11
0
8
11
1
13
0
14
2
0
0
0
14
14
4.0
3.2
1.8
4.3
3.7
3.7
6.4
3.71
3.9
3.5
4.4
5.7
2.2
0.9
2.6
-0.2
3.4
4.5
3.6
3.5
3.6
3.2
3.9
3.7
4.8
3.2
3.8
4.1
3.5
3.6
3.7
3.6
4.0
1.7
2.7'
1.71
4.1
5.1
GB
IR CI Tjan
2470 870 5
3.5
22
0 3 4. 1
237
10 0
3.7
1
0 0 5,4
855
63 2
3.8
537 103 2
3.9
14.6
15.2
12.9
15.9
16.4
15.0
15.6
16.5
15.8
14.5
16.6
16.1
13.9
12.3
13.5
11.3
14.6
14.4
14.5
14.7
14.4
12.5
14.7
14.9
15.5
14.0
15.8
16.2
16.2
14.7
16.3
14.6
16.4
13.2
13.4
13.3
15.7
15.4
1129
892
1678
846
679
990
1156
676
780
1103
758
943
1270
1379
1361
215 1
1085
1265
1091
1063
1127
1910
1, 3, 17
6
10, 15, 16
10
14
3, 16, 17, 19
3, 5, 17
1
7, 19
6
13
11, 13
7
16
16
15, 16
3
10
6
11
11
11
1174
3, 16, 17
1086
3, 10, 16, 17
938 Co 19
1459
14
826
13, 14
734
3
674
1
1073
1, 3
7 11
3, 6
1074
7
707 Co 19
1079
7
1334
11, 16
1333
7
858
3, 5, 6, 17
1042 Co 18
Tj ul Prec Co Br Habitats
14.6 1094
1, 3
16.2
649 Co 19
15.3
965
1, 3, 16, 17
15.5
796
7, 16
15.5
933
3, 16, 17
14.7 1194
3, 17
7
6
7
7
6
7
8
6
8
7
9
8
5
5
6
5
8
8
7
7
8
5
6
5
5
6
7
5
4
5
4
5
8
9
4
7
7
7
5
5
5
8
9
8
7 3
6 5
9 5
8 9
7 10
8 7
5 5
6 5
8 5
7 4
9 4
8 5
6 7
8 3
8 5
9 6
4
7
2
2
6
7
7
8
8
6
5
7
8
8
8
9
4
3
7
7
6
6
7
6
8
5
6
4
7
7
7
7
6
7
6
9
7
7
4
5
3
2
7
6
5
71
6
4
4
5
4
2
4
2
4
2
6
3
2
2
4
4
7
5
7
2
5
6
6
4
3
5
4
3
7
5
L F R N S
4 5 6 6
9 1 3 1
7 4 7 4
8 3 7 2
8 3 8 4
7 3 7 4
f..a
c)
Taxon name
Fam NS CS
RS Chg
Hght Len P1 P2 LF1 LF2
40
p
Gn
30
p
Gn
50
p
Gn
25
p
Gn
30
p
Gn
30
a
Hz
40
p
hc
100
p
Pn
100
p
Pn
100
a
Th
100
b
hc
120
b
hc
150
p
hc
60
p
hc
70
p
hc
100
a
Th
60
a
Th
p
Ch
37
20
p
Ch
6
p
Ch
150
b
hc
35
a
Th
50
a
Th
10
p
Ch
8
p
Ch
60
a
Th
80
p
Ch hc
200
b
hc
200
b
hc
150
b
hc
9
p
Ch
80
p
hc
10
p
Ch
35
b
hc
50
a
Th
5
Ch
13
hc
5
hc
5
hc
10
Ch
60
hc
80
he
80
hc
60
hc
150
hc
80
hc
Clonel Clone2 El E2 C
0
0
0
0
0
0
0
0
0
0
0
0
0
DRi
Ogr
0
0
0
0
0
0
0
0
0
Rhiz2
Rhiz2
0
0
0
0
0
Node2
Rhizl
0
0
0
0
0
0
0
w Node2
0
0
0
Rhizl
0
0
0
W
h
h
h
h
h
h
h
w
w
h
h
h
h
h
h
h
h
sw
sw
w
h
h
h
sw
sw
h
h
h
h
h
sw
Origin
GB
IR
CI Tjan
Tjul
Prec Co Br Habitats
1221
12
1237
11
849
11
1244
11
978
11
707
13
1160
7, 8
760
1
890
1
796
17
989
6, 7
913
17
1099
1, 6
1177
8, 10
1211
12, 13
738
4
805
3, 16
797
7
979
16
2845
15
1120
8, 9
777
3, 4, 17
783
3, 4, 17
1686
15
1465
10
8 10
3, 4, 16, 17
790
3, 16, 17
863
3, 6, 17
863
3, 6, 17
729
1, 3, 16
898 Co 3, 18, 19
921
1, 3
1191
16
1584
7, 16
1151
3, 16
2311
15, 16
1422
11, 12
1337
12, 14
1269
12
1734
7, 16
1373
1, 16
1160
1, 16
1075
1
659
11 .
1114
1, 2
1772
1, 15, 16
5
7
4
4
5
5
2
1
71
7• 6
7
7
8
4
6
9
7
7
71
2
2
7
2
5
5
4
4
3
5
3
2
3
6
4
4
5
4
4
5
7
7
6
3
5
4
5
6
5
9
9
4
6
6
8
9
6
6
1
1
2
3
5
4
4
5
2
7
5
5
3
2
6
3
3
2
1
1
5
5
5
2
2
6
4
8
3
1
6
5
7
2
2
7i
6
5
5
9
4
2
5
6
8
3
4
7
7
7
71
6
5
5
4
4
0
0
0
1
F R N S
7 5
3 2
3 0
6 7 3
8 7 2 0
6 7 2 1
10 5 3 0
Taxon name
Dryopteris filix-mas
N
N
N
N
N
N
N
N
N
N
N
N
NE EN
N CR
N
N
Fam NS CS
Dry o N
N
N
N
N
AN
A R EN
N
N
N
N
N
N
N
N VU
N
N
N
AN
AN
N
N
N
N
N
N
N
AN
AN
N
n
n
n
n
n
n
n
n
s
n
s
n
s
n
r
x
n
n
n
n
n
n
n
n
,
n
n
s
n
r
n
n
r
n
n
n
F
n
n
x
s
n
0.39
0.27
0.32
-0.28
-0.0 1
-0.29
-0.4 1
-0.76
1.42
3.88
0.12
0.07
-0.39
0.38
-0.18
-0.4 1
-0.25
1.66
0.16
0.08
0.26
-0.39
0.19
-0.08
0.47
0.91
0.00
0.02
0.60
0.37
0.37
0.10
0.75
0.36
-0.24
1.07
0.66
-0.11
0.03
0.24
RS Chg
Hght Len
120
120
50
80
60
120
67
90
5 10
5 10
10
60
35
60
8
30
60
45
300
300
110
105
55
125
30
20
10
4
75
150
60
75
75
60
75
75
75
30
80
60
45
42
65
60
22
90
1 P2 LF1
hc
hc
he
hc
he
Th
hc
hc
p Hz
Hz
Hy
hc
hc
Hy
hc
hc
hc
Hy
Hy
Hy
he
he
he
hc
Ch
hc
he
he
hc
hc
hc
hc
hc
hc
he
hc
hc
Gn
Gn
Gn
Gn
Gn
Gn
Gn
Gn
Gn
1
4
3
4 c
6
3
2
5
3
3
6
1
4
3
3
4
6
3
6
5
2
3
3
6
3
4
4
4
5
3
4
3
3
1
5
6
9
7
7
5
5
4
7
6
7
5
7
8
5
8
8
6
2
1
1
8
9
7
7
5
7
7
7
5
7
7
7
7
7
4
3
Rhiz2
0
0
0
0
Rhiz2
Rhiz2
El E2 C
7 6
5 6
4 2
4 3
9 3
LF2 W Clonel Clone2
0
0
0
0
0
0
0
0
Nodel
0
Hy
Nodel
0
Rhiz2
he
Rhiz2
0
Rhiz2
hc
Rhiz2
Rhizl
Rhiz2
Irreg
Irreg
Irreg
0
Rhiz2
Rhiz2
Rhiz2
w Node2
Rhizl
Stol l
Node2
Ch
0
Rhiz2
0
0
0
Rhizl
0
0
0
0
0
NZ
Am
Am
Am
NHem
Origin
GB
2650
2650
308
1
33
370
79
1066
212
33
366
14
962
2577
15
1236
574
889
1696
808
1669
370
572
2530
1359
218
236
1226
2005
2036
371
2630
2347
2417
2070
937
1171
60
1218
86
447
134
235
10
8
2666
IR
928
928
5
1
0
4
0
41
79
19
89
0
330
756
5
267
95
286
424
15
122
41
180
837
236
1
0
423
400
795
0
894
758
812
824
24
0
13
16 1
0
160
9
0
0
0
921
CI Tj an
13 3.5
13 3.5
1.9
0
0 2.7
0 2.1
10 4.0
7 4.7
9 3.8
3 3.7
0 3.6
1 3.6
1.4
0
7 3.7
10 3.5
0 4.6
4 3.1
2 4.0
3 3.7
3.6
3
3 3.7
0 3.3
8 4.6
9 4.4
3.6
13
0 2.7
1.0
0
0 0.9
3.0
2
3.7
9
3.8
12
12 4.4
3.5
10
3.5
8
3.4
3
11 3.8
3.6
0
3.9
7
3.0
0
0 3.6
3.6
0
3.9
3
3.8
0
3.6
0
0 2.8
0 3.6
12 3.5
Tjul
14.5
14.5
12.6
13.4
13.7
16.0
15.9
15.6
14.2
14.71
15.2
13.2
14.0
14.5
15.3
13.5
14.4
14.2
15.1
15.8
14.9
15.9
14.5
14.6
13.2
11.9
11.6
13.6
15.0
15.0
16.0
14.5
14.5
14.3
14.9
15.5
15.9
13.8
15.3
15.7
15.3
16.0
16.3
14.3
16.1
14.5
Prec
1096
1096
1810
1998
1527
785
848
822
1365
1072
924
1348
1348
1098
1130
1323
1182
1259
936
836
981
863
1050
1046
1381
1832
1880
1357
972
971
937
1091
1104
1146
996
893
834
1328
946
808
9 15
774
728
820
721
1087
Br Habitats
1, 2
1, 2
15, 16
1, 2
16
17
4
7
13
13
13, 14
13, 14
11, 12, 13
11
Co 21
11
11
11
13, 14
13, 14
1, 3
Co 19, 21
Co 19
3, 4, 19
10, 12, 15
11, 16
11, 15
14, 15, 16
3, 4, 17
11, 14
3, 16, 17
3, 16, 17
11, 14
11, 14
11
1, 3, 14, 17
3, 17
16
1, 7
1
11
1
1
1
1
3, 4
Co
6 1
7
7
9
9
7
7
8
8
7
7
71
7
6
6
7
71
6
6
7
4
3
8
3
2
3
2
7
71
7
7
7
7
2
6
6
4
6
71
6
6
5
5
71
7
5
8
7
9
7
7
8
5
7i
6
6
4
3
4
6
7
8
6
6
7
1
4
3
3
6
71
5
6
5
3
5
7
5
1
4
4
3
4
L F R N
5 6 5 5
5 6 5 5
7 5 2 2
6 6 4 4
8 5 9, 3
6 5 7 8
9 3 5 5
8 4 7 4
7 10 5 4
71 10 5 5
7 10 7 5
8 9 5 5
8 9 4 1
8 10 6 4
6 9 71 5
9 9 7 2
8 9 7 4
8 11 4 2
Taxon name
NS
N
N
N
N
AN
N
N
N
AN
N
N
N
N
N
N
N
N
AN
AN
N
N
N
N
N
N
N
N
N
AR
N
N
N
N
AN
AR
N
AR
AR
N
N
N
AN
Euph AN
Euph AR
Euph AR
Fam
Equi
Equi
Equi
Equi
Equi
Equi
Equi
Equi
Ranu
Eric
Eric
Eric
Eric
Eric
Eric
Aste
Aste
Aste
Orob
Eno
Cype
Cype
Cype
Cype
Gera
Gem
Gem
Gera
Gem
Bras
Bras
Bras
Bras
Bras
Apia
Apia
Bras
Bras
Cela
Aste
Euph
Euph
VU
VU
VU
CS
-1.18
-0.77
80
20
40
RS Chg Fight Len
100
n 0.42
100
n 0.30
60
n 0.18
60
0.11
s
120
90
n -0.35
180
0.41
n
60
s -0. 12
15
1.59
60
r -0.11
60
n -0.94
120
o
60
o
60
n -0.9 1
80
r -0.07
50
n 0.33
20
r -0. 11
25
2.37
20
1.52
20
0.18
r
60
n -0.79
60
r -0.20
60
n 0.36
50
n -0.36
40
n
40
n -0.11
15
s
20
n 0.38
40
0.47
n
9
9
s
10
n
10
n 0.52
60
-0.02
75
-0.41
60
n -0.80
90
-0.65
60
1.05
n 0.15 600
150
n -0.15
70
n -0.22
40
0.98
hc
Th
Th
1 P2 LF1
Gn
Gn
Gn
Gn
Gn
Gn
Gn
Gn
Gn
Ch
Ch
Pn
Ch
Ch
Pn
b Th
Ch
Ch
Ch
Hy
hc
hc
hc
hc
Th
Th
Th
Th
Th
Th
Th
Th
Th
Th
hc
hc
Th
Ch
Ph
hc
Ch
hc
h Root
h 0
h 0
8
8
LF2 W Clonel Clone2 El
5
Rhiz2
Hy
5
Rhiz2
5
Rhiz2
4
Rhiz2
8
Rhiz2
5
Rhiz2
8
Rhiz2
2
Rhiz2
0
Pn w 0
8
7
Pn w
8
w
7
Pn w 0
7
Pn w Nodel
8
w O
hc
5
0
1
0
0
0
DRa
Rhizl
Rhiz2
Rhiz2
Rhiz2
Ogr
0
0
0
0
0
0
0
0
8
0
0
8
0
8
0
0
Pn sw
7
w Root
7
h Rhizl
7
h 0
7
h Root
Unk
Gard
Eur
3
5
Eur
3
3
3
3 c Eur
3
3
4
Origin
E2 C
6
6
6
6
Eur, As
5
6
3
6
Eur
1
1
1
0
2
1
6
4
SAm
Eur
0
6
6 c
3
6
4
4
2
2
1
IR
847
112
704
35
0
334
426
129
0
0
712
24
10
781
1
80
0
34
55
70
831
14
100
560
167
167
13
33
80
40
11
177
423
25
1
106
80
97
483
4 10
0
2
3.7 16.1
3.9 15.8
3.8 14.9
715
783
973
CI Tj an Tjul Prec
8 3.5 14.4 1116
0 3.0 14.0 1108
8 3.4 14.5 1091
1.8 12.7 1406
0
740
0 4.1 16.3
0 2.9 13.7 1269
950
3 3.8 15.2
3.4 14.0 1224
0
716
3.4 15.6
0
0 5.7 15.9 1011
13 3.5 14.0 1226
0 4.3 14.0 1395
0 4.0 13.7 1593
2 3.4 14.0 1230
0 6.0 15,5 1047
774
5 3.8 15.9
0 -1.2 10.6 1653
904
11 4.6 15.9
1 3.3 14.5 1121
4.4 14.1 1435
3.4 14.1 1194
4.5 15.4 1006
2.7 13.6 1394
3.1 13.6 1307
902
3.8 15.2
1
902
3.8 15.2
1
959
5.0 15.5
996
5.3 15.6
1
880
4.6 15.8
1
997
3.5 14.9
844
3.7 15.5
9 18
3.6 15.2
11 3,4 14.7 1011
809
0 3,9 15.9
856
2 4.6 16.1
1010
11 4.7 152
804
3.7 15.7
2
848
8 4.0 15.6
904
0 3.9 15.4
974
11 3.9 15.2
827
9 4.1 16. 1
838
2 3.6 15.6
0 1
169
1039 125 3
2114 642 12
GB
2494
238
2534
170
2
1561
1248
170
6 14
19
1999
0
0
1962
6
974
10
292
341
8
2134
17
575
1516
1666
1666
74
195
338
359
123
1074
2180
141
46
291
929
907
1254
1715
704
369
3
4
4, 17
Br Habitats
11
1, 11, 14
11
11
6
1, 16
11
11
1, 3, 17
10, 12
10
10, 13, 14
10, 12
10, 12
10
3, 16
15, 16
3, 16, / 7
3, 16
13
12
11
11
12
19
19
Co 19
Co 18, 19
3, 4
3, 16, 17
3, 16, 17
3, 16, 17
3, 16, 17
3, 16
3, 5
Co 19
3, 4
3, 16, 17
1, 3
11
1
3, 4, 7, 19
Co
L
6
71 5
4
6
1
2
2
1
4
4
2
6
5
3
3
3
3
7
3
5
7
5
5
7
6
3
5
4
7
2
6
6
7
6
6
7
7
6
7
7
6
6
7
8
8
6
6
7
8
9
9
8
4
4
4
4
4
3
3
3
3
4
3
4
5
4
5
8
5
3
4
5
1
2
1
1
1
3
2
2
2
1
1
2
2
2
4
7
7
7
8
4
R N S
6 4
3
7 6
6
5 4
8 5
5 5
7 6
8 3
7 6
7
5
6
8
8
6
5
5
3
3
11
F
10
7
8
7
4
8
8
8
5,
Taxon name
Euphorbra hyberna
Euphorbia lathyris
Euphorbia paralias
Euphorbia peplis
Euphorbia peplus
Euphorbia platyphyllos
Euphorbia portlandica
Euphorbia serrulata
Euphrasia anglica
Euphrasia arctica
Euphrasia cambrica
Euphrasia campbelliae
Euphrasia confusa
Euphrasia foulaensis
Euphrasia frigida
Euphrasia heslop-hani sonii
Euphrasia marshallii
Euphrasia micrantha
Euphrasia nemorosa
Euphrasia offi cinalis agg.
Euphrasia ostenfeldii
Euphrasia pseudokerneri
Euphrasia rivularis
Euphrasia rostko viana
Euphrasia rotundifolia
Euphrasia salisburgensis
Euphrasia scottica
Euphrasia tetraquetra
Euphrasia vigursii
Exact,/um pusillum
Fagopyrum esculentum
Fagus sylvatica
Fallopia convolvulus
Fallopia dumetorum
Fallopia j aponica
Fallopia sachalinensis
Festuca altissima
Festuca arenaria
Festuca armoricana
Festuca arundinacea
Festuca filiformis
Festuca gigantea
Festuca huonii
Festuca lemanii
Festuca longifolia
Festuca ovina
Fam
Euph
Euph
Euph
Euph
Euph
Euph
Euph
Euph
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Gent
Poly
Faga
Poly
Poly
Poly
Poly
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Poac
Poac
NS CS
N VU
AR
N
N EW
AR
AR
N
NA V U
NE
N
NE VU
NE
N
N
N
NE
NE
N
N
N
N
NE
NE
N
NE EN
N
N
N
NE V U
N
AN
N
AR
N
AN
AN
N
N
N
N
N
N
N
NA
N VU
N
,
RS Chg_ Hght Len P1 P2 LF1 LF2 W Clone l Clone2 El E2 C
Origin
r
52
p
hc
0
8 2
2.16
120
b
Ch
0
8 3
n -0.35
52
Ch
0
9 1
x -1.49
5
Th
0
9 1
-0.17
30
Th
0
8 3
-0.24
70
Th
0
8 3
n -0.09
35
Ch
0
8 /
r
1.20
65
Th
0
8 3
n
20
Th
0
7 1
n
30
Th
0
5 1
r
8
Th
0
1 1
r
10
Th
0
4 1
n
20
Th
0
5 1
s
6
Th
0
4 1
s
20
Th
0
1 4
r
15
Th
0
4 1
r
12
Th
0
4 1
n
25
Th
0
7 3
n
35
Th
0
7 3
n -1.6 1
30
Th
0
6 3
r
12
Th
0
2 1
s
20
Th
0
7 1
r
15
Th
0
4 1
n
35
Th
0
5 3
r
10
Th
0
4 1
o
12
Th
0
2 3
n
25
Th
0
4 3
n
15
Th
0
7 1
r
20
Th
0
7 1
o
8
Th
0
8 2
-0.53
60
Th
0
Asl ?
n -0.62 3000
Ph
sAf
7 3
-1.31
100
Th
0
6 4
s -0.33
200
Th
0
7 5
1.83 200
Gn
Rhiz2
As2
1.05 300
Gn
Rhiz2
As2
n 0.83
120
hc
0
7 3
S
75
hc
Rhiz2
7 1
o
40
hc
0
7 1
n
1.71
120
hc
0
8 4
n
35
hc
Ogr
7 2
n 0.46
125
hc
0
7 3
a
25
hc
0
7 0
r
66
hc
0
7 1
r
40
hc
0
7 1
n
43
hc
Ogr
5 5
2
956
182
26
1911
248
148
13
375
1095
5
11
970
143
118
20
39
941
1498
2600
85
167
14
29 1
4
0
601
313
33
0
501
2397
2139
69
2060
507
241
108
0
2082
821
1885
0
71
15
1788
GB
IR CI Tj an
137 0 5.2
2 1 13 3.8
65 8
5./
1 8
5.8
52 14
3.8
3.9
0
8 14
5.3
0 4. 1
5
0 4. 1
47
0
3.2
0 2.8
0 4.0
3.2
3
3
0
3.6
0
1.5
0 3.7
0 3.8
19
0 3.0
20
71 3.6
87 12 3.5
1 2.8
3.7
0
0
2.3
12
1 3.7
3.5
3
4.6
7
2.5
10 1
4.7
5.4
6.8
3.8
79
3.5
51
3.7
3.9
68 1
3.7
6
3.7
8
2.6
4.2
6.2
55
7 3.6
6
4 3.0
35
0 3.6
7 6.4
1 3.4
6 4.8
5
4
3.5
Tj ul Prec
14.9 1203
15.8
804
15.6
987
16. 1
969
15.1
950
16.3
733
15.4 1060
16 .3
8 12
15.3 1081
13.6 1315
13.2 2458
12.5 1638
13.8 1267
12.4 1263
11.8 1981
12.6 1695
12.3 1334
13.2 1400
14.8 1037
14.4 1131
12.2 1694
16.1
753
13.2 2450
14.3 1336
12.4 1004
14.7 1134
12.8 1575
14.9 1105
15.3 1263
16.7
7 12
15.8
788
14.7 1061
14.9
971
16.4
784
14.8 1043
15.3
976
13.6 14 10
15.4
828
16.9
796
14.9
980
14.3 1100
15.1
981
16.5
780
15.3 1079
16.2
751
14.5 1109
Co
Co
Co
Co
Co
Co
Co
Co
Br Habitats
1, 3
3, 16, 17
19
19
4, 17
4
18, 19
1, 3
10, 16
6, 11
15
10, 11
7, 8, 16
18, 2 1
15, 16
21
18
8, 10
7, 10
6, 7, 8, 10
16
7
15, 16
6
18
16
10, 15
6, 10
10
19
4
1
3, 4
1, 3
3, 14, 17
3
1, 16
19
19
6, 7
8
1
16, 18
7, 8, / 6
8
7, 8
9
8
3
7
6
6
6
3
8
8
8
8
5
8
8
8
7
8
7
8
8
71
8
9
7'
7
3
0
F R N S
5 5
4 0
5 5 7
6
7 5 3
9
7 5 3
9
0
7
7
0
8
3
5
0
71
0
7
0
8
0
8
0
8
0
8
1
81
0
8
3
8
7
L
La)
Taxon name
Festuca ovina agg.
Festuca pratensis
Festuca pratensis x Lolium
perenne (x Festulolium
loliaceum)
Festuca rubra
Festuca rubra agg.
Festuca vivipara
Filago gallica
Filago lutescens
Filago minima
.
Filago pyramidata
Filago vulgari s
Filipendula ulmaria
Filipendula vulgaris
Foeniculum vulgate
Fragaria vesca
Fragaria x ananassa
Frangula alnus
Frankenia laevis
Fraxinus excelsior
Fritillaria meleagris
Fuchsia magellanica
Fumaria bastardii
Fumaria capreola ta
Fumaria densillora
Fumaria muralis
Fumaria occidentalis
Fumaria offi cinalis
Fumaria parvil lora
Fumaria purpurea
Fumaria reuteri
Fumaria vaillantii
Gagea bohemica
Gagea lutea
Galanthus nivalis
Galega dffi cinalis
Galeopsis angustiffi lia
Galeopsis bifida
Galeopsis segetum
Galeopsis speciosa
Galeopsis tetrahit
Galeopsis tetrahit agg.
Galinsoga par Villora
Galinsoga quadriradiata
Garium aparine
Poac
Poac
Poac
Poac
Aste
Aste
Aste
Aste
Aste
Rosa
Rosa
Apia
Rosa
Rosa
Rham
Fran
Olea
Lili
Onag
Fuma
Fuma
Fuma
Fuma
Fuma
Fuma
Fuma
Fuma
Fuma
Fuma
Lili
Lili
Lili
Faba
Lami
Lami
Lami
Lami
Lami
Lami
Aste
Aste
Rubi
NH
N
N
N
AR
NA
N
AR
N
N
N
AR
N
AN
N
N
N
NA
AN
N
N
AR
N
NE
AR
AR
N
AN
AR
N
N
AN
AN
AR
N
AR
AR
N
N
AN
AN
N
EX
VU
EN
CR
VU
Fam NS CS
Poac
Poac
n
n
n
n
r
n
s
n
s
n
n
n
s
n
s
n
n
n
n
s
n
n
n
n
n
65
72
2.96
72
0.13
44
0.01
20
-0.34
25
-0.9 1
20
-1.14
30
-1.20
35
-0.10
120
-0.07
50
1.17 250
-1.09
30
0.42
40
-0.16
500
0.03
8
-0.73 2500
0.86
30
1.85
150
0.39
75
0.31 100
-0.37
52
1.75
100
0.04
100
-0.34
52
-0.55
471
0.25
60
-0.62
100
-0.5 1
42
9
0.16
25
3.01
22
150
-3.31
50
100
50
-1.82
100
100
-0.6 1
100
0.63
80
1.071 80
-0.09
150
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
a
hc
hc
hc
hc
Th
Th
Th
Th
Th
hc
hc Gn
hc
hc
hc
Ph
Ch
Ph
Gb
Pn
Th
Th
Th
Th
Th
Th
Th
Th
Th
Th
Gb
Gb
Gb
hc
Th
Th
Th
Th
Th
Th
Th
Th
Th
b
Otb
Otb
0
0
0
0
0
0
0
0
0
0
oI2
ol2
iz2
iz l
iz2
iz2
gr
DRa
DRg
DRg
DRg
DRi
7
9
9
8
8
7
8
8
7
8
7
8
7
8
7
7
5
7
5
5
5
7
8
7
7
7
7
9
8
5
7
0
7
3
3
2
Gard
Eur
Eur
Origin
SAm
1
2
3
1
1
3
3
1
1
Eur
4
3 c
3 c
3
Eur
3 c Eur
3
5
2
4
3
5
SAm
SAm
3
4
2
3
3
3
3
2
3
5
4
3
4
6
6
6
RS Chg_ Hght Len P1 P2 LFI LF2 W Clonal Clone2 El E2 C
n -0.15
45
p
hc
h Ogr
3 6
n -0. 16
80
p
hc
h O
5 4
790
2799
2799
801
21
85
788
132
980
2692
578
961
2380
740
712
58
2459
98
367
423
482
307
1200
31
1978
128
191
13
116
1
212
1763
352
6 16
1135
32
999
1725
2508
429
524
2672
80
974 1
974 1
182
0
0
105 1
0
78
967
8
60 1
819
20
56
0
930 1
0
542
330 1
180 11
16 0
368 13
0 0
304 12
0 0
37 2
0 1
0 0
0 0
0 0
36 5
0 1
43 0
138 1
01 0
84 0
409 1
623 4
2 9
5 6
960 14
3.5
3.5
3.5
2.7
3.8
3.71
3.6
3.8
3.8
3.5
3.6
4.1
3.5
3.7
3.8
4.4
3.6
3.6
4.3
4.5
4.2
3.6
4.0
6.5
3.6
3.6
4.1
5.4
3.5
2.9
3.0
3.5
3.7
3.8
3.6
3.7
3.1
3.6
3.5
3.8
3.8
3.6
15.5
14.5
14.5
12.7
16.4
16.4
15.1
16.3
15.5
14.5
15.8
15.9
14.6
15.5
15.8
16.5
14.7
16.3
14.3
14.8
14.9
15.5
14.9
15.8
15.0
16.1
14.8
16. 1
16.2
14.7
15.1
15.2
16.2
15.8
14.9
15.6
14.5
14.8
14.5
16. 1
16.0
14.6
851
6
1104
3, 5, 6, 2/
1104
6
1592
15, 16
647
3, 4
660
3, 4
904
16
687
4
832
3
1105
11
776
7
823
3, 16, 17
1091
1, 7
863
3, 17
881
1
680 Co 19, 21
1069
1
688
6
1223
3, 17
1086
4
997
3, 16
763
4
1004
4
1073
3, 4
9 12
3, 4
703
4
978
3, 4
958 '
3, 4, 17
702
4
1008
16
861
1, 3, 14, 16
919
1, 3, 17
725
3, 17
791
4, 16
1011
4
789
4
1043
4
1025
3
1073
3
721
4, 17
778
3, 4, 17
1080
3, 4, 17
GB
IR CI Tjan Tj ul Prec Co Br Habitats
2718 726 13 3.5 14.4 1118
7, 8
1976 569 4
3.6 14.9
985
6
5
3
4
5
4
4
4
3
2
6
6
5
2
5
4
5
5
5
4
5
6
8
4
4
5
5
5
8
8
6
8
6
4
4
6
8
6
7
8
6
6
8
5
7
7
7
8
6
3
7
6
6
6
6
7
6
8
8
6
6
5
8
7
7
5
6
6
5
3
4
4
7
2
6
3
3
4
6
5
5
6
5
5
5
5
2
7
7
8
4
6
3
7
6
6
7
6
8
4
5
2
5
4
7
5
5
6
4
5
6
71
2
2
2
3
71 6
6 5
6
5
L F R N S
7 5 4 2 0
7 6 6 6 0
Taxon name
Galiurn boreale
Galium conshictum
Galium mollugo
Galiurn odoratum
Gahm palustre
Galium parisiense
Galium pumilum
Galium saxatile
Galturn spurium
Galiurn sterneri
Galium tri comutum
Galium uliginosum
Galium verurn .
Gastridium ventricosum
Gaudinia fragilis
Gaultheria shallon
Genista anglica
Genista pilosa
Genista tind oria
Gentiana nivalis
Gentiana pneurnonanthe
Fan/
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Rubi
Poac
Poac
Eric
Faba
Faba
Faba
Gent
Gent
NS CS
N
N
N
N
N
NA
N
N
AN
N
AR CR
N
N
NA
NA
AN
N
N
N
N VU
N
n
r
n
r
s
n
n
$
s
n
RS
n
r
n
n
n
$
r
n
Chg li ght Len
-0.52
45
40
0.04
112
-0.62
45
0.07
75
-0.57
30
-1.32
35
-0. 15
30
-1.871 100
0.69
30
-4.78
50
-0.14
60
-0.85
75
-0.48
50
45
150
-1.09
50
-0.26
40
-0.77
60
15
-0.3 1
40
1 P2 LF1 LF2
hc
hc
hc
hc
hc
Th
hc
hc
Th
hc
Th
hc
hc
Th
Th
Pn
Ch Pn
Ch Pn
Ch Pn
Th
hc
W
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
w
wi
wo
w
Rhiz2
0
0
0
0
0
Clonel
Rhiz2
Rhizl
Rhiz2
Rhiz2
Rhizl
O
Nodel
Node2
O
Node l
0
Rhizl
Rhiz2
o
Clone2 El E2 C
Origin
5 6
9 1
5 3
7 3
5 4
9 2
7 3
7 2
7 6 c NHem
4 2
8 4
5 5
5 5
9 1
9 2
Am4
7 1
7 3
7 3
1 3
7 4
GB
561
12
1665
1836
2652
63
65
2489
55
273
386
1435
2516
159
47
179
858
23
932
4
130
IR CI Tjan
101 0 2.2
0 5 5.0
0 14
3.5
350 1 3.4
932 9 3.5
0 0 3.7
0 0
3.7
793 9
3.5
0 0
3.9
35 0
2.0
1 0
3.8
102 0
3.2
730 14
3.5
0 2 4.6
0 2 4.5
14 1 3.6
0 0 3.0
0 0
5.0
0 1 3.6
0
-0.5
0
3.7
Tj ul Prec Co Br Habitats
12.8 1580
7, 16
16.5
799
13
15.3
905
7
14.7 1086
1
14.5 1105
11
16.3
693
3
16.2
754
7
14.3 1148
8, 9
16.1
755
4
13.0 1457
7, 16
16.1
736
4
14.9
96 1
11
14.6 1045
7
16.3
839
4, 7
16.2
865
3, 6
14.9 1077
1, 10
14.6 1019
10
15.6 1011
10
15.6
880
6
11.2 1853
15, 16
15.7
806
8 , 10
L F R N S
7 5 7 3
8 9 3 2
7 4 7 4
3 5 71 6
7 9 5 4
8 3 7 2
7 4 8 3
6 6 3 3
71 5 8 5
9 4 7 1
7 4 7 4
7 9 6 4
7 4 6 2
9 3 8 2
8 5 6 6
3 4 4 3
8 5 3 2
8 5 4
8 6 71 2
9 5 7 3
8 7 4 1
3
V
la
Ta xo n na me
Fam
Hordelymus europaeus
Poac
Hordeum distichon
Poac
Hordeum distichon sensi at
Poa c
Hordeum j ubatum
Poa c
Hordeum marinum
Poac
Hordeum murinum
Poac
Hordeum secalinum
Poac
Hordeum vulgare
Poa c
Hornungia petraea
Bras
Hottonia palustris
Prim
Humulus lupu/us
Ca nn
Huperzia selago
Lyco
Hyacinthoides hispanica
Lili
Hyacinthoides non-scripta
Lili
Hydrilla vert icillata
Hyd r
Hydrocharis morsus-ranae
Hyd r
Hydrocotyle ranunculoides
A pia
Hydrocotyle vulgaris
A pia
Hymenophyllum tunbrigense Hym e
Hymenophyllum wilsonii
Hym e
Hyoscyamus niger
So b
Hypericum androsaemum
C lus
Hypericum calycinum
Clus
Hypericum canadense
Clus
Hypericum elodes
Clus
Hypericum hirsutum
Clus
Hypericum humil usum
C lus
Hypericum Iinadilolium
C lus
Hypericum maculatum
Clus
Hypericum montanum
Clus
Hypericum c o l oratura
Clus
Hypericum pulchrum
Clus
Hypericum tetrapterum
Clus
Hypericum undulatum
C lus
Hypochaeris glabra
Ast e
Hypochaeris maculata
Aste
Hypochaeds radicata
Aste
/beds amara
Bras
Ilex aquifolium
A qu i
Illecebrum vedicillatum
Ca ry
Impatiens capensis
Ba ls
Impatiens glandulifera
Bals
Impatiens noli-tangere
Bals
Impatiens pandfi ora
Ba ls
/nu/a conyzae
Ast e
Mula crithmoides
Ast e
NS CS
N
AC
AC
AN
N
AR
N
AC
N
N
N
N
AN
N
N
N
AN
N
N
N
AR
N
AN
AN
N
N
N
N
N
N
N
N
N
N
N
N VU
N
N
N
N
AN
AN
N
AN
N
N
RS C hg._ Hght Le n
s
0. 12
120
75
87
60
s -0.85
37
I -0.04
60
n -0. 19
70
100
s
0 .31
10
n -0 .63
120
n -0 .09
450
n -0 .41
10
60
L
n -0.4 1
50
r
100
n -0 .89
50
20 40
n -0 .53
20
n -0 .54
8
n -0 .87
10
, - 1.38
80
n
0.78
80
0.7 4
60
20
n -0 .46
20 40
n -0 .18
100
n -0 .40
10
r
0 .09
40
n
2 .11
60
n -0 .49
80
n
80
n -0.3 2
60
n -0 .41
60
s -0 .12
60
n - 1.0 1
20
r -0 .10
60
n
0.6 1
60
s - 1.2 1
35
n -0. 16 1500
r -0 .60
20
0 .71
60
1.85
20 0
s -0 .77
60
0 .10
100
n -0 .15
125
e
0 .09
95
P 1 P2 LF 1 LR2
p
hc
a
Th
a
Th
p
hc
a
Th
a
Th
p
he
a
Th
a
Th
p
Hy
p
hc
p
Ch
p
Gb
p
Gb
p
Hy
p
Hy
p
Hy
p
hc
p
hc
p
hc
b
hc
p
Pn
p
C h Pn
a
Th
hc Hy
hc
Ch
hc Ch
hc
hc
hc
hc
hc
hc
a
Th
p
hc
p
hc
a
Th
p
Ph
a
Th
a
Th
a
Th
a
Th
a
Th
p
hc
p
hc C h
w
h
h
h
h
h
h
h
w
w
sw
W
C lona l Clo ne2
0
0
0
0
0
0
0
0
0
brag
Rhiz 2
0
OM
Ot b
DRa
DRg
Stoll
lr reg
Nod e2
Rhiz l
Rhiz l
0
0
Rh iz2
0
Nod e2
0
0
0
Rh iz2
0
Rh iz2
Root
0
Stoll
Stol l
0
0
0
0
0
0
Nod a l
0
0
0
0
0
0
5
3
1
7
9
2
0
0
4
2
8
7
5
8
9
c
7
1
5
4
7
8
7
1
4
3
1
3
3
4
2
3
1
3
4 c
3
2
2
2
3
3
4
6
7
7
7
2
7
7
7
8
5
7
8
7
7
8
8
7
8
8
8
8
1
4
3
9
8
7
E l E2 C
7 3
AO
A m6
As l
Eur
Am
Am
Eur
Cro p
Cro p
C rop
As 2 , A m
Origi n
GB
185
7 14
7 14
349
146
1497
92 6
196
56
46 3
1273
989
847
243 9
2
323
43
20 9 1
197
577
796
1139
702
0
58 3
1276
1732
14
1180
26 9
1906
24 27
2 101
81
270
17
272 5
47 1
235 3
37 '
32 3
1599
21
470
860
12 1
1
76
76
6
0
48
30
8
0
2
0
23 4
76
76 0
1
84
0
842
112
176
87
74 4
63
3
260
10
42 8
0
30 0
0
38 5
887
842
0
5
0
97 7
0
86 1
0
1
286
0
1
0
19
IR
C I Tja n Tjul
0
3.2 15.5
3
3.8 15.4
3
3 .8 15.4
1
3 .5 15.4
1
4 .3 16.4
14
3 .8 15.6
0
3 .8 16.0
1
3 .8 15.6
3
3 .5 14.8
0
3 .6 16.0
8
3 .8 15.8
0
2.7 13.2
12
3.9 15 .5
13
3.6 14.6
0
3.5 14.2
2
3.9 15.8
0
4 .0 16.6
10
3 .7 14.4
0
3 .8 14.0
0
3 .1 13. 1
5
4 .0 15.7
8
4.0 14 .8
2
4 .0 15.6
0
4 .6 14.4
7
4 .2 14.8
0
3 .3 15.3
12
3 .6 14.8
9
5 .5 15.8
0
3.6 15.0
0
3.8 15.6
6
3.6 15. 1
9
3.5 14.3
11
3 .71 14.9
0
5 .6 15.5
14
4 .0 15.9
1
3 .9 16. 1
14
3.5 14 .5
0
3.5 16.3
14
3.6 14.7
0
5 .6 15.7
0
3 .7 16.2
6
3 .7 15. 1
0
2 .6 13.9
0
3.5 15.7
7
3.9 15.9
8
5. 1 16. 1
Prec Co Br Hab itat s
788
1
898
3, 4
898
3, 4
800
3, 5
720
6
8 17
3 , 17
742
5
83 3
3, 4
116 1
16
69 7
13
83 3
3
1472
15, 16
866
1, 3 , 17
1092
1, 9
1555
13
772
13
68 0
13, 14
1126
11
1567
16
1685
1, 16
799
4
1184
1, 3
884
3
1252
11, 14
1194
11
83 2
6, 7
1085
3
1069
16
1016
1, 3 , 16
86 7
1, 3
95 1
7
1148
10, 16
1025
11
1152
11
754
8
7 10
7
1104
6
7 10
7
1079
1
1062
3 , 13
72 3
13, 14
95 7
14
1833
1
82 1
1
82 1
7, 16
869 Co 18
L F
6 4
8 4
8 4
9' 6
9 6
8 4
8 6
9 4
9 2
7 11
6 7
7 6
5 4
5 5
6 12
7 11
7 10
8 8
4 6
5 5
8 4
S 6
5 71
8 9
8 10
6 5
7 6
71 3
6 6
7 4
7 4
6 5
71 8
8 8
8 4
8 4
8 4
71 4
5 5
8 7
7 9
6 8
4 71
4 5,
7 3
9 6
R N S
7 7 0
7 7 0
71 7 0
7 6 2
8 6 4
7 6 0
7 6
71 7
8 1
7 5
7 8
2 2
6 6
5 6
9 3
7 7
7 7
6 3
2 3
3 3
7 9
6, 5
5, 5
2 2
3 2
7 5
4 3
3 2
5 5
8 2
7 5
4 3
6 4
4 2
4 2
8 3
5 3
8 3
5 5
3 2
7 6
7 7
71 6
7 8
8 3
7 5
La
co
Taxon name
Mu la helenium
Mu la salicina
Iris foetidissima
Iris germanica
Iris pseudacorus
(sail s tinctoria
Isoetes echinospora
Isoetes histrix
Isoetes lacustris
Isolapis cernua
(solapis setacea
Jasione montana
Jugbans regia
Juncus acutiflorus
Juncus acutus
Juncus alpinoarticulatus
Juncus ambiguus
Juncus articulatus
Juncus balticus
Juncus bigIumis
Juncus bufonius
Juncus bufonius sensi at.
Juncus bulbosus
Juncus capitatus
Juncus castaneus
Juncus compressus
Juncus conglomeratus
Juncus effusus
Juncus filiformis
Juncus foliosus
Juncus gerardii
Juncus infi exus
Juncus maritimus
Juncus pygmaeus
Juncus squarrosus
Juncus subnodulosus
JUnCUS tenuis
Juncus trifidus
Juncus triglumis
Jun/peni s communis
Kickxia elatine
Kickxia spuria
Knautia arvensis
Kobresia sitnpliciuscula
Koeleria macrantha
Koeleria vallesiana
N
Ju nc
Junc N
Cupr N
Scro AR
Scro AR
Dips N
Cype N
Poac N
Poac N
NS CS
AR
N
N
AN
N
AR
N
N
N
N
N
N
AN
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N EN
N
N
AN
Fam
Aste
Aste
Ind
Ind
Irid
Bras
Isoe
lsoe
Isoe
Cype
Cype
Camp
Jugl
Junc
Junc
June
Junc
Junc
Junc
June
June
Junc
June
Junc
Junc
Junc
Junc
Junc
Junc
Junc
June
Junc
June
Junc
Junc
Junc
Junc
RS Chg_ Hght Len P1 P2 LF1 LF2 W Clonal Clone2 El
-0.80
150
p
hc
0
o
60
p
hc
Rhiz2
7
n
1.47
80
p
hc
Rhizl
8
95
p
hc
Rhizl
n 0.16
150
p
Gn Hy
Rhiz2
8
1.08
150
b p hc
0
n 0.65
15 p
Hy
0
4
r
4
p
hc
0
9
n 0.95
25 p
Hy
0
4
n 0.23
15
a p Th hc
0
9
n 0.53
15
a p Th he
0
7
n -1.08
50
hc
0
7
2400
Ph
w 0
n
1.16
100
he
Rhiz2
7
s
0.01
150
hc
Ogr
9
$ -0. 12
30
hc
Rhizl
4
n
17
Th
0
8
n
1.26
60
hc
IRhiz2
8
s -0.34
45
Gn
Rhiz2
2
s -0.17
12
hc
Ogr
1
n
25
Th
0
6
n
1.13
25
Th
0
6
n 0.34
30 90
hc Hy
Node2 Irreg
5
5
Th
0
8
s -0.40
30
he
Rhiz2
1
n -1.09
30
Gn
Rhizl
7
n 0.84
100
hc
Ogr
7
n
1.06
120
hc
Ogr
8
s
0.79
30
hc
Ogr
4
n
25
Th
0
8
n -0. 13
30
hc
Rhiz2
6
n 0.04
90
hc
Ogr
8
n -0.26
100
Gn
Ogr
8
r
8
Th
0
9
n
30
p
hc
Ogr
7
n 0.15
120
p
hc
Rhizl
8
0.83
40
p
hc
Ogr
n -0.38
30
p
hc
Ogr
1
n -0.38
20
p
hc
Ogr
1
n -0.42
500
p
Ph Ch w O
5
-0.18
25
a
Th
0
8
-0.071 25
a
Th
0
8
n -0.88
100
p
hc
0
7
r
0.58
20
p
he
0
1
n -0.29
50
p
hc
0
7
r
40
p
hc
0
8
4
6
6
3
3
4
6
6
2
3
1
6
3
4
6
6
6
6
3
3
6
4
3
3
6
2
6
4
3
1
2
3
6
1
4
1
4
3
3
5 c
2
E2 C
Am, SAm
Eur?, Asl ?
Eur, As
Gard
Origin
Eur, As l
GB
631
0
728
211
2563
102
178
3
522
242
2038
1076
803
2498
41
53
175
2740
92
37
1986
2736
2250
12
44
430
2622
2753
32
2 18
9 19
1758
390
4
1849
680
1053
177
200
1020
911
622
1707
18
1250
4
IR
100
3
0
1
959
2
32
0
147
2 15
638
420
11
845
28
0
50
956
0
0
724
937
827
0
0
4
798
974
0
88
271
631
196
0
484
224
145
0
0
145
41
0
478
0
266
0
CI Tjan
3 4.0
0 4.7
14 4.2
3 3.9
10 3.7
1 3.8
0 3.2
5 6.6
0 2.8
10 4.8
10 3.5
14 4.0
3 3.7
11 3.5
12
5.4
0 0.7
1 4.4
11 3.5
0
3.3
0 0.5
3 3.6
14 3.5
8 3.5
10 6.3
0 0.1
1 3.6
4 3.5
13 3.5
0 2.3
2 4.3
11 4.2
9 3.8
11 4.7
0 6.7
0
3.1
2 3.9
3 3.6
0
1.0
0 0.9
0 2.8
11 4.1
2 3.9
4 3.8
0 -0.1
7 3.7
0 4.5
Tjul Prec
15.1
973
14.9 1017
16.1
821
16.0
734
14.6 1081
16.0
733
13.0 1613
16.2
829
12.9 1660
14.8 1181
14.4 1149
14.8 1152
15.9
779
14.5 1115
15.7 1031
12.1 1477
14.9
999
144
1108
13.1 1012
11.4 2194
14.6 1102
14.5 1105
14.2 1180
16.1
889
11.2 2291
15.9
746
14.4 1117
14.4 1108
13.6 1530
14.7 1241
14.4 1174
15.2
925
15.1 1081
15.9
965
13.8 1254
15.5
838
14.6 1267
11.5 2162
11.8 2085
13.4 1380
16.0
813
16.2
737
15.2
919
11.5 2044
14.7
989
16.4
868
Co
Co
Co
Co
Co
Co
6, 7
11, 15
7
7
4
Br Habitats
3, 17
16
1
3, 17
11
3, 16
13
16
13
11
11, 14
8, 10
1, 3
11
19
11
19, 21
11
19
11, 15
3, 11, 13, 14
3, / , 13, 14
14
10
15
6, 11
11
8, 11
13
11, 13
21
6, 11
21
3
8, 12
11
1, 3, 13
15
11, 16
7, 10, 15, 16
4
1
1
1
F
R N
6 5
9 3
8 5
6 4
6 6
8 3
5 2
5 1
4 1
5 3
5 3
4 2
8 7
4 2
7 3
7 2
71 5
6 3
5 2
8 2
6 5
6 5
4 2
5 1
7 3
7 5
4 3
4 4
6 4
6 6
7 6
71 5
8 5
4 2
2 2
8 4
5,1 4
2 2
6 2
5 3
6 5
7 5
8 4
6 1
7 2
8 1
LO
•0
Taxon name
Fam NS CS
RS Chg
60
35
20
40
60
50
120
40
3000
3700
30
60
20
300
40
75
120
80
200
120
600
300
100
90
30
0.5
0.4
0.3
1
Hght Len
6
700
75
200
200
300
60
60
30
25
30
35
30
95
4600
b
a
Ph
Ph
Gn
Th
Gn
hc
Gn
Th
hc
hc
hc
Gn
Ph
Pn
Th
hc
Th
Hy
Hy
Hy
Fly
hc
hc
hc
Th
hc
p hc
hc
Th
1 P2 LF1 LF2 W Clonal
Th
Ph
Th
hc
hc
Hy
Ch
hc
Th
Th
Th
hc
Th
Th
Ph
Clone2 El E2 C
Origin
GB
IR
CI Tjan
Tj ul
1134
2, 17
1133
2, 17
955
1, 3
718
4, 7
886 Co 19
782
3, 18
1172
8, 15
742
3, 6, 7
823
11
1079
6
835
3
776
3
942
3, 17, 18, 19
1023 Co 18
875
3, 4, 16
766
13
7 17
4
739
13
986
11, 13
787
13, 14
832
11, 13
1103
6
918
7
960
7
801
3, 4 , 17
786
3, 19, 21
1074
3
629
6
757
3, 17
Prec Co Br Habitats
2175
11, 16
884
3, 17
601 Co 19
741
3, 17
693
3, 16
839
13
855
1
9 11
3, 17
842
3, 4
1097
4
877
4
869
3, 17
1027
3, 4, 17
1059
3, 17
1049
1, 2, 17
7j 6,
7 6
3 6
7 3
9 5
71 4
6 5
8 6
7 9
7 6
71 4
6 5
6 5
9 6
9 4
8 9
7 4
7 11
7 11
7 11
71 12
8 6
8 4
8 5
7 4
8 4
7 4
8 5
9 4
5
5
7
8
7
8
4
7
7
6
8
7
7
7
5
8
7
7
7
7'
71
6
7'
6
7
8
5
7
7
3
3
6
4
6
3
3
6
4
5
2
6
6
8
5
71
4
8
6
7
5
4
3
3
6
6
4
8 3
7'
L F R N S
8 9 6 1
6 5 7 7
8 4 7 6
8 5 7 6
8 4 7 7
6 12 7 6
4 5 7 6
7 5 7 8
7 4 7 6
7 5 7 7
71 5 7 6
5 6 7 8
6 5 7 7
6 4 7 7
7 4 6 3
1
Taxon name
Lob elia dortm ann a
Lobelia urens
Lob ularia maritima
Loiseleuria proc um bens
Lolium multifl orum
Lolium p e ren ne
Lolium te m ulentum
Lon ice ra p ericlym e num
Lonicera xylosteum
Lotus angustissimus
Lotus corniculatus
Lotus glab er
Lotus pe dun culatus
Lotus s ubb illorus
Lud wigia palustris
Lupinus arboreus
Lup inus polyphy l/us
Luron ium natans
Luzula arcuata
Luzula campestris
Luzula forsteri
Luzula multillora
Luzula pallidula
Luzula pilosa
Luzula spicata
Luzula sylvatica
Ly ch nis alpina
Ly ch nis flos-cuculi
Lychnis viscaria
Lycium
Lyciurn barbarum
Lycium ch ine nse
Ly cop e rsicon e scule ntum
Lyc op odie lla in un da ta
Lycopodium annotinum
Lycopodium clavatum
Lycopus europaeus
Ly sichiton am erican us
Lysimachia nem orum
Lysimachia nummularia
Lysimachia punctata
Lys ima ch ia thyrsillora
Lysimachia vulgaris
Lythrum hyssopifolium
Lythrum portula
Lythrum salicaria
N
Jun c
Cary N V U
Cary N
Cary N VU
Sola AN
Sola AN
Sola AN
Sola AN
Lyco N
Lyco N
Lyco N
Lami N
Arac AN
Prim N
Prim N
Prim AN
Ri m N
Prim N
Lyth AR VU
Lyth N
Lyth
N
NS CS
N
N VU
AN
N
AN
N
AR
N
AN
N
N
N
N
N
N
AN
AN
N
N
N
N
N
N VU
N
N
Fam
Camp
Camp
Bras
Eric
Poac
Poac
Poac
Capr
Capr
Faba
Faba
Faba
Faba
Faba
Onag
Faba
Faba
Alis
Junc
Junc
Junc
Junc
Junc
June
June
n
n
n
n
,
s
n
s
s
n
n
$
n
n
n
s
r
1
:
s
r
n
n
n
r
n
n
n
r
n
r
n
RS
n
r
'
n
:
n
Chg Hght Len P1
-0.05 4 p
-0. 19
70
p
2.34
30
a
-0.58
25
p
-1.06
100
a
-0.29
50
p
-4.05
90
a
-0.11 600
p
0.58 200
p
-0.23
30
a
1.09
40
p
-0.55
90
p
-0.06
60
p
0.22
30
a
0. 19
15
1.84
200
150
0.24
50
-0.43
10
-0.18
15
0.25
35
0.28
50
30
-0.35
32
-0.72
25
-0.02
80
20
-0.79
75
0.01
45
0.13 250
250
250
150
a
-0.65
5
-0.38
10
-0.52
15
-0.01
100
110
-0.46
20
-0.02
5
4.62
120
0.38
70
0.22
105
-1.12
15
0.32
8
-0.08
120
P2 LF1 LF2 W Clonal Clone2
Hy
h 0
hc
h 0
p Th hc
h 0
Ch
w Node2
b Th hc
h 0
hc
h 0
Th
h 0
Ph
w 0
Node2
Pn
w 0
Th
h 0
hc
h 0
hc
h 0
he
h 0
Rhizl
Th
h 0
p Th Hy
h Node2
Pn
w 0
hc
h 0
Hy
h Sto12
he
h Rhizl
hc
h Rhiz2
hc
h 0
hc
h 0
hc
h 0
hc
h 0
hc
h Rhizl
hc
h Rhizl
Ch
h 0
he
h 0
Ch
h 0
Pn
w Root
Pn
w Root
Pn Ph w Root
Th
h 0
Ch
h Nodal
Ch
sw Node2
Ch
sw Node2
hc
h Rhizl
hc
h Rhizl
Ch hc
h Node2
hc Ch
h Node2
hc
h Rhiz1
hc Hy
h Rhiz2
hc
h Rhiz2
Th
h 0
Th Hz
h 0
Nodel
hc
h 0
E/ E2 C
Origin
4 3
8 1
0 3
Eur
1 6
Eur
8 3
Eur
8 2
7 4
Eur, As1
8 3
8 5
8 3
7 3
8 2
8 3
Am4
Am4
7 2
1 3
7 3
9 2
3 6
5 5 c
5 4
1 3
7 3
2 3
7 4
7 4 c
As
As
As
SAm
5 3
2 6
5 6
7 4
Am4
7 2
7 3
Eur
4 6
7 5
8 4
7 3
7 5
523
233'
171
948
1689
174
2217
1266
1127
51
1227f
112
1262
1692
CI Tj an Tjul Prec Co Br Habitats
0 3.0 12.9 1635
13
0 4.7 16.1
988
8, 10
13 4.0 15.71 854
18, 19
0
0.9 11.4 1994
15
10
3.6 14.9
968
5
14
3.6 14.5 1096
3, 6
7 4.0 15.6
855
4
14 3.6 14.5 1102
1
0 3.4 15.2
874
1, 3
12
5.7 16.2
952
8
14
3.5 14.5 1104
6, 7
4 3.9 16. 1
732
3, 6, 7
13 3.6 14.7i 1072
11
14 5.8 16.0
987
8
3 4.6 16.6
812
13
11 4.2 15.7
825
3. 19
1 3.2 15.2
828
3, 14, 17
0 3.3 14.71 1229
13, 14
0 0.0 10.8 2044
15, 16
14
3.5 14.5 1100
6
5 4.2 16.2
859
1
7
3.5 14.3 1148
8
0
3.4 16.3
553
1, 11
1 3.2 14.4 1127
1, 2
0
1.0 11.5 2085
7, 15, 16
4
3.3 14.1 1208
1, 16
0 0.5 12.0 1640
15
9 3.5 14.5 1095
11
0 2.2 13.8
926
16
9 3.8 15.7
784
3, 19
3, 19
3, 19
41
4.0 15.6
876
17
18
3.5 14.9 1122
10, 12
0
0.7 11.7 1760
10, 15
81
2.4 13.3 1368
10
347
3.8 15.2
995
11
24
3.9 15.1 1073
13, 14
740
3.4 14.4 1150
1
227
3.7 15.5
895
6, 11, 14
48
3.5 15.1
999
1, 3, 17
0
2.7 14.2 1200
11, 13
288
3.71 15.2
943
11
3
3.9 16.0
777
4
321
3.8 14.8 1084
11, 13
827
4.0 15.0 1024
11
GB
IR
570 , 178
12
0
738
33
182
0
2069: 340
2743 964
341 37'
2622 943
242
16
55
0
280 1 975
509
0
2380 729;
861 8
11
0
341 21
2 15
3
91
3,
22
0
2725 835
309
0
245 1 834
2
0,
2 132 266
189
0
2058 621
2
0
2569 765
28
0
1104
68
L F
8 12
8 8
9 3
9 5
7 5
8 5
7 4
5 6
5 5
8 3
7 4
7 7
7 8
7 5
8 9
9 4
7 5
8 11
9 5
7 4
4 4
7 6
7f 7
5 5
8 5
5 5
8 3
71 9
8 3
8 5
8 5
8 5
71 5
9 9
6 6
7 5
7 8
4 9
5 7
5 7
6 6
8 10
7i 9
8 6
8 9
7 9
R N
5 1
4 2
71 4
2 2
7 7
6 6
8 7
5 5
7 6
4 3
6 2
7 5
6 4
6 5
4 4
7 3
5 5
5 3
2 2
5 2
5 2
3 3
5 2
5 3
3 2
4 4
4 2
6 4
4 2
7 4
7 4
7 4
71 8
2 1
3 3
1 2
7 6
6 8
4 5
5 5
7 5
4 3
71 5,
6 4
5 3
7 5
Taxon name
Mahonia aquifoliurn
Maianthemurn bifolium
Malus domestica
Malus sylvestris sens.lat.
Malus sylvestris sens.str.
MaIva moschata
Ma Iva neglecta
Melva pusilla
Ma Iva sylvestris
Marrubium vulgere
Matricaria discoidea
Matricaria recutita
Matteuccia struthiopteris
Matthiola incana
Matthiola sinuata
Meconopsis cambrica
Medicago arabica
Medicago lupulina
Medicago minima
Medicago polymorphe
Medicago sativa
Medicago sativa
subsp falcata
Medicago sativa subsp.sativa
Melampyrum arvense
Melampyrum cristatum
Melampyrum pratense
Melampyrurn sylvaticum
Merica nutans
Merica uniflora
Mehlotus elbus
Mefi lotus altissimus
Melilotus indicus
Melilotus offi cinalis
Melissa officinalis
Melittis melissophyllum
Menthe aquatica
Mentha arvensis
Mentha pulegi um
Mentha spicata
Menthe suaveolens
Menyanthes trifoliata
Mercurialis annua
Mercurialis perennis
Mertensia maritima
Mespilus germanica
N
AN
AN
N
N
N
N
N
AN
AR
AN
AN
AN
N
N
N
N VU
AR
N
N
AR
N
N
AR
Faba
Faba
Scro
Scro
Scro
Scro
Poac
Poac
Faba
Faba
Faba
Faba
Lami
Lami
Lami
Lami
Lami
Lami
Lami
Meny
Euph
Euph
Bora
Rosa
Papa
Faba
Faba
Faba
Faba
Faba
Bras
NS CS
AN
NA VU
AR
N
N
N
AR
AN
AR
N
AN
AR
AN
AN
NA VU
N
N
N
N
N
N
Fam
Berb
Lili
Rosa
Rosa
Rosa
Maly
Maly
Maly
Maly
Lami
Aste
Aste
Wood
Bras
n
s
$
n
s
n
n
r
r
n
r
n
n
s
-0.49
-0.88
-0.88
-0.58
-0.17
-0.04
-0.20
0.73
-1.59
0.02
1.73
-0.47
-0.11
-1.30
-0.70
1.69
-0.32
-0.04
0.28
-0.65
-0.53
60
90
60
50
60
35
60
60
150
150
40
150
60
60
90
60
30
90
100
30 150
50
40
60
900
RS Chg_ Hght Len
150
1.61
20
0.32
r
1000
0.57 1000
n
1000
n
80
n -0.04
60
-0.22
50
150
-0.30
60
s -2.02
35
-0.49
60
0.92
60
80
0.75
60
r
60
2.36
s
60
n 0.69
50
n -0.43
20
s -1.97
60
s -1.34
90
s -0.56
a
a
a
a
hc
hc
Ph
Th
hc
hc
Th
Th
Th
Th
hc
hc
a hc Th
hc
Th
hc
hc
hc
hc
hc
hc
hc
hc
Gn Hy
Rhizl
0
0
0
0
0
Rhiz1
Rhir l
0
0
0
0
Rhizl
Rhizl
Rhiz2
Rhiz2
Rhiz2
Rhiz2
Rhiz2
h Rhiz2
h 0
w o
h Rhiz2
h 0
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
1 P2 LF1 LF2 W Clonel
w Rhiz2
Pn
h Rhiz2
Gn
Ph
w 0
w 0
Ph
w 0
Ph
hc
h 0
h 0
Th
h 0
Th
a
he
p
h 0
h Rhizl
hc
p
h 0
Th
a
h O
Th
a
h O
hc
p
sw O
Pn
p
h O
hc
b
h O
hc
p
h O
Th
a
h o
a p Th hc
h O
Th
a
h o
Th
a
hc
p
h 0
Rhir l
3
4
4
3
5 c
3
3
3
3
6
3
2
6
2
3
3
7
7
5
4
5
7
7
7
7
5
8
9
5
9
7
2
Eur
Crop
Eur, As 1
Eur?, Asl ?
Eur
Eur?, Asl ?
Crop
c Eur
c
4 c
8
7
7
7
7
7
8
8
8
4
9
9
5
9
7
8
9
8
5 0
Origin
Am4
Gard
3
3
3
3
5 c Eur, As
4
4
As2?, Am?
3
6 c NHem
Eur
1
1
1
2
4
4
2
4 c
5
Clone2 El E2 C
01
17
0
0
323
20
0
246
15
16
12 1
26
36
0
922
428
45
74
0
756
55 1
4
29
1
541
1065
50
62
1696
75
408
1511
913
1122
427
1142
667
119
2475
1965
242
1563
118
1905
793
2214
222
98
16.2
15.8
16.2
16.3
14.3
12.5
13.0
15.0
15.8
15.8
15.7
15.7
15.9
15.7
14.7
14.9
15.9
15.0
15.5
14.1
16.0
14.8
13.4
16.1
3
604
3, 6
768
3, 4, 16
692
3
606
1, 2
1220
1, 16
1532
1, 7, 16
1483
1
1015
3
778
3, 17
785
3, 17
790
3, 17
790
3, 17
854
1, 3
1091
11
1065
4, 11
10 17
6, 13
869
3, 17
942
3
1151
11
1201
3, 4, 17
779
1
1045
1083 Co 19
1, 3, 17
861
Tjul Prec Co Br Habitats
1, 3
786
15.6
1, 2
753
15.0
3
975
15.0
1, 3
975
15.0
3
964
15.1
6
871
15.6
3
781
15.7
3, 17, 19
788
16.0
3, 17
899
15.3
7
861
16.1
3, 4
14.5 1086
3, 4
868
15.5
1
14.7 1108
874 Co 18, 19
15.9
15.6 1089 Co 18, 19
1
14.0 1321
3, 17
781
16.2
7, 17
963
15.0
8
623
16.5
8
801
16.4
3, 6
604
16.2
3.51
3.8
3.6
3.4
3.2
1.5
1.9
3,5
3.7
3.8
3.8
3.7
4.1
5.1
3.7
3.6
4.4
3.5
5.4
3.4
4.0
3.2
3.8
4.4
CI Tjan
2 3.4
0 2.9
8 3.7
11 3.7
3.7
0
10 3.7
14 3.8
1 4.1
14 3.8
0 4.6
3.6
14
11 3.7
3.2
5.0
5.4
3.5
4.2
1
3.8
1
3.9
5.0
1
3.5
4
0
327
598
218
0
70
6
354
0
945
51
4
2
8
77
0
686
0
0
0
IR
GB
991
4
1532
2023
1335
1423
1196
107
1788
46
2677
1588
74
107
19
54
744
2064
54
118
54
L
3
4
4
3
5
5
5
5
3
6
5
5
5
4
8
7i
7
5
8
10
5
6
5
4
3
5
3
2
3
2
3
5
4
7
71
5
6
5
5
6
7
7
6
3
7
7
7
6
7
6
8
8
2
2
7
7
7
7
71
7
7
7
7
71
5
7
6
4
7
7
T
6
F R N S
4 6 5
5 3 3
5 6 7'
5 6 6
5 6 6
3 7 4
4 8 71
5 5 5
4 8 7
5 7 8
5 7 7
5 7 7
8 7 7
3 5 2
3 7 2
5 7 5
5 6 5
4 8 4
3 7 2
4 5 5
4 6 5
-h.
to
Taxon name
Meum athamanticum
Mibora minirna
Milium effusum
Milium vernale
Mimulus
Mimulus guttatus
Mimulus guttatus x luteus (M
x robert sin
Mimulus luteus
Mimulus moschatus
Minuartia hybrida
Minuartia recurva
Minuartia rubella
Minuartia sedoides
Minuart ia stricta
Minuartia verna
Misopates orontium
Moehringia trinervia
Moenchia erecta
Molinia caerulea
Moneses uniflora
Monotropa hypopitys
Montia fontana
Muscari neglect um
Mycelis muralis
Myosotis alpestris
Myosotis arvensis
Myosotis discolor
Myosotis taxa
Myosotis ramosissima
Myosotis scorpioides
Myosotis secunda
Myosotis sicula
Myosotis stolonifera
Myosotis sylvatica
Myosoton aquaticum
Myosurus nnnimus
Myrica gale
Myriophyllum alternifl orum
Myriophyllum aquaticum
Myriophyllum spicatum
Myriophyllum verticillatum
Mynt is odorata
Naj as flexilis
Naj as marina
Narcissus pseudonarcissus
NS CS
N
N
N
N
AN
AN
AN
AN
AN
N
N
N
N
N EN
N
AR
N
N
N
N VU
N
N
NA VU
N
N
AR
N
N
N
N
N
N
N
N
N
NA
N
N
AN
N
N
AN
N
N VU
N
Fam
Apia
Poac
Poac
Poac
Scro
Scro
Scro
Scro
Scro
Cary
Cary
Cary
Cary
Cary
Cary
Scro
Cary
Cary
Poac
Pyro
Mono
Port
Lili
Aste
Bora
Bora
Bora
Bora
Bora
Bora
Bora
Bora
Bora
Bora
Cary
Ranu
Myri
Halo
Halo
Halo
Halo
Apia
Naj a
Naja
Lili
0.01
-0.75
-1.70
-0.42
-0.89
n -0.40
n -0.65
n -0.34
r
0.14
n -1.09
n 0.14
r
1.55
n 0.01
r -0.22
-0.34
n 0.14
n 0.65
n 0. 11
n -0.77
n 0.52
o
s
0.77
n 2. 18
n 0.00
n -0.66
n -0.75
n
1.00
,
n 0.63
n -0.89
-0.25
s
0.48
r
n 0.87
s
o
r
s
r
s
35
180
50
50
40
20
5
6
8
10
15
50
40
12
130
4
30
20
30
100
25
40
25
40
25
57
55
15
20
47
100
8
150
30
97
120
200
250
300
p
p
p
a
p
p
p
p
p
a
a
a
p
p
p
a
p
p
p
a
a
b
a
p
p
a
p
p
p
a
p
p
p
p
p
p
a
a
p
hc
hc
hc
Th
Ch
Ch
Ch
Ch
Ch
Th
Th
Th
hc
hc
Gn
p Th Hy
Gb
hc
hc
Th
Th
hc
Th
hc Hy
hc
Th
hc
hc
hc
Th
Pn
Hy
Hy
Hy
Hy
hc
Hz
Hz
Gb
6
6
2
5
6
7
7
4
8
8
2
3
6
3
6
5
4
3
2
4
6
6
3
4
3
6
4
3
6
3
4
1
1
1
5
4
3
2
2
9
1
1
1
1
4
8
7
8
5
4
7
5
8
7
1
5
7
5
8
7
7
9
4
7
7
7
5
5
c
Clone2 El E2 C
4 3
8 2
5 6
9 1
Node2
Node2
Node2
0
0
0
0
0
0
0
0
0
Ogr
Rhizl
Rhizl
Nodal
Gi b
DRg
0
0
0
0
0
0
Stol l
Stol l
0
Stoll
0
Nodal
0
Rhiz2
Irreg
Irreg
Irreg
Irreg
DRa
0
0
0
Otb
RS Chg. Hght Len P1 P2 LF1 LF2 W Clonal
s -0.40
60
p
hc
0
r -0.01
8
a
Th
0
n 0.31 150
p
hc
0
0
5
a
Th
0
-0.47
50
p
hc
Node2
50
p
hc
Node2
Eur
SAm
Gard
SAm
Am4
Am4 , SAm
Am4
Origin
473, 125
186
5
361
13
294
0
0
1
71 0
76'
0
1
0
139
30
488
24
1990 259,
418
0
2244 897
27
0
288 24
21971 547,
13
0
1302
0
7
0
2577 748
23171 521
2409 757
1174 52
229 1 662
1736 498
0
0
115
0
1690
24
927
0
339
0
976, 553
1390 327
268
2
1409 373
360 130
1152 147,
28 28
4
0
646
0
1162
1144
1099
716
1414
1679
2095
1443
1215
864
982
823
1179
985
817
1198
590
944
1623
1056
1088
1085
823
1037
1237
794
1362
931
747
691
1353
1299
838
907
765
1084
1338
600
920
13, 14
13, 14
13, 14
7
15
15
15
11
7, 16
4
1
8
12
2
1
11
3, 8
1, 16
7, 15, 16
3, 4
6
11
8, 16
11, 14
11
19
11, 14
1
11, 13, 14
4
12
13, 14
13
13, 14
13, 14
3, 17
13
13
3
Tj ul Prec Co Br Habitats
12.9 1459
3, 6
16.1
861 Co 19
15.3
928
1
16.6
7 18
19
14.3 1106
13, 14
14.5 1020
13, 14
0
3.1 14.0
0
2.8 13.6
2
3.3 14.6
1 3.5 16.0
0 4.8 14.2
0 0.8 11.2
0
1.2 11.4
0
0.4 12.1
0
2.7 13.9
11 4.3 15.9
4
3.5 15.0
14 4.2 16.0
6 3.5 14.2
0
2.0 13.0
0
3.7 15.8
14
3.4 14. 1
0
3.3 16.3
0 3.3 15.2
0 -0.1 11.6
8 3.5 14.6
14
3.5 14.4
8
3.5 14.5
14
3.8 15.5
3 3.5 14.7
6 3.3 14.0
2 6.2 17.0
0
1.6 13.2
4
3.4 15.1
0
3.6 16.0
2 3.8 16.3
0 3.4 13.8
3 3.2 13.8
7 4.3 16.1
7
3.7 15.1
0
3.8 15.7
0 3.0 14. 1
0 4.1 13.8
0 3.9 16.1
3 3.7 15.6
GB
IR CI Tja n
164
0 0
1.6
7,
0 10 5.8
1391 91 0
3.6
0
0 2 6.8
1767 237 5 3.2
996 44 4
3.2
7 8
7 9
7 8
9 3
8 4
8 4
8 5
9 9
8 4
7 5
4 5
9 4
7 8
4 5
4 5
71 9
7 3
4 5
8 4
7 5
7 5
7 9
8 3
7 9
6 9
6 7
8 9
6 5
7 8
8 7
8 9
7 12
7 12
7 12
7 12
7 6
6 12
5 12
7' 5
7i
5
5
8
3
7
4
5
7
6
7
4
3
4
6
5
7
7
8
6
5
6
6
6
5
6
5
7
7
6
3
5
5
7
7
7
7
9
6
5
5
5
3
1
1
2
2
1
6
6
3
2
1
2
3
5
5
2
6
3
5
3
6
4
3
4
5
8
5
2
3
3
7
7
71
4 1
6 0
5 0
L F R N S
8 5 4 3
9 3 71 1
4 5 6 5
9 3 6 2
7 9 6 5
7 9 6 6
Orchis laxiflora
Orchis mascula
Orchis militaris
Omhis morio
Orchis purp urea
Orchis simia
Orchis ustulata
Oreopteris limbosperma
Origanum vulgare
Ophrys sphegodes
Taxon name
Nardus stdcta
Narthecium ossifragum
Neotinea maculata
Neottia nidus-avis
Nepeta cataria
Nuphar lutea
Nuphar pumila
Nymphaea alba
Nymphoides pe/tata
Odontites vernus
Oenanthe aquatica
Oenanthe crocata
Oenanthe fistulosa
Oenanthe fluviatilis
Oenanthe lachenalii
Oenanthe pimpinelloides
Oenanthe silaifolia
Oenothera
Oenothera biennis
Oenothera biennis x
glazoviana (0 . x fallax)
Oenothera cambrica
Oenothera glazio viana
Onobry chis
Orionis reclinata
Orionis repens
Ononis spinosa
Onopordum acanthium
Ophioglossum azoricum
Ophioglossum lusitanicum
Ophioglossum vulgatum
Ophioglossum vulgatum
sens.lat
Ophrys apifera
Ophry s fuciflora
Ophrys insectifera
N
N
N
N
N
N
N
N
Orch
Orch
Orch
Orch
Orch
Orch
Thel
Lami
VU
VU
VU
n
sr
n
nr
s
o
s
n
n
nr
n
-0.10
-1.77
-0.18
-0.98
-0.56
-Oi l
-0.72
0.72
0.83
-1.34
65
15
30
90
20
45
50
50
20
40
30
35
45
60
200
70
10
4
30
N
N
N
0.66
-0.82
Ophi
Orch
Orch
n
s
r
n
AR
N
N
N VU
N
0.27
-0.76
-0.45
Aste
Faba
Ophi
Ophi
Ophi
nr
n
100
80
100
15
60
60
AX
AN
AN
N
NA VU
N
100
100
100
150
100
80
Onag
Onag
Onag
Faba
Faba
0.48
-0.36
0.37
1.02
-0.04
0.19
-1.18
100
n
s
n
n
150
50
Onag AN
Apia N
Apia N
Onag AN
N
N
-0.13
0.87
1.02
2.81
-0.35
-0.46
150
150
150
200
Apia
Apia
n
s
n
s
n
N
N
N
N
N
Nymp
Nymp
Nymp
Meny
Apia
Scro
47
30
100
n
x
N EX
AR
Orch
Lami
-0.9 1
-1.23
RS Chg_ Hght Len
n -0.32
45
-0.68
40
Fam NS CS
Lili
N
Poac
a
he
Hy
Hy
Hy
Ch
Gn
hc
Gn
Gn
Gn
Gn
Gn
Gn
Gn
hc
hc hc
Ch
Gn
Gn
Gn
hc
he
he
Th
he
Ch
hc
hc
hc
hc
Hy
Hy
Hy
Hy
Th
p Hz
he
Hy
hc
Gn
he
0
0
0
0
0
0
0
Root
0
0
sw 0
Root
Root
Root
0
0
0
0
sw Rhiz2
0
0
0
0
Nodal
Rhizt
Rhizl
Rhizl
Irreg
Rhiz2
0
0
0
Node2
0
0
Rhizl
8
7
8
7
7
7
8
9
7
7
9
7
7
8
5
9
7
9
7
7
9
8
8
5
4
7
7
7
7
8
7
7
9
7
5
3
3
3
4
3
4
2
3
6
2
3
4
2
1
6
1
4
3
1
2
3
4
6
3
5
5
4
2
1
3
4
1
4
c
c
1 P2 LF1 LF2 W Clona l Clone2 El E2 C
he
Rhizl
1
hc
0
5 3
Am?
Am
Am?
Am, SAm
Origin
0
1
452 0
0 0
340 5
0 0
865 7
183 0
633 12
45 0
101
6
99
24
6
9
2
1148
265
10
1585
179
0
138
931
19 124
0
36
0
0
0
62
1962 475
1474 209
6 182
0
936
264 31
778
5
724
0
72
12
1
0
1474 209
0
0
0
8
0
1
8
1
9
125
2 1
217
0
966 24 1
10
0
265
1664 178 1
614
241
0 0
551 118
3
76
0 0
1185
38 11
1140
68
1511
45
2320
505
1599
235
804
7421
478
15.1
12.5
14.8
16.4
14.7
15.6
14.8
16.0
15.8
15.9
16.0
13.3
16.2
15.1
15.9
15.9
15.9
15.8
16.2
15.3
3.7 15.3
3.5 16.4
15.7
3.8
2.8 13.7
3.9
3,4 15.8
16.3
4.0 16.5
6.4
4.0 16.7
16.3
3.5 14.7
3.6 15.1
4.0 15.7
16.4
3,9
3.7 15.8
3.7
3.6
4.3
6,7
3.6
3.7
4.2
4.0
5,6
3,6
3.8
3.9 15.9
4.4
4.3 16.2
15,4
3.7 16.3
3.9 15.8
3.6
17
3.6
3.5
3.7
3.8
4.0
3.7
3.9
3.5
4.7 15.2
14.8
3.7 15.9
3, 16, 17, 19
6
11
6
3, 16, 17, 19
13, 14
13
13
13, 14
6
11
11, 14
14
11
1 16
7,
3, 7
898
777
697
1323
801
694
748
800
729
1073
964
766
807
803
733
718
1252
848
964
7, 16
7
1, 16
6,
7 7
1
6,
7 11
1, 7, 16
6
7
1. 7, 11
3, 7
17
6,
8
8
6, 7
798
3, 16, 19
921
3, 16, 17, 19
832
3, 16, 19
999
710 Co 718
878
7
777
849
957
661
839
975
1680
1090
603
1047
792
1124
718
771
938
1107
761
GB
IR CI Tj an Tj ul Prec Co Br Habitats
16281 560
7 16 0
3.3
205
3 3
2 13.8
14.0 1296
1226
812
L
4
4
3
6
4
3
4
9
4
5
7
4
5
4
6
6
7
3
4
4
2
4
4
4
7
8
9
4
11
11
11
11
5
10
' 9
10
9
4
4
7
8
4
7
9
8
8
9
7
7
9
8
9
6
8
5
5
7
6
6
6
8
6
6
6
8
7
6
7
6
6
7
6
7
6
8
7
7
8
7
4
2
3
3
2
3
2
3
4
3
2
3
2
7
3
2
2
3
5
3
5
2
3
3
4
3
5
5
4
6
4
4
6
5
6
7
6
5
2
6
0
1
0
0
0
01
0
0
0
3
0
0
0
0
0
0
F R N S
9
2 21 0
7 3
Taxon name
Ornithogalum angustifolium
Ornithogalum pyrenaicum
Ornithopus perpusillus
Ornithopus pinn atus
Orobanch e alba
Orobanche artemisiaecampestris
Orobanch e caryophyllacea
Orobanch e elatior
Oroba nch e he derae
Orobanche minor
Orobanche purpurea
Oroba nch e rapum-genistae
Oroba nch e reticulata
Odi nha s ecun da
Os m un da regalis
Otanthus maritimus
Oxalis acetosella
Oxalis art iculata
Oxalis corniculata
Oxalis de bilis
Oxalis exilis
Oxalis incarnata
Oxalis latifolia
Oxalis pes-caprae
Oxalis stricta
.
Oxyria digy na
Oxytropis campestris
Oxytropis ha il ed
Panicum mill aceum
Papaver argemone
Papaver du bium
Papaver hy bridum
Papaver rhoeas
Papaver s om nife rum
Parapholis incurva
Parapholis strigosa
Parentucellia viscosa
Parietaria j udaica
Paris guadrifolia
Parnassia palustris
Pastinaca sali va
Pedicularis palustris
Pedicularis sylvatica
Pentaglottis sempervirens
Persicaria amphibia
N
N
N
N
N
N
N
N
N
N
N
N
AN
AN
AN
AN
AN
AN
AN
AN
N
N
N
AC
AR
AR
AR
AR
AR
N
N
N
N
N
N
N
N
N
AN
N
Orob
Orob
Orob
Orob
Orob
Orob
Orob
Orob
Pyro
Osmu
Aste
Oxa
Oxa
Oxa
Oxa
Oxa
Oxa
Oxa
Oxa
Oxa
Poly
Faba
Faba
Poac
Papa
Papa
Papa
Papa
Papa
Poac
Poac
Scro
Urti
Lili
Saxi
Apia
Scro
Scro
Bora
Poly
VU
EX
VU
EN
VU
NS CS
AN
N
N
N
N
Fam
Li li
Li li
Faba
Faba
Orob
0.16
-1.79
0.23
-0.35
-0.41
2.54
0.09
0.14
0.64
0.08
-0.68
-0.84
-0.39
-0.88
-1.28
1.81
0.27
.
s,
n
n
n1
n1
n;
ni
n
n
,
n
-0.09
-0.7 1
1.62
-0.40
0.56
-1.49
-0.74
0.01
-0.33
0.20
-0.20
0.50
-0.35
n
r.
r
r
r
n
n
n
r
s
r
n
n
x
n
60
40
75
60
60
45
85
60
5
160
30
10
25
15
20
5
20
20
30
40
30
20
13
100
45
60
50
60
100
10
25
50
52
40
20
180
60
25
100
60 200
RS Chg Hght Len
: 1.05
30
s
0.14
75
n -0.18
10
r
8
s -0.38
25
b
b
P
p
p
p Th
a Gn
Gn
a Gn
p Th
a Gn
Gn
a Gn
Ch
hc
hc
hc
hc
a hc
Gb
a hc
Gb
Gb
Gb
Th
hc
hc
hc
Th
Th
Th
Th
Th
Th
Th
Th
Th
hc
Gn
hc
hc
hc
hc
hc
Hy
hc
Th
Th
Th
Th
Gn
Th
Gn
Th
Rhiz1
0
0
0
0
0
0
0
0
0
0
0
0
0
Rhiz2
0
0
0
0
0
Rhiz2
DRg
0
0
0
0
0
0
0
0
Rhiz1
0
Rhiz2
Rhiz1
Rhiz1
Node2
DRg
Node2
DRa
DRg
klieg
7
4
8
9
5
8
7
8
8
7
7
9
8
7
0
p Th
Gn
E1
8
9
7
P2 LF1 LF2 W Clonel Clone2
Gb
Mb
DRg
Gb
O
Th
0tb
6
2
4
6
4
3
3
3
4
2
3
3
1
2
1
6
5
3
c
3
3
2
3 c
6
2
51
3
4 0
3
2
2
3
Eur
Eur?
Crop
SAm
Unk
SAm
Aus , NZ
SAT
SAm
SAl
As2 , Am
E2 C
Origin
3
Eur
2
0
42
7
2
1336
714
1091
1011
1744
2118
1753
2013
541
998
137
323
146
67
10
396
311
3
16
359
874
1875
357 '
17 12
1600
109
347
165
3171
0
302
0
583
744
88
644
65
32
10
11
5
1
0
27
24
0
0
6
48
402
33
370
190
/
10
631
14
0
0
7
1
9
11
11
14
11
12
8
11
11
10
4
0
0
0
6
6
13
6
14
12
5
11
5
800
0 14
25
0 12
422 30 4
71
0 0
228
7 0
735 576 4
2393
23 791
6 2
3
6
0 0
2671
5
0 0
160 122 11
3
92
4.1
3.2
3.0
3.8
3.3
3.4
3.6
3.7
4.4
3.9
4.3
3.8
4.6
5.1
5.9
3.8
1.7
0.4
2.5
4.0
3.7
3.7
4. 1
3.8
3.7
4.4
5.1
4.5
4.0
4.9
4.1
3.1
1.4
4.2
3.4
5.1
15.6
15.3
13.7
16.0
14.0
14.1
15.2
14.9
15.8
15.8
16.1
15.7
15.8
16.1
16.1
15.7
12.1
11.5
12.9
16.0
15.7
15.0
15.9
15.3
15.3
16.4
15.4
15.7
16.1
16.3
15.7
15.3
12.2
14.5
14.4
16.0
4.5 16.5
3.6 16.4
16.2
4.1
4.9 15.5
7.0 13.7
16.4
4.0
8,
16 10
4, 5
3, 6
10
3, 7
2, 10, 16
1, 11
1,
192, 16
865
877
1236
774
1226
1202
915
986
16, 17
1
11
3, 6, 7
11
10, 12, 14
1, 3, 17
11, 13
861
3, 17, 19
843
3, 17
784
3, 17
782
3, 17
927
3, 17
892
3, 17
861
3, 4
895
3, 17
1940
15, 16
1392
7
1141
7, 18, 19
787
1, 4, 17
759
4
910
3, 4
776
4
877
3, 4
868
3, 4, 17
709 Co 18, 19, 2 1
1129
876 Co 3,
215
774
776
890
666
1568
1225
1138
874 Co
753 Co 18
7 13
3,
782
187
1004
1, 3, 16
832
1493
GB
IR CI Tj an Tjul Prec Co Br Habitats
1139
8 12 3.7 15.5
848
1, 3, 7, 8
33
0 12
0
3,6 15.5
16.2
773
1002
20
3.9
935
81, 3
3
4
4
4
5
5,
4
5
6
4
3
3
4
5
4
5
4
6
6
7
4
4
5
6
51
9
2
6
4
3
5
71 4,
3 61
8 8
7 4
8 8
8 8
6 5
7 10
7
7
7
7
6
6
7
6
7
9
9
9
7
7
7
71
7
9
7
8
7
7
7
71
5
6
4
9
8
8
4
8
7
7
7
5
3
6
6
4
6'
6
6
6
6
6
6
5
8
8
7
6
6
8
71
7
7
7
8
7
3
71
5
5
4
5
8
8
9
6
5
6
3
5
2
2
7
6
2
5
8
4
5
5
5
5
3
2
2
6
5
5
4
6
8
4
6
5
6
2
2
6
3
4
2
4
5
3
2
5
L F R N S
8 3 6 4
5
71 5
4 7 5
4 3
8 3 3
7 2
-C.
G.%
Taxon name
n
AN
N
-0.59
13
12
13
ChQ Hght Len
-0.44
80
-0.4 1
75
-0.04
100
-0.95
80
-0.06
40
-0.90
75
-0.58
30
0.59
150
0.01
70
0.80
30
n -0.15
120
r
50
50
n -0.68
50
-0.34
75
n 0.12
100
r
0.29
200
0.03
100
s -0.07
150
n 0.23
200
-0.32
120
45
n -0.22
40
s -0.30
50
n -0.56
17
n
50
r -0.10
60
n
150
n -0.33
100
270
n 0.43
n 0.45
60
r
20
r
0.07
120
s -0. 16
50
r -0.73
80
4600
5500
0.771 80
n -0.06
95
20
n
13
RS
n
n
n
n
n
s
n
r
VU
VU
VU
EN
CR
CS
NE V U
Fam NS
N
N
N
N
N
N
N
AN
AN
AN
N
N
AN
N
AR
N
N
AR
N
N
AN
AN
N
N
N
N
N
N
N
N
N
N
N
N
NA
AN
AN
AR
N
AN
N
hc
hc
hc
1 P2 LF1
hc
Th
Th
Th
Th
Th
he
Gn
Gn
Gn
Gn
Th
Th
Th
hc
hc
hc
he
he
Hy
Th
Th
Gn
hc
Th
hc
hc
hc
hc
Hy
he
Ch
hc
hc
hc
Ph
Ph
b Th
hc
hc
hc
h Sto12
h Sto12
h Sto12
LF2 W Clonal Clone2
Rhiz1
0
0
0
0
0
Rhizl
Rhiz2
Rhiz2
Rhiz2
Rhiz2
0
0
0
0
0
0
0
0
Gn
Rhiz2
0
0
Rhiz2
Rhir /
0
0
0
0
0
Gn
Rhiz2
0
w 0
h 0
h 0
h 0
w 0
w 0
he
h 0
h 0
h Rhiz2 Sto12
h Sto12
2
3
4
6
8
8
5
5
7
4
8
7
4
5
9
5
2
8
8
7
7
8
6
7
1
7
5
7
4
3
2
3
3
7
8
7
8
Eur
Crop
Eur
As 1
Eur
Eur
Origin
3
1
Eur
Eur?
1
Eur
6
6
3
3
4 c
4
4
6
3
3
3
3
3
5 c Eur, As
Am4
3
5
3
Eur
3 c
c
E2 C
5
6
6
5
5
3
6
E1
5
7
8
7
7
7
2
6
12
21
2
67
71
78
93
6
8
8
75
6
IR
43
740
434
934
93
31
5
145
10
582
569
0
0
0
42
0
0
21
72
2629 89
3
GB
1355
204 1
1881
2578
296
203
446
390
335
1328
1825
6
5
6
401
482
10
184
47
2449
1102
80
10171
33
219
1927
26
2294
2429
2 182
2 115
3
14
55
8
1460
1144
1191
885
1343
3
0
12
0
3.0 14.9
3.5 14.5
3.5 11.5
786
1094
1153
CI Tja n Tj ul Prec
0 3.3 14.8 1021
9 3.7 14.8 1093
10 3.7 15.1
958
12 3.6 14.6 1077
1 3.6 15.2 1003
1 3.8 15.9
780
0
1.6 12.2 1588
0
3.8 14.8 1149
0 2.8 13.9 1000
12 4. 1 15.3
941
2
3.6 14.8 1002
5 5.3 16.7
774
0 3.8 16.5
613
5 5.3 16.7
774
10 4.2 15.8
819
3 4.1 16.2
751
599
0 4.1 16.7
0 2.6 13.5 1127
0 3.6 16. 1
631
6 3.6 14.6 1044
10 3.7 15.5
856
6 4.1 15.9
760
0 2.4 13.1 1515
0 -0.7 10.8 1770
9 4.5 15.3
918
942
4
3.5 15.1
0
3.3 16.2
621
3 3.5 14.8 1023
10 3.5 14.7 1047
11 3.7 14.7 1055
14 3.8 14.8 1048
0 -1.1 10.6 1829
0 5.4 15.5 1279
0 3.9 16.3
806
0 4.1 16.4
826
2 3.2 14.7 1054
1 3.2 14.1 1223
10 3.9 15.9
795
8 3.8 16.0
781
4
3.4 14.9
985
0
3.5 11.5 1153
3
7
16
Br Habitats
6
11, 13, 14
4 , 11
3, 4
13, 14
13, 14
7, 15
3
1, 3
3
14
Co 19
8
8, 19
3, 17
3, 4
Co 19
5
11
11, 14
3, 17
4, 17
1, 16
11, 15
Co 19
3, 6
8
3, 6
3, 6
11
1, 16
10, 15
1, 10
7
1, 3
2
2
3, 4
7
3, 17
3, 16
Co
7
3
4
8
6
3
2
71
5
4
5
4
5
6
7
5
4
4
7
7
6
8
7
6
7
4
4
5
5
7
6
5
4
4
3
7
8
9
2
6
7
6
7
1
2
2
5
6
4
7
2
4
2
3
4
3
5
2
4
3
5
4
2
6
6
7
8 4
2
71 6
7
7
6
4
6
5
6
5
6
6
5
5
6
7
6
5
6
7
8
8
71
10
5
4
3
5
9
4
5
6
5
3
6
8
8
6
4
5
5
7
2
3
3
4
5
5
5
F R N S
7 6 6
6 71 7
Taxon name
Fam NS CS
N VU
N
N
N
NA EX
N
N
N
AN
AN
N
AC
N
N
N
N
N
N
N
N
N
N
N
AN
N
N VU
N
N
N
N
AN
N
N
N
N
NA
N VU
N
N
N
N
N
N VU
AR
N
N
0.40
'
s.
0.16
1.35
0.09
-0.28
-0.79
-1.67
-0.88
-0.3 1
I
n:
n 1 -0.70
0.83
0.63
-0.05
n. 0.21
r:
s -0.48
n.
s ' 1.33
n 0.27
-1.55
n. 0.60
n
n
1.10
r
1.17
r ' -0.04
r , -0.10
n. -0.37
n -0.50
n -1.14
n
0.27
s
0.34
r
n
n
n
n
n
n
n1
s
ni
e
s:
-0.83
-0.76
-0.03
-0.16
-0.3 1
Chg
RS
e
S.
n
n
x
o
n
n
Hght Len
9
10 10
100
70
5
8
3
8
2500
4200
3000
200
6
15
15
15
6
40
50
40
70
20
35
100
50
22
40
30
10
75
100
58
75
70
90
25
10
10
15
25
80
40
80
20
30
30
1 P2 LF1 LF2 W Clonel Clone2
hc
h Sto12
Hy
h Rhiz2
hc
h 0
hc
h 0
hc
h O
hc
h O
hc
h o
hc
h 0
Ph
w 0
Ph
w 0
Ph
w 0
Th
0
hc
0
hc
Rhiz1
he
0
hc
0
hc
Rhiz1
Gn
0
Gn
0
hc
0
DRi
hc
Rhiz2
p Th hc
0
hc
0
DRi
hc
0
hc
Rhiz2
hc
0
hc
0
he
Rhiz2
Th
0
he
0
he
0
hc
Rhiz2
hc
Rhiz2
hc
0
Nodel
hc
0
Th
0
Ch
0
Ch
0
Ch
0
Ch
0
Gn
Rhiz2
Gn
Rhiz2
Gn
Rhiz2
Th
0
Th
0
Th
0
5
3
4
4
5
4
5
5
3
6
6
4
4
Eur
Am4
Eur
Origin
Eur
7 3
1 3
2 6
7 ?
9 1
5 6
5 6
NHem
6 6
6 6
6 4
5 4 c
9 1
5 3 c
8 1
7 2
7 3
7 3
7 5
4 3
6 5
6 6
6 6
4
0
8
8
6
3
7
5
7
1
8
6
8
El E2 C
5 2
7 2
7 3
'7 4
1 4
7 1
7 1
4 6
GB
11
312
584
1938
1
0
500
1523
444
1009
65
137
1445
2804
2766
1295
1243
949
1163
72
83 1
2792
72
169
1063
8
62
1865
54
1960
132
2766
1637
2721
16
16
18
153
2 150
2 176
268
48
10
19371
2030
2605
0
24
109
391
0
74
309
484
116
25
0
10
303
985
975
405
0
308
251
2
0
985
0
3
0
0
0
297
0
0
12
928
423
903
0
0
0
0
717
665
0
0
0
480
819
906
IR
CI Tja n
12
5.2
3
3.6
0 3.6
0 3.6
0 3.0
0 5.1
0 3.9
0 3.0
2 3.3
6 3.7
0 0.8
2 4.0
14
4.1
14
3.5
3.5
14
8
3.8
0
3.5
1 3.5
0
3.4
0
0.6
6
3.7
14
3.5
6 4.6
1 3.1
2
3.6
0 -0.6
0
1.1
5 3.4
13 5.9
5 3.2
2
3.7
10
3.5
2
3.6
11 3.6
0
1.9
14 6.3
1 2.8
0
3.7
11 3.4
3.5
3.6
3.1
1.2
1
3.7f
3.7
1
3.6
Tjul Prec Co Br Habitats
16,1
871
7, 16
14.8 1040
13
15.5
8 19
6
15.0
958
7
13.6
752
11
14.7' 1270
10, 12
13.7 1475
10, 11
13.6 1305
11, 12
14.0 1302
2
15.5
889
1, 2, 17, 19
11.7' 1930
2
15.6
896
3, 4
14.8 1065
3, 6, 18
14.5 1105
6, 7
14.5 1102
3, 5
14.0 1248 Co 15, 2 1
15.6
804
7
14.3 1235
1, 10
14.7 1147
1, 6
11.5 2142
15
15.8
790
7, 8
14.5 1105
3, 4, 5, 6
16.4
741 Co 19
14.5
959
1
15.6
850
3
10.5 1907
15, 16
11.6 2252
15, 16
14.3 1134
6, 19
16.3
935
3
14.8 1015
1
15.2
948
11, 13, 14
14.5 1102
3, 5, 6, 7
14.8 1017
3, 5, 6, 7
14.5 1095
1, 3, 6
13.9 1246
16
16.3
923
3, 4, 17
15.0 1037
7
16.2
783
7
14. 1 1200
8, 10, 12
14.4 1114
7
16.0
829
1
15.1 1091
1, 16
13.1 1061
1
14.9 1008
3, 4
14.8 1039
4, 17
14.6 1073
3, 4, 17
L F R N
8 3 8 2
8 10 4 2
7 5 7 6
71 4 7 3
9 8 8 2
7 8 4 2
8 8 4 2
8 8 6 2
7f 5 5 2
7 3 5 2
7 6 2 2
7 4 7 7
8 6 6 4
71 5, 6 4
7 5 6 7
8 7 6 4
8 4 7 3
6 6 6 2
5 5 71 4
7 5 7 3
7 5 7 5
7 5 6 7
8 3 5 2
5 5 6 5
9 4 7 4
6 5 3 2
7 5 6 3
8 6 6 4
8 4 5 5
4 5 6 5
7 9 7 6
7 5 6 5
7 5 6 5
7 6 6 6
5 5 7 6
9 4 6 4
9 6 9 1
7 3 8 2
8 71 2 2
8 5 6 3
4 5 7 6
5 3 7 3
4 5 5 5
71 5 71 6
7 51 6 7
7 5 7 6
0
0
0
0
0
0
0
0
0
2
0
0
3
0
0
0
0
0
1
0
0
Co
Potamogeton praelongus
Potamogeton pusillus
Potamogeton rutilus
Potarnogeton trich oide s
Potentilla anglica
Potent)Ila ans erina
Potentilla argentea
Potent"Ila crantzir
Po tam og eto n po lyg onifolius
Populus alba
Populus alba x tremula (P. x
cane s ce ns)
Populus nigra sens.lat
Populus tremula
Potarnogeton acutifolius
Potamogeton alpinus
Potamogeton be rchto ldii
Potarnogeton co/oratus
Potamogeton compressus
Potamogeton crispus
Potamogeton epihy drus
Potamogeton filiformis
Potamogeton friesii
Potamogeton gramineus
Potamogeton grarnineus x
lucens (P. x zizii)
Potamogeton grarnineus x
perfoliatus (P. x nite ns)
Potarnogeton lucens
Potarnogeton natan s
Potamogeton nodosus
Potamogeton obtusifolius
Potarnogeton pectinatus
Potarnogeton perfoliatus
Po lys tichurn setiferurn
Taxon name
Polygonum boreale
Polygonum maritimum
Polygonum oxyspermum
Polygonum runvagum
Polypodi urn cambricum
Polypodiurn inte rj e ctum
Polypodium vulgare
Polypodiurn vulgare sens.lat
Polypogon rnonspeliensis
Polystichum aculeaturn
Polystichurn fon ch itis
Sali
Sali
Sall
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
-0.26
0.77
0. 18
0.57
0.11
-0.23
-0.78
-0.2 1
-0.18
0.96
p
p
p
p
p
p
p
p
p
hc
Irreg
Irreg
Irreg
Me g
DRa
Rhiz2
Meg
Irreg
lrreg
DRa
DRa
DRa
Sto12
Sto12
0
0
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
hc
hc
hc
hc
p
p
p
p
p
p
p
n
n
n
r
n
n
n
n
n
n
r
n
n
n
n
s
NH
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Pota
Rosa
Rosa
Rosa
Rosa
250
250
100
250
190
230
300
10 70
300
70
45
100
25
25
30
20
Meg
Hy
120 p
S
NH
Pota
0.25
w Root
w Root
w Root
DRa
lrreg
DRa
Meg
DRa
Rhiz2
lrreg
Rhiz2
DRa
lrreg
Ph
Ph
Ph
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
Hy
n
n
r
n
n
s
s
n
r
s
$
n
DRg
Rhiz2
Rhiz2
Rhiz2
Rhiz2
Rhiz2
Rhiz2
Rhiz2
Rhiz2
Rhiz2
DRa
DRa
DRg
Rhiz2
DRa
7
5
8
5
6
5
7
4
8
4
8
7
5
7
2
7
5
7
4
5
8
5
8
5
4
5
5
4
6
6
6
6
6
2
6
6
3
4
3
6
4
4
4
5
3
6
6
3
5
5
0
6
6
6
Eur
Origin
LF2 W Clonal Clone2 El E2 C
4 2
0
9 1
0
6 3
0
7 3
0
Gn
9 1
Rhizl
7 2
Rhizl
Gn
Gn
5 3
Rhizl
6 3
Rhir l
Gn
9 1
0
7 5
0
4 6
0
9 2
0
Eur, As 1
w Root
I
p
0.97 3000
p
0.65 3000
p
0.88 2000
100 p
0.05
280 p
0.30
60 p
1.66
70 p
0.03
90 p
-1.68
150 p
190 p
0.11
30 p
0.63
150 p
-1.06
80 p
0.67
RS Chg_ Hght Len P1 P2 LF1
Th
a
20
$
Ch
p
20
0.21
r
Th
a
20
n 0.01
Th
a
20
hc
P
40
n
hc
p
40
n
hc
P
40
n
hc
40
n -0.03
p
Th
a
80
0.60
$
hc
p
60
n 0.54
hc
p
30
n -0.76
hc
p
120
1.47
n
Ph
p
2000
AN
N
N
N VU
N
N
N
N
N
N VU
N
N
N
N
AN
NS CS
N
N EN
N
AR
N
N
N
N
N
N
N
Fam
Poly
Poly
Poly
Poly
Poly
Poly
Poly
Poly
Poac
Dryo
Dryo
Dryo
Sail
7
2
7
0
1
1
0
0
0
0
2
1
0
1
1
0
0
0
2
0
0
5
6
4
5
5
7
5
2
5
6
5
6
5
6
2
2
12
12
11
12
12
12
12
10
12
12
12
12
5
7
3
5
14
13, 14
11, 13, 14
14
13
13, 14
13, 14
11, 12, 13
13
13
13, 14
13, 14
6
6
3, 8
7, 16
1257
832
1091
761
994
844
1026
1254
1178
877
1115
739
1045
1083
704
1855
13.6
15.5
14.5
16.4
14.7
15.3
14.6
13.9
14.1
15.1
12.9
16.2
15.0
14.6
16.0
12.0
7
7
7
6
7
6
7
8
7
7
7
6
7
8
8
8
0
4
7 12
13, 14
1138
3.5 14.3
3.5
3.7
3.5
3.9
3.4
3.8
3.5
3.3
3.2
3.6
3.8
3.8
4.0
3.6
3.6
0.7
/
0
1
0
0
5
7
6
6
5
5
5
46
/
5
5
3
6 6
6 8
6 5
7 12
71 12
7 12
7 11
7 12
7 12
8 12
7 12
7 12
7 12
0
0
0
0
1
0
0
N S
6 0
4 3
8 3
5 0
3 0
3 0
3 0
3 0
6 3
5 0
3 0
6 0
6 0
1, 3
3, 14
1, 16
13
13
13, 14
13
13
13, 14
13
13
13
13, 14
L F R
7 5
9 3
9 6
8 4
6 5
5 5
5 5
5 5
8 8
5 5
6 5
4 5
6 6
819
800
1102
702
1188
1027
893
697
893
1288
1109
782
1170
Tjul Prec Co Br Habitats
4
12.4 1084
950 Co 19
16.2
14.8 1102 Co 19
4
738
16. 1
3, 16
15.1 1102
1, 3, 16
14.9 1074
1, 3, 16
14.3 1158
1
14.4 1133
6
661
16.6
1, 16
14.4 1145
7, 15, 16
12.1 1857
1
15.0 1032
3, 17, 19
867
15.3
15.6
15.7
14.5
16.4
14.1
14.7
15.1
15.9
15.2
13.0
13.3
15.4
13.8
3.8
3.6
3.3
4.0
3.1
3.6
4.1
3.4
3.7
4.5
3.6
3.6
3.3
CI Tjan
3.6
0
4 6.0
9 4.6
4 3,9
1 4.4
3.9
12
1 3.5
12 3,5
0 4.3
3 3.2
1.4
0
10 4,1
10 3.6
113
171 I
648 6
0 0
155 0
242 4
275 0
582 5
90 0
146 1
0 0
0 1
733 4
973 12
0 5
0 0
92
100
216
456
2340
15
601
1165
1056
1841
252
806
13
185
1248
2662
323
98
110
66
414
0
142
317
140
0
358
0
63
43
163
0
1
75
0
157
487
523
916
0
314
20
748
116
IR
1145
692
2248
35
5371
1461
142
134
1541
2
16 1
270
473
GB
109
16
305
274
196
1188
1795
2496
45
1618
181
1249
1531
-le
C.
Taxon name
Potent/ha erecta
Potentilla fruticosa
Potentilla ne um ann iana
Potentilla palustris
Potentilla reptans
Potent/Ila rupestris
Potentilla sterilis
Primula elatior
Pri mula farinosa
Primula scotica
Primula veris
Primula vulgaris
Prunella vulgaris
Prunus avium
Prunus cerasifera
Prunus cerasus
Prunus domestica
Prunus laurocerasus
Prunus lusitanica
Prunus padus
Prunus spinosa
Pseudofumaria lutea
Ps e ud orchis albida
Pseudotsuga menziesii
Pteridium aquilinum
Puccinellia dis tan s
Puccinellia fasciculata
Puccinellia maritima
Puccinellia rupestris
Pulicaria dysenterica
Pulicaria vulgaris
Pulmonaria longifolia
Pulmonaria obscura
Pulmonaria offi cina lis
Pulsatilla vulgaris
Pyrola media
Pyrola rninor
Pyrola rotun difolia
Pyrus communis sens.lat
Pyrus cornm un is sens.str.
Pyrus cordata
Quercus cerris
Quercus ilex
Quercus petraea
Quercus robur
R adio/a linoide s
Fam
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Prim
Prim
Prim
Prim
Prim
Lami
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Furna
Orch
Pina
Denn
Poac
Poac
Poac
Poac
Aste
Aste
Bora
Bora
Bora
Ranu
Pyro
Pyro
Pyro
Rosa
Rosa
Rosa
Faga
Faga
Faga
Faga
Lina
NS CS
N
N
N
N
N
N VU
N
N
N
NE
N
N
N
N
AN
AR
AR
AN
AN
N
N
AN
N
AN
N
N
N
N
N
N
N VU
N
N VU
AN
N
N
N
N
AR
AR
NA EN
AN
AN
N
N
N
,
'
e
n
n
1
I
r
s
5
n
s'
n
!
n
n
s
n
s
n
r
s
r,
,
n
n
RS
n
r
s
n
n
r
n
s
s
e
n
n
n
n
Chg Fight Len 1 P2 LF1 LF2
-0.50
25
hc
1.44
100
Pn
-0. 17
10
hc
-0.2 1
50
Gn Hy
-0.62
30
hc
60
hc
-0.30
15
hc
0.01
20
hc
-0.46
5
hc
hc
-0.18
5
-0.32
15
hc
0.16
15
hc
0.60
30
hc
1.29 2500
Ph
3.43
800
Ph
0.90
800
Ph
2.19
800
Ph
4.70
600
Ph
800
Ph
0.58 1500
Ph
0.40 400
Ph
0.59
30
hc
-0.88
20
Gn
5800
Ph
-0.7 1 150
Gn
3.02
60
hc
-0.51
50
hc
-0.27
80
hc
-0.40
40
b Th hc
-0.08
80
hc
-0.551 45
Th
-0.0 1
40
hc
30
he
1.77
30
hc
-0.50
30
hc
-1.09
10
hc
-0.55
7
hc Ch
-0.08
12
hc
1.49 1500
Ph
1500
Ph
400
Ph
2.32 3500
Ph
2.37 2500
Ph
0.14 3000
Ph
-0.60 3000
Ph
-0.87
6
a
Th
Root
0
0
Root
Root
3
5
3
E2 C
4
6
3
6
4
3
2
3 c
5
1
4
3
6
3
Eur
Eur, As 1
Eur
Eur
Eur
Eur
Origin
A m4
7 6
5 4
8 2
5 1
8 1
8 4
7 4
7 1
7 3 c
7 3 c Eur
7 3 c
4 4
4 6
5 4
Eur?, As l ?
Gard
7 1
Eur
9 1
Eur
7 3
7 3
7 3
4
5
7
Clone l Clone2 El
0
5
4
0
7
Rhiz2
5
Sto12
8
0
9
Stoll
7
0
7
0
4
0
4
0
7
0
7
Nodel
6
Root
7
Rhiz2
0
0
Node2
0
Rhiz2
0
0
0
0
0
Rhizl
Rhizl
Rhizl
w Root
w Root
w Root
w
w
w
w
h 0
W
h
w
h
h
h
h
h
h
h
h
h
h
h
w
w
w
w
w
w
w
w
h
h
w
GB
IR CI Tjan Tj ul Prec Co Br Habitats
2700 947 12
3.5 14.4 1118
8
8
7 0 2.8 13.7 1366
16
128
0 0
3.1 14.7
994
7, 16
1672 721 'I 3.4 14.0 1186
11
1882 769 11 3.8 15.1
959
6
01 0
3.0 13.8
4
934
16
2167 742 8 3.6 14.7 1071
1
38
0 0 3.3 16.3
591
1
103
0 0 2.2 13.8 1194
7, 11
42
0 0 3.6 12.4 1026 Co 6, 19
1632 369 6
3 7 15.2
900
6, 7
265 1 934 11 3.5 14.5 1108
1, 16
2783 970 14
3.5 14.4 1106
6, 7
2136 431 7
3.4 14.8 1024
1
900
11 4
3.7; 15.8
803
1, 3, 17
706 339 7
3.9 15.2
961
1, 3
1656 406 8 3.7 15.2
928
3, 17
1253 345 4 3.8 15.3
953
1, 17
468
31 1 3.8 15.5
876
1, 17
1089; 189
2.71 13.8 1240
1
23081 9 17
3.7 14.8 1032
3
1425 39
3.6 15.4
876
3, 17
385 110
2.6 13.0 1534
7, 8
900
13
3.4 15.0 1035
1, 2, 17
2685 964 /
3.5 14.5 1109
1, 9
405
39
4.1 14.9
859
3, 21
97
11
4.5 16.4
722
6
788 201
4.1 14.4 1162 Co 21
148
0
4.5 16.2
756 Co 6, 19
14971 391 1
4.0 15.5
885
6, 11
121
0
4.0 16.5
696
8
21
0
4.7 16.5
8 10
1, 3
1
0
3.2 16. 1
577
1
682
8
3.4 15.3
922
1, 3, 17
691 0
3.3 16.0
664
7
258 47
2.2 13.0 1197
2, 10
558 50
2.6 13.9 1089
1, 16
155
13
2.9 14.2 1072
2, 19
781
6
3.8 15.9
801
1, 3, 17
3, 17
9
0
5.4 15.9 1129
3
1247 42 1
3.8 15.6
862
1, 3, 17
802 22 1
4.1 15.9
824
1, 17, 19
1832 549
3.5 14.6 1136
1
2310 663 1
3.5 14.71 1049
1, 3
548 115 I I
4.2 14.7 1061
10
L F
7 7
8 6
7 3
8 9
71 5
7 4
5 5
4 5
9 8
9 4
7 4
5 5
7 5
4 5
6 5
6 5
7 5
4 6
6 5
5 6
6 5
6 6
8 5
6 6
6 5
8 8
8 7
9 8
9 7
71 71
9 8
6 4
4 6
5 5
71 3
5 4
5 5
6 7
7 5
7l 5
6 5
6 4
6 3
6 6
71 5
8 7
R N
3 2
8 2
8 1
5 3
7i 5
6 2
5 5
7 6
9 2
71 2
7 3
6 4
8 4
6 6
7 6
6 5
7 6
5 6
7 6
6 71
7 6
8 5
6 2
4 4
3 3
7 7
7 7
7 6
7 5
7 4
6 7
6 5
8 7
8 6
8 3
5 2
4 2
7 3
6 7
6 71
5 4
6 6
7 4
3 4
5 4
4 2
0
0
1
S
Ribes alpinum
R ibe s nigrum
Ribes rubrum
Ribes spicatum
Ribes uva-crispa
Robinia pseudoacacia
Romulea columnae
R hy n ch o sp o ra fusca
Taxon name
Ranunculus acris
Ranunculus aquatilis
Ranunculus aquatilis sens.lat
Ranunculus arvensis
Ranunculus auncomus
Ranunculus baud otii
Ranunculus bulbosus
Ranunculus circinatus
Ranunculus ficaria
Ranunculus flammula
Ranunculus fluitans
Ranunculus he derac e us
Ranunculus lingua
Ranunculus muricatus
Ranunculus omiophyllus
Ranunculus ophioglossifolius
Ranunculus paludosus
Ranunculus parviflorus
Ranunculus peltatus
Ranunculus penicillatus
Ranunculus repens
Ranunculus reptans
Ranunculus sardous
Ranunculus sceleratus
Ranunculus trich op hy llus
Ranunculus tripartitus
Raphanus raphanistrum
Raphanus raphanistrum
subsp.maritimus
Raphanus raphanistrum
s ubs p .raphan is trum
Reseda lutea
Reseda luteola
Rhamnus cathartica
Rhinanthus angustifolius
Rhinanthus minor
R hodod e ndron ponticum
R hy nch ospora alba
N
AR
NA
AR
N
AN
N
AN
N
N
N
AN
NA
N
AN
AN
N VU
Bras
Bras
Rese
Rese
Rham
Scro
Scro
Eric
Cype
Cype
Gros
Gros
Gros
Gros
Gros
Faba
hi d
VU
EN
EN
CS
NS
N
N
N
AR
N
N
N
N
N
N
N
N
N
AN
N
N
N
N
N
N
N
N
NA
N
N
N
N
Fam
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Ranu
Bras
r
n
5
n
s
s
n
n
n
0.39
0.69
-0.04
-0.10
-0.49
1.83
-0.43
0.02
0.45
1.76
1.79
-0.12
0.72
60
75
150
600
60
50
500
30
30
200
200
200
200
100
2700
6
80
a
a
b
0
0
0
0
0
0
0
Rhizl
Rhiz2
0
0
0
0
0
Root
Otb
0
LF2 W Clone l
0
0
0
0
0
0
0
Irreg
Otb
Hy
Nadel
1rreg
Hy
0
Hy
Rhiz2
0
Hy
0
0
Rhizl
0
0
Irreg
Sto12
Hy
Sto12
0
0
0
0
hc
0
p he
hc
hc
Ph
Th
Th
Ph
hc
hc
Pn Ph
Pn
Pn
Pn
Pn
Ph
Gn
p hc
RS Chg Hght Len P1 P2 LF1
hc
n 0.30 - 75
p
n
90 a
Hz
n -1.37
90 a
Hz
-3.771 60
a
Th
hc
n -0.33
40
p
n -0.04
60 a
Hz
hc
n -0.48
40
p
Hy
n -0.34
75 p
n 0.16
25
p
Gn
hc
n -0.60
50
p
n
1.96
300 p
Hy
n 0.10
9 23 a p Hz
n
1.70
120
p
hc
40
a
Th
n 0.52
9 25 a p Hz
r
40
a
Th
o
40
p
he
n -0.08
40
a
Th
n
90 a
Hz
n
180 p
Hy
n 0.55
60
p
hc
r
10
p
hc
n 0.24
45
a
Th
n -0.05
60
a
Th
n -0.07
60 a
Hz
s -1.09
9 25 a p Hz
n -1.39
70
a p Th
Nadel
9
5
5
4
5
7
5
7
8
8
8
7
5
5
9
Eur, As 1
Eur
6
2
3
4
Eur, As1
2
3 c
3
Eur
Am6
1
3
3
4
4
4
3
1
Clone2 El E2 C
Origin
3 5
Irreg
7 3
Irreg
6 3
8 4
5 3
Node2 8 3
6 3
7 5
DRa
8 3
7 3
Node2 7 3
Node2 8 2
7 4
0 3
Eur
Node2 8 2
8 3
9 1
6 2
Irreg
6 3
7 3
5 5
4 6
7 3
5 6
Wag
3 6
8 1
8 3
1797
1276
1660
857
90
2629
1966
624
46
50
1749
1874
118
2130
566
2
325
GB
2780
1123
1830
824
1379
361
2259
575
2648
265 1
373
1703
537
18
813
4
0
497
990
706
2784
9
544
1492
1121
79
1862
237 8
0 7
428 12
88 0
0 0
850 6
507 6
407 0
94 0
0 0
213 2
0 2
0 0
312 4
1 4
0 14
3.6
3.6
3.8
3.6
3.1
3.5
3.5
3.6
4.4
2.3
3.4
3.3
2.0
3.4
3.7
6.2
4.7
CI Tjan
13 3.5
4 3.6
6 3.6
3 3.7
1 3.3
5 4.4
3.6
14
0
3.7
14
3.6
11 3.5
0
3.3
8
3.6
0 3.7
0 5.0
1 3.7
2 4.9
4 6.1
11 4.3
5 3.5
0 3.8
14 3.5
0 2.6
12 4.1
6
3.8
6
3.7
0
5.2
13 4.6
108 13
IR
968
157
430
2
226
61
6 16
76
820
938
3
551
210
1
151
0
0
0
151
284
983
0
0
353
259
1
259
15.0
15.7
15.3
15.8
14.6
14.4
14.6
14.0
14.7
14.2
14.9
15.1
13.2
14.8
16.0
16.5
15.1
941
808
879
783
809
1115
1111
1400
1205
1084
995
944
1095
1006
748
842 Co
1093
4
3
3, 17
1, 3
3, 4, 7
6
1, 10, 16
12
12
1, 3, 16
1, 3, 14
1
1, 16
1, 3
3, 17
18, 19
19
Tjul Prec Co Br Habitats
14.4 1106
6
15.2
860
13, 14
15.3
860
13, 14
15.9
742
4
15.1
922
1
15.3
905
13
14.8 1004
6, 7
15.8
753
13, 14
14.5 1084
1, 3
11
14.4 1122
15.4
798
14
14.5 1093
11, 13
15.2
868
11
15.6
864
4
14.7 1180
11
16.5
824
13
16.9
843
6
16.0
834
6
15.2
908
11, 13, 14
15.2
942
14
14.5 1105
3, 6
13.7i 1467
13
15.9
800
3, 6, 13
15.4
861
11, 13, 14
15.2
861
11, 13
15.8 1024
13
15.0 1079
4, 19
L
1
1
I l
1
1
1
1
1
1
1
1
1
F
5
6
5
6
6
2
4
3
1
1
6
6
6
6
6
6
2
7
6
7
8
7
7
6
3
2
3
8
6
7
7
7
6
5
R N S
6 4
7 5
7 5
7 6
6 5
7 6
7 4
7 7
6 6
5 3
7 6
5 5
6 7
5 5
5 4
7 5
6 3
6 5
5 6
8 5
6 7
6 2
6 7
8 8
6 6
6 3
6 6
1
S
vi
t- }
Taxon name
Fam NS CS
RS Chg
Hght Len
60
60
40
77
15
15
3
15
20
10
10
95
36
45
35
38
40
25
40
35
25
40
2500
70
250
1000
800
1500
6
100
100
300
• 40
700
400
300
150
15
1000
600
50
90
80
150
1000
400
1 P2 LF1
hc
Hy
Hy
Pn
Th
Th
Ch
Ch
hc
Ch
Ch
Hy
Th
Th
Th
Th
Th
Th
Th
Th
Th
Th
Ph
Pn
Pn
Ph
Ph
Ph
Ch
Pn
Pn
Pn
Ch
Ph
Ph
Pn
Pn
Ch
Ph
Ph
Th
hc
hc
hc
Ph
Ph
Pn
Ph
Ph
Pn
hc
LF2 W Clonel
h
h
h
w
Clone2 E1 E2 C
Origin
GB
IR
CI Tja n
Tj ul
Prec
Co
Br Habitats
L
F R N
I 7
7
5
12
8
11
4
5
3
4
4
7
1
7
3
7
5
6
2
7
4
6
6
11
6
6
6
6
6
6
6
6
6
5
8
2
3
7
I
5
7
2
3
3
4
2
4
4
5
3
3
5
6
8
4
2
7
7
7
Taxon name
Samolus valerandi
Sanguisorba minor
Sanguisorba offi cina lis
Sanicula europaea
Saponaria officina (is
Sarcocornia perennis
Saussurea alpina
Saxifraga aizoides
Saxifraga cernua
Saxifraga cespitosa
Saxifraga granulate
Saxifraga hirculus
Saxifraga hirsuta
Saxifraga hypnoides
Saxifraga nivalis
Saxifraga oppositifolia
Saxifraga rivularis
Saxifraga rosacea
Saxifraga spathularis
Saxifraga stellaris
Saxifraga tridactylites
Scabiosa columbaria
Scandix pecten-veneris
S ch e uchz e ria palustris
Schoenoplectus lacustris
Schoenoplectus pungens
Schoenoplectus
tabem aemontani
Schoenoplectus ttigueter
Schoenus ferw gineus
Schoenus nigricans
Scilla autumnatis
Scilla Verna
Scirpoides ho los ch oen us
Scirpus sylvaticus
S cleranthu s annuus
S cleranthu s p erennis
Scorzonera humilis
Scrophularia auriculata
Scrophularia no dos a
Scrophularia s corod on ia
Scrophularia umbrosa
Scrophularia vernalis
Scutellaria galericulate
Scutellaria minor
Secale cereale
N
N
N
N
N
N
N
N
N
N
NA
N
N
AN
N
AN
N
N
AC
Cype
Cype
Cype
Cype
Lili
Lili
Cype
Cype
Cary
Cary
Aste
Scro
Scro
Scro
Scro
Scro
Lami
Lami
Poac
EN
VU
VU
CR
VU
NS CS
N
N
N
N
AR
N
N
N
N VU
N VU
N
N VU
N
N
N
N
N
N EW
N
N
N
N
AR
N VU
N
NA
Fam
Prim
Rosa
Rosa
Apia
Cary
Chen
Aste
Saxi
Saxi
Sax/
Saxi
Saxi
Saxi
Saxi
Saxi
Saxi
Saxi
Saxi
Saxi
Saxi
Saxi
Dips
Apia
Sche
Cype
Cype
n
r
r
n
s
n
r
n
n
r
r
n
6
,
n
.
n
6
,
/
r
r
-0.2 1
-0.37
0.75
0.72
0.54
-0.39
0.03
-0.53
-0.37
0.12
0.21
0.02
-2.68
-0.11
0.67
150
125
40
75
20
10
100
120
20
20
50
120
100
100
100
65
50
20
150
60
2 10
0 .47
n
s
n
n
r
r
n
r
o
n
s
n
r
x
o
n
n
n
Chg Hght Len
-0.42
45
-0.16
50
-0.23
120
-0.98
40
0.29
90
-0.22
30
-0.5 1
45
-0.6 1
20
15
-0. 10
10
-0.26
36
-0.30
20
1.39
30
-0.54
20
-0.50
15
-0.45
3
0.19
10
20
40
-0.58
20
-0.12
10
-0.7 1
70
-3.65
50
22
RS
n
n
n
n
Hy
Hy
hc
hc
Gb
Gb
hc
hc
Th
Ch
hc
hc
hc
hc
hc
p hc
hc
hc
Th
0
0
0
0
0
0
Rhizl
Rhizl
0
o
0
Otb
Otb
0
Rhizl
0
o
Rhizl
Rhizl
DRg
1 P2 LFI LF2 W Clonel Clone2
hc
0
hc
0
hc
0
hc
o
Rhizl
hc
Rhiz2
Ch
0
hc
Stoll
Ch
Node2
hc
DRa
DRi
Ch
0
hc
DRa
hc
Rhiz/
hc
Stol l
Ch
Node2
hc
0
Ch
Node2
hc
Rhizl
DRa
Ch
0
hc Ch
Stoll
hc Ch
o
Th
0
hc
0
Th
0
hc
Rhiz2
Hy
Rhizl
Hy
Rhizl
5
8
8
7
4
8
9
7
8
7
7
7
7
8
7
8
7
El
8
8
5
7
7
9
1
1
1
1
7
2
7
4
1
1
1
4
7
1
8
7
8
4
6
7
5
5
3
4
1
1
4
4
3
3
3
2
4
1
Eur
4 c
Eur
4
2
Eur
E2 C
6
4
6
3
3
1
5
3
6
6
3
6
1
1
6
6
6
2
0
3
3
3
4
6 c
4
3
Origin
702
8
3
763
49
310
3
871
983
11
4
1490
2327
68
196
192
1751
803
134
GB
845
1216
946
2025
1246
80
237
422
5
13
975
20
0
406
72
276
21
2
0
440
985
804
780
14
1202
0
192
3
0
484
0 1
29
0
102
69
0
0
504
850
0 1
14
0
2 12
136
71
3.9
5.9
4.6
5.2
3.4
3.6
33
48
3.9
3.6
5.8
3.1
3.3
3.6
4.1
4.0
04
4.2
4.7
IR CI Tjan
4 11 12 4.3
84 6 3.7
12 0
32
634 0
3.5
256 9
3.9
4
1 4.4
21 0
1.6
17 0
1.7
0 0 0.5
0 0 -0.4
6 0 31
16 0 2.4
36 0 5.0
61 0 2.0
1 0
06
22 0
1.8
0 0 -0.3
31 0 4.4
137 0 4.7
61 0
1.8
316 11 3.9
0 0 3,5
94 6 3.8
1 0 2.2
535 0 3.7
0 1 6.1
15.3
16.5
12.4
13.9
16.2
13.9
16.3
15.2
15.2
16.0
16. 1
15.4
14.7
16.0
14.9
15.3
14.9
14.9
15.8
917
11, 13
799
14
1113
11
1261
11
939 Co 10, 18
1103
6, 10
909 Co 19
921
1, 11
883
4, 8, 10, 16
656
8, 16
1097
6
918
11, 14
1069
1, 3
1033
3, 16, 18
868
1, 14
769
1, 3
1055
11
1191
11
852
3, 4
T'ul Prec Co Br Habitats
15.1 1013
11
15.6
844
7
15.1
938
6
14.7 1090
1
15.5
890
3, 17
16.4
702 Co 21
11.9 20 10
7, 15, 16
12.3 1799
7, 11, 15, 16
11.7 2265
15, 16
10.7 2003
15, 16
15.1
853
6, 7
13.2 1253
11
14.6 1334
1, 15, 16
12.8 1697
7, 15, 16
11.5 2099
15, 16
12.1 1980
7, 15, 16
10.8 1941
11, 15
14.3 1312
16
14.4 1328
16
12.3 18 11
11, 15, 16
15.3
921
16
15.6
831
7
15 7
792
4
13.9 1380
12
14.9 1013
13. 14
16.7
869 Co / 9
9 10
8 10
8 9
8 8
9 3
8 5
8 8
6 8
71 4
81 3
8 7
7 8
5 6
71 4
7 9
5 5
7 8
7 9
8 5
L F
8 8
7 4
7 7
4 5,
8 5
9 8
8 6
8 9
6 6
71 5
8 5
8 9
6 7
7 5
6 6,
8 6
6 9
7 4
6 8
8 8
7 2
8 3
7 4
9 9
8 11
8 10
8
7
7
71
6/
5
7
6
4
4
5
7
7
6
7
7
6
4
71
7
7
2
2
1
3
6
6
4
2
2
7
6
6
7
7
5
2
7
R N S
8 5
8 3
6 5
7 5
6 6
8 6
6 3
6 2
7 1
71 1
6' 4
6 2
5 2
6 3
71 3
8 2
5 2
8 4
3 2
5: 3
7 2
8 2
7 4
3 1
71 6
7 7'
Taxon name
Sedum acre
Sedum album
Sedum anglicum
Sedum forsterianum
Sedum rosea
Sedum rupestre
Sedum telephium
Sedum villosum
Selaginella selaginoides
Selinum carvifolia
Sempervivum tectorum
Senecio aquaticus
Senecio cambrensis
Senecio cineraria
Senecio erucifolius
Senecio fluviatilis
Senecio j acobaea
Senecio paludosus
Senecio smithii
Senecio squalidus
Senecio sylvaticus
Senecio viscosus
Senecio vulgaris
Serapias parviflora
Seriphidium maritimum
Serratula tinctoria
Seseli libanotis
Sesleria caerulea
Setaria viridis
Sherardia arvensis
Sibbaldia procumbens
Sibthorpia europaea
Silaum silaus
Silene acaulis
Silene conica
Silene dioica
Silene gallica
Silene latifolia
Silene noctiflora
Silene nutans
Sfi ene ofi tes
Silene uniflora
Silene vulgaris
Silybum marianum
Simethis planifolia
Sinapis alba
Fam
Cras
Cras
Cras
Gras
Cras
Gras
Cras
Cras
Sela
Apia
Cras
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Orch
Aste
Aste
Apia
Poac
Poac
Rubi
Rosa
Scro
Apia
Cary
Cary
Cary
Cary
Cary
Cary
Cary
Cary
Cary
Cary
Aste
Lili
Bras
NS
N
AR
N
N
N
AN
N
N
N
N
AN
N
NE
AN
N
AN
N
N
AN
AN
N
AN
N
NA
N
N
N
N
AN
N
N
N
N
N
N
N
AR
AR
AR
N
N
N
N
AR
N
AR
VU
CR
VU
CS
RS Chg Hght Len
10
n -0.24
20
2.41
5
n -0.2 1
20
1.54
s
35
n -0.4 1
30
2.20
60
n -0.34
10
s -0.76
9
n -0.47
100
r
40
80
n -0.92
30
r
60
2.73
120
n -0.03
150
0.03
125
n 0.11
175
r
100
50
0.771
70
0,09
n
60
0.63
37
n -1.08
30
r
50
n -0.42
70
n -0.2 1
60
r
45
s -0.09
70
0.80
40
n -0.94
5
s -0.75
5
s -0.14
100
n -0.42
10
n -0.47
35
$ -1.05
90
n -0.44
45
-2.78
100
-0.88
50
-2.04
$ -0.39
80
80
r -0.36
28
n -0.39
80
n -1.26
100
-0.07
42
o
90
-0.90
LF2 W Clone')
Node2
Node2
Node2
Node2
0
Node2
0
0
Nodal
0
Stol l
0
0
Pn sw 0
Rhizl
Rhiz2
0
0
0
0
Ch
0
0
0
0
Rhiz2
hc
0
0
0
0
0
0
Node2
0
0
0
Ch
0
0
0
0
0
0
0
hc
0
hc
a hc Th
0
0
Gn
0
Th
p
b
p
a
p
1 P2 LF1
Ch
Ch
Ch
Ch
hc
Ch
hc
Ch
Ch
hc
Ch
p hc
hc
Ch
hc
hc
b hc
hc
hc
a p Th
Th
a
Th
a
Th
a
Gn
p
Ch
p
hc
p
hc
b
he
p
Th
a
Th
a
hc
p
Ch
p
hc
p
Ch
p
Th
a
hc
Th
a
hc
p
Th
a
hc
p
hc
p
Root
Irreg
Clone2
Irreg
Irreg
Irreg
Irreg
8
1
7
7
1
5
5
9
8
7
7
7
5
8
0
5
8
7
7
8
9
7
7
7
5
7
7
0
7
7
7
7
7
2
4
7
Eur
Origin
Eur
3
1
Eur
3
4
Eur, As l
4
4
4 c
SAm
Gard
3
Eur
3
3
1
2
3
5 c
3
Unk
3
6
1
4
3
4
3
2
4
3
4
3 c
2
5
Eur
3
2
3
5
3
6
3
El E2 C
7 3
9 2
7 1
8 1
1 6
GB
2108
1603
988
122
524
1212
1197
211
988
3
500
2304
14
256
1276
184
2725
7
62
1494
1853
1747
2630
1
222
945
4
76
431
1635
133
107
963
236
59j
2514
455
1954
686
52
18
943
1710
552
0
1082
IR CI T an
3.7
504 12
517 13 3.8
393 14 4. 1
0 0 3.6
82 0 21
70 4 3.7
3.5
0 0
1.6
0 0
288 0 2.8
3.4
0 0
3.5
42 2
919 1 3.6
0 0 3.6
8 5 4.4
30 0 3.8
25 0 3.3
982 14 3.6
0 0 3.4
3.4
0 0
3.7
75 4
3.6
238 12
69 1 3.4
942 14 3.6
0 0 6.0
20 1 4.3
0 2 3.8
0 0 3.7
3.1
89 0
27 5 3.9
369, 12 3.9
0 0 0.4
9 8 5.3
0 0 3.6
1.9
9 0
0 10 4.0
263 14 3 4
31 11 4,2
2 11 13 3.6
22 1 3.7
0 10 4.3
3.2
0 0
236 13 4.0
280 8 3.7
64 6 4.0
4 0 5.7
162 10 3.8
T ul Prec Co Br Habitats
16
995
14.9
3, 16, 17
948
15.1
16, 18
14.3 1321
3, 16
14.8 1239
15, 16
12.6 1728
3, 16
905
15.5
1, 3, 16
15.0 1032
11
12.9 1273
7, 11
13.0 1443
11
595
16.1
3, 17
871
15.3
11, 14
14.5 1109
3, 17
804
15.0
3, 17, 18, 19
837
15.9
3, 6
783
15.8
1, 11, 13, 14
14.6 1018
3, 6, 7, 8
14.5 1102
11, 13
573
16.2
12.3 1072
3, 6
3, 17
850
15,5
2, 8, 9
987
14.9
3, 17, 19
920
15.1
3, 4, 17
14.6 1075
6
16.2 1048
783 Co 21
15.8
7
965
15.5
7
645
16.4
7, 16
13.9 1262
3, 4, 17
798
15.9
7, 16
916
15.2
15
11.3 2109
3, 14
15.5 1189
6
743
15.8
7, 15, 16
11.8 1937
8, 19
702
16.0
1
14.6 1065
3, 4
877
15.7
3
911
15,1
4
736
15.8
16
883
15.7
625
16.1
3, 8
14.1 1263 Co 18
3, 6
897
15.3
3
781
15.8
10
15.1 1324
3, 4
832
15.5
L F
8 2
8 3
8 3
7 3
7 6
7 2
7 5
8 9
8 7
7 7
8 2
7 8
8 5
9 3
7 5
7 8
7 4
7 9
8 7
8 4
7 5
8 5
7 5
8 4
9 7
7 6
7 4
7 6
7 4
7 4
8 5
5 7
8 5
8 5
8 3
5 6
7 4
7 4
7 4
5 3
8 3
8 6
7 4
8 4
71 5
7 4
R
7
6
4
5
6
5
7
6
6
8
4
6
7
7
7
6
6
7
6
7
5
7
7
7
8
6
8
8
7
6
4
5
7
6
4
6
5
7
7
8
7
6
8
8
4
7
N S
2 1
2 0
2
1
3
4
5
2
2
4
1
5
7
3
5
7
4
6
7
7
6
6
7
3
6
2
3
2
7
4
3
5
4
1
2
7
5
6
6
4
2
4
5
6
3
6
u-i
u,
Taxon name
Sinapis an eensis
Sison amomum
Sisymbrium altissimum
Sisymbrium offi cinale
Sisymbrium ori entale
Sisyrinchium be rm ud iana
Sium latifolium
Smyrnium olusatrum
Solanum dulcamara
Solanum nigrum
Solanum sarachoides
Solanum tubemsum
Soleiro lia soleirolii
Solidago can adensis
Solidago gigantea
Solidago virgaurea
Sonchus arvensis
Sonchus asper
Sonchus oleraceus
Sonchus palustris
Sorbus anglica
Sorbus aria
Sorbus aria agg,
Sorbus arrane nsis
Sorbus aucuparia
Sorbus bristoliensis
Sorbus de von ien sis
Sorbus domestica
Sorbus e mine ns
Sorbus hibernica
Sorbus intermedia
Sorbus lanc as triensis
Sorbus leptophylla
Sorbus leyana
Sorbus minima
Sorbus p orrigentiformis
Sorbus ps e ud otennica
Sorbus rupicola
Sorbus subcuneata
Sorbus torminalis
Sorbus vexans
Sorbus wilmottiana
Sparganium angus tifolium
Sparganium emersum
Sparganium erectum
Sparganium nata ns
Fam
Bras
Apia
Bras
Bras
Bras
Irid
Apia
Apia
Sola
Sola
Sola
Sola
Urti
Aste
Aste
Aste
Aste
Aste
Aste
Aste
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Rosa
Spar
Spar
Spar
Spar
NS CS
AR
N
AN
AR
AN
N
N
AR
N
NA
AN
AN
AN
AN
AN
N
N
N
N
N
NE VU
N
N
NE V U
N
NE EN
NE
N CR
NE VU
NE
AN
NE
NE CR
NE CR
NE VU
NE
NE VU
N
NE VU
N
NE V U
NE CR
N
N
N
N
RS Chg Hght
, -1 76
90
n -0.19
100
-0,84
100
i -0.2 1
95
-0.24
80
o -1.80
23
s -1.83
200
0.66
150
n -0.11 225
n 0.44
60
60
100
2.36
10
200
200
n -0.89
70
n -0.12
150
n
0.78
150
n -0.42
150
s , 0.18 250
r
200
n
1500
n' 0.82 1000
r
750
n 0.86 1500
r,
1000
s,
1500
r1
500
r
600
o
600
1000
r
500
r
300
r
200
r'
300
5,
500
r
700
s
200
r
1000
n 0.22 1900
r
600
r
600
n
1.66
n
0.61
n 0.48
150
n : -0.13
Len P1 P2 LF1 LF2
a
Th
b
hc
a
Th
a b Th hc
Th
hc
Hy
he
Pn Ch
Th
a
Th
Gn
Ch
hc
hc
he
hc
a
Th
Th
he
Pn
Ph
Ph
Ph
Ph
Ph
Ph
Ph
Ph
Ph
Ph
Ph
Pn Ph
Pn
Pn Ph
Ph
Ph
Pn
Ph
Ph
Ph
Ph
100
Hy
60
Hy
Hy
50
Hy
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
w
h
h
h
h
W
h
h
h
h
h
h
h
h
sw
Cb nel
0
0
0
0
0
0
0
0
Node2
0
0
Rhiz1
Node2
Rhiz1
Rhiz1
0
Rhiz2
0
0
0
0
0
0
0
0
0
0
Root
0
0
0
0
0
0
0
0
0
0
0
Root
0
0
Rhiz2
Rhiz2
Rhiz2
Rhiz2
5
5
2
5
4
3
3
4
1
3
3
1
5
1
1
3
8
8
0
5
7
8
8
7
7
7
7
4
5
7
7
7
7
7
7
7
4
4
7
7
7
7
4
5
7
5
2
1
3
1
1
3
6
6
6
0
4
6
7
7
7
3
8
Clone2 El E2 C
7 4
9 2
Eur
SAm
SAm
Eur
Am
Am
Eur
Eur
Eur
Origin
88
625
773
981
645
2043
2320
2584
2358
51
14
225
341
1
2472
1
32
4
8,
0'
686
9
3
2
1
30
1
98
4,
573
4
1
652
1426
2129
508
GB
2373
790
6 14
21171
864
0
259
803
1918,
1258E
IR
772
0
29
775
75
43
34
327
533
0
0
56
130
24
1
492
806
961
924
0
1
2
75
0
749
0
11
0
0
63
17
0
0
0,
0'
0
0
13
0
0
0
0
207 '
393
766
267
CI Tian Tjul Prec Co Br Habitats
11 3.7 14.7 1012
3, 4 , 17
0 3.9 16.2
756
3
3 3.7 15.6
771
3, 17
13 3.8 15.0
973
3, 4, 17
10 3.8 15.6
815
3, 17
0 4.5 14.4 1290
6, 13
0 3.7 15.9
702
11, 14
13 4.4 15.5
907
3
14
3.8 15.2
945
3, 11, 14
14
3.9 15.9
809
4
4
3.8 16.1
696
4, 17
5
3.9 15.4
883
4
11 4.3 15.7
906
3, 17
3.6 15.6
833
3, 17
3.5 15.5
854
3, 17
3.4 14.2 12 17
10, 16
3.8 14.8 1013
4
1
3.7 14.6 1070
3, 4
1
3.71 14.8 1031
3, 4
3.8 16.4
615
11
3.8 15.2 1082
1, 16
3.8 16.3
774
1
3.8 16.3
780
17
2.5 12.1 1530
16
3.4 14.4 1128
1, 2, 16
4.4 16.5
844
1, 16
5.0 15.4 1171
1, 3
4.5 16.5
857
16
4.4 16.2
878
1
4.5 14.6 1035
1, 3, 16
3.5 15.2
901
1, 3, 17
3.3 14.8 1285
16
2.9 14.5 1501
16
2.5 14.0 1687
16
2.2 14.0 1523
16
3.8 15.2 1273
16
2.5 12.1 1530
16
2.9 13.6 1405
16
4.0 14.9 1380
1
3.9 16.0
804
1
4.2 14.9 1409
1, 16
4.4 16.5
844
1, 16
3.0 12.9 1571
13
3.6 15.0
982
13, 14
3.7' 14.9 1000
11, 13
3.3 13.8 1303
13
L F R N S
8 5 7 7'
7 4 7 5
8 5 6 4
71 4 7 7
7' 4 7 5
8 8 6 3
7 10 7 7
T 5 7 7
71 8 71 7
7 5 7 8
7 4 7 7
7 4 6 7
4 8 7 6
8 5 6 6
8 5 5 6
5 5 4 3
8 6 7' 6
T 5 7 6
71 5 71 71
7 8 7 7'
6 4 8 4
6 4 7 4
6 5 7 4
71 4 4 3
6 6 3 4
6 4 8 4
6 5 6 5
6 4 8 3
6 5 71 5
6 5 7 5
6 6 8 7
8 4 7 3
5 5 7 5
7 5 8 4
6 4 8 3
7' 5 7' 5
7 4 4 3
8 4 7 3
6 5 4 4
4 5 6 5
6 5 4 4
6 4 8 3
8 11 4 2
7 11 7 6
7 ' 10 7 7
7 11 6 3
in
cn
Symphytum orientale
Symphytum tuberosum
Sybnga vulgaris
Tamus communis
Tanacetum parthe nium
Tanacetum vulgare
Taraxacum
Sym phy tum officinale
Taxon name
Spartina alternitiora
Spart ina anglica
Spartina mari tim a
Spergula arvensis
Spergularia bocco ne i
Spergularia marina
Spergularia media
Spergularia rubra
Spergulati a rupicola
Spiranthes aestivalis
Spiranthes ro manz offi ana
Spiranthes spiralis
Spirodela polyrhiza
Stachys alpina
Stachys arvensis
Stachys germanica
Stachys officinalis
Stachys palustris
Stachys palustris x sylvatica
(S. x ambigua)
Stachys sylvatica
Stellaria graminea
Stellaria holostea
Stellaria media
Stellaria media egg.
Stellaria neglecta
Stellaria nemorum
Stellaria pallida
Stellaria palustris
Stellaria uliginosa
Stratiotes ab ides
Suaeda maritima
Suaeda vera
Subularia aquatica
Succisa pratensis
Symphoricarpos albus
Symphytum asperum x
off icinale (S. x up land icum)
0.34
1.83
0.11
4.48
-0.4 1
0.23
-0.23
0.43
AN
N
AN
N
AN
N
AR
N
N
Bora
Bora
Bora
Bora
Olea
Dios
Aste
Aste
Aste
n
n
n
n
n
n
n -0.49
n -0.02
n -0.56
n
n 0.03
n 0.42
n 0.21
1.17
n
n -0.89
n -0.10
1.65
r
n -0.47
s -0.11
n 0.73
n -0.57
1.74
NH
N
N
N
N
N
N
N
N
N
N
NA
N
N
N
N
AN
Lami
Lami
Cary
Cary
Cary
Cary
Cary
Cary
Cary
Cary
Cary
Hydr
Chen
Chen
Bras
Dips
Capr
135
135
70
55
700
400
54
120
30
100
200
30
120
100
100
80
60
50
50
80
60
50
60
40
6
50
RS Chg_ Hght Len
110
-0.22
130
n 0.11
50
s -0.55
30
o -2.30
10
-0.22
10
1.83
n
15
n -0.24
15
n 0.05
10
n 0.30
40
x
25
0.45
s
15
n -0.95
0.8
n -0.18
100
25
-1.17
80
r -0.27
60
n -0.62
0.01 100
n
NS CS
AN
NE
N
N
AN
N
N
N
N
N EX
N
N
N
AN
AR
N EN
N
N
Fam
Poac
Poac
Poac
Cary
Cary
Cary
Cary
Cary
Cary
Orch
Orch
Orch
Lemn
Lami
Lami
Lami
Lami
Larni
Rhiz2
Rhiz2
Rhizl
Rhiz1
0
0
0
Rhizl
0
Rhizl
Rhizl
Stoll
0
0
0
0
Rhiz2
0
0
0
0
Rhir l
0
0
Rhizl
0
w
h
h
w
h
h
h
h
w
h
h
h
h
hc
hc
hc
Ch
Th
Th
Th
hc
Th
hc
he
Hy
Th
Pn
Hz
he
Pn
hc
hc
hc
hc
Ph
Gn
Ch hc
hc
hc
Rhizl
DRa
Nodel
Nodel
Nodel
Nodel
Nodal
Nodel
1
2
4
1
6
4
3
0
3
0
3
6
3
2
3
3
6
E2 C
3
2
5
6
9
5
6
3
7
7
7 4
5 4
7 4
6 5
6 5
7 3
5 3
8 4
5 5
7 3
5 4 c
8 5
9 1
4 6
7 4
P1 P2 LF1 LF2 W Clonel Clone2 El
Rhiz2
hc
p
7
Rhiz2
hc
p
8
Rhiz2
hc
p
6
0
Th
a
9
0
Th
a
8
0
Th
a
8
0
Ch
p
8
0
Th
a
7
0
Ch
7
0
hc
4
0
hc
8
0
hc
8
Frag
Hy
7
hc
0
8
0
Th
7
0
hc
7
he
0
5
Rhiz2
Gn
Origin
Eur
Eur
Eur
Eur
Am
Eur
Eur
Am
CI Tian Tjul Prec
760
0 0 4.1 15.7
904
71 1 4.6 15.7
625
0 0 4.1 16.6
811
0 9 6.3 16.7
901
0 9 5.9 16.3
185 8 4.3 14.6 1094
186 0 4.2 14.6 1125
983
66 13 3.5 14.9
169 13 5.2 15.1 1105
822
0 2 5.6 16.5
44 0 4.4 14. 1 1248
884
117 13 4.4 15.9
768
3.9 15.9
991 2
3.1 14.7 1007
0 0
964
206 13 4.0 15.3
66 1
0 0 3.4 16. 1
940
38 4 3.6 15.4
3.71 14.6 1079
919 9
7
8
6- 6
5
6
6
7
6
7
7
7
6
6
7
7
7
7
5
7
4
6
5
5
4
6
5
6
7
6
6
6
6
7
7
7
3
11
3
1, 3
3, 17
1, 3
3, 17
3
3, 5, 17
15.0
15.4
16.0
13.5
15.3
15.7
14.9
15.0
14.5
3.6
3.7
3.71
2.5
3.6
3.8
3.5
3.5
3.5
8
10
3
0
4
4
11
10
14
399
189
2
0
163
0
413
0
974
1924
1263
442
407
1296
1421
2126
2004
2778
952
871
719
1047
881
850
980
958
1104
6
8
4
6
7
6
7
7
4
4
5
6
6
5
2
2
7
6
7
5
6
6
5
6
6
4
6
5
7
8
8
5
5
6
6
6
6
5
5
5
7
6
4
8
8
11
8
7
11
7
5
7
6
7
5
7
7
6
4
7
7
7
7'
9
9
7
7
5
3, 14
1086
3
1064
6
1065
1
1058
3, 4
1095
3, 4
1095
1, 3
911
1, 14
1027
3, 19
766
11
793
11, 14
1128
13
596
1094 Co 21
620 Co 19, 21
13
1789
6
1124
1, 3, 17
992
14.7
14.7
14.6
14.7
14.5
14.5
15.6
14.2
15.8
15.5
14.4
16.1
14.9
16.4
12.6
14.4
14.9
N S
7
6
5
5
7
6
5
2
5
2
4
3
7
7
5
5
3
7
3.5
3.6
3.5
3.5
3.6
3.6
3.9
2.7
4.0
3.6
3.5
3.5
4.3
4. 1
2.4
3.5
3.6
L F R
9 9 8
9 9 8
9 9 8
7 4 5
9 4 6
8 8 8
8 8 8
8 3 4
9 6 6
9 8 8
8 8 6
8 4 6
71 11 7
7 5 8
8 5 5
7 3 8
7 5 5
7 8 7
4
12
8
7
14
14
0
0
13
1
9
0
6
1
0
2
4
Co Br Habitats
Co 21
Co 21
Co 2 1
4
16, 17, 19
Co 3, 21
Co 21
8, 17
Co 18
11
11
7
13
1, 3
3, 4
3
6, 7
11, 14
1171
806
852
671
962
962
4
0
27
73
798
0
14 1
0
33
934
744
IR
1020
2462
2446
2372
2749
2749
775
432
593
389
2570
65
525
47
324
2633
2067
GB
12
251
571
0
18
680
659
1363
239
1
21
655
535
3
1418
10
1571
2346
T
B
B
B
C
L
R
h
T
T
h O
DRg
DRp
9
1
8 4
7 5
9 1
7 4
7 0
4 6 c
4 6
4 6
8 4
9 1
8 4
7 3
8 4
9 1
3
6
Crop
Eur
Am4
Eur
0
0
Rhizl
0
0
0
0
0
0
Rhizl
Rhizl
Ogr
Rhiz2
0
0
0
0
Node2
0
0
0
w
w
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
h
7
1
Clone l Clone2 El E2 C
Origin
0
7 3
0
7 3
0
Am4
0
3 6 c
0
3 6
0
7 3 c Eur
Rhizl
8 3 c Eur
Rhiz2
8 4
Rhiz2
8 2
Rhiz2
1 6
0
5 4
Rhiz2
5 5
Rhiz2
7 6
0
7 1
0
7 5
0
4 3
0
8 4
0
Am4
Node2
5 3
Node2
7 3
Node2
5 3 c
0
7 4
W
w
h
h
h
h
h
sw
h
h
h
h
h
h
h
h
h
h
w
sw
sw
sw
w
196
84
156
267
11
389
2178,
708
1749
162
1
1553,
514
1205
3
1996
2548
7531
148
1940
3
GB
1881
509
308
99,
26
12
72
24
2322
409'
669
504
171
145
1600
70
9
574
2246
455
9
896
5
1
0
0
11
1
0
824
77
124
62
0
629
0
56
0
381
942
42
5
2 14
0
IR
260
11
20
0,
0
0
2
12
588
23
91
89
54
0
143
0
0
7
486
3
0
20
0
5
0
0
14
2
14
14
6
13
11
4
1
4.0
16. 1
3.3 15.6
0 6 11.6
3.2 14.3
4.0 16.3
3.7 16.2
3.71 14.9
4. 1 15.9
3.6 15.3
3.8 14.2
2.8 14.0
3.2 13.71
1,8 12.6
3.8 15.5
6 5 16.2
3.7 15.1
3 6 14.6
4.0 16.0
4.6 16.3
3.5 15.0
6.4 16.4
780
8 16
1964
1131
739
707
1004
792
87 1
1390
858
1312
1277
84 1
867
939
1072
764
767
943
883
3
1
7, 11
1
6
4
3
3, 6
6
16
12
10, 12
1, 2, 10
8
16
3, 16, 19
6
6
8
3, 5
3, 18
CI Tja n Tj ul Prec Co Br Habitats
7 3 5 15.1
976
1
12 3 4 15.0
949
8
2
3.6 15.2
974
1, 3, 17
0
3 71 16. 1
774
7
0
3.6 16. 1
6 17
13
0
3.6 16.3
767
3, 4, 7, 16
0 4.2 15.7
856
3, 7, 16
I
4.0 15.8
740
11, 13, 14
1
3.5 14.5 1150
1, 9, 16
2.0 12.1 1845
7, 11, 15, 16
3.7 15.8
762
11
3.4 14.0 1185
16, 19
3.9 15.5
869
1, 11
3 6 16.2
762
7
1
3 7 15.3
858
3, 4
2 0 13.5 1314
16
3,3 16.0
732
3, 16
1 3.6 15.4
977
2, 17
1
3.4 14.2 1175
7, 16
3.7 16. 1
752
7
3,3 16.2
606
8
3 5 14.4
15.5 1064
873
2.4
1
7
4
8
3
7
8
7
8
8
2
8
8
5
9
9
8
71
8
9
7
8
R
7
2
6
8
7
8
8
8
4
6
7
6
7'
8
7
6
8
5
6
8
5
6
4
6
5 7
9 7
6 7
3 6
4 8
5 7
5 7
4 7
7 7
9 2
8 2
6 3
3 5
4 5
4 6
4 6
7 7
3 5
5 7
2 5
L F
4 4
8 3
3 8
71 3
7 9
9 2
8 2
7 8
6 4
8 7
7 8
7 4
6 8
8 3
7 4
8 4
8 4
4 5
8 4
8 4
8 2
5 5
5
6
2
7
5
4
71
6
5
3
2
1
3
2
2
4
5,
6
2
6
2
0
1
0
0
0
0
0
1
0
0
0
2
0
0
1
N S
5 0
2 0
4 0
3 0
6 0
2 0
1 0
4 1
3
3
5
3
5
3
6
1
2
4
2
2
2
5
6
tri
co
Taxon name
Trifolium incamatum
subsp.molinerti
Trifolium medium
Trifolium micranthum
Trifolium occidentale
Trifolium ochroleucon
Trifolium om ithop od ioide s
Trifolium pratense
Trifolium repens
Trifolium scabrum
Trifoliurn sguamosum
Trifolium striatum
Trifolium strictum
Trifolium subterraneum
Trifolium suffocatum
Triglochin maritimum
Triglochin pal ustre
Trinia glauca
Tripleurospermum inodorum
Tripleurospermum maritimum
Tnpleurosp en num maritimum
sens.lat.
Trisetum flavescens
Triticum aestivum
Trollius europaeus
Tsuga he terophy lla
Tuberaria guttata
Tussilago farfara
Typha angustifolia
Typha latifolia
Ulex europaeus
(fl ex gallii
Ulex minor
Ulmus glabra
Ulmus minor
Ulmus plotii
Litmus procera
Umbilicus rupestris
Urtica dioica
Urtica urens
Utri cularia australis
Utricularia intermedia
sens.lat.
Utricularia intermedia
sens.str.
Utri cularia minor
0.40
0.20
n
n
n
N
N
N
Lent
Lent
-0.48
-0. 12
0.28
-0.70
-0.65
0.35
1.01
-0.34
0.20
0.20
-0.28
0.75
Lent
r
n
n
n
n
n
n
n
n
s
n
n
n
s
-0.73
n
N
N
AC
N
AN
N VU
N
N
N
N
N
N
N
N
NE
NA
N
N
AR
N
0.3 1
-0.13
-0. 10
0.14
-0.44
-0,22
0.12
-0.84
0.42
-0.18
1.31
-0.39
-0.32
-0.11
-0.53
0.62
n
n
n
r
n
n
a
s
n
n
n
n
s
n
r
n
s
n
n
r
RS Chg
Aste
Poac
Poac
Ranu
Pina
Cist
Aste
Typh
Typh
Faba
Faba
Faba
Ulrna
Ulrna
Ulma
Ulma
Cras
Urti
Urti
Lent
Faba N VU
Faba N
Faba N
Faba N
Faba N
Faba N
Faba N
Faba N
Faba N
Faba N
Faba N
Faba N VU
Faba N
Faba N
Junc N
Junc N
Apia N
Aste AR
Aste N
Fam NS CS
Th
hc
Th
hc Ch
he
Th
hc
hc Ch
Th
Th
Th
Th
Th
Th
hc
hc
hc
Th
Ch hc
p Th hc
he
Th
hc
Ph
Th
Gn
Hy
Hy
Pn
Pn
Pn
Ph
Ph
Ph
Ph
hc
hc
Th
Hy
Hy
Hy
Hy
60
20
20
40
20
45
7
10
45
12
45
20
20
40
20
15
10
5
55
55
20
60
60
60
80
150
60
4600
15
30
300
275
200
150
100
3000
3100
2000
3300
38
150
60
?
5
h DRa
h DRa
?
6
6
4
h DRa
Sto12
3
4
1
4
4
6
1
1
1
3
3
1
3
1
4
4
5
6
3
3
7
9 1
5 4
9 2
7 0
7 3
8 2
7 4
5 4
9 2
9 1
8 3
9 2
9 2
9 1
5 6
5 6
8 3
7 4
3 6
9
5
7
8
7
7
8
7
7
7
7
9
5
8
5
w
h
h
h
h
w
w
w
w
w
w
w
h
h
h
h
0
0
0
0
0
0
Rhiz2
Rhiz2
Rhiz2
0
0
0
0
Root
Root
Root
0
Rhiz2
0
DRa
0
0
0
Node2
0
0
0
Node2
0
0
0
0
0
0
Rhizl
Rhiz2
0
0
0
Hght Len P1 P2 LF1 LF2 W Clonal Clone2 El E2 C
Am4
Crop
Od in
0
208
52
9
0
18
976
981
12
0
31
0
1
0
251
71
3
2050
909
21
1271
281
2745
2798
390
116
848
4
337
96
8371
2196
6
2119
757
4
2
13
12
0
14
14
14
11
6
12
2
13
13
5
5
0
12
3
15
628 377
412
1
0
3.6
13.9
1321
2
2
4
4
8 12
13
11, 12
8 12
2
4
8 12
11, 12, 13
1545
3.1 13.2
8
3
8
6
4
7
4
3
1
6
7
7
3
2
2
6
7
7
6
4
6
7
7
6
3
5
6
7
7
5
3
1
7
7
7
8
5
7
6
5
8 5
7 4
8 5
7 7
6 6
9 2
7 6
8 10
8 10
7 5
7 6
8 6
4 5
5 5
5 5
5 5
6 4
6 6
8 5
7 12
N
4, 17 18, 19
6, 7
3, 4
11, 16
1, 2, 17
10
16
11
11
10
10
10
1
1, 3
3
3
3, 16
3, 14, 17
4, 17
13
1132
900
861
1517
1102
1069
1085
775
958
1090
1099
745
1056
764
667
819
1125
1102
9 14
1120
14,5
15.2
15.6
13.0
14.9
15.5
14.5
15.8
15.1
14.6
15.0
16.4
14.7
16. 1
16.0
15.8
14.9
14.5
15.0
15.0
4.4
3.6
3.8
2.2
3.4
5.71
3.5
3.7
3.8
3.6
4.1
3.9
3.4
3.9
3.4
3.7
4.3
3.6
3.7
4.0
R
2
4
5
2
2
3
5
6
2
6
2
2
2
2
5
2
1
6
6
6
7
8
9
7
9
7
7
9
9
8
9
8
8
8
8
9
8
8
F
51
6
5
6
8
5
7
6
7
7
5
5
4
4
7
6
8
6
6
I.
2
4
5
4
5
6
5
5
3
6
3
2
3
4
7
9
2
5
5
Co 18
6
8, 17
Co 18
3. 6
8
6. 7
6
8
Co 6, 19
8
16
8, 18
8
Co 21
11
7
4. 17
Co 18, 19
883
1008
848
940
595
874
1100
1105
831
757
805
874
805
774
118 1
1145
889
967
1132
16.4
14.8
15.9
15.9
16.3
16. 1
14.5
14.5
15.9
16.5
15,7
15.9
16.2
16.4
14.3
14.2
16.4
14.9
14.5
64
3.4
4.1
6.3
3.4
4.8
3.6
3.5
4.5
4.5
4.0
5.8
4.5
4.8
4.2
3.4
4.9
3.6
4.4
Br Habitats
Co
Prec
T uI
CI T an
3
757i 27
6
1734 36
4
741 5
0
856
0
1
598
4
1
5
7
2618 90
6
776 6
9
1860 68
2518 95 14
918 38 11
4
197
0
2338 60
9
641
0
128
0
13171
784 65 i 14
2773 98 13
1924 28 14
0
5
162
42
27
IR
GB
S
in
so
N
N
N
N
N
AN
N
AR
N
N
AN
AN
AN
N
N
N
AR EN
N VU
N
N
N
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Scro
Capr
Capr
Faba
Veronica an ag allts -aqu abca
Veronica arvensis
Veronica be cca bung a
Veronica catenata
Veronica chamaedrys
Veronica Milormis
Veronica froticans
Veronica he cierif oria
Veronica m on tana
Veronica off icinalis
Veronica persica
Veronica polka
Veronica praecox
Veronica scutellata
Veronica serpyllifolia
Veronica spicata
Veronica triphyllos
Veronica vema
Viburn um lan tan a
Viburnu m opulus
Vicia bithynica
N
Scro
Sc ro
Veron ica alp ina
-0.06
0.80
0.13
-0.82
-0.64
0.37
-0.15
-0.52
n
n
s
r
n
n
s
n
n
s
0.05
0.48
-0.3 1
0.37
-0.50
2.69
0.11
0.57
0.48
-0.84
-0.37
0.07
-0 .29
50
25
45
50
30
5
14
30
15
23
30
10
15
25
10
45
15
15
600
400
60
12
a
a
a
a
a
a
a
a
a
Chg_ Hght Len
20
20
0.47
100
100
0.81
30
-0.6 1
50
0.28
30
-0.39
50
-0. 18
30
-0.67
35
-0.64
175
-0.35
110
2.15
15
-1.86
15
-0.69
15
-0.11
15
-2.55
15
-0 .23
150
-0.12
120
0.94
135
0.27
200
0.35
100
-0.43
67
-0.38
10
a
n
n
n
n
n
s
s
n
s
n
N
n
RS
i
i
n
s
s
g
n
n
n
n
n
N
NA
N
AN
AR
AR
NS CS
N
N
N
N
N
N
N
N
N
N
N
AN
AR
AR
AN
N
AR CR
Scro
Fam
Lent
Lent
Lent
Lent
Eric
Eric
Eric
Eric
Eric
Vale
Vale
Vale
Vale
Vale
Vale
Vale
Vale
Scro
Scro
Scro
Scro
Scro
Verb
Scro
Taxon name
Utricularia ochroleuca
Utricularia stygia
Utricularia vulgaris seno lat.
Utricularia vulgaris sens.str.
Vaccinium microcarpum
Vaccinium myrtillus
Vaccinium oxycoccos
Vaccinium uliginosum
Vaccinium vitis-idaea
Valeriana dioica
Valeriana offi cinalis
Valeriana pyr enaica
Valerianella carinata
Valerianella dentata
Valerianella eriocarpa
Valerianella locusta
Valerianella rimosa
Verbascum lychn ids
Verbascum nigrum
Verbascum pul verulentum
Verbascum thapsus
Verbascum virgatum
Verbe na offi cinalis
Veronica agrestis
Th
Ch
Ch
Th
Th
Th
hc
hc
hc Ch
Th
Th
Ph
Ph
hc
Ch
P2 LF1 LF2
Hy
Hy
Hy
Hy
Ch
Ch Pn
Ch
Ch Pn
Ch
hc
hc
hc
Th
Th
Th
Th
Th
hc
hc
hc
hc
hc
hc
Th
Ch
p Hz Hy
Th
Hy Ch
p Hz Hy
Ch hc
hc Ch
5
0
0
0
Node l
0
0
7
7
7
7
7
9
8
8
5
8
7
5
h Nod e2
0
Node2
Nodel
0
Nodal
0
0
Node l
1
h
h
h
w
w
h
h
h
h
h
h
h
h
sW 0
8
8
7
7
5
7
1
h 0
h 0
Nodel
0
Nodel
Nodel
Node2
Node2
7
9
7
7
7
7
9
7
8
8
0
0
0
0
13
0
0
0
0
0
h
h
h
h
h
h
3
8
0
Eur
Eur
Eur
Eur
Origin
4 c
3 c
4 c
3
6
1
6
Eur
4
Eur, Asl
3 c Eur
4
3
3
3
3
5
3
4
6
4
3
4
3
2
3
3
3
4
2
4
2
5
E2 C
?
?
5
4
6
4
6
6
6
3
5
Clone l Clone2 E1
DRa
?
DRa
?
DRa
5
DRa
7
Node2
4
Rhiz2
4
Node2
4
Rhiz2
2
Rhiz2
2
Sto12
7
0
Rhizl
5
0
W
h
h
h
h
h
w
h
w
w
h
h
h
h
h
h
h
h
h
h
h
h
h
h
9
0
0
0
0
0
0
1
4
88 4 11
346 14
506 0
754 8
722 13
107 8
0 0
576 3
0
459 3
793 14
912 9
249 4
945 14
397 9
177
0
28
0
33
0
71 0
537
0
1854 565
74
0
26 52
1944
1808
2507
2232
1237
6
1877
26
1225
2614
2333
956
2609
2013
1715
35
301 0
77 0
0 0
701 1
246 0
0 0
89 0
0 0
808 0
8 0
66 13
45 4
0 6
238 9
27 0
0 0
0 8
0 0
375 12
13 3
90 10
744
159
100
1886
725
252
938
1171
2408
195
547t
600
59
1358
181
42
479 ,
51
1874
339
8571
3.7
3.5
3.3
3.8
3.6
4.4
3.5
3.71
3.6
3.4
3.7
3.8
3.3
3.4
-0 .5
3.7
3.5
3.6
3.8
3.5
3.6
3 .5
-0.7
3.6
3.8
0.9
3.3
3.0
1.1
2.2
3.2
3.4
2.7
4.3
3.9
4.5
3.9
4.3
4.1
3.9
3.4
3.7
4.2
41
CI Tjan
IR
GB
15.3
16.1
16.2
16.2
15.0
16. 1
14 .5
15.1
14.9
14.4
14.9
15.5
16.3
14.3
11.2
15.1
14.6
14.7
15.6
14.5
14.9
15.0
10.9
14.4
15.3
11.9
13.9
14.0
11.7i
12.9
15.3
145
13.8
15.8
15.7
16.0
15.3
15.9
16.5
16.2
16.1
15.2
15.8
15.8
Tj ul
949
690
6 18
763
10 19
827
1104
918
1029
1125
993
867
618
1129
182 1
899
1077
1034
8 16
1095
990
96 3
1783
1148
874
1339
1243
1229
1897
1474
896
1115
1151
93 1
800
826
925
860
790
742
624
9 18
870
859
Prec
Co
7, 16
3, 4
8
1, 3
1
3
3 , 5, 11
1, 3, 4, 17
1
7, 8
3, 4, 17
4, 17
3, 4, 8
11, 13
15, 16
13, 14
3, 4, / 6
11, 14
13, 14
1, 3, 6
3, 17
3, 4, 17
15
Br Habitats
11, 13
11, 13
13
11
12
10, 16
12
12, 15, 16
2, 10, 15
11
11
1, 3
3, 17
4
3, 16
3, 16, 19
4
3, 7
3
3, 16
3
3, 5, 17, 19
3, 16
8
7
8
7
6
7
7
6
4
6
6
7
8
8
8
7
8
7f
8
6
7
7
8
3
4
2
5
7
4
5
5
6
5
5
4
2
9
5
10
4
10
10
5
6
6
6
L F
8 12
8 12
7j 12
7 12
7 8
6 6
8 9
71 6
6 5
8 8
6 8
5 7
8 4
8 4
8 3
8 4
8 4
71 3
7 4
8 3
7 4
8 4
8 5
7
7
5
7
6
6
6
2
3
1
5
6
4
5
N S
1
2
4
4
1
2
1
2
2
3
5
5
4
4
3
4
3
3
6
5
5
5
6
6 7
5 2
7 7
6 5
6 6
7 8
6 5
7 7
7 2
7j 6
6 6
4 4
7 7
7 5
8 1
5 3
R
3
5
6
7
1
2
2
2
2
6
6
5
8
7
8
6
8
71
7
7
7
5
71
a,
0
Taxon name
Vicia cracca
Vicia laba
Vicia hirsuta
Vicia lathyroides
Vicia lutea
Vicia orobus
Vicia parvillora
Vicia sativa
Vicia sepium
Vicia sylvatica
Vicia tetrasperma
Vicia villosa
Vinca maj or
Vinca minor
Viola arvensis
Viola canina
Viola hilt s
Viola kitaibeliana
Viola lactea
Viola lutea
Viola odorata
Viola palustris
Viola persicifolia
Viola reichenbachiana
Viola riviniana
Viola rupestris
Vio/a tricolor
Viscum album
Vulpia bromoides
Vulpia ciliata
Vulpia fasciculata
Vulpia myuros
Vulpia unilateralis
Wahlenbergia hederacea
Wolffi a arrhiza
Woodsia alpina
Woodsia ilvensis
Zannichellia palustris
Zea mays
Zostera angustifolia
Zostera marina
Zostera noltei
Fam
Faba
Faba
Faba
Faba
Faba
Faba
Faba
Faba
Faba
Faba
Faba
Faba
Apoc
Apoc
\ do
Vio
Vio
\ do
Vio
V io
V io
Vio
Vio
Vio
Vio
Vio
Vio
Vise
Poac
Poac
Poac
Poac
Poac
Camp
Lemn
Wood
Wood
Zann
Poac
Zost
Zost
Zost
NS
N
AN
N
N
N
N
N
N
N
N
N
AN
AN
AR
AR
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
N
AR
AN
N
N
N
N
N
AC
N
N
N
EN
EN
VU
CS
RS Chg.. Hght Len
n -0.37
120
100
n 0.05
80
n -0.36
10
s -0.85
50
s -0.34
60
s -1.05
60
n 0.19
90
n -0.43
60
n -0.7 1
150
n 0.45
60
150
1.49
35
0.48
15
-0.29
40
n -0.87
18
n -0.46
15
r
10
s -1.08
15
n -0.69
20
n -0.19
15
n -0.30
15
r -0.62
25
n 0.20
13
n
1.07
18
r
5
n -1.52
20
n 0.97
100
n
0.18
37
s
0.78
37
s
0.37
55
1.55
62
-0.56
35
n -0.30
5
s -0.03
0.1
8
r
0.11
r -0.10
10
n 0.17
50
300
s -0.68
30
n -0.86
50
s -0.51
12
P1 P2 LF1 LF2 W Clonel
p
hc
Rhir l
a
Th
0
a
Th
0
a
Th
0
a
Th
0
p
hc
0
a
Th
0
a
Th
0
p
hc
Rhizl
p
hc
Rhizl
a
Th
0
a
Th
0
p
Ch
Tip
Ch
p
Node2
a
Th
0
p
hc
0
hc
0
P
a
Th
0
hc
0
hc
Rhiz2
hc
Sto12
hc
Rhiz2
hc
0
hc
0
hc
0
hc
0
p Th hc
0
Ch Pn w 0
Th
0
a
Th
0
a
Th
0
a
Th
0
a
Th
0
p
hc
Node2
p
Fly
Frag
p
hc
0
p
hc
0
p
Hy
Irreg
a
Th
0
p
Hy
Rhiz2
p
Hy
Rhiz2
p
Hy
Rhiz2
Rhiz2
Root
Rhir l
3
3
2
2
2
3
4
4
3
3
c Eur
Eur
Eur
Cro
Origin
4
4
4
3
1
3
3
3
4 c
3
3
5 c
3
3
2
1
1
4
2
Eur
1
4
6
6
6
Crop
? ?
6 6
8 5
7
5
7
8
7
4
7
5
7
7
7
7
7
7
9
9
9
8
9
8
8
2
2
8
7
7
9
7
9
8
5
5
7
8
Clone2 El E2 C
5 5
GB
2647
377,
1925'
501
58
214
136
1930
2615
623
1159
147,
1336
1359
2065
1030.
964,
2
183
536
1340
1880
18
1128
274 1
7
1691
923
1777
109
109
1134
39
314
48
22
14
1196
92
131
296
159,
CI Tjan Tjul Prec
10
3.6 14.5 1083
2 3.71 16.1
734
12 3.7 15.2
926
808
11 3.8 15.2
7 4.8 15.9
844
0 3.0 13.8 1410
695
0 3.9 16.3
991
14
3.7 14.9
4 3.5 14.5 1095
0
3.1 14.3 1119
10
3.9 15.9
804
3 3.8 16.1
757
12 4.0 15.6
873
8
3.7 15.3
893
11 3.6 15.0
940
3.5 14.7 1008
3.7 15.71 795
6.7 16.6
8 14
2
5.0 15.8 1025
2.0 13.3 1281
1
9
3.8 15.7
833
56
3.3 13.9 1262
1
3.9 15.4
826
28
3.8 15.5
869
96 1
3.5 14.4 1112
1,6 13.3 1393
24
3.4 14.6
992
3.8 15.9
788
51 1
3.8 15.0 1006
4.0 16.3
716
1
5.0 15.8
9 11
11 i
3.9 15.6
866
3.6 16.2
705
4
4.1 14.9 1280
4.4 16.6
752
0.4 11.6 237 1
1.3 12.4 2 108
20
3.8 15.5
820
1 1 4.0 15.9
820
27 6 4.3 15.0
926
68 13 4.5 14.6 1156
36 4 4.2 14.9 1020
IR
938
5
285
27
0
15
0
472
945
57
0
2
231
12
33
18
1
Co
Co
Co
Co
Co
Co
Br Habitats
3, 11
4, 17
3, 6
8, 19
18, 19
16
3, 4
6
3, 6
1, 3, 16
3, 4, 6
3, 4
1, 3, 17
1, 3, 17
4
8, 10
7
19
10
7, 8, 16
7
11, 14
11
1
1, 7, 16
7
4, 8, 19
3, 17
3
8
19
3, 17
3, 7
11, 14
13
15, 16
16
13, 14
4, 17
21
21, 23
21
L
7
8
7
8
7
7
7
7
6
7
7
7
5
4
8
8
7
9
7
8
5
7
7
4
6
8
8
7
8
9
9
8
9
6
7
7
7
7
8
7
6
8
F R
6
4
5
3
4
5
5
4
5
5
5
4
6
6
4
4
4
3
6
5
5
9
8
6
5
3
4
5
4
2
3
3
3
8
11
4
3
12
3
12
12
11
N $
5 0
71 0
6 0
3 0
5 1
4
5
4
6
5
6
5
6
7
6
2
2
2
2
2
7
2
3
5
4
2
4
5
3
2
2
3
2
3
7
2
2
7
7
5
6
5
A PPE NDIX
Sources or Mot hei ht
Plant heights have been calculated from a range of sources and have been subject to personal
adjust ment . T he main sources were Clapham,Tut in & Warburg ( 1962), Stace ( 199 1, 1997) and
Sell & Murrell ( 1996). Values that were thought to be too big, especially the height of
procumbent species, were modifi ed using informat ion in Butcher ( 1961).
A nearly complet e set of additional height values was taken from sources including Flora lberica
(Cast roviejo et al., I 986-), a Czech fl ora (Dostal, 1958) and BSBI handbooks such as Crucifers of
Great Britain and Ireland (Rich, 1991). Other such sources included Aeschimann & Burdet ( 1994),
Hutchinson & Thomas ( 1966), Jermy, Chater & David ( 1982), Meikle ( 1984), Page ( 1997) and
Preston ( 1995). The new values were checked against t hose derived fro m the main sources. For
t hose species w here t here was a diff erence of more than a factor of 1.5 between the two
sources, the discrepancy was resolved by making a judgement as to which seemed most suitable
for British conditions. In addit ion, all heights of pteridophytes and of trees and shrubs exceeding
5 m have been critically examined.
A diffi culty w ith creeping or procumbent plants is that the fl oras give shoot lengt h but not plant
height. For several such plants, the canopy height was estimated by Hill, using descriptions and
personal exper ience. Likewise Preston estimated t he heights of Aizoaceae (cf Preston & Sell,
1989).
Many diffi culties in assigning height to a plant remain. The height of a fern is taken to be the
length of its fronds. Scapose plants such as Pinguicula lusitanica may be very small except for their
scape. In several scapose genera (e.g. Bellis, Pilosella, Pinguicula, Plantago, Primula, Pyrola), t he length
of the leaves is used as a measure of size. O n the other hand, rosette plants w ith substantial
infl orescences (e.g. Sempervivum tectorum) have been accorded t he height of their infl orescence
rather t han t he lengt h of t he leaves. It was impossible to be completely consistent. Maximum
height is a general indication of the size of the plant rather than a precise measure.
The result ing table is inevitably a complex result of cited values and personal judgement. It was
clear that aut hors repeat each ot her and that their informat ion is oft en w rong. Overestimates
abound because botanists take pleasure in ext reme values. For example, British authors and Flora
Europaea (Tutin et al., 1964 - 1980) give t he height of Ulmus glabra as 40 m, higher than any ot her
elm. This is clearly unrealistic, but authors' copying of height data has meant that the value 40 m
is constantly repeated. Fort unately Bean ( 1950- 1951) indicates a more realist ic value of 30 m,
w hich has been used here. We have in many ot her cases t ried to drive height s down from
ext reme to typical values, by selecting the publication w ith the lowest values.
De tuition o Ii e orms
Life forms (Raunkiaer, 1934) are an example of what are now called 'plant funct ional types' (cf.
Box, 1996). A modifi ed version of Raunkiaer's system was set out by Clapham,Tutin & W arburg
( 1962), w ho specifi ed a life form for almost all species. Their assignments are a valuable starting
point but contain many inconsistencies. For PLA NTATT, they have been comprehensively
revised. T he categories were part ially redefi ned (Table A l ).
61
Table A I . Life for m categor ies from C lapham ,Tut in & W arburg ( 1962) and revised
categories
Category
(CTW )
Defi nition (CT W )
PLA N TAT T
Defi nit io n (dat ab ase)
Phanerophyte,
mega-, meso-
>8 m
Ph
Mega-, meso- and microphanerophyte
Phanerophyte,
micro-
2-8 m
Ph
(combined with taller phanerophytes)
Phanerophyte,
nano-
25 cm-2 m
Pn
Nanophanerophyte
Chamaephytes
0-25 cm
Ch
Chamaephyte
Hemicryptophytes
W int ering buds at soil
surface
hc
Hemicryptophyte
Geophyt es
Herbs with wintering
buds below t he soil
surface
Gb, Gn
Geophytes
Helophyt es
Marsh plants
(not recognized)
Hydrophytes
Water plant s
Hy
Perennat ing buds submerged during
wint er
Hz
Annual water plant
Th
Annual land plant, including t hose living
on summer mud
Therophytes
Passing unfavourable
season as seeds
The redefi ned categories are generally self-explanatory. A s the database contains plant heights,
the dist inctions between phanerophytes and chamaephytes, and distinct ions within
phanerophytes, are not ent irely necessary. If required, new categories can be derived using
combinations of character's. For example German authors such as Mueller-Dombois & Ellenberg
( 1974) generally allow chamaephytes to be up to 50 cm tall and distinguish a category of dwarf
shrubs (denot ed by Z - Zwergstraucher). These are woody or semi-woody chamaephytes. It
would be possible to reconstruct t his category fro m our database, allowing for t he fact that
ordinary maximum heights of many of these bushes might be up to 100 cm.
Ot her categories included by Mueller-Dombois & Ellenberg ( 1974) are lianes, epiphytes and
errant hydrophytes (unattached water plants). They defi ne Hemicryptophytes as plants that die
back to near the ground in the unfavourable season. Although worked out in some detail, their
scheme remains somewhat tentative and we have not adopted it.
Exam les o how li e- orm criteria are a lied
The criteria given above result in many discrepancies between our defi nit ions of life for m and
those of Clapham,Tutin & Warburg ( 1962). For example, t he distinction between geophytes and
62
hemicryptophytes is oft en hard to apply w ith rhizomatous perennials. In principle, plants whose
new shoots penet rate the surface before the advent of winter are hemicryptophytes, while those
w hose shoot s remain well buried are geophytes. Thus Phragmites australis, with a relatively deep
rhizome (Preston & Croft , 1997), has the life forms perennial Hy and Gn. In Carex acutiformis,
however, many new shoots have already emerged by t he autumn (Preston & Croft, 1997). It is
also less strongly aquatic. It is t herefore given the life for ms hc and Hy.
In general, biennials have been classifi ed as hemicryptophytes, but a few such as Euphorbia lathyris,
which are elongated from the beginning, are chamaephytes.
It is inst ructive to consider some ot her cases (Table A2), w hich will not be discussed in detail.
Table A 2. Som e cases where life-for m ( L F) is difficult t o apply or where our concept
diff ers from t hat of C lapham,T ut in & W ar burg ( 1962)
Links between Ii e orms and lent hei ht
Because chamaephytes and our two categories of phanerophytes are defi ned on the basis of
height, an eff ort was made to defi ne life forms consistently by reference to plant height (Table
A 3). Sometimes, as in Artemisia campestris, t he overwintering buds are at a substantially lower
height t han the height of the mature plant. Therefore, this scheme has not been followed exact ly
but has been used as a guide.
63
These defi nitions are necessarily imprecise. However, knowing that Ca!tuna vulgans has a typical
max imum height of 60 cm suggests that it will often grow as a chamaephyte and oft en as a
nanophanerophyte. This is indeed the case; C. vulgaris is the classic dwarf shrub.
Table A 3. C ham aephyte and phanerophyt e cat egories in relat ion t o plant height
Typical maximum
plant height (cm )
Prim ary life form
3-39
Ch
40-69
Ch
70-299
Pn
300-399
Pn
400-
Ph
Secondar y life for m
Pn
Ph
Cionalit y
Clonality can be defined in various ways. In an enumeration for Cent ral Europe, Klime§ et al.
( 1997) defi ned clonal growt h as synonymous with vegetat ive multiplicat ion, which results in t he
production of new, genetically identical descendants (ramets) w ith the potential to become
independent of t he mot her organism. According to this definition, plants that produce a new
ramet each year but do not spread laterally (for example many orchids) are clonal. Klime§ et aL
recognize 2 1 diff erent clonal t ypes. These are described in Klimet 's excellent website Clopla
(http://wwri butInn.cas.cz/klimes/clopla .htm); all species of Central Europe, including the majority of
British and Irish plants, are assigned to one or more clonal types.
Here we adopt a more familiar defi nit ion of clonal growth, namely vegetative reproduction
combined with lateral spread —i.e. spreading through the formation of clonal patches or clumps,
or through fragmentation and subsequent dispersal of plant part s.
Broad H abit ats
The Broad Habitat Classifi cation, developed as part of the UK Biodiversity Action Plan (Jackson,
2000; UK Biodiversity Group, 1998; UK Biodiversity Steering Group, 1995), provides a
comprehensive framework for surveillance of the UK countryside.We categor ized the habitats of
plants by t heir preferences for Broad Habitats.
In the UK Biodiversity Action Plan (UK Biodiversity Steering Group, 1995), 37 Broad Habitat
categories were defi ned. These included some habitats, such as limestone pavements, that are
quite narrow but were specified because t hey are priority habitats for conservat ion. In a later
publication (UK Biodiversity Group, 1998), the number of terrest rial and freshwater Broad
Habitats was reduced to 17. Limestone pavements were included in t he Broad Habitat inland
rock. Ten coastal and marine Broad Habitats were added shortly aft erwards (UK Biodiversity
Group, 1999). Out of the 27 Broad Habitats, 2 1 support vascular plants (see Table 8, above).
64
Broad Habitats are intended to be comprehensive and exclusive. In other words, every place in
which a vascular plant is found ought to belong to one and only one Broad Habitat. Although
the defi nit ions of Broad Habitats are in many cases obvious, t here was a need for more precise
circumscript ion to defi ne t he boundaries between t hem. For terrest rial and freshwater habitats
this guidance is now available (Jackson, 2000). The guidance not only gives writt en defi nit ions, but
relates Broad Habitat s to the units of t he National Vegetation Classifi cation (NVC; Rodwell,
199 1-2000).
Jackson's (2000) notes for guidance are the basis of the cross-reference and defi nitions used here
(Table A4). In a small number of cases, we have diverged from her interpretations. The basis of
our disagreement was oft en the scale at w hich we wished to view the vegetation. Thus we have
t reated bog pools as st rictly bog features (not as open water) and we treat rock-free maritime
grassland as neut ral grassland (not as supralitt oral rock). Ot her interpretat ions are possible but
result in fl orist ically less homogeneous Broad Habitats.
Table A 4. A ssignment of N V C com munit ies t o Broad H abit at s for t he purpose of
ascribing habit at preferences. Sequential runs of numbers are indicated by hyphens, e.g. 2 123, which signifi es t he sequence 2 1,22,23. Some NVC communit ies were assigned to two Broad
Habitats; w here this is t he case, the community is marked by an asterisk, to show that it is also
listed under another Broad Habitat.
No
N am e
N VC com munities
Broadleaved, mixed and yew
woodland
W I - 17,19
2
Coniferous woodland
W I8
3
Boundary and linear feat ures
W 2 I - 23,24*; OV I 8*, I 9*,2 1*,22*,24*,25*,27
4
A rable and hor t icult ural
O V I - 11, 13, 14- 17, 18* , 19* ,2 1* ,22*,36
5
Improved grassland
MG6*,7; OV 12,23*,25*
6
N eutral grassland
W 24; MC9- 12; MG I - 5,6*; MG8 - I 3; 0 V28*
7
Calcareous grassland
C G I - 14
8
Acid grassland
U I - 6,19
9
Bracken
U 20; W 25
10
Dwarf shrub heat h
H I- 10,12,16,18,2 1; M I5*,16; 0 V 34
11
Fen, marsh and swamp
M4- 14,22-24,25*,26-38; S - I 9,20*,22-28; OV 26,28*,29-
12
Bog
M 1-3, 15*, 17-21,25*
13
Standing wat er and canals
A I - 7,8*,9*,10,( 11- 16)* , 19* ,20* , 21- 24
14
Rivers and st reams
A 8*,9*,( 1I - I 6)*,17,18,19*,20*
15
Montane habitat s
H I 3- 15,17,19,20,22; U 7- I5,18; W 20
16
Inland rock
U 16, 17,2 1; 0 V37-40,4 I*
17
Built-up areas and gardens
0 V 20,23* ,24* ,4 1* ,42
18
Supralitt or al rock
MC 1- 8
19
Supralitt or al sediment
H I I; SD I - 19
21
Litt or al sediment
520,21; SM I -28
65
It is pertinent here to consider the defi nitions, drawing attention to less obvious features and to
where we have diff ered from Jackson (2000).
BH I Broadleaved, mixed and yew woodland
Broadleaved woodland is defi ned as woodland w ith broadleaved trees having at least 20% canopy
cover. In practice, most woodland has closed canopy. Juniper scrub is included in this type if it is
large enough to result in a woodland understorey (W 19). Other scrub types are treated as edge
features (BH3), because hedges are often where they occur. The underscrub community W 24 is
shared between woodland (BH1) and neutral grassland (BH6).
8 H 2 C oniferous woodland
This category is clearly defi ned and corresponds to nat ive pine woodland (W 18) in the NVC.
Stands of planted conifers normally have a subset of the normal broadleaved woodland fl ora
growing under them. There is no corresponding NVC type. Planted conifers have therefore
been under-represented, and the fern Dryopteris dilatata, which is t he commonest vascular plant
in upland conifer plantations, did not appear as a conifer specialist in our fl oristic database, but its
preference was added by us later
BH 3 Boundary and linear feat ures
Apart fro m scrub that forms hedges, boundary and linear features are not directly crossreferenced to the NVC. They include hedges, t ree-lines, walls, earth banks, grass strips and dry
ditches. Note that roads, t racks and railways in urban areas belong to the urban BH I7.
BH 4 A rable and hort icultural
Commercial orchards are included here, as well as fi eld crops. Note that hort iculture here is
defi ned as commercial hort iculture and does not include domestic gardens and allotments, which
are included in BH 17. The NVC communit ies classifi ed as OV 1- 11,13- 19,2 1,22,36 are included
here.
BH 5 Im proved grassland
This has few characteristic species, and is typically dominated by sown ryegrass Lolium perenne or
clover Trifolium repens. OV I2, the Poo annua-Myosotis arvensis community, is indicated in British
Plant Communities (Rodwell, 199 1-2000) as being mainly a disturbed element in improved pasture
and has been included here.
8 H 6 N eut ral grassland
The mesot rophic grassland types MG I-6,8- 13 are the core of t he neutral grassland category.
Coastal grazing marsh is included here. Four of the coastal cliff grasslands, MC9- I2, are not
necessarily or normally rocky and have been included here in BH6. OV28, an inundation
grassland, is shared between BH6 and BH 1I.
BH 7 Calcareous grassland
By convention, this includes all NVC types included in the CG class. As a result, montane
calcareous vegetation is in BH7, along with chalk grassland.
BH 8 A cid grassland
Only lowland and subalpine grassland are included here; vegetation types that would normally be
found above the timberline are in BH I5. Communities U I-6 form the core of this type. The
fern community U 19 is treated in BH8. It is usually a mosaic element in grassland, though it also
occurs on steep banks by rivers and could therefore be included also in the broad defi nit ion (not
used here) of BH I4.
66
BH 9 Bracken
This is small type, defi ned as unwooded land dominated by bracken Pteridium aquilinum. 8H9 has
few characterist ic species, because it is usually a derivative of grassland or forest. Just two NVC
types, U20 and W 25, are included here.
BH 10 Dwarf shrub heat h
T he heath type BH 10 includes vegetation w ith at least 25% cover of Ericaceae or dwarf gor se
Ulex minor. Coastal heath is included in t his categor y, except for dune heath, which is assigned to
BH I9. Dwarf shrub communit ies on blanket bogs are st rictly included in BH I2, but this is a
relat ively minor habitat for them and has not been recognized in t he cross-reference. The
standard heathland communities H 1- 10,12,16,18,21, toget her with the wet heath communities
M 15,16 for m the core of this community. The bizarre 0 V34, a chive M um schoenoprasum
community forming a mosaic element on shallow soils in coastal heath, is included here.
B14 1I Fen, m ar sh and swam p
T his is a very w ide category and includes reedbeds, swamps, tall-herb fens, fl ushes, springs,
marshes, rush-past ures and wet grassland. Mud communities of dried-up ponds and riverbeds
(e.g. 0 V30) have been included here, although they belong more st rictly to BH I3 and BH I4
unless they are very ext ensive (>0.25 ha). Types M4- 14,22-38, all swamp communities except
Scirpus maritimus S2 1, and OV 26,28-33,35 are included. We have excluded the bog-like
communities (Erico-Sphagnion) that are found in very acid lowland valley-mires.
BH I 2 Bog
Bogs are defi ned strictly only as ombrot rophic bogs. However, we have lumped all EricoSphagnion vegetation, including M21, in BH 12 even if it the vegetation occurs in valley mires,
toget her with bog pool vegetation M 1-3. This is to prevent the characteristic species of BH 11
being too heterogeneous. The NVC types included are M1-3, 15,17-21,25.
BH I 3 Standing w at er and canals
O nly aquatic vegetat ion has been cross-referenced to this type, i.e. communit ies A l - 16,19-24.
Mud communities and marginal vegetation are referred to BH 11.
BH I 4 Rivers and st rea ms
This Broad Habitat is t reated in much the same way as BH I3. According to t he strict defi nition,
all vegetation between t he banks of a river counts as belonging to BH I4. For the purposes of
defi ning characteristic species, river-bank vegetat ion has been t reated as belonging to other types
such as inland rock. Thus t he communities included in BH I4 are A8,9,11-20. Several of these
communities are also characteristic of B1-113.
BH 1S Mont ane habit at s
This type is defined by having distinctive arctic-alpine species, provided that they are not
calcicolous. Calcareous montane vegetation is included in BH7. BH 15 includes montane heaths
H I3- 15,17,19,20,22, grassier types U7- I5,18, and a willow scrub type W 20.
BH I 6 Inland rock
This heterogeneous Broad Habitat includes limestone pavement, cliff s, caves, scree, quarr ies and
vegetation on skeletal soils over rock. Communities U 16,17,2 1 and 0 V37-4 1 are assigned to
BH 16.
BH I 7 Built -up areas and gardens
Urban habitats were largely ignored by t he NVC surveyors. Most of the characteristic species
are neophytes, w hile t he commonest species are widespread nat ives and archaeophytes (Hill, Roy
67
& Thompson, 2002). Very few NVC communities are really characteristic of the built
environment. Pavement and wall communities 0 V20,42 are assigned wholly to BH I7;
0 V23,24,4 I are assigned to BH 17 but shared with other Broad Habitats.
BH 18 Supralittoral rock
It is clear fro m the Habitat Statement for supralitt oral rock (UK Biodiversity Group, 1999) that
maritime grassland is not included. For that reason, MC9- 11,12 have been treated as neutral
grassland, BH6. Only MC I-8 are included in BH I8.
BH 19 Supralitt oral sedim ent
This includes strandlines, shingle, machair and coastal dunes. It includes all the NVC types SD I19, toget her w ith the dune heath type H I I .
BH 20 Lit toral Rock
Intert idal rock normally lacks vascular plants and no vascular-plant species or vegetat ion type is
characterist ic of it.
BH 2 I Lit t oral sediment
This includes saltmarshes and saltmarsh pools.
BH 23 Inshore sublittoral sedim ent
One vascular plant. Zostero marina, occurs in this habitat, which is not considered furt her
Initial calculation o s ecies' re erences or Broad H abitats
For the majority of species, habitat preferences were based on quadrat data. Two major datasets
were available, the or iginal data used to derive the association tables in British Plant Communities
(NVC; Rodwell, 199 1-2000) and quadrat samples collected for Count ryside Survey 2000,
CS2000 (Haines-Young et aL, 2000).
The quadrat data used to create the NVC categories were supplied by JNCC as an MSAccess
database. This had been created from fi les in the format of the VESPAN package (Malloch, 1985).
There was no information on the NVC community to w hich each quadrat had been assigned.
Therefore, the computer program TABLEFIT (Hill, 1996) was used to assign 312 16 quadrats to
NVC communities using species' Domin scores. The median goodness-of-fi t was 67%. O nly four
samples had no fi t; t hese were assigned to a community by personal judgement . NVC plant
communit ies were then attributed to Broad Habitats using the cross-reference outlined above
(Table A4). All CS2000 samples had been allocated to BAP Broad Habitat types in the fi eld.
Aft er quadrats had been assigned to Broad Habitats, species' frequencies in each Broad Habitat
were calculated. Frequencies were used to calculate preferences for Broad Habitats. For each
species s and BH h, we calculated an odds-ratio index:
= observed odds for species s in habitat h / expected odds
Qsh
nsh
/ (ns+ "sh + 0.5)
/
esh
Os+ esh)
where
nsh = observed number of occurrences of species s in BH h
ns+ = 1h nsh = total number of occurrences of speCies s
esh = expected number of occurrences of species s in BH h.
68
The expected number of occurrences
esh
= ns N h /
N
+
w here
Nh = number of quadrat samples in BH h
N 4.
= Eh N h
=
total number of quadrat samples.
Species s was deemed to have a preference for BH h if
4.
Qsh
Ext ension and validation o s ecies' re erences
Many rare, scarce or uncommon native species are not represented either in the NVC quadrats
or in CS2000 samples. Furt hermore, t here was the possibility that uneven sampling of Broad
Habitats could have resulted in some w rong indications of preference.
First, we ext ended the list of BH preferences to species not adequately sampled by the NVC o r
CS2000, using t he text prepared for the New Atlas, together with other literature sources (e.g.
Stewart , Pear man & Preston, 1994; W igginton, 1999). Much of t he initial work was done by Bill
Meek (CEH); t he resulting table of preferential species was checked by D.A. Pearman (BSB1), as
well as by ourselves and ot her CEH staff . In addition, preferences were checked against ot her
data in PLA NTATT For example, species attributed to BH I3 (Standing water and canals) and
BH 14 (Rivers and streams) were checked for their Ellenberg moisture (F) values. Those for
which F<8 were scrutinized carefully. Some were excluded from t he list of characteristic species
of these Broad Habitats.
69
A CKN O WL ED GE MEN TS
Much of the work presented here was support ed by funding from the W ildlife and Countryside
Division of Defra and its predecessor DET R. The immediate stimulus was a project to interpret
the New Atlas of the British and Irish Flora (Preston et al., 2003). A requirement for non-spatial
attributes of plants was writt en into the 2002 programme of wo rk for the National Biodiversity
Network (N BN). Informat ion on native status was prepared for the New Atlas and presented
there (Preston, Pearman & Dines, 2002). The Ellenberg indicator values were developed under
the DETR-funded ECO FACT programme and published separately (Hill et al., 1999).
The ot her main ot her source of funding was from our parent body the Natural Enviro nment
Research Council. N ot only have they generously support ed the Biological Records Centre
(BRC) and N BN over the years, but they have allowed us to fi nd additional t ime to work our
data up into publishable form. The geographical classifi cat ion of European distributions (Preston
& Hill, 1997) was developed with support from N ERC's special programme on global
environmental change,TIGER.
Some elements of t his work have been funded by t he Joint Nat ure Conservation Committ ee
through t heir joint sponsorship of BRC. Other aspects were suppor ted by t he Countryside
Council for Wales, English Nat ure, Scottish Natural Heritage, Environment and Heritage Service
(No rthern Ireland) and Plantlife as part of the analysis of New Atlas data.
Publication of this booklet has been jointly funded by CEH and Defra.
We are grateful to the following for particular help:
De fra
Andrew Stott (overall guidance and long-term support)
Debbie Boobyer (Broad Habitats and analysis of New Atlas data)
BSBI
David Pearman (Broad Habitats)
Cambridge University
Peter Sell (unpublished lf ora draft s)
English N ature
Roger Key (Cover photograph)
JNCC
Ian St rachan (NVC quadrat data)
CEH
Bob Bunce (Ellenberg values)
Jon Cooper (liaison with N BN)
Julie Gaunt (graphic design)
Bill Meek (Broad Habitats)
Owen Mountford (Ellenberg values, European fl oras, species attributes)
Peter Rothery (Change index)
Mark Telfer (Change index)
70
B I B L I OGR A PH Y
Aeschimann, D. & Burdet, H.M. ( 1994) Fiore de la Suisse. Editions du Gr iff on, Neuchatel.
Barrow, E.M., HuIme, M. and Jiang,T. ( 1993) A 196 1-90 baseline climatology and future climate
change scenarios for Great Britain and Europe. Part 1: 1961-90 Great Britain baseline climatology.
Climatic Research Unit, N orw ich, 43pp. (plus maps).
Bean,W.J. ( 1950- 195 1) Trees and shrubs hardy in the British Isles. 7 edn.John Murray, London.
Box, E.O. ( 1996) Plant functional types and climate at the global scalej ournal of Vegetation Science,
7, 309-320.
Butcher, R.W. ( 196 1) A new illustrated British lf ora. Leonard Hill, London.
Castrovielo, S. et al., eds. ( 1986-) Flora iberica: plantas vasculares de la Peninsula iberica e Islas
Baleares. Vol. 1-. Real Jardin Botanico, Madrid.
Cheffi ngs, C. (2004) N ew plant status lists for Great Britain. 8581News, 95, 36-43.
Clapham,A.R.,Tut in,T.G. & Warburg, E.F. ( 1962) Flora of the British Isles. 2 edn. Cambridge
University Press, Cambridge.
Dostald . ( 1958) Klic k Opine kvetene CSR. 2 edn. Ceskoslovenske Akademie Ved, Prague.
Ellenberg, H.,Weber, H. E., Dull, R.,W irt h,V ,Werner,W. & Paulissen, D. ( 199 1) Zeigerwerte von
Pfl anzen in Mitt eleuropa. Scripta Geobotanica, 18, 1-248.
Fitt er, A.H. & Peat, H.J. ( 1994) The Ecological Flora Database.Journal of Ecology, 82, 4 15-425.
Gr ime, J. R ( 1979) Plant strategies and vegetation processes. John W iley, Chichest er.
Gr ime, J. P. (2001) Plant strategies, vegetation processes, and ecosystem properties.W iley, Chichester.
Gr imel .R,Thompson, K., Hunt, R., Hodgson,J.G., Cornelissen,J.H.C., Rorison, 1.H., Hendry, G.A.F,
Ashenden,T.W.,A skew,A .R, Band, S.R., Booth, R.E., Bossard, C.C., Campbell, B.D., Cooper,
J.E.L., Davison,A.W., Gupta, PL., Hall,W , Hand, D.W., Hannah, M.A., Hillier, S.H., Hodkinson,
D.J.,Jahli,A ., Liu, Z., Mackey, J.M.L., Matthews, N ., Mowforth, M.A., Neal,A .M., Reader, R.J.,
Reiling, K., Ross-Fraser,W , Spencer, R.E., Sutton, F.,Tasker, D.E., Thorpe, P.C. & W hitehouse, J.
( 1997) Integrated screening validates primary axes of specialisation in plants. Oikos, 79, 259281.
Haines-Young, R.H., Barr, C.J., Black, H.1.J., Briggs, D.J., Bunce, R.G.H., Clarke, R.T , Cooper, A.,
Dawson, F.H., Firbank, L.G., Fuller, R.M., Furse, MI , Gillespie, M.K., Hill, R., Hornung, M.,
Howard, D.C., McCann,T , Morecroft , M.D., Petit, S., Sier,A.R.J., Smart , S.M., Smith, G.M., Stott,
A.R, St uart , R.C. & W atkins, J.W. (2000) Accounting for nature: assessing habitats in the UK
countryside. Depart ment of the Enviro nment,Transport and the Regions, London.
Hill, M.O. ( 1996) Tablefit version 1.0, for identification of vegetation types. Institute of Terrestrial
Ecology, Huntingdon.
Hill, M.O., Mountford, J.O., Roy, D.B. & Bunce, R.G.H. ( 1999) Ellenberg's indicator values for British
plants. ECO FACT Volume 2 technical annex. Institute of Terrestrial Ecology, Huntingdon.
Hill, M.O., Roy, D.B. & Thompson, K. (2002) Hemeroby, urbanity and ruderality: bioindicators of
disturbance and human impact. Journal of Applied Ecology, 39, 708-720.
Hodgson, J.G., Gr ime, J.R, Hunt, K. & Thompson, K. ( 1994) The electronic comparative plant ecology.
Chapman & Hall, London.
Hodgetts, N ., Palmer, M. & W igginton, M. ( 1996) The Pink Book of Plants: lists of vascular and nonvascular plant species which are nationally threatened, localised or protected in Great Britain.
JN CC, Peterboro ugh.
HuIme, M. & Jenkins, G. ( 1998) Climate change scenarios for the United Kingdom: summary report
Climate Research Unit, Norw ich.
Hutchinson, G. & Thomas, B.A. ( 1996) Welsh ferns: clubmosses, quillworts and horsetails. 7 edn.
National Museums and Galleries of Wales, Cardiff .
Jackson, D.L. (2000) Guidance on the interpretation of the Biodiversity Broad Habitat Classifi cation
(terrestrial and freshwater types): Defi nitions and the relationship with other habitat classifi cations.
JN CC Repor t No. 307.JNCC, Peterborough.
Jermy, A.C., Chater, A.O. & David, R.W. ( 1982) Sedges of the British Isles. Botanical Society of the
British Isles, London.
Klim8 , L., Klimegova J Hendriks, R. & van Groenendael,J. ( 1997) Clonal plant architecture: a
comparative analysis of form and funct ion. In The ecology and evolution of clonal plants (eds H.
de Kroon & J. van Groenendael), pp. 1-29. Backhuys, Leiden.
Lindacher, R. ( 1995) PHANART: Datenbank der Gefasspfl anzen Mitteleuropas. Vertiffentlichungen
des Geobotanischen Institutes der ETH, Stiftung RUbel, Zurich, I 25, 1-436.
Macpherson, P., Dickson,J.H., Ellis, R.G., Kent, D.H. & Stace, C.A. ( 1996) Plant status
nomenclature. M I News, 72, 13- 16.
Malloch,A .J.C. ( 1985) VESPAN: FORTRAN programs for handling and analysis of vegetation data and
species distributions. Depart ment of Biological Sciences, University of Lancaster.
Meikle, R.D. ( 1984) Willows and poplars of Great Britain and Ireland. Botanical Society of t he British
Isles, London.
Mueller-Dombois, D. & Ellenberg, H. ( 1974) Aims and methods of vegetation ecology. John W iley &
Sons, New Yor k.
Page, C.N. ( 1997) The ferns of Britain and Ireland. Cambridge University Press, Cambridge.
Palmer, M.A., Hodgetts, N.G.,W igginton, Mj ., Ing, B. & Stewart , N.F. ( 1997) The application to t he
Brit ish fl ora of the Wo rld Conservation Union's revised red list criteria and the signifi cance
of red lists for species conservation. Biological Conservation, 82, 2 19-226.
Perring, EH. & Walters, S.M. ( 1962) Atlas of the British Flora.Thomas Nelson & Sons, London.
Preston, C.D. ( 1995) Pondweeds of Great Britain and Ireland. Botanical Society of the British Isles,
London.
Preston, C.D. & Croft, J.M. ( 1997) Aquatic Plants in Britain and Ireland. Har ley Books, Colchester.
Preston, C.D. & Hill, M.O. ( 1997) The geographical relationships of Brit ish and Irish vascular
plants. Botanical Journal of the Linnean Society, 124, I - 120.
Preston, C.D. & Hill, M.O. ( 1999) The geographical relationships of the British and Irish fl ora: a
comparison of pteridophytes, fl owering plants, liverworts and mosses.Journal of Biogeography,
26, 629-642.
Preston, C.D., Pearman, D.A. & Dines,T.D., eds. (2002) New atlas of the British and Irish Flora.
Oxford University Press, Oxford.
Preston, C.D. & Sell, RD. ( 1989) The Aizoaceae naturalized in the British Isles. Watsonia, 17, 217245.
Preston, C.D.,Telfer, M.G.,Arnold, H.R., Carey, RD., Cooper, J.M., Dines,T.D., Hill, M.O., Pearman,
D.A., Roy, D.B. & Smart , S.M. (2002) The changing fl ora of the UK. DEFRA , London.
Preston, C.D.,Telfer, M.G., Roy, D.B., Carey, RD., Hill, M.O., Meek,W.R., Rot hery, P. Smart,
Smith, G.M.,Walker, K.J. & Pearman, D.A. (2003) Thec hanging distribution of the fl ora of the
United Kingdom: technical report Report to DEFRA on CEH project C0 1093. CEH Monks
Wood, Huntingdon.
72
Raunkiaer, C . ( 1934) The life forms of plants and statistical plant geography. Oxford University Press,
Oxford.
Rich,T.C.G. ( 199 1) Crucifers of Great Britain and Ireland. Botanical Society of the Brit ish Isles,
London.
Rodwell, J.S. ( 199 1-2000) British plant communities. Cambridge University Press, Cambridge.
Sell, P. & Murrell, G. ( 1996) Flora of Great Britain and Ireland. Volume 5 Butomaceae-Orchidaceae.
Cambridge University Press, Cambridge.
Stace, C.A. ( 199 1) New fl ora of the British Isles. Cambridge University Press, Cambridge.
Stace, C.A . ( 1997) New flora of the British Isles. 2 edn. Cambridge University Press, Cambridge.
Stewart, A., Pearman, D.A. & Preston, C.D., eds. ( 1994) Scarce plants in Britain. JNCC,
Peterborough.
Telfer, M.G., Preston, C.D. & Rothery, P. (2002) A general method for the calculation of relat ive
change in range size from biological atlas data. Biological Conservation, 107, 99- 109.
Tutin,T.G., Heywood,V.H., Burges, N A ., Moore, D.M.,Valent ine, D.H., Walters, S.M. & Webb, D.A.
( 1968- 1980) Flora Europaea. Volumes 2-5. Cambridge University Press, Cambridge.
Tutin,I G., Heywood,V.H., Burges, N.A.,Valent ine, D.H.,W alters, S.M. & Webb, D.A. ( 1964) Flora
Europaea. Volume I. Cambridge University Press, Cambridge.
UK Biodiversity Group ( 1998) Action Plan Volume 2:Terrestrial and freshwater habitats. English
Nat ure, Peterboro ugh.
UK Biodiversity Group ( 1999) Tranche 2 Action Plans.Volume S. Maritime species and habitats.
English Nat ure, Peterborough.
UK Biodiversity Steering Group ( 1995) Biodiversity: the UK Steering Group Report. Volume 1,
Meeting the Rio challenge; Volume 2: Action plans. HMSO, London.
W igginton, M.J., ed. ( 1999) British Red Data Books. I Vascular plants, 3 edn.Joint Nature
Conservat ion Committee, Peterborough.
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