G C A H O R T I C U L T U R E Basic Plant Propagation 2012 GCA Horticulture Committee This online manual is an expanded version of the GCA Basic Plant Propagation Handbook. The Handbook is available from GCA Headquarters and on the GCA Website (see page 15). The GCA Horticulture Committee hopes these publications will encourage you to start propagating plants! VEGETATIVE PROPAGATION Basic Plant Propagation 1 G C A H O R T I C U L T U R E Table of Contents ! ! Introduction! ! ! ! ! ! ! ! ! ! ! ! 3 ! General Rules of Propagating!! ! ! ! ! ! ! 3 ! ! Supplies: Basic Tools and Equipment!! ! ! ! ! ! 4 ! ! Vegetative Propagation! ! ! ! ! ! ! 5 ! ! ! Softwood Cuttings! ! ! ! ! ! ! ! ! 5 ! ! ! Rooting Hormones! ! ! ! ! ! ! ! ! 7 ! ! ! Hardwood Cuttings! ! ! ! ! ! ! ! ! 8 ! ! ! Semi-hardwood Cuttings! ! ! ! ! ! ! ! 8 ! ! ! Root Cuttings! ! ! ! ! ! ! ! ! ! 8 ! ! ! Division! ! ! ! ! ! ! ! ! ! 9 ! ! ! Leaf Cuttings! ! ! ! ! ! ! ! ! ! 9 ! ! ! Layering! ! ! ! ! ! ! ! ! ! 10 ! ! ! Grafting! ! ! ! ! ! ! ! ! ! 11 ! ! Appendix! ! ! ! ! ! ! ! ! ! 12 ! ! ! Glossary! ! ! ! ! ! ! ! ! ! 12 ! ! ! Websites! ! ! ! ! ! ! ! ! ! 12 ! ! ! GCA On-Line Links! ! ! ! ! ! ! ! ! 12 ! ! ! Further Reading! ! ! ! ! ! ! ! 13 ! ! ! Propagation Handbook Ordering Information! ! ! ! ! 15 2 ! ! Basic Plant Propagation G C A H O R T I C W H Y P R O PA G A T E General Rules of Propagating PLANTS? U L T Research Variety—Anything Is Possible! A little time spent researching the Growing your own expands your choices. plant species before you start will You can choose flowers for cutting, result in much heirloom varieties of flowers or vegetables, higher success or native plants to provide habitat for rates and the butterflies and birds. Internet makes Preservation this easy. A With an old or rare plant, taking cuttings quick on-line is a way to perpetuate the plant. A plant search, using grown from a cutting will be a clone, a Google or new plant with identical characteristics. another search Because most garden centers offer engine, will “popular plants” that sell in large save time and quantities, propagating from seeds and anguish. Just cuttings provides a chance to grow type in: “propagating” followed by uncommon plants, varieties that comthe name of the plant. Some seeds mercial growers and garden centers do need a cold treatment, some plants not find profitable. are almost impossible to propagate from stem cuttings or from seed, but Club and Zone Plant Exchanges or can easily be grown from a root Challenge Classes at a Flower Show cutting or a simple layer. Learn these skills so you can participate in plant exchanges and the propagation Cleanliness classes for flower shows. This is a fun Disease organisms can survive in the skill to acquire and share with other club bits of soil in used pots; plant viruses members, your children or grandchildren. can be spread from an infected plant You Control Chemical Usage to a healthy plant on your hands or Most commercial growers use chemicals by a knife or scissors. to control insects and disease and Be sure your hands, gloves, chemical growth regulators to keep the containers and tools are clean! plants short. When growing your own plants, you control any use of chemicals Wash with soap and water, then and can grow a sturdy, “green” plant. disinfect tools and containers with 1 TBSP Clorox/ Fun 1gallon of warm water. Growing from seed or cuttings is a great learning experience; you will have great Use new, sterile, soilless mix; this is success, and probably a few failures. You not the time to re-use the will have many plants to share with soilless mix from last summer’s friends! Your gardening skills will container. improve as you work with seedlings and Use Healthy, Well-Watered Plants for newly rooted cuttings, learning when to Cuttings, New or Well-Stored Seeds water and to recognize the signs of pests Many seeds will remain viable for or disease. years if stored in a plastic container in a refrigerator; some seeds must be Basic Plant Propagation U R E fresh. (Research your plant species, and for information on testing for viability.) Water Warm water is best. Cold water will lower the soil temperature, slowing germination or root formation. Water processed through a water softener has toxic amounts of sodium, which will kill plants. In a home with softened water, use untreated water, water from a rain barrel or an outside tap (that does not connect to the water softener). Timing Seed packets (or on-line sites, reference books) will give number of days to germinate, and number of weeks before planting. Check the “last frost” date for your area, and then count backwards on a calendar. Do not start too early! Seedlings will grow quickly, especially during the longer, sunnier days of April and May. For most annuals, 6–8 weeks after germination will produce a healthy, well-rooted plant. Tip: Plants will be much healthier in your garden if planted before they come into bloom. You may not get the instant satisfaction on the day you plant, but a stocky “green” plant will reward you with better garden performance all summer. 3 G C A H O R T I C U L T U R E Supplies Basic Tools & Equipment 1. 2. 3. Sterile growing medium—A commercial soilless mix (e.g. Pro-Mix, Fafard, Sunshine) works well for almost all seeds and cuttings; these are sterile, consistent, and available. (A local nursery growing their own plants often offers a quality soilless mix for sale, or Google a brand name to find a local source.) Avoid mixes with “continuous feeding.” Soilless mixes with up to 40% hardwood bark will help suppress disease organisms in the soil. By using the same mix, you will learn to judge when to water. Shallow containers 2"–3" deep with drainage holes: Bedding plant flats and inserts are useful; shallow pots also work. Deeper containers remain too wet (no air spaces). New roots will rot. 3”-4” containers work best for softwood and semi-hardwood cuttings of woody plants. 5. a. Spray bottle to spritz (mist); b. Watering can with a “rain-nozzle,” or “water-dispersing nozzle,” so the water doesn’t create gullies but is gently spread across the surface. Self-watering trays are a useful option d. Otherwise, large dishpan or aluminum turkey roasting pan for bottom watering. Light: If you will be growing the young seedlings for more than two weeks before planting outdoors, supplemental lighting is useful. Fluorescent lights are ideal (do not need specialty “gro-bulbs”). Fluorescent lights 16 hours per day, (with an 8-hour dark period) are helpful for most plants. Temperature control: You can go hitech, or improvise. There are specialty heat mats; or you may have a warm spot above a radiator, or on top of a refrigerator. A string of tiny Christmas lights under a Pyrex baking dish will create bottom heat. (LED lights will not work; use an old string of incandescent bulbs that give off heat.) 10. Small fan or cold frame: useful options to help harden off seedlings or cuttings For Cuttings 1. Clean cutting tool: one-sided razor, sharp knife, clippers, scissors 2. Rooting compound: Rootone, Hormodin, etc. 3. Potato peeler for “wounding” the bark of a cutting Additional Supplies for Cuttings For Layering: 1. Pebble or toothpick to hold open split in stem Peg (or large hairpin, piece of wire) to hold down shoot in soil 6. Labels, #2 lead pencil, notebook to record 2. 7. Plastic for tenting: Ziploc bag or a dry-cleaner bag, plus supports (almost anything works: wire hangers, popsicle sticks, bamboo stakes, plastic straws) to keep plastic about 4" above seeds or cuttings. Self-watering trays have a lid that serves this function. For Air Layering: Tool for marking rows, transplanting the rooted seedlings or cuttings—a butterknife, dibble, pencil For Grafting: Use warm water: c. 4 4. 8. 9. Fertilizer: use only when seedlings or cuttings are well established; liquid (chemical) fertilizers, timed-release, or organic forms—e.g. diluted sea weed emulsion 1. Spaghnum moss 2. Plastic to hold spaghnum around branch 3. Tape to hold plastic in place 1. Sharp grafting knife (clean) 2. Grafting rubbers (special wide rubber bands available from Nursery Supply stores) 3. Grafting wax or Tree-Kote, freezer tape, strips of Saran Wrap or Paraffin Basic Plant Propagation G C V E G E TAT I V E P R O P A G AT I O N A H O R T I C U L T Softwood Cutting U R E gently to remove excess (which could clog pores). You do not need a lot—just a light dusting. These are cuttings taken from current season’s growth (late spring or Vegetative propagation requires summer): use non-blooming or non- 4. Poke 1/2" holes in soil-mix with a pencil or dibble—not too deep! cutting a small part of a parent fruiting softwood when new growth is Place the cutting in (being plant and promoting root growth hardening or at the “snap stage.” careful not to scrape off the Depending on the plant, cuttings to create a new plant. The new rooting powder as you place it); should be 4"–8", ideally with 2 leaf plant will be genetically firm soil around it. (Make sure nodes. Annuals and tender perennials no leaves are under the soil. identical to the parent plant. are usually shorter, 3"–4", woody They would begin to rot, and plants 6"–8". This method of It can be as simple as placing those disease organisms could propagation works with almost all stems of ivy, pussy willow, or spread to the cutting.) species of annuals and tender coleus in water and seeing roots perennials and many woody plants as form in a week or two, or a very well. slow meticulous process. Before you start, do a quick on-line search to learn what type of vegetative propagation works best for the plant you are working with. Tip: Make sure the “parent plant” is well watered in advance; once the cutting is removed from the parent it cannot absorb more water until roots grow. Success depends on keeping the stem and leaves from drying out before new roots form. Take cuttings in the morning, place in a plastic bag to retain moisture—do not allow to wilt! Basic Plant Propagation 1. Fill container with commercial soilless mix; bedding plant inserts give each cutting its own pocket, preventing the spread of soil-born disease. Shallow pots also work. Water thoroughly; mix needs to be thoroughly moist. Tip: Cuttings longer than 4"–8" seldom take root; without roots to absorb water, the tip of a long cutting loses moisture (turgidity), and within a week, the entire cutting desiccates. 2. Cut a 4"–8" tip from the branch, below a node; remove any lower leaves that would be below ground, cut large leaves, such as rhododendron, in half. Minimize evaporation by minimizing the number and size of leaves. For woody plants, use a potato peeler or knife, very gently peeling a vertical strip at the bottom 1/4" of the stem in two spots, leaving part intact. This is called “wounding” and encourages root development. 3. Pour a small amount of Rootone or Hormodin (see “Rooting Hormones”, p. 6) into a cup. Dip cutting in—then tap Roots will form from the leaf nodes, the area on the stem where the leaves were removed. Drawing by Martha Kemp. 5 G C Tip: Even cuttings taken from the same plant will vary in the length of time to form roots. By using separate containers or cellpaks, it is easier to move the rooting plants from those still needing extra warmth and humidity. 5. Do not crowd cuttings in a single container—leaves should not touch. 6. Continue with your cuttings, insert labels with name and date, then spritz with water. 7. Tent with plastic, keeping it above the foliage, creating a warm, humid micro-climate. 8. Provide bottom heat to keep the soilless mix close to 75°, roots will form more quickly. This may be on top of a refrigerator, a radiator, or using a heat mat. (Disease pathogens thrive in cooler temperatures; providing bottom heat speeds up root formation and slows the growth of disease organisms.) A H O R T I C U L 6 U R E 13. Wait to transplant into a larger there is resistance, roots are container until roots on cuttings forming. To peek, carefully use a are about 1" (ideally a nice ball, dibble or label and lift from with multiple roots). Transplant below. to a 3"–4" container, water well, 11. As roots begin to form, reduce label, and keep out of direct sun the humidity—open the plastic for a few days (to help plant tent, and then remove it after 3– recover from transplant shock). 4 days. You want these new Choose a container in roots to grow into the soilless proportion to the root ball, mix, so water only when mix is providing 2"–3" inches of drying out (lift container to soilless mix around the roots. know when it is dry). 12. Immediately remove any cuttings that have lost all leaves, look squishy or sick, and remove the soilless mix from the immediate area if you have several cuttings in one container. This will prevent disease organisms from the “sick” cutting from attacking the other cuttings. (Or move healthy cuttings to a new container with fresh soilless mix.) Tip: Fertilizing too early or with a full strength dose in the first weeks, before roots are reaching the bottom of the container may injure or even kill your rooted cutting; a plant can only absorb fertilizer when the roots are healthy. (See The Real Dirt, Issue #16, “Fertilizing Indoor Plants.”) 14. Once plant is established, actively growing with healthy roots, use either organic or chemical liquid fertilizer at halfstrength in alternate waterings. 9. Spritz with warm water 2–3 times a week to keep humidity high. Lift containers to check weight, to know if it is drying out. Sub-irrigate (bottom water) if dry. Soilless mix needs to be moist, but not soggy. 10. After two weeks, begin to check for rooting. Some plants may root in two weeks, others may take months. Tug very gently. If T 15. After 3–4 weeks, if the roots have reached the bottom of the pot, cuttings may need to be transplanted to a larger pot. A well-rooted stem cutting of a Begonia cultivar. The cutting was provided with adequate moisture and bottom heat and the results show! 16. If the rooted cutting is now growing actively, fertilize at full strength in alternate waterings. Basic Plant Propagation G C A H O R T I C U L T U R E Other Methods of Vegetative Propagation The easiest methods for propagating plants are usually planting seeds and rooting softwood cuttings. There are, however, many plants that do not flower (and therefore do not have seeds), and the seeds from some plants do not produce a plant with the characteristics of the parent plant (e.g. if you plant an apple seed you will get an apple tree, but the apples it produces have a 99% chance of being very sour and inedible.) Some plants do not have the stems necessary for softwood cuttings (e.g. African violets, some begonias, many succulents). Over the centuries, other methods of propagating plants have been developed, usually because the desired plant will not reproduce from seed or softwood cuttings. The following sections describe propagation from Hardwood and Semi-hardwood Cuttings, Root Cuttings, Division, Leaf Cuttings, Layering and Grafting. Remember, to insure a more successful outcome, research your particular plant’s best method of propagation before beginning. Rooting Hormones The best way to propagate is to use the plant’s natural reproduction ability. Plants produce auxins in their leaves that serve as growth regulators, and can cause dormant cells to divide and become roots. In vegetative propagation, synthetic auxins (rooting compounds) are often used to stimulate the growth of adventitious roots that will form at the base of the cutting. Cuttings of some plants will form roots easily: willows, ivy, coleus. But rooting compounds (auxins) will help with most plants. Roots will form more quickly and the percentage of cuttings that form roots will increase. Rooting Compounds come in various strengths and formulations. Hormodin, Hormex, Rootone, and Hormo-Root all mix IBA or NAA in talcum powder. A light dusting of Basic Plant Propagation one of these powders will promote root growth for most softwood cuttings (cuttings of new growth). Most perennials, many houseplants, some trees and shrubs can be propagated using softwood cuttings, taken in late spring or early summer. For semi-hardwood or hardwood cuttings, a rooting compound with a stronger formulation is usually needed. These are liquid compounds, using IBA in 0.3% up to 1.6% strengths. Search on-line to find the concentration of chemical needed for the genus/species. Tip: Rooting compounds in the right strength will speed root formation, but these same compounds in higher concentrations will burn or even kill your cutting! No matter what type of vegetative propagation you are attempting, always remember to.... ...remove all buds, flowers or seeds from a cutting before attempting to root. All of the cutting’s growth activity needs to be concentrated on forming roots, not setting seed! ...take all cuttings from healthy, well-watered young specimens when possible. It is a well known fact that the “juvenility” of the parent plant has a positive effect on the success rate of cuttings. 7 G C A H O R T I C U L T cuttings in early winter and follow directions for softwood cuttings above, using a There are many different ways to do stronger rooting this type of stem cutting. This is the hormone. Some most basic method appropriate for species will home gardeners. Plants suitable for benefit from hardwood cuttings are most deciduous more extensive wounding of the shrubs and some trees such as willow. base. Read up on Vines such as honeysuckle (Lonicera), the specific grape (Vitus) and Wisteria are good needs of each candidates for hardwood cuttings. species. Semi-hardwood cuttings Semi-hardwood cuttings are most should be treated to bottom heat and often used for evergreens and conifers. a humid environment so tent or enclose the pots in plastic and slowly 1. Take pencil-width cuttings from remove after rooting. It may take 2 or current year’s growth with 3 or 3 months for roots to form. more leaf nodes, in late fall or early winter when plant is dormant and has lost its leaves. If a longer stem is taken to cut into Root cuttings are a good way to smaller pieces make sure each propagate any suckering woody plant cutting has at least 3 leaf nodes such as lilacs (Syringa) and many (around 6” long for most plants). perennials with fleshy root systems Be sure to make cuts indicating stem’s polarity: slanting cut below such as poppies (Papaver). a node for part of stem growing 1. In late winter or very early spring, closest to the roots of the plant, when the plant is dormant and straight cut for top of cutting; you before growth has commenced, don’t want to plant your cutting use a sharp spade, driving it into upside down! the ground one third of the way in 2. Apply hormone powder. from the outermost spread of the branches. 3. Label cutting and tie in bundles and bury outdoors or in cold 2. Cut off a section of roots. Pry frame to callus, cover with 6-8” of them up and pull gently from the sand, and mulch. soil. Rinse soil off roots. Hardwood Cuttings Root Cuttings 4. In spring, plant in ground or pot, straight cut up, with one bud showing. (Note: In climates with mild winters, hardwood cuttings may be planted directly into planting medium and placed outdoors.) Semi-hardwood Cuttings Semi-hardwood cuttings are most often used for non-deciduous plants, both broadleaf and conifers. Take 8 3. Make a straight cut on the proximal end of the root (roots growing closest to plant) and a slanted cut on the distal end of the root (roots growing farthest away from plant). It is very important to mark the roots polarity, as the root growing closest to the plant crown must be at the top of the pot when root cuttings are planted vertically because this is where the new growth will emerge. U R E 4. Cut cleaned root sections into pieces about 2" to 7" long. Trim off any excess, tiny fibrous roots. 5. Dust root cuttings with fungicide or ground cinnamon, a natural and effective alternative to chemicals. Thicker roots can be cut in shorter pieces while thinner, more delicate roots should be on the longer side. 6. The smaller, thinner roots may then be placed horizontally in the propagation bed, and covered with just ½” of growing medium. 7. Thicker roots should be planted vertically, making sure the proximal end of the cutting (end that was growing closest to the parent plant) with the flat top is stuck facing upward and flush with the growing medium in the pot. 8. The slanted cut on the distal end will then be pointing down into the pot. 9. Root cuttings can be left outdoors in their pots or flats in a protected area such as a cold frame or heeled into a plunge bed. Growth should commence in the spring. Preparing Sassafras albidum root cuttings; the cleaned root sections are being dusted with cinnamon, a natural and highly effective fungicide, before being treated with rooting hormone and then laid horizontally in the flat. Basic Plant Propagation G C A H O R T I C U L T Division Leaf Cuttings Whether you are dividing a pot-bound plant or a perennial in the ground, the purpose of division is to create more of the same plant and to improve the vigor of the original plant and all its progeny. Many tropical and succulent plants including Begonia, Sansevieria, and gesneriads such as African violets (Saintpaulia) can easily be propagated from a leaf or a piece of a leaf. There are several different types of leaf cuttings. Most benefit from some kind of bottom heat and some also need the increased humidity that a plastic tent or bag would provide. As with all propagation, research your specific species’ needs before beginning. Division is best undertaken before active growth commences. Don’t be afraid to use sharp implements to cut apart overgrown plants; they will recover and be better off for it. Leaf Without a Petiole This method is used for plants with thick, succulent leaves such as Sansevieria, that are propagated by cutting the long leaves into 3-4” pieces. Insert the cuttings vertically into the medium, making sure the original orientation is the same— basal or proximal end (the end closest to the crown of the plant or the roots) down and the part of leaf that was growing towards the tip, up. Echeveria, African violets and begonias can also be propagated by leaf cutting. Cut a leaf from a plant and remove the petiole (the small stalk that attaches the leaf to the stem). Insert the leaf vertically into the medium making sure that the mid-vein, the vein growing in the center of the leaf, is buried in the rooting medium. New plant(s) will form from this vein. When enough roots are formed the new plants can then be detached from the shriveled up old leaf and potted up. An Aloe brevifolia in need of division. Rooted pups are carefully teased away from the parent plant; any unrooted pieces of succulent plants should be left to heal for a day or two and then placed in a well drained mix to form new roots. Label all new plants with name and date of propagation. Dividing and repotting is also an excellent time to groom the parent plant. Basic Plant Propagation Leaf with Petiole Remove a leaf from the parent plant and include up to one and a half inches of the petiole (the small stalk that attaches the leaf to the stem). Insert the lower end of the petiole U R E Leaf cuttings of various succulent Crassulaceae, all of which are very easy to propagate from leaf and stem cuttings. When an adequate root system has formed, the new plantlets can be potted up on their own. Remove the old leaf at this point. into the medium. One or more new plants should form at the base of the petiole. The amount of time it takes for each species to form roots varies with some taking weeks and others months. The new plantlets can then be severed from the original leafpetiole cutting. The cutting may be left in the rooting medium and used again to produce more plants. Examples of plants that can be propagated by leaf-petiole cuttings include African violets, Peperomia, Episcia, Hoya, and Sedum. Split-vein Detach a leaf from a rex begonia and remove the petiole (the small stalk that attaches the leaf to the stem). Make cuts on several prominent veins on the underside of the leaf. Lay the cutting, lower side down, on the medium. If the leaf starts to curls up, hold the edges down with rooting medium. New plants will form at each cut. 9 G C Layering Layering is a method of vegetative propagation whereby a stem of a plant that is still attached to the plant is induced to produce adventitious roots. The layer is the newly formed, rooted plant after it is severed from the parent plant. This method is the easiest way to root difficult or slow to propagate species including Rhododendron, Magonolia, Hamamelis, and Cotinus as well as many other easier to propagate species. There are several different types of layering. The most commonly used types are: Simple or Ground Layering, Tip Layering, and Air Layering. A H O R T I C U L T up and either potted up or moved to its new home. If you prefer to be extra cautious, cut the new plant from its mother and allow it to remain undisturbed and on its own roots for a full season before moving to a new home. Tip Layering This is very similar to simple layering except the tip of the stem is buried. The buried stem will eventually turn upward and break ground, roots forming at the bend in the stem. This method is often used for raspberries and other members of the Rubus family. Air Layering U R hormone. There should be about 9” of growth about the wounded area. This method is used most often for woody plants such as Magonolia, Gardenia, Camellia, Citus, roses or figs. Whatever method is used, the wounded area must now be enclosed in damp sphagnum moss that has been soaked in water for several hours and squeezed out. 1. Form a handful of the moss and place it around the wounded stem. 2. Wrap with plastic wrap. Make sure no moss is extending beyond the plastic wrap. 3. If you have houseplant that has become too tall and top heavy you might try air Simple Layering layering it. This is an ancient method of propagation, believed developed by the 1. Select a young flexible branch near Chinese centuries ago. For best results, the ground in early spring or select at least a pencil-sized stem in summer. either the spring, with the mature wood 2. Make a cut on the underside of this from last season or late summer, with branch with a sharp knife or razor mature shoots from this season’s growth. 4. blade about 12” from the tip of the The area where the layering is to take stem. place should be at least 6” and no more than 12” from the tip. Plan on making 3. Scrape away a thin sliver of bark the cut just below a node on the stem 5. from the stem. Conversely, a cut may and remove all leaves and twigs 3 to 4” be made in the stem forming a thin on either side of the layer if possible. tongue held open with a small There are 2 different methods of pebble or toothpick. wounding the bark in air layering. 4. You can dust the cut with rooting hormone. 5. Loosen the soil where the wounded branch is to lay. 6. Anchor the branch with a wire pin to hold it in place making sure the exposed area of the stem is firmly in contact with soil. A rock or brick on top may further help hold it in place and mark the layering so it is not inadvertently disturbed. Inspect from time to time, making sure the area where the layering is taking place is getting adequate water. After a full season or two (depending on the species), the layer should be ready to be detached from the parent plant and dug 10 E Hold the plastic in place with electrical tape or whatever material you are using. Tape up the seam in the plastic. (It is very important that the medium remain moist but not wet and that the only water getting into the plastic wrapped moss is what you put in. ) If the layering is taking place in sunlight a loose tent of aluminum foil can be wrapped around the ball. Check periodically to make sure the moss is moist. If it needs water, very carefully open a small section of the top and dribble in some water. Tape back up. Make a slanting, diagonal, upward cut in the stem, about 1 ½” long going a third of the way into the stem. Insert a match or toothpick to hold the cut open and dust with rooting hormone. There should be at least 6” of growth above the cut. This method is used most often for tropical foliage plants or for plants with less woody stems such as Ficus or Dieffenbachia. If you are successful, roots will appear in the moss after several months or up to a year depending on the species. As soon as the rooting medium is full of roots cut the newly rooted stem from the mother plant just below the rooted layer. Remove the plastic and pot up the layer with the roots still encased in the sphagnum moss in a suitable soilless mix. The newly severed plant will probably suffer some “separation Make a 1” ring cut by scoring the stem anxiety” so be sure to place it in a shady with 2 parallel lines about 1” apart and spot and water adequately until it removing the bark in between. You establishes its independence! It is also a may also need to scrape the wounded good idea to put the newly rooted and area to insure the plant does not potted plant into a plastic bag for a few produce callus tissue that will form a weeks to keep humidity levels high and bridge over the wound and prevent promote additional root growth. roots from forming. Dust with rooting Basic Plant Propagation G C A H O R T I C U L T U R E off the scion from its rootstock. Both 2. Make a horizontal cut in the plants will continue to grow. This is under stock so that about 1/4 to one advantage of this method as both 1/3 of the previous diagonal cut is Many different grafting methods have plants can grow on. In other methods, leveled off. been devised, but all involve uniting a one of the plants will not. 3. On this portion, make a vertical piece of stem (the scion) with a When roots have formed (it may take cut in the under stock and hold compatible stock (the rooted plant on a year or more), cut the new plant free open with a knife or shim. Insert which it will grow). The stock generally from the parent plant and move it to the scion so that the cambium should be from a related plant (e.g., the its intended location. layers will meet. Dust with same genus). With any grafting, it is rooting hormone and wrap a crucial that the cambium, or growing Budding or Bud Graft grafting rubber tightly around the layers, of scion and stock are aligned. It Budding or Bud Grafting is a method stock to secure the scion in the often used for fruit trees, usually from under stock. Check to be sure is also desirable for the stock to be the end of June to mid-August, and cambium layers of the scion and larger or at least as large as the scion. there are many, many versions. under stock are aligned. Many trees and shrubs that are dwarf, Probably the method gardeners are weeping, or have unusual color most likely to use is "Shield Budding." 4. Cover graft with large jar or plastic bag supported by stakes so variegations are created by grafting. Select a side bud from the current that the scion does not touch the Cuttings taken from the branches that season's growth of the desired plant, jar, but humidity is retained for and, using a sharp knife, remove the exhibit a natural mutation (witches' optimum development. bud and a thin portion of bark and broom, a shoot with an unusual color cambium. This will be the scion. 5. Place in a shady spot or cover top variegation or weeping characteristic) Make an incision in the bark of the of jar with paper. When scion are grafted onto an under stock. recipient tree (stock) in a "T" shape. begins to grow, raise jar or plastic Grafting is also used on plants to Lift the bark carefully and insert the to let in some fresh air. Check improve the vigor of a weaker growing growth bud, then securely tie it in carefully for any signs of wilting. cultivar by grafting it onto a more place with raffia or grafting rubbers If wilting is seen, cover again to keep it from changing position for vigorous under stock or to produce a completely. Gradually acclimatize two or three weeks, after which it stronger growing plant in a shorter the graft to outside air by slowly should have been a successful graft. removing covering. amount of time. Grafting is used Grafting frequently on fruit trees, roses, tree Cleft Graft peonies, Hamamelis cultivars and other A Cleft Graft is most often used with difficult to grow plants. woody plants such as Camellia or fruit trees. In this case, an under stock with vigorous rooting tendencies is An Approach Graft consists of two selected. The under stock is cut off at rooted plants, one of which will be 6" to 10" from the ground with a grafted onto the other. Both plants are diagonal cut. The scion (selected from rooted, but a narrow strip of bark is the plant which is desired to be removed from each plant at a height perpetuated) is cut from a similar or where the two may be conveniently smaller piece of wood. brought together, wound to wound. 1. Prepare the scion, removing (The stems are sliced to expose the excess leaves so that only 2 or 3 cambiums in equal patterns.) remain. If leaves are very large, Using grafting rubbers, bind the two trim l/2 of each leaf. With a sharp plants together with their cambium knife, trim the bottom of the scion layers touching. When the one being in to "V" shape for insertion in the used as a scion begins to grow, check under stock. to see if the graft is healed. If so, cut Approach Graft Basic Plant Propagation A lesson from professionals: members of a The Lenox Garden Club learn how to graft an heirloom peach variety ‘Highlawn Peach’ to an under stock of Prunus persica ‘Red Haven’, part of the 2013 C e n t e n n i a l Tr e e P r o j e c t . A horticulture workshop lead by experts is a great way to learn challenging forms of vegetative propagation. Photography by MaryEllen J. O’Brien. 11 G C A H O R T I C U L T U R E Glossary Adventitious Roots: roots that form in parts of the plant (stems or leaves) during vegetative propagation. In a growing plant, cells will differentiate to become root cells, stem, leaf or flower cells. When a cutting is taken, new cells in the stem or leaf areas can become root cells, forming new roots. Rooting compounds are used to encourage the development of these new roots. Cambium: the green, or growing layer immediately under the bark, which carries water and nutrients to the entire plant. Dampening-off/Damp off: Fungal diseases that may attack seedlings. Distal End: Refers to the root area furthest from the crown of the parent plant (opposite of Proximal End). Double Dormancy: Multiple cold and warm stratification periods. In Situ: Literally “in place”; here refers to planting directly in the ground where the plant is intended to grow. Node: Area on stem where a leaf or leaves have been attached. Polarity: This is very important with vegetative propagation. Always mark and be aware of which part of the root, cutting or leaf cutting was growing closest to the crown of the plant or the roots; this will determine which end is up when you stick your cutting. Petiole: The small stalk attaching the leaf blade to the stem. Proximal End: Refers to the root area nearest the crown of the parent plant (opposite of Distal End). Radicle Root: The first part of a seedling to emerge from the seed during the process of germination. It is the initial root of the plant. Scarification: Breaking down of the impermeable outer layer of the seed coat to allow water to reach the seed. Scion: a hardwood tip cutting containing growing points (buds) from a plant with desirable characteristics which is grafted onto the under-stock. Stratification: The after-ripening of a seed—can be warm or cold, to mimic the seed’s natural environment and cause germination. Cold Stratification: After-ripening of the seed embryo brought on by a cold, moist period. Websites: The Lady Bird Johnson Wildflower Center at The University of Texas at Austin is an excellent resource for information on native plants and topics such as seed collecting. Plants may be searched on a master database or by region. Go to: www.wildflower.org/plants. The internet offers many other sites to learn about plant propagation, such as: www.theseedsite.co.uk, www.ces.ncsu.edu/depts/hort, www.hort.purdue.edu/ ext/garden_pubs.html#GeneralHort, www.hcs.ohiostate.edu/mg/manual/prop.htm#2. In addition, Master Gardeners all over the country have excellent online resources to help you learn more about propagating in your area of the country. GCA Online Links: GCA Seed Share, GCA Members Collect, Conserve, and Grow Sharing Seed: www.gcamerica.org/ membersonly/docs/seedexchange.htm The Real Dirt, GCA Horticulture Committee Online Publication: www.gcamerica.org/membersonly/docs/ hort-newsletter-archive2/index.htm 12 Growing Ferns from Spores Shirley Meneice, Carmel-by-the-Sea GC www.gcamerica.org/membersonly/docs/hortnewsletter-archive2/03-2005-Spring/page01a01.htm Taking the Plunge: Holding Beds that Work By Susan Deeks, GC of Morristown www.gcamerica.org/membersonly/docs/hortnewsletter-archive2/15-2010-Spring/page009.htm A Cold Frame for Winter Storage of Cuttings and Marginally Hardy Plants By Mercer O'Hara, GC of Englewood www.gcamerica.org/membersonly/docs/hortnewsletter-archive2/04-2005-Fall/page01b01.htm Camellia Grafting Barbara Tuffli, Past President of the American Camellia Society, Woodside-Atherton GC www.gcamerica.org/membersonly/docs/hortnewsletter-archive2/15-2010-Spring/page001.htm Make the Cut with This Easy Workshop Priscilla Twombly, Sasqua GC www.gcamerica.org/membersonly/docs/hortnewsletter-archive2/03-2005-Spring/page02b01.htm Air Layering Sandra Kouwenhoven, Amateur GC www.gcamerica.org/membersonly/docs/hortnewsletter-archive2/03-2005-Spring/page01c01.htm Basic Plant Propagation G C A H O R T I C U L T U R E Further Reading: Ashworth, Suzanne, Seed to Seed: Seed-Saving Techniques for Vegetable Gardeners, 2nd rev. ed. Bir, Richard. E., Growing and Propagating Showy Native Woody Plants. Chapel Hill (NC) and London (GB): University of North Carolina Press. ISBN 0807843660 Bubel, Nancy, The New Seed Starters Handbook. Emmaus (PA): Rodale Press, 1988. 1988. ISBN 9780878577521 Capon, Brian, Botany for Gardeners, An Introduction and Guide. Portland (OR): Timber Press, 1990. ISBN 0881921637 Cullina, William, Native Trees, Shrubs and Vines: A Guide to Using, Growing, and Propagating North American Woody Plants. Boston (MA): Houghton Mifflin Harcourt, 2002. ISBN 0618098585 Cullina, William, The New England Wild Flower Society Guide to Growing and Propagating Wildflowers of the United States and Canada. Boston (MA): Houghton Mifflin Co., 2000. ISBN 9780395966099 For advanced information on germination: Deno, Norman C., Seed Germination Theory and Practice, second edition, 5th printing, 1994. Plus First and Second Supplements to the Second Edition. Selfpublished and has been available through author at 139 Lenor Dr., State College, PA 16801. See also the article about Deno: www.rps.psu.edu/mar96/seed.html Dirr, Michael and Heuser, Charles, Jr. The Reference Manual of Woody Plant Propagation: From Seed to Tissue Culture: A Practical Working Guide to the Propagation of Over 1100 Species. Cary (NC): Varsity Press, 2006, second edition. ISBN 97809423775008. Also second edition: Portland (OR): Timber Press, 2009. ISBN 1604690046 Druse, Ken, Making More Plants, The Science, Art, and Joy of Propagation. New York (NY): Clarkson Potter, 2000. ISBN 051770787X Emery, Dara, Seed Propagation of Native California Plants, 3rd printing. Santa Barbara (CA): Santa Barbara Botanic Garden, 1995. ISBN 0916436039 Gardiner, Jim, The Royal Horticultural Society, A Wisley Handbook, Propagation from Seed. London (GB): Cassell, 1997. ISBN 0304320625 Hartmann, H. T., Kester, D. E., Davies, F. T., Geneve, R. L., Hartmann and Kester's Plant Propagation: Principles and Practices, Upper Saddle River (NJ): Pearson / Prentice Hall, 2010. ISBN 0135014492 Basic Plant Propagation 13 G C A H O R T I C U L T U R E Further Reading (continued): Koch, Henry, with Aird, Paul; Amrose John; and Waldron, Gerald, Growing Trees from Seed. A Practical Guide to Growing Native Trees, Vines and Shrubs. Ontario (Canada): Firefly Books, 2008. ISBN 9781554073634 Loewer, Peter, SEEDS: The Definitive Guide to Growing, History & Lore. Portland (OR): Timber Press, 2005. ISBN 9780881926828 McClure, Susan, Smith & Hawken: Hands On Gardener: Seeds and Propagation. New York (NY): Workman Publishing Co., 1997. ISBN 0761107339 McVicar, Jekka, Seeds: The Ultimate Guide to Growing Successfully from Seed. Guilford (CT): Globe Pequot, 2003. ISBN 9781585748747 Midgley, Jan A. W., Native Plant Propagation. Self-published, 2006: 234 Oak Tree Trail, Wilsonville (AL) 35186. Telephone: (205) 669-4097; email: [email protected] Schmidt, Marjorie G., Growing California Native Plants. Berkeley (CA): University of California Press,1980. ISBN 0520037626. Toogood, Alan, Editor-in-chief (United States editor Ray Rogers), American Horticulture Society Plant Propagation, The Fully Illustrated Plant-by-Plant Manual of Practical Techniques. New York (NY): Dorling Kindersley Publishing, Inc., 1999. ISBN 0789441160 For out of print or used books, try: www.abebooks.com or www.amazon.com 14 Basic Plant Propagation G C A H O R T I C U L T U R E Basic Plant Propagation Basic Plant Propagation Copies of the Basic Plant Propagation Handbook are available for $3.00 each (includes postage) or $2.50 for orders of 25 or more. Ordering information is on this GCA website, under Shop, or mail your request with your check, name and address to: The Garden Club of America 14 East 60th Street New York, New York 10022 Please include your name and shipping address. This handbook was written by the 2011-2012 GCA Horticulture Committee Edited by Dedee O’Neil, Akron Garden Club & Carrie Waterman, Noanett Garden Club Drawings by Martha Kemp, Piedmont Garden Club Photographs by Carrie Waterman, Noanett Garden Club (Unless otherwise attributed) GCA HORTICULTURE COMMITTEE
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