DEVELOPMENT * occurs after both the male and female gametes (sperm and eggs) fuse-FERTILIZATION * the process by which the fertilized egg, the ZYGOTE develops from a single cell into an adult multicellular organism Development I. ANIMAL DEVELOPMENT A) Embryonic Development * the changes a zygote undergoes as it changes into an embryo 1) cleavage-* a series of mitotic cell divisions that increase the number of cells but not the size of the cells * actually, the cell size becomes smaller * leads to the formation of a hollow ball called the BLASTULA 2) gastrulation-* in certain animals, the blastula becomes indented, forming a GASTRULA * the gastrula has 3 layers, sometimes called germ layers: * ENDODERM-the inner layer * ECTODERM-the outer layer * MESODERM-the layer in between the endoderm and the ectoderm 3) differentiation-* the different germ layers in the gastrula develop into special cells, tissues, and organs of multicellular animals ECTODERM: nervous system and the outer skin MESODERM: muscles, skeleton, circulatory and excretory systems, gonads ENDODERM: linings of the digestive and respiratory systems, as well as parts of the liver and pancreas a great look at the early stages of embryonic development awesome site from Univeristy of Utah's Genetic Science Center on stem cells; look at their main page too! Tim and Moby discuss stem cells. * these cells are called embryonic stem cells and can become all types of cells * during differentiation, the stem cells become more specialized for their function * scientists are trying to revert mature, specialized cells back to the stem cell phase (dedifferentiation--see pic below) so they can help out individuals by turning them into a needed type of tissue! examples: 1. healing spinal cord injuries 2. growing new heart tissues 3. forming new muscles for trauma patients 4) growth-* an increase in size or cell number as differentiation continues * early development consists chiefly of the differentiation and growth of cells and tissues The above embryonic development can occur either outside (externally) or inside (internally) the mother. B) External Development * the development of the embryo outside the mother 1) in water Development * in fish and amphibians, the eggs are fertilized externally and then develop externally as well * the embryo's food source is the yolk in the egg * gas exchange for the embryo occurs by diffusion through the thin egg with the watery environment * water also acts as a shock absorber for protection 2) on land * after being fertilized internally, the eggs of birds, reptiles, and a few mammals (like the duck billed platypus) lay their eggs to develop on land ADAPTATIONS OF EGGS a) shell-outermost layer which provides protection and a favorable environment b) yolk-food source of egg; produced by uneven cytoplasmic division during oogenesis (meiosis) c) different membranesi) chorion- the outermost membrane which is under the shell; contains the other membranes ii) amnion- contains AMNIOTIC FLUID which cushions the embryo and prevents cells from sticking to the shell iii) yolk sac- surrounds the yolk; blood vessels penetrate the yolk sac to carry food to the developing embryo iv) allantois- functions as a respiratory surface (exchange of gases); also acts as a storage site for the nitrogenous waste produced, uric acid C) Internal Development * the development of an embryo inside the mother 1) placental mammals * mammal embryos develop within the UTERUS after internal fertilization * the PLACENTA is a special organ made of both embryonic and maternal (mother) tissues * the placenta is the place where the exchange of nutrients, wastes, and respiratory gases occurs by diffusion * an UMBILICAL CORD with blood vessels attaches the developing embryo to the placenta * the bloodstreams of the embryo and the mother never mix! 2) marsupial mammals * these mammals have eggs that are fertilized internally * they develop internally for a short period of time without a placenta * the embryo is then born VERY EARLY IN ITS DEVELOPMENT! * then, the born embryo continues its development externally in a pouch which contains the mammary glandssupplies the nutrients to the young " kangaroos, opossum, and koala bears (not shown) are examples of marsupials II. PLANT DEVELOPMENT * both meiosis (the production of sex cellsgametes) and fertilization (the fusion of gametes) occur in the flower, a plant structure specialized for reproduction Development A) The Flower * may contain both male (stamen) and female (pistil) reproductive organs... * some species might have only one or the other * flowers might have other accessory structures such as: petals- used to attract insects sepals- the leaf of the flower 1) the stamen-* composed of the anther and filament (which looks like a wire) * the diploid (2n) cells of the anther produce the monoploid (n) pollen grains, which is the male gamete * MEIOSIS OCCURS IN THE STAMEN 2) the pistil-* composed of the stigma, style, and the ovary a) stigma- catches pollen (sticky-stigma) is the upper part of the pistil pollination--so that is why they talk about the 'birds and the bees'...they mean plant sex! b) style- connects the stigma to the... c) ovary- monoploid (n) eggs are produced here in ovules; MEIOSIS OCCURS IN THIS PART OF THE plant sperm--I mean pollen! PISTIL B) Pollination * the transfer of pollen grains from the anther (part of the stamen) to the stigma (part of the pistil) * pollination may be accomplished by wind, insects, or birds that carry the pollen * colored petals and nectar can attract different organisms to help pollination occur 1) Self-Pollination * the transfer of pollen on the same plant * can occur in the same flower or from flower to flower of the same plant * uses the genetic information from that same plant but is still sexual reproduction! (two cells are involved) 2) Cross-Pollination * the transfer of pollen of one plant to another plant entirely * allows for the combination of characteristics from both "parent plants" * only one problem...the environment is usually very dry * pollen grains have thick walls to help prevent water loss (dehydration) of the nucleus (found inside the pollen grain) C) Fertilization and Development * after pollination, the pollen grain germinates (grows) on the stigma * as it grows, it forms a pollen tube which extends into the ovary of the pistil Development * the sperm now forms from the nucleus of the pollen and travels down the pollen tube 1) Fertilization in the Flower * when the sperm fertilizes the egg in the ovule (part of the ovary) a zygote is formed * the zygote develops by mitosis into an embryo * the embryo and now matured ovule is called a SEED a) seed coat- a tough protective outer layer that develops from the walls of all about seeds the ovule b) fruit- the entire ripened ovary 2) Development of Embryos * the seed has 3 different parts: a) hypocotyl- becomes roots and lower stem of adult plant b) epicotyl- becomes upper stem and leaves of adult plant c) cotyledon- stored food for germinating plant 3) Germination and Growth *seeds grow under suitable environmental conditions such as: * sufficient moisture * proper temperature * sufficient oxygen for aerobic respiration--NRG! * seeds will grow into mature plants * growth occurs at regions called MERISTEMS
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