BROAD - LEAVED FOREST PLANT COMMUNITIES IN LATVIA DACE SĀMĪTE History of hardwoods in Latvia • 8,000 – 7,800 years ago became Tilia cordata togeather with Alnus incana, A. glutinosa and Populus tremula •7,200 years ago at the begining of Atlantic peroiod Quercus robur, Fraxinus excelsior •6,000 years ago hardwoods became established and reached their maximum distribution, even Fagus syvatica and Carpinus betulus apear in our forests •1,500 years ago the forests covered ~80% of the area of the country and mixed broad-leaved-spruce stands predominate and until 13th century Latvian forest preserved its primeval status well History of hardwoods in Latvia •14-16th century – intensive forest clearings for cropgrowing first devasted the most fertile broad-leved forests •18th century – due to Duchy of Courland where manufacturing and trade rapidly developed, oakwoods was total destroyed •At the old crop-growing teretorys soil fertility foal, and hardwoods were replased by Betula pendula, Popula termula and Picea abies Due to previous forest management in naw days forest composition is more like boreal, than it can be in our climatic and vegetation zone Boreo-nemoral forest zone in Europe Due to – among boreal and nemoral zone, Latvian forests represent nordic coniferous, Europ broad-leaved and mixed stands European broad-leaved forests in Latvia Dominant tree species are Quercus robur 0.3%* Fraxsinus excelsior 0.6% Tilia cordata 0.06% Ulmus glabra 0.03% * - percentage from all forest stands in Latvia oca l lu no sa Va cc in os a M yr ti ll os Hy a lo co m ios a O xa l id os Ae a go M yr p od ti ll io os M sa o yr s ti ll pa os gn oos po a l yt ri c ho Dr sa yo pt er io Ca sa Sp ric ha os gn oos Dr ph a yo ra pt g m er ito io sa so -c ar i co Fi sa l ip en du Va los cc a in os a m M el yr . ti ll os M er a cu m el. ri a M li o yr s a ti ll m os el. a O tu xa r f. l id m os el. a tu rf. m el . Cl ad ino s Broad-leaved forest representation among forest types 60,0 50,0 30,0 ME E 40,0 Lime 10,0 ME Oak ME Ash M 20,0 E E ME E 0,0 Quercus robur – oak •Demand temperate light and worm; •Grow moustly in riverine forests on rich soils •Best represented in eastern part of Latvia (Lubana lake, Pededzes and Aiviekstes river) Quercus robur forest distribution in Latvia 0.1-0.4% 0.5-0.9% 1% and more 20 1 1 1 age 1 1 1 1 1 1 an d 19 18 17 16 15 14 13 1 12 1 11 10 91 20 10 00 0 e 00 90 80 70 60 50 or m -2 -1 -1 -1 -1 -1 -1 40 -1 30 -1 -1 -1 -9 0 0 0 0 0 0 1 600,0 81 -8 -7 -6 -5 -4 -3 -2 -1 0 200,0 71 61 51 41 31 21 11 1 area (ha) Stand age of Quercus robur in Latvia 1400,0 1200,0 1000,0 800,0 Private forest 400,0 State forest 0,0 Fraxinus excelsior - ash •Feel good in shadoved places •Grow on carbonatic and humus rich soils •Offen make stands togeather with oak •Best represented in central and north-eastern part of Latvia Fraxinus excelsior forest distribution in Latvia 0.1-0.4% 0.5-0.9% 1-3.9% 4% and more age 20 1 an d 19 1 18 1 17 1 16 1 m or e -2 00 -1 90 -1 80 -1 70 -1 60 -1 50 -1 30 -1 40 -1 20 -1 10 -1 00 -9 0 -8 0 -7 0 -6 0 -5 0 200,0 15 1 14 1 13 1 12 1 11 1 10 1 91 81 71 61 51 41 -4 0 -3 0 -2 1 -1 0 400,0 31 21 11 1 area (ha) Stand age of Fraxinus excelsior in Latvia 1800,0 1600,0 1400,0 1200,0 1000,0 800,0 600,0 Private forest State forest 0,0 Tilia cordata – small-leaved lime Ulmus glabra – wych elm lime •Lime and elm monodominant forest stands meake very rear (river valleys, lake and bog islands) •Mostly they make mixed stands togeather with oak, ash and spruce elm •Elm feels good at mor shadowed and humus rich places than lime Tilia cordata forest distribution in Latvia Liepāja Jelgava Bauska 0.01- 0.04% 0.05 – 0.09% 0.1% and more Ulmus glabra forest distribution in Latvia 0.01-0.04% 0.05-0.09% 0.1% and more 20 1 1 1 age 1 1 1 1 1 1 an d 19 18 17 16 15 14 13 1 12 1 11 10 20 10 00 0 0 m 00 90 80 70 60 50 or e -2 -1 -1 -1 -1 -1 -1 40 -1 30 -1 -1 -1 -9 -8 0 0 0 0 0 1 120,0 91 81 71 -7 -6 -5 -4 -3 -2 -1 0 40,0 61 51 41 31 21 11 1 area (ha) Stand age of Tilia cordata in Latvia 180,0 160,0 140,0 Private forest 100,0 80,0 60,0 State forest 20,0 0,0 Broad-leaved forest characteristics • Gap disturbance – main factor of stand development • Total change from vernal to summer vegetation • Forest communities are similar to ones in central Europe, the lack of some characteristic species confined to the Fagus sylvatica and Carpinus betula forests are due to geographical limitations Diagnostic species of European broad-leaved forests Tree species Ulmus glabra Acer platanoides Fraxinus excelsior Quercus robur Carpinus betulus Shrub species Euonymus europaea Lonicera xylosetum Padus avium Corylus avellana Brachypodium sylvaticum Mercurialis perennis Carex digitata Milium effusum Lathraea squamaria Phyteuma spicatum Actea spicata Polygonatum multiflorum Adoxa moschatellina Pulmonaria obscura Allium ursinum Ranunculus lanuginosus Asperula odorata Ranunculus cassubicus Corydalis solida Sanicula europaea Corydalis cava Festuca altisima Galeobdolon luteum Dentaria bulbifera Impatiens noli-tangere Stellaria holostea Circaea alpina Matteuccia struthiopteris Circaea lutetiana Carex remota Main biodiversity structures •Multi-layered and multi-aged tree stand •Abundance of cryptogams on stems and trunks •Dead wood in diferent satges of decay •Woodpecker signs and trees with holes •Rich shrub layer •Many gaps •Low admixture of conifers •Trees with trunk diameter more than 30cm Broad-leaved forest syntaxonomy in Latvia Cl. Alnetea glutinosae Br.-Bl. et Tx. 1943 O. Alnetalia glutinosae Tx. 1937 em. Oberd. 1953 All. Alnion Glutinosae (Malc. 1929) Meijer Drees 1936 em.Th. Müll. et Görs 1958 Ass. Carici elongatae – Alnetum Schwick. 1936 (N. Priedītis) Wetland forests with Fraxinus excelsior and Alnus glutinosa Broad-leaved forest syntaxonomy in Latvia Cl. Querco- fagetea Br.-Bl. et Vlieger em Klima 39 O. Fagetalia sylvaticae Pawl. ap. Pawl. et al. 28 All. Alno – Ulmion Br.-Bl. et Tx.ex Tchou 1984 em.Th. Müll. et Görs 1958 SubAll. Alnenion glutinosae – incanae Oberd. 1953 Padus avium – Fraxinus excelsior community Geum rivale – Fraxinus excelsior community Carex vaginata – Fraxinus excelsior community Crepis paludosa – Fraxinus excelsior community Mercurialis perennis - Fraxinus excelsior community (D. Mangale, D.Sāmīte) Broad-leaved forest syntaxonomy in Latvia All. Carpinion betuli Oberd. 1953 Ass. Tilio-Carpinetum Traczyk 1962 All. Carpinion Ass. Querco - Tilietum Laiviņš 1983 Hepatica nobilis – Quercus robur community Calamagrostis arundinacea - Quercus robur community Picea abies - Quercus robur community (D. Sāmīte) Broad-leaved forest syntaxonomy in Lithuania Vaccinium vitis-idaea – Quercus robur (P. Snarskis, A. Jansons) Scorzonera humilis – Quercus robur Lathyrus niger – Quercus robur Pteridium aquilinum – Quercus robur Polygonatum verticilatum Campanula trachelium – Quercus robur Galeobdolon luteum – Quercus robur Oxalis acetosella – Quercus robur Deschampsia cespitosa – Quercus robur Populus tremula – Quercus robur Geum rivale – Quercus robur Filipendula ulmaria – Quercus robur Calamagrostis canescens – Quercus robur Alnus glutinosa – Quercus robur The future of broad-leaved forest • • • • Forest management Wood cultivation on agriculture land Nature protection Game animal influence • PRESENT COMPOSITION AND PATTERNS OF CHANGE CAN BE DUE TO PREVIOUS MAJOR DISTURBANCES AND CHANGES IN LAND USE, WHICH HAPPENED MANY HUNDREDS, EVEN THOUSANDS OF YEARS AGO E. van der Maarel Forest management in broad-leaved forests 300 250 Oak Ash 246,2 242,1 area (ha) 200 150 130,1 100 50 82,5 45,3 35,2 35,0 25,2 16,8 15,7 0,6 0 State Private Final felling State Private Cleaning cutting State Private Sanitary felling Reforestation 60,0 51,2 50,0 area (ha) 40,0 Oak 30,0 20,0 Ash 11,2 Lime 10,0 0,2 1,9 1,3 0,0 Natural Artifical Natural Artifical Artifical
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