9/11/2015 Bio 122 Fall 2015: Lecture 3: Energy Acquisition (Chpt 5) I. Energy Acquisition A. Energy sources B. Obtaining Energy: Autotrophy 1. Chemosynthesis 2. Photosynthesis (C3 pathway) a. Light reaction i. Capture photon ii. Use energy from photon to split water iii. Use electron from splitting water to synthesize ATP & NADPH b. Calvin cycle i. Uptake of CO2 ii. Synthesis of 3-C compound (PGA) iii. Synthesis of 6-C compound (glucose) C. Photosynthesis constraints & solutions 1. Light 2. Temperature a. rubisco – catalyst for 2 competing reactions b. C4 photosynthesis (spatial separation of cycles) i. Light reaction the mesophyll – 4-C compound created ii. 4-C to bundle sheath, providing CO2 & 3-C compound for glucose synthesis c. C3 vs. C4 photosynthesis 3. Water a. CAM photosynthesis (temporal separation of cycles) 4. Nutrients D. Heterotrophy Figure 5.6 Absorption Spectra of Plant Photosynthetic Pigments Photosynthesis: Light Reactions (sunlight required) Chloroplast Light Reactions Dark Reactions 1 9/11/2015 Figure 5.7 Plant Responses to Variations in Light Levels (Part 1) C3: Photosynthesis in mesophyll cells C4: Photosynthesis in 1) mesophyll cells 2) Bundle sheath cells C4 Cycle Calvin Cycle 2 9/11/2015 Figure 5.17 Examples of Plants with the CAM Photosynthetic Pathway Problem 5.1: Some Like it Hot Comparison of C3 and C4 Pathways Simulation model: http://sites.sinauer.com/ecology3e/problem05.html Figure 5.16 Crassulacean Acid Metabolism (Part 1) Figure 5.16 Crassulacean Acid Metabolism (Part 2) Figure 5.15 C3, C4, and CAM Photosynthesis Compared 3
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