Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Sminaire UMMISCO- 29 Avril 2009 STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHOLD PHENOMENA IN STRUCTURED POPULATION DYNAMICS Hassan HBID [email protected] LMDP et UMI-UMMISCO (Cadi Ayyad University, Marrakech,Morocco) April 28, 2009 Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Index 1. Introduction 2. Size Structured Models 3. Age-Structured Models with Moving Boundary Conditions 4. A short review on state dependent delay deferential equations Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Introduction In this work we are interested in the role of state-dependent delayed differential equations in the modelling of structured population dynamics. Our aim is to put in evidence the onset of state-dependent delays in an important class of models: threshold models. These are structured population dynamics models in which the transition of individuals between different stages is due to some quantity reaching a prescribed level. An interesting presentation of this kind of equations can be found in (H.L. Smith and Y. Kuang, Periodic solutions of differential delay equations with threshold-type delays, Contemporary Math. Vol. 129 (1992) 153-176). Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Introduction Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Introduction Although the appearance of differential equations with state-dependent delays goes back to the 19th century, the study of them has recently been reactivated due to their frequent occurrence in a wide variety of situations. These equations can be found in the literature associated with a wealth of applications in different fields, such as ! Electrodynamics (R.D. Driver, 1969), ! Economics (J. Bélair and J.M. Mahaffy, 2001), ! Control (H. O. Walther, 2002) ! Population dynamics (O. Arino, H. H. and R. Bravo de la Parra, 1998) Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Introduction From a mathematical point of view, it is well-known that structured models of population dynamics which are formulated in terms of hyperbolic partial differential equations can be reduced, integrating along the characteristic lines, to a so-called renewal equation for the function of births. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Introduction This process, applied to the reduction of a structured model of a population divided into several stages leads frequently to a delayed equation with a time-dependent delay. This variable delay corresponds to the time spent by the individuals of one stage in passing to the next one. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Introduction If this transition depends on the density of the population, then the delay will be state-dependent. In many cases, as we will show in the examples below, this condition of transition is formulated like an equation which implicitly defines the delay and which has a meaning in the model: the individuals change their stage after a certain quantity reaches a predetermined threshold value. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Introduction In some situations, the magnitude which determines the change of stage depends on the state of a non-structured population x(t), and therefore the threshold condition is formulated as ! t k(x(σ)) dσ = W ∗ t−τ (t) where k : R+ −→ R+ is a known function. There are also other situations where the population is structured by some character in all the stages. In this case, threshold condition should be expressed in terms of a functional F : ! t F (l (·, σ)) dσ = W ∗ . t−τ (t) Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Size Structured Models In (J.A.J. Metz and O. Diekmann, The Dynamics of Physiologically Structured Populations. Lecture Notes in Biomath. 68 (Springer-Verlag, 1986)) the process of modeling the dynamics of populations structured by some physiological variable is described, along with a great amount of examples and applications. With the aim of simplifying, and since it is the context in which the following models are presented, we stick to the one-dimensional case. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Size Structured Models As is fully explained in (J.A.J. Metz and O. Diekmann, 1986) , the model for the dynamics of this population is obtained through making a balance of mass in [xb , xd ], giving rise to a transport P.D.E.: ∂n ∂ (x, t) + [g (x, t)n(x, t)] = Sources − Sinks (1) ∂t ∂x together with a boundary condition which takes into account the entrance of individuals accross the boundary xb : g (xb , t)n(xb , t) = Local arrival rate. (2) The function g (x, t) is the individual growth rate, so that the individual state is determined by the O.D.E.: x # (t) = g (x, t) Hassan HBID [email protected] ; x(0) = x0 . STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Size Structured Models It is well known that solving along the characteristic lines transforms the problem into a renewal equation (see Webb, 1985 and Iannelli, 1994). This equation involves a variable delay τ (x, t) which is the time spent by an individual to reach the state x at time t and, therefore, is determined by the equation ! t g (x(σ), σ) dσ = x xb + t−τ (x,t) where x(σ) is the characteristic line through (x, t). Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Size Structured Models Let us suppose that the population is organized in different stages, say juvenile (J) and adult (A) to simplify, and the transition of an individual from (J) to (A) is made when its size reaches a predetermined threshold value W ∗ > xb . Then the entrance to the adult stage will depend on n(W ∗ , t). In turn, calculating this value for t big enough along the corresponding characteristic line, we get an expression in terms of the density n(xb , t̃), for some t̃ < t. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Size Structured Models As a consequence, a variable delay τ (t) = t − t̃ defined by ! t xb + g (x(σ), σ) dσ = W ∗ t−τ (t) will appear in the model. This threshold condition, in those cases where the growth rate g depends on the population produces in the model a state-dependent delay. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Size Structured Models In Hbid, Sanchez, Bravo de La Parra (M3AS, 2006), we illustrate this idea in different situations. We present a classical model by Nisbet and Gurney that develops completely a formalism to deal with insect populations in which the duration of their different instars is variable and is determined by obtaining a predetermined weight. We also applies these ideas to the modeling of cell populations. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions As we have seen in the previous section, the delay is the time employed by each individual to reach the threshold value of a certain variable, which means in age-structured models that the age of passing from one stage to the next is time-dependent. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions To illustrate these ideas, let us consider a population divided into two stages, the juveniles and the adults, each one being structured by the age in the stage. Denote by l (a, t) (resp. L(a, t)) the density of juveniles (resp. adults) of age a at time t. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions We have the following equations: ∂l ∂l ∂t (a, t) + ∂a (a, t) = −µl (a)l (a, t) ∂L (a, t) + ∂L (a, t) = −µL (a)L(a, t) ∂t ∂a (3) where µl (a) (resp. µL (a)) is the age-dependent mortality rate of juveniles (resp. adults). We point out the fact that the variable a represents the age of juveniles in the function l (a, t) while it is the age of the adults in the function L(a, t). Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions The above two equations have to be supplemented by boundary conditions and initial values. For the juvenile we have l (0, t) = B(t) ; l (a, 0) = 0 where B(t) represents the time density of newborns and the initial condition expresses the fact that no juvenile is supposed to exist at the time t = 0 (or, no one is able to survive beyond a certain period of time). The moving boundary is caused by the expression of B(t) in each model. Let us begin with a simple example (see O. Arino and E. Sánchez, Delays induced in population dynamics, In Mathematical Modelling of Population Dynamics, Banach Center Publications, Vol.63 (2004) 9-46). Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions We define J(t) to be the biomass of juveniles at time t, that is ! aM J(t) := l (a, t) da 0 aM (0 < aM < +∞) being the maximum age of juveniles (that is, individuals which have not reached the threshold value past time aM , will die or never reach the next stage) and let us suppose that B(t) = kJ(t) , Hassan HBID [email protected] (k > 0). STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions The passage from the juvenile to the adult stage is described in terms of the weight function of the juvenile, namely we assume that ∂w ∂w K (a, t) + (a, t) = ∂a ∂t J(t) + C where w (a, t) is the quantity of food eaten until time t by an individual entering the juvenile stage a time units ago. K > 0 is the quantity of food entering the species habitat per unit of volume and per unit of time, which for simplicity is considered to be a constant. The constant C > 0 represents the food (converted into a number of individuals) taken per unit of volume by consumers other than juvenile stage. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions Limitation of food due to density is modeled in a possible way, assuming that the quantity of food available is shared in equal parts by all the individuals occupying the same space at time t. Assuming that w (0, t) = w (a, 0) = 0 (which corresponds to the absence of&juveniles at time t = 0), we arrive at t K dσ We now introduce the threshold w (a, t) = t−a J(σ)+C condition: juveniles turn adult when the food index reaches a prescribed value W ∗ > 0, that is at each time t, a∗ (t) is the unique solution to the equation ! t K dσ = W ∗ (4) J(σ) + C ∗ t−a (t) and so J(t) = ! a∗ (t) l (a, t) da. (5) 0 Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions Solving the equation in l from system (3) along the characteristic lines starting from a = 0, we obtain l (a, t) = B(t − a)e − Ra 0 µl (s) ds = kJ(t − a)e − Ra 0 µl (s) ds which, bearing in mind (5) and accepting for simplicity that µl = 0, provides the following equation for J(t): ! t J(σ) dσ. J(t) = k (6) t−a∗ (t) Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation Age-Structured Models with Moving Boundary Conditions Differentiating in (4) and (6) we reduce the problem to the following system of state-delayed differential equations ' ( J(t − a∗ (t)) + C ∗ # J (t) = k J(t) − J(t − a (t)) J(t) + C (a∗ )# (t) = J(t − a∗ (t)) − J(t) J(t) + C Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation In the sequel, we will consider the two following classes of differential equations with state dependent delay: 1) # x (t) = f (x(t − τ (t))) (7) τ # (t) = h(x(t), τ (t)) where f : R → R, h : R × R → R, and the delay is determined by a differential equation. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Convenient assumptions are made on f and h in order to have t − τ (t) increasing and the function τ (t) bounded, that is, τ (t) ∈ [τ , τ ], where τ is the small delay and τ is the maximum delay. 2) (8) x # (t) = g (x(t), x(t − r1 (xt )), ...., x(t − rn (xt ))) where g : Rn+1 → R, ri : C ([−M, 0]; R) → [0, M], for i = 1, ..., n, and the delays are explicit functions of the solution x(t). Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Equation (7) was discussed by : ! (O. Arino, M. L. Hbid and R. Bravo de la Parra. A Mathematical Model of Growth of Population of Fish in the Larval Stage: Density-Dependence Effects. Math. Biosci. 150: 1-20,1998.) ! to describe a renewal equation of a population of fish. (O. Arino, K. P. Hadeler, and M. L. Hbid. Existence of Periodic Solutions for Delay Differential Equation with State Dependent Delay. Journal of differential equations, 144: 263-301, 1998.) ! to show the existence of oscillating periodic solutions. Equation (8) is studied by Mallet-Paret et al. in (J. Mallet-Paret, R. D. Nussbaum et P. Paraskevopoulos. Periodic Solutions for Functional Differential Equations with Multiple State-Dependent Time Lags. Topological Methods in Nonlinear Analysis, Journal of the Juliusz Schauder Center, 3: Hassan)HBID STATE-DEPENDENT DELAYSperiodic ASSOCIATED TO THRESHO to [email protected] show the existence of oscillating 101-162, 1994. Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Equations (7) and (8) can be written in the following “general generic” formulation: ) ẋ (t) = F (xt ) , t ≥ 0 x0 = ϕ ∈ C (9) where F (ψ1 , ψ2 ) = (f (ψ1 (−ρ(ψ2 (0)))), h(ψ1 (0), ρ(ψ2 (0)))), Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation for all (ψ1 , ψ2 ) ∈ C ([−τ , 0]; R2 ), and ρ is the retraction ρ : R → [τ , τ ] defined by: if a ≥ τ τ ρ(a) = a if a ∈ [τ , τ ] τ if a ≤ τ . for equation (7) and F (ϕ) = g (ϕ(0), ϕ(−r1 (ϕ)), ..., ϕ(−rn (ϕ))), for equation (8). Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation The purpose of this section is to study the aspect of semigroup associated with the solutions of the following delay differential equation: # x (t) = f (x (t − τ (t))) τ # (t) = h (x (t) , τ (t)) (x0 , τ (0)) = (ϕ, τ ) ∈ C 0,1 ([−τ , 0]) × [τ , τ ] := C 0,1 × K (10) Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation where f and h are two functions defined respectively from R into R and from R × R into R, C 0,1 is the Banach space of Lipschitz continuous functions defined from [−M, 0] to R, τ and τ are two positive constants, and as usual xt denotes the history segment of the solution x(t) at time t, xt (θ) = x(t + θ) for all θ ∈ [−τ , 0]. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation The functions f and h satisfy the following assumptions: H1 : ”f and h are two functions locally Lipschitzians,” H2 : ” there exists constants α and β such that, for each x ∈ R |f (x)| ! α |x| + β, ” H3 : ” i: |h (x, τ )| ! 1; ii: h (x, τ ) ≤ 0 and h (x, τ ) ≥ 0 , for each (x, τ ) ∈ R × K .” Hassan HBID [email protected] (11) STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Assumptions H1 and H2 , which we will use throughout the paper without further repetition, are introduced for the next reasons: - H1 guaranteed the existence and the uniqueness of the solution of the equation (10) (O. Arino, K. P. Hadeler, and M. L. Hbid. Existence of Periodic Solutions for Delay Differential Equation with State Dependent Delay. Journal of differential equations, 144: 263-301, 1998.), - H2 serves to obtain an estimation of the solution of the equation (10), - H3 is imposed on h in view to obtain a frame of the delay τ (t) between the two values τ and τ (ii), and to insure that the function t (−→ t − τ (t) is increasing (i). Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Put T (t) (ϕ, τ ) = (xt , τ (t)) , for each (ϕ, τ ) ∈ C 0,1 × K , where (x(·), τ (·)) is the solution of (10) with initial data (ϕ, τ ). Remark : From the existence and the uniqueness of the solution of (10) one deduces that the family (T (t))t"0 constitutes an algebraic nonlinear semigroup on C 0,1 × K . However, the example given in the proposition 3.1 of (M. Louihi, M. L. Hbid and O. Arino. Semigroup Properties and the Crandall Liggett Approximation for a Class of Differential Equations with Satate Dependent-Delays. Journal of Differential Equations 181: 1-30, 2002. ) remains valid to prove that t (→ xt (ϕ0 , τ ) for all τ ∈ K is not strongly continuous from [0, ∞] into C 0,1 . Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Thus we are brought to consider the set E of strong continuity of T (t), it is to say E := + * (ϕ, τ ) ∈ C 0,1 × K : t (→ T (t) (ϕ, τ ) is cont. from R+ into C 0,1 × K (12) Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Theorem Suppose that H1 , H2 and H3 hold. Then, we have: 1. * + E = (ϕ, τ ) ∈ C 1 × K : ϕ# (0) = f (ϕ (−τ )) (13) 2. E is closed, invariable by T (t), for each t ≥ 0, and the restriction of family of operators (T (t))t"0 to the set E, noted again by (T (t))t"0 , is a strongly continuous semigroup. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Proposition Assume that H1 , H2 and H3 are satisfied. If moreover f is of class C 1 , then a) the infinitesimal generator of the semigroup T (t) is the operator Λ defined by: , D(Λ) = {(ϕ, τ ) ∈ C 2 × K : ϕ#− (0) = f (ϕ(−τ )) and ϕ##− (0) = L(ϕ, τ )} Λ(ϕ, τ ) = (ϕ# , h(ϕ(0), τ )). (14) b) the closure of D(Λ) in C 1 × R, is equal to the set E. Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation In this section we consider the next notations: ) · )1,γ is the norm, equivalent to ) · )1 in C 1 , defined by: )ϕ)1,γ = )ϕ)∞ + 1-ϕ# - , for each ϕ ∈ C 1 ∞ γ Vγ is the retraction on the ball of center 0 and radius 1, in relation to the norm ) · )1,γ , defined by: ) ϕ if )ϕ)1,γ ! 1 Vγ = ϕ if not, %ϕ% 1,γ Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Gk and Hk are the functions defined respectively on C 1 × K and R × K by: . .ϕ/ / Gk (ϕ, τ ) = f kVγ (−τ ) , for each ϕ ∈ C 1 and all τ ∈ K . k (15) / . x (16) Hk (x, y ) = h kv ( ), y , for each (x, y ) ∈ R × K . k Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Ak is the operator defined by: * + D(Ak ) = (ϕ, τ ) ∈ C 2 × K : ϕ#− (0) = Gk (ϕ, τ ) Ak (ϕ, τ ) = (ϕ# , H(ϕ(0), τ ). (17) E k designates the set * + E k = (ϕ, τ ) ∈ C 1 × K : ϕ#− (0) = Gk (ϕ, τ ) . (18) , Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Jλ,k designates the operator Jλ,k = (I − λAk )−1 . R is the function defined on C 1 × [−τ , 0] by 0 0 1 exp( θ ) & R (ψ, θ) = λ λ exp − λs ψ (θ) ds. θ Ultimately, one considers the set Bρ defined by Bρ = {(ϕ, τ ) ∈ E : )ϕ)1 ≤ ρ} (19) . Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Theorem Suppose that H1 , H2 and H3 hold. Then, for all T > 0, ρ > 0 and t ∈ [0, T ], one has Lim -Jnλ,k (ϕ, τ ) − T (t) (ϕ, τ )-1 = 0, for all (ϕ, τ ) ∈ Bρ nλ→t where Bρ is the set defined by (19), k = γ0 (ρ, T ) + defined by: γ1 = max {α(ρ + βt) exp (αt) + β, ρ} . Hassan HBID [email protected] γ1 (ρ,T ) , γ γ1 are STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation A short review on state dependent delay deferential equation Theorem Moreover: )T (t) (ϕ1 , τ1 ) − T (t) (ϕ2 , τ2 ))1(ω) ! exp (ω0 t) )(ϕ1 , τ1 ) − (ϕ2 , τ2 ))1(ω) for each (ϕ1 , τ1 ) , (ϕ2 , τ2 ) ∈ Bρ × K . With ω = ω0 = 2lipk (h) + ω. Hassan HBID [email protected] max{α1 ,α2 } 1−α3 and STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO Introduction Size Structured Models Age-Structured Models with Moving Boundary Conditions A short review on state dependent delay deferential equation References ! Arino, Hbid, Bravo de La Parra, 1998 ! Arino, Hadeler, Hbid, 1998 ! Louihi, Hbid, Arino, 2001 ! Louihi, Hbid, 2006 and 2007 ! Hbid, Sanchez, Bravo de La Parra, 2006 ! Hbid, Louihi, Sanchez (in press) Hassan HBID [email protected] STATE-DEPENDENT DELAYS ASSOCIATED TO THRESHO
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