Department of Wildlife Biology and Game Management

Woess, M., Grillmayer, R., and Voelk, F. H. Green bridges and Wildlife Corridors in Austria.
Report: 1-14. Vienna.
Keywords: 8AT/barrier/bear/Canis lupus/Cervus elaphus/connectivity/conservation/corridor/
corridors/dispersal/fragmentation/game/green bridges/greenbridges/home range/landscape/large
carnivores/lynx/Lynx lynx/Malme/management/migration/movement/passageways/railways/red
deer/road/roads/Ursus arctos/wildlife corridors/wildlife management/wolf/wolves
Abstract: Anthropogenic barriers (roads, canals, railways and belts of settlements) may affect
wildlife species with large home ranges and specific seasonal migration traditions in a negative
way. The augmenting constitution of such barriers in Austria and other industrial countries
restricts genetic interchange between various subpopulations at an increasing rate. Additionally,
landscapes of intense agricultural use (lacking structural elements of vegetation cover like bush
belts a.o.) reduce migration. Therefore, still existing migration corridors gain high importance. The
conservation of those traditional and important possibilities for game movement and dispersal
should constitute an explicit part of wildlife management and of a wildlife-considering landscape
management respectively. In Austria, first steps towards resource management of this kind have
been taken most recently with special regard to large predators (bear, lynx, wolf) and red deer:
Two specific projects form the prerequisites for conservation or improvement of still existing
movement corridors.
Department of Wildlife Biology and Game Management
University of Agricultural Sciences
Peter Jordanstrasse 76/9
1190 Vienna, Austria
Title:
Green bridges and Wildlife Corridors in Austria
Keywords:
* WOESS, GRILLMAYER and VOELK
migration, corridors, Austria, passageways, green bridges, roads, red deer, big
game, fragmentation, connectivity, large carnivores, ressource management
Author:
Dipl.-Ing. Mark Woess
Department of Wildlife Biology
and Game Management
University of Agricultural Sciences
Peter Jordanstraße 76
1190 Wien
[email protected]
+431-476544466
Co Authors:
Dipl.-Ing. Dr. Friedrich H. Voelk
Austrian Federal Forests
[email protected]
+431-711454558
Roland Grillmayer
Department for Surveying, Remote
Sensing and Land Information
University of Agricultural Sciences
[email protected]
+431-476545106
Green bridges and Wildlife Corridors in Austria
Introduction
Anthropogenic barriers (roads, canals, railways and belts of settlements) may affect wildlife species with large
home ranges and specific seasonal migration traditions in a negative way. The augmenting constitution of such
barriers in Austria and other industrial countries restricts genetic interchange between various subpopulations at
an increasing rate. Additionally, landscapes of intense agricultural use (lacking structural elements of vegetation
cover like bush belts a.o.) reduce migration. Therefore, still existing migration corridors gain high importance.
The conservation of those traditional and important possibilities for game movement and dispersal should
constitute an explicit part of wildlife management and of a wildlife-considering landscape management
respectively. In Austria, first steps towards resource management of this kind have been taken most recently with
special regard to large predators (bear, lynx, wolf) and red deer: Two specific projects form the prerequisites for
conservation or improvement of still existing movement corridors.
PROJECT “EFFICIENT GREEN BRIDGE INSERTION IN AUSTRIA”
Ordering Party: Austrian Ministry of economic affairs
Problem and Purpose
The Austrian network of motorways is almost complete and fenced off. Only a few kilometres of road
construction are necessary to close the last gaps in the network. Therefore the possibilities of interchange for
wildlife are heavily restricted. One purpose of this study was to supply and provide information for taking
aspects of sufficient permeability in future road construction processes into consideration. The permeability of
Austrians existing network of motorways was examined for some indicator species and recommendations for
measurements referring to reconstitution of a sufficient wildlife-permeability are made. (VOELK et al 2001)
Materials and Methods
The survey focused on forest-favouring game species, which are most sensitive regarding size of home range and
size of passageways. As indicator species we selected rare and mobile big game species (brown bear, lynx, wolf,
moose) and widespread ungulates (wide ranging red deer, additionally chamois and wild boar, subsidiary also
roe deer). Information from and cooperation with hunters and game managers were additional aspects.
Routes and location of important migration corridors in Austria and in the border regions of neighbouring
countries were reconstructed by means of the Austrian map (distribution of forests and settlements) and by
means of multiple observations of big game species in certain regions. (see map 1) We got a lot of information
from locally responsible hunters, the plausibility of which was controlled by several track observations in all four
seasons. The routes of bear, lynx, wolf and moose are well documented within Austria from 1970 to 1999.
(ZEDROSSER AND VOELK 1999, STEINER 1995, MRLIK 1995, AESCHT ET AL 1995)
Map 1: Large carnivores (bear, lynx, wolf), moose and motorways in Austria
The evaluation of migration corridors in Austria has shown that apart from motorways many anthropogenic
factors act as barriers and reduce migration possibilities for game: Agricultural areas without natural woodland,
belts of settlemens, embankment constructions (e.g. Danube), railway lines and others. (see map 2) The
reconstitution of permeability through all these barriers is possible at corresponding expense (greenbridges,
measurements at river banks, compound of biotop in agricultural landscapes) except for one: Wildlife corridors
are irreversibly lost when settlements grow together and become impassable “belts of settlements”.
The evaluation of all existing passageways with potential for wildlife use (overpasses, underpasses, tunnels)
within the network of existing motorways gave an impression of the actual permeability of this network for the
indicator species. A database of The Ministry for Economic Affairs containing all existing overpasses,
Map 2: Spatial distribution of forests and barriers in Lower Austria
underpasses and tunnels (preliminary built for human use) was the basis for our analyses. Passageways with a
width of less than 30 m were preselected. They stayed unconsidered except if they fitted well in the landscape
and were of suitable construction for wildlife species (e.g. near forests, hedges leading towards the passageway,
not completely asphalted). Another preselection was made with the aid of aerial immages and maps containing
forest distribution and distribution of settlements, rivers and infrastructure. For this purpose a special
standardised checklist was developed. Afterwards a terrestrial evaluation of wildlife relevant characteristics of
the remaining passagways (standardised checklists) was carried out. Track controlls at passageways in all four
seasons brought information on movement and seasonal mobility of big game species. Simultaneously, a
questionnaire was handed over to locally responsible hunters and conservationists, which brought additional
information about actual use of the existing passageways by game species. The passageways were split into 5
groups from A (of international importance; useful for all species) to E (only local importance; useful for less
demanding small furred game species).
Results
As a result of these surveys, a national database about wildlife relevant information referring to existing
passageways across the Austrian network of motorways and information about the location of important wildlife
corridors is available now. (see map 3) The evaluated crossing possibilities for wildlife represent the
“permeability ressource” of the Austrian network of motorways. Additionally, the database contains information
about possibilities to upvalue certain passageways and make them more suitable for wildlife.
Map 3: Motorways and crossing possibilities for large mammals in Austria
The registration of potential and important migration routes for genetic interchange is the basis to secure still
existing corridor-ressources and to reactivate corridors, which are actually interrupted by certain barriers.
The study contains recommendations concerning the number, location, distribution, dimension and
constructional design of crossing possibilities across the Austrian network of motorways. Regarding the
necessary minimum permeability, we differentiated between international, regional and local defragmentation.
The necessary permeability was evaluated for each motorway segment between two motorway junctions
seperately. Recommendations concerning the minimum permeability for wildlife are slightly different for the
existing network and for new and future motorway constructions. The stronger criteria apply to the latter.
Motorway segments with insufficient permeability for wildlife could be identified by comparing the desired
values for permeability with the existing permeability (existing passageways). For these segments,
recommendations for constitution of wildlife passageways (green bridges) could be made. They included details
on location, dimension, number and quality of wildlife passageways. (see map 4) To avoid misinvestments other
barriers such as belts of settlements, railway lines or rivers with insurmountable embankments were considered.
These are the defined standards for existing highways:
Passageways Type A (for international habitat defragmentation)
Wherever a wildlife corridor of international relevance crosses a motorway, a passageway for wildlife with a
minimum width of 80 m is needed. (Recommended width 80-100 m)
Passageways Type B (for regional and national habitat defragmentation)
To ensure a minimum permeability and a minimum habitat defragmentation for the sensitive indicator species
at least 5 passageways for wildlife along one highway segment with a minimum width of 30 m are
recommended. Their locations should be away from human settlements and the maximum distance between
neighbouring passageways of type A or B must not exceed 20 km. If the motorway segment is longer than 75
km the number of necessary passageways increases for every started 20 km by 1. (Recommended width 30-80
m)
Passageways Type C (for local habitat defragmentation)
For local habitat defragmentation away from far ranging wildlife corridors, 5 passageways with a minimum
width of 15 m along one motorway segment are necessary. The average distance between passageways of type
A + B + C should not exceed 10 km. (Recommended width 15-30 m)
For very short motorway segments (<5 km , <25km) exceptions are provided.
The developed theoretical values for minimum permeability served as terms of reference but were not strictly
exerted. Corresponding to the specific local means the desired values were handled flexibly. But it would be
risky to refuse any kind of standards. In that case, measurements of habitat defragmentation for wildlife would
be regarded rather a luxury investment than a necessity. During each motorway construction project, the
necessity and amount of measurements of compensation would have to be discussed seperately which would
bring further disadvantages and inefficiency.
Map 4: Need for green bridges in Austria
PROJECT “WILDLIFE CORRIDORS”
This project is an interdisciplinary research project at the University of Agricultural Sciences Vienna. Specialists
in the fields of Remote Sensing, Landscape Ecology and Wildlife Biology are working together in this project. It
started in 1999 and will be completed in November 2001. (GRILLMAYER et al 2000)
Purpose
Within a study area, selected in advance, the applicability of remote sensing techniques for the provision of
information about corridor structures and barriers with relevance for wildlife migration in intensively used
agrarian areas is investigated.
Materials and Methods
Within the connecting belt between the Alps and the Carpathian mountains a part of the main migration route
was selected as investigation area. The study area is located east of Vienna between the floodplains of the
Danube and the Leitha. The area is dominated by intensive agricultural use. Due to the long term of
anthropogenic utilization many kinds of restraints (settlements, road infrastructures, fenced forest habitats a.o.)
for the mobility of the chosen indicator species red deer and wild boar exist. Besides this variety of barriers the
test area also contains structural potentials for corridor spaces, which can be used as starting point for
measurments of improvement.
Aerial image analysis and terrestrial mapping were carried out simultaneously, the resulting database contains
information on landcape elements with barrier characteristics, such with corridor characteristics and information
on categories of landcover. To guarantee the comparability of aerial image analysis and terrestrial mapping, an
interpretation key for the aerial image analysis was developed, which contained the same structure classes as the
terrestrial assesment. The interpretation key was verified and improved after a probational mapping.
For collecting actual information about wildlife migration the following methods were used:
Trail surveys along defined lines, wildlife observation with infrared field glasses and questioning of locally
responsible hunters.
Results
One final result of this study is the preparation of a low cost method to evaluate wildlife species relevant
landscape elements and their characteristics. This method can be utilized in major parts of Austria except for
Alpine areas.
Using the collected data a resistance model was developed, which allows analysis about wildlife movements in
the investigation area. This resistance model enables risk analysis to show if and how future developments in this
region will influence or interrupt the existing migration routes of red deer and wild boar. It will help to prepare
demonstrative future szenarios regarding consequences for wildlife mobility due to regional planning strategies.
At last measurements and instruments of landscape planning and politics but also normative and financial
measurments shall be discussed which help to minimize the risk of the loss of mobility for red deer and wild
boar in this region.
This project shows that basic information about the resource “wildlife corridor” can be collected by use of
remote sensing so that expensive terrestrial mapping can be reduced to a necessary minimum. Especially for the
development of concepts and strategies for regional and landscape planning such basic information is of great
usefulness. Consequently aspects of wildlife management as the defragmentation of habitat fragments can be
considered in time. Planning strategies and concepts can be tested with regard to their fragmentation risk using
the resistance model.
CONCLUSIONS / SUMMARY
An interdisciplinary project in Austria, titled “Wildlife corridors”, examines the applicability of remote sensing
methods and terrestrial surveys to identify corridor structures at different landscape scales. The investigation area
is a part of the connecting belt between Alps and Carpathian mountains. With the collected data and information
from aerial / satellite images and terrestrial surveys a resistance model for the investigation area and the indicator
species red deer and wild boar is developed.
“Efficient green bridge insertion in Austria” is another, recently finished project in Austria which lead to a
concept of green bridges for the Austrian network of motorways. By evaluating existing passageways wider than
30 m and their suitability for wildlife and further by gathering information about the most important migration
routes for red deer, moose and large carnivores the permeability of the network of motorways was investigated.
Both projects reveal explicit measurements of “resource management”, which ensure genetic exchange on the
long term. They are a contribution to sustainable wildlife management in Austria and the border areas of
adjacent countries.
Landscapes and landscape structures that enable the exchange, dispersal and migration of big forest-favouring
game species are resources which densly populated Europe is running short of. Both projects provide
information that is needed for the management of this ressource. Besides the improvement and implementation
of these data the integration in instruments of regional planning and landsape planning are the most important
next steps. Only thereby the ressource wildlife corridor can be taken into consideration in planning processes and
strategies at different scales.
References (cited)
AESCHT, E., AUBRECHT, G., GUSENLEITNER, F. (eds), 1995: Einwanderer. Neue Tierarten erobern
Oesterreich. Stapfia 37 (zugleich Kataloge des OOE Landesmuseums. Neue Folge 84). Linz. 275 pp.
GRILLMAYER, R., SCHACHT, H., SCHNEIDER, W., VOELK, F., WOESS, M., 2000: Forschungsprojekt
“Wildoekologische Korridore” (Entwicklung von fernerkundungsgestuetzten Methoden zur Erfassung und
wildoekologischen Bewertung von Korridoren, insbesondere Gehoelzstrukturen in der Agrarlandschaft, als
Grundlage landschaftsplanerisch - naturschutzfachlicher Planungen). 1. Zwischenbericht. Universaet fuer
Bodenkultur Wien. 36 pp + 24 pp. Appendix.
MRLIK, V., 1995: Evaluierung der Elchpopulation in der Grenzregion von Oesterreich und Tschechien.
Forschungsinstitut WWF Oesterreich. Studie 22. Wien 39 pp.
STEINER, E., 1995: Die Rueckkehr des Elches (Alces alces L.) nach Oesterreich - Chronologie der Ereignisse.
Stapfia 37: 255-267.
VOELK, F., GLITZNER, I., WOESS, M., 2001: Kostenreduktion bei Gruenbruecken durch deren rationellen
Einsatz. Empfehlungen fuer bundesweite Mindeststandards auf Basis der als Wildwechsel angenommenen
Ueber- und Unterfuehrungen an Oesterreichs Autobahnen und Schnellstraßen. Erstellt im Auftrag des
Oesterreichischen Bundesministeriums fuer Verkehr, Innovation und Technologie. Institut fuer Wildbiologie und
Jagdwirtschaft der Universitaet fuer Bodenkultur Wien. 62 pp + Anhang. (Bestellung bei: BMVIT. Tel.: ++43-171100-5527).
ZEDROSSER, A., VOELK, F., 1999: Large carnivores (bear, wolf, lynx), moose and trunk roads in Austria. In:
INFRA ECO NETWORK EUROPE (1999): 5th IENE meeting Budapest, Hungary, 14 - 17 April 1999. Report
of the meeting. Presentations of the participants. Swedish National Road Administration. Borlaenge. 24 - 26.
Further References
BENNETT, A.F., 1991: Roads, roadsides and wildlife conservation - a review.
In: Saunders, D.A. und Hobbs R.J. (eds.), 1991: Nature conservation 2 - The Role of Corridors. Chipping
Norton, Australien. pp 99 - 118.
BENNET, A.F., 1999: Linkages in the Landscape. The role of Corridors and Connectivity in Wildlife
Conservation. IUCN Publications Services Unit, UK.
BUNCE, R.G.H. and HOWARD, D.C., 1990: Species dispersal in agricultural habitats. Belhaven Press.
FORMAN, R.T.T., 1995: Land Mosaics. The ecology of Landscapes and regions. Cambridge University Press.
FORSCHUNGSGESELLSCHAFT FUER DAS VERKEHRS- UND STRASSENWESEN (ed), 1997: Richtlinie
RVS 3.01: Strassenplanung, Umweltschutz, Wildschutz. 9 pp.
GLITZNER, I.,VOELK, F., 1999: Passageways through 1990 km fenced motorways in Austria: Assessment of
number, distribution and quality for big game species (indicator species: red deer). In: INFRA ECO NETWORK
EUROPE: 5th IENE meeting, Budapest, Hungary (14-17 April 1999). Report of the meeting. Presentations of the
participants. Swedish National Road Administration (SNRA). Borlaenge. Sweden. 20 - 23.
GRUBER, F., 1994: Die Veraenderung von Rotwild- und Gamswildverbreitung und der Abschusstendenzen von
1983 - 1993. Forstschutz aktuell Nr. 15 (September 1994), FBVA-Wien: 6 - 9.
HOLZGANG, O., SIEBER, U., HEYNEN, D., VON LERBER, F., KELLER, V., PFISTER, H.P., 2000:
Wildtiere und Verkehr - eine kommentierte Bibliographie. Schweizerische Vogelwarte, Sempach. 72 pp.
HUBER, T., 1995: Luchse in Oesterreich - zurueckgebracht und eingewandert. Stapfia 37 (zugleich Kataloge
des OOE Landesmuseums. Neue Folge 84). Linz. 269 - 275.
KACZENSKY, P., KNAUER, F., HUBER, T., JONOZOVIC, M., ADAMIC, M., 1997: The Ljubljana Postojna
highway - a deadly barrier in Slovenia? J. Wildl. Res. 1 (3): 263 - 267.
KNAUER, F. and KACZENSKY, P., 1999: A habitat connectivity model for brown bears in Slovenia. In:
Waters, Forests and Environment Protection Ministery, National Forests Administrations (ed.): 12th Intern.
Conf. On Bear Research and Management, Abstract-Book, 11; Poiana Brasov, Romania.
KNEITZ, G., OERTER K., 1997: Minimierung der Zerschneidungseffekte durch Strassenbauten am Beispiel
von Fliessgewaesserquerungen bzw. Brueckenoeffnungen. Forschung Strassenbau und Strassenverkehrstechnik
755. Bundesministerium fuer Verkehr. Bonn - Bad Godesberg. 292 Seiten.
MADER, H.J., 1990: Survival and Dispersal of animals in cultivated landscapes: Introduction. Biolog.
Conservation 54 (3); 167-173
MAIZERET, C., A. CAMBY, 1987: Les cerfs et l’autoroute - bilan des observations réalisées en CharenteMaritime dans le cadre du souivi écologique de l´A10. Office national de la chasse - Bulletin mensual 119: 2534.
MINISTRY OF TRANSPORT, PUBLIC WORKS AND WATER MANAGEMENT, 1997: Habitat
Fragmentation & Infrastructure. Proceedings of the international conference on habitat fragmentation,
infrastructure and the role of ecological engineering, 17-21 September 1995, Maastricht and The Hague, The
Netherlands. Delft. 474 pp.
MUELLER, S., BERTHOUD, G., 1995: Fauna/Traffic Safety. Manual for Civil Engineers. École polytechnique
fédérale de Lausanne. Département de génie civil Laboratoire des voies de circulation (LAVOC). 135 pp.
OLBRICH, P., 1984: Untersuchung der Eignung von Wilddurchlaessen und der Wirksamkeit von
Wildwarnreflektoren. Forschung Strassenbau und Strassenverkehrstechnik, Heft 426. Bundesminister fuer
Verkehr, Abteilung Strassenbau. Bonn - Bad Godesberg. 58 pp.
PFEIFER, M., ASTE, C., 1996: Zerschnittene Lebensraeume. Barrierewirkung von Autobahnen und
Schnellstrassen fuer Wildtiere, Leitart Rotwild. Politikum. Josef Krainer Haus Schriften 16 (70): 63 -68.
PFISTER, H.P., HEYNEN, D., KELLER, V., GEORGII, B., VON LERBER, F., 1999: Haeufigkeit und
Verhalten ausgewaehlter Wildsaeuger auf unterschiedlich breiten Wildtierbruecken (Gruenbruecken). Ergebnisse
von Beobachtungen mittels Infrarot-Video-Technik. Schweizerische Vogelwarte, Sempach. 48 pp.
PFISTER, H. P., KELLER, V., RECK, H., GEORGII, B., 1998: Bio-oekologische Wirksamkeit von
Gruenbruecken
ueber
Verkehrswege.
Forschung
Strassenbau
und
Strassenverkehrstechnik
756.
Bundesministerium fuer Verkehr. Bonn - Bad Godesberg. 590 pp. (Bestellungen: Typo-Druck Bonn: 0049-228650906; 258,- DM)
PROMBERGER, C., SCHROEDER, W., 1993: Wolves in Europe - status and perspectives. Proceedings of the
workshop “Wolves in Europe - current status and prospects”, April 1992, Oberammergau. Germany. 136 pp.
RAUER, G., GUTLEB, B., 1997: Der Braunbaer in Oesterreich. Umweltbundesamt Wien. Monographien, Band
88. Wien. 64 pp.
RIGHETTI, A., 1997: Passagen fuer Wildtiere. Die wildtierbiologische Sanierung des Autobahnnetzes in der
Schweiz. Pro Natura. Beitraege zum Naturschutz in der Schweiz Nr. 18. Basel. 46 pp. (Bezug: Pro Natura, CH4020 Basel, Postfach)
SAUNDERS, D.A. and HOBBS, R.J., 1991: Nature Conservation 2: The role of corridors. Surrey Beatty and
Sons, Chipping Norton, Australia.
SCHADT, S., 1997/98: Ein Habitat- und Ausbreitungsmodell fuer den Luchs. Diploma Thesis. Techn.
Universitaet Muenchen – Weihenstephan.
SCHUMAKER, N., 1996: Using landscape indices to predict habitat connectivity. Ecology 77; 1210-1225
SCHWEIZERISCHE GESELLSCHAFT FUER WILDTIERBIOLOGIE, 1999: Wildtierkorridore Schweiz Raeumlich eingeschraenkte, ueberregional wichtige Verbindungen fuer terrestrische Wildtiere im oekologischen
Vernetzungssystem der Schweiz. Schweizerische Vogelwarte, Sempach. 71 pp + Anhang.
SETRA (Service d’études Techniques des Routes et Autoroutes; ed.), 1993: Passages pour la grande faune.
Guide Technique. Bagneux Cedex. 121 pp.
UNIVERSUM (REDAKTIONSBERICHT), 1999: Autobahnen als Barriere fuer Baer, Luchs und Wolf. Wild
auf Wanderschaft. Universum. Wien. Heft 5: p. 77.
VOELK, F., GLITZNER, I., ZEILER, H., REISS-ENZ, V., 1998: Wildwechsel trotz gezaeunter Autobahnen.
Oesterreichs Weidwerk, Heft 1: 14 - 16.
VOELK, F., GLITZNER, I., 1999: Barrier effects on big game due to motorways in Austria. Assessment of 1990
km of fenced motorways. In: International Seminar “Ecological passes for wildlife and roadside afforestation as
necessary parts of modern road constructions (motorways and railway roads)”. Ed.: Jozef CURZYDLO,
Department of Ecological Bases of Environmental Engineering, University of Agriculture. Krakow. 107 - 134.
(ISBN 83-912184-1-4).
VOELK, F., KALIVODOVA, E., KUERTHY, A., GLITZNER, I., KALIVODA, H., WOESS, M., 2000:
Wildtier-Korridor Alpen-Karpaten - slowakischer Teilbereich: Staatsgrenze Oesterreich bis oestlich der
Autobahn E 65. Schlussbericht des Projektes Nr. 29s17 der Aktion Oesterreich-Slowakei. Institut fuer
Wildbiologie und Jagdwirtschaft der Universitaet fuer Bodenkultur Wien und Institut fuer Landschaftsplanung
an der Slowakischen Akademie der Wissenschaften (SAW) in Bratislava. 42 pp + 18 pp Annex.
WARD, A., 1982: Mule deer behavior in reaction to fencing and underpasses on Interstate 80 in Wyoming.
Transport Research Record 859: 8 - 13.
WARD, A.L., CUPAL, J.J., GOODWIN, G.A., MORRIS, H.D., 1976: Effects of highway construction and use
on big game populations. - U.S. Dep. Transportation, Fed. highway Adm. Off. Res. Dev. Rep. FHWA-RD-76174. 98 pp.
WOELFEL, H., KRUEGER, H.H., 1995: Gestaltungsmoeglichkeiten von Wilddurchlaessen an Autobahnen.
Zeitschrift fuer Jagdwissenschaft 41: 209 - 216.
YAHNER, T.G., KOROSTOFF, N., JOHNSON, T.P., BATTAGLIA, A.M. and JONES, D.R., 1995: Cultural
landscapes and landscape ecology in contemporary greenway planning, design and management: A case study.
Landscape Urban Plann. 33; 295-316.
ZEDROSSER, A., 2000: Baer, Wolf, Luchs, Elch und das Autobahn- und Schnellstrassennetz in Oesterreich.
Vertiefungsprojekt fuer Landschaftsplaner ueber Biotopmanagement und Wildtieroekologie (Sommersemester
1999 und Wintersemester 1999/2000). Schlussbericht zu Handen des Instituts fuer Wildbiologie und
Jagdwirtschaft der Universitaet fuer Bodenkultur Wien. 15 pp + Kartensatz.