Geosphere rock & soil spheres evolution biomes Pedosphere Pedosphere: soil properties • Millions of microorganisms • Minerals/Nutrients Porosity Permeability Color Grain size Soil Texture pH - Acid, < 7.0 - Alkaline, > 7.0 – oxygen (O), silicon (Si), aluminum (Al), iron (Fe), calcium (Ca), sodium (Na), potassium (K), magnesium (Mg) • Decaying organic matter (humus) • Void space – water • Void space – air Soil horizons O - surface litter layer • Fresh at surface, decomposed at depth • Humus provides N, K, binds particles, enhances soil moisture retention Pedosphere: soil profiles A - topsoil layer • Beginning of true soil • Dark in color • Mixture of organic / inorganic products • Most productive layer E - Eluviation layer • Light colored • Active leaching – soluble minerals and nutrients carried downward B – Subsoil, Illuviation • Dense • Lower in organic matter than A • Zone of accumulation C - Substratum 1 Atmosphere Universe Biosphere Ecology Galaxies Biosphere Solar systems Planets Earth Ecosystems Communities Organ systems Organs Tissues Populations Cells Protoplasm Molecules Organisms species Atoms Subatomic Particles Ecosystem components Ecosystem dynamics Food chain and trophic levels Oxygen (O2 ) Sun Producer Carbon dioxide (CO2) Secondary consumer Primary (fox) consumer (rabbit) First Trophic Level Second Trophic Level Third Trophic Level Producers (plants) Primary consumers (herbivores) Secondary consumers (carnivores) Heat Heat Fourth Trophic Level Tertiary consumers (top carnivores) Heat Solar energy Producers Heat Heat Soil decomposers Heat Heat Water Heat Detritivores (decomposers and detritus feeders) Ecosystem components Autotrophs (producers) Ecosystem components Quantifying productivity Sun • Light Reactions Gross primary productivity (gC/m2/yr) - Chlorophyll molecules absorb photons - Excited electrons make ATP - Photons split H2O to make H+ and O2 - H+ used to make NADPH - O2 released • Dark Reactions - rate at which producers turn solar energy to biomass Chlorophyll H2O - CO2, NADPH, ATP used to make glucose Light-dependent Reaction Chloroplast in leaf cell O2 Net primary productivity - rate at which producers turn solar energy to biomass minus respiration - energy available to consumers Energy storage and release CO 2 Lightindependent reaction Glucose 2 Ecosystem components Quantifying productivity Ecosystem interactions Quantifying productivity Terrestrial Ecosystems Swamps and marshes Tropical rain forest Temperate forest North. coniferous forest Savanna Agricultural land Woodland and shrubland Temperate grassland Tundra (arctic and alpine) Desert scrub Extreme desert Aquatic Ecosystems Estuaries Lakes and streams Continental shelf Open ocean Average net primary productivity (kcal/m2 /yr) Source: NASA Ecosystem components Heterotrophs (consumers) Ecosystem components Heterotrophs (consumers) • Herbivores • Carnivores Ecosystem components Detrivores (specialized consumers) Ecosystem dynamics Biodiversity What does biodiversity do for us? How do we as a species affect biodiversity? How do we protect biodiversity? 3 Nutrient Cycles Nutrient Cycles: Water Condensation Rain clouds Transpiration Precipitation to land Transpiration from plants Precipitation Runoff Surface runoff (rapid) Evaporation Precipitation Evaporation from land Evaporation from ocean Precipitation to ocean Surface runoff (rapid) Infiltration and Percolation Groundwater movement (slow) Ocean storage Nutrient Cycles: Carbon Nutrient Cycles: Nitrogen Nutrient Cycles: Phosphorus Nutrient Cycles: Sulfur 4 Evolution 4.5 4 3 2 1 0 4.5 4 3 2 4.5 4 today Billions of years ago • Formation of Earth’s crust, atmosphere and oceans Evolution Evolution 3 2 1 1 0 Evolution 4.5 4 today Billions of years ago 3 2 1 • Single-cell prokaryotes form in the sea -Bacteria -Archaea • Single-cell prokaryotes form in the sea • Single-cell eukaryotes form in the seas -Amoebae -Ciliates -Flagellates -Heliozoa Cyanobacteria 2 Amoeba Credit: Micrographia 4 3 2 1 Billions of years ago 0 0 today Billions of years ago • Formation of Earth’s crust, atmosphere and oceans • Small organic molecules form in the sea • Large organic molecule form in the sea • First protocells form in the sea 4.5 today Billions of years ago • Formation of Earth’s crust, atmosphere and oceans • Small organic molecules form in the sea • Large organic molecule form in the sea • First protocells form in the sea • Formation of Earth’s crust, atmosphere and oceans • Small organic molecules form in the sea • Large organic molecule form in the sea • First protocells form in the sea Evolution 0 Evolution today 4.5 4 3 2 Credit: Micrographia 1 Billions of years ago • Formation of Earth’s crust, atmosphere and oceans • Small organic molecules form in the sea • Large organic molecule form in the sea • First protocells form in the sea • Formation of Earth’s crust, atmosphere and oceans • Small organic molecules form in the sea • Large organic molecule form in the sea • First protocells form in the sea • Single-cell prokaryotes form in the sea • Single-cell eukaryotes form in the seas • Multicellular organisms form in the seas, later on land • Single-cell prokaryotes form in the sea • Single-cell eukaryotes form in the seas • Multicellular organisms form in the seas, later on land Red algae Credit: Berkeley, Palomar College Red algae Humans 0 today Credit: Berkeley, Palomar College 5 Evolution: 4.5 4 Homonids 3 2 1 Billions of years ago 0 Phylogeny: study of evolutionary relatedness today Source: Reed et al., PLOS Biology how? Evolution: Natural selection • genetic variability • traits must be heritable • trait must lead to differential reproduction Structural adaptations Fruit and seed eaters Insect and nectar eaters Greater Koa-finch Kuai Akialaoa Amakihi Kona Grosbeak Akiapolaau Credit: NASA Evolution: Natural selection • • • • how? genetic variability traits must be heritable trait must lead to differential reproduction leads to adaptations: heritable traits that enable survival and reproduction - Structural, behavioral/sexual, physiological Evolution: Genetic variation how? • mutations: random changes in the structure or # of DNA molecules in a cell that can be inherited by offspring Crested Honeycreeper Maui Parrotbill Apapane Unknown finch ancestor 6 how? Evolution: Genetic variation • mutations: random changes in the structure or # of DNA molecules in a cell that can be inherited by offspring • gene flow: introduced genes from a different population (same species) how? Evolution: Genetic drift • random selection • allows maladaptive traits in small populations - probability Worms of different color over 4 generations: Evolution: Genetic variation how? • mutations: random changes in the structure or # of DNA molecules in a cell that can be inherited by offspring • gene flow: introduced genes from a different population (same species) • hybridization: 2 distinct taxa crossbreed to produce fertile offspring Evolution • Large gene pool - Increases biological fitness of the species • Small gene pool - Fewer traits for natural selection - Genetic drift has significant affects - Increased potential of extinction - chance Hunted to near extinction, 30,000 to 1,000 in 1900 Evolution Behavioral Physiological Structural Speciation through adaptations Genetic drift Speciation through chance/probability Coevolution Interacting species engage in a back and forth genetic contest Convergent evolution Organisms not closely related, independently evolve similar traits due to similar ecological niches Biomes Speciation due to human selection 7 Biomes Biomes Biomes Average annual precipitation 100–125 cm (40–50 in.) 75–100 cm (30–40 in.) 50–75 cm (20–30 in.) 25–50 cm (10–20 in.) below 25 cm (0–10 in.) 4,600 m (15,000 ft.) 3,000 m (10,000 ft.) 1,500 m (5,000 ft.) Coastal mountain ranges Sierra Nevada Mountains Great American Desert Coastal chaparral Coniferous and scrub forest Biomes Rocky Mountains Desert Great Plains Coniferous forest Mississippi River Valley Prairie grassland Appalachian Mountains Deciduous forest Biomes: # mammal species World Wildlife Fund 142 terrestrial ecoregions "geographically distinct assemblage of species, natural communities, and environmental conditions" Olson, BioScience, 2001 8 Biomes: # endemic mammal species Biomes World Wildlife Fund 142 terrestrial ecoregions Olson, BioScience, 2001 Biomes: # mammal species 9
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