CURRICULUM VITAE KAREN KOVAKA University of Pennsylvania Department of Philosophy Cohen Hall 433 Philadelphia, PA, 19104 phone: 518.651.0298 web: www.karenkovaka.com email: [email protected] AREAS OF SPECIALIZATION philosophy of science, philosophy of biology, environmental ethics AREAS OF COMPETENCE Social and political philosophy, epistemology EDUCATION_________________________________________________________________ University of Pennsylvania PhD (2012-[expected] 2017), Department of Philosophy Dissertation: Disagreement and Developmental Plasticity Committee: Michael Weisberg (Supervisor), Quayshawn Spencer, Alexander Guerrero, Timothy Linksvayer (Biology) The Australian National University Visiting Scholar (August 2016) Boston College BS (2008-2012), Environmental Geoscience, Philosophy PUBLICATIONS______________________________________________________________ Kovaka, K. “Evidence and underdetermination in the developmental plasticity debate.” (conditional acceptance, British Journal for the Philosophy of Science). Kovaka, K. “Different research programs need different individuality concepts.” (invited review, Studies in the History and Philosophy of Science Part C). Warner, M., Kovaka, K. & Linksvayer, T. 2016. “Late-instar ant worker larvae play a prominent role in colony-level caste regulation.” Insectes Sociaux. DOI: 10.1007/s00040-0160501-3. Kovaka, K., Santana, C., Patel, R., Akcay, E. & Weisberg, M. 2016. “Agriculture increases individual fitness.” Brain and Behavioral Sciences 39. Kovaka, K. 2015 “Biological individuality and scientific practice.” Philosophy of Science 82(5):1092-1103. Kovaka CV 1/7 MANUSCRIPTS_______________________________________________________________ Kovaka, K. “Replicators, inducers, and evolutionary novelty.” (revise and resubmit, Biology and Philosophy). Singer, D., Bramson, A., Grim, P., Holman, B., Jung, J., Kovaka, K., Ranginani, A., Berger, W. “Rational social and political polarization.” (under review) Kovaka, K. “Why don't Americans accept climate change?” (in preparation) Kovaka, K. “A defense of relative significance controversies.” (in preparation) Kovaka, K. “Underdetermination in real life.” (in preparation) Kovaka, K. “Characteristics of successful environmental interventions.” (in preparation) FELLOWSHIPS, AWARDS, AND HONORS_______________________________________ 2016 Dean's Scholar (University of Pennsylvania) Academic award granted to 9 graduate students each year 2016 Teaching Certificate (Penn Center for Teaching and Learning) Earned by only 5% of Penn PhD students 2016 President Gutmann Leadership Award (University of Pennsylvania) Academic award granted to 12-15 graduate students each year 2012-2015 Lilly Graduate Fellowship (Lilly Fellows Program, Valparaiso University) National award for teaching-focused graduate students in the humanities. Awarded to 16 students each year. 2013-2016 George W.M. Bacon Fellowship (University of Pennsylvania) Full support for graduate research from the Philosophy Department 2012-2013 Benjamin Franklin Fellowship (University of Pennsylvania) Full support for graduate research from the Philosophy Department 2008-2012 Presidential Scholar (Boston College) Full tuition scholarship granted to 15 students each year Kovaka CV 2/7 CONFERENCE PRESENTATIONS_____________________________________________ What is the signature of plasticity? • PSA 2016 (Philosophy of Science Association Biennial Meeting), November 2016 Interacting inheritance channels • Philosophy of Biology Dolphin Beach, August 2016 Underdetermination in the developmental plasticity debate • Philosophy of Biology at Madison, May 2016 Why don't Americans accept climate change? • Philosophy and Education Workshop, University of Pennsylvania, October 2015 Replicators, inducers, and evolutionary novelty • IHSPSSB 2015 (International Society for the History, Philosophy, and Social Studies of Biology, Montreal), July 2015 • BSPS 2015 (British Society for the Philosophy of Science, Manchester) July 2015 Biological individuality and scientific practice • PSA 2014 (Philosophy of Science Association Biennial Meeting), November 2014 Values, evolution, and predictive ecology • Institute for for Philosophy and Public Policy, George Mason University, May 2014 Superorganisms and biological individuality • ISHPSSB 2013 (International Society for the History, Philosophy, and Social Studies of Biology, Montpelier), July 2013 DEPARTMENTAL PRESENTATIONS___________________________________________ Replicators, inducers, and evolutionary novelty • Penn Philosophy of Science Workshop, May 2015 Values, evolution, and predictive ecology • Penn Philosophy of Science Workshop, May 2014 What counts as niche construction? • Penn Philosophy of Science Workshop, November 2013 In defense of superorganisms • Penn Philosophy of Science Workshop, May 2013 Kovaka CV 3/7 TEACHING__________________________________________________________________ As Sole Instructor Fall 2016 Fall 2016 Environmental Ethics, University of Pennsylvania (introductory undergraduate level) Environmental Ethics, St. Joseph’s University (introductory undergraduate level) As Teaching Assistant (grading and teaching sections): Traditional Courses Spring 2015 Fall 2014 Spring 2014 Fall 2013 Philosophy of Social Science (instructor: Dr. Jan Willem Lindemans) Biomedical Ethics (instructor: Dr. Andrew Mcaninch) The Social Contract (instructor: Prof. Kok-Chor Tan) Knowledge and Reality (instructor: Prof. Daniel Singer) Online Courses Summer 2016 Fall 2014 Introduction to Philosophy (instructor: Dr. Gary Purpura) Revolutionary Ideas (instructor: Dr. Alex Guerrero) Guest Lectures Fall 2015 Fall 2015 Fall 2015 Fall 2014 Fall 2014 Spring 2014 “Experiments, simulations, and observations.” (Philosophy of science) “Species concepts” (Philosophy of biology) “Speciation” (Philosophy of biology)( “The enlightenment case against God.” (Introduction to Philosophy) “A solution to the problem of evil?” (Introduction to Philosophy) “The Communist Manifesto” (The Social Contract) SERVICE_____________________________________________________________________ Referee for: 2016-present 2015-present 2014-2015 April 2014 2013-2014 2013-2015 April 2013 Philosophy of Science, British Journal for the Philosophy of Science, Philosophy of Biology Graduate Representative to the faculty, Penn Philosophy Department Co-leader, Philosophy Club at Philadelphia Futures (college readiness and success program for underserved students) Research Assistant, University of Pennsylvania's Social Responsibility Advisory Committee (SRAC) Organizing Committee, Penn-Rutgers-Princeton Social Epistemology Workshop Departmental Representative, Penn School of Arts and Sciences Graduate Student Government Coordinator, Penn Philosophy of Science Reading Group Session Chair, Penn-Rutgers-Princeton Social Epistemology Workshop Kovaka CV 4/7 OTHER ACADEMIC AND PROFESSIONAL ACTIVITY___________________________ 2016-present 2015-present 2013-2015 2013-2015 Member of the Computational Social Philosophy Lab Member of the Penn Laboratory for Understanding Evolution Member of Timothy Linksvayer’s Social Evolution Laboratory (Penn Biology Department) Graduate Associate, Harrison College House COURSEWORK_______________________________________________________________ *=Audit Philosophy of Science Philosophy of Biology (Michael Weisberg) Philosophy of Science (Michael Weisberg) Biological Concepts of Race* (Michael Weisberg) Biology Advanced Evolution (Joshua Plotkin and Paul Sniegowski) Advanced Ecology (Brent Helliker and Erol Ackay) Evolutionary Ecology* (Timothy Linksvayer) Political Philosophy Topics in Philosophy of Law: Political Authority and Obligation (Stephen Perry) Topics in Political Philosophy: (Kok-Chor Tan) Topics in Philosophy of Law: Epistemology and Democracy* (Alexander Guerrero) Expertise: It's Nature and Uses* (Alexander Guerrero) History of Philosophy Kant I: Critique of Pure Reason* (Rolf Horstmann) Continental Rationalism (Karen Detlefsen) Aristotle’s Theoretical Philosophy (Susan Sauve Meyer) Origins of Analytic Philosophy (Joshua Armstrong) Other Proseminar (Elisabeth Camp) Formal Logic (Scott Weinstein) Topics in Philosophy of Psychology: Epistemic Realism (Gary Hatfield) Topics in Epistemology (Daniel Singer) Contemporary Ethics: Metaethical Rationalism* (Errol Lord) Kovaka CV 5/7 PROFESSIONAL MEMBERSHIPS______________________________________________ American Philosophical Association (APA) International Society for the History, Philosophy, and Social Studies of Biology (ISHPSSB) Philosophy of Science Association (PSA) REFERENCES________________________________________________________________ Michael Weisberg Professor of Philosophy University of Pennsylvania 215.898.0417 [email protected] Timothy Linksvayer Assistant Professor of Biology University of Pennsylvania 215.573.2657 [email protected] Quayshawn Spencer Assistant Professor of Philosophy University of Pennsylvania 215.573.5120 [email protected] Matthew Haber Associate Professor of Philosophy University of Utah 530.848.5579 [email protected] Alexander Guerrero Associate Professor of Philosophy Rutgers University 646.250.1375 [email protected] Daniel Singer (Teaching) Assistant Professor of Philosophy University of Pennsylvania 920.474.6437 [email protected] Kovaka CV 6/7 DISSERTATION SUMMARY___________________________________________________ Biologists have a long history of arguing about the relative importance of organisms and environments in driving evolution. Do organisms passively respond to their environments or actively shape them? Is the environment a filter that removes the least fit organisms from each generation, or also a source of new traits? The most recent incarnation of these debates focuses on developmental plasticity, a developing organism's sensitivity to environmental inputs. All organisms are plastic to some degree. Many can change their sex, morphology, and behavior in response to their environments. The question for biologists is, does the widespread presence of developmental plasticity mean new traits can originate in response to new environmental conditions, and prior to genetic changes? According to one influential hypothesis, plasticity is a significant source of novel traits in evolution. I make three arguments regarding this plasticity-first hypothesis. First, that it has revisionary implications for evolutionary theory. Second, that confirming the hypothesis will require biologists to shift their methodological priorities. Third, that the ongoing debate about the hypothesis is scientifically productive. I argue that if plasticity is a source of novel traits in evolution, we should revise our theory of biological inheritance. In particular, we should abandon the common view that the most important extra-genetic inheritance mechanisms (inheritance mechanisms other than DNA transmission) are the ones that are most similar to DNA transmission. The common view is wrong because plasticity allows for two kinds of inheritance mechanisms: those that pass on adapted developmental resources, such as genes and epigenetic marks, and those that pass on non-adapted resources, such as habitats. These two kinds of inheritance play different evolutionary roles, and biologists have ignored a potentially important class of evolutionary process by focusing so much on the paradigm case. In this, as in other instances, science must proceed with caution when taking one process as the paradigm for understanding others. The plasticity-first hypothesis has been a source of controversy among biologists for several decades. Despite plenty of new data, the main points of disagreement in the debate have hardly shifted. I show that the debate suffers from an underdetermination problem, that is, the present state of the evidence is not sufficient to confirm or disconfirm the hypothesis. This problem arises because of the tremendous difficulties associated with uncovering evidence about the details of ancient evolutionary processes. I argue that at present, biologists are looking for the wrong kind of evidence, and I offer a way to break this longstanding impasse. To uncover evidence that can resolve the debate, biologists need to make use of a richer set of methodological resources, especially formal modeling and experimental evolution. The strategy I propose is also a promising one for resolving stubborn underdetermination problems in other areas of science. The debate about plasticity in evolution is an example of a particular kind of scientific controversy called a relative significance controversy. A relative significance controversy is a scientific disagreement about how to weight the importance of multiple hypotheses that are all part of the explanation of a given class of phenomena. A famous example is the controversy over the relative importance of genes and the environment in determining IQ. Many philosophers and scientists are skeptical about the value of these controversies. They think precise estimates of the importance of genes relative to IQ, or of plasticity-first evolution relative to gene-first evolution, are not scientifically interesting. I respond to their skepticism by offering two reasons to think relative significance controversies are valuable and productive. First, they help scientists identify a kind of underdetermination problem called contrast failure. Second, they are scientists' way of determining the scope of causal patterns, a task that is central to scientific explanation. Kovaka CV 7/7
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