Lundquist and Loheide, Sierra ET vs. Elevation page 1 of 46 1 How evaporative water losses vary between wet and dry water years as a function of 2 elevation in the Sierra Nevada, California and critical factors for modeling 3 Jessica D. Lundquist1 and Steven P. Loheide, II2 4 5 1 6 7 2 Civil and Environmental Engineering, University of Washington, Seattle, WA Civil and Environmental Engineering, University of Wisconsin-Madison, Madison, WI 8 9 submitted September 28, 2010 10 revised January 24, 2011 11 accepted February 11, 2011 12 13 to Water Resources Research, please consider for special Steve Burges Issue 14 15 Corresponding author address: 16 Jessica D. Lundquist 17 Civil and Environmental Engineering 18 University of Washington 19 Seattle, WA 98195-2700 20 Phone: (206) 685-7594 21 Fax: (206) 685-3836 22 [email protected] Lundquist and Loheide, Sierra ET vs. Elevation 23 page 2 of 46 Abstract 24 25 High altitude basins in the Sierra Nevada, California have negligible summer precipitation and 26 very little groundwater storage, making them ideal laboratories for indirectly monitoring changes 27 in evaporative losses between wet and dry years. Dry years typically have greater potential 28 evapotranspiration (ET), due to warmer June and July air temperatures, warmer summer 29 water/soil temperatures, greater solar radiation exposure due to less frequent cloud cover, greater 30 vapor pressure deficit, and longer growing seasons. However, dry years also have limited 31 moisture availability compared to wetter years, and thus actual evapotranspiration is much less 32 than the potential in dry years. The balance of these factors varies with elevation. Here, we use 33 gridded temperature, precipitation, and snow data, along with historic streamflow records in two 34 nested basins of the Merced River, California and a simple model to determine the following: 35 Annual ET increases in wetter years at mid-elevations (2100-2600 m), but this pattern can only 36 be represented in model simulations that include some representation of water transfer between 37 higher and lower elevation soil reservoirs. At higher elevations (>2600 m), greater water 38 availability in wet years is offset by shorter growing seasons due to longer snow cover duration. 39 These results suggest that models seeking to represent changes in ET in mountainous terrain 40 must, at a minimum, include both hillslope processes (water transfer down steep slopes) and 41 snow processes (timing of water and energy supply). 42 Lundquist and Loheide, Sierra ET vs. Elevation 43 44 page 3 of 46 1. Introduction Research has shown that temperatures in the western U.S. have been warming [Bonfils et al. 45 2008; Dettinger and Cayan 1995] and that snowmelt timing has shifted earlier in the season 46 [Stewart et al. 2005]. Both trends are likely to continue [Stewart et al. 2004]. These changes beg 47 the question if, and how, evapotranspiration (ET), which makes up approximately 30 to 50% of 48 the annual water budget in the Sierra [e.g., Jeton et al. 2004], is likely to change under these 49 circumstances. For example, recent modeling studies for projected climate impacts in the 50 Colorado River basin show a range of climate impacts on Colorado River runoff ranging from 51 5% reduction in flows by 2050 [Christensen and Lettenmaier 2007] to 20% reduction in flows by 52 2050 [Milly et al. 2005], with one of the major differences being due to how ET and its relation 53 to snow cover is handled in the models [Hoerling et al. 2009]. ET in cold regions, specifically 54 how ET relates to snow and ice processes, was highlighted by Shuttleworth [2007] as an area 55 with substantial need for more research and understanding. 56 Overall, changes in ET depend critically on the balance between energy and moisture 57 available for evaporation [Monteith and Unsworth 1973; Brutsaert 1982; Jeton et al. 1996]. 58 Warmer temperatures increase potential ET, and an earlier disappearance of the snowpack 59 reduces albedo, warms soil and air temperatures, and increases potential ET. However, as soils 60 dry out earlier [e.g., Hamlet et al. 2007], plants close their stomata and reduce transpiration [e.g., 61 Goldstein et al. 2000], resulting in actual ET that is much less than potential. 62 The mountains of the Western United States, and particularly the Sierra Nevada of 63 California, provide an interesting test case. Here, the majority of precipitation falls in the winter, 64 in the form of snow, and plants depend on stored moisture from snowmelt during the summer 65 growing season. A larger snowpack serves both to potentially decrease ET (through a longer Lundquist and Loheide, Sierra ET vs. Elevation page 4 of 46 66 period of snow cover, low temperatures, high albedo, and plant dormancy) and to potentially 67 increase ET (through a greater summer water supply). 68 It is still uncertain whether ET in this region is likely to increase, decrease, or remain the 69 same with warmer temperatures and earlier snowmelt. Fortunately, interannual climate 70 variations in the western U.S. span a range of precipitation larger than any projections of future 71 mean change [Cayan et al. 2008]. Risbey and Entekhabi [1996] examined data from the 72 Sacramento Basin, CA and found that annual streamflow responded to changes in annual 73 precipitation and not to changes in annual temperature. Thus, any climate impact on ET in this 74 region is likely to propagate from the temperature impact on snowpack. Therefore, we can 75 examine how ET varied in wet (late melt) vs. dry (early melt) water years in the past to gain 76 insight regarding future change and/or variability. 77 Tague et al. [2009] and Christensen et al. [2008] both looked at the differences in ET 78 with elevation in the Merced River above Happy Isles as represented by the RHESSys model 79 [Tague and Band 2004] under current and future climates. They found that transpiration at 80 middle elevations (1800-2150 m) was primarily water-limited, with the most transpiration in wet 81 years, whereas the highest elevations (2600-4000 m) were more energy-limited and responded to 82 temperature. Thus, ET increased under future warming scenarios for only the higher elevations, 83 and decreased for the lower elevations, resulting in negligible changes over the watershed as a 84 whole. 85 Here, we use gridded temperature, precipitation, and snow data, along with historic 86 streamflow records in two nested basins of the Merced River, California and a simple model to 87 address two key questions: 1) How does ET vary across elevations and years in the Sierra Lundquist and Loheide, Sierra ET vs. Elevation page 5 of 46 88 Nevada? and 2) What key processes are essential to include in any model that strives to 89 represent variability in ET across elevations and years in snowy mountain terrain? 90 91 2. Observational Data and Methods 92 2.1 Study Area 93 We focus our study on the Merced River Basin within Yosemite National Park, 94 California, which has been gauged at two locations, Happy Isles (USGS gage 11264500) and 95 Pohono (USGS gage 11266500) since 1916 (Figure 1a). Elevations range from about 1050 m at 96 the Pohono gage to nearly 4000 m at the top of Mt. Lyell. The region receives most of its 97 precipitation from October through May in the form of snow. The basin is underlain by 98 granodiorite, and soils are shallow [Huber 1987]. Because of its long streamflow record and 99 unimpaired flows, the Merced River has been the subject of many prior modeling studies and 100 101 analyses [e.g., Clow et al. 1996; Dettinger et al. 2004; Christensen et al. 2008]. The nesting of the Happy Isles basin within the Pohono basin provides a unique 102 opportunity to parse out different processes occurring at high vs. low elevations. As seen in 103 Figure 1b, all of the contributions to the Pohono gage from elevations above 3200 m come from 104 the Happy Isles sub-basin. If we consider the contributing areas above and below the Happy 105 Isles gage separately, we see that the lower basin has the middle 50% of its contributing area 106 between 2170 and 2580 m, and the upper basin has the middle 50% of its area between 2420 and 107 3050 m elevation (Fig. 1b). The area-weighted mean of the lower basin is 2320 m and of the 108 upper basin is 2750 m. The different elevations of these two basins control where water at the 109 Pohono gage originates from at different times of year (Fig. 2). Winter streamflow peaks 110 originate primarily from the lower basin, where a greater fraction of precipitation falls as rain Lundquist and Loheide, Sierra ET vs. Elevation page 6 of 46 111 than as snow. Late summer streamflow originates almost entirely from the upper basin, the only 112 region with remaining late-lying snow (Fig. 2). Note that the entire system has limited deep 113 groundwater flow, so that streamflow is closely linked to rain and snowmelt processes. 114 115 116 2.2 Data Annual runoff was calculated as the sum of daily USGS streamflow records for Happy 117 Isles and Pohono, from 1 October to 30 September each water year from 1981 to 2007. Both of 118 these gages have a quality classification of “good” (<10% error) [Rockwell et al. 1997]. To 119 emphasize differences between elevations, we subtracted the streamflow measured at Happy 120 Isles from the streamflow measured at Pohono to estimate the contribution from the lower 121 elevations (Fig. 1). From here on, we will refer to the “upper basin” as the areas contributing to 122 the Merced River gauge at Happy Isles, and the “lower basin” as the areas contributing to the 123 Merced River gauge at Pohono with the Happy Isles contribution removed. 124 Precipitation data were obtained from the Parameter-elevation Regressions on 125 Independent Slopes Model’s (PRISM, available at http://www.prism.oregonstate.edu) monthly 126 precipitation maps for water years 1981 to 2007 at 4-km resolution [Daly et al., 1994, 2008]. 127 PRISM uses empirical relationships to account for the influence of elevation, rain shadows, and 128 coastal proximity when interpolating between existing measurement stations. The data from 129 PRISM have been quality-controlled and peer-reviewed, and are widely used in modeling 130 applications [Daly et al., 2002]. This time period (post-1980) was chosen because high altitude 131 snow pillows, which determine snow water equivalent by weighing the snowpack, were first 132 installed in the region around 1980, providing guidance on upper elevation precipitation 133 distributions. For water-year analysis, all of the PRISM monthly maps from October to the Lundquist and Loheide, Sierra ET vs. Elevation page 7 of 46 134 following September were summed. The 4-km maps were then sub-sampled at a 100-m 135 resolution, and all grid cells falling within a basin’s boundaries were summed to represent the 136 total annual precipitation that fell within that watershed. Precipitation in the upper and lower 137 Merced basins was calculated separately. 138 Snow-covered area (SCA) and its rate of change with time were calculated from daily 139 500-m gridded MODIS maps of fractional SCA for water years 2002 to 2007, which were 140 created using the MODSCAG algorithm [Dozier et al. 2008; Dozier and Frew 2009; Painter et al. 141 2009]. Similar to treatment of the PRISM gridded data, these daily grids were sub-sampled to 142 100 m and clipped to represent SCA in the upper and lower Merced basins, respectively. Within 143 each of the two watersheds, fractional SCA was then calculated separately for each 200-m 144 elevation band, ranging from 1200 m to 4000 m. 145 Meteorological data for the 2002-2007 period were obtained from the Dana Meadows 146 Snow Pillow Site (at 3000 m, Fig. 1) just to the north of the two watersheds. Temperature, 147 relative humidity, wind speed, and incoming solar radiation were measured hourly and converted 148 to daily averages and daily maximum and minimum temperatures. Daily precipitation data was 149 measured at the Tuolumne Meadows Snow Pillow Site (2600 m, Fig. 1). 150 151 2.3 Methods: Water balance approach to estimating ET 152 Over longer time-periods, such as the annual average, basin storage in steep basins with 153 154 shallow soils can be considered negligible, such that P = R + ET , (1) 155 where P is total annual precipitation, R is total annual runoff, and ET is total water lost to 156 evapotranspiration and sublimation [Adam et al. 2006]. Because precipitation and runoff are Lundquist and Loheide, Sierra ET vs. Elevation page 8 of 46 157 both easier to measure than ET, ET is typically estimated as the residual of equation (1). The 158 catchment water balance method of calculating ET agrees within 10% of upscaled direct eddy- 159 covariance flux measurements of ET [Miller et al. 2007, at AmeriFlux sites; Wilson et al. 2001, 160 at a 97.5 ha watershed in Tennessee], although errors are potentially larger in more extensive 161 areas of complex terrain. To compare interannual variations in ET at different elevations, we 162 constructed annual water balance estimates for the upper and lower basins of the Merced River 163 (Fig. 1) for 1981 to 2007, using PRISM maps of precipitation and USGS streamflow records, as 164 outlined in Section 2.2. Assuming no net changes in water storage between one year and the 165 next, annual ET was calculated as the difference between precipitation and runoff. 166 167 168 3 Observational Results Fig. 3 illustrates 27 water years (October-September) of area-normalized annual basin 169 precipitation (from PRISM), discharge (from USGS gages), and ET (from the difference between 170 the prior two) for (a) the upper basin (above Happy Isles) and (b) the lower basin (between 171 Happy Isles and Pohono). Area-normalized precipitation varies from 0.6 to 2 meters, and 172 discharge varies from 0.3 to 1.5 m. ET varies much less than either precipitation or discharge, 173 but ranges from 0.3 to 0.5 m in the Upper Basin, and from 0.4 to 0.8 m in the Lower Basin. 174 To better illustrate the dependence of ET on wet vs. dry years, Fig. 4 plots the annual ET 175 shown in Fig. 3 as a function of basin-average annual precipitation. In the Lower Basin, ET 176 increases in wetter years, with a significant correlation (p<0.01) and a best-fit slope of 13%. 177 This suggests that the 2170 - 2580 m elevation range that dominates this Lower Basin is water- 178 limited, with ET responding to increases in precipitation to a greater extent than to changes in 179 temperature. These observations agree with other Sierra studies of ET in this elevation range Lundquist and Loheide, Sierra ET vs. Elevation page 9 of 46 180 [e.g., Risbey and Entekhabi 1996; Jeton et al 1996; Miller et al. 2007; Christensen et al. 2008]. 181 ET in the Upper Basin, which has the middle 50% of its area between 2420 and 3050 m 182 elevation, has no significant trend, although ET appears to decrease in both the wettest and driest 183 years on record, suggesting that it is sometimes energy-limited and sometimes water-limited. To 184 better understand what controls these patterns and what parameters are essential to model them, 185 we create a simple model of basin ET, detailed in Section 4. 186 187 188 189 4 Basin ET Model 4.1 Model Set-up Model representations of ET and water availability vary widely, ranging from relatively 190 simple physical parameterizations in global climate models [Cornwell and Harvey 2007], to 191 extensively parameterized stomatal responses in vegetation canopy models [e.g., Running and 192 Coughlan 1988; Tague and Band 2004]. 193 194 195 We start with the commonly-used basis for ET modeling, the Penman-Monteith equation [e.g., Shuttleworth, 1993; Allen et al., 1998]: ET = Δ ( Rnet − G ) + ρ air c p (e s − e) / ra λρ water (Δ + γ (1 + rs / ra ) ) (2) 196 where Δ is the gradient of the saturated vapor pressure with temperature, des/dT (kPa °C-1), Rnet is 197 the net radiation (W m-2), G is the ground heat flux (W m-2), ρair is the density of air (kg m-3), cp 198 is the heat capacity of air (J kg-1 °C -1), es is saturated vapor pressure (kPa), e is vapor pressure 199 (kPa), γ is the psychometric constant (kPa °C -1), λ is the latent heat of vaporization of water (J 200 kg-1), ρwater is the density of water (kg m-3), rs is the surface resistance (m s-1), ra is the 201 aerodynamic resistance (m s-1), and P is atmospheric pressure (kPa). Lundquist and Loheide, Sierra ET vs. Elevation 202 page 10 of 46 We used the approximations of rs and ra laid out by the FAO (Food and Agriculture 203 Organization of the United Nations) to adjust the Penman-Monteith equation to reference 204 (potential) evapotranspiration [Allen et al. 1998]. We divided the basin into 200-m elevation 205 bands, which were weighted by area, according to basin hypsometry. Daily maximum and 206 minimum temperatures were distributed to the center elevation of each zone assuming a 6.5°C 207 km-1 lapse rate, the average lapse rate for the region found by Lundquist and Cayan [2007]. 208 Wind speed and solar radiation were assumed constant with elevation. Atmospheric pressure 209 was a function of elevation [Allen et al. 1998, their equation 7] but was fixed in time. Measured 210 relative humidity at the Dana meteorological station (Fig. 1) was used to determine the mixing 211 ratio (g kg-1 of water in the air), which was assumed constant with elevation. Albedo was set to 212 0.15, a typical value for open forests [Jarvis et al. 1976; Brandes and Wilcox 2000]. We 213 neglected ground heat flux, which is much smaller than net radiation and can generally be 214 neglected over daily time scales [Allen et al. 1998]. This is especially true in the Sierra Nevada, 215 where soils do not freeze but rather, remain at or near 0˚C throughout the snow season 216 [Lundquist and Lott 2008]. 217 Actual ET was assumed to deviate from potential ET due to three main causes. First, we 218 assumed that ET did not occur at elevations above tree line (>3000 m) when fractional snow 219 covered area (SCA) exceeded a set threshold, where snow cover was represented by direct 220 MODIS observations of fractional SCA (see Section 2.2). Second, we assumed that no ET 221 occurred at any elevation on days when minimum temperatures at that elevation fell below a set 222 threshold, indicating a hard freeze, based on studies showing that low soil and air temperatures 223 inhibit plant metabolic activity [e.g., Troeng and Linder 1982; Tanja et al. 2003; Mellander et al. Lundquist and Loheide, Sierra ET vs. Elevation page 11 of 46 224 2004]. Third, as described in detail below, we assumed that the soil moisture deficit limited 225 summer ET. 226 The formulation of actual ET follows a “bucket” approach, similar to that used by 227 Manabe [1969], Koster and Milly [1997], Rodriguez-Iturbe et al. [1999] and Gedney et al. 228 [2000], among many others. Actual ET for each elevation band was derived by scaling potential 229 ET as a function of soil moisture: 230 ETactual = f (θ ) ET potential (3) 231 [following Feddes et al. 1978; Laio et al. 2001; D’Odorico and Porporato 2006], where θ is the 232 volumetric soil moisture. f(θ) is set to 1 above a critical soil moisture (θc, which depends on both 233 soil and vegetation properties), to 0 below the wilting point (θwp), and varies linearly in between 234 these values; this functional form was found to work best for forests by Brandes and Wilcox 235 [2000]. The model set-up for determining soil moisture is illustrated in Figure 5. Soil water in 236 each elevation band is represented by a simple bucket with root zone depth (Zr), set porosity (n), 237 rate of soil drainage (K), and field capacity (θfc). Water is added to the bucket through liquid 238 precipitation (measured at the Tuolumne Meadows snow pillow, Fig. 1, and determined to be 239 liquid if Tmean>1°C) and through snowmelt, which is calculated as the product of mean daily air 240 temperature times a melt factor, which follows the formulation in Snow-17 [Anderson 1973], 241 using calibration parameters determined by Shamir and Georgakakos [2005], on all days when 242 Tmean>0°C and when snow covered area (SCA) greater than 10% is observed at that elevation by 243 MODIS. Soil water in excess of porosity is assumed to run off via overland flow and be lost to 244 the soil water system. Soil water less than porosity but in excess of field capacity drains from 245 soil storage by the following equation from Rodriguez-Iturbe et al. [1999]: Lundquist and Loheide, Sierra ET vs. Elevation transferout = 246 page 12 of 46 K (θ − θ fc ) (n − θ fc ) (4) 247 In our model, we use this to not only represent vertical drainage from the soil column, but also 248 lateral drainage through interflow and hillslope processes. Thus K, as used here, describes rates 249 of both vertical water movement and downslope lateral transfer, and thus represents a rate of 250 drainage rather than a true hydraulic conductivity. Water ceases draining as soon as soil 251 moisture reaches field capacity. Some portion of this water transferred down slope will enter 252 streams and be removed from the watershed, but some portion will be transferred from the soil 253 zone in one elevation band to that in the next lower elevation band. The percentage of water that 254 is transferred out and that reaches the soil in the next lower elevation band is set by a transfer 255 coefficient (tc). The remaining percentage (1- tc) is presumed to join a stream or river and be 256 exported from the system, unavailable for ET. Thus, soil moisture in one elevation band is 257 defined as follows: Δθ = 258 P + Melt + transferin − ET − transferout Zr (5) 259 where the transfer in is equal to the transfer coefficient times the leak out at the next higher 260 elevation band. Note that the total efficiency of water transferred for the basin as a whole is a 261 function of the number of elevation bands, and thus tc would change as the resolution of 262 elevation bands changes. 263 264 4.2 Design of Modeling Experiments 265 To better understand parameter sensitivity and importance, we ran the model for 5 water 266 years from 2003-2007. We chose this period because a) this period has complete, quality- 267 controlled meteorological data at high elevation to calculate potential ET; b) this period has Lundquist and Loheide, Sierra ET vs. Elevation page 13 of 46 268 quality-controlled daily MODIS information on SCA; and c) this period contains 2 years in the 269 highest quartile of the 91 years of Merced river streamflow (2005 and 2006) and 2 years in the 270 lowest quartile (2004 and 2007), which aid in comparing model sensitivity to wet vs. dry years. 271 We examined which model parameters and assumptions could adequately represent the mean 272 and inter-annual variations in soil moisture, as a function of elevation, in the Sierra Nevada by 273 assessing our model’s ability to fit the following values as determined from the water balance 274 calculations: 1) mean annual ET (2003-2007) for the upper basin (36 cm); 2) mean annual ET 275 (2003-2007) for the lower basin (54 cm); 3) mean annual difference between dry years (2004 and 276 2007) and wet years (2005 and 2006) for the upper basin (3 cm) and 4) mean annual difference 277 between dry years (2004 and 2007) and wet years (2005 and 2006) for the lower basin (-12 cm). 278 Individual values for each component of the water balance for these years are presented in Figure 279 3 and in Table 2. 280 For these runs, we held the following parameters constant: The soil moisture where ET 281 begins to be limited was set at 0.25, except in cases where field capacity (θfc) was less than 0.25, 282 when it was set to equal field capacity. The wilting point was set at 0.10. Porosity was set at 0.4 283 based on basin soil maps [USDA NRCS 2006]. The temperature threshold at which ET stopped 284 was Tmin = -1°C. The fraction of SCA above which ET could not occur above treeline was set to 285 0%. Sensitivity to each of the above parameters was investigated, but was found to be less than 286 the sensitivity to the varied parameters discussed below. 287 A sensitivity analysis was performed on the remaining parameters by modeling a range of 288 reasonable values in all reasonable combinations (determined based on field measurements and 289 basin soil maps, USDA NRCS 2006) to investigate which best fit the water balance observations. 290 The rate of soil drainage, K, varied by orders of magnitude from 0.001 to 1 m day-1. Field Lundquist and Loheide, Sierra ET vs. Elevation page 14 of 46 291 capacity varied from 0.15 to 0.4. The transfer efficiency of water from soil in one elevation band 292 to the next varied from 0 to 75%, and the rooting depth, which is the same as the soil depth in 293 these shallow Sierra soils, varied from 0.25 to 1.5 m. 294 295 4.3 Results 296 The model accurately simulated mean annual ET for both the upper basin and the lower basin 297 with a wide variety of parameter sets (lower basin shown in Fig. 6; upper basin patterns 298 qualitatively similar, not shown). When soil drainage is slow (K ≤ 0.01 m day-1), mean ET in the 299 upper basin is insensitive to field capacity (see horizontal lines in top rows of Fig. 6) because soil 300 moisture stays above field capacity for much of the growing season. Rather, annual ET is most 301 sensitive to the rooting depth (Zr), because this represents the total water available. As the rate of 302 soil drainage (K) increases, other parameters become more important. When drainage is fast 303 (e.g., bottom row in Fig 6, K = 1 m day-1), soil moisture quickly drops to field capacity, so θfc 304 and Zr both influence the mean annual ET. With no water transfer (tc=0), ET increases with both 305 increasing θfc and increasing Zr. 306 As the transfer of water from upper to lower elevations increases, the relationship to θfc 307 becomes more complex, with a local maximum ET near θfc =0.35, with lower ET for both higher 308 and lower field capacities. This occurs because of the interplay between field capacity, porosity, 309 and water transfer in the model, which jointly control drainage and downslope water transfer. 310 Porosity (n) was held fixed at n=0.4 for all model simulations. Soil water in excess of porosity is 311 assumed to run off the surface to rivers and thus is unavailable for ET at lower elevations. Thus, 312 only soil water content both greater than θfc and less than n is available for subsurface transfer to 313 lower elevation soils. As θfc increases, more water is held locally, which acts to increase ET. Lundquist and Loheide, Sierra ET vs. Elevation page 15 of 46 314 However, as θfc approaches 0.4 (the soil porosity), less water is available for transfer to lower 315 elevations, up to the limit of θfc =0.4, where no water is transferred, regardless of what the 316 transfer coefficient is. (Notice that the ET values at θfc =0.4 are identical for all values of tc (Fig. 317 6).) Because transfer of water from higher to lower elevations acts to increase annual ET, this 318 decrease in water transfer offsets the local increase in water availability for field capacities 319 greater than 0.35. 320 For soil rooting depths ≤ 1 m, only model simulations with 50-75% transfer efficiency and 321 relatively fast water movement through the soil calculated annual average ET as high as the 322 observed value (54 cm). Based on soil surveys [USDA NRCS 2006], only isolated meadows and 323 wetlands in the region have soils deeper than 1 m, and most basin areas (forested slopes) have 324 soils shallower than 1 m. These results imply that transfer of water from upslope (higher 325 elevations) to downslope (lower elevations) is essential to provide enough water for the observed 326 ET values in the lower basin. 327 Requiring the model to match the observed differences between wet and dry years further 328 constrains the acceptable parameter sets. In both basins (Fig. 7 and Fig. 8), when drainage was 329 slow (K ≤ 0.01 m day-1, top two rows), there was little difference in annual ET between dry and 330 wet years. This corresponded to an over-prediction of 12 cm for the wet-dry ET difference in the 331 lower basin and an under-prediction of 2.5 cm for the wet-dry ET difference in the upper basin. 332 With slow drainage, the water available each year is set primarily by the rooting depth and does 333 not vary between years. 334 A wide range of parameter sets led to simulations with ET that was relatively constant 335 between wet and dry years. However, the model only simulated greater ET in wet years than in 336 dry years, as observed in the lower basin, when it was parameterized with a high transfer Lundquist and Loheide, Sierra ET vs. Elevation page 16 of 46 337 coefficient, high K, and mid-range θfc (Fig. 7). With these parameters, the model transferred 338 more water to lower elevations to support ET in wetter years (when snow lasts much longer at 339 higher elevations) than drier years. As discussed with the mean annual ET, larger values of θfc 340 that approach porosity result in less overall transfer between elevations, offsetting this effect. 341 Note that the parameters that best fit the observed difference in the lower basin do not best fit the 342 observed difference in the upper basin (Fig. 8). This difference between parameterization of the 343 upper and lower basins is not an unexpected result given large differences in vegetation (Table 3, 344 also see Lutz et al. 2010), slopes, and soils across this elevation gradient. 345 Figure 9 illustrates the processes leading to the model results summarized in Fig’s 6 to 8 346 for parameters that fit the lower basin well (tc = 0.75; K = 1 m day-1; Zr = 0.75 m; θfc = 0.3). For 347 a fast draining soil (K = 1 m day-1), soil moisture stays close to field capacity so long as local 348 snowmelt, or snowmelt transferred in from higher elevations, sustains it (Fig. 9 a, b, c, d, e, f). 349 The rate of snowmelt (so long as snow is available to melt), exceeds the rate of ET by almost a 350 factor of 10 (Fig. 9 c, d, g, h), so Sierra soils are saturated under snowmelt conditions (see Flint 351 et al. 2008 for an example of this in the Merced Basin). In this situation, the primary factors 352 limiting ET are the date ET begins (limited by SCA at elevations above treeline and by minimum 353 temperatures at all other elevations), the date snowmelt stops (both local SCA<10% and 354 combined effects of higher elevation snowmelt and transfer efficiency), the depth of the local 355 soil reservoir (Zr), and the ability of the soil to hold onto water once snowmelt input ceases (K 356 and θfc). For this parameter set, more ET occurs in the lower basin (example, 2300 m elevation 357 band) in the wetter year (2006, Fig. 9g) than the drier year (2004, Fig. 9h) because moisture 358 limitation begins in July in 2004 (Fig. 9e) but not until mid-August in 2006 (Fig. 9f). Lundquist and Loheide, Sierra ET vs. Elevation 359 page 17 of 46 The lowest elevations, where more precipitation falls as rain, are moisture-limited for all 360 reasonable parameter sets, while the highest elevations, above treeline where snowcover lasts 361 late into the summer, are energy-limited for all reasonable parameter sets (Fig. 10a). Only 362 intermediate elevations (2000-3000 m) are strongly sensitive to the parameter set chosen. 363 However, as shown in Fig. 1b, these same intermediate elevations are the ones that dominate the 364 runoff in both the upper and lower watersheds examined. In Fig. 10, these intermediate 365 elevations tend towards water limitation (more ET in wet years), but for other parameter sets (not 366 shown) can tend towards an even balance or towards energy limitation. The majority of 367 parameters result in annual ET at these elevations that stays relatively fixed from one year to the 368 next. With the parameters of tc = 0.75; K = 1 m day-1; Zr = 0.75 m; and θfc = 0.3, the model 369 370 matches the observed mean and variability for the lower basin well (Fig. 10b). However, while 371 it approximates the mean ET in the upper basin, it does not well-represent the upper basin’s 372 interannual variability. This may be due to error in PRISM precipitation variations at the highest 373 elevations due to sparse observations above 3000 m (which would bias the observed patterns) or 374 to high elevation vegetation behavior differing from the parameters set here. 375 376 377 5 Summary and Discussion In the Sierra Nevada, California, annual ET is limited more by moisture availability than 378 energy availability, and thus, annual ET is unlikely to be strongly affected by warming 379 temperatures and earlier snowmelt, with the exception of the highest elevations above treeline, 380 which currently remain snow-covered throughout most of the growing season. These observed 381 water balance results agree qualitatively with earlier modeling studies [e.g., Christensen et al. Lundquist and Loheide, Sierra ET vs. Elevation page 18 of 46 382 2008 and Tague et al. 2009]. The new insight we have gained relates to which model 383 components matter most in recreating these results. Mean annual ET can vary by as much as 30 384 cm depending on the basin soil properties (Fig. 6). When soil properties are not well known, this 385 results in a poorly-constrained parameter space that can be set in a variety of configurations to 386 match observed precipitation and runoff. Fortunately for studies of change in ET between years 387 (such as climate change experiments), the interannual variability in ET is less sensitive to the soil 388 parameterization (Fig. 7 and Fig. 8), with about a 10 cm range in the calculated difference 389 between dry and wet years depending on the parameter set chosen. For this region, with about 390 100 cm annual average precipitation, a 10 cm range corresponds to approximately 10% 391 uncertainty. 392 However, our work here demonstrates that this 10 cm range is primarily controlled by one 393 parameter. Specifically, water transfer is the most important parameter required to model more 394 ET in wetter years in mid to low snow elevations in the Central Sierra. This transfer represents 395 movement of water from higher, cooler locations with snow to lower, warmer elevations with 396 transpiring vegetation, which is an important feature of steep terrain. This downslope transfer of 397 water in our model is intended to represent several physical mechanisms, including the 398 following: interflow along the soil-bedrock interface, perched groundwater flow in glacial 399 moraines, overland flow particularly in regions with exposed granitic domes, groundwater flow 400 through the bedrock fractures, and streamflow which can provide water to downstream riparian 401 areas. In our basin, most of this transfer is likely due to interflow, which is partitioned into base- 402 flow to rivers and shallow down-slope perched groundwater flow. Observations and modeling of 403 groundwater levels in the nearby Tuolumne Meadows indicate that these hillslope water fluxes 404 are an essential component of the meadow groundwater budget [Lowry et al. 2010]. While we Lundquist and Loheide, Sierra ET vs. Elevation page 19 of 46 405 have modeled this water movement as the gross downslope movement of water from one 406 elevation band to the next, each individual process is much more complex, operating at scales 407 ranging from a single hillslope, for processes such as interflow, to basin-wide, for processes such 408 as flow in the bedrock aquifer. Our model demonstrates that while it may not be necessary to 409 simulate all of the complexities of these spatially variable transfer mechanisms, at least a crude 410 representation of downslope water transfer is required to simulate the observed variability in ET. 411 When interpreting our results, we must be clear that estimating ET from the water balance is 412 subject to uncertainty in runoff, to uncertainty in the measurements and distribution of 413 precipitation, and to uncertainty in the assumption that storage does not change between years. 414 As stated previously, the USGS streamflow measurements used here are among the most 415 accurate in the network, with relatively small uncertainty on an annual basis. Characterizing the 416 uncertainty in the precipitation distribution across complex terrain is more difficult, and we 417 believe this to be the largest source of error in our estimates of ET (see Lundquist et al. 2010 for 418 further discussion). To check the robustness of our results to potential errors in basin-wide 419 precipitation, we also employed an alternative approach to estimating ET using precipitation data 420 collected at the Yosemite Valley Park Headquarters and streamflow at our two USGS sites from 421 1916 to 2000. (Reliable precipitation data was not available at this location after 2000.) To map 422 point precipitation to each basin’s areal-average precipitation, we used the mean annual areal 423 precipitation calculated by PRISM divided by the mean annual precipitation measured at the 424 gage for the period of overlap (water years 1981-2000) to calculate a constant weight function 425 for each basin. Despite the assumption of a constant precipitation distribution each year in this 426 alternate estimation of precipitation, the same general pattern emerged as in our 1980-2007 427 dataset. Namely, more ET occurred in wet years than in dry years in the lower basin, while ET Lundquist and Loheide, Sierra ET vs. Elevation page 20 of 46 428 remained relatively constant between wet years and dry years in the upper basin. The pattern of 429 the upper basin reaching a maximum ET in average years, with less ET for both wetter and drier 430 years (as suggested by visual inspection of Figure 4), did not appear in the historic dataset, 431 suggesting that this pattern is likely less robust. Changes in the average annual precipitation at 432 the highest elevations, where there are no gages, would change the average ET estimated by the 433 water balance but would not change the year to year variability. 434 Our analysis assumes that there is limited water storage from one year to the next, and annual 435 changes in storage would introduce error into our estimates of ET. We assumed annual changes 436 in storage are small because the Merced basin is primarily granite overlain with shallow soils, 437 and this is supported by our model with the best-fit parameter set illustrated in Figures 9 and 10. 438 With the exception of 2007, which had a substantial late-summer rain event [Lundquist and 439 Roche 2008], all elevations below 2800 m had soil moisture reduced to the wilting point each 440 year (e.g., Figure 9). Elevations between 2800 and 3400 m had the greatest difference in end-of- 441 water-year soil storage between a dry year (2004) and a wet year (2005), with an average of 15 442 cm more soil water at the end of the wet year. Above 3400 m, there was on average of 2 cm 443 more soil water at the end of the wet year. Weighted by basin area, these changes in soil water 444 represented 6 cm more water stored in 2005/2006 than in 2003/2004 in the upper basin, but only 445 1 cm more water in the lower basin. Therefore, the differences in simulations and observations 446 for the upper basin (Figure 10) may be partially explained by changes in storage in these upper 447 elevations. However, our results for changing ET in the lower basin are not strongly influenced 448 by changes in storage and are robust. 449 450 Our results demonstrating the importance of water transfer to variations in ET are specific to the seasonal snow zone. This differs from prior work showing ET increases with increased Lundquist and Loheide, Sierra ET vs. Elevation page 21 of 46 451 precipitation in rain-dominated regions [e.g., Hickel and Zhang 2006] because water transfer 452 across elevations is less critical when rain falls during the growing season and when precipitation 453 and melt rates do not have strong elevational gradients. Such water transfer is included as part of 454 hillslope processes in some form in most distributed hydrologic models, e.g., DHSVM 455 [Wigmosta et al. 1994; see Lowry et al. 2010 for a discussion of the importance of subsurface 456 water transfer to maintain meadow water levels in steep terrain] or RHESSys [Tague and Band 457 2004], which documented similar ET patterns to those presented here [Christensen et al. 2008]. 458 However, most climate predictions are made with coarser models that neglect this key 459 process [e.g. Koster and Milly 1997; Gedney et al. 2000; Cornwell and Harvey 2007]. Our work 460 suggests that such models will not accurately represent changes in ET in response to changes in 461 precipitation and snowpack timing in steep, snow-fed terrain, and thus, should not be used for 462 this purpose. Specifically, as illustrated in Figure 6 for the lower basin, for a parameter set of K 463 = 1 m day-1; Zr = 0.75 m; θfc = 0.3, a simulation with no water transfer (tc=0) had 25 cm less 464 mean annual ET than a simulation with substantial water transfer (tc=75%). This would result in 465 a 25 cm overestimation of the amount of water available for runoff, which is substantial 466 compared to actual area-normalized runoff (30 to 150 cm). As illustrated in Figure 7, the 467 simulation with no water transfer showed no change in ET between wet and dry years in the 468 lower basin, while the simulation with water transfer (tc=75%) showed 10 cm more ET in wetter 469 years. This would result in error in the model’s estimate of how runoff varies with changing 470 precipitation. While mean errors in ET would likely be compensated for during model 471 calibration, the model without water transfer would not exhibit the correct sensitivity to climate 472 change. Based on these results, a hydrologic model with at least a crude representation of Lundquist and Loheide, Sierra ET vs. Elevation page 22 of 46 473 hillslope processes, i.e., lateral moisture transfer, is essential in hydroclimatic modeling to 474 adequately represent ET in complex terrain with seasonal snow. Lundquist and Loheide, Sierra ET vs. Elevation 475 page 23 of 46 6. Acknowledgements 476 This material is based upon work supported by the National Science Foundation under Grant No. 477 CBET-0729830 and CBET-0729838. Any opinions, findings, and conclusions or 478 recommendations expressed in this material are those of the author(s) and do not necessarily 479 reflect the views of the National Science Foundation. We thank the USGS for their dedicated 480 maintenance of the stream gages, the CA Department of Water Resources for the meteorological 481 data, and Chris Daly at Oregon State for the PRISM estimates of precipitation. 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Lundquist and Loheide, Sierra ET vs. Elevation page 31 of 46 638 8. Figure Captions 639 Figure 1. (a) Locations of gages (Merced River at Happy Isles and Merced River at Pohono) and 640 of meteorological stations (Tuolumne and Dana) in Yosemite National Park, California. (b) Area 641 as a function of elevation for the basin above Pohono gauge, the basin above the Happy Isles 642 gauge (Upper Basin), and the basin area below the Happy Isles gauge that drains to the Pohono 643 gauge (Lower Basin). 644 645 Figure 2. Discharge normalized by basin area for the Merced River at Happy Isles (Upper 646 Basin), for the Merced River at Pohono Bridge (Combined Basins), and for the Merced River 647 contributions in between Happy Isles and Pohono (Lower Basin) for water year 2006. Note that 648 during winter rain storms (near 1 January and 1 March) more runoff per unit area originates from 649 lower elevations, because snow is falling at higher elevations. Late summer runoff originates 650 primarily from higher elevations, both from snowmelt and from high-elevation summer 651 thunderstorms. 652 653 Figure 3. Annual, basin-averaged water balance components for (a) Merced River above Happy 654 Isles (Upper Basin) and (b) for the Merced River above Pohono Bridge and below Happy Isles 655 (Lower Basin) for 1992 to 2000. Evapotranspiration (ET) is estimated as the residual of annual 656 basin precipitation minus runoff. 657 658 Figure 4. Scatterplot of ET in the upper and lower basins as a function of annual basin-average 659 precipitation. For the lower basin, the best fit line has a slope of 13%, and the correlation is 660 significant (p<0.01). Lundquist and Loheide, Sierra ET vs. Elevation page 32 of 46 661 662 Figure 5. Illustration of soil water accounting for determining actual ET in the model. Model 663 details are in main text. 664 665 Figure 6. Model simulation performance relative to lower basin average annual ET for 2003- 666 2007, which is 54 cm, based on water balance calculations. Contour intervals are each 2.5 cm, 667 and a value of zero indicates a parameter set that precisely predicts the mean ET of the lower 668 basin. The y-axis of each plot shows the rooting depth (Zr). The x-axis shows the soil field 669 capacity (θfc). Columns indicate the transfer coefficient (tc), representing the percent of soil 670 water transferred between one elevation band and the next. Rows indicate the value of K. Other 671 model parameters were held fixed as follows: Porosity, n, is 0.4. Melt ceases when SCA drops 672 below 10%, and ET begins at elevations about 3000 m when SCA reaches 0. Plant water stress 673 begins at θc =0.25, and the wilting point occurs at θwp =0.10. 674 675 Figure7. Mismatch between modeled and observed mean ET differences between dry years 676 (2004 and 2007) and wet years (2005 and 2006) for the lower basin. The observed mean ET 677 differences between dry years (2004 and 2007) and wet years (2005 and 2006) for the lower 678 basin is -12 cm based on the water budget calculations (Table 2). The parameter values are the 679 same as in Fig 6. 680 681 Figure 8. Mismatch between modeled and observed mean ET differences between dry years 682 (2004 and 2007) and wet years (2005 and 2006) for the upper basin. The observed mean ET 683 differences between dry years (2004 and 2007) and wet years (2005 and 2006) for the upper Lundquist and Loheide, Sierra ET vs. Elevation page 33 of 46 684 basin is 3 cm based on the water budget calculations (Table 2). The parameter values are the 685 same as in Fig 6. 686 687 Figure 9. Timeseries for a dry year (2004, left) and a wet year (2006, right) for (a, b) fractional 688 snow covered area, (c,d) snowmelt, (e,f) soil moisture, and (g,h) actual ET as measured by 689 MODIS and calculated by the model for a parameter set with tc=75%, K=1 m day-1, θfc =0.3, and 690 Zr=0.75. The horizontal dashed lines in (e ,f) identify the field capacity and wilting point. 691 692 Figure 10. (a) Model-simulated annual ET for individual elevation bands and (b) model- 693 simulated and observed ET for upper and lower Merced Basins for the same parameter set as in 694 Fig. 9. Years are ranked from driest to wettest, where 2007 and 2004 were dry, 2003 was 695 intermediate, and 2005 and 2006 were wet (see Table 2). Lundquist and Loheide, Sierra ET vs. Elevation page 34 of 46 696 9. Tables 697 Table 1. Parameter Values Used in Model Simulation Experiments Parameter θfc θc θwp Zr n K tc SCA- ETstart Freeze Melt at cutoff el>3000 cutoff m Values 0.15- Used 0.4 0.25 0.10 0.251.5 m 0.4 1, 0- 0.1, 0.75 0.01, 0.001 698 0.1 SCA=0 -1°C Lundquist and Loheide, Sierra ET vs. Elevation page 35 of 46 699 Table 2. Annual values from water balance (all in cm). HI indicates the upper basin, and LO 700 indicated the lower basin. Lowest row is difference between dry and wet years. HI HI Precipitation Discharge LO HI ET LO Precipitation Discharge LO ET 2003 114 69 45 118 61 57 2004 92 49 43 96 48 48 2005 142 112 30 152 98 55 2006 158 121 36 166 103 63 2007 60 32 28 67 22 45 mean 113 77 36 120 66 54 -74 -76 3 -78 -65 -12 mean(2004 & 2007)mean(2005 & 2006) 701 Lundquist and Loheide, Sierra ET vs. Elevation page 36 of 46 702 Table 3. Vegetation fractions for different elevation zones of upper and lower basins, based on 703 Yosemite National Park vegetation surveys. Elevation Conifers Broadleaf No data Meadow Barren Lakes (%) (%) (%) (%) (%) (%) 1200-1800 63.3 11.5 0.0 0.0 25.3 0.0 1800-2600 88.3 3.5 0.0 0.2 7.7 0.3 2600-4000 59.3 0.6 0.1 4.8 34.3 0.9 1200-1800 59.3 22.7 0.0 1.7 16.2 0.1 1800-2600 91.7 2.0 0.0 0.5 5.5 0.3 2600-4000 77.0 1.4 0.0 2.2 18.8 0.6 Range (m) Upper Basin Lower Basin 704 Lundquist and Loheide, Sierra ET vs. Elevation page 37 of 46 705 10. Figures 706 707 Figure 1 (a) Locations of gages (Merced River at Happy Isles and Merced River at Pohono) and 708 of meteorological stations (Tuolumne and Dana) in Yosemite National Park, California. (b) Area 709 as a function of elevation for the basin above Pohono gauge, the basin above the Happy Isles 710 gauge (Upper Basin), and the basin area below the Happy Isles gauge that drains to the Pohono 711 gauge (Lower Basin). Lundquist and Loheide, Sierra ET vs. Elevation page 38 of 46 712 713 Figure 2. Discharge normalized by basin area for the Merced River at Happy Isles (Upper 714 Basin), for the Merced River at Pohono Bridge (Combined Basins), and for the Merced River 715 contributions in between Happy Isles and Pohono (Lower Basin) for water year 2006. Note that 716 during winter rain storms (near 1 January and 1 March) more runoff per unit area originates from 717 lower elevations, because snow is falling at higher elevations. Late summer runoff originates 718 primarily from higher elevations, both from snowmelt and from high-elevation summer 719 thunderstorms. Lundquist and Loheide, Sierra ET vs. Elevation page 39 of 46 720 721 Figure 3. Annual, basin-averaged water balance components for (a) Merced River above Happy 722 Isles (Upper Basin) and (b) for the Merced River above Pohono Bridge and below Happy Isles 723 (Lower Basin) for 1992 to 2000. Evapotranspiration (ET) is estimated as the residual of annual 724 basin precipitation minus runoff. Lundquist and Loheide, Sierra ET vs. Elevation page 40 of 46 725 726 Figure 4. Scatterplot of ET in the upper and lower basins as a function of annual basin-average 727 precipitation. For the lower basin, the best fit line has a slope of 13%, and the correlation is 728 significant (p<0.01). Lundquist and Loheide, Sierra ET vs. Elevation page 41 of 46 729 730 Figure 5. Illustration of soil water accounting for determining actual ET in the model. Model 731 details are in main text. Lundquist and Loheide, Sierra ET vs. Elevation page 42 of 46 732 733 Figure 6. Model simulation performance relative to lower basin average annual ET for 2003- 734 2007, which is 54 cm, based on water balance calculations. Contour intervals are each 2.5 cm, 735 and a value of zero indicates a parameter set that precisely predicts the mean ET of the lower 736 basin. The y-axis of each plot shows the rooting depth (Zr). The x-axis shows the soil field 737 capacity (θfc). Columns indicate the transfer coefficient (tc), representing the percent of soil 738 water transferred between one elevation band and the next. Rows indicate the value of K. Other 739 model parameters were held fixed as follows: Porosity, n, is 0.4. Melt ceases when SCA drops 740 below 10%, and ET begins at elevations about 3000 m when SCA reaches 0. Plant water stress 741 begins at θc =0.25, and the wilting point occurs at θwp =0.10. Lundquist and Loheide, Sierra ET vs. Elevation page 43 of 46 742 743 Figure7. Mismatch between modeled and observed mean ET differences between dry years 744 (2004 and 2007) and wet years (2005 and 2006) for the lower basin. The observed mean ET 745 differences between dry years (2004 and 2007) and wet years (2005 and 2006) for the lower 746 basin is -12 cm based on the water budget calculations (Table 2). The parameter values are the 747 same as in Fig 6. Lundquist and Loheide, Sierra ET vs. Elevation page 44 of 46 748 749 Figure 8. Mismatch between modeled and observed mean ET differences between dry years 750 (2004 and 2007) and wet years (2005 and 2006) for the upper basin. The observed mean ET 751 differences between dry years (2004 and 2007) and wet years (2005 and 2006) for the upper 752 basin is 3 cm based on the water budget calculations (Table 2). The parameter values are the 753 same as in Fig 6. Lundquist and Loheide, Sierra ET vs. Elevation page 45 of 46 754 755 Figure 9. Timeseries for a dry year (2004, left) and a wet year (2006, right) for (a, b) fractional 756 snow covered area, (c,d) snowmelt, (e,f) soil moisture, and (g,h) actual ET as measured by 757 MODIS and calculated by the model for a parameter set with tc=75%, K=1 m day-1, θfc =0.3, and 758 Zr=0.75. The horizontal dashed lines in (e ,f) identify the field capacity and wilting point. Lundquist and Loheide, Sierra ET vs. Elevation page 46 of 46 759 760 Figure 10. (a) Model-simulated annual ET for individual elevation bands and (b) model- 761 simulated and observed ET for upper and lower Merced Basins for the same parameter set as in 762 Fig. 9. Years are ranked from driest to wettest, where 2007 and 2004 were dry, 2003 was 763 intermediate, and 2005 and 2006 were wet (see Table 2).
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