j Raptor Res. 27(3):154-160 ¸ 1993 The Raptor ResearchFoundation,Inc. PRESENCE AND DISTRIBUTION A REVIEW OF MEXICAN OWLS: PAULA ENRiQUEZ-ROCHA GEB-Mex, Ap. Postal4-077, 06400 M•xico, D.F., M•xico j. Luis RANGEL-SALAZAR Departamento de Ecologœa Terrestre,CIQRO, Ap. Postal424, 7700 Chetumal,QuintanaRoo,Mdxico DENVER W. HOLT OzolResearchInstitute, P.O. Box 8335, Missoula,MT 59807 U.S.A. ABSTR^CT.--Mexicohasa rich owl assemblage, represented by 27 species. Eighteenof thesespecies occur in theNearcticandNeotropical regions, andninespecies occuronlyin theNeotropical region.Their biology, ecologyand distribution,however,are poorlyknown. We recorded3683 specimens collectedbetween 1840 and 1991 from 11 nationaland 37 foreignmuseums,and reviewedthe literature concerningthese owls.From thesedata we presenta moreunifieddistributionof Mexican owls.Four species, Barn Owl (Tyroalba), Great Horned Owl (Bubovirginianus),FerruginousPygmy-Owl(Glaucidiumbrasilianum) and Burrowing Owl (Speotytocunicularia),have a wide distribution. The Balsas ScreechOwl (Otus seductus) is endemicto the centralPacificregion.Oaxacaand Michoacanhad the highestowl species richnesswith 21 and 19 species,respectively.Aguascalientes, Campecheand Tlaxcala had the lowest owl species richness,four, five,and fiverespectively. The FerruginousPygmy-Owlwas the mostcollected owl, representing30.7% of the specimens, and the UnspottedSaw-whetOwl (Aegolius ridgwayi)the least with 0.19%.Of the Neotropicalspecies, the UnspottedSaw-whetOwl, StripedOwl (Asioclamator),and StygianOwl (A. stygius) are considered endangered by the Mexicangovernment. The screech-owl group (Otus)and the Barred Owl (Strix varia)/FulvousOwl (S.fulvescens -- S. v. fulvescens) have uncertain distributions due to taxonomic uncertainties. Presenciay distribuci6nde los bfihosMexicanas:una revisi6n RESUMEN.--M•xico tieneunaalta riquezadeespecies debfihos,representada pot 27 especies. Dieciocho de estasespecies seencuentran tantoen la regionneirticacomoen la neotropical, y nuevede lasespecies selocalizansoloen la regi6nneotropical.Su biologla,ecologlaasl comosudistribuci6nespococonocida. Nosotroscompriamos 3683 datosdeespeclmenes de bfihoscolectados en Mgxico;conregistrosdesde1840 hasta 1991, de 11 museosnacionalesy de 37 extranjeros,adicionalmenterevisamosintensamentela literatura disponible.Combinandoestosdatos,damosuna distribuci6nunificadade los bfihosde Mgxico. Cuatroespecies, la Lechuzade Campanario(Tytoalba),el BfihoCornadoAmericano(Bubovirginianus), el TecolotitoBajefio(Glaucidium brasilianum) y el TecoloteZanc6n(Speotyto cunicularia)tienen una amplia distribuci6nen el continenteAmericano.E1 TecoloteOjioscurodel Balsas(Otusseductus) es endgmico parala regi6ndelPacificoCentraldeMgxico.Losestados demayorriquezadeespecies fueron Oaxacay Michoacincon21 y 19 especies respectivamente, porel contrariolosestados conmenorriqueza fueronAguascalientes, Campechey Tlaxcalacon4, 5 y 5 respectivamente. El TecolotitoBajefioha sido la especie de bfihomilscolectada, representando el 30.7%delosespeclmenes, y el queregistroconmenos colectas fugel TecoloteAbeteroSurerio(Aegolius ridgwayi)representando tansoloel 0.19%.De lasespecies neotropicales, el TecoloteAbeteroSurerio,el BfihoCornadoCariblanco(Asioclamator)y el BfihoCornado Oscuro(A. stygius)sonconsideradas comoamenazadas por el gobiernoMexicano.E1 grupoOtus asi comoel BfihoSerranoVientrirrayado(Strixvaria)/BfihoSerranoSurerio(S.fulvescens = S. v.fulvescens) tienen distribuciones controversiales debidas a su clasificaci6n incierta. [Traducci6n Autores] Currently, there are 178 speciesof owls recognized in the world (Sibley and Alhquist 1990), of which 27 (15%) speciesoccurin Mexico. This exceedsthe owl speciesdiversity(N = 22) of northern 154 forests,northof 35ø(Norberg1987). Although,there is someoverlap. Owls in Mexico have been little studied and most literature is old (Ridgway 1895, Kelso and Kelso SEPTEMBER 1993 MEXICAN OWL DISTRIBUTION 155 Sonora Chihuahua Coahuila aa Californi, Si Sur Durango dipas 1. Aguascalientes 2. Oistrito Federal Gulf of •,,N• 3. Guanajuato 4.Hidalgo Mexico • 5. Morelos 6. Queretaro IRa :ruz Colinn 7. Tlaxcala Guerrero Pacific Figure 1. Ocean Oaxaca Chiapas Map of the Mexican Republic showinglocationsof 32 states. 1936, Moore 1937a, 1937b, 1941, 1947a, Moore choacfin, EscuelaNacionalde CienciasBio16gicas, Insti- and Peters 1939, Alvarez del Toro 1949, Briggs 1954, Moore and Marshall 1959, Buchanan 1964, Northern 1965). Generally,mostMexican owl specieshavelimitedmorphological descriptions and only approximateddistributions(Friedman et al. 1950, tuto de Historia Natural de Chiapas, Instituto Nacional de InvestigacionesForestalesy Agropecuarias-Bacalar, Instituto Nacional de Investigaciones sobreRecursosB•6ticos,LaboratorioNatural de Las Joyas, Universidadde Guadalajara, Museo de Zoologlade la Facultad de Ciencias, Museo de ZoologlaIztacala, Sa16nde las Aves de Blake 1972, Davis 1972, Peterson and Chalif 1973, 1989, Edwards1989). Unfortunately,thesedistributionsare often controversial.Here, we presenta review of the distribution and relative status of Mex- ican owls based on museum specimen data and available literature. STUDY AREA AND METHODS Mexico (2 million km2)has32 states(Fig. 1) with > 50% of the land mass above 1000 m elevation (Ramos 1985). Its 32 vegetationtypes (Rzedowski 1986), geographiclocation, and topography contribute to a rich diversity of fauna and flora. To obtain information on Mexican owl specimenswe wrote letters to or visited 11 national museums(Colecci6n Ornitol6gicadel Instituto de Biologla,Colecci6nOrnito16gicade San Nicol•tsde Hidalgo, Universidadde Mi- Saltillo Coahuila,UniversidadAut6nomade Baja California Sur and 37 foreignmuseums(Australian Museum, AnnistonMuseumof Natural History, Academyof Natural Sciencesof Philadelphia,Bell Museum of Natural History, British Museum, Carnegie Museum of Natural History, Cornell University Collection,CollectionZoologyMuseum Amsterdam-Nederland, Denver Museum of Natural History, Delaware Museumof Natural History, Fort Hays Museum,FloridaMuseumof Natural History, Forschungsinstitut undNaturmuseumSenckenberg, Harvard Museumof ComparativeZoology,Illinois StateMuseumCollection,KansasUniversity Collection,Los AngelesCaliforniaMuseum-Natural History, LouisianaState UniversityMuseum Zoology,Moore Laboratoryof Zoology-OccidentalCollege, Museo Nacional de Ciencias Naturales-Espafia,Museum of Natural History-Chicago, Museum of Vertebrate Zoology-Universityof Cali- forniaBerkeley,OklahomaMuseumof Natural History, ProvincialMuseum of Alberta, PeabodyMuseum Col- 156 ENRiQUEZ-ROGHAET AL. VOL. 27, NO. 3 lection-YaleUniversity,Royal Ontario Museum, Rijks- museum VanNaturvlijke Historie-Leiden, Santa Barbara Museum ofNatural History, Southwestern College Col- Sr^TESø• 0 lection,San Diego Museum of Natural History, Staatli- chesMuseumftir Naturkunde in Stuttgart, TexasCooperativeWildlife Collection,National Museum of Natural History-SmithsonianInstitute,Universityof Washington, Thomas Burke Memorial State Museum, Universityof Wisconsin Zoological Museum,WesternFoundation Vertebrate Zoology Collection,ZoologicalInstitute of the Academyof Sciences-URSS).Additionally, we reviewed the existingliteraturefor informationconcerning Mexican owls.For vegetationtypes,we followedRzedowski(1986). State boundarieswere the unit usedto delineatedistributionsof the owls. We applied the taxonomyusedby AmadonandBull (1988) for mostof the owl species which occurin Mexico. We followed Marshall et al. (1991) for recentchangesof VermiculatedScreech-Owl(Otusguatemalae)to Variable Screech-Owl(O. atricapillus),and A.O.U. (1991) for genericchangeof the BurrowingOwl from Atheneto Speotyto. We includedthe Fulvous Owl (Strixfulvescens) asa subspecies of theBarredOwl (Strix varia) = S. v.fulvescens (Edwards1989, J. Marshall pers. comm.). RESULTS O0 0 Bcs 0 0 0 O0 0 c.,. c.•s co^ cot OF O0 0 0 O0 0 0 0 O0 0 O0 O0 0 O0 0 XO 0 XO O0 X O0 O0 0 0 O0 O0 O0 0 O0 O0 TAB 0 0 0 O0 0 0 0 •^• 0 O0 TLAX O0 0 0 0 O0 0 X O0 000 0 0 0000 O0 0 O0 OX OXO0 0 0000 0 O0 0 0 000 0 O0 0 OOOOXO 0 O0 XO XO 0 O0 XO O0 0 OOX XO0 0 O0 000 00000 O0 O0 O0 XOOO0 0 O0 000 0 0 O0 X O0 000 000 O0 O0 0 XOO0 0 0 0 OOX XO 0 O0 000 0 XOO0 OXO 0000000 OX O0 O0 OXXOOO0 0 O0 O0 0 X O0 0 0 0 OOOX O0 0 0 OX son 0 O0 0 0 0 o•oo 0O0 0 0 O0 000 O0 0 O0 O0 0 O0 o•x O0 O0 0 000 0 •oR O0 XO 0000000000 O0 O0 XO 0 X O0 0 0 0 OOXO X 0000 0 0 0 X X 0 0000 O0 O0 X O0 O0 0000 O0 0 0 000XO 0 X 0 0 0 0 0 O0 0 0000000 0 O0 OX OOXX We compiledand analyzeddata on 3683 speci0 0 O0 0 0 X X mensof Mexicanowls.Twenty-sevenMexicanowl Yuc 0 0 0 0 000 species fromboththe Nearcticand Neotropicalzoogeographic regionsare represented. Eighteenspecies Figure 2. Presenceof owl speciesby state in Mexico. occurin the Nearcticand Neotropicalregions,while nine speciesoccuronly in the Neotropicalregion. The BalsasScreech-Owl (Otus seductus)is endemic to Mexico's Pacific slope region. The Bearded Screech-Owl(Otus barbarus)and UnspottedSawwhet Owl (Aegoliusridgwayi) are found only in Chiapas. The greatestnumberof museumspecimens were of the FerruginousPygmy-Owl (Glaucidiumbrasihanum) and least number of the UnspottedSawwhet Owl. The richestowl assemblage camefrom Oaxaca and Michoacan with 21 and 19 species, respectively. SpeciesAccounts Barn Owl (TALB) Tyroalba The most widely distributedowl in Mexico, it inhabitatsforest,openareas,villagesand cities,and islands from sea level to 3500 m in elevation. One Codesare describedin the text. (O = museumrecordsand X = literature records.) sula. Blake (1972) and Grossmanand Hamlet (1988) consideredthem widely distributed throughout Mexico. Flammulated Owl (OFLA) Otusfiamrneolus Flammulated Owl inhabits highlandsand pine forestfrom 2100-3000 m elevation.It is migratory in southern Sinaloa, central Mexico and south throughthe highlandsof Chiapas.Eighty-onespecimens from 16 stateswere recorded(Fig. 2). Additionally,it has beenreportedfor Distrito Federal (Ridõway 1914, Friedmanet al. 1950, Blake 1972) and Nuevo Le6n (A.O.U. 1983, Contreras-Balderas 1992). Eastern Screech-Owl (OASI) Otus asio Eastern Screech-Owlinhabits pine forest, pineoak forest and riparian habitats from sea level to 1500 m elevation.One hundredand sevenspecimens from 19 stateswere recorded(Fig. 2). Its distribution hundred forty-one specimensfrom 31 stateswere recorded(Fig. 2). Friedman et al. (1950) also reportedBarn Owls fromHidalgoandQuintanaRoo, as have we (Enrlquez-Rochaand Rangel-Salazar is restricted to northeastern and southern Mexico, pers. ohs.). Paynter (1955) and Hartiõ (1979) did which agreeswith Marshall (1967) and Rowley not report the Barn Owl from the Yucatan Penin- (1984). SEPTEMBER1993 MEXICAN OWL DISTRIBUTION Western Screech-Owl (OKEN) Otuskennicottii Western Screech-Owl inhabits dense pine and 157 elevation.One hundredand six specimensfrom 16 stateswererecorded(Fig. 2). Althoughreportedfrom pine-oakforests,openhabitats,and semi-desert veg- Campeche and Tabasco (Friedman et al. 1950, etation from 0-2400 m elevation. One hundred and Paynter 1955, Storer 1961, Davis 1972), no speciforty-eight specimensfrom 15 stateswere recorded menswere located.This is the only speciesfrom the (Fig. 2). There is confusionregardingspeciesand Otusgenusfound in the Yucatan Peninsula. subspecies distributionsdue to taxonomicchanges (Marshall 1967). Our data conflictwith Blake (1972) CrestedOwl (LCRI) Lophostrixcristata Crested Owl occurs in southern Mexico and inwho reportedthis species in Queretaroand Hidalgo. Its distribution is northcentral, southwestern, and habitstropicalevergreenand semi-deciduous forest, Baja California. riparian areas, partial clearings, and foothills. It rangesfrom sealevelto 1000 m elevation.TwentyBalsas Screech-Owl (OSED) Otus seductus two specimens from four stateswere recorded(Fig. BalsasScreech-Owlinhabits tropical deciduous 2). Grossmanand Hamlet (1988) reportedit from forestand arid tropicalscrub,and the edgesof cropthe Yucatan Peninsula, but we could not confirm land. It rangesfrom sea level to 1200 m elevation. this. Twenty-five specimensfrom two stateswere recorded(Fig. 2). It hasalsobeenrecordedin Guerrero Spectacled Owl (PPER) Pulsatrixperspicillata (Marshall 1967, Peterson and Chalif 1975, A.O.U. SpectacledOwl inhabitstropical evergreenand 1983). This, the only endemicMexican owl, is cur- semi-deciduous forests,riparian areas,partial clearrently found only in Michoacan and Colima. ings, and forest edges.It rangesfrom sea level to 900 m. Thirty-one specimens from three stateswere PacificScreech-Owl(OCOO) Otuscooperi recorded(Fig. 2). It appearsrestrictedto southern Pacific Screech-Owl inhabits tropical lowlands, Mexico, but Grossmanand Hamlet (1988) report mangroves,open forests,palm groves,and swamps. it for the Yucatan Peninsula. We are uncertain of It occursfrom sealevel to 90 m elevation.Twenty- this. three specimens from six stateswere recorded(Fig. 2). It appearsto exist only along the Pacificslope Great Horned Owl (BVIR) Bubovirginianus in southwestern Mexico. Great Horned Owl inhabitsa wide diversityof habitatsfrom forestland to open areas.It ranges WhiskeredScreech-Owl(OTRI) Otustrichopsis from sea level to 2700 m elevation. Two hundred Whiskered Screech-Owl inhabits highlands, in thirty-six specimensfrom 26 stateswere recorded pine-oakforest,pine forestand dry oak. It ranges (Fig. 2). Brodkorb(1943) reportedit for Tabasco, from 1200-5000 m elevation. One hundred and but no specimens exist.The presentdistributionenninety-threespecimens from 20 stateswere recorded compasses almostthe entirecountryand its presence (Fig. 2). Additionally, it has been reported from in Tabascoseemslikely. Hidalgo and Puebla (Davis 1972, A.O.U. 1985). It has a wide distribution but is absent in the Yucatan Mottled Owl (CVIR) Ciccabavirgata and Baja California peninsulas. MottledOwl inhabitstropicallowlandforest,open tropicalforest,and foothills.It rangesfrom sealevel BeardedScreech-Owl (OBAR) Otusbarbarus to 2100 m elevation.Four hundred and one speciBeardedScreech-Owlinhabitspine and pine-oak mensfrom 22 stateswere recorded(Fig. 2). It has forestfrom 1400-1800 m elevation.Only nine speca widedistribution,butis notyetreportedfromBaja imens,all from Chiapas,wererecorded(Fig. 2), but Peninsula and northcentral Mexico. Ridgway (1914) reportedit southof Chiapasin the Black and White Owl (CNIG) highlandsof Guatemala. It has the most restricted distributionof any Mexican owl. Ciccabanigrolineata Black and White Owl appearsto have discontinVermiculated Screech-Owl (OGUA) uousdistributionwithin Mexico. It inhabitstropical Otusguatemalae evergreenand semi-deciduous forests,forestedge,or VermiculatedScreech-Owlinhabitsriparian ar- partial clearings.It rangesfrom sealevelto 1200 m eas, tropical semi-deciduousand deciduousforests, elevation.Forty-six specimensfrom five stateswere and forestedges.It rangesfrom sealevelto 1500 m recorded(Fig. 2). Our museumdata agreewith the 158 ENRiQUEZ-ROCHA ET AL. VOL. 27, No. 3 literature (Ridgway 1914, Friedman et al. 1950, not been reported from Mexico's peninsulasor the Blake 1972, Davis 1972, Petersonand Chalif 1973, northcentraland central plateaus. A.O.U. 1983, Edwards 1989). Paynter (1955) reFerruginousPygmy-Owl (GBRA) portedone specimenfrom Quintana Roo, and we Glaucidium brasilianum have additional specimensfrom there. FerruginousPygmy-Owl inhabitslowlandriparBarred Owl (SVAR) Strix varia ian forest,forest edge, secondgrowth, and thickets Found in pine and pine-oak forests,and ranges from sea level to 1500 m elevation. One thousand from 1500-3000 m elevation.Thirty-nine specimens one hundred thirty-one specimensfrom 23 states from 10 stateswere recorded(Fig. 2). Someauthors were recorded(Fig. 2). It has not beenreportedin (Ridgway 1914, Peters1940, Friedmanet al. 1950, the Baja California Peninsula. Friedman et al. Davis 1972, A.O.U. 1983) reportedit for Puebla, (1950), and A.O.U. (1983) report it for Distrito but we locatedno museumspecimens.Alvarez del Federal and Coahuila but no museum specimens Toro (1980) reportedthe FulvousOwl (Strixful- exist. vescens)from the Cloud Forest of Chiapas, but we Elf Owl (MWHI) Micrathenewhitney• identifiedonlyonemuseumspecimen fromthe Cloud Elf Owl inhabitssaguarodesertsandarid tropical ForestasStrixvariafulvescens. Six specimens in three scrub from sea level to 1800 m elevation. One hunmuseums identified as Strix varia were reclassified dred sixty-sevenspecimensfrom 16 stateswere re- as S. v.fulvescens by J.T. Marshall (pets.comm.). corded(Fig. 2). This includesthe RevillagigedoIsAlso,two otherspecimens from Oaxacawere iden- landsin the Pacific Ocean. It has alsobeenreported tified asS.fulvescens, but they shouldbe reclassified for the state of Mexico (Peters 1940, Blake 1972), as S. v. fulvescens. and Tamaulipas(Grossmanand Hamlet 1988), but SpottedOwl (SOCC) Strix occidentalis no specimensexist. It doesnot occurin southern SpottedOwl hasa discontinuous distribution,in- Mexico. habiting pine and pine-oakforestsfrom 1200-2500 Burrowing Owl (SCUN) Speotyto cunicularia m elevation.Twenty-sevenspecimensfrom eight Burrowing Owl inhabitsopenlands,grasslands, stateswere recorded(Fig. 2). It has been reported prairies, and tropical scrubfrom sealevel to 1800 in Baja California (Ridgway 1914, A.O.U. 1983, m elevation.Two hundredseventy-seven specimens Edwards 1989, Peterson and Chalif 1989), but no from 27 stateswere recorded(Fig. 2). It has a wide specimensexist from there. A specimenfrom San distribution and occursthroughout the country in Luis Potosiand one sightrecordfrom Durango do suitable habitat (Grossman and Hamlet 1988). exist (R. Clark pets. comm.),but we were not able to confirm this. Northern Pygmy-Owl(GGNO) Glaucidiumgnoma Northern Saw-whet Owl (AACA) Aegoliusacadicus Northern Saw-whet Owl inhabits foothills with pine and pine-oakforestsfrom 1800-2800 m elevation.Twenty-six specimens from 11 stateswere recorded(Fig. 2). It occursin northcentraland central Mexico, and hasbeenreportedfrom Chihuahua, Jalisco,and Veracruz(Ridgway1914, Friedmanet NorthernPygmy-Owlinhabitspine-oakforestand scrubvegetation,from 1800-3700 m elevation.One hundredeighty-eightspecimens from 25 stateswere recorded(Fig. 2). It has also been reportedfrom Colima (Ridgway 1914, Friedmanet al. 1950), but al. 1950, Blake 1972, Davis 1972). no museum specimensare known. UnspottedSaw-whet Owl (ARID) Aegoliusridgwayz Glaucidium minutissimum UnspottedSaw-whet Owl inhabitscloudforest, LeastPygmy-Owlinhabitstropicalforest,forest pine, and pine-oak forest from 2100-3000 m eleall from Chiapaswere edge,plantations,and barrancasfrom sea level to vation.Only sevenspecimens, 1800 m elevation.One hundred thirty-nine speci- recorded(Fig. 2). It has been reportedpreviously mens from 14 stateswere recorded(Fig. 2). The from Chiapas(Moore 1947b,AlvarezdelToro 1980, specimendata agreewith the distributionsgivenby Davis 1972, Petersonand Chalif 1973, A.O.U. 1983, Davis (1972) and Petersonand Chalif (1989). It has Enriquez and Rangel-Salazar (pers. obs.). The LeastPygmy-Owl(GMIN) SEPTEMBER 1993 MEXICAN OWL DISTRIBUTION 159 A.O.U. (1983) reported a questionablespecimen Owl (Edwards 1989, J. Marshall pers. comm.); record from Oaxaca. however,field work is still neededto verify this. The UnspottedSaw-whetOwl, StripedOwl, and Striped Owl (ACLA) Asiaclarnatar StygianOwl are listedas endangeredby the MexStripedOwl inhabitsdensetropicalevergreenforican government (SEDUE 1991). The Balsas est,forestgaps,and forestedgefrom sealevelto 900 Screech-Owl, Bearded Screech-Owl, Crested Owl, m elevation.Twenty-six specimensfrom 3 states SpectacledOwl, Black and White Owl, Barred Owl were recorded(Fig. 2). It occursprimarily in southand SpottedOwl are being consideredfor endaneasternMexico,but alsoin Veracruz,Chiapas(Blake geredstatusbecauseof their low numbers,restricted 1972, Davis 1972, Petersonand Chalif 1973), and distribution, and habitat loss. Oaxaca (Binford 1989). Museum collectiondata can help delineategeoStygianOwl (ASTY) Asiastygius graphic distribution and estimate relative abunStygian Owl inhabits densecloud forest in the dance.Lack of museumcollectionsmay indicatea mountainsand rangesfrom 1500-3000 m elevation. rare, uncommon,or easily overlookedspecies.MuSeventeenspecimensfrom nine stateswere recorded seumdata canbe importantalsofor comparingpres(Fig. 2). It hasalsobeenreportedin Chiapas(Fried- ent and historical distributions. Threatened and enman et al. 1950, Davis 1972, Alvarez del Toro 1980, dangered species datain collections arealsoimportant for historic location sites. On the other hand, for A.O.U. 1983) and the Yucatan Peninsula (A.O.U. 1983, Edwards 1989), but, this doesnot agreewith somemuseumspecimens the accompanyingdata are our dataor that of Paynter(1955) andHartig (1979). incompleteor completelywrong. Often date of collection and other important information were not Long-earedOwl (AOTU) Asiaatus included. Long-earedOwl inhabitspineandpine-oakforest This is the first attempt in Mexico to draw inand riparian forestfrom sealevel to 1800 m elevaformationtogetheraboutthis taxonomicgroup.We tion. Forty-one specimensfrom 13 stateswere rehope this paper acts as a catalystto spur further corded(Fig. 2). Museum data and the literature are investigationsinto Mexican owl distribution, status, in closeagreement(Ridgway 1914, Friedmanet al. biology,ecology,and conservation. 1950, Blake 1972, A.O.U. 1983). There are band ACKNOWLEDGMENTS recoveriesof long distancemigrantsto Mexico from Saskatchewan,Canada (Houston 1966), Montana, We are very grateful to the museums that provided U.S.A. (D. Holt pers.obs.),and Minnesota, U.S.A. information. We thank J.T. Marshall for commentson the manuscriptand additionalinformation.We thank R.J. (D. Evans pets. comm.). Clark, C.D. Marti, H. Mikkola andan anonymous referee Short-earedOwl (AFLA) Asiafiamrneus for commentson the manuscript. Short-earedOwl inhabitsgrasslands, open trop- LITERATURE CITED ical forests,prairies, and marshesfrom sea level to ALVAREZDEE TORO, M. 1949. Striped horned owl in 1500 m elevation.Twenty-nine specimensfrom 14 southern Mexico. Condor 51:232. stateswere recorded(Fig. 2). It has also been re1980. Las Aves de Chiapas. Gob. del Edo. de ported from Jalisco, Oaxaca, Puebla, Veracruz, Chiapas.Tuxtla Gutierrez, Chiapas,M6xico. Quintana Roo and Yucatan (Friedman et al. 1950, AMADON, D. AND J. BULL. 1988. Hawks and owls of the world: a distributional and taxonomic list. Proc. Hartig 1979, A.O.U. 1983, Binford 1989, MacWest. Found. Vertebr. Zool. 3:295-357. Kinnon 1992). AMERICAN ORNITHOLOGISTS' UNION. DISCUSSION The taxonomyand distributionof mostMexican owls is still incomplete.For example,subspecies of the screechowls (Otus)groupdiscussed by Marshall (1967) werereportedonly fromeasternMexico. Our museum data, however, indicate that someof these subspecies occur in western Mexico. The Fulvous Owl has beenconsidereda subspecies of the Barred 1983. Check-list of North AmericanBirds.6th. Ed. Am. Ornithol. Union, Washington, DC U.S.A. 1991. Thirty-eight supplementto the American Ornithologists'Union Check-list of North American Birds. Auk 108:750-754. BINFORD, L.C. 1989. A distributionalsurveyof the birds of the Mexican stateof Oaxaca. Ornithol.Monogr.43. Am. Ornithol. Union, Washington, DC U.S.A. BLAKE,E.R. 1972. Birdsof Mexico. Univ. ChicagoPress, Chicago, IL U.S.A. 160 ENRiQUEZ-ROCHAET AL. BRIGGS,M. 1954. Apparent neotenyin saw-whet owls of Mexico and Central America. --AND BRODKORB,P. 1943. Birds from the Gulf lowlands of southern, Mexico. Misc. Publ. Mus. Zool. Univ. Mich. America. and conservation 1964. The Mexican races of the least pygmyowl. Condor66:103-112. CONTRERAS-BALDERAS, A. 1992. Second record of the Flammulated Owl in Nuevo Leon, Mexico. Wilson Bull. 104:375. DAVIS,L.I. 1972. A field guideto the birds of Mexico EDWARDS, E.P. 1989. A field guideto the birdsof Mexico. Ernest P. Edwards. Sweet Briar, VA U.S.A. FRIEDMAN, H., L. GRISGOM AND R.T. MOORE. 1950. Distributionalcheck-listof the birdsof Mexico. Cooper Ornithol. Club, Berkeley,GA U.S.A. GROSSMAN, M. AND.J. HAMLET. 1988. Birds of prey of the world. Bonanza Books, New York, NY U.S.A. HARTIG, H.M. 1979. Las Avesde Yucat•tn. Fondo Edit. Yucat•tn, M6xico. HOUSTON,C. 1966. SaskatchewanLong-earedOwl recoveredin Mexico. Blue Jay 24:178. KELSO L. 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