11/3/2010 Geography 1001: Climate & Vegetation Limiting Factors & Microenvironment Dr. Barnard Physiognomy Life Zones & Plant Life Forms 1 11/3/2010 Raunkiaer life forms (based on perennating bud) Biotic Ecosystem Operations (Function) • Producers, Consumers, and Decomposers – autotrophs and heterotrophs • Examples of Food Chains and (Complex) Food Webs • Efficiency in a Food Web Equilibrium, Stability & Diversity • Stable conditions – hard to change its conditions – When changed, may return back to original stable conditions • Unstable conditions – Easy to change its conditions – When changed, may never return back to original stable conditions Equilibrium, Stability & Diversity • Stability – Inertia stability and resilience – Disturbances • A discrete & stochastic event that disrupt (any) part(s) or (any) component(s) of a biotic community. • Chronic disturbances – A large changes in ecosystem processes in response to a continuous & directional variation in the environment. • Neutral intermediate 2 11/3/2010 Stochastic & Chronic Disturbances Number of species Low---------------------------High Species-Area Curves Small------------------------------------------------Large Size of area Number of species Low---------------------------High Number of species Low---------------------------High Latitudinal gradients 0-------------------------------------------------------90 Latitude Intermediate disturbance hypothesis Low---------------------------------------------------High Disturbance 3 11/3/2010 Ecological Succession What is ecological succession? • Ecological Succession – Primary and Secondary succession – Ecological Succession • Terrestrial succession • Aquatic Succession Photo: D. Kulakowski Photo: A. Holz Primary Succession Why does ecological succession occur? Photo: A. Holz Figure 19.25 4 11/3/2010 Mount St. Helens Mount St. Helens Mount St. Helens Mount St. Helens 5 11/3/2010 Secondary succession Mount St. Helens Photo: A. Holz Figure 19.27 Ecological Succession Aquatic Succession – Forest development (4 stages) What about Beavers? • • • • Stand initiation Stem exclusion Understory re-initiation Old growth – Gap phase dynamics – Continuous regeneration mode – Early vs. Late stages of succession characteristics Figure 19.30 6 11/3/2010 Forest development of whole stands Stand initiation Photo: A. Holz Stem exclusion Photo: D. Kulakowski Understory reinitiation Photo: D. Kulakowski 7 11/3/2010 Old growth Forest development & regeneration modes at fine-scale • Gap phase dynamics – Tree regeneration is dependent on canopy openings created by the death of the former canopy occupants (e.g. canopy opening diameters of c. 4 m to 40 m). E.g. New England forest, Tropical Rain Forests, etc. Photo: A. Holz Forest development & regeneration modes at fine-scale • Continuous regeneration mode (more rare) – Shade tolerant species can complete their life cycle beneath a relatively closed forest canopy. E.g. Tropical Rain Forests, some Aspen stands. Early vs. Late stages of succession • Attributes • Characteristics Photo: A. Holz Photo: A. Holz 8 11/3/2010 Early Stages of Succession Late Stages of Succession Plant Biomass Small Large Plant Longevity Short Long Well dispersed Poorly dispersed Low High Litter and Soil Living Biomass and Litter Role of Decomposers in Cycling Nutrients to Plants Minor Great Rate of Net Primary Productivity High Low Plant Species Diversity Low High Attribute Seed Dispersal Characteristics of Dominant Plants Photosynthetic Efficiency of Dominant Plants at Low Light Site of Nutrient Storage Take home messages • Ecosystems are dynamic and change is to be expected Does ecological succession ever stop? Terrestrial Biomes • Biogeographic Realms • Earth’s Major Terrestrial Biomes • Long intervals between disturbances results in a broad range of natural conditions • There is no single snapshot that can describe how a healthy ecosystem should look 9 11/3/2010 Biogeographic Realms Biogeographic Realms • Transition Zones • Terrestrial Ecosystems Figure 20.1 Earth’s Major Terrestrial Biomes • • • • • • • • • • • Equatorial and Tropical Rain Forest Deforestation of the Tropics Tropical Seasonal Forest and Scrub Tropical Savanna Midlatitude Broadleaf and Mixed Forest Needleleaf Forest and Montane Forest Temperate Rain Forest Mediterranean Shrubland Midlatitude Grasslands Deserts Arctic and Alpine Tundra Major Terrestrial Biomes Figure 20.4 10 11/3/2010 Rainforest Canopy Rainforest Canopy Figure 20.5 Amazonian Rain Forest Figure 20.5 Buttressed Trees and Lianas Figure 20.6 Figure 20.6 11 11/3/2010 Deforestation of the Tropics Figure 20.7 12
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