1 1. Abstract Vegetation maps play an important role in understanding vegetation dynamics at Mono Lake, California. These maps act as a basis for studying succession, helping establish correlation between vegetation and other environmental influences, and providing important indications of how plants respond to disturbances. Current vegetation maps for Mono Lake do not measure up to the standards of vegetation maps that are accurate, easily understood and of high quality. Employing Rabbitbrush (Chrysothamnus nauseous) and Big Sagebrush (Atremisia tridentata) as study species, ground-truthing methods were utilized to take flora samples at Mono Lake, California. Results showed that along a 500 meter transect, moving away from the lakeshore, both the percent cover and number of individuals of Rabbitbrush decreased where as Big Sagebrush increased. Topographic data was also gathered along the transect, and found to be increasing, with the exception of a few level data points that represented old beach ridges. Finally, the study examined how current Geographic Information Systems (GIS) techniques aid in the making of effective vegetation maps and how they could be applied at Mono Lake. 2 2. Introduction Mono Lake is an interesting area to do research, due to its unique geology, wildlife, and political history. Mono Lake is a terminal saline lake, located in the Great Basin and Range Province of the Western United States (Figure 1). Figure 1. The boxed in area represents Mono Lake’s location in the Great Basin (Hart, 1996) The area’s geologic past includes volcanism and fault activity, which is still present today, and glaciation. Mono’s wildlife is very distinctive; the lake is home to the alkali fly and brine shrimp, which are found only a few places in the world. Politically, its past is full of controversy, which revolves around the taking of Mono Lake’s freshwater tributaries by the city of Los Angeles. Mono Lake is an appropriate place to conduct a study concerning GIS and vegetation mapping because since the early 1900’s, the state of California has been taking water from the Mono Basin. The city of Los Angeles Department of Water and Power began diverting Mono Lake’s tributaries in 1941. As a result, the volume of the lake was halved and the salinity doubled. In 1962, the lake had dropped 25 vertical feet and in 3 1995 had dropped 40. However, the perseverance of dedicated people and the help of the courts, enabled Mono Lake to be regain many of its tributaries. In 1993, it was ordered that the lake level eventually be restored to 6, 390 feet or higher (Hart, 1996, www.monolake.org.). Developing a vegetation database and using GIS would be beneficial as well as time saving to the mapping of Mono Lake plant communities. With the lake levels increasing on a yearly basis, it would easy to update maps and make changes that correspond to those taking place in the environment. Using GIS techniques for mapping would also be an efficient way to keep track of the fluctuating lake levels. The objectives of the study conducted at Mono Lake relating to vegetation mapping and GIS are: 1. Conduct vegetation samples in order for the researchers to learn the types of groundtruthing that has to accompany map making, and get a feel for the amount of time it would require to do a vegetation survey. 2. Provide background information on current vegetation present and Mono Lake, GIS, and how vegetation mapping and GIS techniques have been integrated. 3. Discuss the results and draw conclusions. 3.0 Background Information 3.1. The Vegetation of Mono Lake There are at least seven types of vegetation previously described in studies done at Mono Lake, which current maps reflect (Figure2). These vegetation types have been described from personal observations made by Burch et al (1977) and by Vorster (1985). Figure 2(next page) Current vegetation map of Mono Lake(Mono Lake Committee, 1999) 4 However, Constantine (1993) also categorized the vegetation into six categories (Figure 3). The major difference in the categorization of these studies is that Burch (1977) and Vorster (1985) considered transition zones, areas where major vegetative communities overlap and Constantine (1993) did not. Figure 3. A Current Plant Distribution of Mono Lake (Constantine, 1993) The major types of plant communities present at Mono Lake are as follows as described by Constantine (1993) are as follows: (See also Figure 2 attached as the previous page) 3.1.1) Alkaline Sink Scrub/ Alkaline Herb - This community is characterized by alkaline crusted soils and low moisture availability. The vegetation is primarily herbaceous and halophytic (salt-loving) in nature. The dominant species are Rabbitbrush (Chrysothamnus nauseosus), Greasewood (Sarcobatus vermiculatu), and Saltgrass (Distichlis spicata). 5 3.1.2) Freshwater Marsh/ Wet Marsh – This type of community is characterized by the water table at the surface originating from non lake sources, a presence of standing water, and nutrient rich soil that is oxygen deprived. The dominant species are Monkey Flower (Mimulus guttatu), Common Cattail (Typha latifolia), Foxtail Barley (Hordeum jubatum) and other grasses and sedges. 3.1.3) Sagebrush Scrub/Dry Scrub - This community is the most common in the Mono Basin. It is characterized by a sparse herbaceous layer and a shrub cover of > 5%. Due to the volcanic activity in the area, the soils are usually deep, well drained, and nonalkaline. Plants in this community must be able to tolerate a variety of extreme conditions and most are equipped with large deep taproots. The dominant species are Big Sagebrush (Artemisia tridentata), Bitterbrush (Purshia tridentata), Giant Blazing Star (Mentzelia laevicaulis), and a few Desert Peach (Prunus andersonii). 3.1.4) Pinyon-Juniper Woodland - Juniper woodland typically occurs at higher altitudes, where the climate is colder and wetter. Woodlands are classified by being open forests where trees and intermixed with shrubs and grasses. The presence of Pinyon and Juniper is dependent on the moisture availability, altitude, and soil depth. The dominant species in this community are Pinyon Pine (Pinus monophylla), Utah Juniper (Juniperus osteosperma), Mormon Tea (Ephedra virdis), and Cut-leaf Mahogany (Cercocarpus ledifolius). 3.1.5) Jeffrey Pine Forest - This plant community does not occur directly near the lake, but in the surrounding mountains. The largest stand of Jeffrey pine in the world occurs near the southeast end of the lake. The dominant species are Jeffrey Pine (Pinus jeffreyi), Lupin (Lupinus spp), and Prickly Phlox (Leptodactylon pungens). 3.1.6) Riparian Forest - These forests occur along the streams that run down the mountains and into Mono Lake. The soils are layers of fine and coarse material that has been deposited by varying stream levels. This community is characterized mainly by 6 deciduous species. Plants in this community must be able to withstand periodic flooding. These communities also provide great habitat for wildlife. The dominant species are Quaking Aspen (Populus tremuloides), Black Cottonwood (Populus trichocarpa), and Coyote Willow (Salix exigua). 3.2 The Importance and Use of Vegetation Maps Vegetation is a fundamental aspect of the environment, animals use it as habitat, it is a renewable resource that provides food and fuel for humans, and it is very influential in affecting other earth processes such as the carbon and nitrogen cycles. Vegetation maps are important because they can model the current vegetation of an area and can be used to make predictions about future vegetation changes. Vegetation maps provide in part for the basis of understanding the world around us and for solving complex problems, such as management. Therefore, it is important to be able to map vegetation thoroughly. However, it is important to remember that any vegetation classification whether it be in map form or not, is a simplification of a complex reality of gradients and mosaics in vegetation (Whittaker, 1970). Many vegetation maps of the past and present have traditionally focused on what the map communicates rather than analysis. Demers (1991) explains that the purpose of most maps is to produce a visual thematic pattern that corresponds to the vegetation classification employed. There is not much quantitative map information available except for measurements taken manually because the classification process often combines numerous vegetative attributes in a given area (Demers, 1991). In the past, vegetation maps were constructed based on observations of mappers and scientists who surveyed the vegetation using a variety of field methods. Consequently, the maps were not as accurate, were fairly subjective, and it took a long time to make them. For example, in Mammoth Cave National Park, scientists spent three years 7 surveying all the vegetation (University of Idaho, 1999). However it is important to note that no single method of mapping vegetation is best because the methods used are largely determined by the purposes for which the map will be used (Kiichler, 1988). Many maps have been made using time consuming visual interpretation of aerial photographs, planimeters, and field-collected data. As time has progressed, vegetative cover has been determined by spectral data from satellites, which can produce maps much faster but far less detailed. Most manual mapping with satellite data is done with film products that are not optimally suited for mapping heterogeneous natural vegetation mainly because the film has already been radiometerically and geometrically rectified (University of Idaho, 1999). Problems that occur can be terrain shadowing and poor spectral separation of cover types, which requires additional data and high quality products. In the long, this run means higher costs (University of Idaho, 1999). There are problems with correcting spectral separation because many types of vegetation have the same spectral signature and therefore usually receive the same class label even though these communities should be classed as separate entities. At this point, Geographic Information Systems (GIS) can be very useful. 3.3 A Brief Overview of Geographic Information Systems (GIS) Geographic Information Systems are computer based systems for storage, display, and manipulation of spatial data (Clarke, 1997). GIS is useful for vegetation mapping because it can perform complicated overlays and spatial analysis that historically have been time consuming and very difficult. GIS databases make it possible to model, combine, and analyze spatial relationships between data sets for the characterization and mapping of plant communities 8 The common use of GIS is the storage of thematic data layers, such as vegetation that can be superimposed onto other data layers. GIS data is commonly stored in a computer in two forms: raster or vector. Vector data represents spatial information with lines in coordinate space and stored as points, line, and polygons (Figure 4) and can have textual information stored with them in accompanying databases. Lines are stored as strings of coordinates and spatial relationships can be computed if needed. The vector model permits the closest digital approximation to the original map and also retains spatial relationships such as network linkages, object areas, perimeters, and shared boundaries (Clarke, 1997, University of Idaho, 1999). Raster data divides space into fields and assigns each field a unique value, usually displayed in a square lattice or grid (Figure 4). Raster structure is convenient for imaging systems such as digital satellite imagery. (Figure 5) (Clarke, 1997 and University of Idaho). Figure 4. A represents a heterogeneous area modeled in a vector type and B represents the same area modeled in a raster type (University of Idaho, 1999). 9 Figure 5. The boxed in area represents Mono Lake in the satellite image (www.monolake.org). Current vegetation mapping is utilizing integration of GIS techniques, satellite data, software programs, and ground-truthing. Recently, scientific studies being conducted have illustrated the usefulness of GIS techniques in vegetation mapping. A study in the foothills of San Diego used a combination of ARC/INFO (a well-known GIS package) and a statistical analysis program called S-Plus to map vegetation (S-Plus, 1999). The team of professors and students analyzed Landsat Thematic Mapping satellite data to map general vegetation categories, and to obtain more detailed vegetation types, they collected field data for each location. ARC/INFO was then used to bring all the data together and to determine the most probable vegetation growing at each site on the map (S-Plus, 1999). In Norway, mapping plant communities is accomplished using infrared aerial photographs, GIS, and digital elevation models (DEM). The study utilized groundtruthing in which vegetation was sampled systematically along transects that covered all the major plant communities. A Global Positioning System (GPS) unit was also used to determine the positions of the samples, which ensured that the sample area could be 10 precisely matched to the corresponding area on the infrared aerial photographs. The infrared photographs were then transformed into a raster format with a scanning device. This information was then inserted into a GIS. Nilsen, et al, (1999) explained that the modeling method operates by crosslinking field data (i.e. the defined plant communities) and the raster layers of the GIS. Probability models were built to establish links between GIS data layers, some of which were derived from the DEM, and plant communities resulting from classification of field data. 4 Methods 4.1 Study Area The study area was located on the south/southeast side of Mono Lake, just west of Navy Beach, southeast of Lee Vining, California. The plant communities sampled are classified as Alkaline Sink Scrub and Big Sagebrush Scrub (Figure 6). Figure 5. The boxed in area represents the location where vegetation samples were taken. (Hart, 1996) 4.2 Field Methods One continuous transect 500 meters in length was run south from a point on the shore of Mono Lake, west of Navy Beach. Circular quadrats with a radius of 5.6 meters 11 were used to sample shrub vegetation (Kent and Coker, 1994). One circular quadrat was set down at random intervals based on a random number table (Zar, 1974). In each quadrat the number of both Rabbitbrush (Chrysothamnus nasueous) and Big Sagebrush (Artemisia tridentata) were counted and the percent cover of each species was estimated. Topographic data was also gathered along the transect using a laser level (See Appendix A). Global Positioning Systems data was gathered using a GPS unit and an exact position for each quadrat was established. 4.2.1 Vegetation Description As a result of the limited time for this study, only one transect was sampled. Two plant species, Rabbitbrush (Chrysothamnus nauseosus) (Figure 7) and Big Sagebrush (Atremisia tridentata) (Figure 8) were considered in the study. These two species were chosen because of their abundance and because they are easily recognized. Figure 7. Rabbitbrush (Chrysothamnus nauseosus) ( Bowers, 1993) 12 Figure 8. Big Sagebrush (Artemisia tridentata) (Bowers, 1993) 5. Results 5.1 Vegetation Data Results from the vegetation samples are displayed in Table 1. The data reveals that the percent cover and number of individual Rabbitbrush (Chyrsothamnus nauesous) was highest closest to the shoreline and decreased with increasing distance from the shore. The percent cover and number of individuals of Big Sagebrush (Artemisia tridentata) was lowest closest to the shoreline and increased with increasing distance from the shore. However, quadrats 2,7, and 10 did not follow the general trend. Table 1. Represents Vegetation Data Collected in the Field Quadrat Number Point of Quadrat on Transect in Meters % cover of Rabbitbrush 1 2 3 4 5 6 7 8 9 10 11 0 13 43 81 101 157 195 218 223 244 273 0 70 35 85 72 45 20 15 10 10 <1 % Cover of # of # of Big individuals of individuals of Sagebrush Rabbit brush Big Sagebrush 0 0 <1 0 0 5 0 0 0 11 30 0 21 19 32 36 25 7 16 11 15 2 0 0 2 0 0 14 0 1 1 4 12 13 12 13 14 15 16 17 18 296 328 364 397 456 485 496 0 <1 0 0 0 0 0 35 55 70 50 75 81 79 0 1 0 0 0 0 0 17 10 23 27 30 33 29 5.2 Topographic data A topographic profile is presented in Figure 9. Results showed that topography increased with increasing distance form the shoreline. However, there were areas where the change was very small. These locations represent old beach ridges and manifested themselves in the form of terraces. Location along transect (m) Topographic Profile 500 400 300 200 100 0 1 5 9 13 17 21 25 29 33 37 41 45 49 53 57 Height (m) Figure 9. Graph represents topographic profile 5.3 Global Positioning Systems (GPS ) Data Raw GPS data was also gathered. The exact position of every qaudrat sampled was taken using a GPS unit. Figure 10 shows an image of the transect with each circle representing a quadrat. 14 Figure 10. Represents the raw GPS data gathered along the transect. 6. Discussion The results from the vegetation samples taken at Mono Lake showed a density gradient of vegetation moving away from the lake shore. The occurrence and percent cover of Big Sagebrush (Atremisia tridentata) increased from the shore of the lake and Rabbitbrush (Chrysothamnus nauseous) decreased. This could be due to Rabbitbrush being more salt tolerant and having higher water demands than Big Sagebrush. It is important to note that the time that elapsed while gathering this information was over two hours and covered only a small portion of the vegetation present. Time is a factor for people who make maps to consider when taking on the daunting task of ground-truthing a large area of vegetation. Another factor that could have influenced the results is topography. Results showed that topography also formed an increasing slope along the transect. An increased slope may have increased the vegetation’s exposure to higher amounts of wind and weather elements. Big Sagebrush is a hearty plant and may be more tolerant to harsh 15 weather conditions, thus being able to thrive in elevated areas. Along the topography gradient there were three spots where the ground leveled out and vegetative cover decreased. These areas represent remnants of a previous lake shore or beach ridge. The reduced amount of vegetation may possibly be due to an increased salt and mineral deposit left behind by evaporated lake water. Raw GPS data was gathered so that if needed, locations of field sites could be directly located on a map or satellite imagery. The GPS positions allow direct comparisons of existing map information to actual field locations and thus map discrepancies can be easily resolved. 7. Conclusion The field methods conducted in the study gave the researchers a clear idea of how ground-truthing methods for vegetation mapping are conducted, the methodology, and the amount of time involved. The study showed that Big Sagebrush(Artemisia tridentata) and Rabbitbrush (Chrysothmanus nauseous) form a distribution gradient moving away from the shore of Mono Lake. Topography also showed a gradient, increasing south of the shoreline. The was study conducted to see how GIS technology aids advancing the process of vegetation mapping. After examining Mono Lake, it is evident that it would be a good place to utilize GIS techniques to make high quality, accurate, and easy to update vegetation maps 16 Appendix A Location along transect(m) Actual Height (m) 0 3 5 10 15 18 21 25 30 32 34 38 41 44 45.8 46 50 60 70 80 120 130 140 150 160 170 180 190 200 210 220 230 237 240 244 250 260 270 280 290 300 310 320 330 340 350 360 370 0 0.11 0.01 0.18 0.44 0.49 0.54 0.71 1.04 1.44 1.57 1.8 2.17 2.45 2.93 2.69 3.19 3.55 3.93 4.3 4.54 5.24 5.94 6.22 6.81 7.75 8.43 8.73 8.73 8.94 9.32 9.66 9.99 10.38 10.99 10.97 11.56 12.49 12.82 13.38 13.95 14.34 14.84 15.01 15.33 15.82 16.07 16.5 17 380 390 400 410 420 430 440 450 460 500 16.7 16.95 17.29 17.36 17.44 17.59 18.13 20.18 20.78 18.63
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