Harold N . Moldenke.
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and other authors ) as synonyms of Priva, are actually synonyms
of Gastelia Cav. ), a distinct although closely related genus of southern
temperate South America, which also, as Junell has pointed out,
shows considerable relation to Boucliea.
Priva is a member of the subfamily Verbenoideae, tribe Priveae
Briq., according to B r i q u e t ' s excellent classification of the family
in Engler and Prantl's "Die Natürlichen Pflanzenfamilien" ). In this
tribe also belongs the genus Dipyrena Hook., which differs in having
small, alternate, entire, 1-nerved leaves, the calyx in fruit open at
its apex, and the spiciform inflorescences much abbreviated. It is
a native of Mendoza, Argentina. In this tribe also belongs the genus
Castelia referred to above, which differs in being a group of glabrate,
more or less fleshy xerophytes with smooth fruits, flowers irregularly
scattered or whorled in the inflorescences, and reproducing mostly
by tubers, rarely producing fruit.
S c h a u e r , in 1847, divided Priva into two sections: 1. Eupriva
Schau, ["calyx fructifer inflatus, capsulam laxe ambiens. Cocci biloculares, dispermi"], including P. laevis, P. leptostachya, P. bahiensis,
P. dentata, and P . echinata, and 2 . Aparinaria Schau, ["calyx fructiger
capsulam arete ambiens et cum ea bipartibibs. Cocci loculo altero
abortivo uniloculares, monospermi"], including P . hispida and P .
aspera. B o c q u i l l o n , in 1863 ), combined the genus Dipyrena Hook.
[Wilsonia Gill, et Hook.], mentioned above, with Priva, although
he admits that its one known species differs from all other known
species of Priva in many respects. He thought, however, that the
differences are not sufficient in weight to warrant a new generic
name. This is also the view of S c h a u e r , B r i q u e t , and K o b u s k i
in regard to the genus Castelia. B o c q u i l l o n divided Priva, there­
fore, into 2 sections: 1. Eupriva Schau, sens, e m e n d . , characterized
b y (he says) prickly fruit and opposite penninerved leaves, and in­
cluding all of Schauer's Eupriva and Aparinaria species, and 2 . Di­
pyrena (Hook.) Bocq., characterized b y the fruit not being prickly
and b y alternate 1-nerved leaves. Briquet, in 1895, recognizing about
10 species in the genus, divided it into 3 sections: 1. Castelia (Cav.)
Briq., 2 . Eupriva Schau., and 3. Aparinaria Schau., keeping Dipyrena
distinct as a separate genus. The genus Blairia Houst., as inter­
preted b y G a e r t n e r and b y some other workers and as accepted
b y these authorities, is equivalent to the section Aparinaria of
S c h a u e r . G a e r t n e r , however, erred by including in Blairia such
other species as Phyla nodiflora (L.) Greene and Lippia javanica
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3
4
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2
) Junell, Symb. Bot. Ups. 4, 39—43, 1934.
) Anal. Cienc. Nat. Madrid 3, 134, t. 30 (1801), Ic. et Dec. PI. 6, 60, t. 583,
1801.
3
) Engler et Prantl, Nat. Pflanzenfam. 43a, 144 t 155, 1895.
) Adansonia 3, 212 [Bocq. Rev. Verb. 116], 1863.
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