Freshwater Biology (2003) 48, 1147–1160 Droughts and anti-droughts: the low flow hydrology of Australian rivers T. A. MCMAHON*,† AND B. L. FINLAYSON*,‡ *Cooperative Research Centre for Catchment Hydrology, and Centre for Environmental Applied Hydrology, The University of Melbourne, Victoria, Australia †Department of Civil and Environmental Engineering, The University of Melbourne, Victoria, Australia ‡School of Anthropology, Geography and Environmental Studies, The University of Melbourne, Victoria, Australia SUMMARY 1. Droughts are not easily defined other than by culturally driven judgements about the extent and nature of impact. Natural ecosystems are adapted to the magnitude and frequency of dry periods and these are instrumental in controlling the long term functioning of these systems. 2. In unregulated rivers, low flows are derived from water in long-term storage in the catchment, commonly as shallow groundwater. Four types of low flow sequences are evident for representative rivers from each of the seven flow regime zones in Australia and an arid zone stream: perennial streams with low annual flow variability that have seasonal low flows but do not cease to flow; perennial streams with high annual variability that cease to flow in extreme years; ephemeral streams that regularly cease to flow in the dry season; and arid zone streams with long and erratic periods of no flow. 3. Although Australian rivers record runs of consecutive years of low flows longer than would be expected theoretically, the departures from the expected are not statistically significant. Trends and quasi-cycles in sequences of low-flow years are observed over decadal time scales. 4. Examples of the effects of river regulation on low flows in southern Australia indicate that, while in detail the impacts of regulation vary, in general regulation mitigates the severity of low flows. 5. It is our contention that the indigenous biota of Australian rivers are adapted to the naturally occurring low flow conditions and that, while there is considerable scientific interest in the effects of climate change on stream ecology, such studies have little practical relevance for the management of indigenous biota in unregulated rivers. 6. The changes brought about by the regulation of rivers are much more rapid and dramatic than those which might occur as a result of climate change and it is possible to develop management procedures to mitigate them. In regulated rivers, the real problem may be ‘anti-droughts’ – the removal of significant natural low-flow events from the flow pattern. Keywords: anti-drought, Australian rivers, drought, low flows, regulated rivers Introduction Correspondence: Brian L. Finlayson, School of Anthropology, Geography and Environmental Studies, The University of Melbourne, Victoria 3010, Australia. E-mail: [email protected] 2003 Blackwell Publishing Ltd Drought is a culturally loaded word in our society. Objective definitions have proved elusive and mostly they are defined in terms of their social, political or economic impacts (Coughlan, 1985). Further, 1147 1148 T.A. McMahon and B.L. Finlayson droughts are usually categorised as ‘meteorological’, ‘hydrological’ and ‘agricultural’ droughts. Keyantash & Dracup (2002) have reviewed the many indices which have been proposed for measuring the severity of droughts in each of these three categories and point out that precise quantification is ‘a difficult geophysical endeavour’ (p. 1167). Defining drought as a phenomenon of the natural environment and attempting to measure severity is even more difficult. Keyantash & Dracup (2002) based their assessments on two agricultural regions, where impact on agricultural production provides the culturally relevant scaling factor for defining a drought. In natural ecosystems, individuals and communities must be capable of surviving, in some way, the protracted dry periods that are a natural feature of the system. It could be argued that long dry spells (droughts?) in the natural environment, while they may cause high mortality rates among plant and animal species, are nevertheless one of the components which control the longterm functioning of these systems. Any particular stream will have an expected pattern of low flows, which depends on its seasonal hydrological regime type. Haines, Finlayson & McMahon (1988) identified 15 regime classes globally. Ten of these are found in Australia and seven form spatially contiguous zones which can be used as the basis for regionalisation of Australian streams (Fig. 1). The nature of the low-flow periods experienced in these streams includes regular periods of no flow or very low flow that is a normal part of the regime (e.g. the low winter flows of rivers in the Extreme Late Fig. 1 Zones of seasonal regime types in Australia after Haines et al. (1988) and the locations and designations of stream gauging stations whose records are used in this paper. 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 Low flows in Australian rivers Summer type) and flows that are lower than expected for the time of year. These regime types occur across a range of climatic moisture zones from permanently humid to permanently arid. Consider, for example, the range of climatic moisture status conditions found across the zone shown on Fig. 1 as having an Extreme Late Summer runoff regime. This zone includes rivers that are perennial and rivers that flow only following heavy rain, but they share the common characteristic that the period of most flow is generally in late summer. The low flow behaviour of streams is further complicated by position in the stream channel network. At the upper end of channel networks (the first order streams of Strahler, 1952), channels may carry flow only immediately following rainfall. The active channel network extends upstream during wet periods in all regime and climate types. Thus the length and severity of low flow periods, and frequency of cease-to-flow, typically decrease with distance downstream in the channel network and, therefore, increasing catchment area. Bedrock type also mitigates the nature of low-flow periods. Rivers draining catchments containing major aquifers may continue to flow during long dry periods when neighbouring streams on different bedrock have ceased to flow. Further, in catchments containing aquifers, springs may keep sections of the channel network flowing more or less permanently as described by Lake (2003). In addition to low-flow characteristics determined by regular seasonal conditions and catchment characteristics, many rivers experience longer-term fluctuations as trends and quasi-cycles. There are sequences of consecutive years when flows are significantly below (or above) the long-term average. These result from a range of climatic mechanisms at the global scale that affect Australia, including the El Niño Southern Oscillation (ENSO) (Allan, Lindsay & Parker, 1996); the Central Pacific sea surface temperature oscillations (Power et al., 1999); and those associated with the Southern Ocean, known as the Antarctic circumpolar wave (White & Peterson, 1996). Relationships have also been observed between the Indian Ocean sea surface temperatures and rainfall in Australia (Smith, 1994). River regulation impacts on the nature and extent of low flows in river systems. In Australia, as for the rest of the world, the flows in many river systems are no longer natural throughout the whole catchment 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 1149 (McCully, 1996; Smith, 1998). The construction of storage dams and diversion weirs and pumping by riparian water users introduce impacts on the patterns of natural flows which vary according to the type of regulation involved and the location in the river basin (e.g. McMahon & Finlayson, 1995). The impact of large storage reservoirs depends on the use to which the water is put. Many irrigation dams in Australia are located upstream of the irrigation area and the water is transferred downstream along the natural river channel. This produces a major change of seasonal flow regime but little change to total flow, because water is released down the channel when irrigation demand is high, which, in the major irrigation districts of southern Australia, is usually the period of natural low flow. Where the water is diverted out of the system by inter-basin transfer or for use in urban water supply, the impact on flow downstream of the dam is a major reduction in the amount of flow as well as a change in seasonal regime. In both cases there is also a change to flow variability. Where the dam is located in an area of high local runoff, the impact of the dam will be mitigated downstream by flows from unregulated tributaries. Diversion weirs are commonly used on smaller streams and typically can have a major effect on low flows while the high flows pass over them. This is also the case with riparian pumping schemes, which are sometimes associated with a weir constructed to create a pool from which to pump. It is the case that most hydrological analysis techniques have had their origin in engineering hydrology and have been developed for engineering uses such as the development of water resources or prediction in hydrology. It is almost certainly the case that had the development of such techniques been carried out by biologists seeking to use them to analyse biological problems, there would probably be a quite different suite available. Our purpose here is to use the existing toolkit of techniques to try to explain the low flow behaviour of rivers in ways which may be relevant to river ecology. In this paper, we explore the nature of low flows in rivers using examples derived from Wullwye Creek, New South Wales, a typical cool-temperate stream with relatively low annual discharge variability by Australian standards. Then we define and describe low streamflow, characterise low-flow sequences, the 1150 T.A. McMahon and B.L. Finlayson frequency of low-flow events, the length of runs of low flows and the nature of trends and quasi-cycles for a representative stream from each of the seven flow regime zones, and the Todd River, Northern Territory, which represents a stream in the most arid part of Australia. We then examine the effects of river regulation on low-flow periods, using the Snowy and Lachlan Rivers as examples, and introduce the concept of ‘anti-drought’, i.e. the provision of low flows that are larger and more persistent than those which occur naturally. Methods Flow data for Wullwye Creek at Woolway (station 222007), catchment area 520 km2, southern New South Wales (NSW; Fig. 1) were obtained from the NSW Department of Land and Water Conservation. From the 40 years of record, we have chosen 6 years of continuous data, from 7 July 1978 to 21 April 1984, to illustrate some characteristics of low flows. Streamflow hydrographs are characterised by two types of flow, commonly referred to as quickflow and baseflow. Quickflow can be taken to represent the surface runoff which occurs during and immediately following rainfall, while baseflow comes from the slow drainage of the saturated storages in the catchment, usually local groundwater. Low flows are mainly baseflow, and the rate of baseflow is proportional to the remaining volume of saturated storage that is connected to the relevant stream reach. When flow is supplied from these stores, points on the hydrograph will plot as a straight line when discharge is plotted on a log scale and time on a linear scale. As a general rule, where the hydrograph begins to plot as a straight line, surface runoff has ceased. Surface runoff is assumed to have begun when the hydrograph rises. Recession coefficients are calculated for a 73-day period during the relatively wet summer of 1978/79 and for the much drier summer of 1980/81 in Wullwye Creek. Baseflow recession curves are described by: Qt ¼ Q0 Kt ; ð1Þ where Qt, Q0 are the respective flows at time t and at time 0 (when surface runoff ceases), K is called the recession constant and t is time since surface runoff ceased. For daily data, a typical value of K ¼ 0.95 (Pilgrim & Cordery, 1993). For each of the seven seasonal flow regime types, we have selected a ‘typical’ stream, for which a reasonably long flow record is available (Table 1), and we use these seven stream flow records to characterise the pattern of low flows for each regime type in the absence of significant flow diversion or regulation. The Todd River is included as an example of streams in the more arid parts of Australia which frequently cease to flow. The analyses that follow are based on monthly and annual flows. The mean annual runoff of the eight catchments varied from 31 to 1040 mm (Table 1). The range of variability of annual flows observed for Australian streams is adequately represented by the coefficients of variation (CV) of annual flows of these catchments (McMahon et al., 1992). The lag one auto-correlation coefficient (r1) is a measure of the persistence of the flow or, in other words, the likelihood of successive years of low flows or successive years of high flows. Values shown in Table 1 are typical of Australian streams (McMahon et al., 1992) except for two large values, 0.27 and 0.35 for the Todd and Thomson Rivers, respectively. Table 1 Characteristics of stations used in this study. Regime refers to the pattern of flows through the year as defined by Haines et al. (1988) River Station no. Gauging station Regime Catchment area (km2 ) Period of record (years) MAR (mm) CV r1 Coranderrk Ck Thomson Styx Barron Ord Broken Queanbeyan Todd 229115 225210 206001 110003 809302 404200 410701 060046 No. 2 Weir The Narrows Jeogla Picnic Crossing Coolibah Pocket Goorambat Googong Wigley Gorge Early spring Moderate winter Moderate late summer Early autumn Extreme late summer Extreme winter Moderate autumn Extreme late summer 18.6 518 163 220 46100 1920 873 360 1909–99 (91) 1886–1970 (85) 1919–90 (72) 1926–99 (74) Modelled (87) 1917–67 (51) 1913–74 (62) 1962–95 (34) 1040 510 700 640 83 120 130 31 0.28 0.42 0.57 0.57 0.71 1.00 1.06 1.50 0.06 0.35 0.17 0.04 0.01 0.10 0.07 0.27 MAR, mean annual runoff; CV, coefficient of variation of annual flow; r1, lag one serial autocorrelation. 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 Low flows in Australian rivers Low-flow events can be classified by the average recurrence interval (return period) of low-flow sequences of consecutive days, months or years. For low-flow events of <12 months duration, the analysis is relatively straightforward as the sequence can be treated in the same way as the annual or partial flood series (Gordon, McMahon & Finlayson, 1992). For the major low-flow events of longer than 12 months, the fundamental issue is the requirement for the assumption of independence of an n-consecutive-years event (say 3 years) drawn from a record length of N years (say 60 years) of annual streamflow data. In the example of the 3-year event, there are N/n (60/ 3 ¼ 20) largely independent sequences but N ) n + 1 (60 ) 3 + 1 ¼ 58) overlapping and highly correlated sequences. When we choose a specific n-years event from N years of historical data, we are essentially sampling the n-years event from the overlapping sequence. Thus the analysis must take this high correlation into account and a procedure to do this has been developed by Srikanthan & McMahon (1986). The steps in the procedure have not been listed here, but note that the simplest technique assumes that the annual flows can be represented by either a Normal or a Gamma distribution. Taking annual flow below the median as the measure of low flow, we calculated the deviations from the median for the seven representative Australian streams. From these data we could compile frequency distributions of lengths of runs below the median. We then superimposed theoretically derived frequency distributions of expected runs below the median (as a function of the length of data) in order to compare the extent to which the distribution of runs of low flows in these representative Australian streams differ from that which would be predicted theoretically. The method follows the derivation by Yevjevich (1982) but is modified to take into account the finite length of the data record and assumes the annual flows are independent variables, an assumption which holds true for five of the analysed streams. Flow data from the Snowy River in eastern Victoria and the Lachlan River in south-central NSW, each of which is regulated by a large storage reservoir, are used to demonstrate the nature and magnitude of the impacts of regulation on low-flow behaviour. The Snowy River is a coastal river in southeastern Australia with a catchment area of 13 400 km2 and a natural mean annual flow of 2020 · 106 m3. Flow from 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 1151 the upper 14% of the catchment is diverted into the Murray and the Murrumbidgee Rivers, inland flowing streams from which the water is extracted for irrigation farming. At the diversion point, the Jindabyne Dam, only 1% of the original flow is left in the river for local riparian users (Finlayson, Gippel & Brizga, 1994). The Lachlan River, more than 900 km in length, drains a catchment of 84 700 km2 and provides irrigation for 500–1000 km2 annually. The Lachlan system consists essentially of a linear stream controlled by one major reservoir (Wyangala Dam with a storage capacity of 1220 · 106 m3), a very small dam and two off-river storages. The annual flows are characteristic of highly variable western flowing NSW streams. For this system, irrigation water is taken directly from the river or from regulated effluent streams. Wyangala and the other storages were constructed to provide more reliable supplies. We compare the characteristics of low flows before the dams were constructed with the flows as regulated by the dams. The analyses differ from those used in our regional survey above. First, we compare annual flow totals and compare daily flow duration curves before and after the dams were completed. Results Nature of low streamflow Any record of streamflow will exhibit consecutive periods of low flow of various lengths. For example, the early years of the 40-year record of flows in Wullwye Creek at Woolway contained many fewer extended periods of low flow than the latter period (Fig. 2a). This inconsistency through time is an important feature of Australian rivers. The flow during the selected 7-year period (1978–84) at Wullwye Creek was characterised by a high-flow period in winter and low-flow period in summer, but, while the interannual pattern was consistent, the magnitude of the flows was not. For example, the mean daily winter (July– September) flow of 0.006 mm day)1 for the year 1982 was 1% of that for 1978, while the mean daily summer flow of 0.0001 mm day)1 for 1982/83 was 0.2% of the mean daily summer flow for 1978/79. These yearto-year extremes illustrate the range of hydrologic conditions that aquatic flora and fauna encounter even in a system of relatively low variability. 1152 T.A. McMahon and B.L. Finlayson Fig. 2 Daily streamflows for Wullwye Creek at Woolway (222007), southeastern Australia, from (a) 25 March 1949 to 21 June 1999, (b) 7 July 1978 to 21 April 1984, (c) 9 November 1978 to 20 January 1979 (linear scale), (d) 9 November 1978 to 20 January 1979 (logarithmic scale), and (e) 1 November 1980 to 19 January 1981 (logarithmic scale). Periods of missing data in panel (a) are indicated by bars below the zero line. The separation of surface and baseflow is important in understanding the source of water for the stream. Over the 73 days of flow in Wullwye Creek during the wet summer of 1978/79, 58 days (79%) were baseflow (A-B¢, B-C¢, C-D¢ and D1 onwards), yielding 66% of the total flow (Fig. 2c). Because the immediate source of this water is different to the source of surface runoff, the water quality of the flows may also be very different. From eqn (1), K values (recession constant) for A, B and C in Fig. 2d are 0.92, 0.92 and 0.89, respectively. In the case of D1, D2 and D3, where there are three different recessions in sequence, the values are 0.82, 0.90 and 0.96, respectively. Given the slopes of recessions A, B and C, it is probable that the recession beginning at D1 is a result of interflow rather than groundwater discharge, where interflow is defined by Mosley & McKerchar (1993) as ‘…rapid subsurface flow through pipes, macropores and seepage zone in the soil’, and Pilgrim & Cordery (1993) suggest that for daily data K for interflow ranges from 0.8 to 0.9. An 80-day sequence of daily flows during the much drier summer of 1980/81 included five minor stream rises (Fig. 2e). The recession coefficients were 0.77, 0.72, 0.63, 0.60 and 0.56. These were much steeper than the D2 and D3 values observed in the summer of 1978/79. Furthermore, the slopes became progressively steeper over the 80 days. In mid-November it 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 Low flows in Australian rivers 1153 took 9 days for the discharge to reduce by an order of magnitude but by mid-January it took only 4 days. From a hydrologic point of view, this indicates that the main sources of groundwater during the recessions in the summer of 1978/79 were dry, so that the recessions in 1980/81 were only from localised groundwater (probably bank storage) and as the flows were so low, evaporation from the stream, together with transpiration from riparian vegetation, was greater than the groundwater discharge (point potential evaporation in the area typically is about 6 mm day)1; Wang et al., 2001). The recessions shown in Fig. 2e were for extremely severe conditions and illustrate the reducing effectiveness of rainfall as the catchment dries out. Over this 80-day period, the average discharge from the 520 km2 catchment was only 0.08 m3 s)1 (equivalent to 0.4 mm month)1 runoff) which for an average flow velocity of, say, 0.5 m s)1 and an average stream width of 10 m, the depth of flow was only about 16 mm. Under these conditions, small falls in discharge will make relatively large reductions in wetted channel area and thus habitat availability for aquatic organisms. At higher flows, quite large recessions in baseflow will have much less impact on habitat availability, because they mainly reduce water depth and these differences are mitigated by channel shape (Fig. 2c). Characterising low flow sequences Figure 3 shows time series of annual streamflows of three of the streams from the seven chosen to represent the regime zones of Australia. Coranderrk Creek has a low annual CV and the Queanbeyan River a high CV, and the Todd River represents the arid zone. Differences in streamflow variability are clearly apparent and other steams in Australia grade between these extreme cases. By examining the flows of the streams in the seven climatic zones in Australia (Fig. 1), we have identified four general types of low-flow sequences that may be of particular relevance to stream ecology. The first is for perennial streams that have low annual variability and, given the climatic zones in which they are located, even under extreme low-flow conditions would not be expected to cease to flow. Three streams fall into this category: Corranderrk Creek (CV ¼ 0.28), Thomson River (CV ¼ 0.42; Fig. 4a), and Barron River (CV ¼ 0.57). 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 Fig. 3 Annual streamflow series for (a) Coranderrk Creek at No. 2 Weir, (b) Queanbeyan River at Googong, and (c) Todd River at Wigley Gorge. The second type is illustrated by the Queanbeyan River (CV ¼ 1.06), which is a perennial stream but in extreme years cease-to-flow can occur (Fig. 4b). The Styx River (CV ¼ 0.57) and the Broken River (CV ¼ 1.0) are also representatives of this type. The third type is represented by the Ord River (CV ¼ 0.71; Fig. 4c), which is an ephemeral stream where cease-to-flow occurs during the winter each year. Note also that while flow occurs every summer, there are substantial differences between successive summers. The fourth type of flow sequence, illustrated by the Todd River (CV ¼ 1.5; Fig. 4d), is a highly variable ephemeral arid-zone stream. While the Todd has an Extreme Late Summer regime, there is much more year-to-year variability in the regime than is the case for the Ord (Fig. 4c). While most of the large flow 1154 T.A. McMahon and B.L. Finlayson Fig. 4 Monthly streamflows for the (a) Thomson River at The Narrows, (b) Queanbeyan River at Googong, (c) Ord River at Coolibah Pocket, and (d) Todd River at Wigley Gorge. events in the Todd occur in late summer, there are some years with no summer flows at all (e.g. 1969/70; 1979/80) and some years when the major flow events do not occur in late summer (e.g. 1986). This behaviour is typical of streams that have their headwaters within the semiarid and arid zones of inland Australia though in terms of water resources they have only minor local significance. Frequency of low-flow events The computed rank 1 (i.e. the lowest in the record) 1-, 2-, 3- and 5-year low-flow sequences of five Australian rivers are given in Table 2. Note that for the Ord and Styx, neither the Normal nor the Gamma distribution was applicable and the average recurrence intervals could not be computed using this method. These results can be used as the basis for making probabilistic statements about the likelihood of occurrence of events of these magnitudes and enables observed events to be scaled within the record. For example, the lowest 1-year flow at Coranderrk is an extreme event when compared with, say, the lowest 1-year flow for the Barron. Clearly this Table 2 Average recurrence interval of lowest n-years flows. (Could not be computed for the Styx and the Ord) Recurrence interval of the n-years low flow River Station no. 1 2 3 5 Corranderrk Ck Thomson Barron Broken Queanbeyan 229115 225210 110003 404200 410701 370 99 14 79 19 250 270 60 370 58 470 67 140 300 110 110 140 170 75 150 analysis is heavily dependent on the appropriateness of the probability distribution being used but it does provide a consistent procedure which can be used in ecological studies to assess the ‘importance’ of observed low-flow events to the system. Length of runs of low flow Another relevant property of low flows is the lengths of runs of low-flow years, defined here simply as years with flows less than the mean, although other definitions are possible (Fig. 5a,b). Note that the 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 Low flows in Australian rivers 1155 Fig. 5 Above and below median streamflows for (a) Coranderrk Creek, and (b) Queanbeyan River, and theoretical and actual frequency distributions of runs below the median for (c) Coranderrk Creek and (d) Queanbeyan River. magnitude of deviations below the mean for the Queanbeyan River are small compared with Coranderrk Creek, indicating a much more skewed distribution of annual flows in the case of the latter. If this behaviour can be described using a theoretical model, then it is possible to predict the likelihood of occurrence of runs of low flows of any given length. This approach is commonly used in water resources analysis but clearly also has ecological significance. For the rivers used in the present study the maximum run lengths are considerably above theoretical expectations from an independent series, but the differences are not statistically significant at the 5% level: Coranderrk has 2 · 4 years of consecutive low flows, Thomson 1 · 7 years, Styx 2 · 6 years, Ord 1 · 6 years, Broken 1 · 8 years and Queanbeyan 1 · 8 years. Such long periods of low flow are more likely to influence the structure of aquatic communities and their physical habitat. The longer periods would be seen by Erskine & Warner (1988) as being part of their drought- (and flood-) dominated regimes, which they argue play an important role in 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 determining channel form and behaviour, at least in the coastal streams of NSW, Australia. Trends and quasi-cycles Coranderrk Creek and the Queanbeyan River, rivers with the lowest and highest CV, respectively, in our seven station sample, are used to illustrate the quasicycles in these records. Again, as we move from the least to most variable streams, the quasi-cycles become stronger, with extended periods up to several decades of low and/or high flows. Some of the cycles are particularly dramatic. The Queanbeyan River (Fig. 6b) exhibited only small variations during the period from 1920 to the early 1940s and this was followed by a dramatic shift to larger annual flows such that the period from 1947 to 1966 yielded more than double the mean annual flow of the previous decades. Such major shifts over decadal time scales are noteworthy features in Australian river flow records and more likely to be of particular relevance to the ecological functioning of these systems. 1156 T.A. McMahon and B.L. Finlayson Fig. 6 Moving average plots of annual streamflow in (a) Coranderrk Creek and (b) Queanbeyan River. Although we have not formally analysed these records for trend here, slight trends are apparent in the case of both the Queanbeyan River and Coranderrk Creek although in opposite directions (Fig. 6). Interestingly, the record for Wullwye Creek, which is not regulated and was chosen more or less randomly just to illustrate baseflow recessions earlier in this paper, shows a dramatic downwards trend over the period of record (Fig. 2a). Further discussion of this type of behaviour in Australian rivers can be found in Finlayson & McMahon (1991). Regulated river systems The Lachlan River at Cowra is regulated by the Wyangala Dam and as the river is used as a conduit to deliver irrigation water, there is no significant effect on total flow but the seasonal regime has changed somewhat (Fig. 7). Consistent with this use of the river, the summer flows are higher post-regulation and the winter flows are lower as the dam is being refilled for the next irrigation season (Fig. 7b). While Fig. 7 suggests that the effect of regulation is marginal, the daily flow duration curves suggest otherwise (Fig. 8). There is evident a dramatic Fig. 7 Annual streamflows (a) and mean monthly streamflows (b) for the Lachlan River at Cowra. difference in the flow duration curves before and after regulation for the Lachlan River at Cowra and Booligal, with the most significant differences being at the low-flow end of the curves. At Cowra, for example, the 95% low-flow post-regulation is three orders of magnitude larger than the natural flow. At Booligal prior to regulation, the river ceased to flow for 25% of the time, whereas in the present regulated regime the river rarely ceases to flow. More details are given in Panta et al. (1999). In the case of the Snowy scheme, water is diverted out of the catchment. When the scheme was designed, a small river valve was built into the Jindabyne Dam large enough only to provide stock and domestic water to riparian landholders living just downstream of the dam. The first gauging station is 25 km downstream of the Jindabyne Dam at Dalgety. Here mean annual flow before the dam was 1187 · 106 m3 and after the dam this was reduced to 68 · 106 m3. Daily flow duration curves for the Snowy at Dalgety for the predam period (1950–67) and the postdam period (1968-99) show that low flows have been 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 Low flows in Australian rivers 1157 indicator of baseflow contribution in natural streams and in regulated rivers is an indication of the persistence of low flow. For the Snowy at Dalgety, this ratio has tripled in value following regulation, from 0.22 to 0.66. Discussion and concluding remarks Fig. 8 Daily flow duration curves for Lachlan River at (a) Cowra and (b) Booligal for regulated and unregulated conditions. reduced by around an order of magnitude (at Q95 from 0.1 mm day)1 to 0.007 mm day)1; Fig. 9). While the low flows are lower, they are nevertheless persistent and this is a feature of this type of regulation. Where flows are diverted out of the system, releases are made to provide for riparian users and these tend to be more reliable than the natural low flows they replace. The ratio Q90/Q50 is an Fig. 9 Daily flow duration curves for the Snowy River at Dalgety before and after the construction of the Jindabyne Dam. 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 In this paper we have described the characteristics of low flows in Australian streams. There is reason to believe that this pattern of flow variability, or something like it, has persisted for a very long time on the Australian continent. In discussing the strategies used by Australian freshwater biota in coping with low flows, Lake (1995) points out that the ‘richness of refugial strategies is in contrast to the paucity elsewhere… Such adaptive strategies have had a long evolutionary development in Australia.’ One of the major drivers of climatic variability in Australia is the ENSO effect and Kuhnel et al. (1990) have shown the extent and magnitude of its impact on streamflow variability. Nicholls (1989) points out that climate variability in Australia is amplified by ENSO such that variability is higher than in other areas of the world with equivalent climates not influenced by ENSO. He uses a range of animal adaptations to climate variability to argue that much of the Australian biota is adapted to variability and that this adaptation is more complete than in other areas of the world. The conclusion can therefore be drawn that the influence of ENSO in the Australian climate is of sufficiently long standing to have influenced the course of species evolution. The influence of flow variability and low-flow behaviour on fish distribution can also be seen at the level of individual river systems. As pointed out above, flow tends to become more reliable with distance downstream (and increasing catchment area) in a river system. A number of studies of fish distribution in Australian river systems have pointed to a downstream increase in fish species richness (Lake, 1982; Hortle & Pearson, 1990; Gehrke & Harris, 2000). Our discussion of the low-flow behaviour of Australian rivers has not been exhaustive; there are many techniques and procedures for analysing low flow behaviour which we have not discussed, particularly spell analysis (Donald, Nathan & Reed, 1999), which may be of particular interest to river 1158 T.A. McMahon and B.L. Finlayson ecologists. Typical flow sequences consist of many small low-flow events and few major ones (Fig. 5). The probability of these events can be determined from the flow record and for this reason we encourage aquatic ecologists interested in studying the impacts of low flows on in-stream biota to locate their study sites near gauging stations with a long flow record so that the periods they have studied can be located in the context of the long-term hydrological record of the stream. Where ecological studies are located in ungauged reaches or catchments, it is possible to estimate characteristics of the flow behaviour (R.J. Nathan, unpublished data; Nathan & Weinmann, 1993; McMahon et al., 2002) although this lacks the precision of real flow records. The difficulty in determining whether a significant low-flow period has started, and of predicting how long and how severe it will be, has meant that these periods are not well described ecologically (Lake, 2000). Lake (2000) has argued that there is a need to improve understanding of disturbance in aquatic ecosystems, especially faced with global climate change, which is predicted to have a significant impact on the frequency of extreme events (Whetton et al., 1993). While this may be true, the identification of a climate change signal in the Australian streamflow records has so far proved elusive (Gan, McMahon & Finlayson, 1990; Chiew & McMahon, 1993). Flow forecasting is becoming available which will allow some level of prediction of impending low-flow periods (Chiew et al., 2000). Drought is a poorly defined word and many discussions of drought get bogged down in unproductive attempts to define it. The point we have tried to make here is that the flows of natural river systems encompass a range of flow conditions from extremes of high flows to extremes of low flows. These are not disasters when viewed from the perspective of the natural ecosystem. They may cause major disturbances in those systems but any communities or species unable to cope with that magnitude and frequency distribution of flow events would seem to have no place in the system in the long term. Ecological processes in rivers are controlled by flow variability (Puckridge et al., 1998) and low flows are an inherent part of the pattern of variability. Of much more significance in this context is the impact of river regulation on low-flow behaviour. Unlike changes driven by climate change, the impacts of river regulation on flows are dramatic, usually quite sudden, and clearly discernible in the records. Even if the most extreme predicted climate change scenarios were to eventuate, they would be small in the scale of their impact and rate of onset when compared with the changes that have already occurred as a result of regulation. It is for this reason that a better understanding of the impact of low flows on river ecosystems is needed. There is little that river managers can do to deal with climatically driven changes to flows in unregulated systems but on regulated rivers, where there are major changes already in place, the capacity exists to manipulate flow patterns so as to provide ecological benefits. In our regulated rivers, we have replaced a natural distribution of low flows with an artificial distribution that lacks many natural low-flow events. In the terminology of this special issue, some of the ‘droughts’ have been removed from the regulated rivers and replaced with what we term ‘antidroughts’ – periods that have higher flows than would be expected naturally. Furthermore, in some periods, high natural flows have been replaced by periods of considerably lower flows as a result of water extraction or stream regulation. We contend that ‘drought’, however defined, is not a problem for the management of Australian freshwater ecosystems, though there is clearly scientific interest in understanding the way these systems cope with such events and recover from them. The issues for management are the ‘anti-droughts’ and induced droughts that now characterise most, if not all, of our regulated rivers. Acknowledgments We are grateful to our colleagues Drs Senlin Zhou and Murray Peel who processed data for us and to the Department of Natural Resources (Queensland), Ecowise Environmental Ltd, Melbourne Water Corporation, Theiss Environmental Services and the Water and Rivers Commission (WA) for providing streamflow data. The final form of this paper owes much to the efforts of Drs Paul Humphries and Darren Baldwin of the Murray-Darling Freshwater Research Centre, Albury. 2003 Blackwell Publishing Ltd, Freshwater Biology, 48, 1147–1160 Low flows in Australian rivers References Allan R., Lindsay J. & Parker D. 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