METADATA Title 0.01° stack of climate layers for continental analysis of biodiversity pattern, version 1.0 DATASET NAME Individual attributes are raster datasets named as listed in the summary table below Custodian CSIRO Ecosystem Sciences Jurisdiction Australia DESCRIPTION Abstract This metadata statement documents a related set of datasets (attributes) derived from a common source and software tools. These data provide rasterised layers of climatic variables hypothesised to explain spatial patterns in biological diversity at continental scales for use with statistical modelling tools (e.g. see methods described in Elith et al. 2006). Specifically, these data were derived for modelling the compositional pattern of multiple species with environmental factors such as climate, soil and topography using the statistical technique Generalised Dissimilarity Modelling (Ferrier et al. 2007) applied to continental Australia (Williams et al. 2010a; b). Monthly climate layers were derived using version 3 of the national 9 second DEM (Hutchinson et al. 2008), resampled to 0.01 degrees, and the ESOCLIM module of ANUCLIM version 6.0 (beta) (Hutchinson et al. 2000 and T. Xu pers. comm.) with the Australian Climate Surfaces version 2.1 (Hutchinson and Kesteven 1998). The derived climate variables are monthly mean values for minimum temperature, maximum temperature, precipitation, solar radiation, evaporation, wind and others. Some of these monthly variables were used to generate growth indices (using the GROCLIM module of ANUCLIM) and a number of other variables related to mean and extreme conditions of atmospheric and soil moisture conditions and climatic seasonality (Williams et al. 2010a; b). The Australian Climate Surfaces version 2.1 for the ANUCLIM-BIOCLIM package are interpolated 75-year means (1921 to 1995) for rainfall and temperature and 25-year means for other variables (1970-1995), representing the period prior to the onset of measureable climatic warming. These data for GDM analysis were masked to consistently define data/nodata values and supplied in DIVA-GIS floating-grid format in the WGS84 geographic reference system. Spatial extent and mask is the same as the substrate/terrain grids described elsewhere (Williams 2010a). This metadata statement describes the data in a formal metadata style. More detail is given in the project report “Harnessing Continent-Wide Biodiversity Datasets for Prioritising National Conservation Investment” (Williams et al. 2010a; b). Two other metadata statements describe the substrate/terrain grids (Williams 2010a) and biotic attributes (Williams 2010b) which derived from National vegetation mapping. OTHER METADATA ANZCW0703011541: GEODATA 9 Second Digital Elevation Model Version 3 (Hutchinson et al. 2008) Stein J. (2008) Metadata: Environmental attributes compiled for the continental GDM analysis. Fenner School of Environment and Society, The Australian National University, Canberra. Williams K. J. (2010) 1km resolution terrain and substrate layers for continental analysis of biodiversity pattern (metadata for digital spatial data). CSIRO Sustainable Ecosystems, Canberra. Williams K. J. (2010b) 1km resolution vegetation attribute layers for continental analysis of biodiversity pattern (metadata for digital spatial data). CSIRO Ecosystem Sciences, Canberra. GEOGRAPHIC EXTENT Geographic Extent Name(s) Australia GEOGRAPHIC BOUNDING BOX North Bounding Latitude -9.0 South Bounding Latitude -43.8 East Bounding Longitude 153.64 West Bounding Longitude 112.9 DATA CURRENCY Beginning date 1921 Ending date 1995 Dataset Status Complete Maintenance and Update Frequency Not planned Data Format DIGITAL - ESRI Arc/Info binary GRID (floatgrid format) and DIVA-GIS (http://www.diva-gis.org/) Lineage Monthly climatic layers were computed with the ESOCLIM module of ANUCLIM version 6.0 (beta) (Hutchinson et al. 2000) for each grid cell of a 0.01 degree resolution DEM derived by resampling the national 9 second DEM version 3 with the Arc/Info RESAMPLE function using bilinear interpolation. Annual growth indices were computed using the GROCLIM module of ANUCLIM, a derivative of GROWEST (http://fennerschool.anu.edu.au/publications/software/growest.php). These and other climatic variables are described below. The values of each layer were consistently expanded around the coastline using the focalmean function in Arc/INFO GRID with neighbourhood shape defined by a circle of radius two cells with DATA values. In each case, the original values of the grids were retained and values were extrapolated outwards from the coast, or to fill data voids where applicable. The data were expanded around the coast because of the coarseness of the 1km grid which results in removal of areas of coastline when viewed at finer scales. Data were masked to consistently define data/nodata values across all climatic and substrate/terrain layers (for the latter, see Williams 2010a). A coastline mask with resolution 0.01 degrees (OZCSTM_1KM) was generated from the current extent of the Australian coastline and surrounding islands defined by the GEODATA Coast 1:100,000 topographic vector data series (Geoscience Australia 2004) (details given in Williams et al. 2010a). This coastline incorporates some minor land and island locations that are not yet captured by the 9-second digital elevation model (Hutchinson et al. 2008). It was created to mask model outputs to better reflect the GEODATA coastline, rather than the expanded coastline mask. Annual Climate Statistics A wide range of bioclimatic indices (climatic factors that influence living organisms) are considered relevant to the distribution of biodiversity, encompassing essential resources (water) or conditions determining growth and survival (frost, heat, drought, wind) or linked to disturbance events (e.g. fire, cyclone, flood). As a result, a large number of bioclimatic indices have been developed for modelling biodiversity patterns (Busby 1991; Nix 1986). Currently 35 bioclimatic indices have been incorporated into the ANUCLIM software package (Hutchinson et al. 2000) as derivatives of monthly mean climatic variables. A key feature of these bioclimatic indices is the two-way combination of climatic factors such as the mean temperature of the driest quarter, where the driest quarter may be any consecutive 12 week period within the year. By ignoring the time of year, seasonal variation in conditions that equate across wide geographic ranges are automatically taken into account. While many of the bioclimatic variables appear meaningful, some can result in unusual spatial patterns with arbitrary sharp boundaries. To avoid this spatial patterning we generated a suite of simple climatic statistics as the minimum and maximum of the monthly values—maximum temperature (°C), minimum temperature (°C), diurnal temperature range (°C), rainfall (mm), evaporation (mm), rainfall-modified solar radiation (MJ/m2/day), wind run (km/day) and wind speed (m/s at 9am and 3pm)—for testing as predictor variables in models of biological response along with other climatic factors. Some of these climatic statistics replicate bioclimatic indices (e.g. Bioclim variable 5 is maximum temperature of the warmest period). The wind datasets (wind run and wind speed at 9am and 3pm) were compiled from 9-sec resolution monthly variables generated by Williams et al. (2006) to map suitable areas for macadamia horticulture. These data were created using version 2 of the 9sec digital elevation model in GDA94 geographics with ANUCLIM version 5.1. The monthly data were projected to WGS84 geographics and resampled to 0.01 resolution using the cubic convolution algorithm. Day length A commonly used indirect factor in biodiversity modelling, correlated with many underlying causal factors, is the geographic gradient in north-south or latitudinal dimensions. The latitudinal gradient in terrestrial species richness is commonly known as Rapoport’s Rule (Gaston 2000; Stevens 1989; Taylor and Gaines 1999; Willig et al. 2003). Such positional factors are not determinants of biodiversity pattern, but correlate with a number of potential causal environmental factors (Austin 1980; 2002; Gaston 2000). A slightly improved derivative of the northsouth factor related to the energy budget and influencing plant phenology is day length (e.g. Rathcke and Lacey 1985). Day length is a rescaling of the latitudinal gradient that is more relevant to biological responses. We therefore applied the day length model defined by Forsythe et al. (1995) in which the timing of sunrise/sunset is defined when the centre of the sun is even with the horizon. These parameters represent the period of effective light for photosynthesis, applicable to ecological studies (Brock 1981; Running and Coughlan 1988). We developed monthly estimates of day length based on the 15th day of each month within a year. Two indices were selected to describe day length – the shortest day (minimum in June) and the longest day (maximum in December). Table 1. Definition day length indices. Label DL_P0 Description Day length Units Definition Hours The monthly length of day in hours where sunrise/sunset occurs when the centre of the sun is even with the horizon, using the calculations in Forsythe et al. (1995). Growth Indices (courtesy Janet Stein, March 2008) The growth Index (GI) is produced with the GROCLIM program from the ANUCLIM V5.1 package using weekly estimates of long-term mean monthly rainfall, temperature, radiation and evaporation (Table 1). The index essentially rates the potential for plant growth between 0 (completely limiting) and 1 (non-limiting). Four GI layers were produced, one for each of the annual mean GI for C3 microtherm, mesotherm and macrotherm plants and for C4 megatherm plants (using different default optimal temperature ranges as defined in the ANUCLIM manual online at: http://fennerschool.anu.edu.au/publications/software/anuclim/doc/groclim.html) (Hutchinson et al. 2000). GI is the product of the three component indices (temperature, light and moisture), the latter (moisture index) is one of the 35 BIOCLIM parameters. The moisture index was computed for a default clay loam soil type with 150mm available water capacity. The GI grids provide a useful summary of climatic factors relevant to plant productivity. Table 2. Definition of growth indices. Label Description Units C4GI Growth index C4 megatherm plants index MEGAGI Growth index C3 macrotherm plants index MESOGI Growth index C3 mesotherm plants index MICROGI Growth index Definition Annual mean growth index for C4 megatherm plants, computed for a clay loam soil type with 150mm available water holding capacity, as the product of three component indices (temperature, light and moisture) using an optimum temperature of 32°C and range 10-45°C. Annual mean growth index for C3 macrotherm plants, computed for a clay loam soil type with 150mm available water holding capacity, as the product of three component indices (temperature, light and moisture) using an optimum temperature of 28°C and range 10-38°C. Annual mean growth index for C3 mesotherm plants, computed for a clay loam soil type with 150mm available water holding capacity, as the product of three component indices (temperature, light and moisture) using an optimum temperature of 19°C and range 3-36°C. Annual mean growth index for C3 microtherm Label Description index C3 microtherm plants Units Definition plants, computed for a clay loam soil type with 150mm available water holding capacity, as the product of three component indices (temperature, light and moisture) using an optimum temperature of 10°C and range 0-20°C. Atmospheric Water Availability Measures of atmospheric humidity (relative humidity and vapour pressure deficit) were computed using published meteorological calculations applied to monthly ESOCLIM variables: minimum and maximum temperature (overnight minimum and daytime maximum), wet and dry bulb (9am and 3pm) and dew point (9am and 3pm) (see http://www.bom.gov.au/climate/cdo/about/definitions9and3.shtml for definitions of 9am and 3pm climatic statistics). Relative humidity (RH) and vapour pressure deficit (VPD) require estimates of actual and saturation vapour pressure at the same temperature. Depending on available meteorological data, there are different ways to estimate relative humidity and vapour pressure deficit (Allen et al. 1998). As saturation vapour pressure is related to air temperature, it can be calculated from the air temperature. Although it is not possible to directly measure actual vapour pressure it can be derived from relative humidity or dewpoint temperature with known parameters for atmospheric pressure, latent heat of vaporisation and the psychrometric constant. Dewpoint temperatures can be calculated from measurements of wet and dry bulb temperatures (psychrometer) when air temperatures are greater than 0ºC. We used these monthly measures of dew point temperature, wet and dry bulb temperatures to calculate vapour pressure deficit following the equations outlined in Allen et al. (1998). For relative humidity we adopted the equation used by the Australian Bureau of Meteorology (after Abbott and Tabony 1985). We used the psychrometric data method above to estimate actual vapour pressure, with altitude derived from version 3 of the 9sec DEM (resampled to 1km resolution) to estimate gridded values of atmospheric pressure. Based on the 9am and 3pm monthly calculations of VPD and RH, we generated the overall mean, minimum and maximum values. Essentially RH and VPD have the same data origin in actual and saturation vapour pressure; one is a ratio of actual to saturation values and the other, a difference. Table 3. Definition of atmospheric water availability indices. Label Description Units Definition VPD215 Vapour pressure deficit 3pm KPa The monthly difference between the amount of moisture in the air (actual vapour pressure) and how much moisture the air can hold when it is saturated (saturation vapour pressure) at that temperature (3pm). VPD29 Vapour pressure deficit 9am KPa The monthly difference between the amount of moisture in the air (actual vapour pressure) and how much moisture the air can hold when it is saturated (saturation vapour pressure) at that temperature (3pm). RHU215 Relative Humidity 3pm % Approximate monthly relative humidity defined as the amount of moisture in the air expressed as a percentage (ratio) of the amount of moisture present if the air was saturated at that temperature (3pm). RHU29 Relative Humidity 9am % Approximate monthly relative humidity defined as the amount of moisture in the air expressed as a Label Description Units Definition percentage (ratio) of the amount of moisture present if the air was saturated at that temperature (9am). Other atmospheric moisture indices in current use in the literature were also calculated. These include the aridity index, dryness index and precipitation deficit. The precipitation deficit has been used to broadly compare the hydrological regimes of sites and evaluate climate change implications (Harmsen et al. 2009). A positive value indicates water in excess of crop water requirements and a negative value indicates a deficit in terms of crop water requirements. An alternative expression is the aridity index which is the ratio of precipitation to evaporation (UNEP 1992 cited in Middleton and Thomas 1997), or the inverse ratio as the dryness index (Zhang et al. 2004). The choice of index to include in an analysis will depend on the alternative hypotheses of biological stress (e.g. due to moisture limitations) or productivity factors driving the patterns of distribution among the biota in question. Table 4. Definition of aridity and precipitation deficit indices. Label ARID ADEF Description Aridity index Precipitation deficit Units Definition Dimensionless The monthly ratio of precipitation to potential evaporation (pan, free-water surface). A numerical indicator of the degree of dryness of the climate at a given location. Adapted from the index proposed by UNEP (1992; cited in Middleton and Thomas (1997)). Mm The monthly difference between precipitation and potential evaporation (pan, free-water surface), without accounting for soil buffering capacity on water availability (after Harmsen et al. (2009), adapted from De Pauw (2002)). Also known as water deficit or hydrological deficit. Values are negative when evaporation demand is greater than rainfall indicating a water deficit. Soil Water Availability Soil moisture and atmospheric humidity are important drivers of biotic responses and therefore commonly used as physiologically-relevant predictors in models of biodiversity distribution to improve their generality. To more effectively capture the interaction between soil and climate, we applied a simple tipping bucket water balance model, similar to that used in GROCLIM to estimate soil water availability using the soil water holding capacity attribute derived from the 1:1M to 1:3M Atlas of Australian Soils (see metadata: 1km resolution terrain and substrate layers for continental analysis of biodiversity pattern). A continuous soil-water retention function (for details see Walker and Langridge 1996) allowed spatial variation in soil water holding capacity and water extraction to be incorporated. For simplicity, we used a constant soil water retention function scaled between field capacity and wilting point. The water balance model takes monthly inputs of rainfall and evaporation, initialised with soil water holding capacities at half their maximum levels and equilibrated over two annual cycles after which the monthly outputs—soil moisture content (volumetric water content in mm), surplus water (mm) and water deficit (mm)—are retained. The soil moisture content is converted into physiological units of water potential (varying between field capacity at 0 and wilting point at -1.5 MPa or 15 bars). Indices of actual evapotranspiration to potential evapotranspiration (PWAT) or to pan evaporation (EAEO) were also generated for consistency with the alternative published calculations (Hackett 1988; Specht and Jones 1971). The effectiveness of one measure over the other depends on how well seasonal variation in canopy resistance to water loss is estimated. In this case a constant coefficient of potential evapotranspiration was used (PETCF=0.9), which assumes canopy leaf areas are seasonally constant. Table 5. Definition of water balance indices. Label WPOT Description Soil water potential WDEF Soil water deficit SPLS Soil water surplus EAEO PWAT Units Definition MPa The monthly soil volumetric water content in units of pressure potential between field capacity (0 bars) and wilting point (-15 bars / -1.5 MPa), derived from a model of water balance. Mm The monthly residual evaporative demand that is in excess of soil moisture at wilting point (-15 Bars) including rainfall, derived from a model of water balance. Very negative values represent a particularly marked soil water deficit Mm The monthly precipitation that is in excess of maximum soil water holding capacity including evaporative demand, derived from a model of water balance. Crop factor dimensionless Water Stress Index % Trends in Seasonal Climatic Variation The monthly ratio of actual evapotranspiration to potential (pan, freewater surface) evaporation. Adapted from the index by Specht and Jones (1971). Actual evapotranspiration is an output of a water balance model, and potential evaporation is an input (ie not adjusted by the coefficient of potential evapotranspiration, see PWAT). This ratio represents a water stress index and has been termed the ‘crop factor’ (Doorenbos and Pruitt 1975). The monthly water stress is the ratio of actual to potential evapotranspiration expressed as a percentage. High water stress occurs when values are low or zero, low water stress occurs with higher values. Adapted from the index by (Hackett 1988). Actual evapotranspiration is an output of a water balance model, and potential evapotranspiration is the pan evaporation (free-water surface) adjusted by the coefficient of potential evapotranspiration (PETCF). A constant PETCF value of 0.9 was used for all months which assumes constant leaf area index. Many species respond to the regular progression in seasons utilising environmental cues to trigger particular phenological responses that result in a productivity advantage (growth, reproduction) or to avoid extreme conditions (drought, frost). For example, many tree species are able to adjust, through acclimation, their optimum temperature for photosynthesis with changes in environmental temperatures with a lag of days to weeks (Hikosaka et al. 2006). Rapid rates of change in environmental temperature, for example inland regions versus coastal regions, may limit this acclimation response and trigger frosttolerance responses such as dormancy instead. Different species therefore occupy different environments related to their adaptive capacity through various degrees of specialised phenology responses to environmental cues. While environmental differences may appear marginal, phenological phenomena are often very sensitive to small variations in climate, especially temperature which exhibits less inter-annual variability than rainfall. Phenology has therefore become of particular interest as an indicator of climate change (e.g., Edwards and Richardson 2004; Walther et al. 2005). To capture some of the climatic cues potentially correlated with phenology responses, we generated variables for seasonal rates of change (trends) in temperature and rainfall as the difference between months in a forward progression of the seasons (i.e., January values minus December values, February values minus January values, etc). Intra-annual change in rainfall patterns were standardised by the number of days in an average month. Table 6. Definition of seasonal climatic indices. Label Description RTI Change in seasonally varying minimum temperatures RTX Change in seasonally varying maximum temperatures RPREC Change in seasonally varying rainfall Units Definition °C/day The monthly average daily difference in minimum temperatures between successive months (i.e., January values minus December values, February values minus January values, etc), representing increments of change in the seasonal progression of overnight temperatures. °C/day The monthly average daily difference in maximum temperatures between successive months (i.e., January values minus December values, February values minus January values, etc), representing increments of change in the seasonal progression of daytime temperatures. mm/day The monthly average daily difference in rainfall between successive months (i.e., January values minus December values, February values minus January values, etc), representing increments of change in the seasonal progression of rainfall, standardised by the average number of days in a month. Rainfall Seasonality Rainfall across Australia is strongly seasonal and a significant determinant of biodiversity patterns influencing the timing of faunal reproduction and the balance between annual and perennial life cycle strategies among plants. A winterdominated rainfall regime occurs in the south and a summer-dominated regime in the north. Uniform rainfall occurs across much of New South Wales, parts of eastern Victoria and southern Tasmania. The monsoonal rains of the extreme north fall between November and March and are often unreliable. More than 80% of the continent has at least three months each year that are without effective precipitation, resulting in drought conditions when this occurs during periods of high temperature. In previous studies of plant distributions in north-eastern NSW, an index of rainfall seasonality was used with considerable success (Austin 1998). This index was derived as the ratio of summer to winter rainfall where summer is defined as the total rainfall over the three months December to February and winter as the total rainfall June to August, beginning with the solstice months. For continental Australia, we derived a composite rainfall seasonality index to emphasise the north-south gradient in conditions from summer to winterdominated rainfall, and applied the same framework in developing an index of intra-seasonal variability beginning with the equinox months (spring-autumn dominated rain events) (Equation 1). For comparison, we also generated an index of annual rainfall seasonality based on the 6 warmest months (October to March) and 6 coolest months (April to September) and the simpler form of these indices as previously used in the NSW studies (Austin 1998). Because summer rainfall volumes are often many magnitudes greater than winter rainfall volumes, two forms of the index were developed, one of which used the logarithm of rainfall. Preliminary analyses indicate that the two seasonal indices (SRAIN1 and SRAIN2) were the most effective predictors (Williams et al. 2010a). Equation 1: Derivation of rainfall seasonality indices from 12 months of total rainfall. Alternative form of equation does not use the logarithm of rainfall. Warm season rainfall (RONDJFM) is the total of rainfall in the six months from October to March Cool season rainfall (RAMJJAS) is the total of rainfall in the six months from April to September Summer rainfall (RDJF) is the total of rainfall in December, January and February Winter rainfall (RJJA) is the total of rainfall in June, July and August Spring rainfall (RSON) is the total of rainfall in September, October and November Autumn rainfall (RMAM) is the total of rainfall in March, April and May Seasonality of rainfall (summer-winter) (SRSW) (SLRAIN1): log ( R + 1) SRSW = + 10 DJF − 1 ; else, log ( R + 1 ) 10 JJA log10 ( RJJA + 1) = − +1 log10 ( RDJF + 1) if, RDJF ≥ RJJA, then: SRSW Seasonality of rainfall (spring-autumn) (SRSA) (SLRAIN2): if, RSON ≥ RMAM, then: log ( R SRSA = − 10 MAM log10 ( RSON log10 ( RSON + 1) SRSA = + − 1 ; else, log10 ( RMAM + 1) + 1) +1 + 1) Annual seasonality of rainfall (warm-cool) (SRWC) (SLRAIN0): if, RONDJFM ≥ RAMJJAS, then: log ( R + 1) SRWC = + 10 ONDJFM − 1 ; else, log10 ( R AMJJAS + 1) log ( R + 1) SRWC = − 10 AMJJAS +1 log10 RONDJFM + 1) Alternative form does not use logarithm of rainfall (SRAIN1, SRAIN2, SRAIN0) Table 7. Definition of rainfall seasonality indices. Label SLRAIN0 SRAIN0 SRAIN0MP Description annual (log) rainfall seasonality index annual rainfall seasonality index annual rainfall seasonality ratio SLRAIN1 summer or winter (log) rainfall season SRAIN1 summer or winter rainfall season Units Definition Dimensionless Annual rainfall seasonality is an index derived from two ratios of precipitation. The ratio of warm (Oct-Nov-Dec-Jan-FebMar) to cool (Apr-May-Jun-Jul-Aug-Sep) season log-rainfall totals (minus 1) are assigned positive values when rainfall in the warm season is greater than rainfall in the cool season. The ratio of cool to warm season log-rainfall totals (plus 1) are assigned negative values when rainfall in the cool season is greater than rainfall in the warm season. Dimensionless Annual rainfall seasonality is an index derived from two ratios of precipitation. The ratio of warm (Oct-Nov-Dec-Jan-FebMar) to cool (Apr-May-Jun-Jul-Aug-Sep) season rainfall totals (minus 1) are assigned positive values when rainfall in the warm season is greater than rainfall in the cool season. The ratio of cool to warm season rainfall totals (plus 1) are assigned negative values when rainfall in the cool season is greater than rainfall in the warm season. Dimensionless Annual rainfall seasonality is the ratio of warm to cool season precipitation, where warm season precipitation is defined as the sum of Oct-Nov-Dec-Jan-Feb-Mar precipitation and cool season precipitation is defined as the sum of Apr-May-Jun-JulAug-Sep precipitation (based on Austin 1998). Dimensionless Summer or winter rainfall seasonality is an index derived from two ratios. The ratio of summer (Dec-Jan-Feb) to winter (Jun-JulAug) log-rainfall totals (minus 1) are assigned positive values when rainfall in summer is greater than rainfall in winter. The ratio of winter to summer log-rainfall totals (plus 1) are assigned negative values when rainfall in the winter is greater than rainfall in summer. Dimensionless Summer or winter rainfall seasonality is an index derived from two ratios. The ratio of summer (Dec-Jan-Feb) to winter (Jun-JulAug) rainfall totals (minus 1) are assigned positive values when rainfall in summer is greater than rainfall in winter. The ratio of Label Description SRAIN1MP Solstice rainfall seasonality ratio SLRAIN1 SRAIN1 SRAIN2MP Spring or autumn (log) rainfall season Spring or autumn rainfall season Equinox rainfall seasonality ratio Units Definition winter to summer rainfall totals (plus 1) are assigned negative values when rainfall in the winter is greater than rainfall in summer. Dimensionless Solstice rainfall seasonality is the ratio summer to winter precipitation, where summer precipitation is defined as the sum of Dec-Jan-Feb precipitation and winter precipitation is defined as the sum of Jun-Jul-Aug precipitation (Austin 1998) Dimensionless Spring or autumn rainfall seasonality is an index derived from two ratios. The ratio of spring (Sep-Oct-Nov) to autumn (Mar-AprMay) log-rainfall totals (minus 1) are assigned positive values when rainfall in spring is greater than rainfall in autumn. The ratio of autumn to spring log-rainfall totals (plus 1) are assigned negative values when rainfall in the autumn is greater than rainfall in spring. Dimensionless Spring or autumn rainfall seasonality is an index derived from two ratios. The ratio of spring (Sep-Oct-Nov) to autumn (Mar-AprMay) rainfall totals (minus 1) are assigned positive values when rainfall in spring is greater than rainfall in autumn. The ratio of autumn to spring rainfall totals (plus 1) are assigned negative values when rainfall in the autumn is greater than rainfall in spring. Dimensionless Equinox rainfall seasonality is the ratio spring to autumn precipitation, where spring precipitation is defined as the sum of Sep-Oct-Nov precipitation and autumn precipitation is defined as the sum of MarApr-May precipitation (based on Austin 1998) Inter-Annual Climatic Variability (temperature) The seasonality, growing season lengths, climatic extremes and stochastic (interannual variability) nature of rainfall and temperature are strong determinants of breeding patterns and reproductive success among Australian fauna and flora, especially in arid and Mediterranean regions (e.g., Lamont et al. 2007). Although biologists have long-recognised the importance of unpredictability in rainfall on the patterns of distribution of biota and migration events, there have been few attempts to develop predictors using the historical climatic data (although see Zimmermann et al. 2009). Historical climatic data record temporal patterns in the intensity and duration of extreme events linked to gradients in productivity or stress across continental Australia. Indices of average extreme cold or hot conditions were derived from the 5km gridded values of daily climate over 50 years (c.1955-2005) (Jeffrey et al. 2001) resampled to 1km using the CUBIC option with RESAMPLE in ARC/Info GRID. The temperatures of the hottest or coldest day per month were averaged over 50 years to define the absolute mean monthly maximum or minimum temperatures respectively. This is a topic requiring further research and analysis that is currently under development (M. Hutchinson and J. Kesteven in prep.). Table 8. Definition inter-annual temperature variability indices. Label Description TMAXABS mean absolute maximum temperature TMINABS mean absolute minimum temperature Units Definition ºC The monthly mean absolute maximum temperature derived from the hottest day of each month over 50-years (1955 to 2005) of 5km gridded daily climate (Jeffrey et al. 2001) ºC The monthly mean absolute minimum temperature derived from the coldest day of each month over 50-years (1955 to 2005) of 5km gridded daily climate (Jeffrey et al. 2001) Dataset Naming Conventions The following attributes were compiled from the monthly climatic variables and are represented as individual raster datasets. Derivation method, units and variable definitions are given in the lineage, above. In each case, the mean, minimum and maximum values of the 12 monthly outputs were compiled for GDM analysis. Dataset labels were postfixed ‘I’ or ‘MIN’ for minimum, ‘X’ or ‘MAX’ for maximum, and ‘M’ or ‘MEAN’ for the average where relevant, except for growth indices (only the mean annual statistic was derived). Other statistics of the 12 monthly variables were also computed – standard deviation and coefficient of variation – but are not included here. For modelling biodiversity patterns, minimum and maximum statistics are recommended. Annual mean statistics are often highly correlated with monthly maximum values and as the integral of the mean monthly variation throughout a year often result in complex spatial patterns. Many related statistics, for example water balance and atmospheric water deficit, will be highly correlated. Decisions about choice of variables to use in a model need to be made according to knowledge of the biology or ecology of the target species, hypotheses of drivers of distribution patterns, and by taking into account multicollinearity among environmental variables. Table 9. Raster dataset names and descriptive labels for climatic attributes. Attribute name (raster dataset name) C4GI MEGAGI MESOGI MICROGI RHU215_I RHU215_X RHU215_M RH2X RH2I RH2M Descriptive labels Mean annual growth index C4 megatherm plants Mean annual growth index C3 macrotherm plants Mean annual growth index C3 mesotherm plants Mean annual growth index C3 microtherm plants Minimum month relative humidity at 3pm (%) Maximum month relative humidity at 3pm (%) Mean annual relative humidity at 3pm (%) Maximum month relative humidity (%) Minimum month relative humidity (%) Mean annual relative humidity (%) Attribute name (raster dataset name) VPD29_I VPD29_X VPD29_M VPD2X VPD2I VPD2M ADEFI ADEFX ADEFM ARID_X ARID_I ARID_M EVAPI EVAPX EVAPM RAINI RAINX RAINM RPRECX RPRECI SLRAIN0 SRAIN0 SRAIN0MP SLRAIN1 SRAIN1 SRAIN1MP SLRAIN2 SRAIN2 SRAIN2MP EAEO_X EAEO_I EAEO_M PWAT_X PWAT_I PWAT_M SPLS_X SPLS_I SPLS_M WDEF_X WDEF_I WDEF_M WPOT_X WPOT_I WPOT_N RADNI RADNX RADNM Descriptive labels Minimum month vapour pressure deficit at 9am (KPa) Maximum month vapour pressure deficit at 9am (KPa) Mean annual vapour pressure deficit at 9am (KPa) Maximum month vapour pressure deficit (KPa) Minimum month vapour pressure deficit (KPa) Mean annual vapour pressure deficit (KPa) Maximum month precipitation deficit (mm) Minimum month precipitation deficit (mm) Mean annual precipitation deficit (mm) Maximum month aridity index Minimum month aridity index Mean annual aridity index Minimum month evaporation (mm) Maximum month evaporation (mm) Mean annual evaporation (mm) Precipitation of the driest month (mm) Precipitation of the wettest month (mm) Mean annual rainfall (mm) Greatest rainfall difference between successive months (mm/day) Least rainfall difference between successive months (mm/day) annual (log) rainfall seasonality index annual rainfall seasonality index annual rainfall seasonality ratio summer or winter (log) rainfall season summer or winter rainfall season Solstice rainfall seasonality ratio Spring or autumn (log) rainfall season Spring or autumn rainfall season Equinox rainfall seasonality ratio Maximum month crop factor Minimum month crop factor Mean annual crop factor Maximum month soil water stress index (%) Minimum month soil water stress index (%) Mean annual soil water stress index (%) Maximum month soil water surplus (mm) Minimum month soil water surplus (mm) Mean annual soil water surplus (mm) Maximum month soil water deficit (mm) Minimum month soil water deficit (mm) Minimum month soil water deficit (mm) Maximum month soil water potential (MPa) Minimum month soil water potential (MPa) Mean annual soil water potential (MPa) Minimum month rainfall-modified solar radiation (MJ/m2/day) Maximum month rainfall-modified solar radiation (MJ/m2/day) Mean annual rainfall-modified solar radiation (MJ/m2/day) Attribute name (raster dataset name) MAXTI MAXTX MAXTM MINTI MINTX MINTM RTI_X RTI_I RTX_X RTX_I TMAXABSM TMAXABSX TMINABSI TMINABSM TRNGA TRNGI TRNGX TRNGM WINDRI WINDRX WINDRM WINDSP9X WINDSP9I WINDSP9M WINDSP15X WINDSP15I WINDSP15M WINDSPI WINDSPX WINDSPM DL_P0_I DL_P0_X DL_P0_M Descriptive labels Maximum temperature coolest month (°C) Maximum temperature hottest month (°C) Mean annual maximum temperature (°C) Minimum temperature coldest month (°C) Minimum temperature warmest month (°C) Mean annual minimum temperature (°C) Maximum difference in minimum temperatures (°C/day) Minimum difference in minimum temperatures (°C/day) Maximum difference in maximum temperatures (°C/day) minimum difference in maximum temperatures (°C/day) Mean annual absolute mean maximum temperature (°C) Maximum month absolute mean maximum temperature (°C) Minimum month absolute mean minimum temperature (°C) Mean annual absolute mean minimum temperature (°C) Annual Range Temperature (°C) Minimum month diurnal temperature range (°C) Maximum month diurnal temperature range (°C) Mean annual diurnal temperature range (°C) Minimum month of wind run (km/day) Maximum month of wind run (km/day) Mean annual wind run (km/day) Maximum month of wind speed at 9am (m/s, 9am) Minimum month of wind speed at 9am (m/s, 9am) Mean annual wind speed at 9am (m/s, 9am) Maximum month of wind speed at 3pm (m/s, 3pm) Minimum month of wind speed at 3pm (m/s, 3pm) Mean annual wind speed at 3pm (m/s, 3pm) Minimum month of wind speed at 9am or 3pm (m/s) Maximum month of wind speed at 9am or 3pm (m/s) Mean annual wind speed at 9am or 3pm (m/s) Shortest day length (hours) Longest day length (hours) Mean annual day length (hours) Accuracy Assessment of the accuracy of version 3 of the DEM is still underway but is likely to be at least as good as version 2. The standard elevation error of version 2 was found to vary between about 7.5 and 20m but may be as high as 200m in very steep or complex terrain (Hutchinson et al. 2001). Climate estimates are derived from long term monthly mean climate surfaces (Australian Climate Surfaces version 2.1) through the ESOCLIM module of ANUCLIM. Rainfall and radiation surface estimates have a relative standard error of about 10%. Maximum and minimum temperature surface estimates have a standard error of 0.2 to 0.4 °C (Hutchinson et al. 2000). CONTACT INFORMATION Contact Organisation (s) CSIRO Sustainable Ecosystems Contact Position Dr Kristen J Williams, Research Scientist Mail Address GPO Box 1700 Locality Canberra State ACT Country AUSTRALIA Postcode 2601 Telephone 02 6246 4213 Facsimile n/a Electronic Mail Address [email protected] Metadata Date 15-07-2010 DATA TYPE Spatial representation type RASTER PROJECTION Map projection GEOGRAPHIC Datum WGS84 Map units DECIMAL DEGREES SCALE Scale/ resolution 0.01 degrees Use Limitation For internal use within DEWHA, Atlas of Living Australia spatial portal, ANU Fenner School and CSIRO only. Not to be distributed. Acknowledgement required. Acknowledgements These data build upon an original set of National environmental variables and metadata compiled by Dr. Janet Stein of the ANU Fenner School (Stein 2008). Monthly climatic variables from ANUCLIM were generated by Tom Harwood of CSIRO Entomology with technical support provided by Tingbou Xu (ANU Fenner School). 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