Journal of Archaeological Science 39 (2012) 2272e2279 Contents lists available at SciVerse ScienceDirect Journal of Archaeological Science journal homepage: http://www.elsevier.com/locate/jas Water availability and landuse during the Upper and Epipaleolithic in southwestern Syria Knut Bretzke a, *, Philipp Drechsler a, Nicholas J. Conard a, b a b Eberhard Karls Universität Tübingen, Institut für Ur- und Frühgeschichte, Abt. Ältere Urgeschichte und Quartärökologie, Burgsteige 11, 72070 Tübingen, Germany Senckenberg Center for Human Evolution and Paleoecology, Universität Tübingen, Schloss Hohentübingen, 72070 Tübingen, Germany a r t i c l e i n f o a b s t r a c t Article history: Received 31 October 2011 Received in revised form 27 February 2012 Accepted 28 February 2012 Paleolithic research often assumes that environmental conditions played a major role in shaping human evolution. To study this relationship we present a spatially explicit approach based on the assumption that the distribution of water within the landscape is an essential component of local environmental conditions. Here, we analyze the relation of wetness and human landuse patterns from the Upper Paleolithic (UP) and Epipaleolithic (EP) of Western Syria. In particular the spatially explicit character of the approach enables the detection of a significant change in landuse patterns during the UP and EP accompanied by a significant shift in the wetness characteristics of the preferentially used areas. These results are discussed against the background of published data on climatic conditions in order to identify both a possible time frame and triggers for this change in landuse. While we conclude an increased influence of natural conditions on the spatial behavior for the UP, we suggest an additional influence of cultural circumstances in shaping EP spatial behavior. For the region studied we argue that the bounded pattern observed during the UP changes to a spatially flexible pattern during the Late Natufian. Ó 2012 Elsevier Ltd. All rights reserved. Keywords: Spatial analysis Human landuse TWI PAW Paleolithic 1. Introduction Paleolithic research often places a high level of importance on the environment’s role in determining Paleolithic populations’ evolutionary and cultural development (Banks et al., 2008; Binford, 2001; Collard and Foley, 2002; Foley, 1995). A variety of studies has been published using a broad range of approaches. The most common approach analyzes archaeological site distributions with respect to environmental variability (Bretzke, 2008; d’Errico and Sanchez-Goni, 2003; Gamble et al., 2004). In addition, archaeologists increasingly realize the potential of GIS based models for studying the relationship between humans and their inhabited environment (Banks et al., 2008, 2006; Burke et al., 2008). Regardless of the approach chosen, one shortcoming is the disagreement between the spatial resolution of the archaeological and climatic data. While archaeological finds can be located precisely using GPS or high resolution maps, paleo-environmental information taken from climate proxies or climatic simulations usually considers average developments over large areas and therefore yields low spatial resolution. Hence, available climatic data often underestimates local variation, a factor which is particularly important for the distribution of vegetation that forms the basis for animal and human life. Especially in arid regions, the vegetation pattern is closely bound to the availability of water. Therefore, knowledge about the local distribution of water in the landscape is a critical prerequisite for the assessment of humaneenvironment interactions. Here, we describe a method that transforms low resolution precipitation data into high resolution surface-wetness distribution data at a regional scale. We assume water availability as a critical factor shaping the spatial distribution of plants and animals, and therefore human biotic resources. Then we compare the wetness distribution with landuse patterns from the Upper Paleolithic (UP) and Epipaleolithic (EP) of Western Syria to characterize the relationship between human activities and the wetness pattern in the landscape as an indicator of the environment’s role in the organization of human spatial behavior. 2. Materials and methods 2.1. Modeling the spatial distribution of wetness * Corresponding author. Eberhard Karls Universität Tübingen, Department of Early Prehistory and Quaternary Ecology, Burgsteige 11, 72070 Tübingen, Germany. Tel.: þ49 69 75421549; fax: þ49 7071 295714. E-mail address: [email protected] (K. Bretzke). 0305-4403/$ e see front matter Ó 2012 Elsevier Ltd. All rights reserved. doi:10.1016/j.jas.2012.02.033 The availability of surface and subsurface water at a specific point in the landscape is primarily a function of the amount and distribution of precipitation and local topography. Information K. Bretzke et al. / Journal of Archaeological Science 39 (2012) 2272e2279 about the amount and distribution of precipitation can be either obtained from isohyet maps or calculated from climatic station data. The most complete information about local topography exists as digital elevation models (DEMs). The spatially continuous information of DEMs about topography can be used to determine the amount of water that potentially flows together at any given point in a landscape. This concept is realized in the ‘Topographic Wetness Index’ (TWI), a well-established secondary topographic attribute introduced by Beven and Kirkby (1979). It assumes that topography controls the movement of water in sloped terrain and hence, the spatial pattern of soil moisture (Schmidt and Persson, 2003). As a consequence, TWI also accounts for water available on the surface. While high TWI corresponds to low lying areas, low TWI values occur in steep, diverging areas. The TWI applies likewise for subsurface water movements driven by terrain gradient (Barling et al., 1994; Sørensen et al., 2005). One of the characteristics of arid and semi-arid regions is that most precipitation does not infiltrate the sedimentary column where it falls, but drains toward lower areas, even where the surface inclination is slight (Schulz, 2000). The greatest concentration of water can be found in the flat areas between hills where sediments and water collect. With these characteristics, the TWI can be used to differentiate potentially moister areas with higher herbal biomass from drier areas with less vegetation. The major drawback for the application of the TWI with respect to the distribution of surface water is the assumption of a homogenous spatial precipitation pattern over the entire region of analysis. While this assumption is valid for some regions of the world, other regions show variable precipitation patterns on a regional scale dependent upon topographic features. To overcome this weakness, we add the spatial variability of precipitation. We achieved this by multiplying a separate data layer containing information about the spatial distribution of precipitation. The resulting formula defines Plant Available Water (PAW) in a specific region: PAW ¼ TWIln PRECpresent (1) where PRECpresent represents the present precipitation. The consideration of a spatially explicit precipitation pattern produces a more realistic picture of the distribution of moisture in the landscape than the TWI alone. 2.2. Modeling past human landuse Since we see the most direct interaction of humans with their environment in the context of gathering and collecting, we consider human activities away from the main living sites. But how can we gain information from this area? One possibility we have explored over the past years (Bretzke, 2008; Bretzke et al., 2011; Conard et al., 2010) necessitates a new way of viewing cultural remains in the landscape. If we change our perspective from identifying sites to the distribution patterns of single artifacts, we access data on human spatial activities across the entire landscape (Dunnell and Dancey, 1983). Our investigations follow an off-site approach, as introduced by Isaac (1981) and Foley (1981), and assume that the accumulation of cultural remains in the landscape is a function of the intensity of its use. Therefore, areas with high artifact densities denote intensively used areas, while low artifact densities indicate locales exploited to a lesser degree. This leads us to conclude that by discerning the spatial distribution of artifact densities, we can posit inferences regarding past human behavior in particular settlement spaces. Information about the patterning of artifacts in the landscape derives from archaeological survey data. The systematic collection 2273 of artifacts and the estimation of artifact densities specific of cultures at all surveyed localities provide a dataset suitable for this analysis. Applying the “Inverse Distance Weighted” interpolation technique (Conolly and Lake, 2006), the density information of artifacts gained from spatially separated localities can be transferred into a spatially continuous dataset that better represents the activity distribution (Bretzke, 2008). 2.3. Case study We apply this approach to archaeological data from the Tübingen-Damascus Excavation and Survey Project (TDASP in German) (Conard, 2006). The TDASP study area in the Damascus Province about 50 km northwest of Damascus (Syria) is situated between the Anti-Lebanon Mountain range and the Syrian Desert. An impressive cliff line formed of resistant Oligocene limestone represents the border between areas of higher elevation, up to 2300 m, and lowland areas down to 800 m (Fig. 1). The study area is further characterized by a steep gradient in precipitation. While substantial precipitation falls along the western faces of the Anti-Lebanon Mountains, precipitation decreases markedly toward the eastern flanks and the hinterland. Enzel et al. (2008) argue that this overall pattern has not changed considerably since the last glacial/interglacial period. Thus, we can use the present precipitation pattern to create a simplified model of local environmental conditions. The dataset on the spatial distribution pattern of present precipitation in the study area was created by a thin-plate spline interpolation of isohyets from a Syrian precipitation map (Syrian Arab Republic, 1977). Finally, data on the topography derives from a conveniently processed DEM that was based on maps from the shuttle radar topography mission (United States Geological Survey, 2007) (Fig. 2). The TDASP study region is a highly differentiated landscape with characteristics that likely encouraged Paleolithic settlement in the region. Lithic raw material of good quality is readily available from primary and secondary deposits in many areas of the study region (Dodonov et al., 2007). Another advantageous characteristic is the occurrence of perennial water sources along the cliff line. Today active artesian springs can be found near the base of the cliff line in vicinity to the modern villages of Ma’aloula, Jaba’deen and Yabroud (Fig. 1). These springs are associated with deeply incised canyons, which not only confirm the existence of water sources at these locations for long periods, but also provided easy passage for people and animals between the lowlands and the highlands. The field work of TDASP builds on previous work in the Damascus Province by Rust (1950), Suzuki and Kobori (1970), Solecki and Solecki (1987/1988), and Bakdach (2000). Before TDASP began working in the Damascus Province, the only stratified Paleolithic assemblages known were those from the sites of Yabrud (Rust, 1950; Solecki and Solecki, 1966). During 12 years of research in the Damascus Province, TDASP conducted excavations in Baaz Rockshelter, a multi-period site with layers from early and late UP and Late Natufian (Conard, 2000; Conard et al., 2006a,b; Deckers et al., 2009), Kaus Kozah Cave, a site containing Middle and Epipaleolithic artifacts (Conard and Masri, 2006), Ain Dabbour, a site with two Geometric Kebaran layers (Hillgruber, 2010), and Wadi Mushkuna, a Middle Paleolithic site currently under excavation (Fig. 1). Since 1999 TDASP has conducted intensive surveys in the Damascus Province that represent a significant record of Paleolithic data in Southwestern Asia (Conard, 2006). Today, the collection contains chipped stone artifacts from 598 areas (Fig. 3). The record includes all major periods from the Lower Paleolithic to the Neolithic (Table 1). Within our survey area of ca. 500 km2, we collected artifacts using pedestrian transects through defined areas 2274 K. Bretzke et al. / Journal of Archaeological Science 39 (2012) 2272e2279 Fig. 1. Geographic setting of the study area and location of sites mentioned in the text. 1) Baaz Rockshelter, 2) Kaus Kozah Cave, 3) Ain Dabbour, 4) Yabroud, 5) Wadi Mushkuna. in the landscape. Over the course of 8 seasons of survey, we were able to sample approximately 15% of the surface, including samples from all major landscape forms and topographic settings. Our decision about the location of areas to sample was guided by the aim of covering the entire area of study. We use the term localities to identify areas where artifacts were collected. At each locality, the survey team documented the presence or absence of every cultural entity and the respective artifact density (Fig. 4). One surveyed locality may contain artifacts from different periods. To use one case study, we limit the presented data to the UP (ca. 40e20 ka) and the EP (ca. 20e10 ka) and consider 181 localities containing UP artifacts and 97 containing EP artifacts (Fig. 4). In 45 localities EP and UP artifacts occur together. 410 localities lack artifacts from both periods and were used as negative evidence. A sub-division of the entities UP and EP into specific archaeological cultures is usually not possible due to the low chronological resolution of the survey finds and the scarcity of chrono-culturally sensitive artifacts at the majority of our survey localities. However, a distinction between the more general entities UP and EP is possible. Guided by our knowledge from the stratified assemblages of Baaz Rockshelter, Kaus Kozah Cave and Ain Dabbour we classified the surface finds into UP or EP using primarily technological characteristics of cores and blades/bladelets (Hillgruber, 2010). Edge damage and patina are additional characteristics used to support the assessments based on technological criteria. The integrity of the artifact pattern identified in the landscape is confirmed by geomorphological studies that suggest stable land surface conditions and only minor dislocation of artifacts in the landscape (Dodonov et al., 2007). Furthermore, in 52 localities EP finds are accompanied by Middle Paleolithic artifacts but lack UP finds. This observation suggests that the absence of UP is not due to taphonomy but rather reflects actual differences in spatial behavior between EP and UP. 3. Results We successfully transformed low resolution precipitation data into a high resolution model of wetness distribution using (1) and readily available topographic data (Fig. 2). The calculated PAW indices represent wetness in the landscape dependent on the degree of local precipitation and topography (Fig. 5). Very high PAW values indicate a high amount of local wetness often in the flat parts Fig. 2. Depiction of data used to model the distribution of wetness. A) Digital elevation model (DEM), derived from SRTM data, dark areas indicate high values, B) Topographic Wetness Index (TWI), calculated from A after Beven and Kirkby (1979), color code see A, C) local precipitation pattern, derived from precipitation data of the Syrian Arab Republic (Syrian Arab Republic, 1977). K. Bretzke et al. / Journal of Archaeological Science 39 (2012) 2272e2279 Fig. 3. Distribution of surveyed localities. A total of 598 localities were sampled to collect data on the spatial behavior of prehistoric populations. One locality may contain cultural remains from different chrono-cultural entities. of the landscape, but also in runoff channels. Areas with increased amounts of water are represented by medium PAW values, whereas dry areas have low PAW values and occur in areas with relatively steep slopes. A comparison of similar terrains from the eastern and western part of the study area indicates higher PAW values in the west. This results from the regional precipitation pattern (Fig. 2C) where higher values in the northwest decrease toward the southeast. To analyze the relation between human groups and the distribution of wetness, we used data from 598 surveyed localities to model UP and EP human spatial behavior. The results show that a spatially compact cluster of UP activity areas gives way to a pattern characterized by a wide distribution of loosely connected activity areas during the succeeding EP (Fig. 6). Hence the modeling of human spatial behavior suggests a profound change in the way the inhabited landscape was used. The reasons for such a change may be manifold, but changes in environmental conditions may have played a major role. We test this hypothesis by analyzing the relationship between landuse patterns and the distribution of wetness as represented by Table 1 TDASP survey record. Summary of sites based on chrono-cultural entity and artifact density. These sites are present at 575 localities documented since 1999. No artifacts were found at 23 surveyed localities, which leads to the total number of 598 survey localities. A locality may contain sites from more than one cultural group. * each of these sites contained only one handaxe. Lower Paleolithic Handaxes Middle Paleolithic (Levallois) Middle Paleolithic (non-Levallois) Upper Paleolithic Upper or Epipaleolithic Epipaleolithic Post-Epipaleolithic Indeterminate Total Low density Medium density High density Total number of sites 30 (37%) 16* (94%) 143 (40%) 36 (44%) 0 160 (45%) 16 (20%) 1 (6%) 51 (14%) 82 17 354 9 (38%) 12 (50%) 3 (13%) 24 65 (36%) 7 (88%) 55 (57%) 11 (41%) 60 (79%) 396 92 (51%) 1 (13%) 33 (34%) 10 (37%) 14 (18%) 358 24 (13%) 0 9 (9%) 6 (22%) 2 (3%) 112 181 8 97 27 76 886 2275 the PAW index. We overlay the wetness distribution raster (Fig. 5) on the UP and EP landuse patterns (Fig. 6) and deduce the wetness characteristics of the activity areas for both periods. The wetness distribution is represented by raster data consisting of cells of 90 by 90 m. The EP landuse pattern covers activity areas of 3020 cells of the wetness raster. These cells have a mean PAW value of 36.6 with a standard deviation of 5.4. In contrast, the UP landuse pattern covers 4585 cells of the wetness raster. Here the mean PAW value is 38.8 with a standard deviation of 4.7. The range of the samples is blurred by few extreme outliers (UP: n ¼ 8; EP: n ¼ 1). To improve the analytical value of the PAW samples, we eliminated these outliers. This results in an UP range of 31.5 and an EP range of 35.5 for all remaining values, indicating an increase of about 13%. Hence we observe an increase in the diversity of the wetness characteristics of the activity areas. The differences between both samples were tested using a ManneWhitney rank sum test. According to the test results a statistically significant difference (p < 0.01) exists between the UP and EP. This marks a pronounced shift in the way humans used the landscape, from the preferential use of wet areas during the UP to the incorporation of dry areas with low PAW values during the EP. This change in the wetness characteristics can be visualized using kernel density estimation (Baxter et al., 1997). We use here an Epanechnikov kernel and a bandwidth of 2 for the PAW values of either entity (Fig. 7). Greater kernel density values indicate a greater frequency of values at respective PAW values. The results show a significant shift toward smaller values from UP to EP. While the peak PAW value in the UP is about 38, the PAW peaks at about 34 for the EP. 4. Discussion Our results indicate a change in landuse patterns from UP to EP that is accompanied by a significant change in the wetness characteristics of the used areas. Within the same landscape, activities of UP peoples were restricted to wet areas, while the inhabitants of the region during the EP expanded their landuse to include dry areas. This leads us to conclude that a significant change in the subsistence strategy occurred during the EP. The widespread, unconnected activity areas deduced from the EP landuse model suggest the evolution of new landuse strategies. In addition to an increased PAW range for these activity areas, this observation points to a habitat expansion and a rise in the resource diversity potentially available to the EP population. Furthermore, significantly lower PAW values of the areas used during the EP, compared to the UP, indicate a degree of independence from wet parts of the landscape. Our conclusions on changes in the subsistence strategy are drawn from the computational models and surface finds. Despite this new approach to the question of landuse and subsistence, our results are in good agreement with results from more traditional approaches using data from archaeological excavations. For example, studies on cultural remains from Baaz Rockshelter indicate an increase in the diversity of both the prey spectrum (Napierala et al., in press-a-b) and the lithic tool assemblages between the UP and the EP layers there (Barth, 2006; Hillgruber, 2010). The good agreement between our survey results and those from Baaz Rockshelter indicates that our approach using PAW as an analytical tool can be implemented as an independent line of evidence and as a possibility to add a new spatial perspective in the analysis of the interaction between people and environments. The TDASP survey documents that artifacts from the EP are consistently found in areas of higher altitude (Conard et al., 2006d; Dodonov et al., 2007). Moreover, they are often located on prominent and often highly exposed elevated points in the landscape. 2276 K. Bretzke et al. / Journal of Archaeological Science 39 (2012) 2272e2279 Fig. 4. Distribution of artifact densities used to model landuse. A) Distribution of artifact densities and negative evidence for the UP; circle size indicates density; white points represent localities without UP artifacts (negative evidence). B) Distribution of artifact densities and negative evidence for the EP; symbols similar to A. Areas of higher altitude are located in the northwestern part of the TDASP study region and characterized by higher precipitation. Despite higher precipitation, the wetness is less concentrated in this part of the landscape due to the lower potential for the accumulation of runoff. Our results show that EP populations were able to use areas with less wetness and were thus potentially able to use areas situated upstream and at higher elevations more systematically than UP populations. Hence it can be concluded that the increased degree of independence from wet areas is an adaptation that allowed the successful expansion of EP spatial activities into elevated areas. Elevated areas in the TDASP study region are mainly located on the eastern flank of the Anti-Lebanon Mountains, a region where wild stands of cereals potentially grew (Conard et al., in press; Deckers et al., 2009). Based on the conclusion that EP populations were able to access elevated areas, we can conclude that EP populations were also able to access this important resource (see Figs. 4B and 6B). Mortars and pestles found in EP contexts at Fig. 5. Distribution of wetness based on the estimation of PAW. Dark areas indicate an increased amount of accumulated surface and subsurface water. White areas are relative dry. Baaz Rockshelter (Conard, 2000), Kaus Kozah Cave (Conard and Masri, 2006) and several survey localities (Conard et al., 2006c) also suggest that the EP inhabitants of the region occasionally processed cereals, in addition to pigments, nuts and perhaps other materials (Hillgruber, 2010). Although there are no direct indications for the regular use of cereals, the evidence from our landuse models points to the likelihood that EP populations in Western Syria may have accessed wild stands of cereals. It remains uncertain during which period of the EP the change in landuse occurred. The EP can be subdivided into the Early (ca. 22,000e16,000 cal BC), the Middle (ca. 16,000e13,100 cal BC) and Late EP (13,100e9600 cal BC) (Goring-Morris et al., 2009). While the Early EP contains the Kebaran, the Geometric Kebaran belongs to the Middle EP. The Late EP is comprised of the Early and Late Natufian. Hillgruber’s (2010) study of EP lithics from Western Syria points to a paucity of evidence for Early Natufian groups in this region. This observation narrows the options for when the shift in landuse took place and suggests that this change in landuse occurred either during the Early/Middle EP or near the start of the Late Natufian. The frequent occurrence of backed artifacts and microliths during the Kebaran and Geometric Kebaran (Hillgruber, 2010) marks a noticeable technological difference between the Early/ Middle EP and the Late UP, where such pieces occur less frequently. Moreover, these artifacts indicate new hunting technologies, which may have dictated changes in landuse. Despite these differences in the material culture, archaeozoological studies suggest that, like their UP predecessors, Kebaran and Geometric Kebaran populations were mobile hunter gatherers relying on a similar spectrum of prey (Banning et al., 2001; Bar-Oz and Dayan, 1999; Stiner, 2005; Stutz et al., 2009). Given the similarities in the economic behavior between the Late UP and the Early/Middle EP, we suspect that the main shift in landuse may postdate the Early/Middle EP. Although the chronological resolution of the survey finds is too low to be certain, we hypothesize that the most significant shift in landuse in TDASP study region occurred with the start of the Late Natufian. An argument for the hypothesized shift in landuse with the advent of the Late Natufian may be reflected in the changing material culture. The archaeological record of the Late Natufian in our study region is characterized by a variety of differences in material cultural compared to the UP and Early/Middle EP. The Late Natufian archaeological record contains evidence of residential structures (Conard, 2000), pestles and mortars (Conard et al., 2006a,b), and a greater diversity of hunted game (Napierala et al., in press-a-b). Since these data suggest multiple differences in K. Bretzke et al. / Journal of Archaeological Science 39 (2012) 2272e2279 2277 Fig. 6. Landuse model for the Upper Paleolithic (A) and the Epipaleolithic (B) based on data from the Tübingen-Damascus Excavation and Survey Project. Black areas indicate areas of increased artifact density. lifeways and subsistence, the Late Natufian seems to be a better candidate for changes in landuse as seen in our results. This is in keeping with many publications on subsistence during the EP, which suggest that a significant change occurred during the Natufian (Baadsgaard et al., 2010; Bar-Yosef, 1998; Gilead, 1988, 1991; Munro, 2003; Stiner, 2001; Stutz et al., 2009). Usually this change is demonstrated for the Early Natufian, but since the Early Natufian is poorly documented in the TDASP study region (Hillgruber, 2010), the Late Natufian is the most plausible candidate in our regional context. We plan to test this hypothesis in the future by conducting detailed studies of the lithic assemblages from the survey localities with the goal of establishing improved chronological control of the history of occupation and landuse in the TDASP survey area. After identifying the probably period in which landuse changed, we can go one step further and ask the next question. What triggered this development toward the use of drier, often elevated, areas? The Late Natufian is often cited as an example of human response to environmental change, which suggests a natural trigger. Indeed, there is ample evidence that the environmental conditions changed dramatically during the Younger Dryas (YD), a dry spell chronologically associated with the Late Natufian (Bar-Matthews et al., 1999; Frumkin et al., 1999). In the southern Levant the YD has often been seen as a trigger for the decline of large scale villages in a semi-sedentary settlement system during the Early Natufian toward smaller and more mobile groups during the Late Natufian (Bar-Yosef, 1987; Bar-Yosef and Valla, 1990). However, new data challenges this model (Deckers et al., 2009; Henry, in press; Lev-Yadun and Weinstein-Evron, 2005; Napierala kernel density 0.10 EP UP 0.08 0.06 0.04 0.02 0 30 35 40 45 50 PAW Fig. 7. Kernel density estimates of the PAW values deduced from the Upper Paleolithic (UP) and Epipaleolithic (EP) landuse models. The shift to lower PAW values during the EP indicates an altered landscape use. et al., in press-a) by suggesting that the environmental change in some parts of the Levant was of small magnitude, if present at all. Our data from the TDASP study region do not document a sharp environmental decline during the YD. Trout remains from the Late Natufian assemblage of layer III in Baaz Rockshelter (Napierala et al., in press-b) suggest a sufficient supply of water to a perennial river system in the vicinity of the site. Furthermore, Deckers et al. (2009) present evidence suggesting that woodland vegetation was not fully established during the preceding Bölling/Alleröd interstadial. Instead it is probable that trees and shrubs were only thinly scattered over the study region. A scenario also concluded for the YD in our study area. The archaeobotanical data suggest a degree of continuity in the environmental conditions between the Early EP and the Late EP and argues against a noticeable impact of the YD, and thus against a natural trigger for the initiation of changes in EP landuse. Indeed, nearly constant environmental condition between the Early EP and the Late EP on the one hand, and a significant change in the landuse on the other, suggest an increased importance of the social-economic behavior, as opposed to natural environmental causality, in shaping patterns of landuse. The landuse pattern deduced for the UP stands in stark contrast to the EP pattern. We interpret the strong relationship between wet areas and UP activities bounded within one part of the landscape as a response of the UP populations to decreased precipitation (Bar-Matthews and Ayalon, 2003). If we consider the importance of water for the development of the biosphere, we can conclude that under a regime of decreased precipitation, in addition to the areas in the immediate vicinity of perennial springs, only those areas where surface and subsurface water accumulates would offer adequate biomass to allow for an archaeologically visible human presence. Such areas serve as magnets, leading to a concentration of activities within a restricted part of the landscape, as visible in the distribution of artifacts from the UP. Although arguing here for natural causes being mainly responsible for the shape of the UP landuse pattern, we do not exclude the possibility that the landuse pattern could also be influenced by the social environment, as discussed by Bretzke (2008) in more detail. The discussion above highlights the potential of a spatially explicit approach to the study of human behavior with respect to environmental characteristics. The strength of this approach is its quantitative character that provides the possibility of adding a spatial perspective to the study of the interaction between humans and their environment. 2278 K. Bretzke et al. / Journal of Archaeological Science 39 (2012) 2272e2279 5. Conclusion This paper demonstrates the potential of spatially explicit studies of human landuse. Based on readily available data on topography and precipitation we introduced an approach to transform low resolution precipitation data into high resolution surface-wetness data to model local variability in ecological conditions. By comparing the wetness distribution pattern with landuse patterns of UP and EP human groups from the Western Syria, we identified significant differences in the use of the landscape between both periods. While landuse during the UP was characterized by a contraction of activities into relatively small, wetter parts of the landscape, EP landuse featured the exploitation of a broader range of areas with different environmental characteristics including dry and elevated parts of the landscape. Furthermore, the spatially explicit and quantitative character of our approach allows us to investigate the response of human landuse to changing ecological conditions. We conclude from our survey results and the discussion of the climatic background that the UP pattern is largely influenced by natural conditions. In contrast, the significant change in landuse, which likely took place during the Late Natufian, cannot entirely be explained by changes in the natural environment. The available climatic data lack evidence for a significant change in the local environmental conditions during the Late Natufian. Hence we assume an additional trigger probably resulting from changes in the social-economic environment. Acknowledgments We are grateful to the Syrian General Directorate of Antiquities and Museums and especially to our colleagues B. Jamous, M. Maqdisi, M. Masri, M. Hamoud and S. Muhesen for providing access to the study area. We thank A.W. Kandel for discussion and proofreading. We are also deeply indebted to A. Dodonov, who made great contributions to the geoarchaeology of the TDASP project. 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