Origin of Life • How did life begin? • What were the earliest organisms like? • Endosymbiosis • Multicellular organisms? • Invasion of land Europa, one of Jupiter’s moons History of Life on Earth How did life begin? • Spontaneous generation refuted • Francisco Redi • Louis Pasteur Early Earth • Sources of organic compounds • Extra-terrestrial sources: meteorites • Contain amino acids, small amount • Concentrated in tide pools Miller & Urey experiment • • • • • • • • Closed system Simulate conditions of early earth as suggested by Oparin & Haldane Sparks for lightning Identified: Amino acids, organic acids, hydrocarbons Further work has synthesized all major groups of molecules. May not have contained necessary gases Deep sea vent and volcanos RNA • RNA: Two properties • Genetic material, self replicating • Enzymes: catalytic function • Ribozymes Monomer to Polymers • Abiotic synthesis on hot clay surface • Spontaneous polymrerization • Protobionts (microspheres) • Agitation of proteins and lipids • Steps appear to be biologically possible and plausible First Cells •Anaerobic prokaryote •Heterotrophic •No nucleus •3.5 billion years old •Photosynthetic bacteria evolved •O2 evolving organisms Endosymbiont Hypothesis • Formation of Eukaryotic cells • 2.1 billion years ago • Aerobic bacterium • Evidence for Mitochondria • DNA • Membranes • Ribosomes • Amoeba with aerobic bacterium Plant Cells Formation of plastids similar to mitochondria Multicellular organisms • Colonial organisms, similar cells • Multicellular algae • Differentiation • Cellular specialization • Division of labor Animal Diversity Squid • Arose in PreCambrian era • ~1 billion years ago • Fossils, 610-540 million years ago • Invertebrates • Major body plans formed at Cambrian period Ammonites Trilobites Nautilus Invasion of the land • Adaptations for life on land: • Gravity • Breathing • Water • Movement • Reproduction Thinking about the chapter • • • • • • • • 1. What is the evidence that life might have originated from nonliving matter on early Earth? What kind of evidence would you like to see before you would accept this hypothesis? . 2. If the first cells with aerobic metabolism were so much more efficient at producing energy, why didn’t they drive cells with only anaerobic metabolism to extinction? . 3. Explain the endosymbiont hypothesis for the origin of chloroplasts and mitochondria. 4. Name two advantages of multicellularity for plants and two advantages for animals. . 5. What advantages and disadvantages would terrestrial existence have had for the first plants to invade the land? For the first land animals?
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