HARDWOOD PLANTATIONS SOUTHERN F. W, ONTARIO VON ALTHEN ONTARIO FOREST SAULT CANADIAN OF LABORATORY MARIE/ INFORMATION DEPARTMENT REGION RESEARCH STE, OF ONTARIO REPORT FORESTRY FISHERIES DECEMBER O-X-2 SERVICE AND FORESTRY 1970 Copies of this reportj which is a reprint of a report dated September 196Sj may be obtained from Director3 Ontario Region, Canada Department of Piisherieu and Forestry3 Box 4903 Sault Ste. Marie3 Ontario. Frontispiece. High-value walnut trees grown in a small plantation near a farmyard. ABSTRACT The development of hardwood plantations discussed. Height and diameter growth of in southern Ontario is 17 tree species is listed and the most interesting plantations are described in some detail. Mean annual height growth of the most commonly sampled species is evaluated in regard to soil texture, moisture, the plantation site. rooting depth and previous use of Recommendations are made for planting site selec tion of the most commonly sampled species. TABLE OF CONTENTS Page INTRODUCTION 1 THE STUDY AREA 2 COLLECTION OF DATA 2 ANALYSIS OF DATA 4 RESULTS 4 . Black Ash White Ash Basswood White Birch Butternut Catalpa Black Cherry Rock Elm 7 7 10 12 12 13 13 14 White Elm Shagbark Hickory 14 16 Black Locust 16 Silver Maple Sugar Maple Burr Oak Red Oak White Oak Carolina Poplar Black Walnut 17 18 23 23 26 27 27 CONCLUSIONS 31 REFERENCES 33 APPENDIX A 34 INTRODUCTION The hardwood forests of southern Ontario are among the most potentially productive, most accessible, and most poorly managed forests of Ontario. For over 100 years they have been a source of high quality timber and many local industries are directly dependent upon them as a source of raw material. Over two-thirds of the forest area is held in private ownership, mostly in the form of small woodlots, and this has made it impossible to regulate the cut on a sustained yield basis. Cutting has usually been done on a high grading basis in which a con tractor obtains the right to cut all usable trees regardless of their maturity or value in the future development of the stand. Many woodlots have been logged three or four times and are now grossly under stocked or contain traes of no commercial value (Fingland et at. 1964). At least 50 percent of all woodlots are permanently or occasionally pastured by cattle which destroy natural regeneration and cause exces sive soil compaction. Awareness of the growing scarcity of good quality timber and the interest of individual land owners in the creation and maintenance of productive hardwood forests, combined with the government policy of supplying planting stock at a nominal charge, has been responsible for planting of millions of hardwood seedlings during the last four decades. From 1936 to 1951, the last year for which detailed figures have been published, an average of two million hardwood seedlings or cuttings have been distributed annually. The apparent failure of many hardwood plantations, compared to the usually more successful coniferous plantings, has resulted in a curtailment of hardwood planting stock production in the provincial nurseries and relatively few new plantations have been established during the last 10 years. Although it is generally acknowledged that good hardwood plan tations are more difficult to produce than those of coniferous some very successful plantations do exist. However, species, little information has been published on the actual performance of different hardwood species planted on various sites in southern Ontario. A hardwood plan tation survey was therefore carried out in 1963 and 1964; summarizes the most important findings, this report provides a detailed description of the more interesting plantations, and makes recommendations for planting site selection of the most commonly planted species. The author is a Research Scientist, Sault Ste. Marie, Ontario. Forest Research Laboratory, THE STUDY AREA The area of study covered by this report is that part of Ontario that lies south of a line connecting the south shore of Georgian Bay with the eastern end of Lake Ontario. It includes one of the most heav ily populated and industrialized areas in Canada. Intensive agriculture prevails wherever the land is suitable for farming, and timber stands generally consist of small- to medium-sized woodlots owned individually. The climate of this region is greatly influenced by the Great Lakes and is generally milder than the northern or eastern parts of the province. Average and extreme temperatures and precipitation of the study area are presented in Table 1. Landform consists of smooth lacustrine plains in the west, changing eastward to undulating bevelled till plains or mor aines with lacustrine deposits between the ridges. The main regional soil types are grey-brown podzolics, dark grey gleisolics, or brown podzolics (Hills 1959). Nearly all of the commercially important hardwood trees of Canada are found in the region and many species reach their northern botanical limit here (Hosie 1969). COLLECTION OF DATA A check of the records of planting stock shipments in district offices of the Ontario Department of Lands and Forests supplied the addresses of plantation owners, the location of proposed plantings, and the number of seedlings shipped. Only orders of at least 3,000 hardwood seedlings shipped in any one year or 5,000 seedlings in aggregate were included in the lists. During the survey additional plantations were found as a result of information received from foresters familiar with their districts or from private landowners who had obtained seedlings from commercial nurseries or had used wildlings from their woodlots. A total of 296 locations were visited, each plantation was in spected and the current owner questioned in regard to plantation history, former land use, planting site preparation, planting method, cultivation, and silvicultural treatments. If sufficient information could be obtained from the current or former owners, and survival and growth were deemed sufficient to warrant an intensive investigation, one or more samples were taken. Each sample consisted of a tally of not less than 10 dominant and co-dominant trees per species. Height of all sample trees was measured by Abney level to the nearest foot and diameter at breast height was measured by tape to the nearest 1/10 inch. Survival was estimated to the nearest 10 percent, whereas stand quality was evaluated in regard to stem straightness, relative branchiness, and size of crowns. At least one soil pit was dug in each plantation and the profile was described together with notes on land form, aspect, minor vegetation, or any observations of special interest. Of the 296 properties visited, 68 plantations were selected for intensive study and 190 samples were recorded. Table 2 lists by numbers Table 1 Regional climates of southern Ontario (Putnam and Chapman 1938) Climatic region Altitude (ft) Mean annual temperature (°F) Mean winter temperature (°F) Mean spring temperature (°F) Mean summer temperature (°F) Mean fall temperature (°F) Extreme low temperature (°F) Extreme high temperature (°F) Daily range of temperature (°F) Avg date of last frost in spring Avg date of first frost in fall Avg length of frost-free period (days) Beginning of growing season End of growing season Avg length of growing season Avg annual precipitation (days) (in.) Avg annual snowfall (in.) Avg rainfall April 1 - Sept 30 (in.) Avg summer rainfall (June, July, Aug) P-E index (June, (in.) July, Aug) Frequency of droughts Percent possible sunshine in growing season and percentages the reasons for rejection of 228 properties. Table 2 Reasons for rejection of plantations from survey Reason for rejection No. Failure* 86 38 21 I 13 5 5 ° ~ 228 «>0 Ownership changed, present owner uninformed Plantation destroyed by real estate and road 62 building Trees used in landscape planting Too young to evaluate 21 15 Planted in natural woodlot and impossible to identify Erosion control planting Miscellaneous Total Percent Li 9 ' * Most frequently due to planting on unsuitable sites and animal browsing ANALYSIS OF DATA For each species in each sample, average height and diameter at breast height were computed as well as mean annual height growth from seed. If not stated otherwise, average height and diameter mentioned in this report refer to dominant and co-dominant trees only. The influence of planting method, cultivation, or early stand treatments could not be evaluated statistically since all information was based on the recollec tion of work carried out 20 to 50 years ago. Because of this a descrip tion of the growth performance of individual species in various planta tions was chosen to emphasize the factors that appear to be most respon sible for the variations in plantation development. Where considered use ful, comparisons between species were made towards a better understanding of the interactions of factors within the same stands. For the most commonly sampled species, a subjective evaluation of plantation success or failure was made, taking into account survival, growth rate, form development, and high grade timber production. RESULTS Success or failure of hardwood plantations must be evaluated by different standards than those applied to coniferous plantations, since quality of the individual hardwood tree is of paramount importance. A successful hardwood plantation must nor only have satisfactory survival and growth, but quality and form must make the trees acceptable for either sawlogs or veneer bolts since no market currently exists in southern Ontario tor small or poorly shaped trees. The only notable exception is black locust which is valued for fence posts. In nearly all hardwood plantations sampled, height growth varied widely within the same plantation and often changed abruptly from one small area to another. Investigation of the plantation soils nearly al compaction, or moisture ways showed that a change in soil texture, depth, regime had affected height growth. For example, a trench dug through an area occupied by a group of good red oak trees, which were the lone sur vivors of a large plantation, showed a pocket of deep sandy soil under the good oak, changing to a shallow, stony soil over compact clay in the area of failure. Selection of a suitable planting site must therefore be regarded as the most important factor in the establishment of success ful hardwood plantations (von Althen 1964). Suitable planting sites are generally found in natural woodlots with deep fertile, undisturbed soil, in farm orchards, abandoned vege table gardens, or other highly fertile areas such as moist bottomland along streams and creeks and lower slopes with deep, fresh soil. Areas unsuitable for hardwood planting include abandoned agri cultural and pasture land of low fertility, dry, exposed ridges, or any land with a shallow topsoil and a compacted or stony sub-soil. As a rule any land considered to be too poor for agriculture is also too poor for good hardwood growth. Site preparation and cultivation are necessary prerequisites for successful hardwood plantation establishment because any prolonged period of slow growth during the early life of the plantation greatly increases the danger of seedling loss or permanent injury owing to suppression by weeds, rabbit browsing, or ground frosts occurring after leaf flushing. Site preparation may be in the form of complete or partial cultivation or chemical eradication of competing vegetation. Plowing and tilling of the total plantation area is by far the best method of site preparation because it offers tages not obtainable by any other method: the deep-rooted perennial species; it loosens improved aeration and water infiltration; the following advan it destroys all weeds including the soil and provides and it adds plant material which improves the nutrient status and organic content of the soil. Complete mechanical site preparation also provides favourable conditions the most for subsequent chemical weed control because regrowth of weeds can generally be prevented by applying herbicides dosages small enough to be tolerated by seedlings. Glossary of scientific names of in the highly susceptible hardwood species appears in the appendix. Where plowing of the total area is not feasible, strips several feet wide should be plowed and the trees planted in the overturned soil. On light-textured soils or those with few weeds, site preparation may take the form of chemical eradication of weeds on the total area or in strips at least 5 feet wide. However, these methods are generally much less effective in promoting seedling growth than complete mechanical site preparation. Weed control during the first few years after planting depends greatly on the intensity of site preparation with the most intensively prepared sites requiring the least weed control. The additional cost of intensive site preparation may, therefore, be partly or totally offset by reduced weeding costs. Manual and mechanical weed control methods have proven very successful but the frequency of operations required for effec tive control makes these methods rather expensive. Herbicides offer more economical control, but many hardwood species are highly susceptible to damage by the dosages necessary for effective control. Tree girdling by mice and browsing by rabbits can cause heavy damage in hardwood plantations. Poisoning of the mice and application of rabbit repellents can provide some protection but these methods are rather expensive for the degree of protection they provide. By far the best protection against rodent damage is the elimination of weeds. prives the animals of shelter and food and makes This de the plantation a hostile environment for rodent survival and reproduction. Deer browsing damage has been unimportant in southern Ontario owing to the relatively small number of deer in the area. However, in regions with a dense deer population, serious consideration must be given to this problem. It should be obvious to every plantation owner that cattle should never be allowed to graze in a hardwood plantation since they not only damage the trees by browsing, but also break or bend many saplings and severely compact the soil. Where protection from livestock and wild life cannot be guaranteed, hardwood planting should not be attempted. Many of the sampled plantations had been established at a very close spacing. stock, The reasons for dense planting were smallness of planting fear of low survival, and the desire for early canopy closure to eliminate competition from herbaceous vegetation. Not only had the es tablishment of such plantations been very costly, but high survival had frequently resulted in extremely dense stands and various degrees of stagnation. Thinning at an early age could have avoided stagnation and the resulting loss in diameter increment, but very few plantations had received any silvicultural treatments. Changes in ownership were found to be a major reason for the sometimes rather obvious neglect, but lack of knowledge of growth requirements was also widespread. Similar conditions prevailed in some mixed plantations where a less valuable tree species, such as black locust, was allowed to dominate the more valuable black walnut or white ash trees. In other mixtures consisting of alternate rows of white and rock elm, poor growth of the latter species accidentally allowed the white elm trees to reach maximum growth rates. Most tree planters have little or no control over the quality of planting stock, but seed zones give the grading of nursery stock and the buyer a guarantee that his planting under average conditions. 1 or 2 years nursery. Hardwood planting stock is mostly old when distributed and is seldom transplanted in the Good stock should be hardy, root system. the establishment of stock will be suitable for sturdy and have a well-branched During transport from the nursery and storage at the planting site the stock must be protected against desiccation or any other injury which could preclude planting success. Black Ash Black ash has rarely been planted in southern Ontario because its growth and form characteristics make it inferior to white ash. plantation sampled was planted by spade in ploughed furrows, The only at a spacing of 6 x 6 feet on former pasture land with a dense cover of weeds and grasses. The soil was a moist clay loam changing to clay at a depth of 15 inches. After 22 growing seasons black ash survival was only 20 per cent; height and diameter ranged from 16 to 22 feet and 2.1 to 3.3 inches, respectively, and mean annual height growth was 0.9 feet. In comparison, on the same site, mean annual height growth of white elm was 1.0 feet, silver maple 0.9 feet, and basswood 0.7 feet. White Ash White ash has been the most commonly planted hardwood tree in southern Ontario and 47 plantations were sampled, ranging in age from 19 to 43 years. On the basis of subjective evaluation, 9 plantations were classified as good, 26 as acceptable and 12 as poor. Best growth was al ways associated with fresh to moist soils of sandy loam to clay loam texture and rooting depth of 15 inches or more, but an increase in avail able soil moisture appeared to be able to compensate for a decrease in possible rooting depth. The species appeared to be well suited for planting on former fertile soil, but on unfertile or agricultural land with a deep, moist, dry sites growth was generally slow and tree form was poor. Plantings in openings of natural woocllots were generally more successful than those on former agricultural land of the same soil type. Rabbit browsing was a problem in some plantations but white ash seedlings appeared to be less susceptible than many other hardwood species, notably the oaks. Site preparation and cultivation after planting appeared to have increased Table 3 Growth of planted white ash as influenced by site conditions and planting treatments Age froo Avg Dominant and co-dominant trees Mean atra Former Depth Moisture Texture in. regime Total + fertilization 2 yr clay loam 22 fresh woodlot none none clay loam 14 moist good woodlot none none sandy loam 24 dry 2.0 fair cropped furrows none loam 34 moist 1.7 good cropped furrows 3 yr sandy loam 24 dry 1.6 fair pasture total 3 yr clay loam 16 fresh 1.4 good cropped furrows 3 yr clay 18 fresh 1.5 fair pasture furrows none loam 22 fresh 1.2 fair cropped furrows none silty clay 16 fresh 0.8 poor cropped furrows none sand 20 dry 0.7 poor cropped furrows none silty clay 8 0.7 poor pasture furrows none sandy loam 11 Survival dbh Avg ht ht growth Tree 20 70 5.1 55 2.7 excellent cropped 2.0 good 2.0 % ln# ft Soil of rooting zone Cultivation planting se«»d r Planting ft form land use site preparation after fresh dry Remarks best growth along creek 09 survival and early height growth in areas of heavy competition grass and weed (Table 3). The best plantation sampled was planted by spade at a spacing of 4x4 feet on fully cultivated agricultural land with a fresh clay loam soil kept, (Fig. 1). Although no record of the amount of fertilization was the owner recalled that nitrogen, potassium, and phosphorus were applied to the land shortly before planting. Full cultivation was maintained during the first 2 years after planting but no silvicultural treatments were applied thereafter. For 20-year-old white ash, height ranged from 49 to 59 feet and diameter from 3.3 to 7.1 inches. Mean annual height growth was 2.7 feet. With a survival of 70 percent, the stand is in urgent need of thinning. White ash planted in alternate rows with black locust in another section of the same plantation was completely dominated by the locust, average height and diameter being only 26 feet and 2.4 inches, respec tively, compared to 54 feet and 5.7 inches for black locust. An example of poor white ash growth due to unsuitable conditions was recorded in a 26-year-old plantation (Fig. 2). The trees had been planted by spade in ploughed furrows at 4 x 4 foot spacing on former pasture land with a rolling topography and dry shallow topsoil of sandy loam over medium fine sand. and diameter averaged Survival was only 30 percent, while height 17 feet and 1.9 inches, respectively. Eight years previously the white ash had been underplanted with white pine which has grown well and which shows internodes of up to 2.5 feet. pine establishment and early growth was probably aided by Although white the open ash overstorey, pine, nevertheless, appeared to be much better suited for afforestation of this site. Growth of white ash was nearly always satisfactory and often good in natural woodlot plantings. In a 31-year-old plantation of white ash and white elm established in a clearcut area of a natural woodlot, with a moist soil of clay loam, mean annual height growth of white ash was 2.0 feet while diameters ranged from 4.5 to 8.2 inches. White ash has frequently been planted in alternate rows with white spruce, Scots, red, and white pine. Few of these mixed plantations were fully successful because of differences in species requirements. One of the better plantations was a mixture of white ash and other hard woods planted in alternate rows with Scots pine. Trees had been planted by spade at a spacing of 6 x 6 feet on former pasture land with a fresh clay loam soil. At 43 years of age, height growth of white ash was best with a range of 54 to 71 feet compared to 55 to 66 feet for Scots pine. However, average diameter of Scots pine was 9.4 inches compared for white ash. to 7.4 A more representative example of mixed plantation development was recorded in a 24-year-old plantation of alternate rows of white ash, 10 Figure 1. White ash plantation established 20 years previously on soil of deep3 fresh clay loam. Avg. height 55 feet. Figure 2. White ash planted 26 years pre viously on dry, shallow sandy loam. Avg. height 17 feet. White pine underplanted 8 years previously. white spruce, white pine, and red pine. Trees had been planted by spade in ploughed furrows at a spacing of 6 x 6 feet on former pasture land with a sandy loam soil. Although survival of all species was fair, height growth of white ash varied widely whereas heights of conifers were rather even. In areas of good growth, ash was superior to all conifers but averaging the total 5-acre plantation, conifer growth was much better. Mean annual height growth of "good" ash was 1.0 foot compared to 0.8 foot for all three coniferous species. Average diameter was 3.3 inches for ash and spruce, 4.5 inches for white pine, and 4.6 inches for red pine. Basswood Basswood has been recommended as a plantation tree with great potential, but little is known about its site requirements. Relatively few. seedlings have been planted in southern Ontario and only five plan tations ranging in age from 22 to 79 years were sampled, four of which were evaluated as good, one as acceptable and one as poor. 11 In a 25-year-old mixed hardwood plantation of four rows per species, planted on fully cultivated agricultural land of sandy loam texture at a spacing of 4.5 x 4.5 feet, average height and diameter were 47 feet and 6.2 inches, respectively, with a mean annual height growth of 1.9 feet. Tree form was good (Fig. 3). Figure S. Basswood planted 25 years previously on soil of deep sandy loam. Avg. height 47 feet. In a single tree mixture of basswood and white ash, planted by spade on former agricultural land with a sandy loam soil, both species appeared to be evenly matched after 25 growing seasons. Since the orig inal spacing was 3.5 x 2 feet and the plantation was never thinned, approximately 40 percent of the trees had grown well while another 40 percent were badly suppressed, and the rest were missing. Dominant and co-dominant basswood trees were 24 to 39 feet high with diameters ranging from 3.5 to 6.8 inches compared to heights of 25 to 44 feet and diameters of 3.5 to 4.7 for white ash. The oldest plantation sampled had been planted 79 years pre viously at a spacing of 4 x 4 feet. The main species was sugar maple, with basswood and white ash scattered throughout the stand. Soil was a sandy loam near the surface changing to coarse sand at a depth of 22 inches. 11.5 to Height of basswood ranged from 67 to 83 feet with diameters of 19.4 inches. Height of sugar maple was nearly equal but the largest diameter was only 11.6 inches. 12 White Birch No extensive plantations of white birch were discovered during the survey but the species has been planted intermittently and six plan tations ranging in age from 18 to 26 years were sampled, three of which were evaluated as good, one as acceptable and two as poor. When white birch was planted on abandoned pastureland with dry shallow soil of a sandy loam texture, survival was less than 10 percent after 26 growing seasons, and average height was only 23 feet. A slightly better sur vival of 20 percent and average height growth of 25 feet were recorded in a 20-year-old plantation established by machine planting in furrows on former agricultural land with a 21-inch-deep loamy clay soil. In an 18-year-old mixed plantation of alternate rows of white birch and several other hardwood species machine planted in furrows, on dry pasture land with a loamy clay soil, birch had a survival of 80 percent while all other species failed completely. Average height and diameter were 27 feet and 3.4 inches, respectively, and mean annual height growth was 1.5 feet. ! Best survival and growth were recorded in a 20-year-old pure white birch plantation established on fully cultivated agricultural land at a spacing of 8 x 5 feet. The soil consisted of a plough layer of grey-brown clay loam over reddish-brown clay loam and changing to dark reddish-brown sandy clay loam at a depth of 16 inches. penetration was 32 inches. Deepest root Survival was 70 percent and height and diam eter ranged from 37 to 46 feet and 4.4 to 7.2 inches, respectively. Mean annual height growth was 2.0 feet. Stem form was good, with natural pruning to a height of 12 feet. Butternut Butternut has been planted sparingly in southern Ontario because its growth and wood characteristics are inferior to those of black walnut. Only four plantations ranging in age from 25 to 33 years were sampled. Best growth was found in a small plantation established in an understocked portion of a natural sugar maple-hemlock stand. Seedlings had been planted by spade without site preparation at a spacing of 8x8 feet in fresh clay loam soil. After 25 years, average height and diameter were 52 feet and 6.3 inches, respectively, and mean annual height growth was 2.1 feet. Natural pruning has been fair with some large, dead branches persisting. At the time of planting, butternut seedlings had also been planted in an adjacent, open field on soil belonging to the same series. Two years later red pine was planted between the rows of butternut to act as a nurse crop. After 25 years, average height and diameter of the 13 field-planted butternut trees were 26 feet and 3.3 inches, respectively, and mean annual height growth was only 1.0 feet. pine trees, feet high and had an average diam on the other hand, were 31 eter of 6.3 inches. The 22-year-old red Early thinnings favoring the butternut probably would have increased diameter growth. In a 25-year-old plantation, average height of butternut was 38 feet and average diameter 4.7 inches on soil of sandy loam, whereas on compact clay loam within the same plantation, height and diameter growth were only 30 feet and 3.5 inches, respectively. Catalpa Catalpa is definitely not a plantation tree in southern Ontario even though it is highly valued as an ornamental tree in landscape plant ing. The only plantation believed to exist in Ontario was planted in 1912 in a clearcut of a natural woodlot with a clay loam soil belonging to the Perth series. Planting stock was imported from a nursery in the United States and seedlings were planted by spade on ploughed ridges a spacing of 8 x 8 feet and were cultivated for at least 1 year. at The owner had been led to believe that catalpa could produce fence posts in 12 years and railway ties in 20 years, but the first fence posts of 8-inch diameter were actually cut in 1940. When the stand was sampled in 1964, heights and diameters ranged from 39 to 59 feet and 5.2 to 14.9 inches, respectively, with an average height of 52 feet and a diameter of 8.2 inches. Tree form was fair with generally short but straight stems and large crowns starting at a height of only 20 feet above ground. Natural regeneration has produced a dense second storey of good native hardwood saplings 10 to 25 feet high which are expected to produce a fully stocked stand when the catalpa are cut. Blaok Cherry Black cherry is of little importance as a plantation tree in southern Ontario and only two planting sites were visited: one planta tion showed fair growth while the other was a total failure. Black cherry machine planted in alternate rows with white ash, white pine, and white spruce at a spacing 6x7 feet on former pasture land had a survival of 70 percent after 22 growing seasons. Soil con sisted of a 9-inch-deep plough layer of dark grey clay loam over yellowish-brown clay loam, changing to brown clay at a depth of 14 inches. Form was very poor with half of the trees resembling small bushes with multiple leaders, probably as the result of repeated rabbit browsing. Heights and diameters ranged from 23 to 34 feet and 1.8 to 5.3 inches, respectively, with an average height of 27 feet compared to 34 feet for ash and pine. 14 Of many thousand black cherry seedlings planted by machine between rows of red pine 10 to 15 feet high on soil of dry loamy sand, not a single tree survived. This complete failure was probably the result of several factors, including unsuitable soil, excessive competition from the well established pines, and extensive rabbit browsing. Rook Elm The few rock elm seedlings planted have invariably failed to produce trees of acceptable size and form. After 25 years, trees planted on fully cultivated agricultural land with a sandy loam texture, had an average height and diameter of 37 feet and 4.5 inches, respec tively. Tree form was poor. In a 23-year-old plantation of two rows of rock elm alternating with one row of white elm, rock elm survival was less than 20 percent and the best tree was only 24 feet high compared to 59 feet for the best white elm. White Elm White elm has never been a common plantation tree in southern Ontario and its susceptibility to the Dutch elm disease, caused by Ceratoayatis ubni (Buism.) C. Moreau, has made planting completely inad visable at present. Fifteen plantations ranging in age from 20 to 41 years were sampled during the survey. Since plantation development has been greatly influenced by the severity of attack of the Dutch elm disease, it was impossible to evaluate success or failure of individual planta tions on the basis of survival and growth rate. However, best growth appeared to be associated with moist soils of silty clay or clay loam texture. On sandy, dry soils, on the other hand, growth was always slow and tree form was generally poor. Best growth was measured in a 23-year-old plantation of one row of white elm alternating with two rows of rock elm, at a spacing of 6 x 3 feet. Since the rock elm had grown very poorly and many of the white elm trees had died, final spacing was approximately 18 x 10 feet. Soil con sisted of silty clay over compact clay at a depth of 18 inches. Average heights and diameters ranged from 50 to 59 feet and 7.6 to 10.6 inches, respectively. Mean annual height growth was 2.5 feet. Planting of white elm at a spacing of 8 x 8 feet in openings of a heavily cut woodlot with a moist, deep soil of clay loam texture pro duced trees of good form, whose average height and diameter were 59 feet and 8.7 inches, respectively, at 31 years of age (Table 4). Table 4 Growth of planted white elm as Influenced by site conditions and planting treatments Age Dominant and co-dominant trees from seed vr Avg Survival % Hean ann dbh Avg ht in. ft ht growth Former Tree land ft form use 57 2.5 fair 59 1.9 32 Planting site Cultivation after Soil of rooting zone Depth Moisture preparation planting Texture in. regime pasture furrows 2 yr silty clay 18 fresh good woodlot none none clay loam 14 moist 1.6 fair cropped total 2 yr clay loam > 17 fresh 36 1.4 fair cropped total 1 yr sandy loam 16 dry 31 1.2 poor cropped total 3 yr clay loam 12 fresh 27 0.8 poor pasture furrows none sand 20 dry Remarks best growth 16 In mixed plantations of white elm, white ash, and silver maple, the elm trnes was always slightly lower than that of average height of the other two species while average diameter was equal or slightly larger. Shagbark Hickory Shagbark hickory occurs naturally in many southern Ontario woodlots but has seldom been planted. In the only plantation sampled, the species had been planted in single tree mixture with red oak, sugar maple, and basswood on fully cultivated agricultural land of Burford loam. Spacing was 7x3 feet and cultivation was maintained for 2 years after planting. The height and diameter of 27-year-old hickory ranged from 33 to 46 feet and 3.9 to 5.4 inches, respectively. Mean annual height growth was 1.4 feet for hickory, sugar maple, and basswood, and 1.8 feet for red oak. Black Locust Black locust is not a common plantation tree in southern Ontario but has been planted occasionally, mostly in the form of shelterbelts around farms or fields or for erosion control on steep slopes or land of low fertility. Three plantations were sampled during the survey, in age from 8 to 20 years. When planted on infertile land, growth were poor; but on deep, fertile soil, ranging survival and growth ranged from acceptable to very good. The average height of black locust planted in four rows along the edge of a field at a spacing of 4 x 4 feet on soil of fresh clay loam over compact clay was 23 feet after eight growing seasons. Average diameter was 2.5 inches and mean annual height growth was 2.8 feet. Crowding and excessive drainage appeared to be the main causes of poor growth of black locust planted 17 years ago in alternate rows with black walnut on former pasture land with a dry sandy loam soil. Spacing was 6x3 feet. High survival and lack of thinning has resulted in a very dense canopy with none of the trees being able to express dominance. Average height of black locust was 27 feet and average diameter 3.8 inches, compared with black walnut growth of 22 feet and 2.8 inches. In a mixed plantation of alternate rows of black locust and white ash planted 20 years previously on fully cultivated agricultural land with a clay loam soil and cultivated for 2 years after planting, black locust completely dominated the stand (Fig. 4). Average height and diameter were 54 feet and 5.7 inches, respectively. Form was good with long slender boles, and few branches to a height of 30 feet. attack were found. No signs of locust borer 17 Figure 4. Black locust and white ash planted in alternate rows 20 years previously on soil of deep, fresh clay loam. Avg. height of locust 54 feet. Silver Maple Silver maple has been planted extensively in southern Ontario because the species is well suited for afforestation of poorly drained areas too wet for farming. Twenty-two plantations ranging in age from 15 to 43 years were examined; but since most plantings were restricted to land unsuitable for farming, plantation size was generally less than 3 acres. Evaluation of plantation success revealed that only three planta tions could be classified as good, three as acceptable and 16 as poor. Although height growth was often considered satisfactory, tree form was generally poor, making the trees unsuitable for sawlog utilization. Spacing as close as 3 x 3 feet did not prevent profuse sprouting from the base. To produce trees of merchantable size and acceptable form, it will, therefore, be necessary to reduce clumps to single stems and even then many trees will tend to grow crookedly or lean excessively. 18 Moist bottomland is the natural habitat of silver maple in southern Ontario and plantations established on such sites generally showed satisfactory growth. However, some plantings in fresh to moist soils of silt, clay, or loam texture also grew well. Plantings on dry exposed sites or soils of coarse texture, however, never produced trees of acceptable size or form. On moist sites, even dense ground competition had little effect on seedling survival and growth; while on drier sites, cultivation for at least 2 years after planting will be necessary to assure satisfactory establishment. Although the species is presently of little value as a timber tree, it can be used to good advantage as a nurse crop for more valuable hardwood species. Because of its rapid growth, it has also been found useful for shade tree planting in moist corners of pastures or near barn yards where shade is required for cattle. The best plantation sampled grew on bottomland near a creek which flooded the valley for short periods during spring. Soil was a structure less, deep silty clay. For 23-year-old silver maple, average height was 57 feet and average diameter 8.2 inches. Tree form was fair to good (Table 5). Good growth was recorded in a silver maple stand planted at the bottom of a steep slope on soil of moist clay loam (Figure 5). For 43year-old silver maple, average height and diameter ranged from 69 to 90 feet and 9.5 to 15.4 inches, respectively, with a mean annual height growth of 1.9"feet. However, most trees were crooked or leaning and will produce little timber of sawlog quality. Planted in mixture with white spruce on dry soil of medium fine sand, silver maple survival was less than 20 percent after 35 years while mean annual height growth of the poorly shaped trees was 1 foot. Sugar Maple Planting of sugar maple has often been attempted but since the majority of plantations have been marked failures, the belief is widely held that the species cannot be planted successfully. The survey has shown that although sugar maple is more difficult to plant than almost any other native hardwood species, planting, nevertheless, is quite feasible. Eleven plantations ranging in age from 23 to 79 years were sampled and four plantations are discussed in some detail. Evaluation shows that two plantations may be classified as good, two as acceptable, and seven as poor. Size of planting stock appeared to be of great importance in the establishment of successful plantations. Large wildlings transplanted frcm nearby woodlots were generally more successful than small nursery-giown stock. Table 5 Growth of planted silver maple as influenced by site conditions and planting treatments 20 Figure 5. Silver maple planted 43 years previously on soil of moist olay loam. Avg. height 81 feet. Tree form poor. While natural regeneration is nearly always abundant under the canopy of a mature sugar maple forest, plantation establishment in the open is difficult because the seedlings appear to be very susceptible to heat and frost damage. Furthermore, the seedlings' inability to compete with herbaceous vegetation and their susceptibility to rabbit and deer browsing appear to be the main obstacles to successful planting. In a 25-year-old mixed hardwood plantation of four rows per species, planted on fully cultivated agricultural land of sandy loam texture at a spacing of 4.5 x 4.5 feet, survival of sugar maple was lowest of all species planted (Table 6). Only 40 percent of the original number of seedlings planted were alive and less than 10 percent are expected to grow into trees of acceptable form. Tree height ranged from 26 to 41 feet with diameters of 2.8 to 5.5 inches. In comparison, basswood and silver maple in the adjacent rows were 60 to 65 feet high. An even lower survival was recorded in a mixed 23-year-old hardw od plantation of 10 rows per species. Less than 10 percent of the sugar maple trees had survived after years of rabbit browsing and com petition from weeds and grasses. Comparative survival of white ash was 80, black walnut 70, and red oak 50 percent. The remaining sugar maple trees ranged in height from 17 to 30 feet with diameters of 1.2 to 3.0 inches. Stem form was generally acceptable but because of their open growth all trees had exceptionally large crowns. Table 6 Growth of planted sugar maple as influenced by site conditions and planting treatments 22 Use of large natural seedlings resulted in high survival in a 26-acre sugar maple plantation established on fine, sandy soil of an abandoned strawberry plantation. Natural seedlings 4 to 6 feet high, from the adjacent woodlot were planted at an even spacing of 12 x 12 feet, and hay and hardwood slash were spread over the soil to prevent drifting and to help in the establishment of a grass cover. Thirtyeight years after planting survival was still 90 percent. Heights ranged from 30 to 51 feet and diameters from 4 to 9 inches. Tree form was only fair because of short boles and bushy crowns, but 2 years previously the owner had successfully started to tap the larger trees for production of maple syrup. Figure 6. Sugar maple planted 79 years previously on soil of sandy loam. Avg. height 71 feet. The oldest plantation sampled was planted 79 years previously at a spacing of 4 x 4 feet on former agricultural land with a sandy loam soil changing to coarse sand at a depth of 22 inches (Fig. 6). Origin of planting stock could not be determined but it is believed that natural seedlings were used. In 1947, 63 years after planting, the stand received a light thinning. Since the survival has been very high, the stand would probably have greatly benefited from repeated thinnings started at an early age. Height and diameter ranged from 53 to 83 feet and 5.6 to 11.6 inches, respectively. Tree form was 23 excellent, with straight stems free of branches to a height of 30 feet. Diameter growth, on the other hand, has been retarded by the very close spacing which still averages less than 6x6 feet. Burr Oak Burr oak has been recommended for planting on adverse sites, but few trees have been planted in southern Ontario and the species was sampled in only two plantations. In the first plantation, burr oak had been planted on abandoned farm land with a topsoil of clay loam changing to compact clay at a depth of 17 inches. After 19 growing seasons survival was only 30 percent while average height and diameter were 13 feet and 2.1 inches, respective ly and mean annual height growth was 0.8 feet. In a 30-year-old plantation,of alternate rows of Scots pine and burr oak planted at a spacing of 4 x 4 feet on sandy loam soil, survival of both species was over 80 percent. Height and diameter of burr oak ranged from 35 to 51 feet and 5.1 to 9.2 inches, respectively, mean annual height growth being 1.4 feet. The height of Scots pine equalled that of the oak but the average diameter was nearly 80 percent higher. All oaks had very straight stems, but despite the close spacing, small adventitious branches covered nearly the total length of all stems. Red Oak Red oak is a rapidly growing timber tree of good quality and is one of the most commonly planted hardwood trees in southern Ontario. Of the 13 plantations sampled ranging in age from 23 to 60 years, six were classified as good, three as acceptable, and four as poor. Best growth was always associated with deep, well-drained soils ranging in texture from loamy sand to loam, whereas trees planted on soils of very coarse or very fine texture or those with a shallow topsoil and a compacted subsoil never grew well. When planted on suitable soils, the species appears to be well adapted for reforestation of cleared land. However, rabbit control is an absolute prerequisite to successful plantation establishment. In sevtiral reforestation areas visited during the survey, rabbits had either killed the seedlings or by repeated browsing of annual shoots had held total height to less than 4 feet. In mixed plan tations of red oak and pine, oak survival was generally low because of the difficulty of rabbit control (Table 7). In a single tree mixture of red oak, sugar maple, shagbark hickory, and basswood planted 25 years previously on fully cultivated agricultural land with deep loam soil, red oak growth was greatly sup erior to that of all other species (Figure 7). Height and diameter Table 7 Growth of planted red oak as influenced by site conditions and planting treatments to 25 ranged from 36 to 57 feet and 3.5 to 9.6 inches, respectively, while mean annual height growth was 1.8 feet. Tree form was Twenty-three years after red oak, white ash, good. and black walnut had been planted in several rows per species on old pasture land with a deep, loam soil, height and diameter of red oak ranged from 27 to 54 feet and 2.9 was to 6.3 inches, respectively, 1.6 feet compared to 1.5 and mean annual height growth for black walnut and 1.3 for white ash. In a mixed plantation of several rows of red oak, silver maple, white elm, and white ash, black walnut, planted on former agricultural land of fairly compact clay loam texture changing to very compact clay at a depth of 16 inches, red oak growth was greatly inferior to that of all other species. was only 1.2 feet, After 25 growing seasons, mean annual height growth compared to 1.7 feet for black walnut and 1.6 feet for white ash. Figure 7. Red oak planted 25 years previously in mixture with other hardwoods on deep loam soil. Avg. height 48 Figure 8. Pure red oak plan tation established 23 years previously on soil of deep sandy loam. Avg. height 41 feets good form. feet. One of the best red oak plantations sampled was a pure stand, established 23 years previously on former agricultural land of deep sandy loam (Fig. 8). Twenty years after planting approximately 200 trees per acre had been pruned to a height of 17 feet. inal spacing of 4 x 4 feet and a survival of 70 percent, not thinned because a sufficient number of With an orig the stand was trees had achieved dominance. 26 The height and diameter ranged from 33 to 48 feet and 4.0 to 6.8 inches, respectively, inches. for an average height of 41 feet and a diameter of 5.4 This growth compared favorably with that of an adjacent white pine plantation of equal age, having an average height and diameter of 38 feet and 6.9 inches, respectively. The oldest red oak plantation sampled was established 60 years previously by direct seeding. Acorns had been sown in several rows the middle of a white pine plantation established at deep, well-drained soil of medium fine sand. was 67 feet compared to 59 for white pine. and pine were 8.1 and 9.2 inches, in the same time on The average height of oak The average diameter of oak respectively. Tree form of oak was good to excellent. White Oak White oak is of little importance as a plantation tree in southern Ontario because of its relatively slow growth and its suscep tibility to rabbit browsing. However, the species has been planted occa sionally and three plantations were sampled in which white oak occurred as a minor species. In a 19-year-old plantation, white oak had been planted in mixture with white ash, silver maple, burr oak, and black walnut on abandoned farmland with a topsoil of dark grey, friable clay loam changing to brown clay loam at a depth of 9 inches, and to yellowish 17 inches. White oak survival was only 20 brown very compact clay at percent while average height was 12 feet and mean annual height growth was 0.6 feet, compared to silver maple. growth; 1.1 feet for ash and 0.8 feet for burr oak and Black walnut was a total loss owing to very stunted its height averaged 3 to 4 feet. Growth of white oak was mjch better in a mixed plantation of sugar maple, white ash, and black walnut planted 20 years earlier in an old, poorly regenerated clearcut. loam over very compact clay. of 4 x 2 feet and lack of Soil consisted of 21 inches of clay All trees had been planted at a spacing thinning and dense ingrowth of blue beech resulted in a very crowded stand. Height and diameter of white oak trees ranged from 17 to 33 feet and 1.3 to 3.4 inches, respectively, while mean annual height growth was 1.2 feet compared to 1.3 feet for the two other species. Best growth of white oak was recorded in a 43-year-old mixed plantation of Scots pine, white ash, black walnut, silver maple, and white oak, listed in order of survival. than 10 percent is attributed The poor oak survival of to heavy rabbit browsing. was a dark brown friable clay loam to a depth of less Plantation soil 12 inches underlain by 6 inches of lighter colored clay loam and changing to medium brown, 27 compact clay at a depth of 18 inches. Very few roots were found below this depth. Height and diameter growth were surprisingly even for all species with the best Scots pine tree reaching a height of 66 feet and a diameter of 11.3 inches, while the best white oak was 71 feet high and 12.8 inches in diameter. However, only one such tree was found and the average height and diameter of white oak were 56 feet and 7.9 inches, respectively. Mean annual height growth of oak was 1.3 feet compared to 1.5 for white ash and 1.4 for Scots pine. Carolina Poplar Nearly all poplar plantings in southern Ontario have been in the form of windrows or afforestation of either swampland or very dry, sandy areas. No large forest plantations were found but a few height and di ameter data were collected in windrows or erosion control plantings. Carolina poplar cuttings planted 26 years previously in a single row along the eastern side of a silver maple plantation on soil of fresh clay loam grew to a maximum height of 88 feet and a diameter of 13.5 inches. Average height and diameter were 65 feet and 10.0 inches, respec tively, compared to 56 feet and 6.7 inches for silver maple. On the slope of an artificial dam constructed of stony clay fill, height and diameter of 21-year-old poplar trees ranged from 46 to 64 feet and 5.0 to 10.1 inches, respectively, with healthy trees of acceptable form. All plantings on very dry, sandy soil failed to produce trees of acceptable size or form and most stands started to deteriorate at an age of 20 years or younger. No natural regeneration of any kind was found in these plantations and the soil was at best only partially covered by patches of sparse grass or dead branches. With the deterioration of the poplar stand, the site will again be subject to erosion. Poplar there fore appears less suitable for afforestation of such sites than jack pine which was planted occasionally on the same site and was generally growing well. Black Walnut Black walnut is unquestionably the most valuable timber species in Ontario. Trees of good form and diameters of 20 inches and larger are in great demand as veneer logs and command prices up to 10 times higher than those paid for sugar maple or yellow birch logs of equal quality. Because of the great demand and high value, black walnut has been planted extensively throughout its range. During the survey 43 plantations ranging In Jif»ii from 14 1am lu;iL Ion r Ins.s 11 i oil to 85 years were sampled on a varLety of soLls. IJ of these plantations as good, 19 as acceptable, 28 and 11 as poor. Woodlot plantings were generally more successful than plantations established on former agricultural land belonging to the same soil series. Of the six woodlot plantings sampled, three were classified as good, two as acceptable, and only one as poor (Table 8). Most plantation failures could be traced directly to unsuitable soil and moisture conditions of the planting site. Since black walnut is a very demanding species, good growth can be expected only on rich, moist, deep, fertile soil which is found in natural woodlots, valley bottoms and along creeks. On the farm, the most suitable planting sites appear to be orchards or other highly fertile areas such as abandoned vegetable gardens or moist but well-drained draws. It is completely inadvisable to plant black walnut seedlings in abandoned fields or pas tures of low fertility or on dry exposed ridges or shallow soils. The most successful planting was found in a natural woodlot where seedlings had been planted by spade in openings of a mature stand of sugar maple, silver maple, white elm, and white ash (Fig. 9). At 48 years from seed, average height and diameter were 72 feet and 8.8 inches, respectively, with the best tree reaching a height of 90 feet and a di ameter of 15.3 inches. Form was excellent with clear boles to a height of 30 feet. The woodlot soil belongs to the Perth series and consists of 8 inches of dark brown clay loam over 7 inches of greyish-brown clay loam changing to a dark brown clay, mottled and compact. consists of a compact, mottled, below a depth of 15 inches. pale brown clay. The C horizon No roots were found The site is imperfectly drained and pools of water cover the surface until early summer. The woodlot is well managed, has never been pastured, and since the owner still uses fuelwood on his farm, he has removed all dead, dying, or poorly shaped trees, thereby releasing the walnuts. Black walnut seedlings, while intolerant of overhead shade, appear to demand protection when planted in the open. plantation established on good, soil, black walnut seedlings pine trees, In a 20-acre fully cultivated farm land of clay loam grew only a few inches per year until red planted 3 years later between the rows of walnut, started to provide protection (Fig. 10). After 25 growing seasons, average height and diameter of walnut were 27 feet and 3.2 inches, respectively, com pared to 30 feet and 5.4 inches for the 22-year-old red pine. Since the trees were planted at a spacing of 4 x 3 feet and had never been thinned, crowding limited crown development of many black walnut saplings, but enough trees maintained their position in the canopy to be able to grow into a fully stocked stand if removal of red pine trees is not too long delayed. At the time of open-field planting, a small walnut plantation was established in an understocked area of growth has been much better the adjacent woodlot. than in the open field. diameter of black walnut were 49 feet and 5.5 inches, mean annual height growth being 2.0 feet and to good. Here, Average height and respectively; tree form ranging from fair Table 8 Growth of planted black valnut as Influenced by site conditions and planting treatments to 30 Figure 9. Excellent black wal nut planted 47 years previously in a natural woodlot on soil of moist clay loam. Avg. height 72 feet. Figure 10. Alternate rows of black walnut and red pine, planted respec tively 3 25 and 22 years previously on soil of fresh clay loam. Avg. height of walnut 27 feet. Figure 11. Poor, deteri orating black walnut, planted 38 years previously on soil of 11-inch deep clay loam over very compact clay. Figure 12. Trees of the same plantation as Figure 11 but growing in 21inch-deep soil of clay loam over compact clay. Avg. height 24 feet. Avg. height 48 feet. 31 The effect of rooting depth and spacing on height and diameter growth of planted black walnut is shown in Table 9. plots were measured in the same plantation (Fig. plots number Three 1/10-acre 11 and 12) but while 1 and 2 were located well inside the plantation, plot 3 represented only trees growing in the outside row bordering a pasture. All trees had been planted 38 years previously in a fully cultivated field at an even spacing of 10 x 10 feet and clean cultivation had been maintained for 5 years after planting. The soil belonged to the Schomberg series which is characterized by a dark clay loam topsoil underlain by lighter colored clay loam over a very compact clay which appears to be impenetrable to tree roots. Table 9 Influence of soil depth and spacing on black walnut growth Rooting Plot no. 1 depth in. Avg Avg Survival % ht ft diam in. 70 24 4.4 11 Stand quality deteriorating, poor form, many dead branches, gnarled tops. 2 21 70 48 7.4 healthy, good form, long boles, small crowns. 3 21 100 47 12.0 healthy, fair form, short boles, large crowns. The difficulty of obtaining reliable information on plantation establishment and stand history has limited the survey stands. to a few mature The oldest trees measured had been planted 85 years previously in an abandoned nursery with a deep clay loam soil. The height and diam eter ranged from 84 to 95 feet and 15.0 to 23.5 inches, respectively. Tree form was fair to good, some trees having clear boles to a height of 45 feet. CONCLUSIONS The results of the survey lead to the conclusion that it is not at present economically feasible to establish large hardwood plantations. The few excellent plantatiors, on the other hand, can be planted successfully if prove the proper conditions that hardwoods are met. Unfortu nately, our knowledge of "proper conditions" is so limited that hardwood reforestation is mostly carried out by costly and wasteful. trial and error and this is both 32 The survey results also indicate that hardwood trees planted in woodlot soils have succeeded much more frequently than those planted on former agricultural or pasture land. It therefore appears reasonable to concentrate present day hardwood planting at the rehabilitation of de vastated, understocked woodlots and those small areas near creekbeds and on lower slopes which are characterized by deep, moist, fertile soils. Intensive research on the other hand should be directed toward increasing our knowledge of species requirements, in regard to soil, moisture and nutrition as well as protection from exposure and browsing damage. Since the first printing of this report, 6 years ago, intensive research (von Althen 1969a,b; 1970) has produced sufficient information to recommend hardwood afforestation if the following conditions can be met: (1) a deep, fertile, moist but well drained planting soil; (2) complete mechanical site preparation; (3) large, sturdy planting stock; (4) large, deep planting holes and careful planting; weed control; and (6) fertilization where necessary. (5) effective 33 REFERENCES Fingland, J.A. et at. 1964. Report of the committee on private forest lands in southern Ontario. Ontario Professional Foresters' Association, Hills, G.A. 1959. p 13. A ready reference to the description of the land of Ontario and its productivity. Dept. Hosie, R.C. Lands Forests, 1969. Native trees of Canada. Forest. , Can. Putnam, D.F. and L.J. Sci. Agr. Forest. 1969. . 1969. Seventh edition. Dept. Fish. 380 p. 1938. The climate of southern Ontario Hardwood planting problems and possibilities in Can. Dep. Forest. Publ. No. 1043. Proposed planting stock grades for hardwoods Can. Dep. Fish. Forest. Inform. Rep. Artificial regeneration of black walnut. plaiting and container-planting. Inform. . of Res., Ont. 9 p. of seeding, Forest. Rep., Div. 401-446. planted in Ontario. O-X-106. Serv. Chapman. 18: (8): von Althen, F.W. 1964. eastern Canada. . Prelim. p 142. 1970. Rep. O-X-107. Can. Dep. Studies Fish. 8 p. Methods for successful afforestation of a weed infested clay soil. Forest. Chron. 46(2): 139-143. ■■• t " XIQNHddV 34 APPENDIX A Common and Botanical Names Mentioned in Text Black ash Fraxinus nigra Marsh. White ash Fraxinus americana L. Basswood Tilia amerioana L. White birch Betula papyrifera Marsh. Butternut Juglans ainerea L. Catalpa Catalpa speaiosa Ward. Black cherry Prunus serotina Ehrh. Rock elm Ulmus thomasii Sarg. White elm Ulmus amerioana L. Shagbark hickory Carya ovata (Mill.) K.Koch Black locust Robinia pseudoaoaoia L. Silver maple Acer saooharinum L. Sugar maple Acer saocharum Marsh. Burr oak Querous macrocarpa Michx. Red oak Querous rubra L. White oak Querous alba L. Carolina poplar Populus oanadensis Moench Black walnut Juglans nigra L. Eastern hemlock Tsuga oanadensis Eastern white pine Pinus strobus L. Jack pine Pinus banksiana Lamb. Red pine Pinus resinosa Ait. White spruce Pioea glauoa (Moench) Voss (L.) Carr
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