Soil-forming factor 4: Living organisms All plants and animals (from microorganisms to humans) affect soil formation. Living organisms add organic matter - a key component of soil - through the breakdown of litter and the decomposition of roots. Microorganisms, such as fungi and bacteria, facilitate chemical exchanges between roots and the soil to produce essential nutrients. Both animals and plants allow moisture and gases to seep into deeper layers along burrows and root channels. Humans can impact soil formation through land management practices that disturb natural processes and change the chemical and physical characteristics of the soil. Cultivation practices and burrowing animals mix soil from different horizons, especially from the organic-rich surface layers. The nature of biological activity in the soil is governed by climate, topography and soil characteristics such as depth, texture, structure and chemistry (e.g. pH, salinity). Ecoregions are classified by the presence of biomes, which are major plant communities determined by rainfall and climate. Forests, grasslands (including savannah and shrubland) and deserts are distinguished by climate (e.g. tropical, subtropical, temperate) and water conditions. In addition, forests are divided into conifers, broadleafed or mixed. Biomes Tropical and subtropical moist broadleaf forests Tropical and subtropical dry broadleaf forests Temperate broadleaf and mixed forests Temperate coniferous forests Tropical and subtropic grasslands, savannas and shrubland Ecoregions and biomes Flooded grasslands and savannas Ecoregions can be defined as relatively large units of land or water containing a distinct assemblage of natural communities and species, with boundaries that approximate the original extent of natural communities prior to major land-use change. The limits of ecoregions generally follow continental boundaries or major barriers to plant and animal distribution (such as the Himalayas and the Sahara). Africa south of the Sahara Desert, including the island of Madagascar, lies in the Afrotropic ecoregion. With the exception of Africa's southern tip, the Afrotropic ecoregion has a tropical/subtropical climate. The Sahara and northern Africa lie in the Palearctic ecoregion. Montane grasslands and savannas Mediterranean forests, woodlands and scrub Deserts and xeric shrublands Mangroves Lakes Above: Map showing the major biomes of Africa as defined by the World Wide Fund for Nature (WWF). The continent is characterised predominantly by deserts and savannah. (WWF/JRC) [15]. Below: Land cover map of Africa. This map was produced for the year 2000 using data collected by sensors on satellites. (JRC) [16, 16a] Land cover The term 'land cover' is used to describe the vegetation covering the surface of the planet (which can also be bare ground or unvegetated). It is important to distinguish between the terms ‘land cover’ and ‘land use’. For example, a land cover of mixed shrubs and grass could be a park, an orchard or in an African context, savannah. This map on the lower-right shows the principal land cover types of Africa for the year 2000 as mapped by sensors on-board satellites orbiting the Earth [16]. The map shows that the central regions are covered by extensive tropical forests (dark green) which merge to the north and south with open woodland and grasslands or savannah (brown and orange). In between, the light green areas denote a mosaic of cropland and forest that indicates the human alteration of natural vegetation patterns. With increasing aridity, the savannah grades into open and sparse grasslands (yellow colours). Where water is more plentiful, rainfed and irrigated agriculture exists (pink). These areas are particularly evident in Zimbabwe, around Lake Victoria and along the Mediterranean, East African and South African coast. The map clearly depicts the enormous extent of the vast Sudd wetlands of southern Sudan (light blue), the Okovango Delta in Botswana and the swamp forest of the Congo Basin. The distinctive red patches are the evaporative salt flats and pans of the desert margins. Land cover Land-cover classes with a dominant tree layer Land-cover classes with a dominant shrub or grass layer mixed with agriculture Dominant agriculture Closed evergreen forest Open deciduous woodland Rain-fed agriculture Degraded evergreen forest Closed shrubland with sparse trees Irrigated agriculture Closed deciduous forest Closed shrubland Swamp forest Open shrubland with sparse trees (>65% tree cover) (5-15% tree cover) Mangrove Open shrubland (<5% tree cover) Bare rock Mosaic forest – croplands Closed grassland Salt hardpans Mosaic croplands – other woody vegetation Open grassland with sparse trees Water bodies Closed deciduous woodland Open grassland (>65% tree cover) (>65% tree cover) (40-65% tree cover) (15-40% tree cover) (5-15% tree cover) Orchards (<5% tree cover) Other land-cover classes Bare soil (5-15% tree cover) (<5% tree cover) Sparse grassland Swamp bushland and grassland Introduction | Soil Atlas of Africa 19
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