Tectonophysics 502 (2011) 175–195 Contents lists available at ScienceDirect Tectonophysics j o u r n a l h o m e p a g e : w w w. e l s ev i e r. c o m / l o c a t e / t e c t o The evolution of the Danube gateway between Central and Eastern Paratethys (SE Europe): Insight from numerical modelling of the causes and effects of connectivity between basins and its expression in the sedimentary record K.A. Leever a,⁎, L. Matenco a, D. Garcia-Castellanos b, S.A.P.L. Cloetingh a a b Netherlands Research Centre for Integrated Solid Earth Science (ISES), Faculty of Earth and Life Sciences, VU University, De Boelelaan 1085, 1081 HV Amsterdam, The Netherlands Instituto de Ciencias de la Tierra Jaume Almera (ICTJA-CSIC), Group of Dynamics of the Lithosphere (GDL), Solé i Sabarís s/n, 08028 Barcelona, Spain a r t i c l e i n f o Article history: Received 15 June 2009 Received in revised form 4 December 2009 Accepted 9 January 2010 Available online 20 January 2010 Keywords: Connectivity Sedimentary basins Numerical modelling Paleogeography a b s t r a c t The Pannonian and Dacic Basins in SE Europe are presently connected by the Danube River across the South Carpathians, to which they are in a back-arc and foreland position respectively. Part of the Paratethys realm during the Neogene, open water communication between the basins was interrupted by the Late Miocene uplift of the Carpathians. Different mechanisms have been proposed for the formation of the Danube gateway: capture of the upstream lake or an upstream river or incision of an antecedent river. Estimates on its age range from Late Miocene to Quaternary. A related issue is the effect of the large Mediterranean sea level fall related to the Messinian Salinity Crisis on the Paratethys subbasins, specifically the “isolated” Pannonian Basin. In a synthetic numerical modelling study, using a pseudo-3D code integrating tectonics, surface processes and isostasy, we addressed the causes and effects of changes in connectivity between two large sedimentary basins separated by an elevated barrier. Specifically, we aimed to find the expression of connectivity events in the sedimentary record in general and the consequences for the evolution of the Pannonian–Dacic area in particular. We studied a range of parameters including the geometry and uplift rate of the barrier, downstream sea level change and lithosphere rigidity. We found that changes in connectivity are expressed in the sedimentary record through their effect on base level in the upstream basin and supply in the downstream basin. The most important factors controlling the response are the elevation difference between the basins and the upstream accommodation space at the time of reconnection. The most pronounced effect of reconnection through lake capture is predicted for a large elevation difference and limited upstream accommodation space. Downstream increase in sediment supply is dependent on the latter rather than the reconnection event itself. Of the parameters we tested, the rigidity of the lithosphere was found to be of major importance by its control on sediment loaded subsidence and generation of accommodation space. A downstream sea level change is unlikely to induce capture, but may affect the upstream lake level by enhancing incision in a pre-existing gateway. In the Pannonian–Dacic region, the mechanically weak, continuously subsiding Pannonian lithosphere allowed accommodation of significant volumes of continental sedimentation and as a consequence, transfer of excess sediment to the downstream Dacic Basin was only gradual. The Messinian sea level fall in the Dacic Basin could have been recorded in the Pannonian Basin only if a connection between the basins already existed. More detailed modelling of river incision taking into account lateral differences in erodibility in the South Carpathians will be required to give better time constraints on the formation of the Danube Gateway. © 2010 Elsevier B.V. All rights reserved. 1. Introduction Paratethys, extending from the foreland of the Alps to the Aral Sea (Fig. 1a), was a large brackish epicontinental sea that was separated from the world oceans during the progressive closure of the Tethys Ocean and associated rising of the Alpine chain. In Southeast Europe, ⁎ Corresponding author. Presently at: Department of Geosciences, University of Oslo, P.O. Box 1047, Blindern, 0316 Oslo, Norway. E-mail address: [email protected] (K.A. Leever). 0040-1951/$ – see front matter © 2010 Elsevier B.V. All rights reserved. doi:10.1016/j.tecto.2010.01.003 formation and Late Miocene uplift of the Carpathians disrupted open water communication between the Central/Western and Eastern Paratethys, corresponding to the Pannonian Basin and the Dacic Basin respectively (Fig. 1). The separation of the basins led to the evolution of separate faunas and different biostratigraphies (e.g. Rögl, 1996) making stratigraphic correlations problematic. The Pannonian Basin to the west and the Dacic Basin to the east (Fig. 1) are in a back-arc and foreland position relative to the Carpathians, respectively. The Pannonian Basin system formed as a result of back-arc extension during the Miocene (Horváth et al., 2006) and features the 176 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 weakest continental lithosphere of Europe (Cloetingh et al., 2006). The subsidence in the Getic Depression (westernmost part of the Dacic Basin) was due to its transtensional opening during the clockwise rotation of the orogen along the Moesian platform and later orogenic loading (Matenco et al., 1997; Fuegenschuh and Schmid, 2005). Other than by tectonic structuring, the connectivity between the Paratethys subbasins was influenced by climate through its control on sea level. Effects of the major sea level fall in the adjacent Mediterranean Sea related to the Messinian Salinity Crisis (>1000 m, Clauzon, 1978) have been reported in the Paratethys subbasins, both the Black Sea (Gillet et al., 2007), the Dacic Basin (Clauzon et al., 2005; Leever et al., 2009) and the Pannonian Basin (Csató et al., 2007; Leever et al., 2009). Presently, the Pannonian and Dacic Basins are exposed and connected by the Danube River, deeply incised into its gorges across the South Carpathians (Fig. 1b). The morphology of the long-studied feature has been explained by the contrasting models of either stream capture (Peters, 1876), lake overflow (Toula, 1896) or incision of an antecedent river (Cvijic, 1908), while estimates on the age of their formation range from Sarmatian to Quaternary (see Marovic et al., 1997, for a review). These models are fundamentally different in the sense that the “capture model” assumes that the river carved its bed into a preexisting topography by headward erosion, while the “antecedent river model” assumes that river incision was able to keep pace with uplift of the mountains—implying that the fluvial connection between the basins was never lost. Recent modelling work addressed the evolution of gateways across elevated barriers by investigating the factors controlling capture (piracy) of tectonic lakes (Garcia-Castellanos, 2006) and even an ocean (Loget and Van Den Driessche, 2006): capture of the Atlantic Ocean across the strait of Gibraltar reputedly allowed flooding of the Mediterranean, restoring its sea level and ending the Messinian Salinity Crisis. The time required for capture was found to be dependent mostly on the precipitation/evaporation budget in the captured drainage basin (upstream of the barrier), the rate of tectonic uplift at the barrier and the flexural isostatic response of the lithosphere (Garcia-Castellanos et al., 2003; Garcia-Castellanos, 2006) and on the base level in the lower (downstream) basin (Loget and Van Den Driessche, 2006). In this paper we use forward numerical modelling of coupled surface processes and lithosphere response to determine the effect of changing connectivity between two large sedimentary basins on depositional geometries and sediment partitioning. The aim is to define the signature of connectivity events on the stratigraphic record, i.e. what to look for in seismic data. In other words, applied to the study area: are observations of sedimentary architecture from seismic sequence stratigraphy of any use to decide which of the models for formation and evolution of the Danube gateway (capture vs. incision of an antecedent river) is correct? We address questions such as: what is the influence of the strongly contrasting lithosphere strength in the Pannonian and Dacic Basins? Could the effects of the Mediterranean sea level fall of the Messinian Salinity Crisis have extended to the Pannonian Basin, and how? Did it influence the evolution of the Danube Gateway and as such the connectivity between the Pannonian and Dacic basins? 2. Tectonic and paleogeographic setting of Paratethys in SE Europe 2.1. Tertiary tectonic evolution The Pannonian–Carpathian region evolved during Alpine deformations in the tectonic context of the Mediterranean, which is character- 177 ized by highly arcuate plate boundaries resulting from the roll-back and steepening of subducted lithosphere into land-locked remnant oceanic basins (Wortel and Spakman, 2000; Fig. 1a). The South Carpathian orocline, located between the Balkans in the south and the South Carpathians in the northeast (Fig. 1), was structured in a polyphase tectonic history that started in the Middle Cretaceous and continued well into the Miocene (Sandulescu, 1984; 1988; Matenco and Schmid, 1999; Fuegenschuh and Schmid, 2005). Paleogene to Early Miocene rotation of the Cretaceous belt around the western edge of the Moesian promontory led to orogen-parallel extension which culminated in the Late Eocene and large displacement along curved strike–slip faults during the Oligocene to Early Miocene (Cerna Jiu and Timok fault, Fig. 1) Locally, the exhumation induced by the uplift was large enough to be recorded by apatite fission track data (Fuegenschuh and Schmid, 2005 and references therein). In the Moesian foreland plate, the transcurrent motions led to the transtensional opening of the Getic Depression as a dextral pull-apart basin during the Early Miocene. In the course of the Miocene, the tectonic regime changed to transpression, with the strike slip motions in the western Getic Depression gradually changing to thrusting in the east (Rabagia and Fülöp, 1994; Rabagia and Matenco, 1999). Thrusting of the belt onto the Moesian Platform led to additional subsidence in the South Carpathian foredeep. The effective elastic thickness of the Moesian lithosphere has been estimated at 30 km (e.g. Cloetingh et al., 2006). The ongoing Neogene E-ward movement of the internal Carpathian units, driven by slab roll back, led to extension in the intra-Carpathian area, affecting both Tisza–Dacia in the south and the Alcapa block in the north (Ustaszewski et al., 2008). The formation of the Pannonian basin system by rifting and transtension (Horváth et al., 2006) was controlled by three main tectonic processes, gravitational collapse, subduction rollback and asthenosphere updoming, the relative importance of which is still a topic of intense debate (Cloetingh et al., 2006 and references therein). In particular the latter process led to weakening of the lithosphere in the Pannonian Basin, its effective elastic thickness being estimated at 5–10 km by Lankreijer et al. (1997). In the last stage of its evolution, basin formation and extension in the Intra-Carpathian region have come to an end. Structural inversion of the Pannonian basin is currently in progress, driven by the push of the Adriatic plate (e.g. Bada et al., 1999) and evidenced by significant late-stage uplift and subsidence anomalies during Late Pliocene through Quaternary times (Horváth and Cloetingh, 1996). 2.2. Paleogeographic evolution 2.2.1. Biostratigraphic constraints on connectivity Until the Middle Sarmatian (Bessarabian, Fig. 2), the Central and Eastern Paratethys were connected over the South Carpathian orocline (Fig. 1), as evidenced by the brackish water fauna common to Central and Eastern Paratethys found in the intramontane basins on its west flank (Marinescu et al., 1981 and references therein; Gagic, 1997). During the Sarmatian, uplift of the South Carpathian orocline led to the disconnection of the two basins. Open water communication ceased and Central Paratethys continued its evolution as the endemic Lake Pannon, the divergence in the faunal evolution due to the separation defining the onset of the Pannonian stage. In the Dacic Basin the Sarmatian stage continued into the Khersonian (Fig. 2). During the Pannonian stage, corresponding to the Upper Sarmatian and Meotian of the Dacic Basin (Fig. 2), the fauna in the two basins are different (e.g. Rögl, 1996; Rögl, 1999). However, in Meotian sediments Fig. 1. Study area. a. Late Miocene (Messinian) paleogeography of Paratethys and the Mediterranean showing location of gateways and study area. Modified from Popov et al. (2006). b. DEM of the Pannonian–Carpathian area. Blue lines indicate the extent of Lake Pannon (after Magyar et al., 1999a) and the Dacic Basin (after Saulea et al., 1969) at different time steps postdating the mid-Sarmatian disconnection of the two domains (M–U Sm, Middle–Upper Sarmatian; Meot, Meotian; U Pont–Dc, Upper Pontian–Dacian; see time scale in Fig. 2). White lines are drainage divides and indicate the boundaries of the present day Danube drainage basin. Black rectangle represents the modeled area. Within the Danube drainage basin, the Carpathians and northern Balkans divide the Dacic (200 × 103 km2) and Pannonian (600 × 103 km2) realms. Note the difference in the extent of the drainage (and sediment source) areas of the modeled parts of the Pannonian and Dacic basins. The Danube crosses the drainage divide between the Pannonian back-arc and Dacic foredeep realms in the South Carpathian Orocline. 178 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 2. Time scales. a,b. Correlation charts between Central and Eastern Paratethys time scales after Rögl (1999); Steininger et al. (1990); Sacchi et al. (1999) and Vasiliev et al. (2004, 2005). c. Correlation chart of 2nd and 3rd order sedimentary sequences in the Pannonian Basin (Hungary). Data on sedimentary sequences from (1) Transdanubia (Sacchi et al., 1999); (2) entire Hungary (Juhász et al., 1999); and (3) Eastern Hungary, (Ujszászi and Vakarcs, 1993; Vakarcs et al., 1994). Sequences were defined from seismic (1, 3) and well data (2); their ages constrained by magnetostratigraphy or by direct correlation to the Haq et al. (1987) eustatic curve (3). For reference the central Paratethys time scale is shown (left hand panel, ages according to Sacchi et al., 1999; Sacchi and Horváth, 2002). MSC indicates the Messinian Salinity Crisis (5.96–5.33 Ma, Krijgsman et al., 1999). From the lower resolution well data (2), only 2nd order sequences were distinguished, bounded by significant hiatuses. Sacchi et al. (1999) correlate their 3rd order sequences PAN-1 to PAN-4 with the “Late Miocene sequence” of (2) and sequences 5–8 or IV–VII of Ujszászi and Vakarcs (1993) and Vakarcs et al. (1994) respectively; the correlated sequences are marked in light grey shading. The large age variations for the base Pannonian unconformity illustrate the correlation problems even within the Pannonian Basin. The 3rd order sequence boundaries (SB) associated with–according to the authors–the largest base level falls, are indicated with a bold line: SB PAN-3 in Transdanubia at 8.7 My (1) and SB #8 in eastern Hungary (3). K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 in the Dacic Basin, Pannonian-derived Congeria, typical for freshwater environments, have been found (Olteanu, 1979), suggesting a fluvial influx from Lake Pannon into the Dacic Basin. In the Lower Pontian (Eastern Paratethys definition, Fig. 2), the fauna in the Pannonian, Dacic and Euxinic realms again show common characteristics. The reconnection has been attributed to the “Odessian transgression” (Olteanu and Jipa, 2006). Pannonian type fauna are found over the entire Paratethys. However, no Eastern Paratethys forms have been found in the Central Paratethys (Magyar et al., 1999b). 2.2.2. Seismic sequence stratigraphic constraints on base level changes Lake Pannon (Magyar et al., 1999a) inherited its complex bottom topography from the rifting and transtension stage that led to the opening of the Pannonian Basin. Subsidence rates and water depth were therefore different in the individual subbasins. Disconnected from the world oceans during the Pannonian endemic stage (Fig. 2), the water level of the lake was controlled by the balance between evaporation and water supply from contributing rivers and precipitation (GarciaCastellanos, 2006), both controlled by climatic variations (Kázmér, 1990; Juhász et al., 1999). The surrounding mountain chains provided abundant sedimentary influx. The basin was progressively filled by fluvial–deltaic and turbiditic sediments dominantly from northwesterly (palaeo-Danube) and northeasterly directions (palaeo-Tisza) (Vakarcs et al., 1994). The variations in rate of progradation to aggradation of the deltafed shelf-slope systems, led to the recognition of third and fourth order seismic sequences (Vakarcs et al., 1994; Sacchi et al., 1999) and indicate that the relative lake level varied significantly both in space and time. The most detailed sequence stratigraphic studies have been carried out in the Hungarian part of the basin, both to the west of the Danube in Transdanubia (Ujszászi and Vakarcs, 1993; Sacchi et al., 1999) and to the east in the Great Hungarian Plain (Pogácsás et al., 1990; Pogácsás et al., 1992; Csató, 1993; Vakarcs et al., 1994; Csató et al., 2007; Juhász et al., 2007). Estimates of the Pannonian lake level change vary from “tens of meters” to 200 m. Due to the scarcity of reliable age constraints, the correlation between the sequences is not straightforward (Fig. 2). The northwestern part of the Dacic Basin is coincident with the Getic Depression, its subsidence history controlled by transtension and subsequent foreland flexure (cf. Section 2.1). From a detailed study of seismic sequence stratigraphy at the margins of the Getic Depression, Rabagia and Matenco (1999) concluded that the observed base level changes until the Late Miocene were predominantly tectonically controlled, eustatic changes playing only a subordinate role. A regional seismic sequence stratigraphic interpretation in the western part of the Dacic Basin focused on the Latest Miocene–Pliocene basin fill evolution (Leever et al., 2009). In this stage of its evolution, in contrast to the earlier stage and the Pannonian Basin, control by differential tectonic subsidence was less important. A sudden base level fall in the lower part of the Pontian was attributed to the Messinian lowering of the water level in the Black Sea below the threshold of the Dobrogea barrier. Late Pontian sediments associated with a subsequent base level rise transgressively cover the older deposits (Leever et al., 2009). 3. Numerical modelling of the signature of changing connectivity on the sedimentary record We use numerical modelling to study the factors influencing connectivity and determine its signature on the sedimentary record. The conceptual model considers two adjacent basins separated by an elevated barrier, with or without a pre-existing gateway. In the latter case, a fluvial connection is established by capture of the upstream basin as the result of its higher lake level and the erosion of the barrier. Coupled forward modelling of tectonics, surface processes and isostasy is used to determine the response of the system in terms of depositional geometries to parameters such as tectonic uplift, downstream base level 179 changes and flexural rigidity, the importance of which in controlling lake capture and the evolution of tectonic lakes has been established in previous studies (Garcia-Castellanos et al., 2003; Garcia-Castellanos, 2006; Leever et al., 2009). 3.1. The numerical method: TISC TISC is a pseudo-3D (planform) forward finite difference code (Garcia-Castellanos, 2002; Garcia-Castellanos et al., 2003) in which tectonics, surface processes and flexural isostasy are fully coupled. It is designed to study the interaction between surface mass redistribution and the lithospheric response by uplift and subsidence on large temporal (105–106 y) and spatial scales (of an entire sedimentary basin and/or orogenic belt). The rate of surface uplift (and subsidence) is thus a function of the rate of tectonic uplift, erosion and sedimentation, and flexural isostatic rebound. It does not allow–nor do we aim–for detailed reconstructions of morphological evolution or prediction of higher than first order sedimentary sequences. Tectonic uplift (of the barrier, in our case) is kinematically defined. Resulting surface mass redistribution is calculated at time steps of 1000 yr following the stream power-law formulation by Kooi and Beaumont (1994) including short-range diffusive and long-range linear transport functions that represent hillslope and fluvial processes respectively. Fluvial transport is calculated by explicitly calculating the drainage network during the topographic evolution, accounting for the formation of lakes in local topographic minima. These lakes become closed (endorheic) if evaporation in its surface becomes larger than the water they collect. The time step for imposed tectonic uplift and calculated flexural isostatic response is 0.5 Ma. Flexural calculations follow an elastic thin plate model characterized by the effective elastic thickness Te, and account for the loading of sediment and water as well as unloading due to erosion. 3.2. Modelling setup and boundary conditions The model setup and boundary conditions (Fig. 3) are derived from the main characteristics of the study area (Fig. 1). The Pannonian and Dacic basins are in very different stages of their tectono-thermal evolution, and as a consequence have greatly different lithospheric rigidities (Te of 5–10 and 30 km respectively; Cloetingh et al., 2006). The basins are in restricted connection over a topographic high, with only a small elevation difference between them. The drainage area and hence the initial sediment supply in the Pannonian basin were much larger than those of the downstream Dacic basin (Fig. 1). In the model, where the area of both basins is equal, the larger drainage area and sediment supply in the upstream basin are represented by a water and sediment input at the western edge of the model (Fig. 3). The surrounding topography prevents any sediment from leaving the model domain. No vertical movements due to flexure are allowed at the model edges, except on the eastern side. This is based on the assumption of a steady-state topography in the mountains surrounding the basins, while the Dacic Basin has a larger extent eastward than its model counterpart (Fig. 1b). In the model, the sediment loading is larger than the erosion due to the external sediment source, and these flexural boundary conditions are required to prevent excess downward deflection. The parameters used by the surface process model are listed in Table 1 and have been validated in previous modelling studies (e.g. Garcia-Castellanos et al., 2003). The models were run for a period of 15 My. All model runs have a 400 × 200 km model domain and 2 × 2 km grid cell size (Fig. 3). In our reference model, the initial maximum elevation of the surrounding topography is 800 m and the maximum initial depth of the basins is 400 m. Tested variations of this scenario are summarized in Table 2. An example of model evolution is shown in Fig. A1. 180 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 3. Model setup. a. 2 km resolution DEM of the modeled area. b. Initial topography and boundary conditions for the conceptual model series: 400 × 200 km, 2 km grid cells. Topography in this figure corresponds to the initial elevations for Model 2, 5 and 6. The geometry of the barrier separating the two basins is different for the various model runs, see Table 2. All model boundaries are fixed for deflection, only the eastern side is allowed to move freely. A sediment source on the western side of the model (1.3 103 km3/Ma) represents the flux from the large Pannonian drainage area. Sediment is allowed to leave the model at all sides. c. Model setup and parameters in cross section. Qs_ext, external sediment input; P, precipitation rate; E, evaporation rate; S, slope; H, max. elevation of surrounding topography; Hb, elevation of barrier; Hg, elevation of gateway; dH, elevation difference between base levels at time of capture; U, uplift rate. Parameter values are listed in Table 1 and 2. 3.3. Results 3.3.1. Model 1—reference model In the reference model, the basins are separated by a threshold at sea level. The surrounding topography is 800 m (Fig. 3, Table 2). Model results after 15 My are shown in Fig. 4. 3.3.1.1. Rates of sedimentation and erosion. In the upstream basin, sedimentation initially occurs at a higher rate than in the downstream basin (Fig. 4a, b). This is due to the external sediment source that feeds the upstream basin (Fig. 3), while the downstream basin is initially only sourced by erosion from the surrounding mountains. Between 6.5 and 9 My, the relative sedimentation rates are inverted: the K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Table 1 Parameters for the surface process model. Transport and diffusion coefficients and erosion and deposition length scales according to the formulation of Kooi and Beaumont (1994). Parameters Values Model resolution (grid cell size) Transport coefficient Kf Diffusion coefficient Ks Surface processes time step Tectonic/isostatic time step Erosion length scale lf Basement Sediments Deposition length scale lf 2 × 2 km 60 kg/m3 0.05 m2/s 0.05 My 0.5 My 120 km 60 km 25 km upstream basin is filling up and excess sediment is transferred to the downstream basin. In the upstream basin, the sedimentation rates decrease to zero as the basin is completely filled (Fig. 4). The erosion rates show an overall decreasing trend in both basins, due to the declining topography. Erosion rates are higher in the upstream basin, which is surrounded by topography on three sides while the downstream basin is lined by topography only on the N and S side (Fig. 3). The initial decreasing trend in sedimentation rates in the upstream basin (0 to 6 My, Fig. 4b) reflects the decrease in supply from local erosion. The sedimentation rates in the downstream basin, however, do not reflect the local erosion rates: some sediment is transferred from the upstream source into the downstream basin also before the bulk shift in sedimentation between 6.5 and 9 My. 3.3.1.2. Depositional geometries. The increase in sedimentation rates in the downstream basin is evident from the increasing spacing between the 0.5 My time lines from 6 My (Fig. 4c). Also the upstream basin shows locally increased sedimentation rates after 6 My, when looking at the time lines. The local increase is due to the decreased accommodation space and is not reflected in the sedimentation rates integrated over the entire upstream basin (Fig. 4b–c). Sediment loading resulted in flexural isostatic subsidence in the entire model area: more than 400 m in the basins (the basin floor was initially at –400 m, Table 2) and some 200 m on top of the “barrier” (Fig. 4c). In the upstream basin, subsidence led to a concave shape of the initially flat shelf edge trajectory. 181 3.3.2. Effect of the barrier geometry (models 2-3) The effect of barrier geometry is studied in different ways. In Model 2, the basins are connected across an existing gateway, the elevation of which is varied. In the subsequent models, the basins are initially disconnected and the elevation and slope of the barrier are varied. 3.3.2.1. Model 2: elevation of a pre-existing gateway. Model 2 addresses the effect of the initial elevation of a pre−existing gateway in the barrier between the two basins (Table 2, Fig. 3). The results are shown in Fig. 5, with those of Model 1 for comparison. 3.3.2.1.1. Rates of sedimentation and erosion. During the first 7 My, the gateway elevation has virtually no effect on the infill of the basins: the cumulative sedimentation is equal for the three cases (Fig. 5a). Only in the second model stage, after the shift of the bulk sedimentation to the downstream basin, do the models show some difference in sediment volume. The difference with the reference model (in which no barrier is present) is more pronounced than the difference between the individual cases: the bulk sedimentation shift occurs earlier for all cases in Model 2 than in the reference model. Moreover, sedimentation continues afterward, albeit at a lower rate. An initial difference in erosion rates in the upstream basin (Fig. 5b), reflecting the initial difference in base level controlled by the elevation of the outlet of the basin, disappears after 2 My. This is the time required to bring the gateway to an equal elevation for all three models, by allowing the river in the gateway to become graded in a dynamic equilibrium. In the downstream basin, the erosion rates are exactly equal for all three models during the entire model time (Fig. 5b). 3.3.2.1.2. Depositional geometries. The depositional geometries in Model 2, i.e. the shelf edge trajectory and the spacing between time lines, are very similar to the reference model. A major difference is that sedimentation in the upstream basin continues after the bulk sedimentation shift, accommodating a large volume of fluvial sediments (Fig. 5, subparallel subhorizontal time lines). The volume of continental sediments in the three models is different, however; the largest volume being accommodated by the model with the initially highest gateway (Fig. 5c). The fluvial sedimentation and the difference in volumes can be explained from the river profiles at the final time step. The eroding barrier between the two basins is uplifting by flexural isostatic rebound. This is the main difference with the reference model, in which the “barrier” was initially at sea level, was loaded by sediments and, as a Table 2 Model setup. Geometry Max barrier uplift rate U (m/Ma) Te (km) W–E Rate of sea level change (m/Ma) – 2.7 – – 10 10 – – – 10 – – 0.75 2 3 2.7 10 – 800 200 2.7 100 200 400 – – 800 800 200 2.7 – 400 400 – 2 – 30 20 10 5 20–30 10–30 5–30 10 Max elevation H (m) Barrier elevation Hb (m) Gateway elevation Hg (m) Max slope S (degrees) 1 Reference model (Fig. 4) 2 Gateway elevation (Fig. 5) 800 800 0 800 3 Barrier slope (Figs. 6, 7) 400 400 – 0 100 200 – 4 Uplift rate (Figs. 8, 9) 800 0 to 400 5 Te constant (Figs. 10, 11) 800 6 Te varied (Figs. 11, 12) 7 Sea level change (Fig. 13) – 200 100 66.6 50 182 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 result, subsided. The gateway between the two basins is more resistant to erosion than the sediments in the basins (Table 1), resulting in a knickpoint in the river profile. Upstream of this knickpoint, i.e. in the upstream basin, the river accumulates sediments in order to maintain its gradient and keep the ability to cross the barrier (see also Snow and Slingerland, 1990). K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 3.3.2.2. Model 3: slopes of the barrier. In this model, in contrast with the previous models, there is no pre-existing gateway of reduced elevation as a way of communication between the basins. The surrounding topography is 400 m (Table 2). The effect of barrier slope was addressed in three different models, with a slope on the downstream side of the barrier of max 0.75, 2 and 3° respectively. The slope on the upstream side of the barrier is 2°, in order to keep the volume of the upstream basin constant. Sedimentation and erosion rates are shown in Fig. 6, profiles in Fig. 7. 3.3.2.2.1. Rates of sedimentation and erosion. The model with the lowest slopes has the largest exposed area and hence the highest erosion rates in the downstream basin (Fig. 6). The sedimentation rates in the downstream basin are initially equal to the local erosion rates: the only supply to the downstream basin is from the local erosion. The moment of capture (Fig. 6, marked by x) may be recognized from the breakdown of this relationship: sedimentation rates start increasing while the erosion rates keep decreasing. Capture occurs first for the model with the steepest slope (Fig. 6): due to the steeper slope the barrier is narrower and less erosion is required to oversteepen the slopes at the drainage divide. This asymmetry allows the local backward migration and lowering of the drainage divide and thereby capture of the upstream lake—of which the level has risen to the top of the barrier (Fig. 7). It is important to notice that, after capture of the upstream basin, the sedimentation rates in the downstream basin initially increase only gradually. The strongest increase occurs later and is due to the overfilling of the upstream basin. The capture-induced increase in sedimentation rates in the downstream basin is not reflected by a similar decrease in the upstream basin: the water level drop that results from the capture leads to a larger exposed area and therefore to increased supply, balancing the sediment outflow to the downstream basin (Fig. 6). Moreover, the capture time influences the timing of the bulk sedimentation shift, because capture and subsequent lowering of the outlet by erosion lead to lowering of the base level in the upstream basin and reduction of the accommodation space. As such, the sooner the capture, the sooner the upstream basin is filled, and the sooner the sedimentation shifts to the downstream basin. The lacustrine–continental transition in the upstream basin occurs during or after the strongest decrease in sedimentation rates. Compared to the previous model series, only a very small amount of continental sediments are deposited in the upstream basin (compare Fig. 5c and 7), before sedimentation rates drop to zero or even become negative (implying erosion of previously deposited sediments). 3.3.2.2.2. Depositional geometries. The implications of lake capture on depositional geometry are shown in more detail in Fig. 7 for each of the three scenarios. In the left series of panels the sediment geometry in the upstream basin before capture and resulting erosion is shown. The initial progradation, seen from the shelf edge trajectory, is due to the high sediment input close to the source (Fig. 3). The sediments are progressively spread over a larger area and consequently develop a more aggradational character, also due to the larger subsidence in the center of the basin. Capture occurs first for steeper slopes (Fig. 7c). Following capture (time line highlighted in Fig. 7, right hand panels), sediments in the upstream basin are deposited in a downstepping geometry during the ongoing water level fall that results from erosional lowering of the outlet, lasting ∼2 My in all cases. The sediment accumulation area in the upstream basin is reduced by the water level fall, resulting in a local 183 increase in sedimentation rates (see spacing of time lines in Fig. 7, right hand panels) even though the total rate in the whole basin remains largely unchanged. The downstream basin shows higher sedimentation rates (wider spacing of the time lines in cross section, Fig. 7, cf. Fig. 6) after capture. The moment of capture determines the total amount of sediments that can be accommodated in the upstream basin, and therefore the amount of sediments transferred to the downstream basin. The longer the duration of the isolated stage of the upstream basin, with its elevated base level, the more sediment can be accommodated. The duration of the lacustrine stage in the upstream basin is therefore most extended in case of the low slope model (upper panels): the upstream basin is not completely filled even after 15 My. Delay in the moment of capture, i.e. a longer isolated life time of the lake at local elevated base level, leads to a higher degree of overfilling. The resulting high river gradient after capture causes erosion rather than fluvial sedimentation in the upstream reaches in order to attain equilibrium (compare Figs. 5, 7). 3.3.3. Effect of barrier uplift rate (Model 4) In this model, the two basins are initially separated by a barrier at 0 m (as in the reference model), which is subsequently uplifted. The rate of the uplift is varied in the three cases while its magnitude is constant at 400 m (Table 2). The width of the uplifting zone is ∼50 km and has slopes of ∼3°. The base level or maximum lake level in the upstream basin is kept at 200 m by an outlet at the N margin of the upstream basin, simulating the effect of the precipitation–evaporation ratio in the Pannionian Basin, which has a larger area in nature than in the model. The surrounding topography is 800 m. Results are shown in Figs. 8 and 9. 3.3.3.1. Rates of sedimentation and erosion. The conditions and initial model evolution, before the onset of uplift at 2 My, are equal to Model 1. During this stage, some sediment is transferred from the upstream to the downstream basin, shown by the sedimentation rates in the downstream basin that exceed the local supply (Fig. 8b). In the absence of an initial gradient, the two basins are disconnected the moment the barrier rises above 200 m, the maximum lake level of the upstream basin. Uplift of the barrier leads to an increase of the erosion rates in the downstream basin (Fig. 8a), while the erosion rates in the upstream basin decrease. The changes are most pronounced for the models with the high uplift rates. The approximately constant sedimentation rates in the downstream basin during the uplift period for low uplift rates (U = 100 m/Ma, Fig. 8b) reflect the continuous connection between the basins, maintained by the barrier erosion that is able to keep up with the uplift. The sedimentation rates in the downstream basin start increasing gradually after the end of the uplift stage from 6 My on, due to erosional lowering of the barrier and increasing sediment transfer from the upstream basin. For the other models (U = 200, 400 m/Ma), the sedimentation rates in the downstream basin match the local supply rates from the moment of disconnection onward (Fig. 8b). Capture in these models (marked by x in Fig. 8b) directly leads to the onset of the bulk shift in sedimentation between the basins, as shown by the sudden and fast increase of sedimentation rates in the downstream basin. 3.3.3.2. Depositional geometries. Sedimentation patterns are shown in Fig. 9. In the left panels, the time step preceding capture is shown. Note that the basins in the low uplift rate scenario (U = 100 m/Ma, upper Fig. 4. Model 1: reference model. a. Cumulative sediment volumes, derived from local erosion and external sediment input. b. Rates of sedimentation (bold lines) and erosion (thin lines). The sedimentation rate is the rate of variation of the total sediment volume, while the erosion rate represents the total eroded volume per time step, including bedrock and previously deposited sediments. Note that the initial sediment accumulation rates in the upstream basin greatly exceed the erosion rates due to the external sediment input (see Fig. 3) Grey shading in a. and b. marks the period of bulk sedimentation shift from the western to the eastern basin, characterized by rapidly changing sediment accumulation rates and related to the lacustrine– continental transition in the upstream basin (t = 8 My). The (restricted) connection between the two basins allows some sediment transfer from the upstream to the downstream basin from the start, as seen from the sedimentation rates in the downstream basin that exceed the local supply (erosion) rates. c. W–E cross section at the center of the model (y = 0, cf Figure A1) at the final time step (15 My) with time lines for each 0.5 My. The onset of increased sedimentation rates in the downstream basin (t = 6.5 My) is marked by a bold line; the final stage of infill of the upstream basin (t = 8 My) by a heavy broken line. Shelf-slope break indicated by short dash in both basins. 184 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 5. Model 2: effect of gateway elevation. a. Cumulative sediment volumes. For comparison the results of Model 1 are shown. In contrast to Model 1, sedimentation continues in the upstream basin after the bulk sedimentation shift, albeit at lower rates. The transition from lacustrine to fluvial sedimentation is marked by box (cf Fig. 5c). b. Erosion rates. Rates are equal in the downstream basin for all model runs (dashed lines), independent of the gateway elevation, but different in the upstream basin during the first 2 My. c. W–E cross sections at t = 15 My. Lacustrine–continental transition is indicated by bold line, shelf-slope break by short dash. The volume of continental sediments deposited in the upstream basin since 8 My is largest for the model with the initially highest gateway elevation of 200 m (lower panel). K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 185 Fig. 6. Model 3: effect of barrier slope. Sedimentation and erosion rates indicated by bold and thin lines respectively. High slopes correspond to a narrow barrier (see also Figs. 3c, 7), and lead to faster capture of the upstream basin (marked by X). Capture results in increasing sedimentation rates in the downstream basin, which were initially declining and equal to the local erosion rates. Increasing erosion rates in the upstream basin after capture reflect the increased exposed area due to the falling base level. The largest increase in sedimentation in the downstream basin corresponds to the end of lacustrine sedimentation and filling up of the upstream basin (marked by arrows). panels) were repeatedly connected and disconnected during the period of uplift, as incision was able to keep up with uplift. The uplifted, elevated barrier represents a load that causes deflection of the basin floor adjacent to the barrier (compare Fig. 9, upper left panel and Fig. 7, lower left). The sedimentation patterns in the upstream and downstream basins subsequent to capture are similar to those in Model 3 (see Fig. 7). However, in Models 4b and 4c, because the barrier rises to an elevation exceeding the maximum upstream lake level, and because– despite the subsidence induced by the barrier uplift–the available accommodation space preceding capture is smaller (base level at 200 m instead of 400 m), the basin fill is in a more advanced stage at the moment of capture than in Model 3. The capture is followed by an immediate strong increase in sedimentation rates in the downstream basin (lower panels, see also Fig. 8b), instead of the more gradual increase in sedimentation rates characterizing the low uplift rate of Model 4a, and also Model 3 for low slope values. 3.3.4. Effect of flexural rigidity (Models 5–6) In these models, the effect of flexural isostatic response is studied by varying the effective elastic thickness Te. 3.3.4.1. Model 5: constant Te. The initial setup of this model is equal to Model 2 (Fig. 3) with an initial gateway at 200 m elevation in an 800 m surrounding topography. Based on estimates of lithosphere rigidities (Lankreijer et al., 1997; Cloetingh et al., 2006), we varied Te values between 5 and 30 km (Table 2), constant over the entire model area. The results are shown in Fig. 10. In Fig. 10a the sedimentation rates in the upstream and downstream basin are compared for different Te. Onset of the (enhanced) decrease of sedimentation rates in the upstream basin occurs first for the most rigid plate (Te = 30 km, at ∼3.5 My) and results in a very pronounced and sudden increase in sedimentation rates in the downstream basin (Fig. 10a, peak at 5.5 My). The required time to completely fill up the upstream basin is shortest on the highly rigid plate (Figs. 10 and 11). For Te= 30 km, even the downstream basin is filled up by 10 My. For the weaker plates, the onset time of decreasing sedimentation rates in the upstream basin is later, and the change more gradual. Much more time is required to completely fill the basin: the low rigidities allow the generation of significant additional accommodation space by sediment loading. The onset of fluvial sedimentation in the upstream basin occurs just before the peak in sedimentation rates in the downstream basin for all models. Sedimentation rates in the downstream basin start gradually increasing long before the onset of the fluvial stage in the upstream basin. In Fig. 10b, the time to the lacustrine/continental transition and the complete filling of the basin is plotted as a function of Te. In case of a weak lithosphere, the basin infill not only takes much longer to complete, but it is also preceded by a protracted stage of fluvial sedimentation (cf. Fig. 11, upper panels). 3.3.4.2. Model 6: lateral transition in Te. These models address the effect of a lateral (W–E) change in Te. Based on the lithosphere rigidities in the Pannonian and Dacic Basins, the Te in the east is kept at 30 km, while its value in the upstream basin is varied between 5 and 30 km (Table 2). The transition is gradual over a distance of 60 km below the barrier. The results of this model (Fig. 11, lower panel; Fig. 12) are similar to those of the previous model. The rigidity of the lithosphere underlying the upstream basin determines its infill time, and as such the time and rate of the shift of bulk sedimentation to the downstream basin. Peak sedimentation in the downstream basin is again most pronounced in case of a rigid upstream lithosphere. The bulk sedimentation shift occurs ∼0.5 My earlier for all scenarios than in the previous model, because of reduced subsidence of the downstream margin of the upstream basin. 3.3.5. Effect of downstream base level change (Model 7) In this model, the basins are initially disconnected by a 400 m high barrier (Table 2). A sea level fall of 200 m starts at t = 0 My and proceeds at different rates, until the initial level is reached again at 8 My (Fig. 13a). In case of the fastest sea level fall, this leads to a 6 My lowstand period. The 8 My time interval was chosen based on the results of Model 3, which has the same setup except the sea level change, and where capture is predicted at t = 7.5 My (Fig. 7). The results are presented in Fig. 13b, which shows only the sedimentation and erosion rates in the downstream basin. The models show an initial increase in erosion and sedimentation rates, because the falling water table leads to the exposure of a larger area. The rate of change reflects the rate of the sea level fall. The sedimentation and erosion rates decrease again during the subsequent sea level rise. 186 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 7. Model 3: effect of barrier slope (continued). Cross sections for Model 3. Vertical scale in m, horizontal in km. The left panels show the time step (in My) preceding the capture of the upstream basin. The corresponding time line is marked by a bold line in the right hand panels (cross sections at the model end time, 15 My). The other highlighted (long dash, dotted) time lines represent the end of base level fall and the final stage of lacustrine sedimentation in the upstream basin respectively. For the low slope scenario (S = 0.75°), the western basin is not completely filled after 15 My. The shelf-slope break is indicated by a dashed line in all panels. Note the difference in depth to basement in the upstream basin and the (apparent) increase in sedimentation rates in both basins after capture. Further discussion in the text. Capture of the upstream basin (marked by arrows, Fig. 13) occurs only after the sea level has returned to its initial level and is likely triggered by the loading effect of the rising sea level which subdues the incised barrier. From these results it follows that a base level fall does not accelerate the capture of the upstream basin. To the contrary, the capture occurs 0.5 to 1 My later than in Model 3 (without sea level fall). 4. Discussion 4.1. Controls on depositional geometry and sediment partitioning The individually tested parameters in the different model runs lead to marked differences in the patterns of sedimentary infill in the two basins. However, we found that the tested parameters are not critical by themselves, but only to the degree by which they influence the true parameters that control the downstream sedimentation rates in relation to changes in connectivity. These are the elevation difference and upstream accommodation space at the time of capture, outlined in Fig. 14. The difference between the water levels in the two basins before connection has a large effect on the depositional geometries in the upstream basin since it determines the magnitude of the capture-induced upstream base level fall by erosional lowering of the outlet. The elevation difference is dependent on the barrier geometry and uplift rate: a large elevation difference at the time of capture is more likely for a wide barrier, high uplift rates and low erodibility (Model 3, 4). The remaining accommodation space in the upstream basin at the time of capture determines the time lag for the bulk shift of sedimentation to the downstream basin (e.g. Model 1, 2 in Figs. 4 and 5). The remaining upstream accommodation space is not an independent parameter, it is dependent on the timing of capture and moreover it correlates with the K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 187 Fig. 8. Model 4: effect of uplift rate. a. Erosion rates. Uplift period is indicated by bars. The vertical scales for the respective basins are offset by 100 m for better distinction. b. Sedimentation rates. Crosses mark moment (period) of capture. Legend as in 8a. elevation difference between the basins. Generation of accommodation space by sediment loading is enhanced by low Te values (Models 5 and 6). These parameters are discussed in more detail below for different stages in basin evolution: the disconnected (endorheic) and connected lacustrine stages and the subsequent (upstream) continental stage. Though the modelling setup was based on the characteristics of the Pannonian–Dacic realm, the general results may be extended to other areas where elevated topography divides two sedimentary basins, such as the Ebro Basin (foredeep to the Pyrenees) and the adjacent extensional Valencia trough (Garcia-Castellanos et al., 2003), separated by the Catalan Coastal Ranges. 4.1.1. Disconnected stage and capture of the upstream endorheic lake 4.1.1.1. Upstream basin. The elevation difference between the basins after capture (and therefore the magnitude of the base level fall) is determined by the equilibrium gradient of the river connecting them which is a function of the erodibility and the width of the barrier. If the upstream basin is still in the deep lacustrine stage with ample accommodation space at the time of capture, the capture-induced base level fall results in the deposition of a strongly progradational series with a downstepping geometry (e.g. Model 4, Fig. 9). In case of a large elevation difference and/or limited remaining accommodation space at the time of capture, the sedimentation rates in the upstream basin are locally significantly increased because of the reduced sediment accumulation area (Fig. 9) and capture may lead to accellerated infilling of the remaining accommodation space, ending the lacustrine stage. 4.1.1.2. Downstream basin. The sedimentary response to capture in the downstream basin is mainly sensitive to the remaining upstream accommodation space. If the upstream basin is captured when it still has ample accommodation space, the response in the downstream basin is limited regardless of the elevation of the upstream basin. Though the overall increase in sedimentation rates in the downstream basin after capture is small in these cases (e.g. Model 3, Fig. 6: low uplift rate in 188 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 9. Model 4: effect of uplift rate (continued). Cross sections. Left panels show the time step preceding capture; right panels represent the final time step (15 My). Sediments deposited during the uplift stage are shaded. Vertical scale in m, horizontal in km. In the right panels, the highlighted time lines represent the time step preceding capture, the end of base level fall in the upstream basin, and the final stage of lacustrine deposition (solid, long dash and dotted lines respectively). The capture-triggered increase in sedimentation rates in the downstream basin is much more pronounced for the higher uplift rates (200, 400 m/Ma): the moment of capture coincides with the last stages of lacustrine infill of the upstream basin. No fluvial sedimentation is accommodated in any of the models. Model 4, Fig. 8b), the effect is focused at the inlet and will be locally significant (Model 3, Fig. 7). In contrast, the bulk sediment shift to the downstream basin will be triggered directly by capture if the remaining upstream accommodation space is small at the time of capture, and/or sufficiently reduced by the capture-induced base level fall (high uplift rates in Model 4, Fig. 8). The latter requires a large pre-capture elevation difference between the basins. In other words, lake capture is expressed in the sedimentary record by affecting base level in the upstream basin and supply in the downstream basin. The most significant increase in downstream sedimentation rates, however, is not necessarily directly linked to the capture event (Figs. 6 and 14). 4.1.2. Connected stage The upstream accommodation space is controlled by changes in the lake level as discussed above and, in the absence of tectonic subsidence, the subsidence due to sediment loading. The latter is strongly dependent on the rigidity of the lithosphere below the upstream basin: a weak lithosphere (low effective elastic thickness, Te) allows more subsidence and results in the generation of a larger accommodation space (Model 5). A strong lithosphere supporting the upstream basin leads to a sudden increase in the supply to the downstream basin, while a low Te results in a gradual increase in downstream sedimentation rates (Model 5 and 6; Figs. 11–13). 4.1.3. Continental stage in the upstream basin: fluvial incision and deposition The (post-capture) continental evolution of the upstream basin and the gateway is determined by fluvial processes which are a function of the difference between the actual and the equilibrium river gradient: the river tends to its equilibrium gradient and does so by either incision or deposition (e.g. Kooi and Beaumont, 1994). The K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 189 Fig. 10. Model 5: effect of lithosphere rigidity. a. Sedimentation rates for different values of effective elastic thickness (Te). Transition from lacustrine to fluvial sedimentation in the upstream basin is marked by stars, the horizontal bars indicate the preceding period of increasing sedimentation rates in the downstream basin. Erosion rates in the downstream basin are shown for reference. b. In the upstream basin: transition time from lacustrine to fluvial sedimentation, and fill-up time as a function of Te. t0 is the minimum time required to fill the basin and is a function of the basin volume and the supply rate. Hatch: period of fluvial sedimentation. response–incision or deposition–is determined by elevation and the degree of overfill of the upstream basin at the lacustrine–continental transition. If capture occurs in a late stage, when the upstream accommodation space is nearly filled, the upstream basin will be overfilled and exhumed, a process much dependent on the elevation difference between the basins. The larger the elevation difference, the larger the degree of overfill and the larger the subsequent river incision (Model 3, 4). If capture happens in an earlier stage, with ample upstream accommodation space, fluvial sedimentation is likely in case of low lithosphere rigidities (Model 2 and Model 6, 7 in Figs. 11 and 12). The volume of continental sediments is influenced by the initial elevation of a preexisting gateway. The results from Model 2 show that, due to the erosion-resistant barrier, the initial elevation of the gateway (and as such, upstream base level) controls the upstream accommodation space and corresponding continental sedimentation in the last stage of basin fill evolution, causing differences in the total deposited volumes (Fig. 5). This is despite the fact that the same gradient is established between the upstream and downstream basin in all three models within the first 2 My (Fig. 5b). Even after capture and completion of the lacustrine infill of the upstream basin, the barrier keeps controlling the upstream base level by forming a knickpoint in the river long profile. The mechanisms underlying the knickpoint formation promote upstream fluvial sedimentation as can be seen by comparing the results from Model 1 and 2. In Model 2, the barrier, uplifting by flexural isostatic rebound in response to erosion, separates two depocenters in the model area. In order to maintain its gradient across the erosion-resistant uplifting barrier, the river responds by upstream deposition (see also Snow and Slingerland, 1990). The larger the upstream generation of accommodation space by subsidence, the larger the accumulation of fluvial sediments. This is well expressed by the results of Model 6 (Figs. 10 and 11), where low Te values lead to large subsidence in the basins (and higher uplift of the barrier), accommodating a large volume of continental sediments compared to the high Te values. In case of Model 1, the initial elevation of the barrier is at sea level over its entire length. It subsides with the sediment loaded adjacent basins, and is buried by sediment itself. The wavelength of the subsiding area is different in this case, even though the rigidity of the supporting lithosphere is the same in both models. The river is able to maintain its gradient across the former barrier without any deposition or incision and is consequently at grade over its entire length. 4.2. Implications for the evolution of the Pannonian–Dacic area Since the Sarmatian, Lake Pannon and the Dacic Basin existed as two water masses at a relatively small elevation difference (in comparison 190 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 11. Model 5, 6: lithosphere rigidity. Cross sections at 15 My. Vertical line marks change in orientation from W–E to S–N. Bold line indicates onset of fluvial sedimentation in the upstream basin, short dash marks the shelf-slope break. a. Model 5, Te = 30 km. b. Model 5, Te = 5 km. Note the difference in volume of continental sediments with Fig. 5a. c. Model 6, Te west = 5 km, Te east = 30 km. Te transition over 60 km across the center of the model is marked schematically at the bottom of the panel. with e.g. the Ebro Basin and the Mediterranean, see Garcia-Castellanos et al., 2003), separated by an elevated barrier. The modelling results allow some inferences on the evolution of the Pannonian and Dacic Basins as a function of their connectivity. As discussed above (Section 4.1, Fig. 14), the modelling predicts that in a system such as the Pannonian–Dacic region, changes in connectivity may be recognized mainly from base level changes in the upstream basin and supply changes in the downstream basin, though the two may be separated by a time lag. 4.2.1. Pannonian Basin Constraints on base level evolution have been derived from sequence stratigraphy (see Section 2). The rapid retreat of the coastline K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 191 Fig. 12. Model 6: lateral change in lithosphere rigidity. Sedimentation rates for different Te in the western (upstream) basin, Te in eastern basin is constant at 30 km (Table 2). Transition to continental sedimentation in the upstream basin is marked by x. The lithosphere rigidity below the upstream basin influences the expression of the peak sedimentation in the downstream basin, even though its Te is equal in all model scenarios. of Lake Pannon between 6.5 and 4.5 Ma, as reconstructed by Magyar et al. (1999a) (Fig. 1) closely matches in time the 2nd order sequence boundary observed in the Hungarian part of the Pannonian Basin (Fig. 2) and suggests that this sequence boundary is indeed associated with an absolute lake level fall (rather than reduced tectonic subsidence). Estimates of its magnitude vary from 10s of meters (Sacchi et al., 1999) to 200 m (Vakarcs et al., 1994). The hiatus associated with this boundary has been dated at 5–6 My (Csató et al., 2007). Both capture (restoring connectivity) and enhanced incision by an antecedent river will affect the Pannonian lake level. In case of lake capture, the magnitude of the lake level fall would be determined by the relative elevation of the Pannonian Lake with respect to the downstream Dacic Basin at the time of capture (cf Model 3). Alternatively, in case of an existing fluvial connection, the adaptation of the river gradient in the Danube gateway to the MSC-related lowering of the sea level in the Dacic Basin would lower the Pannonian lake level as well (see also Tari et al., 1992). This raises the question, which mechanism caused the “Messinian” sequence boundary in the Pannonian Basin? The lake capture model assumes that the Sarmatian uplift defeated the river crossing the barrier and created an endorheic Pannonian Lake. Once disconnected, any lake level changes were due to local climatic factors (i.e., the balance between water influx and evaporation in the drainage area; see e.g. Juhász et al., 1999). Capture of the Pannonian Lake, restoring connectivity to the Dacic Basin, would lead to a base level fall. A capture-induced Pannonian lake level fall needs not be associated with a dry climate. To the contrary, capture was likely triggered by an increase in the precipitation/evaporation ratio in the Pannonian Basin, inducing an initial rise in the lake level. This rise might trigger reconnection by “overspill and eventually lead to a lake level fall instead (see also Garcia-Castellanos (2006) for a more detailed discussion on the parameters influencing lake capture). Large uplift rates delay or even inhibit the reconnection (Figs. 8 and 9), while the rise of the upstream lake level is a prerequisite (GarciaCastellanos, 2006). Lake level changes in the Pannonian Basin in the order of 10s of meters (Sacchi et al., 1999) imply that the Sarmatian uplift of the South Carpathian Orocline was limited, otherwise capture is impossible. However, Sarmatian apatite fission track ages along the Danube gorges which are interpreted to result from a tectonic uplift phase (Bojar et al., 1998) argue for a much larger uplift, in which case a major rise in the Pannonian lake level would have been required for capture to occur. These data are in conflict and contradict the capture model. However, lateral differences in erodibility in the South Carpathian orocline along the major strike slip faults (such as the Cerna–Jiu fault, Fig. 1) may have contributed to keeping the actual “barrier” at reduced elevation by facilitating river incision despite large tectonic uplift. In addition, rapid localized incision would reduce the isostatic rebound due to erosional unloading and thus accelerate capture. Sea level fall does not accelerate lake capture in our particular synthetic scenario. Water unloading related to sea level fall produces a barrier rebound that delays the capture process (Fig. 13), in a way identical to what Govers et al. (2009) propose for the onset of the MSC in the Mediterranean. Therefore, even if the lake level fall of the Pannonian Basin were induced by capture, it is difficult to attribute it to the sea level fall in the Dacic Basin during the MSC. Additional modelling giving more precise estimates on the balance between uplift and incision, and taking into account lateral differences in erodibility is required before the capture model can be discarded. According to the antecedent river model, the fluvial connection between the basins was maintained despite the Sarmatian uplift because the incision of the Paleo-Danube could keep up with the uplift in the belt. The Pannonian lake level was controlled by the elevation of the outlet, and kept at a more or less constant level depending on the balance between (isostatic) uplift rate of the barrier and river incision. By calculating the water balance in the Pannonian drainage area (Fig. 15), the endorheic lake size that would be attained if the Danube were blocked at the South Carpathian orocline can be predicted. The calculations show that for the present day range of precipitation and evaporation values, the predicted lake size exceeds even the maximum extent of Lake Pannon during the Sarmatian (Fig. 1, Magyar et al., 1999a). The Early Late Miocene precipitation in Western and Central Paratethys is estimated at 1200 mm/y (Böhme et al., 2006) and would lead to an even larger endorheic lake size (Fig. 15). These numbers suggest that after the Sarmatian uplift closed the open water connections between the Pannonian and Dacic Basins, a fluvial connection should have evolved in order to maintain the lake size. The low salinity of Lake Pannon during its endemic Pannonian stage is an additional argument for a persistent fluvial connection between the two basins (Kázmér, 1990) even if the outflow occurred along subsurface karstic channels (Menkovic and Koscal, 1997) and the surface expression of the Danube Gorges is younger. A pre-Pontian fluvial connection may explain the occurrence of freshwater Congeria of Meotian age derived from the Pannonian Basin in the western part of the Dacic Basin (Olteanu, 1979). The water balance calculations 192 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 13. Model 7: downstream base level change. a. Sea level curve in downstream basin for extreme scenarios (cf Table 2) b. Sedimentation and erosion rates in downstream basin compared with results from Model 3 (same setup; no sea level fall). Increase in sedimentation and erosion rates reflects increase in exposed area due to base level fall. Arrows mark capture time for the three presented models. (Fig. 15) therefore prompt further investigation of the relationship between endorheism and endemism, but suggest that endorheism is not a prerequisite for the Pannonian endemism. A permanent open-water connection would be in conflict with the well established endemism in the Pannonian basin during the Pannonian/Meotian stage (Fig. 2) since it would equilibrate the entire fauna between the two basins. Considering the decreasing salinity also in the Dacic Basin since the Sarmatian (Popov et al., 2006), even a fluvial connection between the basins might have led to fauna equilibration. This does not occur until the Pontian and would be an argument against the antecedent river model. If the Pannonian lake level lowering resulted from the deeper incision of an existing river into its gateway due to the large MSC-related base level fall in the Dacic Basin (Clauzon et al., 2005; Leever et al., 2009), the two events of base level fall can be directly correlated. This would constrain the time of the Pannonian lake level fall to the Middle Pontian (of the Eastern Paratethys), i.e. at ∼5.5 My (Vasiliev et al., 2004; Vasiliev et al., 2005). 4.2.2. Dacic Basin The models predict that the onset of a fluvial connection between Dacic and Pannonian lakes should be marked by locally increased sedimentation rates in front of the Danube gateway in the Dacic Basin, even if the reconnection occurred before the upstream Pannonian Lake completed its infill and attained its fluvial stage. A much more pronounced increase in sedimentation rates would be associated with the overfilling of the Pannonian Basin. The overfilling and bulk sediment shift can be directly related to capture only in the special case of a small remaining upstream accommodation space in combination with a large elevation difference (see Section 4.1, Fig. 14). No strong indications of enhanced sedimentation in front of the Danube gorges with respect to other locations are found in published seismic data in the western Dacic Basin prior to the Pontian (Rabagia and Matenco, 1999; Tarapoanca et al., 2007; Leever et al., 2009). This fits with the model predictions (Model 5, 6). The weak Pannonian lithosphere would allow ongoing generation of accommodation space in the upstream basin even after the end of the lacustrine stage, while K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 193 Fig. 14. Factors controlling sedimentary response to lake capture. no pronounced increase in sedimentation rates is to be expected in the Dacic Basin, even if a connection between the basins exists. Also the limited elevation difference between the basins would inhibit a strong signature of a potential reconnection (Fig. 14). Within the limitations of the 2D seismic data, enhanced sedimentation in front of the Danube gorges may be inferred starting with the Middle Pontian sea-level drop, coeval with the Messinian event (Leever et al., 2009). The enhanced sedimentation in the front of the Danube indicates that the Pannonian basin has reached an overfilled stage and that a weak, accommodating Pannonian lithosphere (Fig. 11c) no longer explains the behaviour of the system. A likely mechanism to explain the change in its “strength”–or its reduced sink capacity–would be the inversion of the Pannonian basin starting in the latest Miocene (∼6 Ma, Vrabec and Fodor, 2004; Leever et al., 2009). Modelling results from Jarosinski et al. (this volume) predict uplift in the entire Pannonian basin and show concentration of strain and differential vertical movements at the limit between the basin and the Dinarides, in close proximity with the Danube basin outlet. To determine the relative importance of sea level fall (in the Dacic basin) and basin inversion (leading to changes in lithosphere response to loading in the Pannonian basin and differential vertical motions around the South Carpathian orocline), additional data are required, such as estimates of sediment volumes for both basins and more detailed exhumation analysis of the gateway area in the South Carpathian orocline. Further modelling studies of river incision and gateway formation should take into account the effect of intraplate stresses (Cloetingh et al., 1985) and lateral changes in erodibility. 5. Conclusions Numerical modelling showed that changes in connectivity between two adjacent sedimentary basins are expressed in the sedimentary record through its effect on lake level (accommodation space) in the upstream basin and sediment supply in the downstream basin. The key factors controlling the sedimentary response are the upstream accommodation space and the elevation of the upstream basin at the time of capture. We found that lithosphere rigidity, through its control on accommodation space, is of major influence on sediment partitioning and depositional geometries. In the configuration of our model, downstream sea level changes will only affect the upstream basin if a gateway already exists. Lake level changes in the upstream basin, affecting depositional geometries by changing accommodation space, are most pronounced for a large pre-capture elevation difference between the basins. Depending on both the elevation difference and the remaining accommodation space, the capture-induced lake level fall may lead to locally strongly increased sedimentation rates and final infilling of the upstream basin. In addition to the elevation difference between the basins and assuming that the climatic conditions are steady and well constrained, the rigidity of the lithosphere is the most important parameter influencing the post-capture sedimentary evolution. It controls both the timing and the rate of the bulk sediment shift from the upstream to the downstream basin. A strong upstream lithosphere will lead to a fast and sudden shift while for a weak lithosphere the response is more gradual. The capture event itself will not lead to a strong downstream change in sedimentation rates, unless it triggers the sediment overfill of the upstream basin. This is more likely to occur for a large elevation difference between the basins. Sea level fall does not accelerate lake capture in our particular synthetic scenario. Water unloading related to sea level fall produces a barrier rebound that delays the capture process (see also Govers et al., 2009). As for the Pannonian and Dacic Basins, the modelling results suggest that, after the Sarmatian uplift of the South Carpathians, the fluvial connection between the basins was never lost. Calculations of a water balance for the Pannonian drainage area and the low salinity of Lake Pannon during the endemic Pannonian stage (Kázmér, 1990) also argue in favor of the antecedent river model. This model may or may not be in disagreement with the overall endemism of Pannonian and Meotian faunas. Alternatively, occasional incursions of Pannonian faunas in the 194 K.A. Leever et al. / Tectonophysics 502 (2011) 175–195 Fig. 15. Equilibrium endorheic surface for Lake Pannon. a. Water balance within a drainage area. b. Equilibrium surface (Aw) of a hypothetical endorheic Lake Pannon for different precipitation (P) and evaporation rates (at land, El), as a function of evaporation at the lake (Ew). Curves (a) and (b) are based on the average present day outflow Qout of the Danube at the Iron Gates of 5450 m3/s (http://www.grdc.sr.unh.edu/). If zero evaporation on land is assumed, this yields an average annual precipitation of 287 mm/y over the Pannonian drainage area (a). The actual present-day annual precipitation is 600 mm/y in Hungary (Vituki, 2002), corresponding to an El of 313 mm/y (b). Assuming the average Miocene precipitation values (1200 mm/y, Böhme et al., 2006) leads to a much larger estimate of endorheic lake size (c, d). Heavy horizontal dashed line represents maximum Middle Miocene surface area of Lake Pannon (Magyar et al., 1999a); the surface area in the present-day Pannonian drainage area below 100 and 200 m (derived from SRTM DEM) is shown for reference. These equations show that, within the range of present day evaporation rates at Lake Balaton, an endorheic lake is unlikely for any of the considered P/El ratios and suggest that a permanent outflow must have existed. Dacic basin may be explained by overspill events rather than by a permanent connection. The Messinian sea level fall in the Dacic Basin resulted in deeper fluvial incision into the Danube gateway, which caused the lowering of the Pannonian lake level; the connection between the basins was permanent from then on. The lowering of the Pannonian lake level marked the end of the 2nd order “Late Miocene sequence” (sensu Juhász et al., 1999), enhanced the final infilling of the basin and increased the sedimentation rates into the Dacic Basin. 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