5/3/16 Australopithecines ¨ ¨ ¨ ¨ ¨ ¨ Group of species with the following broad characteristics: Fully bipedal Intermediate teeth Brains slightly larger than apes Existed 4.5 – 2 mya Many species co-existed Genus Homo Australopithecines Early hominids Australopithecines !! 2 5/3/16 CLADOGENESIS OF HUMANS Homo Graciles Robusts Early hominids Gracile Vs Robust More prognathic Smaller molars Bigger canines/incisors Dished face Bigger molars Smaller canines/incisors Australopithecines!!!! More! ¤ ¤ ¤ Australopithecus afarensis This is one of the IMPORTANT ones Found in Hadar, Ethiopia (East Africa), dating back 3.4 to 3.0 million years ago. n More apelike in the skull (small brain and jutting-out face). n Considerable evidence of bipedalism. n Teeth still more primitive than genus Homo. 3 5/3/16 Type specimen: Lucy One of the most complete early hominid skeletons ever found Discovered by Don Johanson of ASU in 1974 Provide key data: Height – 3’8” Weight – 60-70 lbs Sexual dimorphism Age at death : 18-30 3.2 mya Lucy Human vs. Lucy Male afarensis vs. Lucy Sexual dimorphism is a product of sexual selection 4 5/3/16 Don Johanson AKA FSLHNTR Richard Leakey Brother of Maeve Laetoli Footprints Laetoli Southern Ethiopia 3.6 mya 2 individuals walking bipedally through ash - short stride length - no sign of knuckle prints 5 5/3/16 Just moseyin’ along Australopithecus africanus Australopithecus africanus Australopithecus africanus is found exclusively in South Africa. Dates: 3.5-2.2 mya Face: Less prognathism than previous versions Skull: less bell-shaped 6 5/3/16 Hominid Site: Sterkfontein Cave site in South Africa Contains the remains of four different A. africanus individuals All were dinner for some predator OR TRY THIS Homo Graciles Kenyanthropus Robusts Australopithecus anamensis AUSTRALOPITHECUS BOISEI É Australopithecus boisei is found in East Africa. É Sometimes referred to a Paranthropus boisei É Characteristics Ð “Hyper” Robust Huge cheek teeth Ð Large sagittal crest Ð 7 5/3/16 Note manly Fu Manchu mustache Masticatorymuscles temporalis masseter GORILLA SKULL Sagittal crest 8 5/3/16 MOLARIZARION OF THE PREMOLARS human afarensis boisei ECOLOGY OF THE ROBUST LINEAGE Ò Paleoecology É Edaphic grasslands and associated wetlands for hard food items É Adaptation Ò Robusts were highly specialized to their environment and food source POSSIBLE FOOD FOR ROBUSTUS Ò Almost entirely vegetables Ò Harder/tougher foods Ò Nuts, hard seeds, fibrous tubers 9 5/3/16 OR WAS IT?!!? Ò New research (2009) on boisei examined “usewear” of teeth É Use wear are patterns of marks and abrasions left by some hard substances Ò This research did not produce the results we would expect Ò What does this mean? É Anatomy suggests was [boisei] could eat, but not necessarily what it did eat EXTINCTION OF ROBUSTS Ò Robusts persist for ~ 1.3 million years (2.5 – 1.5 mya) Ò World-wide climate grows colder during this time Ò Grasslands dry out Ò Change may have occurred too quickly for robusts to adapt Ò Completely disappear from the fossil record Genus Homo ICE AGE Australopithecines Early hominids 10 5/3/16 TheIceAge DidNOTlooklikethis Pleistocene Epoch ICE AGE – series of warm and cold periods (8-10 degrees cooler on average) ◦ Lasts from 2 million years ago until 10,000 years ago Impact ◦ More water locked in ice than now ◦ Less water = lower sea levels (up to 400 feet in some places) 11 5/3/16 IceAgepoint#1– itwasn’talwayscold IceAge#2:Morewaterwaslockedinice. Sealevelswere200-300feetlower 12 5/3/16 Milankovich Cycles • Changestotherotation andrevolutionoftheearth aroundthesun – Eccentricity – Tilt – Precession • Evenminorchangescan havemajorimpacts • Thesearecycles– not singleevents HOMO! 13 5/3/16 Genus Homo Australopithecine s Early hominids HOMO LINEAGE Ò Definitely incorporated meat into diet in both south and east Africa Ò Co-existed with robust species Ò First appear 2.5-2.0 mya Ò Associated with stone tools Ò Existed HOMO DERIVED CHARACTERISTICS Ò Larger cranial capacity ( all > 500 cc) rounding of cranium Ò Facial prognathism is further reduced Ò Increased 14 5/3/16 HOMO SPECIES Ò H. Ò H. Ò H. Ò H. Ò H. Ò H. Ò H. Ò H. Ò H. Ò H. habilis rudolfensis georgicus ergaster erectus antecessor heidelbergensis neanderthalensis floresiensis sapiens The Origin of the Genus Homo Homo habilis oLived in Africa between at least 2 and 1.44 mya. ¡The first discovered specimens were found at Olduvai Gorge in Tanzania in East Africa. Homo habilis Physical Characteristics ◦ 500 cc brain ◦ Teeth – almost completely humanlike ◦ Still short like a chimp 15 5/3/16 The Handy Man Cranial capacity – 500 cc Teeth – almost completely human-like Most primitive of all Homo ◦ long arms ◦ Short legs ◦ Short stature overall (4.5 ft fully grown) ◦ Smaller face than Australopithecus Homo habilis Cultural Behavior ◦ Developed the Oldowan tradition— the oldest known stone tool culture. ◦ Tools were made from flint, obsidian, or quartz. Oldowan Chopper 16 5/3/16 Marrow consumption Highly nutritious Inaccessible to most carnivores First occurrence of hominids regularly eating meat Scavenging or Hunting? If Homo habilis is hunting animals, then we would expect: cut marks on bone first then carnivore marks second What we get is the exact opposite Last month! (MW) SUN MON TUES WED 10 Great Apes 12 Great Apes 17 Human Ancestors 19 Human Ancestors 24 Human Ancestors 26 NO CLASS 1 Neanderthals 3 Modern humans Final Quiz 8 Final exam @1:00 Final homework 10 THUR FRI SAT Thanksgiving Holiday 17 5/3/16 Modern Homo sapiens sapiens Homo sapiens neandertalensis Homo sapiens sapiens TIME Archaic Homo sapiens Homo ergaster / erectus Homo habilis Australopithecus More than one species? ◦ It is hard to accurately determine species differences for genus Homo H. habilis H. rudolfensis Theproblemis… Homo speciesOVERLAPintime Homohabilis Homo rudolfensis Homoheidelbergensis Homoerectus 3.0 2.0 1.0 0 MYA 19 5/3/16 H. rudolfensis ◦ Lived in Africa about 1.9 mya. ◦ Its cranial capacity is larger than H. habilis (400-800 cc), but more similar to australopiths in certain facial features. ◦ Evolutionary relationships between species of early Homo are very difficult to pin down. 1.8 million year old fossil 1.8 million years ago New discoveries…in the Middle East Dmanisi Hominids Several individuals Very physically variable…but they seem to be part of the same populations Multiple overlapping species of hominids 20 5/3/16 Homo erectus / ergaster 1.9 – 0.04 mya “Upright Man” – so named b/c they were some of the first bipedal hominids identified Brain size – 8501100 cc Low forehead Brow ridges Youngest (37,000) Oldest 1.9 my Homo ergaster Homo erectus Homo Erectus ¡ Has a larger brain than earlier hominids, and there was a slight increase in size over time. ¡ The overall brain case is still smaller than modern humans ¡ Brow ridges also developed over the eyes, possibly for support for chewing forces, or as facial or eye protection. 21 5/3/16 Homo Erectus Physical Characteristics ¡ Jaws were still large but smaller than those of earlier hominid. ¡ Body proportions are similar to modern humans. ¡ Limb proportions reflect a modern bipedal form, and suggest endurance running. Disadvantages Advantages Metabolically expensive Intelligence Lose heat more quickly Toolmaking and tool use Can complicate child birth Technological skill Problem-solving ability Language ability Social intelligence Advantages must outweigh disadvantages The evolution of larger brains 22 5/3/16 Acheulean Tools Acheulean Tools Used high quality stone Long distance exchange or transport Hunting, not scavenging SUN MON TUES 5 Neanderthals WED THUR FRI SAT 7 Modern Humans Final Quiz 12 Final Exam (13) Final Project Due 23 5/3/16 800,000 years ago Homo ergaster Homo erectus Archaic Homo sapiens Development and spread of early H. sapiens (500,000) 24 5/3/16 Homo sapiens neandertalensis Homo erectus Homo sapiens (archaic) Homo floresiensis Homo sapiens sapiens SPECIATION (200,000 years ago)! Homo floresiensis Speciation Homo neanderthalensis Homo sapiens Homo heidelbergensis (?) Homo erectus 25 5/3/16 Archaic Homo sapiens Has been found at sites in Europe, Africa, and Asia. Date from 800,000 to 200,000 years ago, a period of frequent ice ages, below average temperatures, and spreading glaciers over large continental areas. Cranial capacity – 1100-1200 cc Atapuerca, Spain 400,000 years ago Cave system in Spain First humans in Europe First evidence of burial culture? Hominids on Ice! Archaic H. sapiens in Europe had to deal with cold temperatures, ice, and dramatic weather changes 26 5/3/16 Temperature over the last 400k years Inter-glacial…its warm! Now Glacial…its cold! 27
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