Geomagnetic Polarity Time Scale (GPTS) • Berggren et al., 1985 • Berggren et al., 1995 • Gradstein et al., 2004 ODP Site 1207, northwest Pacific • Drilling Objectives: – P/E Boundary – K/T Boundary – Aptian/Albian 1 ODP Site 1207 ODP Site 1207, northwest Pacific Age/Depth 2 Correlation of strata by fossils BIOSTRATIGRAPHY Correlation of rock strata based on fossil characteristics – oldest method of determining relative age • Application – – Marine & Terrestrial Strata Relative vs. Absolute time • Basis - Evolution 1. Species appear & disappear (geologically instant) 2. Rapid distribution 3 Controlling factors on the Distribution of Organisms in the Rock Record 1. Evolution Controlling factors on the Distribution of Organisms in the Rock Record 2. Paleoecology – Environmental restrictions/preferences – facies controlled organisms - restricted to certain sedimentary environment e.g. molluscs - restricted to near shore environments Transgressive sequence (deepening upward) time 3 time 2 time 1 4 Controlling factors on the Distribution of Organisms in the Rock Record 3. Preservation (Taphonomy) – present, but not preserved (e.g. Burgess Shale) Biostratigraphic Zonations Biozone - basic unit defined or characterized by presence of fossil taxa that permit it to be distinguished from adjacent rocks • Stratigraphic Nomenclature (applies to most fossil groups) – range- stratigraphic distribution in a given section – first appearance datum (FAD) - immigration or evolution – last appearance datum (LAD) - extinction or emigration – range zone - all of the strata containing a specific species or group of species. – Taxon range zone-FAD to LAD of a single species 5 Biozones • formally named, defined, and assigned type sections Fossil ranges (species A-D) Biozones Teilzone: local FAD & LAD of a taxon Fossil ranges (species A-D) 6 Biozones Taxon Range Zone: Global FAD & LAD of that taxon Requires inference - Global correlation of horizons bearing the fossil taxon Fossil ranges (species A-D) Biozones Concurrent Range Zone: Intersection of the ranges of two or more taxa (globally defined) A B Interval Zone: two successive FADs or two successive LADs (globally defined) B A A B 7 Biozones Assemblage Zone: Characterized by 3 or more taxa in natural assemblage – fuzzy boundaries, as FADs and LADs aren't simultaneous. Fossil ranges (species A-D) Biozones Lineage Zone: defined by the evolutionary first appearance of successive taxa in a lineage (related) C B A Abundance (Acme) zone: concentration of a particular taxa 8 Additional Zonation Schemes Hypothetical Stratigraphic Section Composite Ranges (locally) • How are the absolute ages of these zones determined? 9 Potential Pitfalls in Biostratigraphy 1. 2. 3. 4. 5. 6. Reworking and Hiatuses • bioturbation: various models , downslope reworking, currents • artificial range truncation of FAD or LAD. Preservation (Taphonomy) • Dissolution, oxidation, etc. Bioprovinciality • Restricted range Migration (Emigration) • environment changes • Evolve Sampling and/or counting biases • Artificial Range truncation • Last occurrence precedes extinction 1. Low temporal sample resolution 2. alternating good and bad preservation, Taxonomic inconsistencies (misidentification) 10 Major Marine Macrofossils Graptolites - plankton Conodonts - nekton • Paleozoic • Widespread • High rates of evolution Dicranograptus nicholsoni Planktonic or epiplanktonic graptolite 11 Index Fossils: abundant fossils that are characteristic of a formation or a period of time • characteristics of ideal index fossils: – distinctive – widespread – abundant – independent of facies – evolve rapidly – short ranging Plankton 12 Estimated Mean Species Duration (Myr) calcareous nannofossils (coccoliths) • phytoplankton – autotrophic, Unicellular marine algae – Phylum-protists Coccolithophore Calcidiscus leptoporus 13 calcareous nannofossils (coccoliths) • Shell - CaCO3 – Shields-elliptical to circular (1 to 15 µm) – envelop the cell, form a sphere-post mortem disintegration usually breaks down the shell into individual shields Coccolithus pelagicus temperate Emiliania Huxleyi calcareous nannofossils (coccoliths) • Advantages 1. High rates of evolution short stratigraphic ranges 2. Morphologically more simple (easier to identify) 3. cosmopolitan species rapid dispersal rates not as diachronous 4. Abundant; reduced opportunity for sampling biases 5. Quick/simple method Braarudosphaera bigelowi 14 Cruciplacolithus phylogenetic lineage • Appear at the K/T boundary – C. Danicus • dominant genus Paleocene & Eocene Discoaster Evolution D. saipanensis D. multiradiatus • Appear in mid-Paleocene • Evolve toward less robust bodies and fewer rays 15 Eocene calcareous nannofossils Biogeography Relative Abundances as a function of latitude & depth in the N. Atlantic Climate Zones • Sub-arctic/Sub-antarctic (2-6°C) – E. huxleyi • Transitional (6-12°C) – R. clavigera • Subtropical (14-28°C) – U. irregularis • Tropical (>28°C) – G. oceanica – E. huxleyi (eq. upwelling) 16 17 Planktonic Foraminifera • Zooplankton, single cell Protozoa • Calcite Shells Morozovella Velascoensis Late Paleocene Globorotalia opima continuosa (Miocene) Globigerina yeguaensis (Oligocene) 18 Dinoflagellates • Mainly planktonic • Photoautrophic/Heterotrop hic • Virtually all aquatic environments, but mainly marine • Organic cysts • Abundant in coastal and high-latitude environments/ sediments Enneadocysta partridgei • Eocene antarctic endemic LO near E/O boundary 19 Dinocyst Ecology Yes No Integrating biozones from a variety of depositional environments? • Sections interfinger – Boundaries between facies • Coastal Regions • Correlation Methods independent of facies – Magnetostratigraphy – Chemostratigraphy? 20 Biostratigraphy in Continental Sediments Challenges • Organisms tend to be highly provincial – Plants – Vertebrates – Invertebrates (confined to water) • Fossil abundances extremely low – Dilution – Preservation North America Land Mammal Age (NALMA) Orohippus (Eocene) Hyracotherium (Eocene) 21 Graphic Correlation 1. Standard Reference Section – – 2. 3. 4. Thickest Most complete Fossil Ranges vs. depth Select second section Fossil Ranges (SRS) vs. depth Graphic Correlation 6. Cross plot the sections – 7. Common FO/LO Line of Correlation (LOC) – Regression (best fit) Are all the correlation points reliable? 22 Graphic Correlation • Determine relative Sedimentation rates • distance = rate x time d=rxt r = d/t 23
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