BLUMEA
37
227-237
(1992)
Miscellaneous notes on Southeast Asian Gramineae. VII
J.F. Veldkamp
Rijksherbarium/Hortus Botanicus, Leiden,
The Netherlands
Summary
In the
Gramineae
family
A. diminuta
reduced,
cies is described
is described in
species
(D. echidnae)
and
Aristida
andA. tenuisetulosa
cumingiana
combination
a new
made
and Saccharum
(P. vallatoria)
The
(S. elegans).
while
(A. papuana),
to A.
holathera;
in
(D. pyramidalis);
identity
and
two species
Deyeuxia
new
minuta
Lachnagrostis (L. avenacea), Neyraudia (N. mezii), Oryza (O.
made in
mites
a new
toA.
are
spe-
combinations
are
silvatica), Phrag-
var.
application
a new
amboinica L.
of Poa
is discussed.
Aristida
(Mez) C.
Aristida diminuta
of its
cause
fore
A.
simple
Trin. &
E. Hubb.
it is
described
originally
cumingiana
awn
in
occurs
as
Otherwise it is
Stipa.
a
and is said
similar
very
differ from this
to
Various authors have therefore
longer spikelets.
Africa from West
tropical
to
in the section Aristida and it
quite exceptional
Rupr.
L.
regarded
as
to
the somewhat
of A.
variety
a
there-
the three-awned
mainly by
it
East. Be-
was
cum-
ingiana.
After
came
having
gone
fair number of
a
and include those
usually reported
Moreover,
26092
to
based
be
on
Aristida
Arisdda
taria
The presence of
L).
in the whole
expected
this feature is
merely
cumingiana
trichodes
Trin. &
Nees
Walp.,
in
Ann.
Rupr.,
Hook,
Bot.
a
area
for A. diminuta
spikelets
(see
of A.
awn
a
wider range than
also
Napper, 1965).
have been made in Luzon: BS
may be
and
cumingiana
Co
s.n.
regarded
a
without prove-
as
an
distinction
occasional
as
a
species
Rupr.
Mem.
f.,
single
it be-
cumingiana,
optimistic.
too
Trin. &
cumingiana
trichodes
of A.
specimens
and their parts have
from Tanculan, Bukidnon, Mindanao, and
(Fenix)
in
spikelets
given
collections with one-awned
two
(both
nance
sport
through
clear that the dimensions of the
Acad. St. Petersb.
J.
Bot. & Kew Misc. 2
3
(1852) 753,
Syst.
nom.
VI,
7
(1842, preprint!)
141.
(1850) 101, nom. superfl.
superfl.
—
—
—
Lectotype: Cuming
Chae-
Aristida
671
(LE,
holo; K).
Aristida
per
Stipa
4
delicatula
1830
diminuta
(1949)
A.
Rich.,
Mez in Feddes
480.
Aristida
—
Appl. 13 (1966) 51,
L
Tent. Fl.
(K, L, P), Quartin
Continued
from
nom.
Blumea
Dillon
Repert.
Abyss.
s.n.
17
cumingiana
superfl.
36
—
(1991)
2
(1850) 393,
non
Kunth,
1829.
—
Syntypes:
Schim-
(P).
(1921)
208.
Trin. &
—
Rupr.
Aristida
diminuta
var. diminuta
J.
C.E.
Felix,
J.
Hubb.,
Type: Buchanan 561 (B, holo, extant?; K).
181.
Kew Bull.
Agric. Trop.
Bot.
228
Vol.
BLUMEA
Aristida cumingiana Trin.
805.
1933)
—
Type:
&
Aristida cumingiana Trin.
1933) 48.
—
Rupr.
var.
Holmes
in
auct., non Lam.:
Pilger,
Notizbl.
Bot. Gart. Berlin-Dahlem
11
(March
(B, holo, extant?; K).
uniseta
Stent & Rattray,
4931
Eyles
(K),
Icon. 5
Cav.,
Bot. 85
Domin, Bibl.
holathera
926.361-524, Kew
Fill
Proc. Rhodesia Sc. Assoc. 32
(May
165 (K).
(1799) 43,
f.
468,
t.
1, quoad Nee
s.n.
(B, MA).
(1915) 340.
—
Domin
Type:
s.n.,
2/1910 (L, holo,
no,
9404; BRI).
neg.
tenuisetulosa
tenuisetulosa
var.
2468
1992
1,
holathera Domin
Aristida
Aristida
&
Syntypes:
Aristida capillacea
Aristida
var. reducta
Rupr.
Schlieben
No.
37,
Mez in Feddes
(Pilger)
in
Pilger
Fl.
(1921)
17
Repert.
Perkins, Fragm.
147.
Philipp. (1904)
—
146.
Aristida
—
stipoides
R.Br,
Merrill
Lectotype:
329
(B, holo, PNH, extant?).
—
BS 2334
Type:
for further
(1992)
Distribution
Habitat
Uses
(widely spread
may be
find
dunes,
awns
weakly
exposed
and
&
papuana
holathera
Aristidae
as
(1980)
said that the articulation
to
indistinct,
was
Simon
the
(1992)
resulting polymorphic species
I
Philippines.
Such
occur.
were
distin-
disjunct distribution;
Veldk.,
Domin
in
may
therefore be
expected
to turn
up in
nov.
spec.
similis,
it
well.
as
partibus spiculae
in
c.
0,5-0,7-plo brevioribus,
glumis aristisque longioribus
differt.
—
columna
non
NGF 49315
Type:
Foreman) (L, holo; A, BRI, CANB, E, K, LAE, M).
acuminate,
mm.
1-1.5
long (incl.
awn
base,
c.
30
Distribution
-
-
10-12
glumes
articulated at
Habitat
m
Panicle erect,
mucronate; lower
upper
Note
Queensland), Malesia:
Pampanga, Zambales).
long drought.
in dunes dwarfed forms will
areas
ab A. macroclada Henrard
0.5-1.7
central
to
A. tenuisetulosa var. arenarioides.
Perennial. Culms 0.25-0.45
mm
spray and
to
articulated. In the
distinctly
to
dry
Henrard
Note the curious
articulata,
resistant
differences with A. tenuisetulosa from the
no
Aristida
from W Australia
tardily disarticulating. According
not or
were
New Guinea and the Moluccas
long;
(1929) 108,114.
sandbinder.
a
articulation. Lazarides
an
guished by
by
58
Rijksherb.
(1928) 627] (PR, holo; B, PNH,
The type material of the Australian Aristida holatherahas columns without
-
of
Coastal sand
-
and that the
(Henty
54B
Ilocos Norte & Sur, La Union,
Recommended as
-
Notes
a trace
In
Meded.
Rijksherb.
synonymy.
Australia
-
Philippines (Luzon: Bataan,
can
Henrard,
Meded.
arenaria auct., non R. Br.
Simon
awn
arenarioides
var.
(Mearns) [see Henrard,
extant).
not
Aristida
See
Mez
tenuisetulosa (Pilger)
Aristida
-
mm
Seasonally
Very
similar
while the column is
not
glumes
mm
mm
long;
Papua
lower internodes
long,
6-7
mm
long.
by
c.
lateral
New
mucro
inundated
to
A.
0.8-2
mm
more or
long.
Glumes
cm.
mm
mucro
long.
0.5-1
mm
Lemma 6-7
Column contorted,
less
cm
reverse,
not
straight, divergent,
Anthers
c.
1.25
mm
long.
Western Prov., Morehead.
woodland,
holathera,
articulated,
subequal,
28-30
awns
Guinea,
4
long, l(-3)-nerved,
1-nerved,
Awns
Blades 7-12.5
glabrous.
10-13
long callus), margins overlapping.
mm
5.5-8
long,
lax, few-spikeled,
but
a rare
c.
15
m
differing
altitude.
in the much smaller
conditionin A. holathera.
spikelets,
J.F.
Veldkamp: Miscellaneous
notes
DEYEUXIA
echidnae Veldk.,
spec.
Deyeuxia
laminas
Conspicue propter
setiformes
purascentem, glumas 4,5-5
truncato
longum apice
parvam
1
(c.
1.5-1.8
by
long
mm
as
the
1
c.
Habitat
Note
spines
lemmate
longis
longiores,
2,25
c.
lemmatis
2.25
-
of the
6-9
mm
mm
insertam
lemma
longis,
intra
pur-
3-3,2
mm
aristam erectam
glumas
inclusam.
together, longest
Blades involute, stiff,
long.
laxly contracted,
apex acuminate.
11
c.
ones
cm
3.5
c.
Rachilla-process
long, smooth,
straight,
awn
30
by
—
long,
cm
spikelet.
Anthers
c.
screes
without other
for its
quill-like,
1.25
c.
Glumes
0.67-0.7th,
(Wilhelm).
4450
vegetation,
m.
tuft
(the
echidna), long ligules,
ant-eater or
as
1.25
c.
long.
mm
stiff leaves
with
erosely 4-dentate,
apex truncate,
1
axis
8-spikeled.
c.
long,
mm
7-9
diam.,
mm
inserted in the upper
New Guinea, Chimbu Prov.
Papua
spiny
mm
long hairs,
mm
Conspicuous
-
relative
satis laxam
longas, paniculam
mm
long (excl. awns), purple-brown, cleistogamous.
Bare rock
-
pilis
long. Ligules
enclosed in the
long,
mm
mm
spikelet,
Distribution
the
m
diam. Panicles
hairs. Lemma 3-3.2
long
callus with
c.
0.3
c.
mm
4.5-5
Spikelets
Beauv.
ex
6-9
rigidas, ligulas
0,67-0,7 superiore parte
lowermost branches 2
smooth,
229
Gramineae
(Womersley) (L, holo; LAE, US).
Perennials. Culms
cm
Asian
nov.
4-dentato callo
in
longam)
mm
NGF 8971
Type:
Clarion
longas spiculam aequantes
mm
erose
Southeast
on
reminiscent of
vaguely
rather open,
purplish pani-
cle, and the truncate, erosely 4-dentate lemmas with relatively long callus hairs and
straight, weak,
Calamagrostis
enclosed
awns.
arundinacea Roth
—
When Schrader
nation Arundo silvatica he excluded several
dinacea,
but
accepted
latter's reference
arundinacea, the
a
superfluous
Beauvois
it
was
the
and
Jansen
would be
out to
be
a
arundinacea based
(1829)
cited both
created
to
liest
to
no
so
arun-
exclude the
previous
Arundo
Arundo silvatica is
replaced epithet.
analysis (Art. 44.1).
which is valid,
This
precludes
Deyeuxia
as
any
arundi-
arundinacea L. and Arundo silvatica Schrader,
and
legitimate
name
two
taxon
superfluous,
Deyeuxia sylvatica
adopted,"
"is
as
may
slightly changing
the Linnean type, whatever that may
turn
1).
other
heterotypic
Host
‘villosa
that
’,
in the
same
being
argued
Agrostis
to
never
the "final
epithet
that
by
been made.
the
the
type
use
villosa Vill.
the rules of
rank." The combination
D. villosa has
be
synonyms,
(1809). According
automatically typified by
but it
and
Under normal circumstances the combination
typified by
adopted
of the
(Art. 72.1.c)
a nomen novum
Art. 33, Note
he should have
have been
and
the combi-
Agrostis
specifically
Linnaeus' combination, e.g.
Calamagrostis pyramidalis
vatica is therefore
7.13
on
Agrostis
make
valid, legitimate,
(Art. 11.3)
permitted,
with Linnaeus'
Deyeuxia sylvatica.
(Art. 7.11,
and
not
heterotypic Deyeuxia arundinacea,
illustration with
However, Kunth also cited
(1786)
not
was
of
(1952).
perhaps intending
spelling
epithet
homotypic
an
He did
arundinacea L. (1753). As there was
of another
accompanied by
Kunth
and
Agrostis
use
name
that of Willdenow (1797).
(1812) published
later Deyeuxia
nacea
to
(1806) proposed
previous applications
of the
of the
priority
of the
ear-
Deyeuxia syl-
According
name
to
which
orthographic
Art.
ought
vari-
BLUMEA
230
of Schrader's
ant
Vol.
"the author of the
epithet
No.
37,
1992
1,
superfluous
has
name
Agrostis
given
arundinacea L. is the
that Kunth had the Linnean
If it is
that Kunth did
argued
could not have done
tis villosa is
genus,
considered
now
Whatever the
the
and
synonymy
is considered
to
cannot
the
Pannoniae
Bruck an
Icon. Descr. Gram.
der Leitha
superfl.
PI. 1
Schrader,
Prim. Fl.
Fl. Germ.
Galic.
1
61.
(1753)
Calamagrostis
—
Bot. Neerl.
1
presumably
in
(1952) 470,
Hb. Linn.
Deyeuxia sylvatica
(1806) 218,
1
auct.,
Austr. 4
(1809) 70,
nom.
non Beauv.
t.
Roth,
Lachnagrostis Trin.,
call-
species
of
a
Agros-
different
and
superfluous
is
can-
villosa is removed from
when
arundinacea L.
Agrostis
heterotypic
caus-
synonym
Fl. Chil.
Fund.
t. 49.
—
"In
Type:
sylvis
f.
4,
7,
superfl.,
nom.
Fl.
agrostis Scop.,
superfl.
nom.
nec
superfl.
Host
—
(1812, 160, t. 15,
f.
2,
—
Cam.,
1
ed. 2
(1789)
89.
(1771) 88,
—
(1809) (prius postve?).
arundinacea
Deyeuxia
lla-d).
—
Arundo
Calamagrostis sylvatica
—
Cinna
Jansen,
Acta
not
resolved,
Lachnagrostis
Desv. in
Lectotype:
Kunth.
Agrost. (1820) 128,
(1853) 320.
6
(1809) 28,
(LINN).
non
—
L
10.
—
Trin.
Agrostis
Lectotype: L.filiformis
L. sect.
Trin.
(Forst.) Sprengel ex
[=
L.
avena-
(Gmelin) Veldk.].
The differences between
Lachnagrostis
garded
as
are
distinct
the type
motypic
Agrostis
by Rugolo
synonym of
non
Agrasar
Nat.
128,
cf.
t.
10,
see
ex
(1787).
IDC
Paschatis.
Veldkamp (1982).
superfl.
7440;
comb.
171.
—
and
(1990)
Trin.
Gmelin and
(1791)
1
nom.
microfiche
Zeelandia et Insula
further synonymy,
2,
Vill.
Adans., Deyeuxia Beauv.,
the
1982). Recently
& Molina
(Gmelin) Veldk.,
Gmelin, Syst.
Willdenow 2208, B,
Nova
avenacea
Agrostis filiformis
Fund. Agrost. (1820)
1036),
de
Agrostis
avenacea
avenacea
(1786) 9,
Agrostis L., Calamagrostis
only slight (see Veldkamp,
generally L. filiformis Sprengel
Lachnagrostis
For
and
Tent. Fl. Germ.
LACHNAGROSTIS
As
of De-
generally
nov.
Arundo
—
arundinacea
agrostoidea Beauv., Agrost. (1812) 147,
cea
was
because he
type
Junio, Julio" (W, holo?).
...
Agrostis arundinacea L., Sp.
Gay,
the
by Kerguelen (1975).
Agrostis
required epithet
Veldk., comb.
...
Besser,
e.g.
Host is then the remaining
(Host)
pyramidalis
silvatica
type,"
time,
Deyeuxia.
Calamagrostis pyramidalis Host,
nom.
of
combination:
next
Deyeuxia
different
a
(1791),
be followed,
to
type
arundinacea L.
Agrostis
to
of
basionym
J. Gmelin
the
provide
in
belong
Calamagrostis pyramidalis
ing
is
Kerguelen
the
of the synonymy
while the material
(Art. 7.13),
argumentation, Deyeuxia sylvatica
be maintained. If
not
be the
(Vill.)
different
a
unknown in his
being
Kunth, applies
non
to
villosa
Calamagrostis
indicate
"definitely
the type concept
but auct.,
Deyeuxia sylvatica
most
case
in mind when he coined the combination.
species
not
is that ofAgrostis villosa
yeuxia sylvatica
ed
that,
It is clear from
prescribed type.
indicated a
definitely
different type," i.e. that of Schrader's Arundo silvatica. In the latter
—
(1820)
a new
genus
& Forde
Edgar
is
cited,
Hb.
(1991).
but this is
combination is
a
ho-
required.
nov.
Avena
ftliformis Forst.,
Fl. Ins.
Lachnagrostis filiformis Sprengel
—
and
has been re-
Type:
Forster
s.n.
(ex
Thunberg 2597, UPS,
cf.
Austr.
Trin.,
ex
Sprengel
in
microfiche
Hb.
IDC
J.F.
Miscellaneous
Veldkamp:
notes
NEYRAUDIA
based the
(1959)
Conert
new
Thysanolaena
The combination
Panicum
acariferum Trin.,
Homem.
(1819),
Nees'
of
species
this and
which is
a
he
so
72. However,
epithet
to
be used
mezii
Neyraudia
non
Nees:
specim.
nom.
e
Nees,
'Promontorio
superfl.
—
in
Mez,
Feddes
Nov. Act Acad. Caes.
Syng-Moon'.
Type: Meyen
s.n.,
(1962,
Repert.
—
with 2n
at most a
lower third
mm
it
17
as
=
as
name
to
under Art.
Thysanolaena
mezii
synonym. Therefore the
a
correct one.
(1921)
86.
Thysanolaenaacarifera
—
Nat. Cur.
19, Suppl.
1
auct.,
(1843) 181, quoad
Bot. Jahrb. 78
(1959) 240,
oral
(1963)
apparently they
seems
or
48.
=
Awns up
long.
Spikelets
less.
to
2
—
cm
wide. 2n
=
long.
—
Ligules
=
48. Otherwise the differences
an
ascending
1.6-2.15
1-4
mm
long,
mm
2.15-2.85
1-8
mm
glabrous.
land: Moa
1.9-2.95 times
as
3.6-6.4
Spikelets
to
O.
mm
2.5
minuta
wide, 1.45-2.05 times
cm
or
minuta
var.
upper part, naked in the
as
hairy. Spikelets
long. Sikkim, India,
long
as
3.6-4.6
Sri Lanka
Sumatra, Malaya, Java, Borneo, Philippines (Bohol, Leyte,
Mindanao, Mindoro, Negros), Celebes,
(Halmaheira, Buru);
are
upper part, naked in the
wide,
descending
a
long, glabrous
Awns sometimes absent, up
S Vietnam; Malesia:
not
recognized [Dr.
long. Philippines (Luzon, Samar, Leyte, Bohol)
Spikelets elliptic,
24.
be
warranted:
oblong,
Ligules
can
have different chromosome numbers,
comm.]. They
status
have maintained that O. minuta
Duistermaat in the herbarium could
species.
24, O. minuta with 2n
varietal
wide. 2n
lower half.
to
to a
L.
Panicles with the lowermost branches with
mm
a new
he cited
a.
—
Honda.
nov.
Panicles with the lowermost branches with
long
on
non
description, belongs
Neyraudia acarifera Conert,
between the two, but in the field
officinalis
and
distinct
two
are
Vaughan (1RRI, Manila)
slight
based
(1820),
July 1831 (B, holo), China, promontorio Syng-Moon.
and Tateoka & Pancho
1963)
officinalis
distinguish
O.
was
Roxb.
minuta Presl
Tateoka
and O.
as
specimen,
ORYZA
Oryza
on
combination exclusively
given
Veldk. is the
Leop.-Carol.
—
Nees)
latifolia (Hornem.)
Latin
new
have
to
comb.
(Janowski) Veldk.,
Thysanolaena mezii Janowski
his
available,
Nees'
(Janowski)
a
Nees
ex
latifolia
Thysanolaena
applied
was
Janowski, which is also typified by
combination Neyraudia mezii
231
&
acarifera (Arn.
for Melica
for which he gave
have been considered
might
an
novum
Conert
Gramineae
f.
Am. & Nees
acarifera
synonym of
Hook. f.
Neyraudia
Asian
Nees, 'excl. syn.'.
ex
a nomen
specimen, however,
Hook.
combination Neyraudia
Arn. & Nees
Thysanolaena acarifera
Southeast
on
New Guinea
Island) (O. officinalis)
Lesser Sunda Islands
(Merauke,
Western
b.
Prov.),
O.
(Flores),
Moluccas
Australia
minuta
var.
(Queens-
silvatica
BLUMEA
232
No.
37,
1992
1,
minuta
var.
a.
Vol.
Oryza
minuta
Oryza
manilensis
Presl,
Rel.
Haenk.
1
(1830)
J.
Philipp.
208.
Bot. 3
Sc.,
Type:
—
Haenke
219.
(1908)
s.n.
(PR, holo).
Type: BS
2194
(Ramos) (PNH, holo,
lost: BO, K, US, W).
Oryza latifolia
b.
auct.,
silvatica
var.
Type:
Poilane
silvatica
var.
839
Wall,
Oryza officinalis
Desv.
(Camus) Veldk.,
Desv.
Oryza latifolia
—
non
comb.
Camus,
nov.
Mus. Nat. Hist. Nat. 27
Bull.
ex
Watt,
Diet. Econ. Prod. 5
Note but the
auct.,
non
(1891) 501.
422.
officinalis Tateoka, BoL Mag. Tokyo 75 (1962)
Oryza latifolia
Type:
—
Oryza officinalis
—
Wallich 8635
41,
t.
18.
Watt
subsp.
(CAL, holo; K, W).
Desv.
The collection of Moa Island has very reduced
awns,
of the fertile lemma is that of O. minuta and
pubescence
f.
(1921) 456,
(P, holo).
only
c.
0.8
long,
mm
that of O. meyer-
not
iana, with which it could be easily confused.
PHRAGMITES
Linnaeus
1759)
(1754,
made the combination Arundo vallatoria Pluk. for Canna
The Arundo vallatoria used
palustris Rumph. (1743).
for
it
a name not
in Rheede
similar
australis
to a
(Cav.)
This leaves
drawing
of
It grows
along
16
(L.)
t.
6),
a name
curiously
'tubu sala' referred
a
Themeda,
to
et
not
(Lobelius, 1581)
but he did cite Plukenet
1988).
In his Introduction Pluke-
eitherArundo donax L.
There is
a
apparendy
a
no
Phragmites
few
possibilities
not
in the
(in litt.).
hollow culm, and
a
a
a
crude
in-
large plume-like
remain
as to
its
identity.
known East of Flores and grows
on
regions.
solid culm. Rumphius knew this
mentioned by Merrill
(1917).
resemble
as
'tubu sala'
Hasskarl
with which he
cymbaria (L.) Roxb.,
not even
or
specimen
the type of the combination. It is
as
Henrard is
Anthistiria
but this does
al.,
Dr. C.E. Jarvis
with
Only
Thysanolaena latifolia (Hornem.)
to
to
seasonal
Saccharum spontaneum has
21,
foot,
water.
rocky places, avoiding
De 1'Obel
from India orientalis from Alexander Brown. He said
Rumphius' plate
to
by
mentioned,
from Malabar, there called ‘Peacaram-
plant
('nostrati similis').
arundinacea
Neyraudia
a
Nicolson
(BM), according
with
us
not
was
European reed, presumably
tall grass, 12
a
florescence.
plants
Steudel
Plukenet Herbarium
sunny,
(see
,
said that he had had
was
Europe
who used the combination for
(1700)
boopu’
net
australis from
Phragmites
Adans.
(I.e.:
(1866) thought
possibly
meant
Rumphius' plate.
Honda has
solid culm.
a
Rumphius
seems
not
have known this.
This leaves
us
with what is
generally
known
as
Phragmites
Linnaeus' combination has been neglected since
Hasskarl
thought Rumphius'
which he
presumably
Merrill
(1917)
and
meant
plant
its
Fosberg (1981)
in Malesia this
del, based
Arundo karka Retz.
must
that it
publication,
identical with Eulalia
Miscanthus floridulus
single species
on
was
karka
must
refer
be what is known
(1786).
A
new
as
by
even
japonica
(Labill.) Warb.],
to a
(Retz.)
Phragmites.
Phragmites
Steudel.
himself.
Trin.
[with
but I agree with
As there is
karka
only
(Retz.)
combination is therefore
a
Steu-
required:
Miscellaneous
J.F. Veldkamp:
vallatoria (Pluk.
Phragmites
Arundo vallatoria
Type: Rumph.,
Arundo
karka
Pluk. ex
Herb.
Retz.,
(1840) 144;
Obs. Bot. 4
Trin. ex
Type: Koenig
4
Steudel,
in
s.n.
Herb.
(1743) 20,
21.
(1786)
I.e.
Hb. Retz.
Southeast
on
L.) Veldk., comb.
ex
[Stickman] L.,
Amb.
notes
—
Erianthus
Raizada
var.
moved: Art.
—
—
Type:
This
was
L.,
Mant. 2
(1771)
originally published
"amboinica Poa
panicula
Phoenix amboinica
4
115.
(1759)
—
Nomencl.
Arch.
ed.
(1798)
37.
Handb. Fl. Java 2
(1928)
42.
1,
1
2,
3
Roth,
—
L.
nov.
Jeswiet ex
Ned. Ind.
21
Not indicated
POA
Poa amboinica
Acad.
(Retz.) Steudel,
karka
Trichoon
comb.
elegans
Arch. Suikerind.
elegans Mukherjee, Lloydia
63.3).
Amoen.
(LD, holo).
Jeswiet var.
(Anderss.)
elegans Riimke,
karka
Phragmites
324.
elegans (Jeswiet) Veldk.,
ciliaris
Erianthus
233
nov.
(1754) 15;
SACCHARUM
Saccharum
Gramineae
5.
t.
(1841)
Amb.
Asian
(1958)
Backer,
(1934)
178
(ref.
229.
—
Erianthus
to Saccharum
benghalense
—
(Roxb.)
procerus
to be
re-
(BO, holo).
L.
AMBOINICA
557.
as
follows:
secunda coarctata, culmo tereti.
Rumph.
montana.
amb. 6. p. 19.
t.
7. f. 3.
Habitat in India.
Similima Poae compressae, sed Culmus
Flores
paniculati,
verticillati. Flores
teres.
quinqueflori.
Antherae rubrae. Pistilla alba."
Several papers and
notes
have been
tion of Poa amboinica, which
All agree that Linnaeus had
a
are
published
the anthers red, and the
seem to
him
exist
anymore.
which he grew the
plant.
a
Nearly
quite
species
reduced
plant
of the
specimen (Fischer,
resemblance
white.
a
have noticed the
is described and
applica-
living plant
is
were
or
species
suggested by
was
5-flow-
it does
probably
was
1934; Furtado, 1937),
new
that it
to note
spikelets
Unfortunately
to
Rumphius' plate
depicted there, Sorghum
where the
Poa compressa is nil.
shiny
able
that it
spikelets
are
as
not
sent to
seed from
from India in
the remarks
on
the
black sessile
is,
a
nitidum (Vahl) Pers.,
a
panicle
If Linnaeus had had
spikelets.
is erroneous, that
strictly 2-flowered, usually
fertile floret. In S. nitidum, moreover, the
to
the
discussing
stigmas.
Andropogoneae,
to one
so was
that the
generally agreed
all noticed that the reference
different
culms,
He cited this collector for many
the Mantissa. That he may have had
colour of anthers and
terete
stigmas ('pistilla')
It also has been
either as
by Konig,
the years
available and
specimen
with
very similar to Poa compressa but
ered,
over
summarized below.
this,
is
effuse,
he would
while the
certainly
BLUMEA
234
Thus there
are two
and the reference
Retzius
he deemed
to
to
ciliaris
do
and
and seeds
to
compared
it
probably
again (' amboinicea'
stems
(L.)
an
,
he did
Merrill
Trin.,
a
Trin.
gorically
mentioned an
refer
Linnaeus'
to
Poa ciliaris L.
Poa
name
or, at
to
(1759),
am-
of
least,
Retzius and
now
idea that Linnaeus had
Steudel
(1840)
between the
be
to
was
Eragrostis
argued,
as
done
the removal of the
Andropogon
figura
subvar.
(bold-face mine)
and
figure
name
specimen)
to
and
descriptio
(1937),
ab
that this
for Amboina, but
Eragrostis
by Steudel,"
“Poa amboinica Linn, is based
from which the
(1917)
element.
major
to
...
being merely
Eragros-
names.
transfer of the specific
a
his
ut et
Furtado
by
Rumphian
serratus
Rumphius'
or
and Druce
description (and
was
an
make the combina-
corrected).
haec enim
est
to
(' amboinensis’)
variant
said “Poa amboinica Linn.... is reduced
stated
description
ex
discrepancy
lectotypification by
(1917)
this
by Linnaeus,
...
(1889)
not
interpretation
Banks. Retzius noted that the culm
to
orthographic
hac maxime aliena est." It may be
Hackel
taken,
of the
collection
same
named
plausible
the
reference: "Errore factum
Rumphius
constitutes
one so
him and caused Trinius and Steudel
Retzius also noted the
the
was
description
from India under the
Konig
either of the
specimens
name
amboinica
Eragrostis
so
from which the
R. Br.
(L.)
species before
tion
1992
1,
that Linnaeus had communicated the binomial
suggested
sent
compressum"
From this remark
tis
from
specimen
a
be identical with the
Rottboell under that
"evidenter
plant
No.
of these elements.
has
(1934)
37,
Two camps have evolved for the
Rumphius.
on one
who thereafter
Konig,
plants
elements: the actual
received
(1786)
boinica. Fischer
to
to
each based
name,
Vol.
species
be
must
wholly
interpreted,"
amboinensis
and then
on
cate-
Rumphius'
and made the
combination Andropogon amboinicus.
From the above it will be clear that these remarks
say
that this is the
Andropogon
two
species
same
serratus
were
Effectively
Rumphius
Linnaeus'
not
(1934)
adopted
from
serratus
subvar.
serratus
them
as
was
gave
but
according
pointed
a
same
out
by
to
phrase
agreed
but for
Fischer concluded that the
Eragrostis riparia.
some
one
(I.e.: 521),
and
species, Sorghum
a
(1938)
nitidum.
the Linnean name
by
it with
differing lectotypification
pointed
out
that, since the epithet
presumably,
is
was
be retained for the
specimen
that
Eragrostis riparia (Willd.) Beauv.,
and
his request,
compared
a
trifling difference in the form of the
correct name
thought
previously lectotypified
such
to
what Hackel called
(1937).
at
with the type of
specimen
two
Code,
it should,
name,
or
lectotypified
Retzius
discussion in which he
Rumphius'
matched Retz's
as
the
Rumphius' plant from Amboina. Pilger,
found that the
forms of
two
Furtado
s.l.,
Hackel
major
Merrill continued
untrue.
and his A. amboinicus. Hubbard
s.s.
way
are
Thunb.
Merrill and many later authors have
reference in the
as
Fischer
genuinus
regarded
description,
allowed,
var.
Andropogon
involved: A.
followed Hackel and
the
as
for Poa amboinica Retz.,
non
L.,
panicle.
should be
J.F.
Veldkamp:
Fischer remarked that
identical
plant
ium
a
of Desmostachya
synonym
D.
Salisbury,
e
seminibus
Another cultivated
scribed
boell under the
hand
"
Furtado
(1937)
followed. The
unsuitability
and
nymy
two
Andropogon
Shaw
Airy
(not
a
as
Indian
species
ment
not
that
to
not to
In
revision of
be
Application
Steudel,
2)
a
the
not
be
a
in-
Rott-
lectotype should
adduced
successfully
be
against
amboinicus
Sorghum
R. &
based
Thunb.
(Thunb.
serratum
S.,
on
and the
amboinicum,
(the typical form)
serratus
un-
erroneous.
Poa amboinica in the syno-
not
Holcus
marks
citing
names
incorrect
or
(the
(1784), actually pub-
ex
O. Kuntze
Murray)
Thunb.,
serratus
now
contemporary Recommendation
the basis of
as
synonyms in
new
Since
by Hubbard).
species
should be
by Furtado,
Eragrostis riparia
Art. 32.2: "The
typified by
the
or
disagreeable,"
indigenous
adopted
in the
epithet
name
and its
because the
Australian
not
Example
species
sorghums
to
of
be
not
should
would be
an
geograph-
reference.
invalid argumay be
species
rejected merely
3: "The
does
an
a
they
the
Rumphian
said above, and is still
as
should be
names
combinations:
in Amboina and since Linnaeus
rejected merely
are
a
used
whatever," 62.1: "an epithet may
name
be-
Scilla peru-
grow in Peru."
Lazarides
et
al.
(1991)
used
Hubbard.
three options:
of the Linnean
species usually
Application
nation
invalid
nitidum and referred for the synonymy
To conclude there
1)
but
name-bringing
(see
source
inappropriate
a recent
an
geographical
refuted
viana L. is
Sorghum
to
in 1796.
by citing Andropogon
Andropogon
(Thunb.)
occurring
under the Code
it is
"Hort.
possibly
by Koenig
sent
was
published by Salisbury
name cannot
followed Fischer and,
already
taken from any
cause
have either
type' (quotation
was
as
of Brachiaria.
epithet ‘amboinica’,
This
cynosuroides,
1785
of the latter, younger synonym is then
by Murray,
analogous
species
Poa
as
inscribed in
was
that this
amboinicus (L.) Merr., but
serratum
(1947)
'indicate the
ical
S.
Rule!), argued
regarded
a
Retz. fascic. 4." It is identifiedin another
...
followed Merrill
use
He also cited
name.
earlier
slightly
a
states
and grew
ad D. Banks missis."
Konigio
a name
of the trivial
distinguished
precluded by
considered
It
should have coined the combination Sorghum
so
forms he
lished
Smith
apparently by
be allowed. The first author to select
nitidum. The
atypical form)
is
not
(1938) apparently
He cited
Konig
choice.
already published
der Sorghum
235
summarized previous literature and said that Merrill's and Fischer's
should
Hubbard
a
Poa amboinica
name
Gramineae
with the Lund sheet is in the BM
plant agreeing
interpretation
an
India Or.
pulchella Salisbury,"
Poa
as
ex
Asian
conserved in the J.E. Smith herbar-
bipinnata (L.) Stapf.
R. Brown, "Mr. Schumacher
by
is
specimen
misidentified
127.104)
Southeast
on
notes
also had obtained seeds from
Salisbury
that of Retzius. A
to
sheet
(LINN;
Miscellaneous
of the Linnean
yet made,
to
epithet
known
as
epithet
replace
to
an
Eragrostis,
E. amboinica
(L.)
Trin.
ex
Eragrostis riparia.
to
a
Sorghum,
the well-known
S. amboinicum (L.),
Sorghum
nitidum (Vahl)
a
combi-
Pers.
BLUMEA
236
of the
3) Rejection
of
presence
epithet altogether,
Konig bearing
the
As
as
treatment
Eragrostis
an
is best served
stability
a nomen
1,
1992
own
mere
grown from seeds ob-
diagnosis
from his
that Linnaeus
conjecture
it is
at
inspired
most an
all.
at
the last
by
is
name
is absent, and the
specimen
I
option,
incertae sedis under
at
intend to cite
present
and
Eragrostis
Eragrostis
in the
Sorghum
am-
forthcoming
for the Flora Malesiana.
For those who
are
grostis riparia,
convinced and insist
not
the Retzius
Those who feel that
easy
No.
Linnaeus'
the Linnaean
In fact,
species.
same
that he had
guess
boinica
as
37,
specimens (Retzius, Salisbury, Schumacher)
tained from
had had
Vol.
opportunity
Publishable
fame and
to create
the
use
it in the
to use
seems an
should be
application
to some
Unit)
specimen
of Retzius for Era-
sense
appropriate neotype (Furtado,
to
the
1937).
element now have
Rumphian
above
arguments given
as
an
LPU
an
(Least
amboinicum.
Sorghum
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H.K.
Airy Shaw,
1947.
of
Typification
new
derived from
names
persons
or
places.
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Kew
2:
35-37.
Palisot de.
Beauvois, A.M.F.J.
a-d.
Druce, G.C.
Club
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&
Edgar, E.,
1916, Suppl.
F.R.
Fosberg,
L.,
torea
1981.
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a
Furtado,
C.X.
Gmelin,
J.F.
d'une
nouvelle
agrostographie: 147, 160,
L
f.
15,
11
The
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Agrostis
in
published
name.
naturae,
Andropogoneae, in
1866. Neue
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Linn. Kew
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1937. The nomenclature
J.K.
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Bot.
Soc. Exch.
2: 621.
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Poa
1791. Systema
E. 1889.
Hasskarl,
1991.
M.B. Forde.
Fischer, C.E.C. 1934.
Hackel,
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