Floral Formulas and Floral Diagrams Floral Formulas Floral formula is a means to represent the structure of a flower using numbers, letters and various symbols. A floral formula is a compact form of summarizing information about the flower and its anatomy Floral Diagrams Floral diagrams are stylized cross sections of flowers that represent the floral whorls as viewed from above. Rather like floral formulas, floral diagrams are used to show symmetry, numbers of parts, the relationships of the parts to one another, and degree of connation and/or adnation. Such diagrams cannot easily show ovary position. 3 Floral Formulas Floral formulae were developed in early 19th century and used for the first time on a textbook by to describe characteristics of floral families. Sachs (1873) used them together with floral diagrams, he noted their advantage of being composed of "ordinary typeface". Organogenesis of Flowers (1973) takes advantage of floral formulae and diagrams to describe the ontogeny of 50 plant species (embryonic development). . 4 Floral Formulas Modern textbooks use floral formulae as a descriptive. Floral formulas (formulae) are useful tools for remembering characteristics of the various angiosperm families. Their construction requires careful observation of individual flowers and of variation among the flowers of the same or different individuals. 5 Floral Formula Symbol 1 The first symbol in a floral formula describes the symmetry of a flower. (*) Radial symmetry – Divisible into equal halves by two or more planes of symmetry. (x) Bilateral symmetry – Divisible into equal halves by only one plane of symmetry. ($) Asymmetrical – Flower lacking a plane of symmetry, neither radial or bilateral. 6 Floral symmetry Floral Formula Symbol 2 The second major symbol in the floral formula is the number of sepals, with “K” representing “calyx”. Thus, K5 would mean a calyx of five sepals. Floral Formula Symbol 3 The third symbol is the number of petals, with “C” representing “corolla”. Thus, C5 means a corolla of 5 petals. 8 Floral Formula Symbol 4 The fourth symbol in the floral formula is the number of stamens (androecial items), with “A” representing “androecium”. A∞ (the symbol for infinity) indicates numerous stamens and is used when stamens number more than twelve in a flower. A10 would indicate 10 stamens. 9 Floral Formula Symbol 5 The fifth symbol in a floral formula indicates the number of carpels, with “G” representing “gynoecium”. Thus, G10 would describe a gynoecium of ten carpels. 10 Floral Formulas Using the 4 whorls of floral parts as our reference Calyx (sepals) = K Corolla (petals)= C Androecium (stamens) = A Gynoecium (pistils) =G Basic Floral Formula *, K5, C5, A∞, G10 Radial symmetry= *, 5 sepals in the calyx= K5 5 petals in the corolla =C5 Numerous (12 or more) stamens = A∞ 10 carpels = G10 12 Floral Formulas Number of Parts K5 = calyx of 5 sepals C6 = corolla of 6 petals A10 = androecium of 10 stamens G2 = gynoecium of 2 carpels C0 = corolla missing A4[5] = androecium usually of 4 stamens, rarely 5 Floral Formulas Symmetry K = calyx actinomorphic (radially) C = corolla actinomorphic Kz = calyx zygomorphic (bilaterally) Cz = corolla zygomorphic Floral Formulas Number of Parts (a little more complicated) A4+2 = androecium of 6 stamens, 4 in 1 set, 2 in another G2-3 = gynoecium of 2 or 3 carpel Ax = androecium of a low unstable number (about 1020) stamens G∞ = gynoecium of high unstable number, more than 20 carpels Floral Formulas Fusion of Parts (Connation): is indicated by a circle around the number or the number inside parentheses. The number represents the the parts involved. K4 = calyx of 4 separate sepals C3 = corolla of 3 separate petals K(4) = calyx of 4 fused sepals G(2) = gynoecium of 2 fused carpels G4 = gynoecium of 4 separate carpels Floral Formulas Fusion of Parts (cont.) A5G2 = androecium and gynoecium fused by their upper parts K C A = calyx, corolla, and androecium united into a hypanthium Floral Formulas Ovary Position A line below the carpel number indicates the superior position of the ovary with respect to other floral parts. A line above the carpel number indicates the inferior position of the ovary with respect to other floral parts. G = ovary superior G = ovary inferior; flower epigynous -G- = ovary half-inferior G = ovary either superior or inferior Floral Formulas Given: 5 1+(2)+2 9+1 1 K Cz A G Floral Formulas K5 Cz1+(2)+2 A9 +1 G1 Translation Calyx of 5 separate sepals Corolla zygomorphic (bilaterally symmetrical); of 5 petals, 2 fused together, 2 free and forming a pair, and a 5th different from the others Androecium of 10 stamens, 9 joined by their filaments, the 10th free Gynoecium unicarpellate, the ovary superior Interpreting Floral Formulas Practice (5) K (5) 2 or 2+2 (2) Cz A G Interpreting Floral Formulas Practice 3-5 K 5 ∞ (5-8) C A G Interpreting Floral Formulas Practice 4 K 4 4+2 (2) C A G Interpreting Floral Formulas Practice 4 K 4 8 (4) C A G Interpreting Floral Formulas Practice (5) K (5) 5 (2) C A G Interpreting Floral Formulas Practice (5) K 5 2+2 (2) Cz A G Writing Floral Formulas Practice Given the following description, write the floral formula: Flowers bilaterally symmetrical 5 fused sepals 5 fused petals 4 stamens, 2 in one set and 2 in another Superior ovary, 2 fused carpels Writing Floral Formulas Practice Given the following description, write the floral formula: 4 sepals 4 petals 8 stamen Calyx, corolla, androecium form a hypanthium Inferior ovary, 4 fused carpels Writing Floral Formulas Practice Given the following description, write the floral formula: 3-5 fused sepals 5 petals More than 20 fused stamens Superior ovary, 5-8 fused carpels Writing Floral Formulas Practice Given the following description, write the floral formula: 5 fused sepals 5 petals 10-many stamens, sometimes 4 Superior ovary, 1 carpel Writing Floral Formulas Practice Given the following description, write the floral formula: 5 fused sepals 5 petals, zygomorphic 10 stamens Superior ovary, 1 carpel Writing Floral Formulas Practice Given the following description, write the floral formula: 4-5 fused sepals 4-5 fused petals, zygomorphic 4 stamens in 2 sets of 2 Hypogenous, 2 fused carpels Floral Diagrams Floral diagrams are stylized cross sections of flowers that represent the floral whorls as viewed from above. Rather like floral formulas, floral diagrams are used to show symmetry, numbers of parts, the relationships of the parts to one another, and degree of connation and/or adnation. Such diagrams cannot easily show ovary position. 33 Floral Diagram Symbols I 34 Floral Diagram Symbols II 35 Sample floral diagrams 36 Sample Floral Diagrams Described 37
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