OSR Yield Physiology & Oilseed Yield Enhancement Network Sarah Kendall, ADAS Outline • Yield Physiology • Oilseed Yield Enhancement Network Components of yield Yield = number of seeds x seed weight Components of yield 7 3 t/ha 4 t/ha 5 t/ha existing crops Seed weight (mg) 6 5 4 3 2 50000 100000 Seeds/m2 150000 Seed number determination Determined by photosynthesis during a 200-300oCd period after flowering (2-3 weeks) Pod & seed numbers 30 Seeds per pod 25 20 15 10 5 0 0 5000 10000 pods/m2 15000 Pod & seed numbers 30 100000 80000 20 60000 15 40000 10 20000 5 0 0 5000 10000 pods/m2 0 15000 seeds/m2 Seeds per pod 25 Effects of canopy size on yield Canopy size (GAI) Pods/m2 Seeds/pod Seeds/m2 Thousand seed weight (g) Seed yield (t/ha) Final crop weight (t/ha) Harvest index Optimum canopy 3 6800 14 95,200 4.8 4.8 14.5 30 Large canopy 5 8400 10.8 90,720 4.8 4.4 15.5 26 Determination of seed size Seed Filling Seed size Seed number 14 12 seed Biomass (t/ha) pod wall 10 8 stem mid-flowering Leaf 6 4 2 0 15-Mar 14-Apr 14-May 13-Jun 13-Jul Duration of seed filling 70 Days of seed filling 65 60 55 50 45 40 35 30 15 17 19 21 Mean temperature (oC) 23 25 Duration of seed filling 70 Days of seed filling 65 60 55 50 45 40 35 30 15 17 19 21 23 o Mean temperature ( C) 25 Photosynthesis 1 Photosynthetic rate 0.8 0.6 0.4 0.2 0 Leaf Pod Stem Dense canopies • Fewer leaves • Poor light penetration to leaves & lower pods • Increased lodging • Less seed filling Survey of OSR rooting (RLD) • 40 crops • 2004 – 2013 • Av. rooting inadequate below 40 cm www.adas.uk White et al., 2014 Survey of OSR rooting (RLD) • 40 crops • 2004 – 2013 • Av. rooting inadequate below 40 cm www.adas.uk White et al., 2014 Oil & protein contents • Protein formed early • Oil formed late • (increases from 15% to 45% during 2nd half of seed filling) • Oil content reduced by high temperature, disease, frost • Oil contains twice as much energy as starch or protein Oilseed YEN Aims: • To identify arable innovators and • Support their innovating Activities: 2. Competition 3. Analysis 4. Networking 1. … open, absolute & % of potential yield for field & plot … understand yield drivers & constraints … idea sharing, conference Ultimately … • Platform for industry-science interaction … & synergies • Accelerate yield improvement Bridging the Technology Gap Bio-physical potential ? 9 OSR Yields in UK tonnes per hectare 8 7 6 5 ‘Technology Gap’ = new systems to increase capture +/or conversion of ‘Technology Gap’ = resources new systems to increase capture +/or conversion of resources ? ? 4 3 2 1 0 2000 2005 2010 2015 2020 Understand yield formation Water Solar Energy Resources mm TJ / ha Yield = x Capture % % x Conversion tonnes / TJ t/ha/100mm Understand yield formation • OSR yield formation is quite well understood (Berry & Spink, 2006) • Yield is sink limited • must maximise number of seeds set • Optimum canopy size at flowering • Prolonged canopy survival to increase oil • Few stem reserves translocated to seed • Ultimately maximising capture & conversion of primary resources (light & water) is critical • Some phases of growth more important than others Estimate Potential Yields Wheat 2015 • Develop a simple, but plausible, model of yield and oil content • Use model to estimate potential yields • Seed yield, oil content, gross output • Previous ADAS papers have estimated; • Potential seed yield with current technology: 6.5 t/ha • Ultimate yield potential for UK of 9 t/ha • Current record OSR yield: 6.5 – 7 t/ha • Quantify how yield potential affected by: • Location; Season; Soil type • Improve model as understanding increases t/ha Yield Competition • Quantify maximum yields possible with current technology • Break world records! • Identify what crop traits and husbandry treatments associated with high yields • Identify yield limiting factors • Calculate return on investment of high yielding crops • Competitions for; • Best overall gross output • Best % of potential gross output Yield Competition – Information collected • Soil description • Weather data (Morecs or local station) • Photos for plant establishment and early light capture • Hopefully also satellite images • Key developmental dates • Stem extension, flowering, senescence • Whole crop sample ~2 weeks pre harvest • Total biomass, harvest index, nutrient uptake • Seed sample off combine • oil, protein, seed size • Husbandry details Yield Competition – Calculations • Potential yield for entrant’s field • Description of the entered crop • Yield components • Oil/protein • Total biomass, Harvest index • Nutrient uptake • Most likely factors constraining yield • E.g. light capture or water capture • Gross margin • Using standard figures Yield Competition – Analysis • The OSR YEN dataset will have great value • Which will increase each year • Data base will need careful management • Identify the most common factors constraining yields in the UK • Enabling research to be correctly focussed • Identify husbandry practices associated with high yields • Identify which plant traits must be improved Summary • Canopy management important for maximising yield – optimal pod number and maximum seed number. • Oilseed YEN initiated to identify innovators and increase understanding of yield drivers. • Estimate potential yields and understand yield limitations. 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