Teaching Guide to Exoskeletons Overview: ● ● ● ● ● Roles of An Exoskeleton ○ a protective covering over the body ○ a surface for muscle attachment ○ a water-tight barrier against desiccation ○ a sensory interface with the environment. It is a multi-layered structure. It is a “non-cellular structure”, made of chitin ○ When chitin combines with the protein sclerotin it creates a multi-layerd exoskeleton with the basic properties of plastic: tough yet light and flexible. The exoskeleton is divided into smaller sections/plates called “sclerites”. These are fused together to form distinct body regions and their appendages. They are joined by membranes of pure chitin or are separated by grooves called sutures. This allows for greater flexibility like a knight's armor. These plates are NOT the separation between body segments, they can be seen on the underbelly of some beetles. Sometimes called “the secret to insect success” Water Proofing: ● The epicuticle is the outermost part of the cuticle. Its function is to reduce water loss and block the invasion of foreign matter. The innermost layer of epicuticle is often called the cuticulin layer, a stratum composed of lipoproteins and chains of fatty acids embedded in a protein-polyphenol complex. An oriented monolayer of w ax molecules lies just above the cuticulin layer; it serves as the chief barrier to movement of water into or out of the insect's body. In many insects a cement layer covers the wax and protects it from abrasion. ● The waterproof wax covering is secreted by glands embeded in the lower levels of the exoskeleton. Helps to retain body moisture. Pros & Cons of Exoskeletons Pro Con Protection Small Size Body Part Regeneration Transformation Coloration Camouflage Unique modifications Molting Limited to Small size Arthropod/Beetle Strength: The exoskeleton penetrates the body forming a complex framework of internal supports that provide attachments for powerful muscles that drive the mouthparts, legs and wings. Exoskeletons are sensory too, not just antennas: On beetles the outer layer of the exoskeleton (cuticle) are “setae”. They sometimes are hair like in nature and are sensory in nature. They transmit tactile and odor information to the nervous system. Can also be protection if clumped in strategic places to reduce abrasion and wear and tear. Blue Death Feigning Beetles: The wax filaments that they secrete makes them blue! These wax filaments are secreted by glands whose spigots open at the tips of the knoblike bumps on the wing covers (elytra). The filaments spread over the upper surface of the body, creating a meshwork that functions as deep sea diving suit in reverse, sealing off the beetle from its dry surroundings to keep water inside the body. Scarab Beetles: These guys eat ant larva and ant colonies can be dangerous places to be as an intruder. Scarab beetles have adapted by having exoskeletons that allow them to tuck away sensitive antennae and mouthparts to keep them away from angry ants. Also, the surface of their exoskeleton has nooks and crannies, like an english muffin, that absorbs the odor of the ants nest so they don’t alarm the ants. (They also have glands that let off an odor that calms angry ants!) Why an exoskeleton: For terrestrial animals, endoskeletons (internal skeletons like a humans) are more efficient for animals of a larger size. Exoskeletons are more efficient with a smaller body size. From an engineer's POV an insect is a set of hollow tubes. For the weight, a tube is more resistant to buckling than is a solid rod. But, as an animal is bigger and bigger, hollow tubes can no longer support that weight. Skeletons of larger animals have more stress from impacts and there is cushioning from soft matter around the skeleton.
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