Plant Anatomy and Nutrient Transport Chapter 42 Tracheophytes • • • • • • • • • • • Tracheophytes – plants w/ vascular tissue Tracheophytes show indeterminate growth Divided into 2 groups: Angiosperms (flowering plants) Gymnosperms (non-flowering plants) Shoot – one or more stems w/ leaves Meristematic tissue – growing tissue at stem or root tip Primary growth – growth in length Secondary growth – growth in width Apical meristems are found in all green plants Lateral meristems – found w/in stems, causes increase in width Tracheophytes Growth Patterns • Annuals – grow for one year, reproduce & die • Biennials – grow for 2 years, reproduce 2nd year & die • Perennials – grow for many years, reproduce for many years & die • Angiosperms are divided into two groups (monocots and dicots) based on the structure of their flowers, leaves, vascular tissue, roots, and seeds Monocots & Dicots • Monocots have one cotyledon (seed leaf) – e.g. grasses, lilies, palms, orchids • Dicots have two cotyledons (seed leaves) – e.g. deciduous trees (drop leaves in winter), bushes, many garden flowers Types of stems • Herbaceous stems – small, soft nonwoody stems. ex. Tomato • Usually found in annuals & biennials • Woody stems – vascular plants w/ large woody bodies ex. Trees, shrubs Stem Function • Support plant • Translocation of materials between roots & leaves • Storage of food & water • Some photosynthesis in herbaceous stems • Some help produce new plants • Hold leaves in position, max exposure to sunlight Stem Structures • Cuticle – waxy, waterproof coating • Epidermis – cells under cuticle, aid in protecting stem • Vascular bundles – composed of collenchyma cells • Pith – storage & support tissue near center • Monocots no pith, dicots do Plant Tissues • Collenchyma – column shaped cells w/ thick walls. Support soft stems as well as xylem & phloem. Living • Parenchyma – cells w/ round x-section, thin walls. Living • Pith in herbaceous dicots contains parenchyma cells w/ chloroplasts • Sclerenchyma – non-living cells, w/ lignin Arrangements of Vascular Bundles • Herbaceous monocots – bundles arranged randomly ex. Corn • Herbaceous dicots – bundles arranged in a circle ex. Bean Woody Stems • Tissues arranged in rings • Vascular cambium – forms new plant tissue. Phloem toward the bark, xylem toward the middle • Primary xylem & phloem – ring of bundles @ outer edge of stem for transport • Secondary xylem – develops into wood of the stem Woody Stems • Bark & epidermis protects against disease & mechanical injury • Cork cambium – beneath epidermis forms cork or outer bark • Pith – heartwood at center, stores water & minerals • Annual rings – separates summer & spring wood • Lenticels – cracks in outer bark, allow gas exchange • Twig – finer branches of tree, bear flowers, fruits Twig Structure • Buds – groups of bud scales (modified leaves) • Leaf scars – regions where leaves have fallen off • Terminal buds – become leaves at tip of stem (from apical meristem) • Lateral buds – become leaves at side of stem Roots • 3 basic types of roots: • 1. Taproot – thick, fleshy root. Stores water & starch. Ex. Carrot • 2. Fibrous roots – threadlike roots ex. Grass • 3. Adventitious – help prop up the plant ex. corn Root Functions • • • • 1. Anchor plant into soil 2. Store food material 3. Transport materials 4. Absorb water & minerals • 5. Absorb O2 in aerated soils Root Structures • Root cap – layer of dead cells at root tip, protection • Endodermis – contains pericycle. Controls movement of materials • Pericycle – meristematic cells, produces new cells when needed • Meristematic region – cells actively divide • Elongation region – area where cells become longer • Differentiation region – area where cells differentiate & become mature • Root hairs – root epidermis projections. Aid in Root Relationships • Symbiotic relationships help plants acquire nutrients • Mycorrhizae (root-fungus symbiosis) – Fungus helps plant extract and absorb minerals – Plant supplies fungus with sugars, amino acids, and vitamins • Legumes (peas, clover, soybeans) house nitrogen-fixing bacteria in nodules (swellings) on their roots – Plant supplies bacteria with sugars – Bacteria supplies plant with usable nitrogen Transport • Capillary action – water creeps up xylem (small dia.) due to cohesion & adhesion • Only will pull water up several cm • Transpirational pull – loss of water through leaves pulls water up through the plant • Major way of upward transport • Root pressure – pressure of water entering roots forces water up the plant Xylem • Conducts water and minerals from roots to shoots • Tracheids – long, thin, thick walled, dead cells • Vessels – short, wide, thick walled, dead cells • Fibers – long, thick walled, dead cells, strengthen xylem • Sclereids – variable shaped, thick walled, dead cells, strengthen xylem • Parenchyma – un-specialized living cells. Store starch, help w/ lateral conduction Phloem • Conduct sugars, hormones, aa & some minerals • Sieve cells – large, column-like living cells. Cytoplasm is continuous w/ another sieve cell • Cytoplasmic bridges allow direct pathways from roots to leaves • Companion cells – Small, column-shaped, living cells. Care for sieve cells • Phloem fibers – support phloem structure Mass Flow • Mass Flow – aka pressure flow or solution flow. Idea that osmotic pressure w/in sieve cells force mass of solutes to flow along a phloem gradient for mass. • Differences in water pressure drive sap through phloem sieve tubes • Sugars are transported from sources to sinks
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