SUCCESSION TO OLD GROWTH IN PENNSYLVANIA 9 OVERVIEW Students learn about forest succession, the characteristics of old-growth forests and about some of the old-growth forests that remain in Pennsylvania. In groups, students design a travel brochure about a specific old-growth forest in Pennsylvania. BACKGROUND Change Over Time: Processes that Affect the Forest Succession Like all ecosystems, forest ecosystems are ever-changing. After a natural or human induced disturbance the developmental sequence or stages of re-growth that a forest follows is known as succession. During succession, the forest adapts to the environment and simultaneously adapts the environment to itself. As the plant community changes the animal community changes with it in response to changing habitat conditions. Succession is often referred to as either primary or secondary succession. Primary succession occurs immediately after a catastrophic disturbance that leaves a site without any plant or animal life and sometimes without soil. An example of primary succession in parts of northern Pennsylvania would be what occurred after the receding of the glaciers about 10,000 years ago. However, if the disturbance is less severe, only disrupting rather than destroying the biotic community, then secondary succession follows. Examples of such disturbances include fires, windstorms, landslides, floods, disease, insects, invasive plants, the effects of grazers and browsers, and timber harvesting. Pioneer species that are shade-intolerant generally predominate in early successional stages after a disturbance in a forest. Since they germinate, grow quickly, and are the first to inhabit the area, they are called pioneer species. Examples include aspen, grasses and forbs. Pioneer species are replaced after a time by shade-tolerant, longer-lived plants. The seedlings of these shadetolerant trees become established beneath the forest canopy and eventually penetrate the canopy and become overstory trees. Some examples of these species in Pennsylvania’s forests include sugar maple, American beech and eastern hemlock. Section II — Activity 9 After a disturbance, and if it is not disturbed again, the forest stand will proceed through five general structural growth phases: seedling, sapling, pole, mature or sawtimber, and finally old growth. There are a limited number of old growth forests remaining in Pennsylvania — about 15,000 acres — scattered in about fifty locations throughout the state. Those that do exist are mainly hemlock and white pine. Most of these remnants are found within Pennsylvania’s state park system, the Allegheny National Forest, or are held by private conservancies. PROCEDURES 1. Hand out copies of the Student Page “Successional Stages” and discuss the process of ecological succession with students. If possible, take students outside to an area that illustrates several stages of succession. 2. Explain that during succession the forest adapts to the environment while simultaneously adapting the environment itself. Through the process of succession a forest will proceed through five general structural or growth phases: seedling, sapling, pole, mature or sawtimber, and finally old growth. There are certain plant and animal species that are more likely to be found at each successional stage. 3. Ask students how they would define “old-growth forest.” Ask whether they know of any locations in the United States where old-growth is located. Have students use the old growth poster for locations and information. Are they aware of any oldgrowth forest in Pennsylvania? Have they ever been in an old-growth forest? 4. Share with students that the definition of old growth is complex and varies from region to region within the United States. In general though, GRADE LEVELS: Grades 5–10 OBJECTIVES: Students will (1) review the process of forest succession, (2) discuss the characteristics of an oldgrowth forest, (3) research a specific old-growth forest in Pennsylvania, and (4) design a travel brochure about that forest. PENNSYLVANIA PROPOSED ENVIRONMENT & ECOLOGY STANDARDS ADDRESSED: 4.2.7 C. Explain natural resource distribution. 4.6.7 A. Explain the flows of energy and matter from organism to organism within an ecosystem. B. Explain the concepts of cycles. C. Explain how ecosystems change over time. 4.2.10 C. Analyze how man-made systems have impacted the management and distribution of natural resources. 4.6.10 A. Explain the biotic and abiotic components of an ecosystem and their interaction. B. Explain how cycles affect the balance in an ecosystem. C. Analyze how ecosystems change over time. ESTIMATED DURATION: Two to three 50-minute class periods. MATERIALS NEEDED: Books, etc. for research on old-growth; paper; writing and art materials needed to create a travel brochure. Succession to Old Growth in Pennsylvania 89 5. 6. 7. 8. old-growth forests contain the oldest and largest individuals of certain tree species. In Pennsylvania they are usually eastern hemlock and occasionally white pine. These trees can be 300–500 years old or more. Oldgrowth forests also tend to have trees of varying heights, creating a multilayered canopy. Because of the age of the forest, dead trees are an important component of an old-growth forest, both those fallen and those still standing as snags. Old-growth forests provide a unique habitat that certain species of plants and animals rely on for survival. Briefly discuss the controversy revolving around whether or not to log oldgrowth forests in the Pacific Northwest. Explain to students that in Pennsylvania, while we don’t have much remaining old-growth forest, most of what we do have is protected as part of our State Park system, state forest, national forest system, or by private conservancies. Divide students into four groups. Explain that each group will research a different old-growth forest in Pennsylvania and then design a travel brochure about that forest. The purpose of the travel brochure will be to inform people about this particular old-growth forest, to encourage people to visit the forest, and to inform of its unique value. Some of the needed information can be obtained from travel guides and other books about Pennsylvania. In addition, you could write to the agency that manages the forest. Students should also include some visuals in the brochure; if actual photographs of the forest are not obtainable, perhaps a series of drawings (real or abstract) can be produced. A map and directions of how to get to the forest would also be important to include. Assign each group one of the following old-growth areas to research. Give students time to research their forest and develop their brochure. Glens Natural Area, Rickets Glen State Park Luzerne County Large track of old growth forest of eastern hemlock, white pine and mixed oak; many of the virgin hemlocks are more than 500 years old. 90 Succession to Old Growth in Pennsylvania Forest Cathedral Natural Area, Cook Forest State Park Clarion & Forest Counties Old-growth stand of white pine and eastern hemlock; many trees are over 300 years old. Forrest H. Dutlinger Natural Area, Susquehannock State Forest Clinton County Old-growth stand of virgin eastern hemlocks. Heart’s Content Scenic Area, Allegheny National Forest Warren, McKean, Elk & Forest Counties 121-acre tract of virgin white pines, eastern hemlocks and American beech. 9. When the brochure is completed, have each group share with the class their travel brochure and what they learned about their old-growth forest. Display the brochures in the classroom. EXTENSION ■ Have students design a poster to accompany their brochure. (Or divide the students into eight groups rather than four; have two of the eight groups research the same old-growth forest, but one group designs the brochure and the other group designs the poster for that forest.) ■ Create a bulletin board display about old-growth forests or have the students create one. The brochures and/or posters developed by the groups can be included as part of the bulletin board. ■ Have students research some of the plant and animal species that depend on mature and old-growth forests in Pennsylvania. Some examples would include the northern flying squirrel (Glaucomys sabrinus), northern goshawk (Accipiter gentilis), and pileated woodpecker (Dryocopus pileatus). ■ If possible, take students on a field trip to an old-growth forest in Pennsylvania. ■ Have students research the details of the old-growth controversy in the Pacific Northwest. ASSESSMENT Have students share their brochures in presentation form to other classes and/or to their family members. Section II — Activity 9
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