Winners of the Nobel Prize for Physiology or Medicine Year Winner 1901 Emil Adolf von Behring Allvar Gullstrand Alexis Carrel Charles Richet Robert Bárány Jules Bordet Schack August Steenberg Krogh "for his discoveries relating to immunity" "for his discovery of the capillary motor regulating mechanism" Archibald Vivian Hill "for his discovery relating to the production of heat in the muscle" Otto Fritz Meyerhof "for his discovery of the fixed relationship between the consumption of oxygen and the metabolism of lactic acid in the muscle" Sir Frederick Grant Banting & John James Rickard Macleod Willem Einthoven "for the discovery of insulin" "for the discovery of the mechanism of the electrocardiogram" Johannes Andreas Grib Fibiger Julius Wagner-Jauregg Charles Jules Henri Nicolle Christiaan Eijkman Sir Frederick Gowland Hopkins Karl Landsteiner Otto Heinrich Warburg Sir Charles Scott Sherrington & Edgar Douglas Adrian Thomas Hunt Morgan "for his discovery of the Spiroptera carcinoma" "for his discovery of the therapeutic value of malaria inoculation in the treatment of dementia paralytica" "for his work on typhus" "for his discovery of the antineuritic vitamin" "for his discovery of the growth-stimulating vitamins" "for his discovery of human blood groups" "for his discovery of the nature and mode of action of the respiratory enzyme" "for their discoveries regarding the functions of neurons" "for his discoveries concerning the role played by the chromosome in heredity" 1903 Niels Ryberg Finsen 1904 Ivan Petrovich Pavlov Robert Koch Camillo Golgi & Santiago Ramón y Cajal Charles Louis Alphonse Laveran Ilya Ilyich Mechnikov & Paul Ehrlich Emil Theodor Kocher 1910 Albrecht Kossel 1911 1912 1913 1914 1915 1916 1917 1918 1919 1920 1921 1922 1923 1924 1925 1926 1927 1928 1929 1930 1931 1932 1933 "for his work on serum therapy, especially its application against diphtheria, by which he has opened a new road in the domain of medical science and thereby placed in the hands of the physician a victorious weapon against illness and deaths" "for his work on malaria, by which he has shown how it enters the organism and thereby has laid the foundation for successful research on this disease and methods of combating it" "[for] his contribution to the treatment of diseases, especially lupus vulgaris, with concentrated light radiation, whereby he has opened a new avenue for medical science" "in recognition of his work on the physiology of digestion, through which knowledge on vital aspects of the subject has been transformed and enlarged" "for his investigations and discoveries in relation to tuberculosis" "in recognition of their work on the structure of the nervous system" "in recognition of his work on the role played by protozoa in causing diseases" "in recognition of their work on immunity" "for his work on the physiology, pathology and surgery of the thyroid gland" "in recognition of the contributions to our knowledge of cell chemistry made through his work on proteins, including the nucleic substances" "for his work on the dioptrics of the eye" "[for] his work on vascular suture and the transplantation of blood vessels and organs" "[for] his work on anaphylaxis" "for his work on the physiology and pathology of the vestibular apparatus" 1902 Sir Ronald Ross 1905 1906 1907 1908 1909 Rationale 1934 George Hoyt Whipple, George Richards Minot & William Parry Murphy "for their discoveries concerning liver therapy in cases of anaemia" 1935 Hans Spemann "for his discovery of the organizer effect in embryonic development" 1936 Sir Henry Hallett Dale & Otto Loewi 1937 Albert Szent-Györgyi von Nagyrapolt 1938 Corneille Jean François Heymans 1939 Gerhard Domagk 1940 1941 1942 Carl Peter Henrik Dam 1943 Edward Adelbert Doisy 1944 Joseph Erlanger & Herbert Spencer Gasser "for their discoveries relating to chemical transmission of nerve impulses" "for his discoveries in connection with the biological combustion processes, with special reference to vitamin C and the catalysis of fumaric acid" "for the discovery of the role played by the sinus and aortic mechanisms in the regulation of respiration" "for the discovery of the antibacterial effects of prontosil" "for his discovery of vitamin K" "for his discovery of the chemical nature of vitamin K" "for their discoveries relating to the highly differentiated functions of single nerve fibres" 1945 Sir Alexander Fleming, Sir Ernst Boris Chain & Howard Walter Florey "for the discovery of penicillin and its curative effect in various infectious diseases" 1946 Hermann Joseph Muller Carl Ferdinand Cori & Gerty Theresa Cori, née Radnitz 1947 Bernardo Alberto Houssay 1948 Paul Hermann Müller Walter Rudolf Hess 1949 António Caetano Egas Moniz "for the discovery of the production of mutations by means of X-ray irradiation" "for their discovery of the course of the catalytic conversion of glycogen" "for his discovery of the part played by the hormone of the anterior pituitary lobe in the metabolism of sugar" "for his discovery of the high efficiency of DDT as a contact poison against several arthropods" "for his discovery of the functional organization of the interbrain as a coordinator of the activities of the internal organs" "for his discovery of the therapeutic value of leucotomy (lobotomy) in certain psychoses" 1950 Philip Showalter Hench, Edward Calvin Kendall & Tadeusz Reichstein "for their discoveries relating to the hormones of the adrenal cortex, their structure and biological effects" 1951 Max Theiler 1952 Selman Abraham Waksman Sir Hans Adolf Krebs 1953 Fritz Albert Lipmann John Franklin Enders, Frederick Chapman Robbins & Thomas Huckle 1954 Weller 1955 Axel Hugo Theodor Theorell "for his discoveries concerning yellow fever and how to combat it" "for his discovery of streptomycin, the first antibiotic effective against tuberculosis" "for his discovery of the citric acid cycle" "for his discovery of co-enzyme A and its importance for intermediary metabolism" "for their discovery of the ability of poliomyelitis viruses to grow in cultures of various types of tissue" "for his discoveries concerning the nature and mode of action of oxidation enzymes" 1956 André Frédéric Cournand, Werner Forssmann & Dickinson W. Richards "for their discoveries concerning heart catheterization and pathological changes in the circulatory system" 1957 Daniel Bovet 1958 1959 1960 1961 1962 1963 1964 1965 1966 1967 1968 1969 George Wells Beadle & Edward Lawrie Tatum Joshua Lederberg Arthur Kornberg & Severo Ochoa Sir Frank Macfarlane Burnet & Sir Peter Brian Medawar Georg von Békésy Francis Harry Compton Crick, James Dewey Watson & Maurice Hugh Frederick Wilkins Sir John Carew Eccles, Sir Alan Lloyd Hodgkin & Sir Andrew Fielding Huxley Konrad Bloch & Feodor Lynen François Jacob, André Lwoff & Jacques Monod Peyton Rous Charles Brenton Huggins Ragnar Granit, Haldan Keffer Hartline & George Wald Robert W. Holley, Har Gobind Khorana & Marshall W. Nirenberg Max Delbrück, Alfred D. Hershey & Salvador E. Luria "for his discoveries relating to synthetic compounds that inhibit the action of certain body substances, and especially their action on the vascular system and the skeletal muscles" "for their discovery that genes act by regulating definite chemical events" "for his discoveries concerning genetic recombination and the organization of the genetic material of bacteria" "for their discovery of the mechanisms in the biological synthesis of ribonucleic acid and deoxyribonucleic acid" "for discovery of acquired immunological tolerance" "for his discoveries of the physical mechanism of stimulation within the cochlea" "for their discoveries concerning the molecular structure of nucleic acids and its significance for information transfer in living material" "for their discoveries concerning the ionic mechanisms involved in excitation and inhibition in the peripheral and central portions of the nerve cell membrane" "for their discoveries concerning the mechanism and regulation of the cholesterol and fatty acid metabolism" "for their discoveries concerning genetic control of enzyme and virus synthesis" "for his discovery of tumour-inducing viruses" "for his discoveries concerning hormonal treatment of prostatic cancer" "for their discoveries concerning the primary physiological and chemical visual processes in the eye" "for their interpretation of the genetic code and its function in protein synthesis" "for their discoveries concerning the replication mechanism and the genetic structure of viruses" Earl W. Sutherland, Jr. Gerald M. Edelman & Rodney R. Porter Karl von Frisch, Konrad Lorenz & Nikolaas Tinbergen Albert Claude, Christian de Duve & George E. Palade David Baltimore, Renato Dulbecco & Howard Martin Temin Baruch S. Blumberg & D. Carleton Gajdusek Roger Guillemin & Andrew V. Schally 1977 Rosalyn Yalow 1978 Werner Arber, Daniel Nathans & Hamilton O. Smith 1979 Allan M. Cormack & Sir Godfrey N. Hounsfield "for their discoveries concerning the humoral transmittors in the nerve terminals and the mechanism for their storage, release and inactivation" "for his discoveries concerning the mechanisms of the action of hormones" "for their discoveries concerning the chemical structure of antibodies" "for their discoveries concerning organization and elicitation of individual and social behaviour patterns" "for their discoveries concerning the structural and functional organization of the cell" "for their discoveries concerning the interaction between tumour viruses and the genetic material of the cell" "for their discoveries concerning new mechanisms for the origin and dissemination of infectious diseases" "for their discoveries concerning the peptide hormone production of the brain" "for the development of radioimmunoassays of peptide hormones" "for the discovery of restriction enzymes and their application to problems of molecular genetics" "for the development of computer assisted tomography" 1980 Baruj Benacerraf, Jean Dausset & George D. Snell "for their discoveries concerning genetically determined structures on the cell surface that regulate immunological reactions" Roger W. Sperry David H. Hubel & Torsten N. Wiesel 1982 Sune K. Bergström, Bengt I. Samuelsson & Sir John R. Vane 1983 Barbara McClintock "for his discoveries concerning the functional specialization of the cerebral hemispheres" "for their discoveries concerning information processing in the visual system" "for their discoveries concerning prostaglandins and related biologically active substances" "for her discovery of mobile genetic elements" "for theories concerning the specificity in development and control of the immune system and the discovery of the principle for production of monoclonal antibodies" "for their discoveries concerning the regulation of cholesterol metabolism" "for their discoveries of growth factors" "for his discovery of the genetic principle for generation of antibody diversity" "for their discoveries of important principles for drug treatment" "for their discovery of the cellular origin of retroviral oncogenes" "for their discoveries concerning organ and cell transplantation in the treatment of human disease" "for their discoveries concerning the function of single ion channels in cells" "for their discoveries concerning reversible protein phosphorylation as a biological regulatory mechanism" "for their discoveries of split genes" "for their discovery of G-proteins and the role of these proteins in signal transduction in cells" 1970 Julius Axelrod, Ulf von Euler & Sir Bernard Katz 1971 1972 1973 1974 1975 1976 1981 1984 Niels K. Jerne, Georges J.F. Köhler & César Milstein 1985 1986 1987 1988 1989 1990 1991 1992 1993 1994 Michael S. Brown & Joseph L. Goldstein Stanley Cohen & Rita Levi-Montalcini Susumu Tonegawa Sir James W. Black, Gertrude B. Elion & George H. Hitchings J. Michael Bishop & Harold E. Varmus Joseph E. Murray & E. Donnall Thomas Erwin Neher & Bert Sakmann Edmond H. Fischer & Edwin G. Krebs Sir Richard J. Roberts & Phillip A. Sharp Alfred G. Gilman & Martin Rodbell 1995 Edward B. Lewis, Christiane Nüsslein-Volhard & Eric F. Wieschaus "for their discoveries concerning the genetic control of early embryonic development" 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 2006 "for their discoveries concerning the specificity of the cell mediated immune defence" "for his discovery of Prions - a new biological principle of infection" "for their discoveries concerning nitric oxide as a signalling molecule in the cardiovascular system" "for the discovery that proteins have intrinsic signals that govern their transport and localization in the cell" "for their discoveries concerning signal transduction in the nervous system" "for their discoveries of key regulators of the cell cycle" "for their discoveries concerning 'genetic regulation of organ development and programmed cell death'" "for their discoveries concerning magnetic resonance imaging" "for their discoveries of odorant receptors and the organization of the olfactory system" "for their discovery of the bacterium Helicobacter pylori and its role in gastritis and peptic ulcer disease" "for their discovery of RNA interference - gene silencing by double-stranded RNA" Peter C. Doherty & Rolf M. Zinkernagel Stanley B. Prusiner Robert F. Furchgott, Louis J. Ignarro & Ferid Murad Günter Blobel Arvid Carlsson, Paul Greengard & Eric R. Kandel Leland H. Hartwell, Sir Tim Hunt & Sir Paul M. Nurse Sydney Brenner, H. Robert Horvitz, & Sir John E. Sulston Paul Lauterbur & Sir Peter Mansfield RichardAxel & Linda B. Buck Barry J. Marshall & J. Robin Warren Andrew Z. Fire & Craig C. Mello 2007 Mario R. Capecchi, Sir Martin J. Evans & Oliver Smithies "for their discoveries of principles for introducing specific gene modifications in mice by the use of embryonic stem cells." 2008 Harald zur Hausen, Françoise Barré-Sinoussi & Luc Montagnier 2009 Elizabeth H. Blackburn, Carol W. Greider & Jack W. Szostak 2010 Sir Robert G. Edwards "for his discovery of human papilloma viruses causing cervical cancer" "for the discovery of how chromosomes are protected by telomeres and the enzyme telomerase" "for the development of in vitro fertilization" 2011 2012 2013 2014 Bruce A. Beutler, Jules A. Hoffmann & Ralph M. Steinman Sir John B. Gurdon & Shinya Yamanaka James E. Rothman, Randy W. Schekman & Thomas C. Südhof John O'Keefe, May-Britt Moser & Edvard I. Moser "for their discoveries concerning the activation of innate immunity" "for the discovery that mature cells can be reprogrammed to become pluripotent" "for their discoveries of machinery regulating vesicle traffic, a major transport system in our cells" "for their discoveries of cells that constitute a positioning system in the brain"
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