Climate and Pattern in Terrestrial Ecosystems Comparing the effect of climate on species diversity in 5 LTER sites Five different biomes Biomes are distinguished by their predominant plants Biomes are associated with particular climates with variations in temperature and precipitation Tropical forests include rainforest and dry forest Deserts’ water loss usually exceeds precipitation Temperate forests rainfall averages 650 - 3,000 mm Grasslands annual rainfall 300 - 1,000 mm Urban desert rainfall augmented by irrigation A Note about Scale It’s important to consider the scale at which questions are investigated Spatial Scale (Separated by space): • As large as galaxies, biomes, countries, states, etc. •As small as animals within same forest, insects on same plant, microclimates above leaf vs. below leaf •Can also be units of measure (i.e. meters vs. feet, volume vs. area, etc.) Temporal Scale (Separated by time): •As large as centuries, decades, years, etc. •As small as hours, seconds, milliseconds, etc. Spatial scale 95 90 85 80 75 70 65 60 55 50 45 40 35 December November October September August July June May April December November October August July June May April March Minimum February F Maximum September •Different patterns 95 90 85 80 75 70 65 60 55 50 45 40 35 January •Separated by space March H. J. Andrews Research Forest •H. J. Andrews Research Forest in Oregon (bottom) •Same biome Minimum February •Harrison Experimental Forest in Mississippi (top) •Same country Maximum January F 2003 Average Maximum and Minimum Temperatures of two temperate forests in the United States: Harrison Experimental Forest Temporal scale 1995-1999 65 60 55 F Maximum and Minimum Temperatures at H. J. Andrews Research Forest in Oregon y = 0.0679x + 56.569 Maximum 50 Minimum 45 y = -0.2412x + 40.91 40 35 1995 •5-year data set (top) •10-year data set (bottom) 1996 1997 1998 1999 1995-2004 65 60 •Same forest y = 0.6394x + 55.316 55 •Separated by time Maximum F •Same site 50 45 40 •Different patterns Minimum y = 0.3422x + 39.322 35 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 H.J. Andrews Temperate Forest LTER These forests are among the tallest and most productive in the world with many plant and animal species. Lower elevation forests are dominated by Douglas-fir (Pseudotsuga menziesii), western hemlock (Tsuga heterophylla), and western redcedar (Thuja plicata). Higher Elevation forests are dominated by Pacific silver fir (Abies amabilis). H. J. Andrews Temperate Forest Maximum and Minimum Temperatures (mean) at H. J. Andrews 1958-2004 65 60 The climate at the H. J. Andrews is cool and wet in winter and warm and dry in summer. Maximum 50 Minimum 45 40 35 1958 1960 1962 1964 1966 1968 1970 1972 1974 1976 1978 1980 1982 1984 1986 1988 1990 1992 1994 1996 1998 2000 2002 2004 F 55 H.J. Andrews Total Precipitation Very large winter precipitation leads to significant soil water surpluses and runoff Average Monthly Precipitation H. J. Andrews Research Forest 1958-2004 16 14 Summertime precipitation is low to nonexistent inches 12 10 8 6 4 2 December November October September August July June May April March 0 February precipitation is about 2,500 mm at low elevations, falling mainly as rain at low elevations and as snow at upper elevations. January Annual Combined Data Mean Precipitation and Temperature at the H. J. Andrews Research Forest 1958-2004 16 85 80 14 75 12 70 65 Precipitation 60 F 8 55 6 50 45 4 40 2 35 December November October September August July June May April March 30 February 0 January inches 10 Maximum Minimum Variations in temperate forests 90 6 80 5 70 60 Precipitation 50 3 F inches 4 40 Maximum Minimum 30 2 20 1 10 December November October September August July June May April March February 0 January 0 85 16 80 14 75 12 70 65 Precipitation 60 F 8 55 6 50 45 4 40 2 35 December November October September August July June May April March 30 February 0 January inches 10 Maximum Minimum Average monthly temperatures and precipitation at two temperate forests: Hubbard Brook in New Hampshire, USA (top) and H. J. Andrews Research Forest in Oregon, USA (bottom) Discerning patterns Based on the patterns at the Hubbard Brook and H. J. Andrews forests: What would you expect the forest soils to be like? What effect might the patterns have on the types of soil invertebrates present at each site? Would Would you expect the plant communities to be similar? you expect patterns between a temperate forest and a tropical forest to differ? How might precipitation influence the components mentioned above?
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