Vol. L] 19113 J COOKE, $pe•OfBin•Flight. SPEED COMPILED OF BIRD BY MAY 309 FLIGHT. THACHER COOKE, DEFINrrE recordsof the speedof bird flight are sometimeshard to find. Quite a few suchrecordsexist; but they are scattered, manyin publications not readilyaccessible.In orderto facilitate furthers•udyof thisinteresting subjectit seems desirable to bring togethertheserecordsfor North Americanspecies,where they will be readilyavailable. Scatteredthroughsportsman's magazinesare many so-called recordsof the speedof game birds, most of which are estimates basedon calculationsthat were at best largely guesswork. In recent years, however,the air-speedindicator and automobile speedometerhave given meansby which individualsof many specieshave been definitelytimed. Also studieshave beenmade of birds flying acrossmeasureddistances,timed by stop watches and theodolites. These are furnishinga fairly definite understandingof the rate at whichbirdsof differentkindsfly, and in generalthe speeds sodeterminedare muchlowerthan wasformerly supposed. Some of the smaller Passeres have been found to make less than 20 milesper hourin ordinaryflight, andrecordsof ducksand geese averagedlittle over 40 miles per hour. At the other extremeis the hunting Peregrine. Portal, an experienced falconer,estimates this bird'saveragemaximumspeedof level flight throughstill air as 62 milesper hour, and somehave estimatedthat it strikesits prey at 150milesperhour. McLean timeda Duck Hawk hunting over a 400-yardfieldin California,whoseaveragespeedwas over 165milesperhourandwhosegreatestspeed was180milesperhour. Extremeas thesespeedsseem,they arepossiblyexceeded by some of the Swifts. Ten yearsagoCol. RichardMeinertzhagen published'in'The • Ibis' a very comprehensive article in which he collectedand ana- lyzed the availableinformationon this subject. He reported,as the experience of aviators,that geesecan acceleratevery little, but that duckswhenpressedcan speedup to about 60 milesper 310 COOKE, Speed ofBirdFlight. [Auk [JLGy hour. He concludedthat birds have two speeds,"a normal rate which is used for every-daypurposesand also in migration,and an aeeeleratedspeedwhieh in someeasesnearlydoublesthe rate of their normal speed." This acceleratedspeed,however,cannot be maintainedfor any length of time. Several reeent articles on the meehanies or aeronauties of bird flight are not ineluded in the following annotated bibliography whieh shows the soutees and nature of the material summarized in the appendedtable. BASSETT,FRANK N. 1921. The Speedof a Flying Dove. Condor,vol. 23, p. 190-191. [BRUETTE,WILLIAM]. 1917. Birds and Aviators. Forest & Stream, vol. 87, p. 603. (Ducks average65• milesper hour when flying upwards;69 milesper hour when flying horizontally [air speed?].) CLARKE••r. EAGLE. 1912. Studiesin Bird Migration. London,Gurney & Jackson,2 vols. 8vo. II. p. 29. (Recordsof skylark and starling.) CLAYTON,H. HELM. 1897. The Velocity of a Flight of Ducks obtained by Triangulation. Science,New Series,vol. 5, p. 26. (The Height and Velocity of the Flight of a Flock of Geesemigratingnorthward. Ibid. p. 585-586. Measured by theodolites.) GLADSTONE, HuGa S. 1922. Record Bags and Shooting Records. London, H. F. & G. Witherby, 240 pp., 8vo. (Chapter, "The Speedof Birds," p. 173-197, quotes many published records and discussesthe effectof wind on speedof flight.) GRINNELL,GEORGEBIRD. 1881. Flight of Birds. Forest & Stream, vol. 17, p. 247. (Data for pigeons.) 1901. American Duck Shooting. New York, Forest & Stream Pub~ lishing Co., 623 pp., 8vo. p. 509-510. (A flock of Pintails flew somedistanceparallel to a train going52 miles per hour.) 1903. Timing the Flight of Birds. Forest & Stream, vol. 41, p. 375. (Data for pigeons,pintail, and starlings.) HARRISSON,T. H. 1931. On the Normal Flight Speed of Birds. British Birds, vol. 25, p. 86-96. (Speedscarefully timed by auto or motorcycle speedometer. Data for 36 species. Bibliography.) HA'ireS,SAMUELP., JR. 1020. Speedof a Flying Hummingbird. The Auk, vol. 46, p. 116. COOKE, Brd 311 HOLLAND,i•Y P. 1924• HOWFast Can a Quail Fly? OutdoorLife, vol. 53, p. 70. (Aviators say that at 65 miles per hour they can overtakethe fastestducks.) How Fast Can a Quaff Fly? Ibid., p. 151. (Timed with stopwatch from time bird flusheduntil secondbarrel was fired; estimatedabout 48 milesper hour.) JONES,Lr•ros. 1927. lqighwayMortality and Speedof Flight. Wilson Bulletin, vol. 39, p. 8-10. (Long-billedCurlew and flock of gulls timed by auto.) LONGS•I•EE?,R. J. 1930. Noteson Sp•d of Flight of CertainWater Birds. The Auk, vol. 47, p. 428-429. (Twelvespecies timedby auto speedometer.) MARTIN, EDWARDT. 1916. The Speedof Ducks. Forest& Stream,vol. 86, p. 1147. (Diving ducksfasterthan puddleducks;teal not sofast as Canvasback and Scaup.) McLEAN, I). I). 1930. The Speedof Flight in Certain Birds. The Gull, vol. 12, no. 3. (Discusses the speedof 13 species timedby autospeedometer.) MEINERTZHAGEN, RICHARD. 1921. Somepreliminaryremarkson the Velocity of Migratory Flight amongBirds, with specialreferenceto the PalaearcticRegion. Ibis,p. 228-238. Reprintedin Smithson]an Reportfor 1921, p. 365-372. subject.) (The most comprehensivediscussionof the MUNSON,EDWARDL. 1930. Timing the Ducks. Field & Stream, vol. 35, p. 18-20, 70. (Tests made by airplane, chasingducks;i.e. gives the maximum speedof whichthe birdswerecapable;five species.) [MuRPHY,WILLIAM W.] 1905. How Fast do Birds Fly? Forest & Stream, vol. 65, p. 330. (Engineerof fast train has raced birds flying parallel to or Pm•z•s, aheadof his engine;five species.) JOHNC. 1922. A Natural Historyof the Ducks. Vol. I, p. 22-23. (Somediving ducks faster near ground than Mallard-like species. Extra speedof teal moreapparentthan real. 40-50 milesper hour, commonspeedin migration; 55-60 milesper hour possibleby somespecies.) POR•, C. F. A. 1922. The Speedof Birds. Field (London),vol. 139,p. 233-4. (Speed is partly individual, one partridgeof a coveywas seento fly 15 per cent fasterthan the otherswhenall wereat full speed. Author, an experiencedfalconer, gives average maximum speedof levelflight throughstill air of seventeenspecies.) 312 COOKE, Speed ofBirdFlight. Auk [July E. P. R. 1913. How Fast Do Ducks Fly? Forest & Stream, vol. 80, p. 41. (Conductorsaid train often raced ducksand at 50 milesper hour the train usually won.) RADCLIFFE,C. r. 1922. The Speedof Birds. Field (London),vol. 139, p. 234. (The heaviestbird of a giventype is the fastestwhenit getsgoing. Quickerflushand riseof smallerbird deceptive. No bird can beat a PeregrineFalcon.) RITTER, C. B. 1910. Speedof Birds and Animals. AmericanField, vol. 73, p. 200. (Observationsof a locomotiveengineer.) ROBINSON,•I. W. 1922. What is the FastestBird? Field (London),vol. 139, p. 138. (ConsidersMerganser going down wind one of the fastest of all birds.) WETMORE,ALEXANDER. 1916. The Speedof Flight of CertainBirds. Condor,vol. 18, p. 112113. (Sevenspeciestimed by auto speedometer.) WHITE, FRANCISBEACH. 1927. Birds and Motor Cars. The Auk, vol. 44, p. 265-266. 1929. Birds and Motor Cars. The Auk, vol. 46, p. 399. (Speedsof thirteen speciesof Passeres.) WOOD. HAROLD B. 1923. The Speedof Flight of Birds. BirdsLore,vol. 25, p. 121. (Recordsof eightspecies timedby auto speedometer.) 314 CookE, Speedof Bird Flight. '-0 Auk July VoL L• 19aa J 0oor,•,,•peed ofBirdFli•hi. 315 316 Coo•,Speed ofBird Flight. Auk [j•ly
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