Physics 1230: Light and Color Exam 4 cancelled: Exam extra credit assignment will be due Wed. at 5PM Extra credit to improve exam scores! Final HWs: Due today, Tuesday, 5PM Lecture 14: The retina and brain, image signal processing. Reading: Chap. 9,10 Color perception. 1 The remaining lectures: We are here • Ch. 7 (Retina and visual perception), • Ch. 9 & 10 (color & color perception). 2 Ch. 7 – Visual Perception We are here • • • • • • • Parts of the visual processing system Lightness and brightness Retinal processing: Lateral inhibition Hermann grid Receptive field Motion illusion Craik O‟Brien illusion & simultaneous lightness contrast • Other optical illusions 3 The Retina: Detecting the light and processing the images The retina and optic nerves are recognized as actually parts of the brain (like your olifactory bulb in the nose). They start development IN the brain and migrate… Has 108 nerve endings to detect image rods, for high sensitivity (night vision) cones, for color and detail, 7 million optic nerve = 106 transmission lines fovea, region of best vision (cones) More nerves in your retina than some creatures have in their entire brains. Processing Power. 4 Rods and cones • Rhodopsin, a photochemical, responds to light It is destroyed and reformed. Signal goes to a synapse, a gap between nerve cells • There are 3 kinds of cones for 3 colors red, green, blue (more later). A great deal is understood about how the individual cells of the retina receive light, respond to light, and transmit signals. 6 Rods and cones Example: Rhodopsin and photosensitivity Photo-responsive membrane protein is known in atomic detail Light drives a change in molecular shape. Opens/closes membrane We will skip Most of cellular detail BECAUSE… 7 Does our understanding of the individual rods, cones, and other cells of the retina do much to explain this?: (A) Creitanly (B) Myaby Not Mcuh We need to understand how NETWORKS of cells WORK TOGETHER to let us perceive. 8 Layers of the retina Light 9 Layers of the retina 10 See text fig. 7.2 Layers of the retina are CROSS Connected From the following article: Neurobiology: Bright blue times Russell G. Foster Nature 433, 698-699(17 February 2005) doi:10.1038/433698a a, The rods (R) and cones (C) convey visual information to the ganglion cells (G) through the bipolar cells (B). Horizontal cells (H) allow lateral connections between rods and cones. Amacrine cells (A) allow lateral connections between bipolar and ganglion cells. The optic nerve is formed from the axons of all the ganglion cells. A subset of ganglion cells (MG cells) also detects light directly; for this, they require the photopigment melanopsin, as now confirmed1, 2, 3. b, Light, via melanopsin, causes changes in Ca2+ levels in MG cells9 (a fluorescent Ca2+ indicator was used here). Counterintuitively, light passes through the transparent ganglion layer to reach the rods and cones. 11 Connections and cross connections are MOST important. Photoreceptors: rods and cones connected to the bipolar cells connected to the ganglion cells, funnel “data” through axons into the optic nerve sideways connectors (these help with analysis) horizontal cells, next to the photoreceptors amacrine cells 12 Clicker question The arrow points to: A. Photoreceptors B. Horizontal cells C. Bipolar cells D. Amacrine cells E. Ganglion cells 13 Clicker question The arrow points to: A. Photoreceptors B. Horizontal cells C. Bipolar cells D. Amacrine cells E. Ganglion cells 14 Optic chiasma and brain structure Brain damage on the left side hurts vision on the right side. 15 See text fig. 7.3 Brain anatomy Optic chiasma Left field of view goes to right brain Right field of view goes to left brain from both eyes Visual cortex is where you “see” Brain damage at this location hurts vision. 16 Clicker question If the left side of your brain is injured, you might lose vision in your A. left eye B. right eye C. left field of view D. right field of view E. some loss in left and right field of view 17 All this „hardware‟ allows us to perceive the world and function in it. Many complicated sub-systems have developed. Let‟s study a few to get some insight into how vision works. 18 Interesting collective behavior 1: We detect RELATIVE Lightness, not total Brightness Brightness: amount of light Lightness: property of a surface newspaper = 0.65 (reflectance) printer paper = 0.84 photo quality paper = 0.90-0.99 Total amount of light is far less important than the relative amount of light, particularly as compared with nearby objects. Demo with room lights. 19 Lightness and brightness Lightness constancy: brain and eye correct for amount of light so that white, gray, and black look the same independent of brightness. Weber‟s law: we think lightness is equally spaced when the ratios are equally spaced Example: lightness 0.5, 0.25, 0.125 look equally spaced. These numbers are ½, ¼, 1/8 etc. The spacing that looks equal is not 0.9, 0.8, 0.7, etc. 20 Demo: Lights on or lights off Retinal processing that allows Relative Lightness sensitivity: Amacrine and horizontal cells “turn down” the signals from areas adjacent to bright areas. “Lateral Inhibition” 21 See text fig. 7.5 “Receptive field” The rods/cones and local cells are connected in a group: Center of group causes nerves to fire if illuminated. Surrounding group causes nerves to STOP firing if they are illuminated. Nerve cell fires rapidly Nerve cell doesn‟t fire Nerve cell doesn‟t fire Nerve cell fires only a bit 22 See text fig. 7.12 Receptive field (again) Called LATERAL INHIBITION 23 The yellow is the region receiving light. See fig. 7.11 Because of LATERAL INHIBITION, Edge detection is enhanced Full illumination: Not much nerve activity. Half illumination gives bigger signal 24 Lateral inhibition along with relative lightness cause: Simultaneous lightness contrast Craik O‟Brien Illusion Contrast at the edge affects your perception of center. Are the small gray patches below identical? A) YES B) NO 25 See fig 7.7 Craik O‟Brien Illusion Simultaneous lightness contrast These are the patches without the surround. 26 Simultaneous lightness contrast (again) “Checker shadow illusion” Which square is lighter in shade, square A or square B? 27 Simultaneous lightness contrast “Checker shadow illusion” Slide them together and compare. A is surrounded by light squares and B is surrounded by dark squares in the previous slide. 28 29 Hermann grid illusion: dark areas are from lateral inhibition 1 3 2 The red areas show the receptive field. Lateral inhibition is greater at 1 than at 2. The fovea has a smaller receptive field. So the lateral inhibition is the same everywhere in the white area. 30 White space is larger than receptive field 31 It is blacker away from a corner where there is more inhibition. 32 33 The music A. Kitaoka 34 Does the center stripe have constant lightness? Or is the center stripe darker in the middle and at the ends? A) Constant B) Darker in middle and ends 35 The center stripe has constant lightness. 36 Clicker question A white sheet of paper continues to look white as the light level is reduced. We call this effect: A. Simultaneous lightness contrast B. Lateral inhibition C. Weber‟s law D. Lightness constancy E. Edge enhancement 37 Clicker question The bands of gray look lighter on their right side because of: A.Simultaneous lightness contrast B. Lightness constancy C. Weber‟s law D. Lateral inhibition E. Both A and D 38 Victor Vasarely, Zebras. The black/white boundaries outline the necks. The artist has made use of the tendency of the eye to find lines. The regions of color don‟t have edges, but appear to. Picasso 40 French artist George Seurat used edge enhancement by lateral inhibition to make figures stand out sharply Lighter just before edge Darker just before edge 41 El Greco 42 The white crosses are an illusion. Victor Vasarely, artist. The edges of the squares seem lighter because of the dark surrounds. 43 Interesting collective behavior 2: We expect a 3D world, lit from ABOVE: Our perception of relative lightness changes based upon Location and Shape! 44 Example: Which is the darker patch, A or B? A B 45 A B Previous experience effect: Here, the eye is “fooled” into thinking the light is from above. The panel “A” has lots of light, so it must be really dark. But “B” must be lighter because it is in the “shade.” 46 Which creature is larger? A) The little one in front B) The big one in back C) They are the same size. Previous experience in tunnels tells us that the creature in back is further away, and hence must be larger. 47 http://www.michaelbach.de/ot/index.html “Previous experience” interprets these flat images as being from 3-dimensional boxes. The shadows tell us what is a “floor” and what is a “wall.” 48 Victor Vasarely, artist Size constancy: Are all the vertical lines the same height? A) Look different to me B) Look the same to me 49 50 http://www.michaelbach.de/ot/index.html Interesting collective behavior 3: Sensitive to a MOVING World. Time and motion important. Fatigue: prolonged stimulation (staring at a lamp) causes a weaker response and a negative afterimage. Successive lightness contrast: a gray object looks darker after looking at white. Positive afterimage: We see a flash as a bright spot after it has gone away. Over stimulated nerves keep firing. 51 Successive lightness contrast Negative afterimage Stare at this for 30sec., then stare at the next slide. 52 53 Stare at this, stare at the next slide. 54 55 56 57 Fatigue and Involuntary eye movement Eye movement moves the image around so that new areas are stimulated. Without eye movement, images fade. This has been verified by experiments that fix the image on the retina. Eye movement causes wavy lines to appear as though in motion, because the afterimage interferes with the moved image. 58 59 The eye is moving all the time. It corrects for motion… IF there are edges, but not if edges are absent. The only difference between the center and edge is the lack of any feature to “focus” on. http://www.michaelbach.de/ot/mot_eyeJitter/index.html 60 Other illusions There are many optical illusions with varying explanations. Many are poorly understood. 61 Are the blue shades the same? Lateral inhibition cannot explain this! 62 63 Lateral inhibition alone does not explain this effect, the Munker-White illusion. 64 http://www.newworldencyclopedia.org/entry/Muller-Lyer_illusion Müller-Lyer illusion Which arrow is longer? 65 http://www.michaelbach.de/ot/index.html Müller-Lyer illusion This is the back corner of a room, it is further away, hence it must be larger. 66 Big Moon Illusion 67 What you remember. Actual Frankfurter illusion While focused on the background, hold your two index fingers horizontally in front of your eyes, not touching. A piece of finger will appear to float in space. 68 http://www.michaelbach.de/ot/sze_Frankfurter/index.html Are the lines straight? Hering Illusion 69 Does the square have straight sides? 70 http://www.michaelbach.de/ot/ang_hering/index.html Does the square have straight sides? 71 72 Poggendorff Illusion Are the lines continuous and straight “behind” the yellow columns? 73 http://www.michaelbach.de/ot/ang_poggendorff/index.html Poggendorff Illusion 74 http://www.michaelbach.de/ot/ang_poggendorff/index.html Art that mimics 3-d. 75 Motion after effect Motion channel, keeps firing after watching a moving object, causing motion aftereffect. http://www.michaelbach.de/ot/mot_adaptSpiral/index.html The following are from Akiyoshi Kitaoka Department of Psychology, Ritsumeikan University, Kyoto, Japan: http://www.psy.ritsumei.ac.jp/~akitaoka/saishin27e.html 76 Motion illusion (A. Kitaoka) Note that each green circle is rotated slightly from its neighbor. As your eye jumps around, it sees the circles rotation. 77 Are the ropes tangled? 78 from Akiyoshi Kitaoka Motion illusion (Kitaoka) 79 Motion illusion (Kitaoka) 80 81 Snakes - Akiyoshi Kitaoka 82 Dead Snakes - Akiyoshi Kitaoka Motion effects from Michael Bach‟s web page • • • • • Silhouette illusion Motion induced blindness Motion aftereffect (Waterfall illusion) Spiral aftereffect (motion channel activated) Breathing square http://www.michaelbach.de/ot/index.html 83 A good place to stop today 84
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