BLUMEA
45
403-425
(2000)
Notes on Malesian Fabaceae
(Leguminosae-Papilionoideae).
7.
The
genus Millettia
Adema
Frits
Nationaal
Herbarium
Nederland,
2300 RA
Universiteit Leiden
Leiden, The
P.O.
branch,
Box 9514
Netherlands
Summary
The
Millettia
genus
described:
Millettia;
the
as
An identification
words
and M.
15
species
list and
an
revised
is
already
in
use
recognised for
are
index
of
names
Solomon
: Millettia,Malesia,
for the Flora
Malesiana
velutina
glabra. Pongamia
epitheton velutina
tina Adema. In total
Key
& Arn. is
Wight
M. borneensis
in
the
Millettia,
area
area. Two
(C.T. White) Verdc,
a new
name
concerned. A
is
species
are
is transferred
to
M. velve-
proposed:
to the
key
new
species
is
given.
included.
are
Islands.
Introduction
Millettia was
M.
species:
species
described
have been added
Geesink
(1981
1984)
a,
Dunn)
and
and their
can
sect.
the
cation list and
sect.
notes.
an
glabra,
and
,
are
A
key
sect.
to
all
species
127
Dunn
Typicae
following
one new name,
names are
two
species
new
many
in 15 sections.
other genera and unitedPongamia
treatment
new
+ sect.
Macrospermae
the genus, both sections,
species
will be described,
velvetina Adema, is
M.
described in full; the other
index of
accommodate
& Am. Since then
also Adema, 2000). In Malesia two sections
Millettia (=
Dunn. In the
to
accepted
to
will be reviewed and discussed. Two
species
synonymy and
Fragiliflorae (see
Fragiliflorae
species
new
(1834: 263)
splendens Wight
this genus. Dunn (1912)
to
be found:
M. borneensis and M.
Only
& Am. and M.
transferred several sections
Vent, with Millettiasect.
of Millettia
& Arnott
by Wight
rubiginosa Wight
species
are
proposed.
mentioned with full
of Millettia in Malesia is
given;
an
identifi-
included.
MILLETTIA
Millettia
Wight
&
Am.,
Jungh. (1852) 247;
Pflanzenfam.
Bot. 5
3,
3
(1916) 361; Merr.,
R. Geesink in
Man.
Hook.f.,
(1894) 270; Prain,
(1910) 70;
595; Verde.,
Prod. Fl. Penins.
Baker in
Ind. Or.
J. As. Soc.
Dunn,
J. Linn.
Soc.
Enum.
Philipp.
Flow. PI. 2
New
Polhill
&
Guinea
Raven,
Millettia
Wight
& Arn. sect.
Millettia
rubiginosa Wight
1
(1834) 263,
Fl. Brit. Ind.
Leg. Syst.
(1912)
Typicae Dunn,
& Arn.
Allen
2
Soc.
in
&
Engl.
in
Miq.,
Prantl,
(1897) 358; Merr., Philipp.
Backer &
&
(1981) 257;
J. Linn.
cons.; Benth.
123; Gagnep.,
(1923) 278;
1
nom.
(1876) 104; Taub.
Beng. 66,
Bot. 41
Leg. (1979) 332;
Adv.
2
Fl.
G6n.
J.
PI.
Nat.
Sci.,
Indo-Chine
Bakh.f., Fl.
Java
1
2
(1964)
Allen, Leguminosae (1981) 435;
Leiden Bot. Ser.
Bot. 41
(1912)
8
134. —
(1984)
102.
—
Type species:
404
BLUMEA
Jard. Malm.
Pongamia Vent.,
&
Bakh.f.,
Backer &
298;
(1803)
Fl. Java
28,
t.
—
2
R. Geesink
Millettia
Wight
& Arn. sect.
indica
Vent.
Taub. in
(1876) 594;
Type species: Galedupa
—
2000
2,
Merr.,
nom. cons.;
Type species: Pongamia glabra
Galedupa Lam., Encycl.
No.
45,
(1964) 616; Verde., Man.
1
Allen, Leguminosae (1981) 543;
259.
Vol.
in
Polhill
P.
[=
Engl.
Enum.
&
Flow. PI. 2
Philipp.
New Guinea
Raven,
(1923)
Leg. (1979) 311;
Adv.
1
Leg. Syst.
Allen
(1981)
pinnata (L.) Pierre],
&
Prantl,
Nat. Pflanzenfam.
3
3,
(1891)
344.
Lam.
J. Linn. Soc. Bot. 41
Fragiliflorae Dunn,
(1912)
136.
—Type species
(1912)
136.
—Type species
not indicated.
Millettia
& Am. sect.
Wight
Macrospermae Dunn,
J.Linn. Soc. Bot. 41
not indicated.
Trees, shrubs,
lianas. Leaves
or
imparipinnate, rarely pinnately trifoliolate; stipules
present, caducous; stipellae present
in
panicles;
or
± tubular, teeth
standard with
keel
Stamens
fixed, in
to
the
inconspicuous
petals slightly
diadelphous,
mono- or
sect.
glabrous,
ellipsoid
or
dehiscent. Seeds
Distribution
—
subsessile
More than
c.
15
b.
a
100
case
Calyx
or
not.
campanu-
wings adhering
with
lateral
a
to
the
pocket.
with basal fenestrae; anthers basito annular or
with 2
(or 3)
to
glabrous upwards.
in the
for M.
reported
SECTIONS OF
button-shaped,
pulvinus
at
hairy
Axillary
buds flattenedovoid
or
rarely
±
tropics
10-lobedand adnate
several ovules;
Pods
style
strap-shaped
indehiscent
or
to
tardily
oblong.
of Africa and Asia into the
pinnata
and several African
MILLETTIA
enclosed
peduncle.
by
IN
species
MALESIA
the budscales.
Standard with
two
Inflorescences
basal callosities.
sect.
without a
ties. Anthers
tightly
the base of the
Anthers
rescences
wings, usually
'quadrate'
to
pseudo-
than the tube. Corolla:
with auricles;
not, dehiscent,
species
opposite.
or
1981).
buds
Axillary
with
or
caducous.
slightly longer
stipitate,
ones
species.
KEY TO THE
la.
to
in lower half and
hairy
to
rarely
lateral
pseudoracemes
the base; flowers fascicled
Disc inconspicuous
Root nodules have been
(Allen & Allen,
up
in the former
transversely ellipsoid
—
Pacific; in Malesia
Note
or
semi-circular, compressed
or
racemes,
at
shorter than the
Fragiliflorae hairy.
hypanthium. Ovary
almost
pulvinus
a
without basal callosities,
or
keel
petals;
Leaflets entire,
panicles,
usually caducous; bracteoles present, usually
Bracts present,
late
with
Fragiliflorae
sect.
absent.
or
and/or terminal,
Inflorescences axillary
pulvinus
to
at
globular, loosely
the base of the
enclosed
peduncle.
by
Fragiliflorae
the budscales. Inflo-
Standard without callosi-
glabrous
sect.
Millettia
Section Millettia
Section Millettia.
Millettia
Section
—
Trees, shrubs,
the budscales.
pulvinus
very
Typicae■Dunn,
Macrospermae Dunn,
Millettia sericea
no
Section
rubiginosa Wight
rarely
or
Soc. Bot. 41
Bot.
(1912)
(1912) 134, nom. illeg. —Type:
J. Linn.
Soc.
41
136.
—
Lectotype (here designated):
Benth.
lianas.
Axillary
buds flattened ovoid
Inflorescences axillary
at the
with
J. Linn.
& Am.
or
terminal
to
globular, loosely
pseudoracemes
or
enclosed
by
pseudopanicles,
base of the peduncle. Standardwithout callosities. Anthers glabrous,
some
hairs.
F. Adema:
Distribution
—
Notes
Malesian
on
Fabaceae
SE Asia from India and Sri Lanka
-
Millettia
405
S China, Indochina, Thailand
to
and W Malesia.
Note
The
—-
Macrospermae
is the presence
In Malesia four
species belong
1. Millettia chrysamaryssa
Millettia
Note
as
This
—
M.
anthers (0.7-1.1
5
0.4
by
cm
upright-oblong,
25
by
Fig.
1. Millettia
24
by
16
Wight
a
& Arn.
2. Millettia
galliflagrans
ex
289.
—
Type:
Hassk.
rather
(AA, L, SINU),
found in the material identi-
surprisingly,
3
by
differs
chrysamaryssa
mm vs.
0.6-0.7
2
mm vs.
2-3.5
by
mainly
from
2-3 mm), the size of the
0.4-0.6 mm), the
pods (flattened,
14
thickened, transverse-ellipsoid, 25-45 by 10-
account
sect.
Millettia.
(a:
Maxwell
Adema, 1999.
Pods.
a.
78—107 ;
b:
Mus. J. 13
enigmatic species.
Suppiah)
may
(Vent.) Wight
Jungh. (1852) 248; Miq.,
J. Linn.
78-107
M.
chrysamaryssa Adema;
King’s
It is
specimen,
belong
Wight
& Am.
Fl. Ned.
Soc. Bot. 41
coll.
b. M.
sericea
709).
Whitmore
3. Millettiasericea (Vent.)
sericea
Maxwell
12.4.1978.
full
known from the type
FRI 17804 (T.
Dunn,
a
(1970) 136; Tree
(RF. Cockburn), Malaysia, Johore, Kluang F.R.,
Note —A rather
Millettia
was,
galliflagrans Whitmore, Fed.
only
(1999)
River,
mm vs.
by
& Am.
now
and
differ-
unimportant
an
thickened, 5-7.5 by 1.5-3.5 by 1-2 cm) and the seeds (flattened,
(Vent.) Wight
FRI 7551
9
1
Fig.
—
stipules (5 by
0.4-0.6
by
vs.
Typicae’ (= Millettia)
As this is
sections should be united.
two
earlier botanists. Millettia
by
4-13 mm). See for
Millettia
Novon
species
new
sericea
stipellae.
this section:
to
Selim
M. sericea in the size of the
by
absence of
Adema
chrysamaryssa Adema,
Malaysia, Perak, Padang,
fied
or
in other sections of.Millettia, the
ence
'
difference between Dunn's sections
only
to
ex
this
Hassk.,
1
small
tree
1
297.
(1972)
—Type
(n.v.).
with unifoliolateleaves.
which I have
& Am.
Ind.
a
Fl. Mai.
at foot of G. Blumut
not
seen as
Up
to
yet. The specimen
species.
ex
Hassk.
Cat. Hort.
—
1b
Bogor. (1844) 283;
(1855) 153; Prain,
(1912) 169; Gagnep., Fl.
Fig.
J. As. Soc.
Gen. Indo-Chine
2
Benth. in
Beng. 66,
2
(1916) 383;
Miq.,
PL
(1897) 88;
K.
Heyne,
406
Nutt. PL
2
s.n.
1
Diet.
2
298.
(1972)
(P?,
Herb.
—
Blume
Java
Jussieu?),
Millettia sericea
154.
Blume
Millettia sericea
(1861)
Millettia
Java
s.n.,
300.
s.n.
Millettia
Millettia
1
(1928)
(1964) 596; Whitmore, Tree
(1803)
28.
t.
Type:
—
De
Fl.
Lahaye
Hassk.
ex
& Am.
L
& Am.
L
Miq.,
Fl.
ex
Hassk. forma
(L 908.120-317)],
Ned.
unknown
Fl. Ned.
forma subcordata
(L 908.120-311)],
[holo
—Type
Miq.,
Ind.
154.
(1855)
1
—
Java.
Hassk.
ex
284.
obtusa
var.
Ind.
1
Fl. Ned.
Miq.,
Ind.
1
(1855)
Fl. Ned. Ind.
1
(1855)
Moluccas.
brachycarpa Miq.,
Sumatra.
156.
(1855)
—
Lectotype (here designated):
& Am.
ex
Hassk.
& Arn.
Eerste
Ind.,
major Miq.,
Fl. Ned.
Ind.,
Eerste
bijv.,
Sumatra
(K).
Hassk.
ex
—Type: Teijsmann
Fl. Ned.
var.
Sumatra
s.n.,
bijv.,
var.
Miq.,
aurata
Sumatra
(n.v.).
Sumatra
(1861)
s.n.,
Fl. Ned.
300.
—
Ind.,
Eerste
bijv.,
Type: Diepenhorst
s.n.,
(n.v.).
sericea
(1897)
1
[holo L (L 908.120-407)].
300.
turgida Miq.,
Sumatra
Java
Malm.
Bogor (1844)
Type: Teijsmann
(1861)
Jard.
& Am.
(Vent.) Wight
sericea
Sumatra
ex
Fl. Siam. Enum.
(1922) 584; Craib,
(n.v.).
[holo
(Vent.) Wight
—
1
2000
2,
Bakh.f., Fl.
(L 908.120-305)],
s.n.
argentea Zipp.
Zippelius
L
(Vent.) Wight
Type:
—
Millettia
Blume
&
Cat. Hort.
(Vent.) Wight
Type:
—
[holo
s.n.
Millettia sericea
154.
Penins.
No.
45,
Pongamia sericea Vent.,
(Vent.) Wight
sericea
Type:
Fl. Mai.
(1935) 1472; Backer
Dalbergia angustifolia Hassk.,
Millettia
Vol.
(1916) 280; Ridl.,
394; Burkill,
Mai.
BLUMEA
88.
—
(Vent.) Wight
& Arn.
Hassk.
ex
1764
Syntypes: Griffith
var.
malaccensis
(K), Maingay
518
Pram,
J. As.
Soc.
Beng. 66,
(A, K, L), Goodenough
1706
2
(n.v.),
Malaysia.
Notes
—
1. Stone 15598 from
different in other characters.
similar
to
M.
has rather small, thick leaflets, but is
Malaysia
6809 from Sumatra is in
Lorzing
the
chrysamaryssa; however,
of this
pods
some
quite
aspects
specimen clearly
not
belong
to
M. sericea.
2. Several
Miq.').
18-19
from Java have ± obtuse
specimens
The fruits of these
by
5-10
specimens
mm). However,
or
rounded leaflets ('var.
appear rather well
the seeds
are
flat
or
developed (size:
indented and
viable.No differences could be found in other characters. These
are
obtusa
50-55
probably
specimens
are
by
not
included
in M. sericea.
3. The type of M. sericea forma subcordata
by
is
Miq.
in New Guinea or in the Moluccas. However, this
given
as
specimen
collected by
was
probably
Zippelius
collected
Blume in Java.
4. A
The
specimen
specimen
with
was
a
Korthals label (Korthals
probably
collected in Sumatra
s. n.,
Borneo
A) gives
as were most
as
the
of Korthals'
locality.
specimens
of M. sericea.
5. 'Var. malaccensis Prain': Of the syntypes
(A, K, L)
were
Maingay
seen.
M. sericea; however,
one
518 is
a
Griffith
mixed collection:
sheet also bears theannotation
Another sheet is also numbered
The Leiden and Harvard
Maingay 1184, and
sheets
belong
M.
to
sheet also bears the label Maingay 508. No
a
'
1764
duplicates
and
at
Maingay
Kew
518
belong
to
11846, Feb 61868, Malacca'.
third sheet also
chrysamaryssa,
lectotype
(K)
Maingay
1182.
of which the Leiden
has been chosen.
4. Millettia solomonensis Verdc.
Millettia
Type:
solomonensis
Schodde
Bougainville.
&
Verde.,
Craven
Kew
3805
Bull.
32
(holo K;
(1977) 247;
iso
Man. New
Guinea
Leg. (1979)
A, BRI, CANB, L, LAE), Papua
New
332.
—
Guinea,
F. Adema:
Notes
1. Verdcourt
—
Notes
Malesian
on
Fabaceae
refrained from
(1977)
including
sections of Millettia.The absence of callosities, the
M. solomonensis
is
similar
quite
2. The
is needed
8 ft
to see
high shrub,
Fragiliflorae Dunn,
Backer &
(1923) 298;
Allen
311;
1
&
specimens.
flowers
J.
Linn.
NGF
13393 is
material
described
described
as
as
a
Fragiliflorae
Soc. Bot.
1
(1912) 136; Merr.,
41
Enum.
Man.
(1964) 616; Verde.,
R. Geesink
Leiden Bot. Ser. 8
Lectotype (here designated):
103.
(1984)
—
in Polhill
Philipp.
New Guinea
Allen, Leguminosae (1981) 543;
(1981) 258;
more
is
(Verdcourt, 1977).
Fl. Java
Bakh.f.,
Much
involved. NGF 31614
one taxon
Section
Section
of the
from Verdcourt (1977).
copied
it also differs in leaves and fruits.
deep purple-red
one
and M. sericea.
than
more
in
species
Millettia. In inflorescence and flower characters it
sect.
of the fruits and seeds have been
ifthere is
this
407
fruits and seeds indicate that
large
several small differences between the
are
shrub with
to
chrysamaryssa
M.
descriptions
3. There
an
belongs
to
Millettia
-
&
Flow.
PI. 2
Leg. (1979)
Raven, Adv. Leg. Syst.
Millettia merrillii
Perkins.
Jard. Malm.
Pongamia Vent.,
pinnata (L.)
Trees.
Pierre
Axillary
=
buds
large
with
—
hairy (at
Note
Millettiais conserved
—
The section
1
(rarely
Fragiliflorae
or
over
large
can
and thick. B:
group
consists of 3
carpa and M.
or
more
the
species:
imparipinnata (5- vel)
nata infra sericea ad fere
glabra.
—
Typus:
or
conserved
as
two
(Brummitt, 1984).
name
groups: A: Pinnata- group: Ovules
to
indehiscent; seeds usually
Ovules 3-10
(rarely less), usually small,
1,
(rarely 2); pods
flattened.
PINNATA-GROUP
new
species
S 27606
nov.
—
Millettia borneensis, and M.
(Soepadmo
1-seminalis
&
Fig. 2a, b,
7-foliolata. Foliola
Pseudo-racemi
(vel 3). Legumina plerumque
Raya,
brachy-
elliptica
axillares
iso
ad obovata breviter acumi-
pedunculo basi pulvinato.
indehiscentia
Smith) (holo L;
3a
circum
A, K),
semen
Ovula
tumescentia.
Borneo, Sarawak,
Bukit
1.4.1969.
shrubs, 4-50
wood white
to
m
high,
ochre.
Stipules obliquely elliptic,
at
Standard with
the base).
pinnata.
Arbor. Folia
low,
at
Inflorescences
Pacific Islands, Australia, Mas-
Merrillii/Xylocarpa-group:
5. Millettia borneensis Adema, spec.
Trees
hairs
some
peduncle.
Pongamia (Geesink, 1981b; Brummitt, 1984);
status
be divided into
A.
2
least
3); pods rather thick, tardily dehiscent
flattened, dehiscent; seeds 3
This
the base of the
Tropical Asia, throughout Malesia;
however, Pongamia has retained its
rather
enclosed in the budscales.
at
Islands.
carene
2
Type: Pongamiaglabra Vent., nom. illeg. [= Pongamia
pulvinus
a
basal callosities. Anthers
Distribution
—
pinnata (L.) Panigrahi],
button-shaped tightly
axillary, pseudoracemes
2
28.
(1803) t.
Millettia
c.
dbh 7-80
cm.
Bark
Twigs
terete, 3-4
4.5
2
by
mm,
mm
grey,
inner
bark white
diam., thinly hirsute
or
to
pale yelglabrous.
caducous, outside sericeous, inside sericeous
base. Leaves (5- or) 7-foliolate. Petiole 3-5
cm
long, striate, thinly
hirsute
to
gla-
408
Fig.
BLUMEA
2. Millettia
Adema;
M.
c,
d.
M.
Wight
Fragiliflorae
brachycarpa Merr.;
xylocarpa Miq. (a,
98642).
& Arn. sect.
Vol.
b: SAN 25299;
e.
c,
M.
45,
No.
2,
2000
Dunn. Leaflets
from below,
pinnata (L.) Panigrahi; f,
d: SMHI
1560; e:
g. M.
SAN 25767;
f,
g:
a, b. M. borneensis
velvetina
Adema; h,
NGF 10747 ;
h,
i.
i: KEP
F. Adema:
3. Millettia
Fig.
carpa
f. M.
Merr.;
Wight
M.
c.
& Arn. sect.
1917 ;
f: FRI
long. Leaflets:
4-9
flat
as
mm
or
long,
cm
mm
;
c:
long, striate, thinly
Pods.
-
Millettia
M. borneensis
a.
Merr.
&
Rolfe;
e.
NBFD 9455 ; d: PNH
hirsute
narrowly triangular,
c.
to
3
dry,
nerves
2-3.5
by
cm,
3-16
0.7
to
to
base cuneate, apex
8
or
9
409
Adema;
b. M.
M. merrillii
11130 ;
e:
brachyPerkins;
Ridsdale
mm,
glabrous; pulvinus
&
long, thinly
3-5
hirsute, midrib and
mm
acumen
hirsute
long.
to
lateral
apart;
1-9
mm
±
Bracts
ovate, 0.6-0.8
nerves
mostly
long; pulvinus
glabrous; peduncle
to
the
outside hirsute, inside sericeous.
the flowers
mm
(shortly) acuminate, acumen
more
mm
5-7
index 2.2-2.7, ± papyra-
cm,
side, 5-13
per
index 1.4-3.3,
cm
glabrous; pulvinus
2-flowered. Bracts
hirsute
obovate, 7-11.5 by 3-5
(thinly) sericeous, rarely
sunken above,
by
to
when
upwards
long. Inflorescences
long, thinly
(1- or)
cm
both surfaces
slightly
Dunn.
Fabaceae
d. M. ahernii
SAN 60019 ; b: SMHI 363
the terminal, 3.5-9
4-6
Fragiliflorae
terminal elliptic
often rolled
Malesian
32637).
brous; rachis 6.5-10.5
ceous,
on
pinnata (L.) Panigrahi;
xylocarpa Miq. (a:
Reynoso
Notes
by
0.4-0.5
3-6
brachyblasts
Brachyblasts
mm,
outside
410
BLUMEA
sericeous, inside sericeous
pedicel, elliptic
or
glabrous. Calyx
long;
mm
c.
'axe-shaped',
Keel
uous.
ceous
outside
3.5
by
inside
mm,
claw 3-3.5
2
c.
long,
mm
Stamens
almost absent, up
1.5-2 mm
to
0.1
ovules 2
long;
sericeous
stipe
cm,
2
c.
Seeds discoid to
oblong
valves
long,
mm
to
ventral
along
3
c.
at
long;
mm
with
mm,
margin
5-6
3.5-4
long,
mm
long,
mm
hairs
at
least in upper part,
18-20 by
outside seri-
mm,
pocket,
1.5-3.5
hirsute
by
1
or
sericeous; stipe
upwards
inflated, ± sharp-edged,
to
5.5-8.5
15-16
c.
long,
mm
the base. Disc annular
by
thick, outside glabrous
mm
inconspic-
3
sericeous in lower part,
half-moon-shaped,
transversely ellipsoid,
at
to-
elliptic
by
above lateral
upright,
some
glabrous
blade
apex, auricles
6.5-7
Pods green, flattened
1.7
c.
top of
base. Standard: claw 2-3
with basal fenestrae, free
part
(or 3); style
around the seed,
claw
boat-shaped,
high. Ovary
mm
at
retuse, outside sericeous,
mm,
blade ±
long,
at
hirsute, inside glabrous; lobes 5,
outside and inside sericeous
glabrous; stigma terminal, capitate.
bulging
to
glabrous. Wings:
mm;
Bracteoles
long.
mm
standard green
rather wide, auricles
10 mm
c.
5
c.
sericeous
10
by
anthers 0.6-1.0 by 0.5-0.6
glabrous;
or
margin,
8
c.
petals:
pocket
high.
mm
long,
(light) purplish,
ventral side, inside
on
lateral
mm
2000
2,
0.2-0.4 mm, outside sericeous, inside sericeous
by
blade ± orbicular, 9.5-10
wards the base and
or
4
Corolla
inconspicuous.
No.
45,
Pedicels
glabrous.
or
ovate, 0.4-0.6
to
Vol.
2-3.5
or
with
by
0.3-1.2
some
hairs.
4-8 mm,hilum 1—1.5
mm
long.
Distribution
Habitat &
sand
or
Sumatra, Peninsular Malaysia, Singapore, Borneo.
—
Ecology
(Disturbed) primary forest,
—
limestone. Altitude up
Fruiting: April
Notes
—
2. This
to
5
flat
in the
that often curl
do
pods
m.
Flowering:
is in
contain
not
3. Two
4.
several aspects rather
dense indumentum, the
more
upwards
in
tum
are
from the Bornean
Bornean
first group
are
far
are
plants
278.
Notes
Type:
—
1.
BS 9578
This
on
are
to
and
species
are
pinnata.
of section
It differs
thinner leaflets
sharp-edged pods
probably tardily
are recorded
Singapore
M.
usually
with
dehiscent. In
Fragiliflorae.
with
(pale)
orange flowers.
and Sumatra differ in indumen-
seem to
tending
to
become patent. The
the lower surface of the leaflets with the hairs
be
in flower and
no
pods
differences; however, all plants from the
are
really
needed for
a
final decision. For
have been included here.
Merr.
brachycarpa Merr., Philipp.
—
similar
narrower
± hirsute with the hairs
sericeous
6. Millettia brachycarpa
Millettia
to
Otherwise there
seen so
the present these
rivers. Soil:
In the first group the lower surfaces of the leaflets
specimens.
sericeous
specimens
appressed.
other
most
from Peninsular Malaysia
from Peninsular Malaysia,
(very) thinly
along
June, October, November.
the smaller flowers and the
drying,
differ from
they
specimens
Specimens
to
seeds, the undeveloped seeds
ripe
smaller, thinner seeds. As in M. pinnata the pods
this character
swamp forest,
March
insects.
developing
species
new
especially
100
November.
to
1. Very
filled with up
to
—
J.
Fig. 2c, d,
Sci.,
Bot.
(A, K, L), Philippines,
species
is
a
Philippine
10
3b
(1915) 17;
endemic.
2. Red exudate has been observed in several
3. Inflorescences of BS 29185
are
Enum.
Philipp.
Flow. PI. 2
(1923)
Palawan.
Only
foundin Luzon and Palawan.
specimens.
sometimes once branched.
F. Adema:
7. Millettia pinnata
Millettia
PI.
t.
Enum.
384; Merr.,
Backer &
271.
(1917)
Leg. (1979)
—
Robinia
mitis
A
Fl. Ned. Ind.l
xerocarpa
Jard. Malm.
&
210.
Hatusima
Notes
2.
(1907)
(1928) 494;
Man.
Herb. Amboin.
Prelim.
(L.) Kurz,
(L.) Merr., Philipp.
J.
(A),
This well-known
1245
J.
Bot. 5
Sci.,
Herb. Timor.
(1916) 306; Gagnep., Fl.
2
(1910)
—
101.
Flow. PI. 2
1797.
(1935)
208.
(1858)
Bot. 5
Philipp.
Rep.
(1910)
unknown.
—
mitis
Pongamia
Merr.
(L.)
Pongamia pinnata (L.)
—
(1923)
298.
(1845) 2; Hassk.,
Verz.
(1835) 448;
Gen. Indo-Chine
—
Type
unknown.
Bogor. descr.,
Hort.
(n.v.).
Bot.
56
Mag. (Tokyo)
(1942)
567.
40 km inward from
Guinea, Sennen,
—
Type:
widespread species really belongs
to
Kanehira
8.3.1940.
Nabire,
Millettia
sect.
1981a, b and Adema, 2000.
in New Guinea. The
are
and the indumentumof the keel
(Kostermans & Soegeng
2
Moritzi, Syst.
Hatus.,
Dutch New
See Geesink,
ed.
Sci.,
Enum.
Type: Zollinger
&
illeg.; Descaisne,
nom.
(1922) 593; Burkill, Diet.
& Moritzi in
Very variable, especially
petals
1
(1972) 303; Verde.,
mitis
Pongamia
—
mitis
Nutt. PI. 2
Bogor. descr.,
size, and indumentum of leaflets; there
the
Enum.
Cytisus
5
(1747) 184.]
illeg.
nom.
Pongamia
Heyne,
(Hassk.) Merr.,
—
12587
1.
—
1
—
Cochin.
117.]
t.
5
(1803) t. 28,
K.
Hort.
novoguineensis Kaneh.
Fragiliflorae.
—
(Hassk.) Merr., Philipp.
Merr. var. xerocarpa
(1858)
Siam.
Fl. Mai.
210.
(1989)
3.
(1743) 183,
Fl. Mai. Penins. 1
Pongamia grandifolia Zoll.
ed. 2
3
49.
(1855) 147;
2(1916) 441; Ridl.,
Millettia
1
Fl. For.
Pongamiapinnata (L.) Merr., Interpr. Rumph.
(1763) 1044,
(1875)
Fl.
(1923) 298; Craib,
Herb. Ambon.
Rumph.,
Pongamia xerocarpa Hassk.,
var.
2
(1964) 616; Whitmore, Tree
—
Distr.
Bilaspur
unknown.
—Type
Pongamia glabra Vent.,
Miq.,
1
312.
PI. ed. 2
L., Sp.
Veg. Pegu, App.
101.
Fl.
411
Millettia
-
Pongamiapinnata (L.) Pierre,
—
Herb. Ambon.
Nussanive
e
Panigrahi & Murti,
Type: Pluk., Phyt. (1690) 104, f.
[Malaparius Rumph.,
[Malaparius
3c
—
Flow. PI.
Philipp.
Fl. Java
Bakh.f.,
New Guinea
For.
in
Fabaceae
Fig. 2e,
741.
(1753)
Malesian
on
(L.) Panigrahi
pinnata (L.) Panigrahi
pinnatus L., Sp.
Notes
varies
species
in
greatly
shape,
often also small differences in the size of
petals.
One
group
of New Guinean
specimens
975, LAE 50088, McDonald <6 Ismail 3706, NGF 39315)
has much smaller fruits with wrinkled walls. Further field studies may show the existence
of
more taxa.
3.1 observed dark
Uses
—
(red)
sap in
stems
small branches
are
used
as
In Malesia this group consists of 7
ahernii Merr.
(1910) 71; Dunn,
278.
Type:
—
Rolfe, Philipp.
J. Linn.
BS 3373
Soc.
Enum.
cavitensis
179; Merr.,
Philipp.
Flow.
Philipp.
J.
PI. 2
J.
and
glabra
is described
as
new.
3d
Bot. 3
(1908) 103; Merr., Philipp.
(1912) 181; Merr.,
Sci.,
Twigs
UP
of which M.
Fig.
Sci.,
constructions.
used in local medicine.
(Ahern's collector) (K), Philippines,
Bot. 5
(1923)
Sci.,
Bot. 5
Flow. PI. 2
(1923)
Merr., Philipp.
Enum.
J.
Bot. 41
canariifolia Merr., Philipp.
178; Merr.,
Millettia
&
—
housing
are
I/XYLOCARPA-GRO
species
8. Millettia ahernii Merr. & Rolfe
Millettia
walls.
pod
toothbrush. Leaves
B. MERRILLI
Millettia
and
The hard wood is used for temporary
Enum.
—
Syntypes:
(1910) 72; Dunn,
279.
J.
Flow.
Sci.,
PI. 2
Bot. 5
(1923)
Luzon.
(1910) 71; Dunn,
279.
Philipp.
—Type:
J. Linn.
BS 4711
J. Linn.
Merrill 4181
Soc. Bot. 41
(n.v.),
4727
Soc. Bot.
(1912)
(K).
41
(1912)
(L, P), Philippines,
Luzon.
Notes
with
the
—
only
a
pedicels
1. Rather similar
slight overlap.
and the
to
M.
merrillii but
Also similar to M.
slightly larger
flowers.
larger
tenuipes
in size of leaflets and fruits
which differs in the
length
of
412
Fig.
e.
BLUMEA
4. Millettia
keel
petal
glabra
from
Adema.
outside;
a.
Habit;
f. staminal
b.
tube;
Vol.
45,
flower; c.
g.
No.
2,
2000
standard from
inside;
pistil (all De Vogel 4488).
d.
wing
from
outside;
F. Adema:
2. Of the syntypes of M.
do
3.
&
Soejarto
smaller
Fabaceae
coriaceous and
the seeds
as
9. Millettia glabra Adema,
ad 3
m
nov.
spec.
alta. Folia
De
Vogel
Treelet, up
4488
3
to
pinnatae similis
m
21
by
midrib and
glabrous,
apart; lateral mostly
1.6-3; pulvinus 4-7
2-4
c.
by
elliptic
Calyx
1
mm,
4
c.
long;
0.20-0.25
long,
mm
hairs. Corolla white
mm
blade orbicular,
mm
long.
3
mm,
Keel
petals:
by
0.2-0.5
claw
Ovary
5-6
at
Distribution
with eroded
Note
most
—
with
—
lumps
Very
glabrous.
some
hairs
Moluccas
—
8-12
hairs
long;
long,
c.
1
mm
primary
or
M.
the section
pinnata,
Podocarpae
10. Millettia merrillii Perkins
Millettia merrillii
Merr.,
Enum,
Perkins, Fragm.
Philipp,
K), Philippines,
—
Fl.
Flow. PI. 2
Luzon.
2-6
index
cm,
mm
Bracts ± linear,
Bracteoles
long.
rim
with
glabrous.
short
some
outside sericeous,
mm,
mm
long;
glabrous,
pocket
mm
2-3
mm
long,
the base. Disc annular,
or
elliptic,
auricles up
to
0.5
mm, out-
auricles inanthers 0.6
long, glabrous;
ovules 4
glabrous
blade
6.5-8.5 by 3
'boat-shaped',
inconspicuous.
5; style 4-6
open disturbed forest. Soil:
to
100
differing
m.
mm
trifoliata
and differs
Fig.
long,
sandy
(3),
the al-
species
Dunn from Burma. This
species
greatly
or
5).
The
or
Dec.
Asian
(4
only
in various characters.
3e
Philipp. (1904) 81; Dunn,
(1923)
clayey
Flowering: Sept.,
in the number of leaflets
and the number of ovules
of Millettia described with 3 leaflets is M.
to
by
base. Standard: claw 1-2.5
claw 2.5-3
long;
7.5—
long,
(Halmahera).
Dense
to
mm
Pods and seeds unknown.
of limestone. Altitude up
similar
8-27
long, glabrous; peduncle
glabrous,
greenish
long,
cm
4-6 per side, 9-40
inconspicuous.
free part 2-3
at
nerves
cm
apex, inside
blade
caducous.
outside sericeous, inside
9-11
by
at
especially
glabrous.
totally glabrous calyx,
belongs
mm,
acumen
Pedicels 5-7
ventral side, lateral
mm
Typus:
(broadly) elliptic,
ovate, 5.5-12.5
rather
glabrous. Wings:
mm
along
the base,
Ecology
above,
to
standard with
long, sericeous; stipe
mm
with few hairs
Habitat &
terminal
1.5-10
long,
0.1-0.3
some
3
c.
Stamens 10-13
mm,
elliptic
mm
emarginate,
outside with
side and inside sericeous
conspicuous.
1
by
tinged violet,
—
rachis 1.5-3.5
glabrous;
teeth inconspicuous, both sides
towards the side and base, inside
7.5-9.5 by
± sunken
nerves
outside sericeous, inside
triangular,
to
minor. Ovula 4 vel 5.
long. Leaflets:
mm
c.
longo,
mm
pilis paucis brevi-
margo
diam., glabrous. Stipules
mm
long. Inflorescences
mm
difference
only
acumine 8-27
inconspicuorum
± smooth,
long,
the terminal,
long, glabrous; pulvinus
cm
2.5
as
pods
7 cm).
by
index 1.8-2.3, base cuneate, apex acuminate,
cm,
both sides
mm
2-5
glabrous; pulvinus
3-9
cm
± cordate;
4
plurimum partium
terete, 3-10
high. Twigs
here. The
belong
lateral: 15
(late) elliptica,
dentium
differs in
specimen
ones
Halmahera.
(holo L), Moluccas,
Leaves 3-foliolate. Petiole 2-5
± smooth,
sed
These
seen.
fully grown).
are not
may
cm,
Fig.
—
3-foliolata. Foliola
utrinque glabra. Calyx utrinque glaber,
bus. Corolla M.
8.5
by
17
obtuse, lowest
more
4. Alston 16740 from Halmahera (Moluccas)
(terminal:
were
understood here.
as
(but probably underdeveloped
Arbuscula
413
of BS 4727
canariifolia only fragments
more
is the size of the leaflets
Millettia
-
6297 is included here. However, this
Reynoso
leaflets
aspects:
Malesian
on
differ much from M. ahernii
not
some
Notes
279.
—
J. Linn. Soc.
Bot. 41
(1912) 180;
Lectotype (here designated): Merrill
1387
(A,
414
BLUMEA
Millettia
foxworthii Merr., Philipp.
179; Merr.,
Enum.
PL
2
No.
45,
Bot. 5
Sci.,
J.
Flow.
Philipp.
Vol.
2000
2,
J. Linn.
(1910) 72; Dunn,
179.
(1923)
—
BS
Type:
740
Soc.
Bot. 41
(1912)
(Foxworthy) (A, K, L),
Philippines.
Millettia
capillipes Dunn, Philipp.
FB
Type:
Notes
seen
18549
—
only
2. Of the
Bot.
6
(1912) 316;
J. Linn.
of M.
type
specimen
I have
capillipes
specimens (Adduru 181,
11. Millettia platyphylla
Merr.
platyphylla
279.
Warburg 12578,
chosen
was
seen
ex
Merr.
Dunn,
BS
fragments
have
76897)
180. —
the
as
12891) I have
lectotype.
in K. These do
deviate
not
longer,
narrower
leaflets.
Dunn
ex
J. Linn.
Soc. Bot. 41
(1912) 140; Merr.,
C.M.
Lectotype (here designated):
—
(1912)
of M. merrillii.
specimens
(1923)
Soc. Bot. 41
Luzon.
1. Of the syntypes (Merrill 1387, 1625;
3. Several
PI. 2
Sci.,
Merrill 1387 (A, K). This
from other
Millettia
J.
(K), Philippines,
Weber 1123
Enum.
Philipp.
Flow.
(A, K, PNH?), Philippines,
Mindanao.
Notes
—
1. This
Leyte, Samar,
2. The
of this
pod
is
species
a
endemic collected in the islands
Philippine
Surigao,
and Mindanao.
is still unknown.
species
12. Millettia pterocarpa Dunn
Millettia
168
J. Linn. Soc. Bot. 41
pterocarpa Dunn,
Millettia
glaucescens
Notes
2. As
—
far
1.
auct.
Only
as
non
Kurz:
3. The
wings
are
perpendicular
the valves of the
13. Millettia
Millettia
the
on
than in
to
pod
—
pods
as
flowers and the
an erect
longer
Note
species.
—
The
2
(1897)
362
to
me
observe, this species
seems to
be
fruits.
winged
winged
Millettia species
in the Derrisgroup
continuation of the
are
wings
they
±
are
orientated
on
the fruit
parallel
to
sutures.
J. Sci.
30
(1926)
398.
—
TYpe: BS
44803
(Ramos
&
Edaho) (A,
Luzon.
a
flowering specimen.
length
of the
In many aspects rather
pedicels,
the
slightly
larger
anthers.
Pierre
var.
(C.T. White)Verde.,
Brass 841
Beng. 66,
Merr.
14. Millettia velvetina Adema,
Type:
pod;
known from the type,
Pongamiapinnata (L.)
Lectotype (here designated): Wray
of the Derris- group. In Millettia
M. ahernii but different in the
to
velutina
with
of this and similar
the valves of the
tenuipes
Only
J. As. Soc.
permits
species
winged species
tenuipes Merr., Philipp.
Note
Prain,
material
K, PNH?, SING), Philippines,
similar
—
known from Peninsular Malaysia: Perak.
the scanty
different from other Millettia
differently
175.
(1912)
Perak.
(K), Malaysia,
nom.
velutina C.T.
Kew
Bull.
(A, BRI, K), Papua
epitheton
nov.
32
White, J.
New Guinea.
velutina was
Arnold Arbor.
(1977) 250;
already
Man. New
(Non:
used
by
Millettia
Dunn
10(1929)
Guinea
222.
—
Pongamia
Leg. (1979)
velutina
312.
—
Dunn, 1912.)
(1912: 149)
for
a
Chinese
F. Adema:
15. Millettia xylocarpa
Millettia
xylocarpa Miq.,
Backer &
Millettia
Bakh.f.,
dehiscens
&
(Koord.
Millettia
techini
Millettia
—
Notes
iso
(holo K;
(1972)
1.
—
Soc.
297.
1
key)
or
J. As.
is, according
Soc.
Java 2
the latter
most
2
Fl. Mai.
small and
has
slightly
Diet.
as
in
The
only
rather
are
soon more
decipiens
differ
to
follows: M.
as
the key, however,
with ±
cannot
the
overlapping
known
or
to
of M.
1749 in
material)
hairs
are
Prain
are
(1912)
Scor-
—Type:
Linn.
Soc.
Bot. 41
Polo Kamiri.
because
glabrous.
less
Dunn
to
in the
(1912,
and M.
seems
to
more
pubinervis
be variable;
The
M.
M.
the
fruiting
already
decipiens:
pubinervis:: 'pubes-
persistence
of the
stipules
stipules
also
persistent
are not
material. As the differences
united here. The type of M.
are
the
Prain
decipiens
holotype
of M.
visible. As
as
the
no
keep
label
specimen
xylocarpa.
mm.
M.
the
are
They
slight.
The syntypes
than M.
also differ in
pubinervis elliptic
pubinervis:
shape
to ovate.
species separate.
clearly
at
of the
I think
reads 1747, which
Kew with
a
letter
by
was
as
the
corrupted
Prain attached
to
holotype.
xylocarpa
explained
are
to
pubinervis
longer pedicels
Scortechini 1749 for the collection chosen
original
the label. The
copying
triangular,
small
too
3.5-7
has almost
in the first
note
glabrous leaflets, only
this is what may be
few
are
here united into
remaining
expected
other differences between this material and the
dehiscence/decipiens’ -material. All species
M.
J.
hemsleyana: 'glabra',
be checked in
pubinervis
(1897) gives
decipiens
old leaflets. There
called
297.
(1935) 1472; Whitmore, Tree
'ubiquum sericeum'),
of the former have
M.
mm;
the sheet has been chosen
5. The
2
hemsleyana
persistent stipules,
species
the flowers: M. dehiscens
Although
lectotype
Soc. Bot. 41
(1972)
smaller flowers.
that these differences
4.
41
BO, K, L).
J. Linn.
1
according
differencebetween M.
species
specimens
M. dehiscens 10-13
bracts
iso
dehiscens
Soc. Bot.
densely hirsute; however,
are
3. The differences between M. dehiscens and M.
(and only
J. Linn.
in the hairiness of the flower parts: the former
only
in all leaves. Both characters
are
Millettia
—
(1897) 90, 360; Dunn,
(1928) 390; Burkill,
parts
(1912)
Malaysia.
key, however, 'plerumque glabrum').
vary
Soc. Bot. 41
Dunn, the persistence of the stipules: in the former until anthesis, in
to
'sparse pubescence' (in
to
(1912) 239;
'hairy'. Dunn, however, in his descriptions gives M. decipiens
Dunn describes the three
seems
96.
Dunn,
(holo L;
the latter early caducous. The hairiness in the corolla parts
cens' (in the
J. Linn.
(1895)
(1897) 360;
Koorders 4105
Beng. 66,
1
and M.
hemsleyana
glabrous,
Peninsular
and leaves
Young twigs
'sparse pubescent'.
as
2
Bot. 41
Lectotype (Dunn, 1912): Wray 3370 (K), Malaysia, Perak,
and Whitmore (1972)
less
Booms.
Beng. 66,
Soc.
Type: Kurz 1778, Burma (n.v.).
—
Beng. 66,2 (1897) 90,360; Dunn,
A, K),
415
(Leg. 188) (K), Java, Soerabaja.
s.n.
(1873) 68; Dunn,
378.
(1922) 585; Whitmore,Tree
of the looseness of the hairs
2. Millettia
(1916)
Soc.
Siam. Enum.
—
2
Beng. 42,
2
Millettia
-
J. Linn.
(1855) 157; Dunn,
Valeton, Bijdr.
J. As.
J. As. Soc.
hemsleyana Prain,
1
1
Fabaceae
3f
Lectotype (here designated):
(1912) 585; Craib, Fl.
Fl. Mai.
&
Fl. Mai. Penins.
1747
Ind.
Malesian
(1964)595. —Type: Horsfield
J. As.
Koord.
decipiens Prain,
172; Ridl.,
1
Valeton) Prain,
172.
(1912)
Fl. Ned.
on
Fig.
—
Fl. Gen. Indo-Chine
177; Gagnep.,
Pongamia
Miq.
Fl. Java
pubinervis Kurz,
Notes
one,
in rather
‘pubinervis/
which has
to
be
416
BLUMEA
KEY TO THE
Vol.
45,
No.
2000
2,
SPECIES OF MILLETTIA
IN MALESIA
la. Leaves 2-11-foliolate
2
2. M.
b. Leaves unifoliolate
2a.
Lianas, rarely shrubs ((M. solomonensis). Inflorescences without
base of the
peduncle;
ties. Anthers
b. Trees,
shrubs.
3a. Leaflets
Peninsular
Malaysia,
dull, thinly
Pods
phous.
by
2
b. Pedicels
c.
by
16
5-7.5
by
4-13
mm
compressed,
6a. Ovules
1-2
obovate
or
or
3-4.5
cm.
by
12-25
—
mm
Seeds
on
25-45
—
or
spongy,
by
7-11
mm.
Pods
Twigs
to
to
6-12
long.
5-6
glabra
valves 0.7-2.7
indehiscent,
Seeds
woody.
transversely ellipsoid,
mm
rather
7
or
'quadrate',
mm
thick, woody. Seeds flattened,
10-17
by
1-3
by
7-15
9
mm
....
sericeous
to
or
Pods dehiscent or indehiscent, valves
mm.
ovate, lower surface
—
spongy.
valves
glabrous, only
11-16 by 9-13
Coastal, rarely inland
sericeous, rarely
hirsute.
leathery
mm.
elliptic
mm
to
Seeds 4-8
to
valves
hirsute
woody.
mm
. ..
mm
long.
Philippines
hirsute.
Calyx
(then pedicels 2-10
14-16
by
tardily
pinnata
thick.
—
Peninsular
± sericeous. Blade of standard 10-
or
Seeds 10-15
obovate, 2.5-13 by 1-8
to
Pods
7. M.
5. M. borneensis
glabrous
hairs
when young
± hirsute. Blade of standard 9.5-10 by
leathery.
Borneo
b. Terminal leaflets obovate,
—
by
glabrous.
9. M.
mm
broadly
dehiscent,
appressed
blade of standard
Calyx
10-25
8
to
Pods indehiscent,
9a. Terminal leaflets
by
24
Pods thick,
by
and leaflets
mm
thick.
—
6. M.
7-12
c.
chrysamaryssa
diadelphous.
obovate, lower surface glabrous
to
b. Lower surface of leaflets
with few
diadel-
oblong,
mixed with trifoliolateones. Leaflets
midrib and veins. Blade of standard
Malaysia, Sumatra,
11
Stamens
long.
±
compressed,
transversely ellipsoid,
(broadly) elliptic.
4-15
by
8a. Lower surface of leaflets
by
—
6
to
dehiscent, valves fibrous
mm.
3).
Inland
b. Terminal leaflets elliptic
strigose
2 (or
monodelphous, rarely
Seeds
cm.
3-8. Pods dehiscent, valves 0.6-0.7
Blade of standard 7-11
11
0.4
rarely (M. pinnata))
transversely ellipsoid, oblong,
10
by
Stamens
5. Moluccas (Halmahera)
10-35
woody.
hirsute. Ovules
or
ovate
7a. Terminal leaflets elliptic
or
long.
2, rarely 3. Pods (tardily) dehiscent
b. Ovules
5
Sumatra,
—
3. M. sericea
thick, leathery, ± fibrous
thick,
mm
14 by 5
c.
long. Calyx
by
hairy
4. M. solomonensis
6-7
b. Leaves with 3 leaflets. Leaflets
Ovules 4
strigose
1. M.
mm
3
(1 or) 2 (or 3)
4
brown
mm
4
the
.
solomonensis)
the base;
at
sericeous below. Ovules 3-6.
silvery
densely,
5a. Leaves with 5-11 leaflets,
elliptic
or
at
Java
to
long. Calyx
mm
1.5-3.5
to
the base ( M.
Solomon Islands
Bougainville,
4a. Pedicels 5-9
at
pulvinus
a
Standard with basal callosities. Anthers
shiny, densely golden
b. Leaflets
hairs
some
pulvinus
a
Standard without basal callosi-
brachyblast.
Inflorescences with
brachyblast.
per
with
glabrous, rarely
rarely
flowers
5-10 flowers per
galliflagrans
12-17
9.5-18
mm
by
2.5-4
mm
cm,
mm
long),
or
(then pedicels
2.5-5
cm,
Blade of standard 15-18
by
lower surface
long
glabrous
or
and blade of standard
calyx
6-7
10-30 mm
lower surface
11-18
Philippines
brachycarpa
mm.
mm
long)
long
....
and
10
thinly strigose.
Pedicels 8-13
11. M.
mm
platyphylla
F. Adema:
10a. Pedicels 10-30
16
mm
b. Pedicels 2-10
Pods 2-15
mm.
2-3
by
1-3
by
1 la. Pedicels 10-14
mm
b. Pedicels 25-30
mm
6-7
mm
Millettia
-
417
Blade of standard
long.
2.5-4 mm
(for
cm
Fabaceae
14-16
12-
by
(not known for M. tenuipes)
cm
long. Calyx
mm
Malesian
on
long. Calyx
Pods 15-21
mm.
Notes
M. velutina
long.
Blade of standard
long.
Blade of standard
11
Blade of standard 7-12
long.
only
pods
young
14-15
are
12-16
by
by
12
.
known)
mm.
6-12
Ovules 4-8
8. M. ahernii
16
c.
17
by
Ovules 8
mm.
13. M.
12a. Lower surface of leaflets hirsute,
glabrescent.
Peninsular
—
tenuipes
Malaysia, Java,
Borneo, Lesser Sunda Islands, Papua New Guinea
b. Lower surface of leaflets
glabrous
with few
or
13
hairs.
appressed
7 leaflets. Lower surface of leaflets hirsute, hairs
or
attached. Pedicels 2-4
not
known.
mm
Pods 2-6.5
long.
1
by
straight,
attached
loosely
Pedicels 3.5-7
remain).
11-15 by
1-2
mm.
mm
(in
long.
Peninsular
—
curly,
±
firmly
young).
Seeds
14. M. velvetina
b. Leaves 7-11-foliolate. Lower surface of leaflets ±
rather
rather
(all
cm
New Guinea
Papua
—
Philippines,
10. M. merrillii
Lesser Sunda Islands
13a. Leaves with 5
—
old leaflets
Pods 8-14
hirsute,
only
1.5-2.5
by
glabrescent,
few hairs
Malaysia, Java, Borneo,
cm.
hairs
the midrib
along
Seeds
12-16
by
Lesser Sunda Islands
15. M.
xylocarpa
DUBIOUS SPECIES
Millettia longipes Perkins,
Fragm.
Enum.
(1912) 178; Merr.,
Fl.
J. Linn. Soc. Bot. 41
Philipp. (1904) 80; Dunn,
Flow. PI. 2
Philipp.
279.
(1923)
—
Syntypes:
Warburg
12094, 12095, 12112.
Up
to
now
fragments,
the
no
Warburg
12112:
12095:
the
consisting
of
a
impossible
various
to
rather
this
as
sheet
and
a
a
does
are
the
preceding
a
belong
not
no
this
second sheet
pod
a
with
12095. Without
having
to a
a
to
which of
three draw-
panicle-like
Millettia species.
of three
drawing
belong
it is also
a
it
specimen,
may
lectotype;
to
by
K. These
at
indication
accompanied
drawing
a
one,
to
fragments
some
bear
complete flowering specimen
leaflet. The
choose
pod valve,
fragments
really belongs
Warburg
to
one
The
specimen
M. merrillii. On
pod
probably belongs
flowers and
they belong.
sheet
same
the second
species, probably
it is
a
quite clearly
on
to
two
collections
ings: Warburg
rescence,
added
other material has been found than
leaflets,
two
pods,
ifthe valve
a
represents
annotated
inflo-
Warburg
Millettia
Warburg
12112
Millettia species; the leaflet
seen
impossible
any
to
other
tell
to
material
original
which
species
the
parts belong.
Millettia stipulata Dunn, Philipp. J. Sci., Bot. 6
(1912) 316;
J. Linn. Soc. Bot. 41
181; Merr., Enum. Philipp. Flow. PI. 2 (1923) 280. —Type:
FB 11140
(1912)
(Aquilar),
Philippines, Luzon, Angat.
From this
to
a
Millettia
duplicate
most
a
seen
leaflet and
species; however,
of the
other
I have
species
of an inflorescence,
type
only
a
of a
twig
with
in K
fragments
of Millettia.
accompanied by
stipules.
several characters need
is still needed. The
Philippine species
fragments
some
part
to
seem to
These
a
drawing
parts may belong
be clarified. A
complete
indicate differences with
418
BLUMEA
Millettia
The
239.
is
There
also
1
number of
specimens
No.
45,
2000
2,
(1855) 156; Dunn, J. Linn.
Java
seen
by
Bot. 41
too
to
belongs
to
a
Millettia
M. sericea.
be identifiedwith any of the
cannot
of Millettia. However, the material is
that
me
specimen belongs
that
Soc.
Bantam.
,
a
from
specimen
III 3997 17. This
Zollinger
are
species
Ind.
Type: Zollinger?, Java,
—
only Zollinger
species
Fl. Ned.
zollingeri Miq.,
(1912)
Vol.
scanty
to
be described as
new
existing
species.
1. SAN 55901, Sabah.
This
specimen belongs
with 2 ovules
to
From this and the other
species
has scattered lenticels. In all
wall
are
(seemingly)
absent
Fragiliflorae
sect.
to
M.
the group
to
brachycarpa.
of the Pinnata- group it differs in the fruit wall, which
species
or
and within that section
The thickness of the seeds leads
(Pinnata- group).
of the Pinnata-group the lenticels of the fruit
confined
to
the
sutures
of the fruit.
2. Native collector 204, Sarawak; native collector 689, Sarawak; Korthals
s.n.,
Kali-
mantan.
The first
has inflorescences with young pods; the second has young
specimen
the thirdhas very young flower buds. All have leaflets
2 ovules;
panicles;
These
pods probably winged.
young
specimens probably belong
to
from those of other Millettia species with
Kooy 821,
This
to
Millettia. The inflorescences differ
winged
specimen belongs
to one
of the
As the flower size
may represent
a new
4. SF 22586 (R.
The fruits
xylocarpa.
seems to
be
species
a
larger
greatly
of these
the
M.
Fragiliflorae.
It is rather similar
leaflets.
Fragiliflorae,
this
specimen
species.
specimen
are
quite
different from the fruits of the
species
specimen
has
true
panicles
is rather
pachycarpa,
pods
as
pods
valves without visible lenticels. This
species
rather similar to those of
inflorescences. SF 22586 has
incomplete
which has
are
it is hard to
with almost
with many scattered lenticels, whereas SF 22586 has
nous
identity
Henderson), Malaysia, Pahang.
of this
cinerea. The latter
As
of sect.
useful character in the
Millettiafound in Peninsular Malaysia. The
to
the
However, it differs from both in the larger flowers;
from M. merrillii it differs also in the
similar
now
Lesser Sunda Islands, Timor.
M. merrillii and M.
panicle.
fruits. For
is unclear.
specimens
3.
pods,
glaucous below; axillary pseudo-
pods
specimen
identify.
glabrous,
a
pseudo-
In Millettia it is
reticulate valves
with smooth,
may represent
of
Callerya
a
densely
new
veluti-
species.
5. PNH2591 (V.R. Marabalal).
This
There
specimen
are no
is in
indumentum rather similar
other data available than:
Philippines,
to
M.
alt. 75
velvetina of New Guinea.
m,
17.V. 1948.
F. Adema:
Notes
on
Malesian
EXCLUDED
Millettia atropurpurea (Wall.)
Fabaceae
Millettia
-
419
SPECIES
Benth., PI. Jungh. (1852) 249; Miq., Fl. Ned. Ind. 1
(1855) 157.—Type: Wallich 5910 (K), Amherst. = Callerya atropurpurea (Wall.)
Schot. See Schot
(1994).
Millettia koordersii Backer
ex
Koord.-Schum., Syst. Verz.
Koorders 26430 (L), Java.
Type:
=
Sophora wightii
1,
128 (1911) 43.
1,
—
Baker. See Backer & Bakhuizen
den Brink (1964).
van
Millettia litoralis Dunn,
Philipp.
J. Sci.,Bot. 6
(1912) 316;
181; Merr., Enum. Philipp. Flow. PI. 2 (1923) 279.
250 (K),
The Kew
Benth. The
Philippines, Mindanao,
of the type
duplicate
structure
J. Linn. Soc. Bot. 41
De Vore
Type:
—
&
(1912)
Hoover
St. Cruz.
has been identifiedas: Derris cf. scandens
specimen
of the inflorescences and flowers
seem to
confirm this identifi-
cation.
Millettia ? luzoniensis A.
Gray,
Bot. U.S.
1
Expl. Exped.
(1854) 456; Merr., Philipp.
J. Sci., Bot 3 (1908) 82; Dunn, J. Linn. Soc. Bot. 41 (1912) 239. —Type unknown.
=
Gliricidia
sepium (Jacq.)
Steud. See Merrill (1908: 82;
Millettia monticola Kurz, J. As. Soc.
Montaban.
=
Beng. 42,
Derris monticola (Kurz)
2
(1873)
1923: 280).
67.
Prain. See Prain,
Type probably
—
J. As.
Soc.
from
Beng. 66,
2
(1897) 361.
Millettia
rostrata
(Hassk.) Miq., Fl. Ned. Ind. 1 (1855) 155.
Hassk., Flora 2, Beibl. (1842) 53. —Type: Hasskarl
rostrata
—
Java
s.n.,
(n.v.).
Type: Dorgelo
The type and
1770 (L), Java, Kediri, baai
only specimen
leaflets, inflorescence
(L), Borneo.
=
of this
species
van
Dalbergia
Blume ex
Fordia
Miq.,
(1964) 596.
Popoh.
consists of
fragments only:
and flowers. Identification leads
axes,
Millettia splendidissima
s.n.
=
Hassk.
Millettia rufa Backer, Blumea 5 (1945) 511; Backer & Bakh.f., Fl. Java 1
—
rostrata
Dalbergia
to
Fl. Ned. Ind. 1 (1855) 156.
splendidissima (Blume
ex
loose
petioles,
Derris spec
—Type:
Miq.) Buysen.
Korthals
See
Buysen
(1988).
Millettia
—
thyrsiflora (Benth.)
Pongamia
Zaga
PI.
thyrsiflora Benth.,
Syntypes: Philips
corallaria
Pohon
Rumphius?
Fl. Ned. Ind.
1
s.n.,
(1855)
Ormosia calavensis Azaola
PI. As.
elliptica (Wall.) Benth.
Fl. Ned. Ind. 1
Malacca (n.v.).
(1855)
=
156.
Aganope
(1971).
149.
Herb. Amboin. 3 (1743) 175,
Pongamia elliptica Wall.,
Derris
Jungh. (1852) 249; Miq.,
Penang (n.v.); Griffith
Polhill. See Polhill
Miq.,
Rumph.,
=
s.n.,
ex
t.
Blanco.
Rar. 3 (1832) 20,
t.
—
[Corallaria latifolia
seu
110.] —- Type: The plate by
See Merrill (1917).
237.
—
Type:
not
indicated.
=
BLUMEA
420
Pongamia
grandiflora Vent.,
Jussieu 15666 (P).
Pongamia grandifolia
Ind. 1 (1855) 147.
=
Vol.
Jard. Malm.
Derris
No.
45,
2000
sub
(1803)
elliptica (Wall.)
2,
Zoll. & Moritzi in Moritzi,
Syst.
28, in obs.
t.
(Geesink
Benth.
Verz.
Herb.
Type:
—
litt.).
Fl. Ned.
(1845) 2; Miq.,
Type: Zollinger 1245, Java, Propoli (n.v.).
—
in
Millettiapinnata
=
(L.) Panigrahi.
Fl. Ned. Ind. 1
Pongamia horsfieldii Miq.,
Patjitan (n.v.).
=
Derris
Pongamia hypoleuca Miq.,
ris
(1855) 149. —Type: Horsfield
elliptica (Wall.)
Fl. Ned. Ind. 1
s.n.,
Java,
Benth. See Kew Index.
(1855)
148.
—Type:
indicated.
not
=
Der-
elliptica (Wall.) Benth. See Kew Index.
Pongamia paniculata Wight
not
indicated.
=
Derris
heyneana
Nom. ed.
Pongamia piscidia Steud.,
Millettia
Pongamia
Benth., J. Linn. Soc. 4, Suppl. (1860)
ex
taiwaniana Matsum.
=
—
Type:
2, pt. 2 (1840) 380.
Type:
—
indicated.
not
=
See Kew Index.
piscidia Wight.
indicated.
110.
Benth. See Kew Index.
ex
Hayata,
Millettia pachycarpa
Ic. PI. Formos. 3
Benth. See T.C.
(1913)
79.
—Type:
not
& H. Ohashi, Flora of
Huang
Taiwan, ed. 2, 3 (1993) 337.
in Hook., Bot. Misc. 3
Pongamia triphylla Wight
Derris trifoliata Lour.
Pongamia uliginosa DC.,
(1833)
41.
t.
—
Type
unknown
=
See Kew Index.
Prod. 2 (1825) 416.
—
Type:
indicated.
not
=
Derris tri-
foliata Lour. See Kew Index.
Pongamia
volubilis Zoll. & Moritzi in Moritzi,
Ind. 1 (1855)
(Wall.)
Pongamia
149.
—
Syst.
Verz.
(1845) 3; Miq.,
Fl. Ned.
Derris
elliptica
148.—Type: Zollinger 325, Java, Tjidurian (n.v.).
=
Benth. See Kew Index.
volubilis Zoll. & Moritzi
var.
glaucophylla
Type: Zollinger ?, Java, Tjidoerian (n.v.).
=
Miq.,
Derris
Fl. Ned. Ind.
1
(1855)
elliptica (Wall.)
Benth.
ACKNOWLEDGEMENTS
The loan of material
is
used in
gratefully acknowledged.
at Kew
smit
new
for their kind
Sachs made
species
the
this
study
hospitality
drawings.
from the herbaria
A, BO, FRI, K, L, LAE, SING,
to thank the director and
I like
and
help during
Dr. J.F.
the members of the
my visit there. Mr. J.H.
Veldkamp kindly
translated
the
van
and WAG
Leguminosae
group
Os and Mrs. A. Wal-
English diagnoses
of the
into Latin.
REFERENCES
Adema,
F.
1999. Notes
maryssa,
Adema,
a new
on
species
Malesian Fabaceae
from Peninsular
F. 2000. Millettia and
Nature
Journal.
(Leguminosae-Papilionoideae).4.
Malaysia.
Novon 9:
Millettia
chrysa-
289-291.
Pongamia (Leguminosae-Papilionoideae)in
Malesia. The
Malayan
F. Adema:
O.N. & E.K. Allen.
Allen,
nodulation.
C.A.
1981. The
MacMillan
Brummitt, R.K.
Backer,
1984.
Publishers
Malesian
on
of the
A
Millettia
-
book
source
of
421
characteristics,
and
uses
London.
for
den Brink.
van
Fabaceae
Leguminosae.
Ltd.,
of the Committee
Report
& R.C. Bakhuizen
J.R.M. 1988. Revision
Buysen,
Notes
Lordia
genus
Spermatophyta:
1964. Llora of Java
27. Taxon
1. P.
33:
298.
Noordhoff, Groningen.
(Papilionaceae: Millettieae).Blumea
33: 239-
261.
S.T.
Dunn,
1912. A revision
R.
Geesink,
(eds.),
1981a. Tribe
Advances
R.
Geesink,
Vent.
1981b.
R.
Proposals
1908. The
Philipp.
conserve Millettia
to
E.D.
Millettiearum.
1917. An
Leiden
Philippine plants
et Am. J. Linn.
Botanic
Royal
Soc.
In: R.M.
(1964).
W. & A. and revise
Bot.
Polhill
123-243.
41:
& PH.
Raven
Kew.
Gardens,
the conservation
ofPongamia
30: 327-329.
Bot. Ser.
collected
8.
the Wilkes United States
by
Exploring Expedi-
Bot. 3: 73-84.
Sci.,
J.
Wight
Hutch.
245-260.
Legume Systematics 1:
1984. Scala
Merrill, E.D.
Merrill,
Millettia
genus
Tephrosieae (Benth.)
(Leguminosae-Papilionoideae). Taxon
Geesink,
tion.
in
of the
6.
of
interpretation
Herbarium Amboinensis. Bureau
Rumphius's
of
Printing,
Manila.
Merrill,
E.D.
(Leguminosae).
Prain,
D.
Schot,
A.M.
of Philippine
1923. An Enumeration
Some observations
Polhill, R.M. 1971.
Bull.
Kew
1897. Order 38.
on
of
1977. New
B.
T.C.
Wight,
taxa of
Endl.
39:
of
Beng. 66,
Malaya
from New Guinea.
1
M.
chrysamaryssa
M.
galliflagrans
485: 7
Peninsulae
24394:
—
7;
Bakar
5862:
7;
27276:
7;
Beccari
9.
M. pterocarpa
borneensis
5.
M.
13.
M.
M.
brachycarpa
14.
M.
velvetina
7.
M.
pinnata
15.
M.
xylocarpa
8.
M. ahernii
16.
M.
Achmad 39:
1291: 3—Alston
26759:
3;
4396:
7
13442:
7;
28802:
7;
286:
—
3;
3062:
7
7;
—
17319:
7;
van
16985:
7;
29892:
7;
30581:
3070:
Blume
5;
2903
(Balajadia): 7;
7
Boden
10811: 7
1567:
—
7;
Bruggeveen
(Sablaya): 11;
689
Kloss
7926:
204
(Daboy): 16;
3;
181:
848:
spec.
10
—
1081:
3;
72: 7
Atasrip
17364:
7;
Aet 453: 7
129:
307:
7;
Aet &
—
1961:
1; 1193:7;
—Atjeh
6497:
15;
31094:
7;
7;
—
3285
17483:
3;
36129:
21437:
7;
3522: 7
—
7
1;
Idjan
2090:
7
—Atmodjo
—
Haglen 903:
7
—
(Foxworthy): 10;
1076:
3380
14062:
22967:
7;
7;
—
21930:
BRUN 5149
1633:
2218: 7
2490
(Ramos): 10;
7;
7;
—
—
(Melegrito): 7;
3766
27742:
(Kamis): 7;
5151
7;
841:
14;
7— Brooke
(Ashton):
(Agama): 7;
2178
7
7
7;
25409:
7;
Brass 821:
7;
3686:
9727:
(Ashton): 7;
587
bb
—
3;
35063:
Beaman
(Polonio): 7;
139: 7
(Bermejos): 6;
1450
—
7;
7
24399:
7;
7
(Tandum): 7;
Branderhorst
245
32894:
Beguin
1787
15191:
24068:
7;
7
20770:
7;
Backer & Van Slooten
Bartlett
—
(Balajadia): 7;
14796: 7
19440:
7;
—
21834:
7; 8209:7; 14061:7;
&
3
7
32880:
BNBFD
(native coll.): 16;
740
7;
den Brink
3487:
(Balajadia):
1150:7; 1225:7;
—
Adduru
Alvins
31079:
1312: 3
(Kamis): 7;
—
—
150:7
7;
28066:
7;
Bakhuizen
16983:
7;
1169: 7
—
tenuipes
3.
16958:
3;
7;
16740: 8
—Andrews 31:
9335:
25800:
worthy): 7;
platyphylla
6.
4048
7;
M. merrillii
M.
1728:
BS 58
glabra
M.
10.
12.
2494
8095:
list
11.
—
225-251.
Orientalis. London
solomonensis
Bloembergen 4376:
—
Indiae
32:
Lumpur.
M.
—Aumeeruddy
3;
Bull.
Kuala
M. sericea
11857: 7
8958:
Whitfordiodendron)
3.
Amdjah541: 5
Backer
and
4.
620:7—ALM
—
Kew
Longman, London,
1.
1834. Prodromus Florae
2.
2: 21-75.
(including Padbruggea
Identification
AbangMuas
Dalbergieae-LonchocarpineaeBenth.
1-30.
Leguminosae
1972. Tree Flora
R. & G.A.W. Amott.
J. As. Soc.
Callerya
(Papilionaceae: Millettieae). Blumea
Whitmore,
in
25: 259-273.
Leguminosae.
1994. A revision
Verdcourt,
Plants 2: 278-280.
Flowering
limits
generic
613
(Ramos):
7
(Fox-
10;
3373
BLUMEA
422
4727:
(Ahern's collector): 8;
11548
(McGregor): 6;
16316
23233
6;
27313
39824
(Ramos): 7;
41449
(Ramos): 7;
Edano): 10; 75191 (Ramos
7
—
BSIP 5666
9874
(Gafui
7
4
al.):
et
3321
(Schram): 7;
5864
7;
15540
15;
4040
21287:
828:
De
3
no.
5
—
Coode
Curtis
—
3146:
9;
717
5072
Ebalo
345:
916
7;
10;
13334:
10;
FB
&
5:
555:
677:
7;
9
3773
7;
(Curran):
10064
1038
13162
—
FRI
120
(F. Ng): 7;
5305
(Chan): 15;
23644
98907
(Ismail):
632:
7
395:
(Wagadje): 7;
der
et
16774
al.): 4;
(Van
4;
164:
857
2743
Leden):
(Versteegh): 7;
7839
8556
9214
(Vink): 7;
14898
(Schram): 7;
(Schram): 7;
266
32637
Forbes
Clemens 64:
16
7;
(Foxworthy):
127:
Craven
—
2380: 7
7;
—
939:
Derry
12690:
10;
—
7;
8048:
7;
& Schodde
J.J.F.E. de Wilde
15061:
3;
1288
21344:
3;
3
—
15.
13035:
7;
18549:
11485
3469
10;
10;
13181:
20242
3103a:
3;
2090
3848
(Whitmore): 15;
4321
6844
14471
12094
(Chan):
: 15;
(Wong & Khairuddin): 15;
27040
36471
22700
Fraser
(Whitmore): 7;
(Kochummen):
(F. Ng): 15;
27049
71201
1;
(Whitmore): 15;
7567
(Cockburn): 7;
(F. Ng): 15;
(Saw): 7;
—
(Whitmore): 3;
20889
15;
7
(Whitmore): 1;
15149
(Everett): 15;
19917
7559
(Chan): 7;
—
(Ahmad): 15;
3332:
995
(Whitmore): 15;
10
(Tamesis):
(Kalong): 15;
3;
(Kochummen): 1;
(Kochummen): 3;
(Ahern's coll.):
(Hutchinson): 7;
5072
(Bidier): 15;
2868:
564
20740
2040
23817
(Putz): 7;
4180
(Mahamud): 15;
7;
1424
(Borden): 7;
(Ahern's coll.): 8;
1809:
(F. Ng): 15;
(Everett): 1;
23679
15
6166:
7;
(Naus): 15;
17188
(Whitmore): 15;
14159
3126
(Hamid): 7;
13613:
3;
428:
15;
(Whitmore): 7;
15;
2155:
15;
3373
(F. Ng): 7;
5309
—
et al. 6436:
(Whitford): 7;
4040
5): 15;
1862
2306
12776:
(Borden): 7;
14226
—
15;
3142: 7.
(Loh Hoy Shing): 15;
(Putz): 7;
(Walton): 1;
1279
15
(Cockburn):
17238
132:
Elmer 5695:
—
Eyma
Guard
3473
(Loh): 15;
29102
Gjellerup
al.):
(Gafui
2967: 3.
3;
Voogd
Berger
(Curran): 7;
(F.
866:
Chin 2154: 7
Wilde-Duyfjes
—
(F. Ng): 15;
10632
(Whitmore): 15;
3
(F. Ng): 7;
5118
13441
De
Den
(Whitmore): 15;
(Kochummen): 7;
(Loh): 1;
de
(Walker):
2091
(Cockburn): 7;
844:
3;
—
15;
—
Coode, Wong,
—
(Foxworthy): 15;
1837
(F. Ng): 7;
15
(Ahern's coll.): 10;
12588
23315
(Symington): 3;
1061
7
(Clark): 7;
2890
3126
no.
(Arshad): 15;
163: 7
—
Elsener 216:
(Borden): 7;
F.M.S. Field
et
(Buwalda): 7;
5361
7524
(v.d. Rijdt): 7;
12281
(Iwanggin): 7;
684
(Versteegh): 7;
8157
15087
(Ramos):
Burkill & Shah
—
2443
(Kooiman): 7;
4746
(Mauriasi
et
&
(Ramos
80571
(Mauriasi
1
27164
(Ramos): 10;
47011
al.):
et
BW
—
(Versteegh): 7;
(v.d. Rijdt): 7;
10 —Elbert 3279: 7
—
(Whitford): 7;
2044
16120
5257: 7
2380
7522
(Abd. Aziz):
5811:
4488:
9;
Smeets 5695: 7
10
9594
(Reillo):
(Ramos):
(Sulit): 7;
(Ramos): 7;
(Mauriasi
19289
22382
33358
13;
Edano):
Babala): 4;
al.): 4;
7;
(Kalkman): 7;
(Ahmad):
1211: 7 —W.J.J.O. de Wilde &
Demoulin
et
25672
(Fenix): 8;
7.
21288:
3135:
14420
4658:
7;
10198
(Iwanggin): 7;
&
al.): 4;
(Koster): 7;
7872
&
80378
Burkill & Holttum 8662:
—
(Malinka): 7;
(Schram): 7;
C.F. Field
—
867: 7
7;
3444
(Moh. Nang): 15;
5;
Vogel
(Rappard): 7;
7005
7
(Agama): 6;
(Cenabre): 7;
(Nakisi
15672
(Wester): 10;
(Phasis): 7;
11391
(Merrill): 6;
10;
21602
(Ramos
(Mauriasi
3280:
1226
10120
(Koster):
3093:
7693:
(Koster): 7;
7870
(Iwanggin): 7;
15925
et
(Loher):
19153
29967
(Ramos): 10;
8233
(Mauriasi
Buwalda
—
(Iwanggin): 7;
(Schram): 7;
Cantley
10
1223
(Brouwer): 7;
76897
Edano): 8;
Biinnemeijer
—
14245
24233
44803
10;
(McGregor):
&
11789
al.): 4;
et
193:
Burley
—
(Cenabre): 6;
(Mauriasi et al.): 4;
15913
al.): 4;
(Barros): 6;
29185
9578
(Ramos): 10;
(Escritor): 7;
24078
(Whitmore's coll.): 4;
(Runikera
8061
(Ramos): 7;
7; 20884
(Reyes):
(Alambra, Caulas): 7;
(Ramos): 7;
2000
2,
(Otanes): 10; l8539(Alvares): 10;
20637
(Tarrosa): 7;
No.
45,
(Ramos): 8;
13255
(Merrill): 8;
17937
(Reillo): 7;
19539
10;
5221
8;
Vol.
13385
15724
21746
(F. Ng): 15;
(Wyatt-Smith): 7;
7.
—
Goetghebeur 3496: 7;
3801:
7
—
Goodenough
1841:
3
—
Griffith
1764:
3;
1781: 7.
Hallier 7:
5;
Hardial
181:5;
656:
Hirano
—
324:
670:
7;
& Hotta
Hoogland 4359: 7;
Iboet
12:
Jacobs
—
7;
527:
4739: 7
Jeswiet
7
—
—
7;
3
1107:5;
—
1098: 5
5146:
Ismail
4313: 7
Haviland
7;
—
—
5;
14
Haniff & Nur 6937:
3
2142:
437:
386:
Hollrung
5229:
—
7
—
Horsfield
7
—
Hellwig
880:
Holmberg
Leg.
27:
—
7;
40:
3
7
Haniff Shah
7
—
Hoogerwerf
—
—
10351:
Henderson
Hose
499:
7;
15
—
18425: 7
120:
581:
7
—
5.
5-55: 7.
Jacobs & Mendoza
1002: 3
—
529:
7878:
Junghuhn 64: 7;
7
—
223:
Jacobson
3;
260: 7.
2154:
3
—
Jaheri
319: 7
—
Jensen
37: 7
F. Adema:
Kairo
207:
230:
7
7
—
Budin
&
82:
66424
Saw): 7;
15
6844:
3;
—
4105:
7;
17669:
15;
7;
821:
7;
23834:
16
5975:
17670:
7;
—
3
Lane-Poole
216: 7
—
314:
1
519:
27417
7
19118:
15;
317:
Kuswata
15
&
(T.
14191:
7;
Kooy 553:7; 811:
21241 A:
81:3
7; 21641:
Kostermans &
—
Krukoff 4459: 7
—
3;
(Gadoh
Koorders
—
12969:
—
15;
7
P.):
7;
5804:
5;
1791
KL
—
12968:
7;
(429,219):
18650:
15;
3235
P.): 1;
12967:
7;
775: 7
7;
98549
Kertosono
—
5625:
7;
al.
(Aziz
(Lindong): 15;
15
(bis?):
et
Karim
—
98820
4169:
3;
7
KEP 51906
—
(Wyatt-Smith): 7;
3163
3179:
&
(T.
51948
al.): 7;
et
Zainudin & Bedul
256:
7
10;
6809:
Lobb
—
2416:
77222
13097:
3;
4145:
310:
15
1
2417:
10;
—
—
10;
Ledermann
2572:
4
Kuntze
10;
5
7322:
7;
Loher
—
5123: 7
(Stone):
Lam 2499: 7
—
7205:
1636: 7
17085:
7;
53704
Lelean): 7;
(Womersley):
Loeters
—
16856:
3;
&
(Streimann
(Kerengaet al.): 7;
2415:
3;
71201
Kerr
7;
Kostermans &
—
Wirawan
74291
Lewis
5866:
3;
&
3
13034:
7;
1513:
7;
199: 7.
15;
10;
3012
10835:
7;
—
Kjellberg
—
Kosterfnans 9156:7;
(Wiakabu
—
2414:
10;
5661:
3;
7
15
1740:
5;
423
(S. Chelliah): 15;
7; 17673:7; 24242: 3;
Latiff,
—
15
4163:
12587:
1221:
7;
98668
Umbai): 15;
7;
Millettia
-
(Wyatt-Smith): 7;
1420:
& Anta 662:
Katik): 7;
13257:
2413:
5616:
7;
3
109:
50088
&
(Conn
Leeuwenberg
2267:7;
17672:
1;
10718:
anak
Kostermans
—
(Floyd): 7;
66052
3;
4162:
7;
Kostermans
15 —Kuswata
LAE 6579
—
(Gadoh
681:
(Rahim Ismail):
709:
3;
7366:
Korthals 873:
—
71119
Fabaceae
& Hatusima
Kasim
—
(S. Chelliah): 15;
104266
3;
7;
Kanehira
—
(Walters): 7;
4159:
15;
975: 7
Soegeng
7;
1794
Umbai): 3;
4096:
7299:
3;
Malesian
on
351: 7
coll. 401:
King's
7019:
7
7;
98642
(Rahim Ismail): 7;
1982:
anak
268:
7;
(Rahim Ismail): 15;
98907
2234:
Kajewski
Karta
—
Notes
—
Lorzing
—
7;
1022:
5163:
Lutjeharms
—
7
2266:
7;
5427: 7.
Mahmud
817:
W.
1437:
302:
1633:
10;
428:
NBFDA61
7
3
W.
A 524
10747
14;
18432
14;
25686
7
(Streimann
&
&
(Womersley
32229
&
Maskuri
Ismail
3831:
7;
7
1387:
8 —
4181:
10;
938: 7
3706:
—
10;
Merrill
Monod de Froideville
37599
(Katik): 7;
47330
(Johns): 7;
49643
&
6284
(Rahman): 7;
1553
21999
100:
7;
(Henty):
(Millar
8619
&
27511
7;
29848
(Brown): 7;
Smith): 7;
17406
22233
(Havel):
(Gilison):
(Streimann): 14;
(Coode & Katik): 7;
37109
44278
31614
(Womersley): 7;
39315
Vandenberg): 7;
&
(Womersley
(Havel): 7;
(Leach): 7;
(Mann): 7;
7794
(Cuadra):
(Kadir): 7;
& L.S.
(Sayers): 7;
(Henty):
34356
43288
17395
1440
A 2613
(Dadswell
Simmonds): 14;
27135
A
(Cuadra): 7;
(Kadir): 7;
(Haas): 7;
Croft): 7;
(Vandenberg & Mann): 7;
(Streimann
&
46782
(Streimann): 7;
7.
10340: 7.
8338:
2591
7
—
7890
18721
(Pop): 6;
39048
(Kondo
80783
(Gutierrez
87842
(Reynoso): 7;
117681
52: 7
—
&
&
261
—
Panoff
(Mendoza): 7;
39400
91213
et
117924
(Stone, Reynoso
—
&
Pfeiffer 48:
(Mendoza
(Steiner): 7;
40487
91496
7
—
PNH
81912
10;
—
150096
Pullen
38168
42505
(Allen):
7373:
7
—
7
(Frake): 7;
(Reynoso): 6;
97890
7;
(Edano): 8;
(Mendoza): 7;
87788
(Palo): 10;
(Edano): 10;
11130
(Frake): 7;
(Cordero & Espirito):
10
1136
Convocar): 7;
38021
(Madulid etal.): 8;
Romero):
&
(Gutierrez):
(Mendoza): 10;
(Mendoza): 7;
al.): 10;
7
(Umali): 7;
81904
Espirito): 7;
192:
10357
36455
(Conklin): 7;
Edano): 7;
(Gutierrez
PPI
1220: 7
Paijmans
(Marabal): 16;
13318
7;
—
1001:7; 1368:7;
—
1230
(Henty & Lelean): 7;
(Isles
Vandenberg): 7;
A
(Womersley): 7;
42287
Paie
8
&
A 2576
(Puasa): 7;
(Frodin): 7;
Kairo): 7;
Ogata
127: 7
(Wright): 7; 13393:4;
29469
Kairo): 14;
Merrill
2921:
10;
5;
(Yap Tan): 7;
(Womersley
19392
26633
al.): 7;
et
McDonald
—
—
(Cuadra): 7;
4842
7106
(Streimann & Lelean): 7;
(Gilison
20:
NGF 159
—
11113
(K.J. White): 14;
2801:
10;
Mondi
720
A 2462
(Cuadra): 7;
(Floyd): 7;
6579
—
7
1517: 7.
(Enggoh):
(Ridsdale & Lavarack): 4;
37196
15: 7
(Kadir): 7; A
A 4062
10559
2673:
7;
Mitchell
5505: 7
8521:
Majaducon
—
82-291:
5;
& Vermeulen
2036:
7;
601: 7
(K);
80-130:
1;
(Kadir & Kambatan): 5;
(Mair): 7;
27779
—
3
(A, L),
Muat & Grimes
—
(Cuadra): 7;
Gray): 7;
1
Meijer
1968:
10;
(Agama): 7;
1845
518:
78-107:
7;
—
(Puasa): 7;
1698
A 3185
9455
77-186:
Blancoana958:7
7;
A
Maingay
—
5569:
Meijer
spec.
7;
3
Maxwell
—
—
(Mendoza):
Powell
Purseglove
30:7;
4127:
1;
4346: 7.
Rahmat si Boeea
240?): 10;
1362:
15;
Ridsdale
RRI4:
3
3950:
740:
2592:
1090:
—
15;
10
RSS
3;
7635:
1085:
7;
7;
3645:
—
183
3
7;
Ramlanto
—
1243
(=
4753:
Ridsdale &
(Corner):
Vidal
15;
8402:
Reynoso
4.
282:
7; 421:
1243?): 10;
5;
1917:
11508:
10
—
7
—
Rau 422: 7
1258: 7
—
Ridley
15; 11509:5;
Robinson
243
—
RPVF 240
64:
14677:
7;
164:
15;
7;
(=
Vidal
251:
15113:
15
(PI. Rumph. Amb.): 7
7;
—
—
424
S
BLUMEA
16037
Chai
7
al.): 5;
Sabana
31706
7
—
62375
72514
84364
(Arsid): 7;
96408
&
&
&
123489
63794
115218
(Amin): 7;
129303
Schodde
Schodde
—
8303
(Haniff): 1;
(Haniff Shah): 15;
12454
(Burkill
15532
(Haniff): 3;
19813
Kloss): 7;
21929
24959
15;
&
34552
Samsuri
3553:7
438
3845:
15
7347:
—
Sudin &
—
800:
5
7;
Valeton
—
5003SPK:
58:
2439: 7
13600:
7
—
—
10
7
3;
—
10
5027: 7
—
Royen
3240:
1085:7; 1238:7;
10;
Kade
—
7817:
& Mahmud
7
8158:
10;
15
648:
Tuyama
7
—
—
Sider
20160:
Symington
363
(Ridsdale):
1560
8;
8
(Rids-
Soejarto&
Soejarto
—
&
Soejarto, Reynoso,
Soepadmo &
—
40788
Shah &
Soejarto & Reynoso
—
10; 6972:7
9160:
Stone & Mahmu
—
8576:
10
(Corner):
M.
6 —
et al. 6972:
& Fernando
(Kiah): 3;
& T.J. Whitmore
1062B:
6;
3;
(Boden
(Henderson):
31622
(Ridsdale): 6;
Soejarto
8157:
19204
15 —
& T.C.
15
—
Haniff): 3;
(Sinclair): 3;
1810:
3;
15
(Holttum): 3;
&
24270
(Corner): 16;
919:
(1749):
20417
39046
Santos
4206:
(Boden Kloss):
Haniff): 3;
1062A:
& Edrada 6602:
—
15;
14758
Sidiyasa
(Podzorski): 6;
15598: 3
Unesco
Leach
7
—
7;
Verheijen
1243:
—
Majaducon
Takasi
2304: 7
129:
—
& Noor
126977
—
15016:
Suhaima
15060: 3
7;
—
3.
Teijsmann
4256
HB:
7491:
3;
7;
7.
Hulstijn
7
439: 7
4009:
(Burkill
(Kalong): 15;
10827:7 —SMHI147
Soepadmo
Hong Fan
(Frodin,
Balgooy
M. Shah
15
9399
(Spare): 16;
(Amin): 5;
3101:
Holttum): 3;
29453
(Corner): 5;
36333
114799
(Henderson): 15;
Soejarto, Gaerlan, Segal
—
—
20242
102497
al.): 7;
1747
&
88746
Toyuk): 7;
(Amin): 7;
7;
12452
84107
Haya): 7;
et
Scortechini
(Burkill
22678
Soejarto27:7
Madulid &
Sun
13909:
7;
Van
—
3298:
1;
806
10 —
1810:
2468B: 7
7;
Van
246:
—
—Sinclair
1; 14221:7;
—
3667
Van
15
Soejarto, Reynoso, Sagcal
Tailor
(Frodin): 7;
3284:
7393:
30: 3
13894:
1498:
1732:
Awang Kela): 1;
(Henderson): 3;
Tachun): 5;
&
&
122834
126804
(Nur): 3;
16665
Haniff): 3;
(Podzorski): 6;
Soejarto,
14002:
Mayer
13862:
et al.
621
&
bin Kiah
Rutz & Edrada 6502:
205:7 —Stone
908
Shah
Sidek
—
10;
6297: 8
7;
Sagcal,
M.
&
&
Sangkachand
—
(Burkill
11338
(Henderson): 16;
(Daud
(Ridsdale): 8;
Madulid 6172: 8
Tahir
—
8662
25988
—
(Saikeh
(Stone): 7;
(Sigin
(Amin et al.): 7;
7
Han):
al.): 5;
et
67206
(Jumatin
(Amin
109705
(Amin): 7;
4548: 7
4;
(Pawachee
(Corner): 15;
Simaga254:7; 863:7
1800
Fernando
6290:
3
(Ridsdale): 7;
dale): 6;
20166
22586
35662
(Spare): 5;
—
15;
25837
(Holttum): 7;
&
(Burkil
(Holttum): 3;
(Sinclair & Kiah):
6;
16462
92454
1176:7; 1636:7;
(Henderson): 3;
12947
15;
7;
Rahim
(Amin
(Madani): 5;
86721
Haya): 7;
23589
(Burgess): 7;
(Abdul
(Aban Gibot): 7;
(Paul
Ching):
Abdul): 5;
60041
81175
37624
Puan
28418
55901
102477
118135
(W. Meijer):
& Craven 3805:
11192
&
(Amin): 7;
Schmutz
—
66951
90795:
126780
(Ashton): 7;
(Yii
bin
(Kadir
Gombio): 5;
Dewol): 5;
(Amin): 7;
(Haniff& Nur): 15;
Haniff):
(Henderson):
&
103163
130232
149: 7
&
(Amin
Rahman): 7;
(Aban
42289
(Singh): 7;
(W. Meijer): 5;
86397
7;
16859
26314
(Dewol Sundaling): 7;
(Amin): 7;
(Amin et al.): 7;
2733: 7
&
115532
SAN
Kunak): 7;
78142
(Amin): 7;
126013
al.): 5;
et
10150
7;
96602
—
51597
Sindin &
(Dewol
5178: 7 —Saunders
SF 3446
Uji
90201
1
21487
(Ashton): 7;
(Soepadmo & Smith): 5;
(Rena George): 7;
(Amin, Joseph
85573:
(Talib Bidin): 7;
Sumbing): 5;
(Amin
7;
60019
(Binson
17860
(Fabis): 7;
(Burgess): 7;
102844
(Amin): 7;
4700:
40409
252:
25767
(Brand): 5;
Haya): 7;
(Amin): 7;
38972
2000
2,
27606
al.): 7;
et
al.): 7;
et
No.
45,
Haji Bakar): 5;
(Jumati Kadim): 7;
Kodoh): 7;
(Sigin
(Amin
Chai
(Dewol Sundaling): 7;
(Lantoh): 7;
(Dewol
bin
(Paul
Samat bin Abdullah
25299
5;
(Dan
Chai
(Paul
(Muin Chei): 7;
55962
Lantoh): 7;
—
38555
13:
(W. Meijer):
16;
26745
(Anderson): 7;
et
—
16519
(Anderson): 5;
25120
Vol.
Van Niel
5453: 7
8;
1962
Expedition
Students): 14;
5205:
7;
—
3391:
2575:
3979:
7;
7;
169:
4565B:
Van Slooten
3519: 7
122:
7;
207:
4635:
7;
& Backer
—
195:
3;
15
—
UPNG
7.
Van Borssum Waalkes
—
2917:
2573:
Limestone
&
Vidal
5
35063:
240:
10; 2615:6,(K): 7;2651:
10;
768:
7;
Van
—
10;
7
—
740:
2672: 7
1624:
7;
7;
Ooststroom
Van Steenis
7;
—
740bis:
7;
Von Romer
22: 7.
Weber
1123:
& Janzen
258:5
—
Yates
11
Wenzel
—
1404:
8;
1645:
Williams 2787: 7
11;
—
—Worthington 13018:7—Wray
Wugadje
1663:
Zainudin,
—
3415: 7
3;
2637:
10;
Winckel
168:
12;
3294:
17688: 3
548:
15;
11
—
—
Widjaja
4530: 7
Wiriadinata
2364:
3;
3164:
3;
&
3310:
2443: 7.
2366:
Hahid
5
Salleh
—
Yoshida
& Zulkifli
2491: 7
Mohamad
5215: 7
—
Zollinger II
3997
17: 3.
—
Wieringa
Prawiroatmodjo
15;
3608:
15
F. Adema:
Notes
on
Malesian
Fabaceae
Millettia
-
425
Index
Accepted
names
The number
=
excluded
in
appear
after each
reman
name
synonyms
type,
is the number of the
Aganope thyrsiflora (Benth.) Polhill
excl.
excl.
Cytisus pinnatus
L. 7
Hassk.
Hassk.
Benth. excl.
Prain
(Kurz)
Lour.
excl.
sepium (Jacq.)
[p. 404]
Nussanive
Wight
& Arn.
Fragiliflorae
sect.
Macrospermae
sect.
Typicae
Rumph.
Dunn
atropurpurea
ex
7
(Vent.) Wight
excl.
[p.
var.
major Miq.
var.
malaccensis Prain
var.
obtusa
Dunn
407]
[p. 404]
3
dub.
tenuipes
Merr.
13
excl.
canariifolia
Merr. 8
glabra Vent.
chrysamaryssa
dehiscens
grandifolia Zoll.
foxworthii
Valeton)
2
12
15
koordersii
ex
Backer
Koord.-Schum.
Merr.
excl.
7
Merr. 7
(Hassk.)
ex
Benth. excl.
Merr. 7
xerocarpa
(Hassk.)
Pierre
velutina
Merr. 7
7
C.T. White
14
Steud. excl.
taiwaniana
Matsum.
triphylla Wight
& Hatus. 7
uliginosa DC.
velutina
Benth. excl.
pinnata (L.) Panigrahi 7,
excl.
7
xerocarpa
pinnata (L.)
7,
sericea Vent. 3
10
excl.
excl.
ex
excl.
var.
novoguineensis Kaneh.
Merr.
(L.)
piscidia
monticola Kurz excl.
platyphylla
mitis
var.
dub.
luzoniensis A. Gray
piscidia Wight
Kurz
pinnata (L.)
litoralis Dunn excl.
Perkins
(L.)
paniculata Wight
glaucescens auct.
Perkins
mitis
var.
9
hemsleyana Prain
pachycarpa
15
10
Merr.
galliflagrans Whitmore
glabraAdema
Prain
excl.
& Moritzi
excl.
hypoleuca Miq.
&
15
excl.
horsfieldii Miq.
15
& Valeton
7
8
(Koord.
Blanco excl.
ex
407]
Wall. excl.
grandiflora Vent.
1
+
excl.
10
Adema
decipiens Prain
404
Koord.
elliptica
cavitensis Merr.
dub.
Miq.
dehiscence
Merr. 6
Dunn
14
15
calavensis Azaola
brachycarpa
capillipes
3
Adema
corallaria
5
excl.
Miq.
ex
Benth. excl.
Pongamia Vent. [p.
(Wall.) Benth.
3
3
Miq.
4
Blume
Dunn
Ormosia
Miq.
borneensis Adema
Verde.
zollingeri Miq.
8
3
brachycarpa Miq.
xylocarpa Miq.
[p. 404]
volubilis
var.
Dunn
11
ex
Hayata
excl.
excl.
(C.T. White) Verde.
14
Zoll. & Moritzi excl.
glaucophylla Miq.
xerocarpa
3
3
stipulata
velvetina
Hassk.
ex
3
Miq.
turgida Miq.
404 +
& Arn.
3
Miq.
aurata
thyrsiflora
[p. 403]
Merr. & Rolfe
argentea Zipp.
Steud. excl.
Dunn
sect.
merrillii
bold.
excl.
Miq.
var.
splendidissima
7
Malaparius Rumph.
longipes
in
species;
dubious
excl.
solomonensis
Gliricidia
ahernii
=
forma subcordata
excl.
e
Backer
forma
Lam.
Malaparius
(Hassk.)
rostrata
excl.
Galedupa
Millettia
printed
names are
(Millettia)
splendidissima (Blume ex Miq.)
Buysen
new
the text; dub.
sericea
Benth. excl.
monticola
trifoliata
3
excl.
elliptica (Wall.)
heyneana
Fordia
in
rufa
Dalbergia angustifolia
Derris
italics , and
species.
Callerya atropurpurea (Wall.) Schot
rostrata
in
species
Hassk. 7
pterocarpa
Dunn
12
Robinia
pubinervis
Kurz
15
Sophora wightii
mitis L. 7
excl.
excl.
excl.