This article appeared in a journal published by Elsevier. The attached copy is furnished to the author for internal non-commercial research and education use, including for instruction at the authors institution and sharing with colleagues. Other uses, including reproduction and distribution, or selling or licensing copies, or posting to personal, institutional or third party websites are prohibited. In most cases authors are permitted to post their version of the article (e.g. in Word or Tex form) to their personal website or institutional repository. Authors requiring further information regarding Elsevier’s archiving and manuscript policies are encouraged to visit: http://www.elsevier.com/copyright Author's personal copy Quaternary Science Reviews 30 (2011) 1155e1172 Contents lists available at ScienceDirect Quaternary Science Reviews journal homepage: www.elsevier.com/locate/quascirev Deconstructing the Last Glacial termination: the role of millennial and orbital-scale forcings L. Menviel a, b, *, A. Timmermann c, O. Elison Timm c, A. Mouchet d a Climate and Environmental Physics, Physics Institute, University of Bern, Sidlerstrasse 5, CH-3012 Bern, Switzerland Oeschger Centre for Climate Change Research, University of Bern, CH-3012 Bern, Switzerland c IPRC, SOEST, University of Hawai’i, 2525 Correa Road, Honolulu, HI 96822, USA d Département AGO, Université de Liège, Liège, Belgium b a r t i c l e i n f o a b s t r a c t Article history: Received 29 September 2010 Received in revised form 15 February 2011 Accepted 16 February 2011 Available online 11 March 2011 Using an Earth system model of intermediate complexity forced by continuously varying boundary conditions and a hypothetical profile of freshwater forcing, the model simulates Heinrich event 1 (H1), the Bølling warm period, the Older Dryas, the Antarctic Cold Reversal (ACR) and the Younger Dryas in close agreement with paleo-proxy data from different regions worldwide. The ACR can be simulated as the bipolar seesaw response to the AMOC recovery during the termination of H1. However, this study also demonstrates that the amplitude of the ACR can be further amplified by a rapid deglacial retreat of the Antarctic Ice sheets. We suggest that melting from both, the Laurentide and the Antarctic Ice sheets contributed to the sea level rise associated with Meltwater Pulse 1-A (MWP-1A). It is hypothesized that the northern hemispheric source of MWP-1A caused the Older Dryas cooling in the Northern Hemisphere, whereas the Southern Hemispheric source contributed to the ACR. The study also documents that for the majority of paleo-climate proxies considered here, the relative timing can be qualitatively reproduced by the transient modeling experiments. The climate model solution presented here may provide a means to further constrain dating uncertainties of some of paleo-climate proxies during the Last Glacial Termination. Ó 2011 Elsevier Ltd. All rights reserved. Keywords: Last Glacial termination Heinrich event 1 Bølling warming Older Dryas Antarctic Cold Reversal Younger Dryas MWP-1A Earth system model of intermediate complexity 1. Introduction Elucidating the driving mechanisms of glacial cycles requires both an accurate knowledge of the timing and magnitude of external forcings and an understanding of internal climate feedbacks that determine the overall climate response to these forcings. In addition, paleo-climate reconstructions that can be used to independently test the hypothesized mechanisms are essential for ascertaining our comprehension of these changes. Paleo-modeling efforts have played an important role in quantifying the climate response to particular glacial-scale forcings. Using coupled atmosphere-ocean models, these efforts have mostly focused on particular time-slices, such as the Last Glacial Maximum (LGM, 21 ka B.P.) (Bush and Philander, 1998; Liu et al., 2000, 2002; Hewitt et al., 2001; Kitoh et al., 2001; Kim et al., 2003; Shin et al., 2003; Timmermann et al., 2004; Braconnot et al., 2007) or the * Corresponding author. Climate and Environmental Physics, Physics Institute, University of Bern, Sidlerstrasse 5, CH-3012 Bern, Switzerland. E-mail address: [email protected] (L. Menviel). 0277-3791/$ e see front matter Ó 2011 Elsevier Ltd. All rights reserved. doi:10.1016/j.quascirev.2011.02.005 Mid-Holocene Optimum (9e6 ka B.P.) (Joussaume et al., 1999; OttoBliesner, 1999; Liu et al., 2003). Results from these timeslice experiments have been compared thoroughly with existing paleoreconstructions, such as CLIMAP-Project-Members (1981), GLAMAP (Pflaumann et al., 2003) and other proxies (see e.g. Shin et al. (2003) for a careful LGM model-data comparison). This modeling approach has helped to explore the spatial characteristics of past glacial climate change and understand the relative contributions of different external forcing factors (Rind, 1987; Timmermann et al., 2004; Justino et al., 2005). However, in order to understand the physical mechanisms responsible for abrupt climate change and to identify the existence of leads, lags and phase-relationships in the climate system, it is mandatory to simulate the temporal evolution of the system in response to the time-varying forcing factors. Recently the transient modeling framework has been applied to climate models to understand the Late Quaternary climate evolution and its relation to slowly-varying boundary conditions (Jackson and Broccoli, 2003; Felis et al., 2004; Lorenz and Lohmann, 2004; Charbit et al., 2005; Tuenter et al., 2005; Renssen et al., 2005; Lorenz et al., 2006; Author's personal copy 1156 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Lunt et al., 2006; Marsh et al., 2006; Timmermann et al., 2007, 2009a; Timm and Timmermann, 2007; Liu et al., 2009; Timmermann and Menviel, 2009). We recently presented a transient model solution for the Last Glacial Termination using the Earth system model of intermediate complexity LOVECLIM that was driven by changes in greenhouse gases, icesheet topography and albedo and orbitally-induced changes in solar radiation (Timm and Timmermann, 2007). Millennial-scale freshwater forcing was however not considered. More recently, Liu et al. (2009) performed a transient simulation for the period 22 to 15 ka B.P. using a state of the art CGCM (NCAR CCSM3) that also captured Heinrich event 1 (H1) and the transition to the Bølling warm period (Timmermann and Menviel, 2009). The meltdown of the Laurentide and Eurasian ice sheets started around 20 ka B.P. An acceleration occurred during H1 (17.8e15.2 ka B.P.), as massive icebergs broke off from the disintegrating ice sheets. The resulting discharge of freshwater into the North Atlantic (Vidal et al., 1997) reduced surface density and eventually halted the formation of North Atlantic Deep Water for about 2500 years, as suggested by the 231Pa/230Th and ventilation age data from North Atlantic sediment cores (McManus et al., 2004; Gherardi et al., 2005; Thornalley et al., 2011) (Fig. 1a). At the same time, temperatures over Greenland remained low (Alley, 2000) while temperatures over Antarctica started to rise (Jouzel et al., 2007) and atmospheric CO2 concentration increased from 190 to 220 ppmv (Monnin et al., 2001). The end of H1 is marked by an abrupt warming over Greenland and the North Atlantic starting around 14.7 ka B.P. Liu et al. (2009) suggested that this Bølling warming was due to the rapid resumption of the AMOC following H1, combined with the glacial-interglacial increase in atmospheric CO2. Fig. 1. (a) Time series of the maximum of the meridional stream function (Sv) in the North Atlantic for experiment DGNS (green) compared to 231Pa/230Th data (black) from core OCE326-GGC5 (McManus et al., 2004) (filled circles) and from core SU81-18 (Gherardi et al., 2005) (empty circles) as well as ventilation age data (gray) from cores RAPID 10-1P, 154P and 17-5P (Thornalley et al., 2011). The time series of (McManus et al., 2004) was shifted by 200 years backward to better fit the GISP2 data (Alley, 2000). H1 stands for Heinrich event 1, BWP for Bolling Warm Period, OD for Older Dryas and YD for Younger Dryas. (b) Time series of the maximum overturning strength in the North Pacific for experiment DGNS (green) compared to the ventilation ages (yrs) as recorded in marine sediment cores from the Western North Pacific (black). The ventilation ages curve is the smoothed spline interpolation of averaged benthiceplanktic foraminifera ages and projection ages in the Western North Pacific (Okazaki et al., 2010). (c) Time series of dissolved oxygen content (ml/ L) averaged over the California basin (135 W-114 W, 25 N-40 N) for experiment DGNS (green) compared to a composite of two d15N record (black) from the California Basin (marine sediment cores JPC-56 (Pride et al., 1999) and ODP 893A (Emmer and Thunell, 2000) and compared to the Mo/Al data from core ODP 893A (gray circles) (Ivanochko and Pedersen, 2004). (d) Freshwater input (Sv) in the North Atlantic (black) and in the Southern Ocean (green) as applied in experiment DGNS. Time series are 21 year running means for the model outputs. (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Subsequent to the Bølling warming, between 14.7 and 13 ka B.P., the temperature over Greenland showed some pronounced fluc tuations of about 5 C within 400e500 years. Two major cold events were identified during this period in the Norwegian Sea (Lehman and Keigwin, 1992) and in Greenland (Stuiver et al., 1995): the Older Dryas (w14 ka B.P.) and the intra-Allerød cold period (13.5e13 ka B.P.) (Fig. 2a). During the same time, a 2 C surface cooling, known as the Antarctic Cold Reversal (ACR) is reconstructed for Antarctica (Petit et al., 1999; Jouzel et al., 2007). Concurrent to the ACR, sea level rose by about 20 m in about 500 years (Fairbanks, 1989; Stanford et al., 2006). The origin of this rapid sea level rise, referred to as Meltwater Pulse 1A (MWP-1A), has been the subject of an intense debate. While Licht (2004) and Peltier (2005) propose a Northern Hemispheric origin, Kanfoush et al. (2000); Clark et al. (2002); Bassett et al. (2005) and Carlson (2009) advocate the idea of a Southern Hemispheric origin of MWP-1A. Evidence for 1157 a meltwater pulse originating from Antarctica at around 14.6 ka B.P. was found in several marine sediment cores of the Southern Ocean. Ice-Rafted Debris (IRD) layers were observed in sediment cores from the Southeast Atlantic (Kanfoush et al., 2000), the Southwest Pacific (Carter et al., 2002) and from around the Antarctic Peninsula (Noumi et al., 2007). Freshwater discharge in the Southern Ocean is also inferred from depleted oxygen values in planktic foraminifera as well as increased occurrence of marine diatoms (Chaetoceros spp) in a sediment core from the Atlantic sector of the Southern Ocean (Bianchi and Gersonde, 2004). Recent studies by Clark et al. (2002) and Bassett et al. (2005) further suggest that in order to reconcile coral-based sea level reconstructions during MWP-1A with predicted glacial isostatic adjustment, one has to invoke a significant melting of Antarctica. Bassett et al. (2005) estimated that the Antarctic icesheet could have contributed by up to about 15 m sea level rise during MWP-1A. The main potential source regions for this pulse in Antarctica were identified as the Weddell Fig. 2. (a) Time series of spring season (MAM) air temperature anomalies ( C) averaged over Greenland (47 W-0 , 60 N-80 N) for experiment DGNS (green) compared to the temperature reconstruction from Greenland ice core GISP2 (black) (Alley, 2000). IACP stands for Intra-Allerød Cold Period. (b) Summer (JJA) air temperature anomalies ( C) averaged over the Netherlands (0 E10 E, 45 N-55 N) for experiment DGNS (green) compared to chironomid-inferred July air temperature anomalies ( C) from Hijkermeer, the Netherlands (Heiri et al., 2007). (c) SST anomalies ( C) averaged over the Iberian margin (11 W-8 W, 37 N-40 N) for experiment DGNS (green) compared to a reconstructed temperature index (black) obtained by averaging different alkenone-based SST reconstructions from cores SU81-18 (Bard et al., 2000), MD95-2042 (Bard, 2002), ODP-161-977A (Martrat et al., 2004) and MD01-2444 (Martrat et al., 2007). The time axis of the alkenone-based SST reconstructions was shifted forward by 200 years to better fit the model results and the other northern hemispheric paleo-proxies. (d) Winter season (DJF) SST anomalies ( C) averaged over the Cariaco basin (55 W-52 W, 10 N-14 N) for experiment DGNS (green) compared to Mg/Ca SST reconstruction ( C, black) from core PL07-39PC (Lea et al., 2003). As marine productivity is the highest during the winter season, this season was chosen to best represent temperature variations. Anomalies are with respect to pre-industrial time. Time series are 21 year running means for the model outputs. (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) Author's personal copy 1158 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Sea (w9 m) and the Ross Sea (w5 m). Finally, recent modeling studies (Philippon et al., 2006; Pollard and DeConto, 2009) suggest that the accelerated melting of the Antarctic icesheet started around 15 ka, further supporting the notion of significant deglacial freshwater forcing of the Southern Ocean during the ACR. The ACR could be explained in terms of the bipolar seesaw response to the AMOC recovery during the Bølling warming (Stocker, 1998), but there is also the possibility that a near-synchronous rapid melting of the Antarctic icesheet may have contributed to the Southern Hemispheric cooling (Menviel et al., 2010a). The Younger Dryas (YD) transition to colder Northern Hemispheric climates occurred around 12.9 ka B.P. (Rasmussen et al., 2006). During this period temperatures over Greenland, the eastern North Atlantic and the Caribbean Sea dropped by 8 C, 2 C and 4 C, respectively (Fig. 2). It has been hypothesized that the YD was caused by a weakening of the AMOC (McManus et al., 2004) due to an outburst of the late glacial Lake Agassiz into the Arctic Ocean (Tarasov and Peltier, 2005; Murton et al., 2010). Its effect on the Southern Hemisphere still remains somewhat elusive. Our goal is to simulate the Last Glacial Termination using an Earth system model of intermediate complexity, LOVECLIM, and to address the following questions: What forcings are needed to reproduce the dominant climate and biogeochemical features of the Last Glacial Termination? What are the global impacts of H1, MWP-1A and the YD? Are climate data consistent with the notion of a Southern Hemispheric origin of MWP-1A? Is it justified to view the North Atlantic as the main driver of millennial-scale variability during the Last Glacial Termination? Is the relative timing of events in paleo-proxy data from different regions consistent with the relative timing in the model? A suite of transient climate modeling experiments is conducted for the period 18e11 ka B.P. using time-varying radiative forcing due to orbitally-induced solar radiation changes and greenhouse gases. The effects of waning glacial ice sheets on topography and albedo are captured by updating the atmospheric boundary conditions continuously with estimates of the icesheet evolution and albedo. Millennial-scale variability is produced by varying the strength of the deep water formation in the North Atlantic and in the Southern Ocean through the input of idealized freshwater pulses. The paper is organized as follows: in section 2 we provide a description of the experimental setup and the methods employed to analyze the paleo-proxy data. Section 3 is devoted to an extensive comparison between model simulation and paleo-climate proxies covering the period 18e11 ka B.P. The paper concludes with a general discussion and a summary of the main results. 2. Model and experimental setup 2.1. The Earth system model LOVECLIM The model used in this study is the model of intermediate complexity LOVECLIM (Driesschaert et al., 2007; Goosse et al., 2007)1. Our experiments were conducted with version 1.1. of this model. The atmospheric component of the coupled model LOVECLIM is ECBilt (Opsteegh et al., 1998), a spectral T21, three-level model, based on quasi-geostrophic equations extended by 1 see also (Goosse et al., 2010a) for the most recent LOVECLIM version 1.2. estimates of the neglected ageostrophic terms (Lim et al., 1991) in order to close the equations at the equator. The coupling between dynamics and thermodynamics is done via a linear balance equation. The sea-ice-ocean component of LOVECLIM, CLIO (Campin and Goosse, 1999; Goosse et al., 1999; Goosse and Fichefet, 1999) consists of a free-surface primitive equation model with 3 3 resolution, coupled to a thermodynamic-dynamic sea-ice model. Coupling between atmosphere and ocean is done via the exchange of freshwater, momentum and heat fluxes. The terrestrial vegetation module of LOVECLIM, VECODE, (Brovkin et al., 1997) simulates the dynamical vegetation changes and the terrestrial carbon cycle in response to climatic conditions. LOCH is a 3-dimensional global model of the oceanic carbon cycle (Mouchet and Francois, 1996; Menviel et al., 2008b; Goosse et al., 2010a). The prognostic state variables considered in the model are dissolved inorganic carbon (DIC), total alkalinity, phosphates (PO34), organic products, oxygen and silica. LOCH is fully coupled to CLIO, with the same time step. In addition to their biogeochemical transformations tracers in LOCH experience the circulation field and diffusion predicted by CLIO. LOCH computes the export production from the fate of a phytoplankton pool in the euphotic zone (0e120 m). The phytoplankton growth depends on the availability of nutrients (PO34) and light, with a weak temperature dependence. A grazing process together with natural mortality limit the primary producers biomass and provide the source term for the organic matter sinking to depth. The other processes described in the model include remineralization, carbonate precipitation and dissolution as well as opal production. LOCH does not include sedimentary processes but nevertheless takes into account carbonate compensation mechanisms. The atmospheric CO2 content is updated for each ocean time step. For our experiments, however, the atmospheric CO2 was prescribed. The coupled model has been used extensively to study different aspects of Late Quaternary climate (Timm and Timmermann, 2007; Menviel et al., 2008b, 2008a; Timm et al., 2008, 2010; Timmermann et al., 2009a). 2.2. Experimental setup In this study the goal is to simulate the sequence of millennialscale events superimposed on the slowly-varying orbital-scale background conditions during the last deglaciation. For this we perform two transient experiments starting from the background conditions of the Last Glacial Maximum (LGM) (Timm and Timmermann, 2007; Menviel et al., 2008b). The model is then forced for the period 21 ka to 10 ka B.P. with the time-varying evolution of solar insolation (Berger, 1978), icesheet topography (updated every 100 years) (Peltier, 1994), high latitude albedo and atmospheric CO2. The changes in atmospheric CO2 are obtained from the EPICA Dome C ice core, Antarctica (Monnin et al., 2001) on the EDC3 timescale (Lüthi et al., 2008). Other greenhouse gases (CH4 and N2O) are kept fixed at LGM values. In experiment DGNS, a series of freshwater pulses is applied in both the North Atlantic and the Southern Ocean in order to reproduce millennial-scale events (Fig. 1d). The timing and amplitude of the freshwater pulses were set so as to provide a good match with the 231Pa/230Th data from the North Atlantic (McManus et al., 2004) and temperature variations in Greenland (Alley, 2000) on the GISP2 timescale, which is within dating uncertainties of the GICC05 timescale (Rasmussen et al., 2006). Even though a significant amount of freshwater was probably discharged into the Gulf of Mexico during the last deglaciation (Aharon, 2003; Flower et al., 2004; Tarasov and Peltier, 2006; Sionneau et al., 2010), with potential implications for the AMOC (Roche et al., 2007), we do not add freshwater into this region. Given the uncertainties associated Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 1159 Fig. 3. Map showing the locations of the paleo-proxy records used to compare model results. Dark blue stars represent paleo-proxy records of oceanic circulation (McManus et al., 2004; Piotrowski et al., 2004; Gherardi et al., 2005; Okazaki et al., 2010; Thornalley et al., 2011). Light blue stars represent terrestrial and marine temperature records (Petit et al., 1999; Alley, 2000; Bard et al., 2000; Lea et al., 2003; Pahnke et al., 2003; Nielsen, 2004; Kaiser et al., 2005; Samson et al., 2005; Calvo et al., 2007; Heiri et al., 2007; Jouzel et al., 2007; Martrat et al., 2007; Barker et al., 2009; Stenni et al., 2011). Orange stars show the location of paleo-precipitation records (Thompson et al.,1995; Peterson et al., 2000; Baker et al., 2001a; 2001b; Wang et al., 2001; Holmgren et al., 2003; Ivanochko et al., 2005; Wang et al., 2007; Weldeab et al., 2007). Red stars represent biogeochemical paleo-proxy records (Pride et al.,1999; Emmer and Thunell, 2000; Ivanochko and Pedersen, 2004; Sachs and Anderson, 2005; Anderson et al., 2009). (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) with the detailed freshwater routing during the Last Glacial Termination we choose a more idealized approach. To mimic H1, Older Dryas and Younger Dryas in our model, freshwater is released into a broad region in the North Atlantic centered between 55 We10 W, 50 Ne65 N and the Arctic Ocean, respectively. The amplitude of the millennial-scale freshwater forcing amounts to 0.2 Sv during the period 18 ka to 17.4 ka B.P. and 0.25 Sv between 17.4 ka B.P and 15.6 ka B.P. To specifically represent the Older Dryas we apply a freshwater pulse in the North Atlantic for the period 14.4 ka B.P. to 14 ka B.P. with a maximum input of 0.25 Sv between 14.2 and 14 ka B.P. The exact shape of the freshwater pulse is depicted in Fig. 1d. A background freshwater flux was also added in the North Atlantic between 14 and 13 ka B.P. to represent the melting northern hemisphere ice sheets. To mimic the Younger Dryas, 0.25 Sv is applied in the Arctic Ocean (175 We95 W, 67 Ne83 N) between 13 and 12.2 ka B.P. Finally, to represent the ACR, an additional meltwater pulse is prescribed for the area enclosing the Ross and Weddell Seas (163 E11 E, 70 S-80 S). The magnitude of this pulse is 0.15 Sv for the period 14.4 ka to 13.4 ka B.P., after which the pulse linearly decreases to 0 over 1000 years (until 12.4 ka B.P.). To better evaluate the impact of Southern hemispheric meltwater pulses, another experiment similar to DGNS (DGN) was conducted. In DGN, no meltwater pulse is added in the Southern Ocean between 14.4 ka and 12.4 ka B.P2. 2 Unlike some of the experiments described in Timm and Timmermann (2007), experiments DGNS and DGN do not employ any orbital acceleration (Lorenz and Lohmann, 2004). The results of experiment DGNS are compared to a variety of paleo-proxy records (Fig. 3 and Table 1), which represent variations in oceanic circulation, temperature, precipitation and biogeochemical variables in different regions of the world. To obtain a more representative comparison of model outputs with paleoproxy records, when possible, composite time series were generated. These composite time series were produced by first interpolating each paleo-proxy data onto an equidistant grid in time using bicubic splines and then averaging the different paleoproxy records. In the model-paleo data comparison conducted here, we allowed for some dating uncertainties in the proxy records. Optimal match between the local proxy time series and the simulated physical variable was sometimes achieved by shifting the proxy time series by a few hundred years (black arrows in time series figures, Table 2). The shift that we had to apply to obtain a good match between model solution and paleo data was smaller than the respective published absolute dating uncertainties. The time series shifting depends on the timing of the freshwater forcing sequences prescribed in the transient simulations. 3. The deglacial history in the northern hemisphere 3.1. Heinrich event 1 During H1, and as a result of the chosen freshwater forcing the simulated AMOC collapses (Fig. 1a), in agreement with the 231Pa/230Th data from the Bermuda rise core OCE326-GGC5 (McManus et al., 2004). Fig. 1a also displays the 231Pa/230Th data from the Iberian Author's personal copy 1160 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Table 1 paleo-proxy records used to compare the model outputs of this study. Location Core Lat Long Elev (m) Proxy Reference Greenland Greenland South Iceland rise South Iceland rise South Iceland rise The Netherlands Iberian margin Iberian margin Iberian margin Iberian margin Iberian margin North Atlantic Cariaco Basin Cariaco Basin Arabian Sea Gulf of Guinea China North Pacific California basin California basin California basin Peruvian Altiplano Bolivian Altiplano Bolivian Altiplano Brazil South Africa South Australia New Zealand Chilean margin Southeast Atlantic Ocean Southeast Atlantic Ocean New Zealand New Zealand Southern Ocean Southern Ocean Antarctica Antarctica Antarctica Antarctica GISP2 GISP2 RAPiD-10-1P RAPiD-15-4P RAPiD-17-5P Hijkermeer SU81-18 SU81-18 MD95-2042 MD01-2444 ODP-161-977A OCE326-GGC5 ODP 1002 PL07-39PC 905 MD03-2707 Hulu Cave 72.6 N 72.6 N 62.6 N 62.2 N 61.3 N 52.5 N 37.8 N 37.8 N 37.5 N 37.8 N 36.1 N 33.42 N 10.7 N 10.7 N 10.46 N 2.5 N 32.5 N 32e55 N 27 N 34.17 N 34.17 N 9S 16e17.5 S 20.14 S 27.2 S 24 S 36.44 S 36.55 S 41 S 41.07 S 41.1 S 45.5 S 45.5 S 53.2 S 53.2 S 72.8 S 75.1 S 75.1 S 78.27 S 38.5 W 38.5 W 17.4 W 17.1 W 19.3 W 6.3 E 10.1 W 10.1 W 10.1 W 10.7 W 1.6 W 57.35 W 65.2 W 65 W 51.57 E 9.4 E 119.2 E 133e168 E 112 W 120.02 W 120.02 W 77.5 W 68.5e70 W 67.3 W 49.2 W 29.2 E 136.33 E 177.26 E 74.5 W 9E 7.8 E 174.9 E 174.9 E 5.1 E 5.1 E 159.2 E 123.4 E 123.4 E 106.5 E 3207 3207 1237 2133 2303 14 3135 3135 3146 3140 1924 4550 893 790 1586 1295 100 Ice core d18O, borehole Ice core CH4 Ventilation age Ventilation age Ventilation age Chironomids UK’37 231 Pa/230Th UK’37 UK’37 UK’37 231 Pa/230Th 550 nm reflectance Foram Mg/Ca d15N Foram Ba/Ca Speleothem d18O Ventilation age d15N d15N Mo/Al Ice core d18O % freshwater diatom g nat. radiation Speleothem d18O Speleothem d18O UK’37 Modern analog UK’37 Nd Foram. Mg/Ca Alkenone content UK’37 Modern analog Opal flux Ice core d18O Ice core dD CO2 Ice core dD (Alley, 2000) (Brook et al., 1996) (Thornalley et al., 2011) (Thornalley et al., 2011) (Thornalley et al., 2011) (Heiri et al., 2007) (Bard et al., 2000) (Gherardi et al., 2005) (Bard, 2002) (Martrat et al., 2007) (Martrat et al., 2004) (McManus et al., 2004) (Peterson et al., 2000) (Lea et al., 2003) (Ivanochko et al., 2005) (Weldeab et al., 2007) (Wang et al., 2001) (Okazaki et al., 2010) (Pride et al., 1999) (Emmer and Thunell, 2000) (Ivanochko and Pedersen, 2004) (Thompson et al., 1995) (Baker et al., 2001a) (Baker et al., 2001b) (Wang et al., 2007) (Holmgren et al., 2003) (Calvo et al., 2007) (Samson et al., 2005) (Kaiser et al., 2005) (Piotrowski et al., 2004) (Barker et al., 2009) (Sachs and Anderson, 2005) (Pahnke et al., 2003) (Nielsen, 2004) (Anderson et al., 2009) (Stenni et al., 2011) (Jouzel et al., 2007) (Monnin et al., 2001) (Petit et al., 1999) JPC-56 ODP893-A ODP893-A Huascaran Lake Titicaca Salar de Uyuni Botuverá Cave Cold Air Cave MD03-2611 H214 ODP 1233 RC11-83 TN057-21 MD97-2120 MD97-2120 TN057-13PC TN057-13PC TALDICE EPICA Dome C EPICA Dome C Vostok margin core SU81-18 (Gherardi et al., 2005) and the ventilation age as inferred from South Iceland rise sediment cores (Thornalley et al., 2011). Even if all the records show a weakened AMOC at some time during H1, there are some substantial differences among them, which will further be discussed in section 5. The AMOC weakening reduces the poleward Atlantic heat transport at 40 N by 0.5 PW (not shown), which leads to an enhanced Northern Hemispheric cooling relative to the LGM (Figs. 2 and 4 left), in accordance with temperature paleoproxy data from Greenland (Alley, 2000) (Fig. 2a), the North Atlantic (Bard et al., 2000; Calvo et al., 2001; Waelbroeck et al., 2001; Bard, 2002; Martrat et al., 2004, 2007) (Fig. 2c), the tropical North Atlantic (Lea et al., 2003; Niedermeyer et al., 2009) (Fig. 2d), the Mediterranean Sea (Cacho et al., 1999, 2001), Europe (Ivy-Ochs et al., 2006), Asia (Wang et al., 2002; Evans et al., 2003) and Africa (Powers et al., 2005). 818 576.5 576.5 6048 3810 3653 250 e 2420 2045 838 4718 4981 1210 1210 2848 2848 2315 3240 3240 3500 The enhanced meridional temperature gradient in the North Atlantic induces a strengthening of the northeasterly trades and a southward shift of the Intertropical Convergence Zone (ITCZ) (Timmermann et al., 2005) (Fig. 4, middle). In agreement with paleo-proxies, drier conditions prevailed during H1 in Europe (Peyron et al., 2005), the northern part of Africa (Peck et al., 2004; Williams et al., 2006; Lamb et al., 2007; Niedermeyer et al., 2009), the Middle East (Bar-Matthews et al., 2003; Vaks et al., 2003; Shakun et al., 2007; Fleitmann et al., 2009) and the northern part of South America. As shown in Fig. 5a, we simulate dry conditions in the Cariaco basin as soon as the AMOC weakens. The reflectance record from the Cariaco sediments (Peterson et al., 2000) indicates relatively dry conditions between 18 and 16.2 ka B.P., however the drying becomes more intense at 16.2 ka B.P. The reason behind this feature remains unexplained. In the Gulf of Guinea, the Table 2 Temporal shift applied to paleo-proxy records used to compare the model outputs of this study. A positive (negative) shift means that the paleo data had to be shifted forward (backward) in time to match the model simulation. Location Cores Proxy Reference Temporal shift applied (yrs) North Atlantic North Pacific Bolivian Altiplano Bolivian Altiplano Southeast Atlantic Ocean New Zealand Southern Ocean Antarctica Antarctica OCE326-GGC5 231 (McManus et al., 2004) (Okazaki et al., 2010) (Baker et al., 2001a) (Baker et al., 2001b) (Piotrowski et al., 2004) (Sachs and Anderson, 2005) (Anderson et al., 2009) (Jouzel et al., 2007) (Petit et al., 1999) () 200 () 600 400 400 500 () 300 () 200 () 600 () 600 Lake Titicaca Salar de Uyuni RC11-83 MD97-2120 TN057-13PC EPICA Dome C Vostok Pa/230Th Ventilation age % freshwater diatom g nat. radiation Nd Alkenone content Opal flux Ice core dD Ice core dD Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 1161 Fig. 4. (left) Annual SST anomalies ( C, shaded) and current anomalies (m/s, vectors) averaged over 0e200 m for H1eLGM conditions (16e18 ka B.P.), (middle) Annual precipitation anomalies (cm/yr, shaded) and annual wind stress anomalies (Pa, vector), (right) Annual export production anomalies (gC/m2/yr) for H1eLGM and overlaid 0.1 m annual sea-ice contour for H1. Fig. 5. (a) Time series of JJA precipitation anomalies (cm/yr) averaged over the Cariaco basin (55 W-52 W, 10 N-14 N) for experiment DGNS (green) compared with the reflectance (%, black) measured in a marine sediment core from the Cariaco basin (ODP 1002) (Peterson et al., 2000). Lower reflectance has been associated with increased marine productivity due to greater riverine input. (b) Time series of JJA precipitation anomalies (cm/yr) averaged over western Equatorial Africa (15 W-15 E, 4 N-20 N) for experiment DGNS (green) compared with SSS reconstruction (black) from the Gulf of Guinea core MD03-2707 (Weldeab et al., 2007). (c) Time series of JJA/DJF precipitation averaged over China (114 E124 E, 28 N-35 N) for experiment DGNS (green) compared to d18O (permil, black) as recorded in stalagmites of the Hulu cave (China) (Wang et al., 2001). (d) Time series of JJA precipitation anomalies over the Arabian Sea (45 E65 E, 5 N-15 N) for experiment DGNS (green) compared to d15N (permil, black) as recorded in marine sediment core 905 from the Arabian Sea (Ivanochko et al., 2005). Anomalies are with respect to pre-industrial time. Time series are 51 year running means for the model outputs. (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) Author's personal copy 1162 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 southward ITCZ shift leads to drier conditions in agreement with Ba/Ca data indicating greater surface salinity during H1 in that area (Weldeab et al., 2007). The AMOC shut down also leads to a reduced ratio of summer/winter precipitation over China in agreement with d18O records from Chinese caves (Wang et al., 2001; Dykoski et al., 2005) (Fig. 5c). Furthermore, our model solution supports the notion of a reduced summer monsoon over the Arabian Sea (Ivanochko et al., 2005) (Fig. 5d). As a result of the simulated southward shift of the ITCZ (Fig. 4, middle), the reduced Atlantic-Pacific freshwater export (not shown) and the reduced Asian summer monsoon (Fig. 5c), North Pacific salinity increases at the onset of H1, as described in Okazaki et al. (2010). North Pacific waters become dense enough to generate North Pacific Deep Water (NPDW) formation in the Bering Sea and the Gulf of Alaska. Subsequently a deep Pacific Meridional Overturning Cell (PMOC) establishes. Maximum meridional transports in this cell attain values of up to 20 Sv. Southward spreading of well-ventilated NPDW is in accordance with a compilation of radiocarbon data obtained from marine sediment cores that suggest a drop of eastern North Pacific ventilation ages by w1000 years (Fig. 1b) Okazaki et al. (2010). As can be seen in Fig. 4 left, the formation of NPDW leads to a reorganization of surface currents in the Indo-Pacific: the Indonesian Throughflow transport decreases (w-50%) and the warm and saline water of the subtropical Pacific are diverted into the northern North Pacific, supplying the sinking branch of the PMOC (Chikamoto et al., 2010; Menviel et al., 2010b; Okazaki et al., 2010). A slowdown of the Indonesian Throughflow during Heinrich events has also been inferred from a surface and upper thermocline temperature record in the Timor Sea (Zuraida et al., 2009), providing support to our analysis. The formation of NPDW leads to an increase in poleward heat transport in the North Pacific of 0.5 PW at 40 N. In experiment DGNS, the enhanced transport of warm equatorial waters to the North Pacific combined with greater insolation and atmospheric CO2 lead to a slight warming of the ocean surface in the North Pacific during H1 (16e18 ka B.P.) (Fig. 4, left). In agreement with SST paleo-proxies, a warming is observed in the northeastern Pacific where warm waters converge (Sarnthein et al., 2006), while a relatively small area around Japan cools (Harada et al., 2006, 2010) (Fig. 4, left). At the beginning of H1, the ITCZ shifts southward in the Pacific, however as NPDW formation enhances, the ITCZ shifts back to its initial position in the Pacific basin (Fig. 4, middle). In the North Pacific, the formation of NPDW leads to an increase in the dissolved oxygen content (by up to 3 ml/L) in the upper 2500 m, in agreement with benthic foraminiferal assemblages in marine sediment cores from the Northwest Pacific (Shibahara et al., 2007). The well-oxygenated waters flow equatorwards from the North Pacific. The main branch flows as a deep western boundary current but well-ventilated waters are also advected along the California basin. The dissolved oxygen content along the coast of California increases by about 50% down to about 2000 m depth. This is in qualitative agreement with redox sensitive metal content, d15N and total organic carbon records from marine sediment core ODP 893A (Kennett and Ingram, 1995; Emmer and Thunell, 2000; Ivanochko and Pedersen, 2004) as well as with d15N record from marine sediment core JPC-56 (Pride et al., 1999) (Fig. 1c). Despite the fact that changes in d15N can have numerous causes (Somes et al., 2010), the redox sensitive metal content and organic carbon records from the California basin support our hypothesis of greater ventilation during H1. Due to the colder conditions and increased stratification, simulated marine export production is low in an area spanning the North Atlantic, the Atlantic coast of North and Central America as well as the Caribbean Sea, in agreement with paleo-proxy reconstructions (Peterson et al., 2000; Vink et al., 2001) (Fig. 4, right). Due to the stronger upwelling, marine export production significantly increases (þ25%) in the Equatorial Atlantic (Northwest African coast and Brazilian coast) in agreement with paleo-proxy records from the Brazilian coast (Jennerjahn et al., 2004). 3.2. Bølling warming, Older Dryas and Intra-Allerød cold period At the end of H1 (w15.2 ka B.P.), the simulated AMOC recovers quickly regaining a strength of about 27 Sv around 14.6 ka B.P (Fig. 1a). As can be seen in Figs. 2 and 6 (left), the temperatures in Greenland and in the North Atlantic increase abruptly during the recovery phase of the AMOC (15e14.6 ka B.P.), until reaching a maximum during the Bølling Warm Period (BWP). As already discussed by Liu et al. (2009), the rapid transition to the BWP can be explained by a combination of a rapid resumption of the AMOC following H1, an overshooting effect of the AMOC, as well as by the contemporaneous increase in atmospheric CO2 concentrations (Köhler et al., 2010). The amplitude of the simulated increase in spring temperature over Greenland between 15.2 ka B.P. and 14.5 ka B.P. amounts to about 12 C, which is similar to the 13 C temperature difference estimated from GISP2 ice core for the same time period (Alley, 2000) (Fig. 2a). Off the Iberian margin the simulated SST and alkenone-based SST (Bard et al., 2000; Bard, 2002; Martrat et al., 2004, 2007) both increase between H1 and BWP by about 3e4 C (Fig. 2c). In the Cariaco basin however, the simulated 1.5 C SST increase during the winter season is only half of that reconstructed from Mg/Ca data in core PL07-39PC (Lea et al., 2003) (Fig. 2d). The resumption of the AMOC leads to a reorganization of surface ocean currents and to the decline in the formation of NPDW. Fig. 6. (left) Annual SST anomalies ( C, shaded) and current anomalies (m/s, vectors) averaged over 0e200 m for Bølling warmingeH1 conditions (14.6e16 ka B.P.), (middle) Annual precipitation anomalies (cm/yr, shaded) and annual wind stress anomalies (Pa, vector), (right) Annual export production anomalies (gC/m2/yr) for BWPeH1 and overlaid 0.1 m annual sea-ice contour for BWP. Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Such an intense and rapid warming in the North Atlantic at about 15 ka B.P. is likely to have consequences on the massive ice sheets that are still present over North America and northern Europe. During this period, melting of the Laurentide and Fennoscandian ice sheets accelerated (Peltier, 1994; Gregoire, 2010; Thornalley et al., 2010) thus adding freshwater into the North Atlantic. In response to this freshwater perturbation the simulated AMOC weakens from 27 Sv at the peak of the Bølling warming down to about 12 Sv during the Older Dryas (w14 ka B.P.) and levels off at about 18 Sv during the Allerød (between 13.8 and 13 ka B.P.) (Fig. 1a). These variations in simulated AMOC strength are in overall agreement with the ventilation age history recorded in the South Iceland rise (Thornalley et al., 2011). The significant freshwaterinduced reduction in AMOC strength during the Older Dryas is not observed in the 231Pa/230Th records (McManus et al., 2004; Gherardi et al., 2005). Higher resolution 231Pa/230Th records for the period 14.2 and 13.5 ka B.P would be needed to test this assertion. Due to the intermittent AMOC weakening at 14 ka B.P., modeled air temperatures over Greenland drop by about 6 C, summer air temperature decreases by 1.8 C in the Netherlands, SST decrease by about 1.2 C off the Iberian margin and by about 0.7 C in the Cariaco basin (Figs. 2 and 7, left). This simulated cooling is in very good agreement with temperature estimates from Greenland ice core (Alley, 2000) and chironomid-inferred July temperatures from the Netherlands (Heiri et al., 2007). However the cooling in the Cariaco basin SST reconstruction (Lea et al., 2003) is lower than in the model results and no significant cooling is observed in marine sediment cores off the Iberian margin (Bard et al., 2000; Bard, 2002; Martrat et al., 2004, 2007). Over northern North America, the waning of the icesheet and the resulting changes in height induce a local surface warming due to the lapse rate effect, even during the Older Dryas. At about 13.8 ka B.P. the simulated temperatures in the North and Equatorial Atlantic recover to levels equivalent to the Bølling warming, in agreement with SST proxies from that region. Greenland ice cores also record another millennial-scale cold event, known as the Intra-Allerød cold period (IACP, 13.5e13 ka B.P., Fig. 2a) (Stuiver et al., 1995), which is not reproduced in our transient modeling experiments. This millennial-scale cold event was also observed in the Norwegian Sea (Lehman and Keigwin, 1992), on the South Iceland Rise (Thornalley et al., 2010) as well as in terrestrial records from Northern Europe (von Grafenstein et al., 1999; Heiri et al., 2007) (Fig. 2b) and North America (Yu and Eicher, 2001). In addition ventilation age anomalies indicate decreased ventilation south of Iceland during that period (Thornalley et al., 2011). However no significant decrease in North Atlantic SST was reconstructed for that time period according to 1163 higher resolution SST proxies (Cacho et al., 1999, 2001; Bard et al., 2000; Calvo et al., 2001; Bard, 2002; Lea et al., 2003; Martrat et al., 2004, 2007). Given these discrepancies between the proxybased spatial extent of the IACP and the model simulation it thus remains unresolved whether large-scale freshwater forcing was the sole cause for these variations. During the Bølling warm period, the ITCZ swings back to the north, leading to wetter conditions over Europe (Peyron et al., 2005), the northern part of South America (Peterson et al., 2000, North Africa (Peck et al., 2004; Williams et al., 2006; Weldeab et al., 2007; Niedermeyer et al., 2009), the middle East (Bar-Matthews et al., 2003; Vaks et al., 2003; Ivanochko et al., 2005; Shakun et al., 2007; Fleitmann et al., 2009), China (Wang et al., 2001; Dykoski et al., 2005) and Tibet (Kramer et al., 2010) (Fig. 6, middle and 5). Due to the intense warming and retreat of the sea-ice edge in the North Atlantic the marine export production more than doubles in this region (Fig. 6, right). It also increases considerably in the Caribbean Sea (Peterson et al., 2000; Vink et al., 2001). Marine export production however decreases in the North Pacific (5%) due to the reduction of the NPDW formation and in the Equatorial Atlantic (25%) due to the reduction in the strength of the northeasterly trades (Jennerjahn et al., 2004). 3.3. Younger Dryas In two recent studies (Tarasov and Peltier, 2005; Murton et al., 2010) it was suggested that a glacial meltwater pulse into the Arctic Ocean caused the YD cold period. The shut down of the AMOC at 13 ka B.P. (Fig. 1a) leads to a cooling of about 10 C over Greenland, 2 C in the Netherlands, 3 C off the Iberian margin and 1 C in the Cariaco basin (Fig. 2). The simulated relative strength of the overturning in the North Atlantic is weaker than relative changes recorded by the 231Pa/230Th record of the Bermuda rise and Iberian margin sediment cores (McManus et al., 2004; Gherardi et al., 2005). Ventilation age anomalies from the South Iceland rise indicate an interruption of the ventilation in the second part of the YD (Thornalley et al., 2011). Over Greenland, the amplitude of the simulated cooling during the YD is similar to the one estimated from the GISP2 borehole (Cuffey and Clow, 1997; Alley, 2000), but the simulated temperature at 13 ka B.P. (before the YD) is about 5 C warmer than the one estimated in GISP2 (Fig. 2a). This 5 C offset from the observations lasts until the end of the experiment DGNS. Part of this temperature discrepancy between the model and the GISP2 record could be explained by an elevation change at GISP2. Using the relationship between height and d18O (Vinther et al., 2009), a 200 m increase in the GISP2 icesheet would lead to a 1.2 Fig. 7. (left) Annual SST anomalies ( C, shaded) and current anomalies (m/s, vectors) averaged over 0e200 m for ACReBølling warming conditions (14e14.6 ka B.P.), (middle) Annual precipitation anomalies (cm/yr, shaded) and annual wind stress anomalies (Pa, vector), (right) Annual export production anomalies (gC/m2/yr) for ACReBWP and overlaid 0.1 m annual sea-ice contour for ACR. Author's personal copy 1164 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 permil decrease in d18O. Assuming a slope of 2.8 C per 1 permil for the deglaciation, the recorded temperatures would be w3.3 C lower. On top of that the adiabatic lapse rate (0.98 C/100 m) has to be taken into account. A 5 C temperature offset could thus be explained by a 180e200 m increase in the icesheet height at GISP2. The simulated cooling off the Iberian margin is overestimated compared to the estimated SST anomaly (Bard et al., 2000; Bard, 2002; Martrat et al., 2004, 2007), while it is in agreement with a reconstructed 3 C cooling in the Western Mediterranean Sea (Cacho et al., 1999, 2001). In the Cariaco basin, however, the simulated cooling is underestimated. The abrupt YD cooling is also observed in paleorecords from continental Europe (von Grafenstein et al., 1999; Heiri et al., 2005; Heiri and Millet, 2005; Peyron et al., 2005; Genty et al., 2006). Similar to the H1 climate response, the anomalous meridional SST gradient during the YD in the tropical Atlantic leads to a southward shift of the ITCZ. In agreement with paleo-proxies, drier conditions are simulated over the northern part of South America (Maslin and Burns, 2000; Peterson et al., 2000; Haug et al., 2001), Europe (Goslar et al., 1999; Kerschner et al., 2000; Peyron et al., 2005), Northern Africa (Peck et al., 2004; Williams et al., 2006; Lamb et al., 2007; Weldeab et al., 2007; Niedermeyer et al., 2009) and the Middle East (Bar-Matthews et al., 2003; Vaks et al., 2003; Ivanochko et al., 2005; Shakun et al., 2007; Fleitmann et al., 2009). Fig. 8. (a) Time series of the maximum overturning strength in the bottom cell of the Southern Ocean (Sv) for experiment DGNS (green) and DGN (cyan) compared to eNd (black) as recorded in marine sediment core RC11-83 (Piotrowski et al., 2004). Lower eNd values indicate the predominance of NADW at the site of core RC11-83 while higher values indicate the predominance of AABW. The time series of (Piotrowski et al., 2004) has been shifted by 500 years forward. (b) Time series of austral spring (SON) air temperature anomalies ( C) averaged over 70 S-90 S for experiments DGNS (green) and DGN (cyan) compared to temperature reconstructions ( C) from EPICA Dome C ice core (black stars) on the EDC(3) time scale (Jouzel et al., 2007) and from Vostok ice core (black thin line) (Petit et al., 1999) as well as compared to the d18O record from Talos Dome ice core (gray) (Stenni et al., 2011). The time axis of the EPICA Dome C and Vostok temperature reconstructions were shifted backward by 600 years to better fit the model results and the northern hemispheric paleoproxies. Simulated austral spring temperatures were used as suggested by the study of (Timmermann et al., 2009b). (c) Time series of annual SST anomalies ( C) averaged over the Pacific and Atlantic part of the Southern Ocean (170 E10 E, 40 S-55 S) for experiments DGNS (green) and DGN (cyan) compared to a composite of SST reconstructions ( C, black) from marine sediment cores MD97-2120 (Pahnke et al., 2003), ODP1233 (Kaiser et al., 2005), TN057-13PC4 (Anderson et al., 2009; Nielsen, 2004) and TN057-21 (Barker et al., 2009). (d) Time series of annual SST anomalies ( C) averaged over the Southwest Pacific (130 E180 E, 32 S-42 S) for experiment DGNS (green) compared to a composite of SST reconstructions ( C, black) from marine sediment cores MD03-2611 (Calvo et al., 2007) and H214 (Samson et al., 2005). Anomalies are with respect to pre-industrial time. Time series are 21 year running means for the model outputs. (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Reduced summer precipitation is also simulated over China (Wang et al., 2001; Dykoski et al., 2005) (Fig. 5). The weakening of the AMOC during the YD leads to the formation of NPDW. As a result, younger ventilation ages are recorded in marine sediment cores from the North Pacific (Fig. 1b) (Okazaki et al., 2010) and dissolved oxygen content increases along the coast of California (Kennett and Ingram, 1995; Pride et al., 1999; Emmer and Thunell, 2000; Ivanochko and Pedersen, 2004) (Fig. 1c). 4. The deglaciation history in the Southern Hemisphere 4.1. Heinrich event 1 As a result of the simulated AMOC shut down during H1, the AABW cell strengthens, attaining values of up to 12 Sv between 17.6 1165 and 14.8 ka B.P. (Fig. 8a). This intensification of AABW formation is in qualitative agreement with Neodynium data from the South Atlantic core RC11-83 (Piotrowski et al., 2004). Lower eNd values indicate the predominance of NADW at the site of core RC11-83 while higher values indicate the predominance of AABW. Accompanying the NADW decrease and the AABW increase, the model simulates an intensification of the poleward heat transport at 30 S by about 0.6PW. As depicted in Fig. 8, temperatures at high southern latitudes increase almost linearly from 17.6 to 14.8 ka B.P. This warming is a combination of different factors: rising atmospheric CO2 concentrations (Timmermann et al., 2009b), increase of austral spring mean insolation changes affecting the sea-ice extent around Antarctica, and the bipolar seesaw effect leading to a warming of the Southern Hemisphere. Over Antarctica, the simulated air temperature during austral spring follows quite closely the temperature derived from EPICA Dome C ice core dD Fig. 9. (a) Time series of annual precipitation anomalies (cm/yr) averaged over Bolivia (71 W-62 W, 15 S-25 S) for experiment DGNS (green) compared to % of freshwater diatoms (black x) in Lake Titicaca (Bolivia) (Baker et al., 2001b) and to a natural gamma ray profile (c.p.s., gray circles) from the Salar de Uyuni (Bolivia) (Baker et al., 2001a). The time axis of the Lake Titicaca and the Salar de Uyuni records were shifted forward by 400 years. (b) Time series of DJF precipitation anomalies (cm/yr) averaged over Peru (85 W-70 W, 5 S 18 15 S) for experiment DGNS (green) compared to d O record (permil, black) from Huascaran ice core (Peru) (Thompson et al., 1995). (c) Time series of DJF/JJA precipitation averaged over Brazil (60 W-44 W, 20 S-32 S) for experiment DGNS (green) compared to d18O record (permil, black) from stalagmites of Botuverá cave (Brazil) (Wang et al., 2007). (d) Time series of JJA/DJF precipitation anomalies averaged over South Africa (25 E35 E, 20 S-30 S) for experiment DGNS (green) compared to d18O record (permil, black) from stalagmites of the Makapansgat Valley (South Africa) (Holmgren et al., 2003). Anomalies are with respect to pre-industrial time. Time series are 51 year running means for the model outputs. (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 2 Opal production (mol/m /yr) a 17 16 4.2. Antarctic Cold Reversal Simulated surface temperatures at high southern latitudes reach a local maximum around 14.8 ka B.P (Fig. 8bec). The rapid strengthening of the AMOC towards the end of H1 leads to a weakening of AABW formation starting 14.8 ka B.P. (Fig. 8a). The warmer conditions in the Southern Ocean as well as higher sea level around 14.6 ka B.P. (Bindschadler et al., 2003; Peltier, 2005), may have helped to destabilize the Antarctic ice shelves and ice sheets (Flückiger et al., 2006), thus adding freshwater to the Southern Ocean. Proxy-evidence for a meltwater pulse originating from Antarctica at around 14.6 ka B.P. was found in several marine sediment cores of the Southern Ocean and was discussed in section 1. Adding freshwater to the Southern Ocean in our simulation DGNS further weakens the formation of AABW and slows down the Southern Ocean overturning cell to 5 Sv until about 13 ka B P, in qualitative agreement with Neodynium data from the South Atlantic (Piotrowski et al., 2004) (Fig. 8a). In contrast, when no freshwater is added to the Southern Ocean (experiment DGN), the AABW stays moderately strong (w12 Sv). The latter scenario does not agree well with the Neodymium data. Reduced AABW formation in experiment DGNS leads to a decrease in southward heat transport at 30 S of about 0.2 PW. As a result, temperatures decrease at high and mid Southern latitudes (Fig. 7, left). Austral spring temperature averaged over Antarctica are about 0.8 C lower (Fig. 8b), which compares reasonably well with the cooling recorded in the dD data from EPICA Dome C (Jouzel 14 13 12 Southern Ocean 0.15 6 ACR 0.1 4 2 H1 0.05 17 36 b 17 16 16 15 14 15 14 13 13 12 12 2 Export production (gC/m /yr) 15 Due to the warmer conditions during H1 and retreating sea-ice in the Southern Ocean, simulated marine export production increases in these areas by about 15% (Fig. 10 a and 4, right), in close correspondence with the data of Anderson et al. (2009) and Sachs and Anderson (2005). Opal flux (g/cm2/kyr) record (Jouzel et al., 2007). Both show an increase of about 4 C between 18 and 15 ka B.P. At that time, the derived temperature from the EPICA Dome C ice core suddenly increases by about 2 C in 200 years (Fig. 8b). This temperature spike is not simulated by our model. As austral spring insolation is an important pacemaker of the glacial termination in the Southern Hemisphere (Timmermann et al., 2009b), simulated austral spring temperatures were used for the comparison with Antarctic records. In the Atlantic and Pacific sectors of the Southern Ocean (Fig. 8c), the simulated annual mean SST increases by 2.7 C between 18 and 15 ka B.P., which is in agreement with the reconstructed SST increase from a composite of marine sediment cores that we obtained by averaging linearly interpolated alkenone SST reconstructions for cores MD97-2120 (Pahnke et al., 2003), ODP1233 (Kaiser et al., 2005), TN057-13PC4 (Nielsen, 2004; Anderson et al., 2009) and TN057-21 (Barker et al., 2009). At the same time period, the simulated SST increases by 1.9 C in the South Pacific (32 S-42 S, Fig. 8d). A composite of SST reconstructions from marine sediment cores MD03-2611 (Calvo et al., 2007) and H214 (Samson et al., 2005) indicates a larger increase of the order of 3 C, but the overall timing matches well with the transient simulation. The cooling of the North Atlantic during H1 in combination with the warming of the Southern Hemisphere lead to a southward shift of the ITCZ and generally wetter conditions over the Southern Hemisphere (Fig. 4, middle), in agreement with paleoproxies from the Bolivian (Baker et al., 2001a, 2001b; Blard et al., 2009) and Peruvian (Thompson et al., 1995) Altiplano (Fig. 9aeb) as well as with d18O records from South African stalagmites (Holmgren et al., 2003) (Fig. 9d). The South American Summer Monsoon is also quite intense during H1, as suggested by the d18O recorded in stalagmites from Botuverá cave (Brazil) (Cruz et al., 2005; Wang et al., 2007) and our modeling results (Fig. 9c). Southern Ocean 600 32 400 28 Alkenone content (ng/g) 1166 200 17 16 15 14 13 12 Time (ka BP) Fig. 10. (a) Time series of opal production (mol/m2/yr) in the Southern Ocean (52 S-64 S) for experiments DGNS (green) and DGN (cyan) compared to the opal flux record (g/cm2/kyr, black) from core TN057-13 (Anderson et al., 2009). The time axis of the opal flux was shifted backward by 200 years. (b) Time series of marine export production (gC/m2/yr) in the Pacific side of the Southern Ocean (150 E190 E, 42 S-49 S) for experiments DGNS (green) and DGN (cyan) compared to the alkenone content record (ng/g, black) from core MD972120 (Sachs and Anderson, 2005). The alkenone record is shown for the period 18 to 10.4 ka B.P. (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 1167 Fig. 11. Austral spring air temperature anomalies (14e14.8 ka B.P, C) over Antarctica and Southern Ocean for (left) experiment DGNS and (right) experiment DGN. et al., 2007) and Vostok ice cores (Petit et al., 1999) and qualitatively with the d18O from Taylor Dome ice core (Steig et al., 1998). While the EPICA Dome C and Vostok ice core records had to be shifted by 600 years, the d18O record from Talos Dome ice core based on the new GICC05 timescale fits well with our simulation without any time shift. Over the Pacific and Atlantic section of the Southern Ocean (40 S-55 S), the simulated cooling of 1.4 C is in agreement with a reconstructed 1.2 C cooling in the Southern ocean SST index derived in this work from various datasets (Pahnke et al., 2003; Nielsen, 2004; Kaiser et al., 2005; Anderson et al., 2009; Barker et al., 2009) (Fig. 8c). Finally in the South Pacific (32 S-42 S), a simulated SST decrease of 0.9 C compares well with a recon structed SST decrease of 1.2 C (Calvo et al., 2007; Samson et al., 2005) (Fig. 8d). Alkenone-based SST estimates from marine sedi ment core GIK 17748-2 (32.45 S,72.02 W) (not shown) also suggest a similar cooling of 1 C during the ACR (Kim et al., 2002). A part of the cooling in the Southern Hemisphere prior to the addition of freshwater is caused by the reduction of AABW formation at 14.8 ka B.P. When no freshwater is added to the Southern Ocean (experiment DGN), the cooling amounts to 0.3 C over Antarctica and 0.6 C in the Southern Ocean. The cooling at high Southern latitudes in experiment DGNS is more pronounced over the ocean than over Antarctica (Fig. 11). In experiment DGN the cooling is reduced significantly compared to DGNS, particularly over Antarctica and the Atlantic and the Eastern Pacific side of the Southern Ocean. During the ACR the temperature decrease in the Southern Hemisphere leads to relatively drier conditions over most of the Southern Hemisphere (Fig. 7, middle). This is in good qualitative agreement with paleo-proxies that document decreasing precipitation over the Bolivian and Peruvian Altiplano (Thompson et al., 1995; Baker et al., 2001a; 2001b) and a reduction of summer rainfall over South Africa (Holmgren et al., 2003) (Fig. 9). Our model also simulates a weakening of the South American Summer Monsoon over Brazil during the ACR in agreement with the d18O record from Botuverá cave (Brazil) (Wang et al., 2007). Due to the cooling at high southern latitudes around w14 ka B.P., the simulated summer sea-ice edge advances to about 53e57 S in agreement with paleoproxies (Bianchi and Gersonde, 2004) (contour in Fig. 7, right). The extended sea-ice coverage induces a decrease in export production in the Southern Ocean due to light limitation. Both the simulated export production and opal production in the Southern Ocean significantly decrease during the ACR (Figs. 10 and 7, right), in agreement with paleoproxies from the Southern Ocean. Marine sediment core TN057-13 (Anderson et al., 2009) exhibits a decline in opal flux during the ACR, which corresponds to a decrease in diatom-productivity. In addition, a decrease in alkenone content indicating a decrease in marine productivity during the ACR is recorded in marine sediment core MD97-2120 (Sachs and Anderson, 2005). When no freshwater pulse is added to the Southern Ocean (experiment DGN), the simulated anomalies in marine export and opal production obtained are much smaller and last for only 1000 years instead of 1700 years for DGNS (Fig. 10). 4.3. Younger Dryas The shut down of the AMOC and the associated strengthening of AABW formation lead to an enhanced poleward heat transport into the Southern Ocean. Surface temperatures thus increase rapidly in the Southern Hemisphere. In conjunction with an increase of radiative forcing and changes in obliquity the simulated austral spring air temperature increases by 2.6 C between 14 and 12 ka B.P., which is somewhat less than for the temperature reconstructions from the EPICA Dome C and Vostok ice cores (Petit et al., 1999; Jouzel et al., 2007) (Fig. 8b). Both the simulated and reconstructed SST in the Southern Ocean increase by 2 C. However the simulated SST increases more abruptly than the reconstructed one (Fig. 8c). In the South Pacific, the simulated SST increases by 1.6 C compared to a reconstructed SST increase of about 3 C (Fig. 8d). In addition, the simulated SST decreases by 1 C at 11.5 ka B.P., in contrast to the paleo- Fig. 12. Time series of annual terrestrial primary production anomalies (GtC/yr) averaged over the latitudinal band 0e30 S (cyan), 0e20 N (red), 45 Ne70 N (magenta). The green line represents the sum of the cyan, red and magenta lines. The black line represents the time series of the atmospheric CH4 (ppb) as recorded in GISP2 ice core (Brook et al., 2000). (For interpretation of the references to colour in this figure legend, the reader is referred to the web version of this article.) Author's personal copy 1168 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 proxy indices used here. Due to the warmer conditions at high southern latitudes and reduced sea-ice coverage, the export and opal productions also increase strongly in agreement with paleoproxies data (Sachs and Anderson, 2005; Anderson et al., 2009) (Fig. 10). In agreement with paleorecords, wetter conditions prevail over the Bolivian Altiplano (Fig. 9a) (Baker et al., 2001a; 2001b). Wetter conditions are also simulated over the Peruvian Altiplano (Fig. 9b), while the Huascaran d18O records show only a slight increase in precipitation (Thompson et al., 1995). Our model simulates a strengthening of the South American Summer Monsoon over Brazil, in agreement with the d18O record from Botuverá cave (Cruz et al., 2005; Wang et al., 2007) (Fig. 9c). 5. Discussion and conclusion The transient deglaciation experiments performed in this study successfully capture the dominant climate and some biochemical features of the last deglaciation in both hemispheres. This good correspondence between model results and paleo-proxy data suggests that most of the millennial-scale variability recorded during the last deglaciation can be explained by freshwater-induced variations in the deep water formation strength in the North Atlantic, Southern Ocean and North Pacific. Based on our modeling results it is justified to conclude that the North Atlantic is the key driver of millennial-scale variability during the Last Glacial Termination. In this study freshwater was mostly added in a broad area of the North Atlantic, not taking into account differences that would arise from changes in the freshwater routing around the North Atlantic (e.g. through the Norwegian Sea, Labrador Sea, Gulf of Mexico) (Aharon, 2003; Tarasov and Peltier, 2006; Sionneau et al., 2010). Therefore, we did not differentiate between the North Atlantic deep water formation sites, i.e. the Labrador Sea and the Greenland Iceland and Norwegian (GIN) Sea. As seen in Fig.1a, paleo-proxy records from different North Atlantic locations suggest slightly different scenarios. The Bermuda rise 231Pa/230Th data (McManus et al., 2004) and the South Iceland rise ventilation age record (Thornalley et al., 2011) indicate a weakened AMOC already at 18 ka B.P., whereas the Iberian margin 231Pa/230Th data shows a weakened AMOC starting 16 ka B.P (Gherardi et al., 2005). Gherardi et al. (2005) suggested that the Labrador Sea NADW formation site might have weakened before the GIN site, which is actually not supported by the data presented in Thornalley et al. (2011). In the Northern Hemisphere, the Cariaco basin reflectance record (Peterson et al., 2000) is the only dataset indicating a late (w16.2 ka B.P.) initiation of H1. On the other hand, the South Iceland rise data suggest a greater ventilation between 16 and 15 ka B.P., which is not seen in other North Atlantic circulation and temperature records. Even if a strong weakening of the NADW formation most likely occurred during H1, the impact of variations in the North Atlantic convection sites needs to be further investigated. Whether a more detailed knowledge of the freshwater routing into the North Atlantic would help to reconcile the conflicting timing in paleo-proxy data during H1 remains to be shown. A shut down of the AMOC during H1 leads to cold conditions over the North Atlantic while the Southern Hemisphere starts to warm. A related reduction of Atlantic-Pacific moisture transport leads to an increase of North Pacific salinity and eventually triggers deep water Formation in the Bering Sea. This leads to the establishment of a deep Pacific Meridional Overturning Circulation during H1 in agreement with paleo-proxy records (Okazaki et al., 2010). The closing of the Bering Strait is necessary for the formation of NPDW (Okumura et al., 2009). Further North Pacific paleoSST records and modeling studies would be needed to better constrain the associated SST patterns in the North Pacific during H1. The rapid and strong resumption of the AMOC at about 15 ka B.P. leads to the Bølling warming in the North Atlantic and initiates the ACR at high Southern latitudes. The ACR can be simulated by a weakening of the AABW formation, either as a bipolar seesaw response to the strengthened AMOC or as a result of a freshwater input in the Southern Ocean. The knowledge of AABW dynamics is too limited at the moment to conclude whether the bipolar seesaw is sufficient to weaken the AABW or to estimate the amount of freshwater necessary to weaken the AABW for about 2000 years. In agreement with paleoproxies (Sachs and Anderson, 2005; Anderson et al., 2009), our model successfully simulates a decrease in export and opal production in the Southern Ocean during the ACR due to the sea-ice edge advance and colder conditions. It should be however noted that our model does not include iron limitation. Changes in export production therefore do not take into account any possible changes in iron supply to the euphotic zone as well as non-Redfield processes. Concurrently in the North Hemisphere, the Older Dryas (14.4e14 ka B.P) is simulated by a substantial weakening of the AMOC. Our simulation thus suggests that melting from both, Antarctica and the Northern Hemispheric ice sheets contributed to MWP-1A at w14.4 ka B.P. The exact timing, shape and location of the different freshwater pulses contributing to MWP-1A still need to be better constrained to allow for a more quantitative comparisons between climate model simulations and paleo-proxy data. The model results suggest that climate sensitivity to southern ocean freshwater input is largest in the high southern latitudes. Future model-proxy comparisons could thus improve the estimates of the Antarctic meltwater contribution to MWP-1A. The Younger Dryas is represented by a freshwater input into the Arctic Ocean following the hypothesis of Tarasov and Peltier (2005) and Murton et al. (2010). The induced shut down of the AMOC captures most but not all of the features of the YD in the Northern Hemisphere. 231Pa/230Th data of (McManus et al., 2004) suggests that only a weakening and not a complete shut down of the AMOC occurred during the YD. Our model is not able to simulate an intermediate strength of the AMOC. In addition, the amplitude of the cooling observed in GISP2 (Alley, 2000), in the North Atlantic (Cacho et al., 1999, 2001; Bard et al., 2000; Calvo et al., 2001; Waelbroeck et al., 2001; Bard, 2002; Martrat et al., 2004, 2007) and in the Cariaco basin (Lea et al., 2003) can only be obtained in our model as the consequence of a shut down of the AMOC. In the Southern Hemisphere, temperatures increase in phase with the strengthening of AABW formation, starting around 13 ka B.P. We therefore find that, similarly to H1, temperature and precipitation variations between the North Atlantic and the Southern Hemisphere are anti-correlated during the YD, in conformance with the bipolar seesaw mechanism. Fig. 12 shows the time series of simulated changes in terrestrial primary production in different latitudinal bands compared to the atmospheric CH4 as recorded in GISP2 ice core (Brook et al., 2000). Wetlands are a main source of atmospheric CH4 and it has been estimated that tropical region wetlands (20 Ne30 S) were responsible for about 60% of the total wetland emissions, while the high northern latitudes wetlands (45 Ne70 N) were responsible for about 38% (Bartlett and Harris, 1993). Due to the cold and dry conditions prevailing in most of the Northern Hemisphere during H1, terrestrial primary production slightly decreases in the latitudinal band 0e20 N (10%). Wetter and warmer conditions in the Southern Hemisphere lead to greater terrestrial primary production in the latitudinal band 0e30 S (þ20%), which could induce the measured atmospheric CH4 increase. Overall, the simulated terrestrial carbon stock during H1 increases by about 100 GtC. The abrupt increase in atmospheric CH4 at 15 ka B.P. corresponds to the Bølling warming and the associated warmer and wetter conditions in the Northern Hemisphere. The terrestrial primary productivity increases by w20% in the latitudinal band Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 0e20 N and 35% over 45 Ne70 N (Fig. 12). As a result of the drier conditions prevailing in the Southern Hemisphere during the ACR, the terrestrial primary production slightly decreases (5%) in the latitudinal band 0e30 S. Overall, the terrestrial carbon stock increases by about 260 GtC between 15 and 13 ka B.P. Finally, the drier conditions over the Northern Hemisphere during the YD induce a decrease in terrestrial primary production by 60% in the latitudinal band 0e20 N and by 30% in the latitudinal band 45e70 N. The terrestrial carbon stock decreases by only 60GtC during the YD after which it increases by almost 300GtC. In this study we suggest a sequence of changes in the strength of deep water formation in the North Atlantic, the Southern Ocean and the North Pacific, which gives a reasonable representation of the reconstructed millennial-scale climatic variability during the Last Glacial Termination. To a large extent, we find that the relative timing of events observed in paleo-proxy data from different regions is consistent with the relative timing in the model. Ensemble-simulations with some form of data-assimilation (Goosse et al., 2010b; Jackson et al., 2010) could help to put an improved constraint on the freshwater forcing time series and, subsequently, the phase relations among proxy time series. A more detailed study of the climatic influence of the different deep water formation sites in the North Atlantic (similar to (Roche et al., 2010)) would also help to improve the understanding of the deglacial climatic changes. Acknowledgment The authors would like to acknowledge the NOAA NGDC database and thank O. Heiri, K. Holmgren, K. Pahnke and J. Sachs for providing their datasets. The authors would like to thank P. Baker for discussion about his dataset. We thank W. Hiscock for his graphic support. This research was supported by NSF grant No. 1010869. Additional support was provided by the Japan Agency for MarineEarth Science and Technology (JAMSTEC), by NASA and NOAA through their sponsorship of research activities at the International Pacific Research Center. A. Mouchet acknowledges support from the Belgian Science Policy (BELSPO contract SD/CS/01A). This is IPRC publication number 756 and SOEST contribution number 8096. References Aharon, P., 2003. Meltwater flooding events in the Gulf of Mexico revisited: implications for rapid climate changes during the last deglaciation. Paleoceanography 18, doi:10.1029/2002PA000840. Alley, R., 2000. The Younger Dryas cold interval as viewed from central Greenland. Quat. Sci. Rev. 19, 213e226. Anderson, R.F., Ali, S., Bradtmiller, L., Nielsen, S., Fleisher, M., Anderson, B., Burckle, L., 2009. Wind-driven upwelling in the Southern Ocean and the deglacial rise in Atmospheric CO2. Science 323, 1443e1448. Baker, P., Rigsby, C., Seltzer, G., Frisk, S., Lowenstein, T., Bacher, N., Veliz, C., 2001a. Tropical climate changes at millenial and orbital timescales on the Bolivian Altiplano. Nature 409, 698e701. Baker, P., Seltzer, G., Frisk, S., Dunbar, R., Grove, M., Tapia, P., Cross, S., Rowe, H., Broda, J., 2001b. The history of South American tropical precipitation for the past 25,000 years. Science 291, 640e643. Bar-Matthews, M., Ayalon, A., Gilmour, M., Matthews, A., Hawkesworth, C., 2003. Sea-land oxygen isotopic relationships from planktonic foraminifera and speleothems in the Eastern Mediterranean region and their implication for paleorainfall during interglacial intervals. Geochim. Cosmochim. Acta 67, 3181e3199. Bard, E., 2002. Abrupt climate changes over millennial time scales: climate shock. Phys. Today 55, 32e38. Bard, E., Rostek, F., Turon, J.-L., Gendreau, S., 2000. Hydrological impact of Heinrich events in the subtropical Northeast Atlantic. Science 289, 1321e1324. Barker, S., Diz, P., Vautravers, M., Pike, J., Knorr, G., Hall, I., Broecker, W., 2009. Interhemispheric Atlantic seesaw response during the last deglaciation. Nature 457, 1097e1102. Bartlett, K., Harris, R., 1993. Review and assessment of methane emissions from wetlands. Chemosphere 26, 261e320. Bassett, S., Milne, G., Mitrovica, J., Clark, P., 2005. Ice sheet and solid Earth influences on far-field sea-level histories. Science 309, 925e928. 1169 Berger, A., 1978. Long term variations of daily insolation and Quaternary climate change. J. Atmos. Sci. 35, 2362e2367. Bianchi, C., Gersonde, R., 2004. Climate evolution at the last deglaciation: the role of the Southern Ocean. Earth Planetary Sci. Lett. 228, 407e424. Bindschadler, R., King, M., Alley, R., Anandakrishnan, S., Padman, L., 2003. Tidally controlled stick-slip discharge of a West Antarctic ice stream. Science 301, 1087e1089. Blard, P.H., Lavé, J., Farley, K., Fornari, M., Jimenez, N., Ramirez, V., 2009. Late local glacial maximum in the Central Altiplano triggered by cold and locally-wet conditions during the paleolake Tauca episode (17-15 ka, Heinrich 1). Quat. Sci. Rev., doi:10.1016/j.quascirev.2009.09.025. Braconnot, P., Otto-Bliesner, B., Harrison, S., Joussaume, S., Peterschmitt, J.-Y., AbeOuchi, A., Crucifix, M., Driesschaert, E., Fichefet, T., Hewitt, C.D., Kageyama, M., Kitoh, A., Laîné, A., Loutre, M.-F., Marti, O., Merkel, U., Ramstein, G., Valdes, P., L.Weber, S., Yu, Y., Zhao, Y., 2007. Results of PMIP2 coupled simulations of the Mid-Holocene and Last Glacial Maximum-Part 1: experiments and large-scale features. Clim. Past 3, 261e277. Brook, E.J., Harder, S., Severinghaus, J., Steig, E., Sucher, C., 2000. On the origin and timing of rapid changes in atmospheric methane during the last glacial period. Global Biogeochem. Cycles 14, 559e572. Brook, E.J., Sowers, T., Orchardo, J., 1996. Rapid variations in atmospheric methane concentrations during the past 110,000 Years. Science 273 (5278), 1087e1091. Brovkin, V., Ganopolski, A., Svirezhev, Y., 1997. A continuous climate-vegetation classification for use in climate-biosphere studies. Ecol. Modell 101, 251e261. Bush, A.B.G., Philander, S.G.H., 1998. The role of ocean-atmosphere interactions in tropical cooling during the last glacial maximum. Science 279, 1341e1344. Cacho, I., Grimalt, J., Canals, M., Sbaffi, M., Shackleton, N., 2001. Variability of the western Mediterranean Sea surface temperature during the last 25,000 years and its connection with the Northern Hemisphere climatic changes. Paleoceanography 16, 40e52. Cacho, I., Grimalt, J., Pelejero, C., Canals, M., Sierro, F., Flores, J., Shackleton, N., 1999. Dansgaard-Oeschger and Heinrich event imprints in Alboran Sea paleotemperatures. Paleoceanography 14, 698e705. Calvo, E., Pelejero, C., Deckker, P.D., Logan, G., 2007. Antarctic deglacial pattern in a 30 kyr record of sea surface temperature offshore South Australia. Geophys. Res. Lett. 34, doi:10.1029/2007GL029937. Calvo, E., Villanueva, J., Grimalt, J., Boelaert, A., Labeyrie, L., 2001. New insights into glacial latitudinal temperature gradients in the North Atlantic. Results from UK`37 sea surface temperature and terrigenous inputs. Earth Planet. Sci. Lett. 188, 509e519. Campin, J., Goosse, H., 1999. Parameterization of density-driven downsloping flow for a coarse-resolution ocean model in z-coordinate. Tellus 51A, 412e430. Carlson, A., 2009. Geochemical constraints on the Laurentide ice sheet contribution to meltwater pulse 1A. Quat. Sci. Rev. 28, 1625e1630. Carter, L., Neil, H., Northcote, L., 2002. Late Quaternary ice-rafting events in the SW Pacific Ocean, off eastern New Zealand. Marine Geol. 191, 19e35. Charbit, S., Kageyama, M., Roche, D., Ritz, C., Ramstein, G., 2005. Investigating the mechanisms leading to the deglaciation of past continental northern hemisphere ice sheets with the CLIMBER-GREMLINS coupled model. Global Planet. Change 48 (4), 253e273. Chikamoto, M., Menviel, L., Abe-Ouchi, A., Ohgaito, R., Timmermann, A., Okazaki, Y.N., Harada, A.O., Mouchet, A., 2010. Variability in North Pacific intermediate and deep water ventilation during Heinrich events in two coupled climate models. Deep Sea Res. (accepted). Clark, P., Mitrovica, J., Milne, G., Tamisiea, M., 2002. Sea-level fingerprinting as a direct test for the source of global Meltwater Pulse 1A. Science 295, 2438e2441. CLIMAP-Project-Members, 1981. Map and Chart Ser. MC-36. Seasonal Reconstruction of the Earth Surface at the Last Glacial Maximum. Palisades, Ch. LamontDoherty Geological Observatory of Columbia University. Cruz, F., Burns, S., Karmann, I., Sharp, W., Vuille, M., Cardoso, A., Ferrari, J., Dias, P.S., Viana Jr., O., 2005. Insolation-driven changes in atmospheric circulation over the past 116,000 years in subtropical Brazil. Nature 434, 63e66. Cuffey, K., Clow, G., 1997. Temperature, accumulation, and ice sheet elevation in central Greenland through the last deglacial transition. J. Geophys. Res. 102, 26383e26396. Driesschaert, E., Fichefet, T., Goosse, H., Huybrechts, P., Janssens, I., Mouchet, A., Munhoven, G., Brovkin, V., Weber, S., 2007. Modeling the influence of Greenland ice sheet melting on the Atlantic meridional overturning circulation during the next millennia. Geophys. Res. Lett. 34, doi:10.1029/2007GL029516. Dykoski, C., Edwards, R., Cheng, H., Yuan, D., Cai, Y., Zhang, M., Lin, Y., Qing, J., 2005. A high-resolution, absolute-dated Holocene and deglacial Asian monsoon record from Dongge Cave, China. Earth Planet. Sci. Lett. 233, 71e86. Emmer, E., Thunell, R., 2000. Nitrogen isotope variations in Santa Barbara Basin sediments: implications for denitrification in the eastern tropical North Pacific during the last 50,000 years. Paleoceanography 15, 377e387. Evans, M., Rutter, N., Catto, N., Chlachula, J., Nyvlt, D., 2003. Magnetoclimatology: Teleconnection between the Siberian loess record and north Atlantic Heinrich events. Geology 31, 031, doi:10.1130/0091e7613 (2003). Fairbanks, R., 1989. A 17,000-year glacio-eustatic sea level record: influence of glacial melting rates on the Younger Dryas event and deep-ocean circulation. Nature 342, 637e642. Felis, T., Lohmann, G., Kuhnert, H., Lorenz, S., Scholz, D., Pätzold, J., Rousan, A.A., AlMoghrabi, S., 2004. Increased seasonality in Middle East temperatures during the Last Interglacial period. Nature 429, 164e168. Author's personal copy 1170 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Fleitmann, D., Cheng, H., Badertscher, S., Edwards, R., Mudelsee, M., Göktürk, O., Fankhauser, A., Pickering, R., Raible, C., Matter, A., Kramers, J., Tüysüz, O., 2009. Timing and climatic impact of Greenland interstadials recorded in stalagmites from northern Turkey. Geophys. Res. Lett. 36. Flower, B., Hastings, D., Hill, H., Quinn, T., 2004. Phasing of deglacial warming and Laurentide ice sheet meltwater in the Gulf of Mexico. Geology 32. Flückiger, J., Knutti, R., White, J., 2006. Oceanic processes as potential trigger and amplifying mechanisms for Heinrich events. Paleoceanography 21, PA2014. Genty, D., Blamart, D., Ghaleb, B., Plagnes, V., Causse, C., Bakalowicz, M., Zouari, K., Chkir, N., Hellstrom, J., Wainer, K., Bourges, F., 2006. Timing and dynamics of the last deglaciation from European and North African d13C stalagmite profilescomparison with Chinese and South hemisphere stalagmites. Quat. Sci. Rev. 25, 2118e2142. Gherardi, J., Labeyrie, L., McManus, J., Francois, R., Skinner, L., Cortijo, E., 2005. Evidence from the Northeastern Atlantic basin for variability in the rate of the meridional overturning circulation through the last deglaciation. Earth Planet. Sci. Lett. 240, 710e723. Goosse, H., Brovkin, V., Fichefet, T., Jongma, J., Huybrechts, P., Mouchet, A., Barriat, P.-Y., Campin, J.-M., Deleersnijder, E., Driesschaert, E., Goelzer, H., Haarsma, R., Janssens, Y., Loutre, M.-F., Maqueda, M.A.M., Opsteegh, T., Mathieu, P.-P., Munhoven, G., Petterson, E., Renssen, H., Roche, D., Schaeffer, M., Selten, F., Severijns, C., Tartinville, B., Weber, N., 2010a. Description of the Earth system model of intermediate complexity LOVECLIM version 1.2. Geoscientific Model. Dev. (submitted). Goosse, H., Crespin, E., de Montety, A., Mann, M., Renssen, H., Timmermann, A., 2010b. Reconstructing surface temperature changes over the past 600 years using climate model simulations with data assimilation. J. Geophys. Res. Atmospheres 115, doi:10.1029/2009JD012737. Goosse, H., Deleersnijder, E., Fichefet, T., England, M., 1999. Sensitivity of a global coupled ocean-sea ice model to the parameterization of vertical mixing. J. Geophys. Res. 104 (C6), 13681e13695. Goosse, H., Driesschaert, E., Fichefet, T., Loutre, M.-F., 2007. Information on the early Holocene climate constrains the summer sea ice projections for the 21st century. Clim. Past 3, 683e692. Goosse, H., Fichefet, T., 1999. Importance of ice-ocean interactions for the global ocean circulation: a model study. J. Geophys. Res. 104 (C10), 23337e23355. Goslar, T., Balaga, K., Arnold, M., Tisnerat, N.E.S., Kuzniarski, M., Chrost, L., Walanus, A., Wieckowski, K., 1999. Climate-related variations in the composition of the Lateglacial and early Holocene sediments of Lake Perespilno (eastern Poland). Quat. Sci. Rev. 18, 899e911. Gregoire, L., 2010. Modelling the northern hemisphere climate and ice sheets during the last deglaciation. Ph.D. thesis, School of Geographical Sciences, University of Bristol, UK. Harada, N., Ahagon, N., Sakamoto, T., Uchida, M., Ikehara, M., Shibata, Y., 2006. Rapid fluctuation of alkenone temperature in the southwestern Okhotsk Sea during the past 120 kyr. Glob. Planet. Change 53, 29e46. Harada, N., Sato, M., Seki, O., Timmermann, A., Moossen, H., Bendle, J., Nakamura, Y., Kimoto, K., Okazaki, Y., Nagashima, K., Gorbarenko, S., Ijiri, A., Nakatsuka, T., Menviel, L., Chikamoto, M., Abe-Ouchi, A., Schouten, S., 2010. Sea surface and subsurface temperature changes in the Okhotsk Sea and adjacent north Pacific during the last glacial maximum and deglaciation. Deep Sea Res. (part II submitted). Haug, G., Hughen, K., Sigman, D., Peterson, L., Röhl, U., 2001. Southward migration of the Intertropical Convergence zone through the Holocene. Science 293, 1304e1308. Heiri, O., Cremer, H., Engels, S., Hoek, W., Peeters, W., Lotter, A., 2007. Lateglacial summer temperatures in the Northwest European lowlands: a chironomid record from Hijkermeer, the Netherlands. Quat. Sci. Rev. 26, 2420e2437. Heiri, O., Filippi, M.-L., Lotter, A., 2005. Lateglacial summer temperature in the Trentino area (Northern Italy) as reconstructed by fossil chironomid assemblages in Lago di Lavarone (1100 m a.s.l.) Studi Trent. Sci. Nat. Acta Geol. 82, 299e308. Heiri, O., Millet, L., 2005. Reconstruction of late glacial summer temperatures from chironomid assemblages in Lac Lautrey (Jura, France). J. Quat. Sci. 20, 33e44. Hewitt, C.D., Broccoli, A.J., Mitchell, J.F.B., Stouffer, R., 2001. A coupled model study of the last glacial maximum: was the North Atlantic relatively warm? Geophys. Res. Lett. 28 (8), 1571e1574. Holmgren, K., Lee-Thorp, J., Cooper, G., Lundblad, K., Partridge, T., Scott, L., Sithaldeen, R., Talma, A., Tyson, P., 2003. Persistent millennial-scale climatic variability over the past 25,000 years in Southern Africa. Quat. Sci. Rev. 22, 2311e2326. Ivanochko, T., Ganeshram, R., Brummer, G.-J., Ganssen, G., Jung, S., Moreton, S., Kroon, D., 2005. Variations in tropical convection as an amplifier of global climate change at the millennial scale. Earth Planet. Sci. Lett. 235, 302e314. Ivanochko, T., Pedersen, T., 2004. Determining the influences of Late Quaternary ventilation and productivity variations on Santa Barbara Basin sedimentary oxygenation: a multi-proxy approach. Quat. Sci. Rev. 23, 467e480. Ivy-Ochs, S., Kerschner, H., Kubik, P., Schluchter, C., 2006. Glacier response in the European Alps to Heinrich event 1 cooling: the Gschnitz stadial. J. Quat. Sci. 21, 115e130. Jackson, C.S., Broccoli, A.J., 2003. Orbital forcing of Arctic climate: mechanisms of climate response and implications for continental glaciation. Clim. Dyn 21 (7e8), 539e557. Jackson, C.S., Marchal, O., Liu, Y., Lu, S., Thompson, W.G., 2010. A box-model test of the freshwater forcing hypothesis of abrupt climate change and the physics governing ocean stability. Paleoceanography 25. Jennerjahn, T., Ittekkot, V., Arz, H., Behling, H., Pätzold, J., Wefer, G., 2004. Asynchronous terrestrial and marine signals of climate change during Heinrich events. Science 306, 2236e2239. Joussaume, S., Taylor, K.E., Braconnot, P., Mitchell, J.F.B., Kutzbach, J.E., Harrison, S.P., Prentice, I.C., Broccoli, A.J., Abe-Ouchi, A., Bartlein, P.J., Bonfils, C., Dong, B., Guiot, J., Herterich, K., Hewitt, C.D., Jolly, D., Kim, J.W., Kislov, A., Kitoh, A., Loutre, M.F., Masson, V., McAvaney, B., McFarlane, N., de Noblet, N., Peltier, W.R., Peterschmitt, J.Y., Pollard, D., Rind, D., Royer, J.F., Schlesinger, M.E., Syktus, J., Thompson, S., Valdes, P., Vettoretti, G., Webb, R.S., Wyputta, U., 1999. Monsoon changes for 6000 Years ago: results of 18 simulations from the Paleoclimate modeling Intercomparison Project (PMIP). Geophys. Res. Lett. 26 (7), 859e862. Jouzel, J., Masson, V., Cattani, O., Dreyfus, G., Falourd, S., Hoffmann, G., Minster, B., Nouet, J., Barnola, J., Chappellaz, J., Fischer, H., Gallet, J., Leuenberger, S.J.M., Loulergue, L., Luethi, D., Oerter, H., Parrenin, F., Raisbeck, G., Raynaud, D., Schilt, A., Schwander, J., Selmo, E., Souchez, R., Spahni, R., Stauffer, B., Steffenssen, J.P., Stenni, B., Stocker, T., Tison, J., Werner, M., Wolff, E., 2007. Orbital and millennial Antarctic climate variability over the past 800,000 years. Science 317, 793e796. Justino, F., Timmermann, A., Merkel, U., Souza, E., 2005. Synoptic reorganization of atmospheric flow during the last glacial maximum. J. Clim. 18, 2826e2846. Kaiser, J., Lamy, F., Hebbeln, D., 2005. A 70-kyr sea surface temperature record off Southern Chile. Paleoceanography 20, doi:10.1029/2004PA001146. Kanfoush, S., Hodell, D., Charles, C., Guilderson, T., Mortyn, P., Ninnemann, U., 2000. Millennial-scale instability of the antarctic ice sheet during the last glaciation. Science 288, 1815e1818. Kennett, J., Ingram, B., 1995. A 20,000 year record of ocean circulation and climate change in the Santa Barbara basin. Nature 377, 510e514. Kerschner, H., Kaser, G., Sailer, R., 2000. Alpine Younger Dryas glaciers as palaeoprecipitation gauges. Annals. Glacio 31, 80e84. Kim, J.-H., Schneider, R., Hebbeln, D., Muller, P., Wefer, G., 2002. Last deglacial seasurface temperature evolution in the Southeast Pacific compared to climate changes on the South American continent. Quat. Sci. Rev. 21, 2085e2097. Kim, S., Flato, G., Boer, G., 2003. A coupled climate model simulation of the Last Glacial Maximum, Part 2: approach to equilibrium. Clim. Dyn. 20, 635e661. Kitoh, A., Murakami, S., Koide, H., 2001. A simulation of the Last Glacial Maximum with a coupled atmosphere-ocean GCM. Geophys. Res. Lett. 28, 2221e2224. Köhler, P., Knorr, G., Buiron, D., Lourantou, A., Chappellaz, J., 2010. Rapid changes in ice core gas records-Part 2: understanding the rapid rise in atmospheric CO2 at the onset of the Bølling/Allerød. Clim. Past Discuss. 6, 1473e1501. Kramer, A., Herzschuh, U., Mischke, S., Zhang, C., 2010. Late glacial vegetation and climate oscillations on the southeastern Tibetan Plateau inferred from the Lake Naleng pollen profile. Quat. Res. 73, 324e335. Lamb, H., Bates, C., Coombes, P., Marshall, M., .Umer, M., Davies, S., Dejen, E., 2007. Late Pleistocene desiccation of Lake Tana, source of the blue Nile. Quat. Sci. Rev.. Lea, D., Pak, D., Peterson, L., Hughen, K., 2003. Synchroneity of tropical and highlatitude Atlantic temperatures over the last glacial termination. Science 301, 1361e1364. Lehman, S., Keigwin, L., 1992. Sudden changes in North Atlantic circulation during the last deglaciation. Nature 356, 757e762. Licht, K., 2004. The Ross Sea’s contribution to eustatic sea level during meltwater pulse 1A. Sedimentary Geol. 165, 343e353. Lim, G., Holton, J., Wallace, J., 1991. The structure of the ageostrophic wind field in baroclinic waves. J. Atmos. Sci. 48, 1733e1745. Liu, Z., Brady, E., Lynch-Steiglitz, J., 2003. Global ocean response to orbital forcing in the Holocene. Paleoceanogr 18, 1041. doi:10.1029/2002PA000819. Liu, Z., Otto-Bliesner, B., He, F., Brady, E., Tomas, R., Clark, P., Carlson, A., LynchStieglitz, J., Curry, W., Brook, E., Erickson, D., Jacob, R., Kutzbach, J., Cheng, J., 2009. Transient simulation of last deglaciation with a new mechanism for Bølling-Allerød warming. Science 325, 310e314. Liu, Z., Shin, S., Behling, P., Prell, W., Trend-Dtaid, M., Harisson, S., Kutzbach, J., 2000. Dynamical and observational constraints on tropical Pacific sea surface temperatures at the Last Glacial Maximum. Geophys. Res. Lett. 27, 105e108. Liu, Z., Shin, S., Otto-Bliesner, B., Kutzbach, J., Brady, E., 2002. Enhanced tropical cooling at the last glacial maximum by extratropical ocean ventilation. Geophys. Res. Lett. 29, doi:10.1029/2001GL013938. Lorenz, S., Lohmann, G., 2004. Acceleration technique for Milankovitch type forcing in a coupled atmosphere-ocean circulation model: method and application for the Holocene. Clim. Dyn. 23, 727e743, doi:10.1007/s00382-004-0469. Lorenz, S.J., Kim, J.H., Rimbu, N., Schneider, R.R., Lohmann, G., 2006. Orbitally driven insolation forcing on Holocene climate trends: evidence from alkenone data and climate modeling. Paleoceanography 21, doi:10.1029/2005PA001152. Lunt, D.J., Williamson, M.S., Valdes, P.J., Lenton, T.M., 2006. Comparing transient, accelerated, and equilibrium simulations of the last 30,000 years with the GENIE-1 model. Clim. Past. Discuss. 2, 267e283. Lüthi, D., Floch, M.L., Bereiter, B., Blunier, T., Barnola, J.-M., Siegenthaler, U., Raynaud, D., Jouzel, J., Fischer, H., Kawamura, K., Stocker, T., 2008. High-resolution carbon dioxide concentration record 650,000-800,000 years before present. Nature 453, doi:10.1038. Marsh, R., Smith, M.P.L.M., Rohling, E.J., Lunt, D.J., Lenton, T.M., Williamson, M.S., Yool, A., 2006. Modelling Ocean circulation, climate and oxygen isotopes in the Ocean over the last 120,000 Years. Clim. Past Discuss. 2, 657e709. Author's personal copy L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Martrat, B., Grimalt, J., Lopez-Martinez, C., Cacho, I., Sierro, F., Flores, J.A., Zahn, R., Canals, M., Curtis, J., Hodell, D., 2004. Abrupt temperature changes in the Western Mediterranean over the past 250,000 years. Science 306, 1762e1765. Martrat, B., Grimalt, J., Shackleton, N., de Abreu, L., Hutterli, M., Stocker, T., 2007. Four climate cycles of recurring deep and surface water destabilizations on the Iberian margin. Science 317, 502e507. Maslin, M., Burns, S., 2000. Reconstruction of the Amazon basin effective moisture availability over the past 14,000 years. Science 290, 2285e2287. McManus, J.F., Francois, R., Gherardi, J.M., Keigwin, L.D., Brown-Leger, S., 2004. Collapse and rapid resumption of Atlantic meridional circulation linked to deglacial climate changes. Nature 428, 834e837. Menviel, L., Timmermann, A., Mouchet, A., Timm, O., 2008a. Climate and marine carbon cycle response to changes in the strength of the southern hemispheric westerlies. Paleoceanography 23, doi:10.1029/2007PA001604. Menviel, L., Timmermann, A., Mouchet, A., Timm, O., 2008b. Meridional reorganizations of marine and terrestrial productivity during Heinrich events. Paleoceanography 23, doi:10.1029/2007PA001445. Menviel, L., Timmermann, A., Mouchet, A., Timm, O., 2010a. Climate and biogeochemical response to a rapid melting of the West-Antarctic Ice Sheet during interglacials and implications for future climate. Paleoceanography (accepted). Menviel, L., Timmermann, A., Mouchet, A., Timm, O., Abe-Ouchi, A., Chikamoto, M., Harada, N., Ohgaito, R., Okazaki, Y., 2010b. Removing the North Pacific halocline: effects on global climate, ocean circulation and the carbon cycle. Deep Sea Res. Part (accepted). Monnin, E., Indermühle, A., Dällenbach, A., Flückiger, J., Stauffer, B., Stocker, T., Raynaud, D., Barnola, J.-M., 2001. Atmospheric CO2 concentration over the last glacial termination. Science 291, 112e114. Mouchet, A., Francois, L., 1996. Sensitivity of a Global Oceanic Carbon Cycle Model to the circulation and to the fate of organic matter: preliminary results. Phys. Chem. Earth 21, 511e516. Murton, J., Bateman, M., Dallimore, S., Teller, J., Yang, Z., 2010. Identification of younger Dryas outburst flood path from Lake Agassiz to the Arctic ocean. Nature 464, 740e743. Niedermeyer, E., Prange, M., Mulitza, S., Mollenhauer, G., Schefuss, E., Schulz, M., 2009. Extratropical forcing of Sahel aridity during Heinrich stadials. Geophys. Res. Lett. 36, doi:10.1029/2009GL039687. Nielsen, S., 2004. Deglacial and Holocene Southern Ocean climate variability from diatom stratigraphy and paleoecology. Ph.D. thesis, University of Tromsø. Noumi, M., Yokoyama, Y., Miura, H., Ohkouchi, N., 2007. Melting history of the Antarctic ice sheet since the Last Glacial Maximum based on the analyses of sediment cores around the Antarctic Peninsula, Southern Ocean. Quaternary Res. Jpn. 46, 103e117. Okazaki, Y., Timmermann, A., Menviel, L., Harada, N., Abe-Ouchi, A., Chikamoto, M., Mouchet, A., Asahi, H., 2010. Deep water formation in the north Pacific during the last glacial termination. Science 329, 200e204. Okumura, Y., Deser, C., Hu, A., Timmermann, A., Xie, S.-P., 2009. North Pacific climate response to freshwater forcing in the Subarctic North Atlantic: oceanic and atmospheric pathways. J. Clim. 22, 1424e1445. Opsteegh, J., Haarsma, R., Selten, F., Kattenberg, A., 1998. ECBILT: a dynamic alternative to mixed boundary conditions in ocean models. Tellus 50A, 348e367. Otto-Bliesner, B.L., 1999. El Niño/La Niña and Sahel precipitation during the middle Holocene. Geophys. Res. Lett. 26, 87e90. Pahnke, K., Zahn, R., Elderfield, H., Schulz, M., 2003. 340,000-year centennial-scale marine record of Southern Hemisphere climatic oscillation. Science 301, 948e952. Peck, J., Green, R., Shanahan, T., King, J., Overpeck, J., Scholz, C., 2004. A magnetic mineral record of Late Quaternary tropical climate variability from Lake Bosumtwi. Ghana. Palaeogeogr. Palaeoclim. Palaeoecol 215, 37e57. Peltier, W., 1994. Ice-age paleotopography. Science 265, 195e201. Peltier, W., 2005. On the hemispheric origins of meltwater pulse 1a. Quat. Sci. Rev. 24, 1655e1671. Peterson, L., Haug, G., Hughen, K., Rohl, U., 2000. Rapid changes in the hydrologic cycle of the tropical Atlantic during the last glacial. Science 290, 1947e1951. Petit, J., Jouzel, J., Raynaud, D., Barkov, N., Barnola, J.-M., Basile, I., Bender, M., Chappellaz, J., Davis, M., Delaygue, G., Delmotte, M., Kotlyakov, V., Legrand, M., Lipenkov, V., Lorius, C., Pepin, L., Ritz, C., Saltzman, E., Stievenard, M., 1999. Climate and atmospheric history of the past 420,000 years from the Vostok ice core, Antarctica. Nature 399, 429e436. Peyron, O., Begeot, C., Brewer, S., Heiri, O., Magny, M., Millet, L., Ruffaldi, P., Campo, E.V., Yu, G., 2005. Late-Glacial climatic changes in Eastern France (Lake Lautrey) from pollen, lake-levels, and chironomids. Quat. Res. 64, 197e211. Pflaumann, U., Sarnthein, M., Chapman, M., d’Abreu, L., Funnell, B., Huels, M., Kiefer, T., Maslin, M., Schulz, H., Swallow, J., van Kreveld, S., Vautravers, M., Vogelsang, E., Weinelt, M., 2003. Glacial North Atlantic: sea-surface conditions reconstructed by GLAMAP 2000. Paleoceanography 18. Philippon, G., Ramstein, G., Charbit, S., Kageyama, M., Ritz, C., Dumas, C., 2006. Evolution of the Antarctic ice sheet throughout the last deglaciation: a study with a new coupled climate e north and south hemisphere ice sheet model. Earth Planet. Sci. Lett. 248, 750e758. Piotrowski, A., Goldstein, S., .Hemming, S.R., Fairbanks, R., 2004. Intensification and variability of ocean thermohaline circulation through the last deglaciation. Earth Planet. Sci. Lett. 225, 205e220. Pollard, D., DeConto, R., 2009. Modelling West Antarctic ice sheet growth and collapse through the past five million years. Nature 458, 329e332. 1171 Powers, L., Johnson, T., Werne, J., neda, I.C., Hopmans, E., Damste, J.S., Schouten, S., 2005. Large temperature variability in the southern African tropics since the Last Glacial Maximum. Geophys. Res. Lett. 32, doi:10.1029/2004GL022014. Pride, C., Thunell, R., Sigman, D., Keigwin, L., Altabet, M., Tappa, E., 1999. Nitrogen isotopic variations in the Gulf of California since the last deglaciation: response to global climate change. Paleoceanography 14, 397e409. Rasmussen, S., Andersen, K., Svensson, A., Steffensen, J., Vinther, B., Clausen, H., Andersen, M., Johnsen, S., Larsen, L., Dahl-Jensen, D., Bigler, M., Röthlisberger, R., Fischer, H., Goto-Azuma, K., Hansson, M., Ruth, U., 2006. A new Greenland ice core chronology for the last glacial termination. J. Geophys. Res. 111, 1e16. Renssen, H., Goosse, H., Fichefet, T., Brovkin, V., Driesschaert, E., Wolk, F., 2005. Simulating the Holocene climate evolution at northern high latitudes using a coupled atmosphere-sea ice-ocean-vegetation model. Clim. Dyn. 24, 23e43. Rind, D., 1987. Components of the ice age circulation. J. Geophys. Res. 92, 4241e4281. Roche, D., Renssen, H., Weber, S., Goosse, H., 2007. Could meltwater pulses have been sneaked unnoticed into the deep ocean during the last glacial? Geophys. Res. Lett. 34, doi:10.1029/2007GL032064. Roche, D., Wiersma, A., Renssen, H., 2010. A systematic study of the impact of freshwater pulses with respect to different geographical locations. Clim. Dyn. 34, 997e1013. Sachs, J., Anderson, R., 2005. Increased productivity in the subantarctic ocean during Heinrich events. Nature 434, 1118e1121. Samson, C., Sikes, E., Howard, W., 2005. Deglacial paleoceanographic history of the Bay of Plenty, New Zealand. Paleoceanography 20, doi:10.1029/2004PA001088. Sarnthein, M., Kiefer, T., Grootes, P., Elderfield, H., Erlenkeuser, H., 2006. Warmings in the far northwestern Pacific promoted pre-Clovis immigration to America during Heinrich event 1. Geology 34, 141e144. Shakun, J., Burns, S., Fleitmann, D., Kramers, J., Matter, A., Al-Subary, A., 2007. A high-resolution, absolute-dated deglacial speleothem record of Indian Ocean climate from Socotra Island, Yemen. Earth Planet. Sci. Lett. 259, 442e456. Shibahara, A., Ohkushi, K., Kennett, J., Ikehara, K., 2007. Late Quaternary changes in intermediate water oxygenation and oxygen minimum zone, northern Japan: a benthic foraminiferal perspective. Paleoceanography 22, doi:10.1029/ 2005PA001234. Shin, S.I., Liu, Z., Bliesner, B.L.O., Kutzbach, J.E., Brady., E., Harrison, S.P., 2003. A simulation of the last glacial maximum climate using the NCAR CSM. Clim. Dyn. 20, 127e151. Sionneau, T., Bout-Roumazeilles, V., Flower, B., Bory, A., Tribovillard, N., Kissel, C., Vliet-Lanoë, B.V., Serrano, J.M., 2010. Provenance of freshwater pulses in the Gulf of Mexico during the last deglaciation. Quat. Res. 74, 235e245. Somes, C., Schmittner, A., Galbraith, E., Lehmann, M., Altabet, M., Montoya, J., Letelier, R., Mix, A., Bourbonnais, A., Eby, M., 2010. Simulating the global distribution of nitrogen isotopes in the ocean. Glob. Biogeochem. Cycles 24, doi:10.1029/2009GB003767. Stanford, J., Rohling, E., Hunter, S., Roberts, A., Rasmussen, S., Bard, E., McManus, J., Fairbanks, R., 2006. Timing of meltwater pulse 1a and climate responses to meltwater injections. Paleoceanography 21, doi:10.1029/2006PA001340. Steig, E., Brook, E., White, J., Sucher, C., Bender, M., Lehman, S., Morse, D., Waddington, E., Clow, G., 1998. Synchronous climate changes in Antarctica and the north Atlantic. Science 282, 92e95. Stenni, B., Buiron, D., Frezzotti, M., Albani, S., Barbante, C., Bard, E., Barnola, J., Baroni, M., Baumgartner, M., Bonazza, M., Capron, E., Castellano, E., Chappellaz, J., Delmonte, B., Falourd, S., Genoni, L., Iacumin, P., Jouzel, J., Kipfstuhl, S., Landais, A., Lemieux-Dudon, B., Maggi, V., Masson-Delmotte, V., Mazzola, C., Minster, B., Montagnat, M., Mulvaney, R., Narcisi, B., Oerter, H., Parrenin, F., Petit, J., Ritz, C., Scarchilli, C., Schilt, A., Schüpbach, S., Schwander, J., Selmo, E., Severi, M., Stocker, T., Udisti, R., 2011. Expression of the bipolar see-saw in Antarctic climate records during the last deglaciation. Nat. Geosci. 4, 46e49. Stocker, T., 1998. The seesaw effect. Science 282, 61e62. Stuiver, M., Grootes, P., Braziunas, T., 1995. GISP2 d18O climate record of the past 16,500 years and the role of the sun, ocean and volcanoes. Quat. Res. 44, 341e354. Tarasov, L., Peltier, W., 2005. Arctic freshwater forcing of the Younger Dryas cold reversal. Nature 435, 662e665. Tarasov, L., Peltier, W., 2006. A calibrated deglacial drainage chronology for the North American continent: evidence of an Arctic trigger for the Younger Dryas. Quat. Sci. Rev. 25, 659e688. Thompson, L., Mosley-Thompson, E., Davis, M., Lin, P.-N., Henderson, K., Cole-Dai, J., Bolzan, J., Liu, K., 1995. Late glacial stage and Holocene tropical ice core records from Huascaran, Peru. Science 269, 46e50. Thornalley, D., Barker, S., Broecker, W., Elderfield, H., McCave, I., 2011. The deglacial evolution of north Atlantic deep convection. Science 331, 202e205. Thornalley, D., McCave, I., Elderfield, H., 2010. Freshwater input and abrupt deglacial climate change in the North Atlantic. Paleoceanography 25, doi:10.1029/ 2009PA001772. Timm, O., Köhler, P., Timmermann, A., Menviel, L., 2010. Mechanisms for the onset of the African Humid period and Sahara greening 14.5-11 ka BP. J. Clim., 2612e2633. Timm, O., Timmermann, A., 2007. Simulation of the last 21,000 years using accelerated transient boundary conditions. J. Clim. 20, 4377e4401. Timm, O., Timmermann, A., Abe-Ouchi, A., Segawa, T., 2008. On the definition of paleo-seasons in transient climate simulations. Paleoceanography PA2221, doi:10.1029/2007PA001461. Timmermann, A., An, S.I., Krebs, U., Goosse, H., 2005. ENSO suppression due to a weakening of the North Atlantic thermohaline circulation. J. Clim. 18, 3122e3139. Author's personal copy 1172 L. Menviel et al. / Quaternary Science Reviews 30 (2011) 1155e1172 Timmermann, A., Justino, F., Jin, F.-F., Krebs, U., Goosse, H., 2004. Surface temperature control in the North and tropical Pacific during the last glacial maximum. Clim. Dyn. 23, 353e370. Timmermann, A., Lorenz, S., An, S.I., Clement, A., Xie, S.-P., 2007. The effect of orbital forcing on the mean climate and variability of the tropical Pacific. J. Clim., 4147e4159. Timmermann, A., Menviel, L., 2009. What drives climate flip-flops? Science 325, 273e274. Timmermann, A., Menviel, L., Okumura, Y., Schilla, A., Merkel, U., Hu, A., OttoBliesner, B., Schulz, M., 2009a. Towards a quantitative understanding of millennial-scale Antarctic Warming events. Quat. Sci. Rev.. doi:10.1016/ j.quascirev.2009.06.021. Timmermann, A., Timm, O., Stott, L., Menviel, L., 2009b. The roles of CO2 and orbital forcing in driving southern hemispheric temperature variations during the last 21 000 yr. J. Clim. 22, 1626e1640. Tuenter, E., Weber, S.L., Hilgen, F.J., Lourens, L.J., Ganopolski, A., 2005. Simulation of climate phase lags in response to Precession and obliquity forcing and the role of vegetation. Clim. Dyn. 24, 279e295. Vaks, A., Bar-Matthews, M., Ayalon, A., Schilman, B., Gilmour, M., Hawkesworth, C., Frumkin, A., Kaufman, A., Matthews, A., 2003. Paleoclimate reconstruction based on the timing of speleothem growth and oxygen and carbon isotope composition in a cave located in the rain shadow in Israel. Quat. Res. 59, 182e183. Vidal, L., Labeyrie, L., Cortijo, E., Arnold, M., Duplessy, J., Michel, E., Becque, S., vanWeering, T., 1997. Evidence for changes in the North Atlantic Deep Water linked to meltwater surges during the Heinrich events. Earth Planet. Sci. Lett. 146, 13e27. Vink, A., Rühlemann, C., Zonneveld, K., Mulitza, S., 2001. Shifts in the position of the North Equatorial Current and rapid productivity changes in the western Tropical Atlantic during the last glacial. Paleoceanography 16, 479e490. Vinther, B., Buchardt, S., Clausen, H., Dahl-Jensen, D., Johnsen, S., Fisher, D., Koerner, R., Raynaud, D., Lipenkov, V., Andersen, K., Blunier, T., Rasmussen, S., Steffensen, J., Svensson, A., 2009. Holocene thinning of the Greenland ice sheet. Nature 461, 385e388. von Grafenstein, U., Erlenkeuser, H., Brauer, A., Jouzel, J., Johnsen, S., 1999. A MidEuropean decadal isotope climate record from 15,500 to 5000 years B.P. Science 284, 1654e1657. Waelbroeck, C., Duplessy, J.-C., Michel, E., Labeyrie, L., Paillard, D., Duprat, J., 2001. The timing of the last deglaciation in North Atlantic climate records. Nature 412, 724e727. Wang, X., Auler, A., Edwards, R., Cheng, H., Ito, E., Wang, Y., Kong, X., Solheid, M., 2007. Millennial-scale precipitation changes in southern Brazil over the past 90,000 years. Geophys. Res. Lett. 34, doi:10.1029/2007GL031149. Wang, Y., Cheng, H., Edwards, R., An, Z., Wu, J., Shen, C., Dorale, J., 2001. A highresolution absolute-dated Late Pleistocene monsoon record from Hulu Cave, China. Science 294, 2345e2348. Wang, Y., Wu, J., Liu, D., Wu, J., Cai, Y., Cheng, H., 2002. A quick cooling event of the East Asian monsson responding to Heinrich Event 1: evidence from stalagmite d18C records. Sci. China 45, 88e96. Weldeab, S., Lea, D., Schneider, R., Andersen, N., 2007. 155,000 years of West African Monsoon and ocean thermal evolution. Science 316, 1303e1307. Williams, M., Talbot, M., Aharon, P., Salaam, Y.A., Williams, F., Brendeland, K.I., 2006. Abrupt return of the summer monsoon 15,000 years ago: new supporting evidence from the lower White Nile valley and Lake Albert. Quat. Sci. Rev. 25, 2651e2665. Yu, Z., Eicher, U., 2001. Three amphi-Atlantic century-scale cold events during the Bølling-Allerød warm period. Geographie physique et Quaternaire 55, 171e179. Zuraida, R., Holbourn, A., Nürnberg, D., Kuhnt, W., Dürkop, A., Erichsen, A., 2009. Evidence for Indonesian throughflow slowdown during Heinrich events 3-5. Paleoceanography 24, doi:10.1029/2008PA001653.
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