HomonalediFACTSHEET EMBARGOEDUNTILTUESDAY9May2017–10amSouthAfricanTime/4amEDT/9am London/6pmSydney,Australia QUESTIONSANDANSWERS ABOUTTHELESEDICHAMBER 1)Whatdoesthename“Lesedi”mean? Theword“Lesedi”means“light”intheSetswanalanguage,whichisoneofthelanguages commonlyspokeninthispartofSouthAfrica.Theword“naledi”means“star”intheSesotho language,andtheword“Dinaledi”means“stars”. 2)WhereintheRisingStarcavesystemistheLesediChamber?IsitneartheDinaledi Chamber? TheRisingStarcavesystemincludesmorethan2kilometersofmappedundergroundpassages. TheLesediChamberisinadifferentpartofthecavesystemfromtheDinalediChamber,and theshortestroutebetweenthesechambersis145meterslong,involvingmultiplesqueezesand climbs,totallyinthedarkzone. 3)Howwerethefossilsfound? RickHunterandStevenTucker,whodiscoveredthefossilremainsintheDinalediChamberin September,2013,alsoenteredwhatwenowrecogniseastheLesediChamber,buttheydidnot recognisethehomininremainsinthechamberatthattime.Later,astheteamcarriedout fieldworkintheDinalediChamberinNovember2013,thesesameexplorerslocatedthefossil homininremainsintheLesediChamber.TwonewfossilareaswithintheLesediChamberwere discoveredbyHannahHilbert-Wolfduringgeologicalsurveyofthechamber. 4)Whoexcavatedthefossils? ThefossilswereexcavatedbyWitsteamsofscientistsincludingformer“Underground Astronaut”DrMarinaElliott. 5)HowdoestheexcavationteamreachtheLesediChamber?Isitashardtoenterasthe DinalediChamber? TheLesediChamberisapproximately30metersunderground.Toenterthechamber,theteam usesapaththatgoesmorethan80metersfromthesurface,twistingandturningthroughthe cave,withonetightsqueezethatonlysmallteammemberscanfitthrough,andsome dangerousdrops.ThischamberisnotasdifficulttoenterastheDinalediChamber,whichcan beenteredonlythroughthe18-cm(7½inch)-wideentrychute,whichisavertical12meter (40-foot)climb.ButthefossilremainsintheLesediChamberareinasmall,crampedside tunnelthatcanonlyaccommodateasingleexcavatoratatime.Itisaverychallengingsituation fortheteamtoworkin. THEFOSSILSFROMTHELESEDICHAMBER 6)HowmanyhomininfossilshavebeenrecoveredfromtheLesediChamber? Theteamhasrecovered131fossilhomininspecimensfromtheLesediChamber.Byitself,thisis oneofthemostproductivehomininsitesinAfrica. 7)HowmanyindividualsarerepresentedbythefossilremainsintheLesediChamber? TheteamhasexcavatedhomininfossilremainsfromthreeareaswithintheLesediChamber. Basedondifferencesbetweenadultandjuvenileskeletalremains,andtherepetitionofthe samepartsoftheskeleton,thefossilsincludetheremainsofatleasttwoadultsandone juvenileindividual.Fossilscomefromthreeseparateareaswithinthechamber,anditislikely thattheminimumnumberisanunderestimate. 8)Whois‘Neo’? Themajorityoftheremainsrecoveredsofarlooklikepartsoftheskeletonofasingleadult maleindividual,designatedLES1.Itistraditionalinthestudyofhumanevolutiontoname skeletonswhenwecanrecogniseindividuals,andthename“Neo”means“gift”intheSesotho language,oneofthelanguagescommonlyspokeninthispartofSouthAfricatoday.Thename resemblesacharacterfromthefilm,TheMatrix,however,thisresemblanceiscoincidental. 9)HowdoestheNeoskeletoncomparetootherskeletonsinthehomininfossilrecord? Becauseofthepresenceofanearly-completeskullwithotherpartsoftheskeleton,thisfindis trulyexceptional.Theskullandjawincludesfragilepartsofthefaceveryrarelypreservedin otherfossils.Thepreservedpartsofthearms,legs,wristandhand,spine,andribsrepresent everyotherregionoftheskeleton,exceptforthelowerlegsandfeet.Thetwothighbones (femora)aresomewhatdifferentinpartsthatmaysuggestthatoneofthemcomesfroma differentindividual,althoughifso,thetwoindividualswouldbeofverysimilarsize.The remainsareamongthemostcompleteeverdiscovered. 10)HowdoyouknowNeoisanadultmale? ThebonesandteethofNeoallindicatethathehadcompleteddevelopment,andtheteeth exhibitconsiderablewear,showinghewasarelativelyoldindividual.Neo’sskullisabitlarger thananyoftheskullsfoundintheDinalediChamber,andhasprominentattachmentsforthe musclesofthejawandneck,whichtendstobetrueofmaleswithinhumanandotherprimate species. 11)HowcanwebesurethattheskeletalremainsintheLesediChamberbelongtoHomo nalediandnotsomeotherspecies? H.nalediintheDinalediChamberisdistinguishedfromotherspeciesbasedonfeaturesacross theskeleton,fromtheskull,jawandteeth,totheshoulder,hand,spine,pelvis,leg,andfoot. SomeofthefeaturesofH.nalediareunique,whileothersoccurinapatternneverfound beforeinotherfossilorlivingspecies.TheremainsfromtheLesediChamberincludeanearly completeskullandjaw,alltheteeth,partsofthearmsandlegs,partsofthespineandribs, femoraandpelvicremains.Thiswealthofevidencehasenabledtheteamtocarryoutsomeof thestrongesttestseveraccomplishedforafossilskeletoncomparedtootherspecies.Allofthe featuresoftheLesediChamberhomininsareconsistentwithH.naledi,andeveryotherspecies canbeclearlyeliminated. 12)WhyaretherefewerhomininfossilsfromtheLesediChamberthanfromtheDinaledi Chamber? Theteam’sinvestigationofbothchambersisataveryearlyphase,andwedonotyetknowthe totalnumberofskeletalremainsineitherchamber,butlesssedimenthasbeenaccumulated andpreservedintheLesediChamber. 13)AretheDinalediandLesediHomonalediindividualscloserelatives? UntilwemanagetoobtainDNAfromthesefossilsitwillbeimpossibletotestwhetherthetwo groupsincludeclosegeneticrelatives.Allwecansayisthattheyareverymuchalikeandthat makestheirrelatednessseemmorelikely. 14)ArethereotheranimalbonesfromtheLesediChamber? TheteamhasrecoveredfragmentaryremainsofmanyanimalsfromtheLesediChamber. Nearlyallofthesebelongtorodentsorsmallcarnivores,suchassmallcats,foxes,and mongooses.WedonotknowiftheremainsoftheseanimalsenteredtheLesediChamberatthe sametimeasHomonaledi,orwhethertheyenteredatothertimes. ABOUTTHEGEOLOGICALAGEOFTHEFOSSILS 15a)Whatapproachdidtheteamtakeindeterminingthegeologicalageofthefossils fromtheDinalediChamber? Theteamdevelopedastrategythatinvolvedthreeseparatesteps.First,theDinaledichamber wascarefullymappedandthevarioussedimentsandflowstonesinthechamberwereplacedin astratigraphicorder;i.e.theorderofrelativetimingofthegeologicalunitsasinterpretedby theresearchers.Usingthisstratigraphy,geologicalunitsbelowandabovethefossilremains weretargetedfordating,sothattheageoftheunitcontainingthenaledifossilscanbe bracketed.Thirdly,someofthefossilmaterial,namelythreeteeth,wasdateddirectly.Allthree approacheswereconsistentwithoneanother,whichconfirmsourgeologicalinterpretations andprovidesuswithconfidenceinthefinalresults. 15b)Whatdatingmethodsdidtheteamusetodeterminethegeologicalageofthefossils fromtheDinalediChamber? Theteamappliedsixdifferentmethodstodeterminetherangeoftimewhenthefossilswere deposited.Twoofthese(radiocarbonandpaleomagnetism)donotplaceboundariesontheage ofthefossils,althoughtheyaddsomeinformationabouttheageofthesedimentsinthe DinalediChamber.ThemostinformativemethodwasESRdating(ElectronSpinResonance) directlyuponsamplesofH.nalediteeth,whichwascombinedwithU-Thdatingoftheteeth. Theteamalsomeasuredtheamountofuraniumandthoriumwithinsamplesofflowstonefrom theDinalediChamber.Whereflowstonelayerslieabovefossilhomininremains,theageofthe fossilsmustbeolderthantheflowstone.Inthisway,theminimumageofthefossilswas constrainedto236ka.Additionally,theteamfoundsometinyquartzgrainswithinsomeofthe oldersedimentsinthechamber,andusedamethodcalledOSL(OpticallyStimulated Luminescence)toestimatethelengthoftimethosegrainshadbeenburiedinadarkplacewith nolight.TheboundariesofthepossibletimethatthefossilsweredepositedcomefromtheESR anduranium-thoriummethods.Theothermethodsgiveconsistentresults,andconfirmthe interpretationofthegeneralstratigraphyinthechamber. 15c)Whydidittakesolongtoobtainadate? IthasbeenverychallengingtodatetheH.naledifossilsaccuratelyforseveralreasons. Firstly,whilstitispossibletodatethefossilsdirectly(andintheendwediddatethefossils directly),thisisadestructiveapproachwhichweonlywanttofollowinexceptionalcases,and onlyafterthefossilshadbeenfullydescribedandanalysed.Buteventhen,onlyfewdirect datingtechniquesareavailable(inthiscasecombinedU-ThandESR),andthesetechniques haverelativelylargeerrormarginsandtheyrelyonmodelassumptionssoeverythingneeded tobechecked,preferablywithindependenttechniques. Secondly,fossilscanbedatedbytargetingthesedimentsinwhichtheyareembedded.The probleminthecaseofH.nalediisthatthesedimentsarenotyetconsolidated,andthatthereis clearevidenceforrepeatedreworkingofthesedimentsincludingthefossils.Thismeansthatit hasbeenextremelydifficulttodeterminetheexactcontextofthefossils.Therewerealsoa numberoftechnicalissuesrelatingtothespecificconditionsinthecave(forexampleexcessive radonlossinthesediments,andunknownsaroundtheburialhistorybecauseofthe reworking),whichmadeithardtoacquirereliableages. Thirdly,weusedalargenumberoftechniquesandadoubleblindapproachofsomeofthe mostimportanttechniques,whichhasmeantthattheeffortsof10separatelaboratoriesin Australia,EuropeandSouthAfricahadtobecoordinated.Someofthetechniqueshavelong leadtimes,forotherscarefulcoordinationandsamplepreparationwasrequired.Thisin combinationwiththetechnicaldifficultiesinthecavemeantthatittooktimetogetgood results. 16)HowdoesESRdatingwork? Thecrystallinestructureofenamelinteethisquitestableovertime,andcanbetargetedfor dating.Afteratoothisburied,verylowlevelsofnaturalradiationintheteeththemselvesas wellasthesurroundingsedimentsandgroundwatercausesomeoftheelectronsinthecrystals tomovetoahigherenergystate,creatingsmallelectromagneticfieldeffects.Thestrengthof theseeffectsbuildsupovertime.Inthelaboratory,scientistscanmeasuretheseeffects,and usethemtoestimatethetimeatoothhasbeenburied.Thesecalculationsarecomplicatedand requireassumptionsabouttheboundaryconditionsthataffecthowthelevelsofradiation withintheteeth,andtoalesserdegreethesedimentsofthesitechangedovertime.Tohelp constraintheboundaryconditions,theUandThvaluesintheteethweremeasureddirectly, andthisinformationwasmodelledandcombinedwiththeESRcalculations.Finalagesusingthe combinedESRandU-Thtechniqueswereobtainedforamaximumandaminimumage scenario,withthetrueagefallingsomewhereinbetween.Tobesureoftheresults,the measurementswerecarriedoutonthesameteethbytwoindependentlaboratories,sothat resultscouldbecompared.Forthesereasons,therangeofuncertaintyinourestimateofthe geologicalagefortheH.nalediteethisquitelarge. 17)DotheLesediChamberremainscomefromthesametimeperiodasthoseinthe DinalediChamber? TheteamdoesnothaveanyinformationyetfromthegeologyoftheLesediChamberthatsays directlyhowoldthosefossilsare,buttheLesedifossilsareverysimilartotheDinaledifossilsof Homonaledi,justassimilarashumanskeletonsfromasinglepopulation.Thissuggeststhat bothchamberscontainindividualsthatmayhavelivedatroughlythesametime.Theteamis workingnowtodiscovertheageoftheLesedihomininfossilstotestthishypothesis. 18)WhenintimedidthespeciesHomonaledioriginate? TheindividualsofHomonaledifoundintheDinalediChamberlivedbetween236,000and 335,000yearsago.Theiranatomysuggeststhattheirancestorsdivergedfromtheancestorsof modernhumansmuchearlierintime.SomescientificresultshavesuggestedthattheH.naledi lineagemayhaveoriginatedasearlyas2.5millionyearsago,othersaslateasonemillionyears ago.Thewiderangeofuncertaintymeansthatwewillneedtodiscovermuchmoreabout otherprimitivemembersofourgenusbeforewecananswerhowlongH.naledimayhave existed. 19)WhendidHomonaledibecomeextinct? ThefossilsofH.nalediintheDinalediandLesediChambersalmostcertainlywerenotthelast H.naleditohavelived.TheevidencecannotsayhowlongH.naledimayhavesurvived,andthe teamdoesnotruleoutthehypothesisthatH.nalediwaspresentmuchlaterthan236,000 yearsago. 20)Whyisitimportanttofinda“primitive”homininatsucharecenttimeperiod? Itwasoncecommonlybelievedthatno“primitive”speciescouldhavesurvivedtheintensifying competitionfromlarger-brainedhumans,withtheiradvancedsocialandtechnological behaviours.Recentdiscoveriesshowthatthisviewisfalse.Adiversityofhomininspeciesand populationsexistedthroughmuchofthelasttwomillionyears.Verydifferenthomininforms, somelikeH.nalediandH.floresiensiswithsmallbrainsizes,somewithcloserconnectionsto humanssuchasNeanderthalsandDenisovans,continuedtoexist.H.nalediisimportant becauseitshowsnotonlythesizeofthebrain,butalsomanyotheraspectsofskeletal adaptationswerediverse. 21)Withsucharecentgeologicalage,isitpossiblethattheHomonaledifossilscontain DNA? TheteamhasattemptedtoobtainancientDNAfromtheDinalediChamberfossilsbutsofarhas hadnosuccess. 21b)Theagerangeof236-335kaisstillverylong.Willyoubeabletoconstrainthisage furtherinfuture? Yes,workintheDinaledichamberison-going,andasmoregeologicalunitsarebeingdated,the ageofthefossilswillnodoubtbefurtherconstrained. 21c)Youindicatethatthatthedatingwasdoneusingdouble-blindexperiments.What doesthatmean? Toensurethattheageresultsthatwehavepublishedarerobust,wehadseverallabs independentlydatethesamesamplesusingthesametechniques.Thiswasdonewithoutone labknowingtheresultsfromtheotherlabs.Weusedthisapproachforthetechniquesthat weremostcriticalforobtainingadateforthefossils,namelyU-ThandESR.Theresultsfrom thedifferentlabsshowedagreatdealofconsistency,meaningthattheageresultsare reproducibleandthereforerobust.Sucharigorousapproachhasrarelybeenusedbeforein datingfossildeposits. HOMONALEDIANDHUMANEVOLUTIONINSUBEQUATORIALAFRICA 22)WhyisthesubequatorialregionofAfricaimportant? Duringthepastmillionyearsormore,theregionsouthofthetropicalrainforestsofAfricahas beenhometovastandcontiguousareasofgrasslands,savannas,andopenwoodlands,where homininsmayhavelived.Likemanyotheranimalspeciesthatliveinsuchhabitats,modern humanshavetheirhighestdiversityinsubequatorialAfrica.ThediscoveryofHomonaledi suggeststhatthehominindiversityinthisregionwasevenhigherinthepast. 23)IsHomonaledia“relictspecies”? Arelictspeciesisonethatremainsafteramorediversearrayofitsancientrelativesbecomes extinct.FossildiscoveriesandgeneticevidenceshowusthatH.naledilivedatthesametimeas manydiverselineagesofhominins,bothwithinsubequatorialAfricaandinotherpartsofthe world.Today’speoplearethelastsurvivorsofthisancientdiversity.Inthatsense,modern humans(H.sapiens)arearelictspecies.ItisnotclearwhetherH.naledialsohadmanyclose relativesthatearlierbecameextinct,sowecannotsayifitwasarelict. 24)DoesHomonaledishowthatSouthAfricawasakindofbiological“culdesac”, isolatedfromthemainstreamofhumanorigins? No.Africasouthoftheequatorisoneoftheworld’smostbiologicallydiverseregions.During thelasttwomillionyears,thesavannaandopenwoodlandhabitatsforhomininswerelarger andmoreconnectedsouthoftheequatorthaninnortheasternAfrica.Agreaterdiversityin subequatorialAfricaisevidentinthegeneticsofmanyanimalspeciesthatliveinthesehabitats today.H.naledinowhelpstoconfirmthatdiversehomininpopulationsalsothrivedintheareas ofAfricasouthoftheequator.Farfrombeingisolated,thisareanowlookslikethemainstream. 25)DoesthesurvivalofH.nalediindicatealongperiodofevolutionaryisolation,likethat suggestedforH.floresiensis? No.WhileFloresisanislandwithauniqueanimalcommunitythatevolvedinisolation,thisarea ofAfricasharesmostofitsanimalspeciesortheircloserelativeswiththegrasslands,savannas andopenwoodlandsoftherestofthecontinent.H.nalediisnotaproductofevolutionary isolation. 26)WhereelsedidHomonaledilive? TheanatomyofH.naledisuggestsitmayhavebeenwide-ranging,usinglandscapesintheway thathumanshavedone.ButasyettheteamhasfoundevidenceofH.naledionlyinthesetwo localities,andwehavenoevidenceaboutthefullgeographicareaitmayhaveoccupied. 27)DothesediscoveriesshowthatthegenusHomoevolvedinSouthAfrica? Itistoosoontosaywhereourgenusmayhaveevolved.Tenyearsago,itwascommontohear scientistsassumethatthegenusHomomusthaveevolvedinEastAfricaortheGreatRiftValley, becauseoftheimportantfossildiscoveriesfromthatregion.Today,theprimitivenatureofH. nalediandtheaccumulatingevidenceofdiversityofhomininswithinsubequatorialAfricanow suggestthatthisdiverseregionmayhavebeenimportanttoourorigins.Butstill,fossil evidencehascomefromonlyverysmallareasofAfrica.Onlymoreexplorationcananswerthis question. DELIBERATEBODYDISPOSAL 28)HowdidHomonaledigetintothetwochambers? Afterintensiveexplorationandinvestigation,theteamhasfoundnoevidencetoexcludethe hypothesisthatH.naledideliberatelydepositedbodiesintheDinalediChamber.TheLesedi Chamberdiscoveryaddsyetanotherinstance,similarandformandcontenttotheDinaledi Chamber.ThisappearstopresentfurtherevidencethatH.nalediwasusingtheRisingStarcave systemforarepeatedbehavior. 29)Didallthefossilhomininsdieatthesametime,andwastheresomesortof catastrophe? TheoccurrenceofH.nalediskeletalremainswithintwochambers,farseparatedfromeach otherwithnodirectundergroundconnectionandnoeasyconnectionfromthesurface,is strongevidenceagainsttheideathattheywerevictimsofanysinglecatastrophicevent. 30)DidHomonalediburybodiesintheLesediChamberaswellastheDinalediChamber? TheevidencesuggeststhatH.naledimayhaveusedboththeLesediandDinalediChambersina similarway,asplaceswheretheydepositedtheirdead.Atsomeothersites,carnivoreskilled andatethehominins,orthehomininsfellintoa“deathtrap”,ortheirremainswerecarriedor relocatedbywaterandgravitywithincaves.TheevidencefromtheLesediChamberrulesout allofthese,justastheDinalediChamberevidencerulesthemout.Itisareasonablehypothesis thatHomonaledithemselvesenteredthecaveandusedthesechambersfordepositingtheir dead. 31)Wasthecaveequallydifficulttoaccessinthepastasitistoday? TheteamdoesnotyetknowhowdifficultitmayhavebeentoreachtheDinalediorLesedi ChambersatthetimeH.naledilived.TheevidenceshowsthattheDinalediChamberwas alwaysverydifficulttoenterandseparatefromneighboringchambers,andboththeDinaledi andLesediChamberswerealwaysinthedarkzoneofthecave.However,someofthecurrent squeezesandphysicalconstraintsthroughthecavesystemmayhavebeendifferentinthepast. Butnotethatwiththerelativeyoungageofthefossils,theundergroundgeometryofthecave systemswillnothavechangedasmuchaswouldhavebeenthecaseifthefossilshadbeenold. 32)DidHomonaledihavefire? ItisareasonablehypothesisthatH.naledimusthavecontrolledfireinordertorepeatedlyuse areasdeepwithintheRisingStarcavesystem.However,theteamhasnotyetconfirmedany physicalorchemicalevidenceofancientfiresineithertheDinalediorLesediChambers. ControlleduseoffireisknownfromothersitesinSouthAfricainexcessof1millionyearsago, includingSwartkranswhichisjust800metersfromtheentranceoftheRisingStarcavesystem. HOMONALEDIANDOTHERSPECIES 33)WheredoesHomonaledifitwithinthehumanlineage? StudiesoftheskullandteethhavesuggestedtwopossiblescenariosfortherelationshipsofH. naledi.Thespeciesmayhaveemergedneartheoriginofourgenus,Homo,aroundthesame timethatH.habilis,H.rudolfensisandH.erectusfirstexisted.ThiswouldmakeH.naledioneof aninitialradiationofourgenus,orpossiblyanancestororcloserelativeofH.erectus.Asecond scenarioisthatH.naledimayactuallybemorecloselyrelatedtomodernandarchaichumans, makingH.erectusamoredivergentbranchoftheHomofamilytree.Theevidencedoesnotyet allowustoexcludeeitherofthesepossibilities. 34)DoestherecentgeologicalagemeanthatHomonaledicannotbeanancestorofHomo erectusorHomosapiens? SomepopulationsofH.naledilivedbetween235,000and336,000yearsago,butother populationsmayhavelivedmuchearlier,assuggestedbythemanyfeaturesthatH.naledi shareswithmuchearlierfossils.Itispossiblethattheseearlierpopulationsmayhavegivenrise toeitherH.erectusorH.sapiens,orthesespeciesmayhaveemergedfromcommonancestors. 35)CanHomonaledishedanylightonthatotherrecent,controversialfossilspecies, Homofloresiensis? BothH.nalediandH.floresiensiswereprimitivespeciesthatexistedintotherelativelyrecent past.TheteamworkingonH.nalediincludessomeexpertswhohavedirectlystudiedH. floresiensis,andtheteamhasexaminedtheirsimilaritiesanddifferences.Mostly,thefeatures thataresimilarbetweenthetwoareprimitiveanddonotindicateacloserelationship,while eachofthesespecieshasareasofderivedanatomydifferentfromtheother. 36)IsHomonaledia“primitiveHomoerectus”? No.AfewoutsideexpertsmadepublicstatementsimmediatelyaftertheannouncementofH. nalediin2015thatthefossilswerenottrulyanewspeciesbutinsteada“primitiveHomo erectus”.Theseclaimshavenotheldupunderscientificexamination.Sincethatoriginal descriptionwaspublishedlessthantwoyearsago,hundredsofpagesofpeer-reviewed scientificresearchhasbeenpublishedontheDinalediChamberfossils,bothbytheoriginal teamandbyotherindependentresearchers.Allofthisscientificresearchsupportstheteam’s conclusionthatH.nalediisauniquenewspecies.Noscientificarticleshavebeenpublishedthat questiontheteam’sdescriptionofH.nalediasanewspecies. HOMONALEDIANDTHEARCHAEOLOGICALRECORD 37)DidHomonalediusestonetools? TheteamhasnotfoundstonetoolstogetherwiththeremainsofH.naledi.Studiesofitshands suggestthatH.naledihadtheanatomyofacapabletoolmaker.Stonetoolsaremuchmore commonthanfossilhomininremains,andareknownfromsouthernpartsofAfricafrom roughlythesamerangeoftimethatH.naledilived.Butitisnotclearwhichhominin populationsmadethosetools. 38)DidHomonaledicreatetheassemblagesthatarchaeologistshavepreviouslyascribed tomodernhumans? Thestonetoolsthatarecommonduringtheperiodbetween236,000and335,000yearsagoin southernAfricaareattributedtothe“MiddleStoneAge”(MSA)setoftraditions.Archaeologists haveoftenassociatedtheMSAtomodernhumansandtheirimmediateancestors.However, veryfewmodernhumanfossilremainsareindirectassociationwithMSAinthisregion,and nonefromtheearlyMSAtimeofH.naledi.Untilstrongerevidenceplacingfossilhominin remainstogetherwithstonetoolassemblagesisdiscovered,wewillnotknowwhichhominin populationsmayhavemadethesestonetools. 39)DidHomonaledimeetmodernhumans? TheevidencedoesnotsaywhetherHomonaledimetmodernhumans(Homosapiens).The earliestfossilremainsofmodernhumansarethosefromtheOmoKibishregionofEthiopia, nearly200,000yearsold.Nomodernhumanfossilremainsareknownfromsubequatorial Africaasearlyasthis.ItispossiblethatsomepopulationsofHomonaledicameintodirect contactwithmodernhumansortheirancestors,butwehavenoevidenceatthistime. 40)DidHomonaledimeetarchaichumans? ThefossilrecordsuggeststhatsomearchaichumanpopulationsexistedinsubequatorialAfrica duringmuchoftheexistenceofH.naledi.ThisevidenceincludestheKabweskeletalremains fromZambia,andtheFlorisbadskullfromSouthAfrica.Itisnotclearwhetherthese populationslivedinthesameareasasthoseoccupiedbyH.naledi,whethertheywereindirect contactwitheachother,orwhethertheymayhavemadesimilarordifferentstonetool traditions. INTOTHEFUTURE 41)WhatadditionalresearchisbeingdoneonHomonaledi? TheteamcontinuestocarryoutresearchinthelaboratoryandintheRisingStarcavesystem. SincetheinitialannouncementofH.nalediin2015,morethan400pagesofpeer-reviewed researchhavebeenpublishedbytheteam,providingmoredetailedanalysisoftheanatomyof mostpartsoftheH.nalediskeleton,moredetailonthegeologicalandtaphonomiccontextof theDinalediChamber,andtestinghypothesesabouttherelationshipofH.naleditoother homininspecies.Thatscientificproductivityisbeingcarriedforwardwithmorepaperssetfor publicationthisyear,andnewresearchunderwayontheLesediChamberremains. 42)WhataretheprospectsforfindingmoreevidenceofHomonaledifromothersites? TheteamhasbeensurveyingmorepotentialfossilsitesintheCradleofHumankind,andthere areexcellentprospectsfordiscoveringmorenewhominin-bearingcaves.Wehopetomakean announcementofexcitingnewnaledifossilsverysoon.Inaddition,itispossiblethatfragments ofH.naledihavealreadybeenfoundatsitesinotherpartsofAfricabutnotrecognized.Areexaminationoffossilevidenceacrossthecontinentmayprovidemorediscoveries.
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