Volume 18
January 2005
Number 1
T H I S M O N T H'S G U E S T S P E A K E R
Taylor Edwards
University of Arizona
Genomic Analysis and Technology Core
In Search of the World's Largest Frog
7:15 PM
Tuesday, January 18th
Arizona Game and Fish Department Office
555 North Greasewood Road
(between Speedway and Anklam, west of Pima Community College)
Taylor Edwards received his BA in zoology from the
University of California, Santa Barbara, in 1992 and soon
afterwards moved to Tucson where he worked in the
Department of Mammalogy and Ornithology at the
Arizona-Sonora Desert Museum. In 2003, he received his
master’s degree in Wildlife Ecology at the University of
Arizona, School of Renewable Natural Resources studying
desert tortoise conservation genetics under the guidance of
Dr. Cecil Schwalbe. Taylor is an active member of the
Tucson Herpetological Society and served as President in
2003-2004. He currently works in the Genomic Analysis
and Technology Core at the University of Arizona. Taylor’s
main interest is conservation biology and his research has led
him as far off as China and Brazil, but his focus has mostly
been in the desert Southwest he calls home. In 2004,
Taylor visited Dr. Hans-Werner Herrmann and Patricia
Hermann in Cameroon to assist on a project on the world’s
largest frog, Conraua goliath. He will talk about his adventures there and how the project has found its way back to
Arizona.
N E X T M O N T H'S G U E S T S P E A K E R
Goliath Frog (Conraua goliath) caught near Mkonine, Cameroon, 23
February 2004. Weight=3.22 kg (114 oz), snout-vent lenth=330 mm
(13 in).
Charles H. Lowe, Jr. Herpetology
Research Fund
Proposals are being accepted from now until March 1 for
the Tucson Herpetological Society’s Charles H. Lowe, Jr.
Herpetology Research Fund. This fund is for support of
research that contributes to the conservation of the
herpetofauna of the Sonoran Desert. Any current THS
member is eligible. Requests can be up to $500. For further
information, please see the detailed fund information on
page 8 of this issue of the Sonoran Herpetologist.
Roy Averill-Murray
Scientific Assessment of the 1994 Desert Tortoise Recovery Plan
Tuesday, February 15th
Tucson Herpetological Society meetings are open to the public and are held on the third Tuesday of each month starting at 7:15PM
SONORAN HERPETOLOGIST 18(1) 2005
1
B O O K R E V I E W
Iguanas: Biology and Conservation
Allison C. Alberts, Ronald L. Carter, William K. Hayes
and Emilia P. Martins, editors. 2004. University of
California Press, Berkeley, California. xvi + 356pp.,
hardcover. US$ 65.00. ISBN 0 - 520 - 23854 - 0.
James L. Jarchow
Tucson, Arizona
[email protected]
The family Iguanidae includes many of the planet’s
largest, most visually impressive, and (unfortunately)
most endangered lizard species. The iguanas are
notable not only for their large size but also for their
herbivory, island endemism, and intriguing behavior.
This volume is a comprehensive cumulation of recent
studies in iguana biology which covers each of these
areas as well as current and proposed management and
conservation efforts, the latter pertaining to West
Indian species. It is the product of a 1997 Herpetologists’ League symposium, “Biosystematics, Behavioral
Ecology and Conservation of Iguanas” held at the
University of Washington. There are 42 contributing
authors, many among the most prominent names in
iguana biology.
The first chapter, by Gordon Burghardt,
provides a review of the history of iguana research with
an emphasis on behavioral ecology. Burghardt’s
writing style flows well and holds the reader’s interest;
his personal recollections of earlier contributors such as
L. T. Evans and Charles C. Carpenter are highlights.
Burghardt concludes the chapter with a discussion of
future areas of research and provides a number of
suggestions to help students prepare for these studies.
The rest of the text is divided into three parts:
Diversity, Behavior and Ecology and Conservation.
The Diversity section includes a discussion of the
taxonomic organization of the of the iguanas by B. D.
Hollingsworth. Intergeneric and interspecific (within
the genera Sauromalus, Ctenosaura, and Cyclura )
relationships proposed by various investigators using a
variety of morphological and molecular methodologies
are reviewed and suggestions for resolving disparities
are made by the author. This chapter concludes with a
current species checklist.
In the following chapter, C. L. Malone and S.
K. Davis examine nuclear and mitochondrial DNA
analytical methods and discuss their application to
determining not only the evolutionary history of
Caribbean Rock Iguanas, Cyclura spp., but also to
prioritizing conservation efforts for those island forms
that are the most genetically unique.
The two chapters that follow explore the effects
2
SONORAN HERPETOLOGIST 18 (1) 2005
of geographical separation on genetic variation among
populations inhabiting groups of islands. M. E. Welch,
G. P. Gerber and S. K. Davis examine Rock Iguanas
(Cyclura c. carinata) on the Turks and Caicos Islands of
the Greater Antilles using nuclear DNA markers, and K.
Rassmann, M. Markmann, F. Trillmich and D. Tautz
study Land Iguanas (Conolophus spp.) and Marine
Iguanas (Amblyrhynchus spp.) of the Galapagos
archipelago using mitochondrial and nuclear DNA
markers. Of particular interest is that these studies provide
compelling evidence that hurricanes may facilitate an eastto-west migration of Rock Iguanas in the Greater Antilles
and the iguanas of the Galapagos apparently predate
present islands by 5 to 9 million years. In the latter study,
mitochondrial DNA analyses suggest that Galapagos
Marine and Land Iguanas had a common ancestor which
inhabited islands in the archipelago that are no longer in
existence.
The final chapter in this section, by L. C. Hazard,
is a discussion of nasal salt gland function in iguanas. The
author provides evidence, from a review of the literature,
that herbivory and the need to excrete its associated
higher potassium loads without significant water loss
predisposed Marine Iguanas for survival in a saline
environment.
The second section, Behavior and Ecology, begins
with two chapters that explore the variation in and
complexity of head-bob displays by Turks and Caicos
Island Iguanas. E. P. Martins and K. E. Lacy describe and
thoroughly document a distinctive appeasement head
bob exhibited by juveniles and smaller females toward
larger adults, which reduced aggressive reactions by the
larger animals. A. N. Bissell and E. P. Martins examine
variations in Turks and Caicos Island Iguana morphology
and head-bob displays associated with variations in
habitat among twelve different island populations.
The following chapter, by J. A. Rivas and L. E.
Levin, describes an intriguing sexual dimorphism in
antipredator behavior exhibited by juvenile Green
Iguanas (Iguana iguana). Two different experiments
reveal consistent variation in the way male and female
juveniles respond to predators and a greater likelihood of
females surviving predation, respectively. In the first
experiment, males in a group of juveniles reacted to a
simulated falcon with activity which seemingly would
divert a predator’s attention from females, which
generally remained still or hidden. In some cases, males
reacted by covering females with their bodies, apparently
supporting a contention of kin selection and altruistic
behavior. The following chapter, by W. K. Hayes, R. L.
Carter, M. Wikelski, and J. A. Sonnentag, is an observational study of lek mating success of male Galapagos
Marine Iguanas (Amblyrhynchus cristatus) on Caamano,
an islet near Isla Santa Cruz. The authors describe that
like seals, male Marine Iguanas form congregations, or
“leks”, on rocky shorelines during the mating season.
Within these leks, males defend small, symbolic territories
while displaying to attract females. The physical and
behavioral characteristics of 213 lekking males were
compared using bivariate and multivariate analyses to
determine which qualities attracted the most females.
Interestingly, females selected males with the fewest ticks
along with the more predictably wide heads and
vigorous territorial displays.
This section concludes with three chapters
devoted to studies of body size and growth rate. M.
Wikelski and C. Carbone develop a mathematical model
to predict the body size of Marine Iguanas based on the
constraints of ambient temperature range, energy intake,
expenditure, and body mass. C. R. Tracy compares body
size variation among populations of two North American
iguanids, the Common Chuckwalla (Sauromalus ater)
and the Desert Iguana (Dipsosaurus dorsalis). Elevation
was found to influence body size in Common Chuckwallas but not Desert Iguanas; greater rates of gene flow
between Desert Iguana populations (which occupy more
contiguous habitat) was one explanation provided for
this disparity. The last chapter in this section, by J. B.
Iverson, G. R. Smith and L. Pieper, examines the effects
of supplemental feeding by tourists on growth rates of
Allen Cays Rock Iguanas (Cyclura cychlura inornata).
Only a few aggressive individuals (mostly males) were
found to benefit from this practice; populations of this
species appear to be organized in dominance hierarchies
rather than separate territories.
The final section, Conservation, consists of seven
chapters addressing proposed and current conservation
programs to manage and increase
existing populations of Cyclura
species in the Caribbean.
Intensive procedures such as
translocation and head starting are
discussed by C. R. Knapp and R.
D. Hudson, and by A. C. Alberts,
J. M. Lemm, T. D. Grant and L.
A. Jackintell, respectively. Past
translocation efforts are described
as “highly successful” and several
examples are presented. The
authors identify prerelease health
screening and genetic origin of
translocated iguanas, and habitat
assessment with particular
attention to forage availability and
introduced predators as keys to
translocation success. An
experimental study of head
started and released Cuban
Iguanas (Cyclura n. nubila)
indicated human avoidance
behavior, acceptance of natural
forage, growth rates, and
thermoregulatory and social
behavior remained largely
unaffected by the captive
headstart experience. Unfortunately, released animals
were not monitored for survival by radiotelemetry
because they were released on the U. S. Naval installation at Guantanamo Bay.
The next three chapters describe existing
management programs for Jamaican and Bahamian
Iguanas. On Jamaica, introduced predators, especially
the mongoose, but also feral house cats, reduced iguana
population recruitment severely. Headstarting and
exotic predator control have probably prevented, or at
least delayed, the extinction of free-ranging Jamaican
Iguanas (Cyclura collei). Following the standards of
previous chapters, problem statements, management
practices, observations and results are presented in
detail and discussions are thoroughly referenced. This
section concludes with two chapters describing the role
zoos have had in providing human and physical
resources for Cyclura conservation, and the potential
benefits of ecotourism to management programs in the
future.
Although West Indian and, to a lesser extent,
Galapagos Marine Iguanas are the focus of this book,
study methods and observations are applicable to other
species as well. This volume complements the well
known Iguanas of the World: Their Behavior, Ecology
and Conservation, edited by G. M. Burghardt and S.
A. Rand and published in 1982. Like its predecessor,
the current work is an indispensable reference for those
interested in or studying iguana biology.
SONORAN HERPETOLOGIST 18(1) 2005
3
Tucson Herpetological Society Membership Directory
1 January 2005
Melissa Amarello 444 E Seneca St 85705. 245-6901 [email protected] 4,3,9-11,21-35,33 St Dec 05
Sandy Anderson PO Box 2095 Sierra Vista AZ 85636. 520-458-0742 [email protected] 4,3-13,14,1130,32,31 Ind Mar 05
Bob Applegate PO Box 338 Campo CA 91906. 619-478-5123 [email protected] 4-16-G Ind May 09
Arizona Game & Fish Department Nongame Amphibian & Reptile Program 2221 W Greenway Rd Phoenix AZ
85023. Ind Sep 05
Arizona Poison & Drug Information Center PO Box 245079 85724-5079. 626-6016 Life
Roy & Laurie Averill-Murray USFWS Nevada Fish & Wildlife Service 1340 Financial Blvd. #234 Reno NV 89502.
775-861-6362 [email protected] 9,1-14,13,19-32,34,37 Fam Aug 05
Royce Ballinger 13584 N Holly Grape Dr Marana AZ 85653. 520-572-5973 [email protected] 3,9,814,15,10-33,35,34 Ind Mar 05
Dan Beck & Kris Ernest Dept Biological Sciences Central Washington Univ Ellensburg WA 98926. [email protected]
509-963-2886 9,8-14,15,11-33,35,31 Life
Daniel Bell 5725 E 4th St 85711. 750-0770 9,8-19,13-33,35 Ind Aug 05
Emily Bennett 20 Permian Dr Carlsbad NM 88220. 505-785-3185 [email protected] 3,4-14,11,15-30,32,31 Su
Nov 05
David Bertelsen 2503 E Blacklidge Dr 85716. 419-1001 [email protected] G-G-1,2,3 Su Feb 05
Kit Bezy 901 S O’Leary St Apt 104 Flagstaff AZ 86001-6454. [email protected] 8,9-0-0 St Dec 05
Robert Bezy
Bezy* 3776 E Nugget Canyon Pl 85718-2341. 615-8862 [email protected] 9,8-20,15-30,31,32 Ind Dec 05
William Black PO Box 2004 St Cloud FL 34770. 0-0-0 Ind Apr 05
Kevin Bonine
Bonine* & Angela Urbon-Bonine 5762 E Bellevue 85712. 751-1349 [email protected] 3,6,915,19,13-32,33,35 Su Feb 05
Robert Brandner & Susan Basford 4241 N Houghton Rd 85749. 760-0574 [email protected] 9,4,1-14,11,1334,36,37 Con Feb 05
Bayard Brattstom PO Box 166 Wikieup AZ 85360. 928-716-4574 3,4,1-11,14,13-32,41,33 Con Mar 05
Debbie Brewer 2121 W Main St Apt 3009 Mesa AZ 85201. 480-610-0457 [email protected] 9,814,13,12-30,32,36 Ind Feb 05
Mark Brock 5015 E Sagebrush St Apache Junction AZ 85219. 480-288-4506 [email protected] 3,1,916,11,19-32,30,34 Con Sep 05
Danny & Sharon Brower 4921 N Siesta Dr 85750-9601. 749-5257 [email protected] 4,3-17,15,11-33,41,37
Con Oct 05
Philip & Barbara Brown 12001 W Ranchito Verde 85743. 578-6305 [email protected] 1-10,12,13-33
Con Feb 05
Young Cage
Cage* 4519 W Cush Canyon Loop Marana AZ 85653-4431. 579-2223 [email protected] 4,3,1-14,13,1630,36,37 Ind Dec 05
Dennis Caldwell 2109 N 2nd Ave 85705. 624-0198 [email protected] G-10,13,19-G Life
Tom Caldwell 4521 N Paseo Pitiquito 85750. 529-8793 [email protected] 0-0-0 Ind Dec 05
Joshua Capps 1206 E Copper St 85719. 906-1092 [email protected] 2,1,3-13,14,11-34 Ind Sep 05
Chip Cochran 2733 N Norris Ave 85719. 904-2181 [email protected] 4,9,5-15,21,19-39.41.37 St Aug 05
George Coffee 775 14th St Boulder CO 80302. 303-449-0082 [email protected] 4,3,6-14,11,15-30,32,37 Su
Apr 05
Jay Cole & Carol Townsend 6381 W Sweetwater Dr 85745. 743-3402 [email protected] [email protected] 315,20,13-34,37,30 Con Sep 05
Joseph Collet 2330 Guardian Cr St George UT 84790. 435-313-2411 [email protected] 3,1,4-16, 10, 1233,32,30 Ind Jan 05
Paul, Steven & Lorna Condon 8832 E Buckboard Rd 85749. 749-3648 [email protected] 4,3-19,16-33,37
Con Jan 05
Justin & Nancy Congdon PO Box 1128 Douglas AZ 85608. 520-824-3121 [email protected] 2,1,3-15,1432,31,31 Fam Mar 05
John Craig 6882 E Calle Cerca 85715. 731-0833 [email protected] 9,8,6-18,14,13-32,34,36 Su Apr 05
Stephen Cumberworth 11313 Fountain Blvd Oklahoma City OK 73170. [email protected] 9,4,3-14,15,1332,30,41 St Mar 05
Brendan Cummings PO Box 493 Idyllwild CA 92549. 909-659-6053 [email protected] 1,4,713,12,14-32,33,34 Ind Mar 05
Kerrah Cutter 628 E Drachman St Apt D 85705. 792-1367 4,1,9-13,14,19-30,32,36 Ind Sep 05
Anthony Dee & Sarah Studd 1131 E Mitchell St 85719. 882-4698 [email protected] 9,8-11,13,14-33,35,37
Fam Dec 05
Willem De Reuse and Shobhana Chelliah 422 Magnolia St Denton TX 76201. 817-591-8786 [email protected]
4,3,6-20,12,13-33,38,40 Life
Marlis & Michael Douglas Dept Fishery Wildl Biol Colorado State University Fort Collins CO 80523-1474. 970-4917265 [email protected] [email protected] 0-0-0 Fam Apr 05
Stan & Emily Draper 1815 Sir James Cr Salt Lake City UT 84116-1812. [email protected] 4,1,6-16,11,19-40,36,38
Su Apr 05
Taylor Edwards* PO Box 3567 85722. 240-9280 [email protected] 3,6-13,14,15-33,32 Su June 05
Erik Enderson 2627 S Jaguar Rd 85730. 298-8599 [email protected] 4,3,9-19,13,14-32,33,34 Con Dec 05
4
SONORAN HERPETOLOGIST 18 (1) 2005
Jerrold & Martin Feldner 1918 E Redfield Rd Tempe AZ 85283-4221. 480-839-2932 [email protected] 3,413,14,20-32,33,35 Su Feb 05
George Ferguson 2677 W Highcliff Dr 85745. 299-4628 [email protected] 3-20,13-33,35 Ind Sep 05
Philip Fernandez Dept Biol Grand Canyon Univ 3300 W Camelback Rd Phoenix AZ 85017. 589-2575
[email protected] G-G-G Ind Aug 05
Carianne Funicelli 4220 E Kings Rd 85711. 320-9054 [email protected] 8,9,1-13,10,14-32,33,35 Ind Apr 05
Richard Funk Mesa Veterinary Hospital 858 N Country Club Dr Mesa AZ 85201. 480-833-7330 [email protected]
Ind Apr 05
Brooke Gebow 3937 N Camino Sinuoso 85718. 557-9357 [email protected] 8,9-13,11,14-33,35 Con Sep 05
Caren Goldberg 420 E E St Moscow ID 83843. [email protected] 6,3,4-14,13,15-34,33,30 Ind Jan 05
Stephen Goldberg 13713 Sunrise Dr Whittier CA 90602-2548. 562-698-8517 3,4,6-18,14,15-33,42,42,40 Ind Dec 05
Matt Goode 2837 Edison St 85716. 323-5003 [email protected] 4,3,8-15,14,13-34,32,33 Con Feb 05
Bethany Gray P O Box 318 Grand Canyon AZ 86023. 928-638-6450 [email protected] 6,3,7-11,15,14-33,32,34
St Jan 05
John Gray 1904 Bermuda St Apt 12 Long Beach CA 90802-3872. 562-436-5593 [email protected] 1-14,12-33 Ind
Jan 05
Randall & Anna Gray USDA NRCS PO Box 2890 Rm 6158-S Washington DC 20013. 520.432-5479
[email protected] 3,4,6-13,14,12-33,32,30 Fam Dec 05
Harry Greene & Kelly Zamudio Ecol Evol Biol Corson Hall Cornell University Ithaca NY 14853-2701.
[email protected] 4,3-15,20,11-33,34 Con Mar 05
Steve Hale 139 W Suffolk Dr 85704-7140. 742-1274 [email protected] 4,6, 3-15,14,16-31,30,35 Ind July 05
Marc Hammond 3949 E 27th St 85711. 750-9507 [email protected] 1,4,3-11,21,13-3-,33,32 Ind Dec 05
Dave & Billie Hardy Sr 585 S Main Ave 85701. 624-8879 [email protected] 4,3-14,21,19-32,36,37 Hon Life
Trevor Hare 2718 E Croyden St 85716. 321-1462 [email protected] 4-13,19,21-32,31,30 Ind Jan 05
Adam & Adam Harting 7337 E Shoreward Loop 85715-3455. 722-6634 [email protected] 4,3,5-21,11,1032,41,39 Fam June 05
Chip Hedgcock & Kim Morrison 4120 E Oxford Dr 85711. 325-4241 [email protected] G-19,14,1130,32,31 Fam Dec 05
Dave & Miriam Hyatt 6145 E San Bernardino 85715. 885-1195 [email protected] 3-16-32,41 Life
Monique Imberski 915 N Tucson Blvd 85716. 404-2529 [email protected] 9,8,4-13,14,15-30,39,38 Ind Jan 05
Ryan Isaac 3576 W Ironwood Meadows Pl 85742. 579-0322 [email protected] 4,8,9-14,11,16-30,33,37 Ind
Apr 05
Craig & Rachel Ivanyi 2433 W Placita de Ramo 85741. 883-3036 [email protected] 6,4,3-13,11,21-32,30,33.
Fam Apr 05
J. M. Jacobson PO Box 68391 85737. 4,3,9-13,16,12-30,32,34 Ind Oct 05
Jim, Linda & Mike Jarchow Tucson. 877-2626 [email protected] 8,9-18,13,16-33,35 Hon Life
Dennis & Gracie Jex PO Box 1592 Tubac AZ 85646. 520-398-0812 [email protected] 4,3,6-13,16,19-32,30,33
Fam Jan 05
Ernest Jimenez III 4393 W Placita de Christina 85741. 405-9990 [email protected] 4,9,8-21,18,13-32,30,33 Ind Apr
05
Gary Johnson 193 Marian Dr Louisville KY 40218. 502-491-1593 [email protected] 4,2,3-12,11,1934,30,31 Ind May 05
Hill Johnson 5910 N Placita Tecolote 85718. 797-7089 4,9-16,12,13-30,34 Su June 05
Janice Johnson 1800 N Avenida Azahar 85745. 743-3714 G-16,11,12-G Ind June 05
Joel Johnson 13751 Gragston Circle Tampa FL 33613. 405-325-7669 [email protected] 3,9,3-14,13,15-33,34,35 St Jan
05
Cristina Jones & Joseph Beals 962 N Chrysler Rd 85716. 320-1622 [email protected]
[email protected] 3,8,1-16,13,14-30,36,37 Fam July 05
Larry Jones Coronado National Forest 300 W Congress 85701. 670-4864 [email protected] 7,4-11,14,13-30,33,32
Ind Feb 05
Leah Rae Jongsma 12035 E Barbary Coast Rd 85749. 975-1118 [email protected] 9,4,6-16,12,19-33,30,34
St Feb 05
Robert Kane 322 W 57th St #34Q New York NY 10019-3712. [email protected] 3-11-33 Ind July 05
Hans Koenig 7536 E Placita de la Poesia 85750. 577-0089 [email protected] 4,3,1-13,12,16-30,33,39 Ind Oct 05
Norma & Abe Lackow 328 E Desert Tree Dr 85704-7043. 742-7561 4,3-14,13-32,33,34 Fam Mar 05
Travis LaDuc Texas Natural History Collection 10100 N Burnet Rd PRC 176/R4000 Austin TX 78758-4445. 512-2915272 [email protected] 4,9,3-14,20,21-33,30 Ind Oct 05
Kelly Lasater 3823 Oxbow Village Albuquerque NM 87120. 505-463-8182 [email protected] 4,9,8-15,14,19-33,38,39
St Feb 05
Lainie Levick Tucson 760-2494 [email protected] 6,3,4-13,14-32,33,30 Ind Nov 05
Felicia Lincoln Kennett Square PA. 0-0-0 Life
Marcia Lincoln & Victor Ong 6701 N Waycross Rd 85743. 744-4211 [email protected] 4-16-G Fam Jan 05
Robin Llewellyn 2219 E Eastland St 85719. 622-2747 [email protected] 8,9,2-13,14,11-30,32.40 Ind Aug 05
Pat Lopez PO Box 1784 Lakeside AZ 85929. 928-368-4443 [email protected] 3,4,6-13,14-30,32,39 Ind May 05
Jay Loughlin 5051 N Sabino Canyon Rd Apt 1216 85750. 529-8064 [email protected] 3-15,20-32,30,33 Ind
Feb 05
Joseph Marek Jr 10411 Balsamwood Court Laurel MD 20708-3176. 410-792-9840 9,8,4-16,19,14-36,37,35 Ind Sep
05
Brent Martin 501 E Squirrel Trail 85704. 320-9336 [email protected] 0-0-0 Ind Feb 05
SONORAN HERPETOLOGIST 18(1) 2005
5
Tom & Judy McDonald 12356 N Echo Valley Dr Oro Valley AZ 85737. 219-5072 [email protected] 9,821,13,19-32,30,34 Fam July 05
Hugh & Rebecca McCrystal 9140 W Henry 85735. 883-0905 [email protected] 4,9-11,15,20-33,35,37 Fam Oct
05
Danny Mello 5209 San Luis Ave Santa Rosa CA 95409. 707-538-4592 [email protected] 3,4-21,16,14-32,30,33 Ind
Nov 05
Gay Miller 1801 E Spring St 85719. 326-2127 [email protected] 3,4-11,14,15-32,35 Life
Michael Mittleman 5 Damin Circle Saint James NY 11780-1603. 516-361-8606 9,8-16,12,11-G Life
Edward Moll 1440 W San Annetta Dr 85704. 742-6525 [email protected] 2,4,3-13,14,15-40,34,30 Ind Nov 05
Jeff Moorbeck & Jennifer O’Leary 2820 N 75th St Milwaukee WI 53210. 414-774-1739 [email protected]
4,3-19,11,13-34,30,41 Su Apr 05
Douglas & Laurie Moore PO Box 2248 Cortaro AZ 85652-2248. 682-0459 [email protected] 4,8,3-13,10,1530,35,36 Fam Oct 05
Robert Moses 2179 Tahoe Pl Sierra Vista AZ 85635. 520.458.3510 [email protected] 9,4,3-14,13,12-30,32,33 Ind
July 05
Steve & Maxine Murray 11050 E 29th St 85748. 885-5800 [email protected] 1-16,18,11-G Con Feb 05
Erika Nowak 9075 W Antoinette Way Flagstaff AZ 86001. 520-556-0546 [email protected] 8,9-14,13,11-1 Ind
Dec 05
Lee Oler 207 W Dahil Rd 85705. 791-9246 [email protected] 1,2-13,15,18-30,32 Ind Apr 05
Elissa Ostergaard
Ostergaard* 1344 W Delaware St 85745. 792-4310 [email protected] 6,3,4-13,14,15-32,30,34 Ind Sep 05
Dennis Pepe 1029 E Halcyon Rd 84719. 743-7434 [email protected] 4,3,1-12,13,14-33,32,30 Ind Mar 05
Trevor Persons PO Box 365 Brewer ME 04412. 928-853-7599 [email protected] 0-0-0 Ind Jan 06
Stéphane Poulin & Yvonne Gubersky 3828 N Robert Daru Dr 85745. 292-3102 [email protected] 3.414,16,19-32,30,41 Fam Dec 05
Andrew Price 3405 Aldwyche Dr Austin TX 78704. 512-912-7022 [email protected] 3,4-14,15,16 Ind
Nov 05
Dave Prival 71 E 13th St Apt 1 85701. 623-8317 [email protected] 4,5,6-13,14,15-30,35 36 Ind Oct 05
Phil Pugliese 8338 Sarnoff Pl 85710. 298-8310 [email protected] 4,3,9-21,15,18-30,33,31 Su Jan 05
Ali Rabatsky 307-B Dulles Dr Lafayette LA 70506. 337-357-6494 [email protected] 4,1,3-15,14,13-35,33,34
St Dec 05
Jenna Ramsey 1231 25th St #22 San Diego CA 92012. 619-501-7840 [email protected] 1,2,3-13,14,1132,41,39 St June 05
Charles Rau PO Box 195 Portal 85632. 558-2356 [email protected] 4,3-14,19,15-30,33,35 Ind Mar 05
John Reiss 4899 S Quarry Rd Bayside CA 95524. 707-825-0465 [email protected] 867-15,14,20-32,34 Ind Dec 05
Roger* & Dianna Repp 9044 N Valgrind Ln 85743. 318-8210 [email protected] 4,1,3-11,13,16-32,34,35 Hon Life
Jim Rorabaugh 5819 N 77th Dr Glendale AZ 85303. [email protected] 0-0-0 Ind Feb 05
Philip Rosen School of Renewal Natural Resources Univ of Arizona 85721. 319-0470 [email protected] 9,814,13,15-G,31,33 Hon Life
Manny Rubio 9925 W Ina Rd 85743. 682-2988 [email protected] 1,9,8-14,19,12-33,35 Ind Oct 05
Michael Samuels 3113 E First St Su C 85716. 323-9912 [email protected] G-16-G Ind Sep 05
Bill Savary & Jillian Cowles PO Box 815 Vail AZ 85641. 647-3320 [email protected] [email protected]
1,3-14,16,21-32,37 Su Feb 05
Justin & Li Shen Schmidt 1961 W Brichta 85745-1812. 884-9345 [email protected] 4,3,6-21,11,15-34,36
Fam Jan 05
Walter Schoepfle 6297 S Mesa Vista Dr Gold Canyon AZ 85218. 760-9469 [email protected] 4,3,6-14,15,1337,30,31 Ind Aug 05
Michael Schroff 3547 E Flower St 85716. [email protected] 4,5,1-13,14,11-30,32,33 Con Mar 05
Jerry Schudda 8132 N Modena Dr 85741. 744-4274 [email protected] 4,3,9-19,16,10-32,31,36 Ind May 05
Gordon Schuett 5144 W Sweetwater Ave Glendale AZ 85304. 602-938-0562 [email protected] 4-14,11,1532,33 Ind Mar 05
Cecil & Carol Schwalbe 4330 S Escalante Ridge Pl 85730-5125. 290-8392 [email protected] 8,9-13,14,1930,31,32 Hon Life
Chris Scott Tucson. 0-0-0 Ind Apr 05
Jeffrey Servoss US Fish & Wildlife Service 3421 W Royal Palm Rd #103 Phoenix AZ 85021. 602-242-0210
[email protected] 4,3,9-14,13,19-30,31,35 Ind Sep 05
Frank Slavens PO Box 30744 Seattle WA 98103. 206-542-6751 [email protected] 0-0-0 Ind Aug 05
Michael & Mary Ann Smith 6816 E 45th St 85730-2214. 748-2705 [email protected] 8,9-14,16,1532,30,31 Su Dec 05
Russ & Karen Solsky 14060 E Copper Mesa Ct Vail AZ 85641. 207-5844 [email protected] 4,3,1-16,21,11-30,32,33
Fam July 05
Michael Sredl AGFD 2221 W Greenway Rd Phoenix AZ 85023. 602-789-3518 [email protected] 6,3,4-13,14,1130,32,33 Ind Jan 05
Eric Stitt
Stitt* 5409 E Willard St 85712. 319-8934 [email protected] 1,8,9-13,14,15-33,34 St Jan 05
Rolf Strom 728 Remuda Dr Glendora CA 91740. 626-335-2273 [email protected] 4,1,9-14,11,20-34,30,33 Ind Apr
05
John Sullivan 873 Cedar St Pacific Grove CA 93950. 831-333-9620 [email protected] 3,9-19,12,14-33 Su Jan 05
Don Swann 5725 E 4th St 85711. 750-0770 [email protected] 2,6,1-13,14-30,34,41 Ind Jan 05
6
SONORAN HERPETOLOGIST 18 (1) 2005
Allison Titcomb & Chuck Fellows 1811 N Hitching Post Rd 85745. 629-9381 [email protected] 4,3-1132,30,31 Su Mar 05
Marty Tuegel
Tuegel* PO Box 1009 85702-1009. 444-3330 [email protected] 2,1,8-14,11,13-30,33,34 Con July 05
Dale Turner & Julia Fonseca 315 E Elm St 85705. 792-2690 [email protected] 3,4,8-13,14,15-32,33 Fam Oct 05
Daniel Turner Tucson 0-0-0 Life
Timothy Van Hoesen 1005 N Del Valle Pl 85711. 745-1005 [email protected] 3,4-11,14,16-30,37,41 Ind Apr 05
James Verdi PO Box 32 Del Mar CA 92014. 858-499-7607 [email protected] 3,9,8-13,11,14-30,32,33 Con May 05
Robert Villa* 1125 E Missouri St 85714. 741-0087 [email protected] 1,2,4-13,12,16-34,39,41 St Feb 05
Richard & Margie Wagner 11662 E Twin Hills Trail 85748. 751-8031 [email protected] 4,8,9-19,21,1133,35,37 Con Jan 05
Christopher & Elizabeth Warren 4681 N Avenida Ronca 85750-6322. 498-1809 [email protected] 3,2,6-11,13,1432,31,30 Fam May 05
Robert Weaver Dept Biol Sci Central Washington Univ 400 E University Way Ellensburg WA 78926-7537. 509-9257844 [email protected] 4,3,7-14,15,13-35,34,36 St Apr 05
Marvin Weitzel 4357 E Ware St 85711. 750-0725 2,1-16-30,33,34 Life
Cyndy Wicker & Kent Jacobs PMB 283 9121 E Tanque Verde Rd #105 85749-8874. 749-1419
[email protected] 9,1,4-13,11,21-30,32,31 Con Apr 05
Thomas Wiewandt PO Box 5118 85703. 743-4551 9,8-19,14-32,39 Ind July 05
Fred Wilson 1641 N Coral Bells Dr 85745. 743-3920 [email protected] 4,9-16,19,20-31,41,30 Life
Betsy Wirt & Peter Holm 1620 Hoover St Ajo AZ 85321. 520-387-3896 [email protected] G-G-30,31,32 Con
July 05
Blair & Deanna Wolf 13504 Red Yucca Ave NE Albuquerque NM 87111. 505-796-0240 [email protected] 3,4,614,15,11-33,35,36 Fam Dec 05
Chris Wolner 3344 E Bellevue Apt B 85716. 400-3905 [email protected] 3,4,1-13,11,14-32,30,35 Su Feb 05
Bill & Beth Woodin 3600 N Larrea Ln 85750. 0-0-0 Fam Jan 05
Lori & Jesse Jones Woods 1745 E River Rd Su 101A 85718. 325-9977 [email protected] 8,9-10,12,13-33,36,41
Su Oct 05
Brian & Minerva Wooldridge 2626 N Shannon Rd 85745-1017. 299-2303 [email protected]
[email protected] 4,3-15,13,21-30,33,34 Fam May 05
Steve Zimmerman 1011 S 22nd Ct Renton WA 98055. [email protected] 4,9,8-11,14,19-34,33,30 Ind Feb 05
Richard & Frances Zweifel Box 16354 Portal AZ 85632. 558-2236 [email protected] Fam Sep 05
Erin Zylstra
Zylstra* 2813 E Allen Rd 85716. 975-3578 [email protected] 1,4,3-11,19,14-32,39,33 Ind Sep 05
* Officer or Director for 2005
Membership code: Ind= individual, Fam= family, St= student, Su= sustaining, Con= contributing, Life= Life, Hon Life=
Honorary Life
Style explanation for address: double space between lines and no punctuation; for Tucson, ZIP Code only.
EXPLANATION OF CODE USED FOR MEMBER’S INTERESTS
Member’s interests are shown at the end of each entry. Coded by number, interests are divided into three categories and
separated by two hyphens. Members could rank up to three interests with the first number having the highest rank, the
second less rank and the third, still less. Some gave only one or two interests in each category and a few did not list any
(indicated by a “0”). Refer to the categories and their corresponding numbers given below as they appeared on the
Membership Form.
ANIMAL CATEGORY:
1-Tortoises, 2-Turtles, 3-Lizards, 4-Snakes, 5-Crocodilians, 6-Frogs and toads, 7-Salamanders, 8-Amphibians, 9Reptiles, 0-None listed
AREAS OF INTEREST:
10-Art, 11-Behavior, 12-Books, 13-Conservation (primarily), 14-Ecology, 15-Evolutionary biology, 16-Captive
maintenanceand propagation, 17-Molecular biology, 18-Pathology, 19-Photography, 20-Systematics, 21-Venoms and
envenomation
GEOGRAPHICAL AREAS:
30-Arizona, 31-Sonora, 32-Sonoran desert, 33-Southwestern U.S., 34-North America, 35-Mexico, 36-Central America,
37-South America, 38-Europe, 39-Africa, 40-Asia, 41-Australia, 42-South Pacific
¬
SONORAN HERPETOLOGIST 18(1) 2005
7
Charles H. Lowe, Jr. Herpetology
Research Fund
The Charles H. Lowe, Jr. Herpetology Research Fund
was established to support research that contributes to
the conservation of the herpetofauna of the Sonoran
Desert, including the states of Arizona, southern
California, Sonora and on the Baja California peninsula
and gulf islands. Dr. Cecil R. Schwalbe spearheaded the
fund in honor of the many contributions to our
understanding of herpetology in the Sonoran Desert by
Dr. Lowe. The fund was inaugurated at the Current
Research on the Herpetofauna of the Sonoran Desert II
Conference, April 2002.
Eligibility
Any current THS member is eligible to receive awards
from the C.H. Lowe Research Fund. Researchers need
not be affiliated with an institution and need not have
previous experience. Pre K-12 educational institutions
and students are encouraged to apply. THS Board
Members and Lowe Fund Committee Members are not
eligible.
Funding
Beginning in 2004, proposals will be reviewed annually
on 1 March. Decisions to award will be made on the
value of the research being proposed and not on the
experience or status of the person making the request.
No requests will be approved that involve collecting
animals for personal collections or profit. Requests are
not to exceed $500. Although the fund is primarily for
the purchase of equipment, requests can include
personnel and travel costs. All awards are subject to
THS board approval. Awards may not necessarily be
granted every period and are subject to availability of
funds. Funding may be awarded out of cycle under
special circumstances.
Proposals
Requests should include a general description of the
project, its objectives and methods, and time frame for
both equipment use and project closure (not to exceed
2 pages, single-spaced). A detailed budget should be
included (not to exceed 1 page). The C.H. Lowe
Research Fund Committee will review proposals blind
(i.e., cover pages will be removed before proposals are
distributed to reviewers so that reviewers will not be
influenced by knowing who submitted a particular
proposal). Proposals should be submitted to the C.H.
Lowe Fund Committee Chair (Taylor Edwards) with a
cover page including contact information, and four
copies of the project description and budget. Project
descriptions should avoid reference to the applicant to
maintain anonymity during the review process.
8
SONORAN HERPETOLOGIST 18 (1) 2005
Proposals meeting the following criteria have a better
chance of success: focuses on herpetofauna of the
Sonoran Desert; contributes to conservation; contributes
to education; is novel or unique; is not supported by
other means; provides geographic distribution data; and
allows equipment to be shared among multiple projects.
Deliverables:
All awardees are required to submit a project report for
publication in the Sonoran Herpetologist’s “Current
Research News” within six months of the award. This
report will include a description of the project and may
consist of just a few lines to several paragraphs, dependent on the scope of the work. For appropriate projects,
the committee may also encourage submission of a fulllength article to the Sonoran Herpetologist.
Equipment
Equipment:
Equipment already owned by the THS can be requested
at any time (email or snail mail acceptable) from the
C.H. Lowe Fund Committee Chair. Equipment may
not be available if already in use, and equipment already
in use may need to be shared among multiple projects.
Committee members can authorize the use of equipment
without board approval and will balance multiple
requests to the best of their ability.
All equipment purchased from the C.H. Lowe
Fund is the property of the THS and can be withdrawn
from a project at any time upon request of the committee. Expendable items awarded during a project may
remain property of the awardee. It is understood that
damage to and loss of field equipment can occur.
Researchers are asked to treat THS equipment with care
and respect. Equipment lost or damaged on a project
will not necessarily be replaced.
Additional information:
Researchers must adhere to federal and state Fair Labor,
Civil Rights and ADA Regulations. Awardees must be in
accordance with all federal and state laws regarding
wildlife, animal welfare, and land access. Awardees will
be responsible for acquiring the proper permits for
conducting such research (wherever that research may
be conducted), which may be requested by the
committee. Awards will be revoked immediately if
compliance is not met. The THS holds no responsibility
for research deliverables that may be required by other
participating parties on the project. For further
information, please contact:
Taylor Edwards
C.H. Lowe Research Fund, Committee Chair
Tucson Herpetological Society
PO Box 709, Tucson, AZ 85702-0709
http://tucsonherpsociety.org
[email protected]
B O A R D M E E T I N G SY N O P S I S
30 November 2004
Robert L. Bezy, Secretary
Directors Present: Roy Averill-Murray, Bob Bezy,
Young Cage, Taylor Edwards, Cristina Jones, Hans
Koenig, Ed Moll, Roger Repp, Eric Stitt, Don
Swann, Marty Tuegel.
Directors Absent: None.
Members Present: Kevin Bonine, Dennis Caldwell, Bill
Savary.
Guest: Chet Hedden.
Treasurer’s Report – Marty Tuegel
Beginning Balance
$10,205.31
Deposits
345.00
Expenses
436.89
Ending Balance
10,113.42
Speakers Bureau
521.40
Jarchow Award
235.00
FTHL Fund
226.00
C.H.Lowe Research Fund
3,294.00
General Fund
5,837.02
CRHSD (CD)
$7,256.30
Itemized Deposits: Lowe Research Fund $25.00,
Membership $320.00.
Itemized Expenses: Bank Fees $1.79, S. H. Printing
$163.89, Speakers Bureau $271.21.
Committee Reports
Conservation – Dennis Caldwell. Caldwell and
Young Cage attended the Arizona Wildlife Summit
conducted by AGFD to develop their Comprehensive Wildlife Conservation Strategy (CWCS), a 10year vision for managing Arizona’s fish, wildlife and
natural habitats. This effort will enlist input and
partnerships with agency cooperators, sportsman
and recreational groups, conservation organizations,
special interest groups, and the general public.
Under the new State Wildlife Grant Program, each
state will need to produce their own CWCS to be
eligible for this federal granting program. Members
and the general public are encouraged to voice their
primary concerns by logging into the CWCS Web
site: http://ideasciences.com/phpBB2/
azgfsignup.php.
Caldwell has developed the Backyard Frog
Refugia Guidelines for the appendix of the
Chiricahua Leopard Frog Recovery Plan, and will be
attending a 4 day Population and Habitat Viability
Analysis workshop on 6-9 Dec.
Via email the board approved that President
Edwards send a letter drafted by Dennis Caldwell
supporting the Paseo de Las Iglesias Ecological
Restoration Proposal. The letter was sent via email to
the Army Corps of Engineers and the Pima County
Flood District. Information is at: http://
www.dot.co.pima.az.us/flood/envrest/paseo.htm.
The backyard leopard frog pond project with
Saguaro National Park is proceeding, with three new
ponds under construction. Volunteers are needed to
help dig the holes over the next couple weeks.
Via email the board approved (Hans Koenig
and Roy Averill-Murray abstaining).sending a letter
of support for a grant proposal to AGFD submitted
by Rincon institute.
Speakers Bureau – Taylor Edwards and Ed Moll.
1 November – Roger Repp gave a talk to the
Notch Neighborhood group with ca 40 people
in attendance.
17 November – Ed Moll taught a course in
beginning herpetology to a group of eleven
seniors for the Oasis Senior’s education program
at Robinson’s May Department Store in El
Con Mall, including a Power Point programs
on local herpetology and opportunities to
touch and hold live reptiles.
19 November - Paul Congdon and Steven
Congdon gave 3 presentations (50 minutes
each) at Salpointe High School using slides and
3 live snakes with a total of over 100 students
in attendance.
Sonoran Herpetologist - Don Swann. The December
issue and most of the January issue are complete.
The Board approved that a $100 cap be placed on
purchase of gift books for previous service and the
reimbursement for the purchase of a book for the
previous editor.
Program – Roy Averill-Murray. The lineup of
speakers includes:
21 December – Young Cage.
18 January – Taylor Edwards.
The Board is seeking a member to fill the important
position of Program Chair which is being
vacated by Roy Averill-Murray effective
December 2004.
PARC – Roy Averill-Murray. Information regarding
applications to the Lowe Research Fund has been
added to the grants page of the National PARC web
site.
Old Business
Current Research on Herpetofauna of the Sonoran
Desert III – Taylor Edwards. The symposium will
take place on 15-17 April 2005 in Phoenix and
abstracts are due before 1 February.
Board of Directors – Taylor Edwards
Edwards. The Board
discussed the one-year vacancy left by the early
departure of Roy Averill-Murray and approved the
appointment of Cristina Jones to fill this position
New Business
Gathering of the Greens – Taylor Edwards. The
event hosted by the Tucson Audubon Society will
be held on 16 December, 1700-2100 hrs.
Rattlesnake Hibernaculum - Chet Hedden
Hedden. The
board discussed the possible need to protect a
denning site described by Hedden.
SONORAN HERPETOLOGIST 18(1) 2005
9
M E M B E R S H I P
Membership Information
Individual
Family
Student
Barking Frog
(Eleutherodactylus augusti)
from Cochise Co., AZ.
Photo by
Erik F. Enderson.
$20
$25
$14
Sustaining
Contributing
Life
$30
$50
$500
To receive a membership form and recent issue of
Sonoran Herpetologist call (520) 624-8879 or write:
Tucson Herpetological Society
P. O. Box 709
Tucson AZ 85702-0709
New Reptile and Amphibian
Regulations for 2005
In October, the Arizona Game and Fish Commission
approved several new changes to regulations for amphibians and reptiles, including:
Commission Order 41 (Amphibians) – Reduce bag
and possession limits to two for Barking Frogs.
Commission Order 43 (Reptiles) – Reduce bag and
possession limits to four per year or in possession for
Western Diamondbacked Rattlesnakes and Mojave
Rattlesnakes. Add Mexican Garter Snakes, Narrowheaded Garter Snakes, and Ornate Box Turtles to the list
of closed-season species.
Time to Renew Your THS Membership?
I hope this is a helpful reminder to those of you whose membership renewal is
due this month. Please call or email with corrections and errors. 624-8879 or
[email protected]
Dave Hardy Sr.
Membership Secretary
Due in January:
Joseph Collet
Caren Goldberg
Trevor Hare
Dennis & Gracie Jex
Marcia Lincoln & Victor Ong
Phil Pugliese
Michael Sredl
John Sullivan
Richard & Margie Wagner
¬
THS Raffle - January Meeting
Happy Newt Year herpers! Because of the THS
election, the November raffle was cancelled at the last
minute. To make up for it, we are going to have a special
raffle this month. Besides a signed copy of Marty
Crump's book, "In Search of the Golden Frog", and
Chris Scott's book "Endangered and Threatened Animals
of Florida and their Habitats", there will be several
surprise items. So don't miss it.
Raffle tickets are $1.00 each and all proceeds go to THS.
See you at the meeting.
Robin Llewellyn
Raffle Coordinator
10
SONORAN HERPETOLOGIST 18 (1) 2005
Paul, Steven & Lorna Condon
John Gray
Monique Imberski
Joel Johnson
Trevor Persons
Justin & Li Shen Schmidt
Eric Stitt
Don Swann
Bill & Beth Woodin
¬
Membership Update
As of 31 December 2004:
Contributing
Erik Enderson
Tucson
New Member
Ali Rabatsky
Lafayette, LA
Jarchow Conservation Award
Tom Caldwell
Tucson
Erika Nowak
Flagstaff
Speakers Bureau
Tom Caldwell
Tucson
Horned Lizard Fund
Tom Caldwell
Tucson
Erika Nowak
Flagstaff
Charles H. Lowe Herpetological Research Fund
Tom Caldwell
Tucson
Sonoran Herpetologist is the newsletter-journal
of the Tucson Herpetological Society, and is
Copyright 2005. The contents of Sonoran
Herpetologist may be reproduced for inclusion in
the newsletters of other herpetological societies
provided the material is reproduced without
change and with appropriate credit, and a copy
of the publication is sent to the Tucson
Herpetological Society. Occasional exceptions to
this policy will be noted. Contents are indexed in
Zoological Record. A complete set of back issues
are available in the Special Collections area of the
University of Arizona library. They are
accompanied by a copy of The Collected
Papers of the Tucson Herpetological Society,
1988-1991.
Editor
Don Swann, [email protected]
Associate Editor
Dale Turner, [email protected]
Art Editor
Dennis Caldwell, [email protected]
Book Review Editor
Eric Stitt, [email protected]
Distribution
Fred Wilson, [email protected]
Trevor Hare, [email protected]
Membership
Dave Hardy Sr., [email protected]
Information for Contributors
Authors should submit original articles, notes,
book reviews to the Editor, either via email using
an attached word processed manuscript or by
mail to the Society’s address. The manuscript style
should follow that of Journal of Herpetology and
other publications of the Society for the Study of
Amphibians and Reptiles. For further
information, please contact the editor.
Deadline for Sonoran
Herpetologist 18(2): January 15
Officers
President Young Cage, [email protected]
Vice President Eric Stitt, [email protected]
The Tucson
Herpetological
Society is
dedicated to
conservation,
education, and
research
concerning the
amphibians and
reptiles of
Arizona and
Mexico.
Secretary Bob Bezy, [email protected]
Treasurer Marty Tuegel, [email protected]
Directors
Kevin Bonine, [email protected]
Cristina Jones, [email protected] 2005
Elissa Ostergaard, [email protected]
Roger Repp, [email protected] 2004-2005
Robert Villa, [email protected]
Erin Zylstra, [email protected]
Past President Taylor Edwards, [email protected]
Society Activities
Monthly Members Meeting
Program Chairperson (Currently Vacant)
3rd Tuesday, 7:15 PM
Board of Directors Meeting
Last Tuesday of each month (except December), 7:00 PM
Arizona Game and Fish Department Office
555 North Greasewood Road
Speakers Bureau (scheduled presentations)
Taylor Edwards, Director
Ed Moll, Director
Conservation Committee Director
Tucson Herpetological Society
is a registered non-profit
organization.
Dennis Caldwell
Herpetological Information Hotline
Bob Brandner 760-0574
Jarchow Conservation Award
Taylor Edwards, Chairperson
Publications:
Sonoran Herpetologist, Living with Rattlesnakes,
Living with Gila monsters, THS Herp Coloring Book,
THS Collected Papers, 1988-1991
THS Internet World Wide Webpage
http://tucsonherpsociety.org
Erik Enderson [email protected]
For more information about the THS and the reptiles and amphibians of the
Tucson area visit tucsonherpsociety.org
SONORAN HERPETOLOGIST 18(1) 2005
11
Your membership has expired.
This is your only reminder.
Please renew!
January 2005, Volume 18, Number 1
BOOK REVIEW
2
Iguanas: Biology and Conservation
SPECIAL ANNUAL FEATURE
4
THS Membership Directory
SHORT ARTICLE
8
CH Lowe, Jr. Herpetology Research Fund
12
SONORAN HERPETOLOGIST 18 (1) 2005