ISSN 1393 – 6670
Checklist and country status of European bryophytes –
towards a new Red List for Europe
Cover image, outlined in Department Green
Irish Wildlife Manuals No. 84
Checklist and country status of European bryophytes
– towards a new Red List for Europe
N.G. Hodgetts
Citation:
Hodgetts, N.G. (2015) Checklist and country status of European bryophytes – towards a new Red
List for Europe. Irish Wildlife Manuals, No. 84. National Parks and Wildlife Service, Department of
Arts, Heritage and the Gaeltacht, Ireland.
Keywords: Bryophytes, mosses, liverworts, checklist, threat status, Red List, Europe, ECCB, IUCN
Swedish Speices
Information Centre
Cover photograph: Hepatic mat bryophytes, Mayo, Ireland © Neil Lockhart
The NPWS Project Officer for this report was: [email protected]
Irish Wildlife Manuals Series Editors: F. Marnell & R. Jeffrey
© National Parks and Wildlife Service 2015
Contents
(this will automatically update)
Contents ......................................................................................................................................................... 11
Preface
Executive
Summary Preface ............................................................................................................................................................
22
Acknowledgements 2
Executive Summary....................................................................................................................................... 3
Introduction 3
Acknowledgements ....................................................................................................................................... 3
IUCN categories and criteria 5
Application
IUCN criteria Introductionof
...................................................................................................................................................
47
Red Data List categories 9
IUCN categories and criteria ........................................................................................................................ 6
IUCN criteria and categories interpreted for European bryophytes 10
Application of IUCN criteria ........................................................................................................................ 8
References 12
Red Data List
categoriesbryophyte
...............................................................................................................................10
Appendix
1. European
checklists and Red Lists 13
Appendix 2. European bryological societies and journals 26
IUCN criteria and categories interpreted for European bryophytes .........................................................11
Appendix 3. Floras 28
References .....................................................................................................................................................13
Appendix 4. ECCB Country Contacts 29
Appendix 5.
1. Data
European
andRed
RedList
Lists
......................................................................14
Appendix
sheetbryophyte
examples checklists
for potential
species
34
Appendix 6. Checklist and country status of European liverworts and hornworts 44
Appendix 2. European bryological societies and journals .........................................................................27
Appendix 7. Checklist and country status of European mosses 65
Appendix 3. Floras .......................................................................................................................................29
Appendix 8. Taxon references for additions and nomenclatural changes to the European
Appendix 4. ECCB
moss checklist
of Hill et.........................................................................................................30
al. 2006 118
Country Contacts
Appendix 5. Data sheet examples for potential Red List species ..............................................................35
Appendix 6. Checklist and country status of European liverworts and hornworts ................................45
Checklist and country status of European bryophytes
Preface
The Board of the ECCB (European Committee for the Conservation of Bryophytes) met in Zurich 2011
and agreed that the Red Data Book of European Bryophytes (published in 1995) needed to be updated
and replaced, because since its publication, there has been an improvement in our knowledge of the
bryophytes of Europe, especially concerning taxonomy and distribution. Later on, in April 2012, the
8th conference on Bryophyte Conservation was held in the Hungarian Natural History Museum,
Budapest, with 70 participants from 34 countries. The conference decided to prepare a new Red Data
Book of European Bryophytes with a new Red List. ECCB established a Main Workgroup and
Regional Workgroups (NW-, SW-, N-, Central-, SE-Europe and Russia).
The main tasks for the new Red Data Book of European Bryophytes project are: to collate and update
existing information for European countries on the basis of country Red Lists and checklists; to list
candidate species for the Red Data Book; to establish methodologies and protocols for data collection,
species distribution mapping and threat assessments, and to produce and publish a Red Data Book.
The first two tasks have now been completed. A comprehensive table, listing the occurrence and Red
List status of each species, in each European country, has been prepared. ECCB contacts from almost
all European countries have been involved in the work. The 1st Workshop meeting of the new Red
Data Book of European Bryophytes project was held in Ekenäs, Sweden, in 2013. A shortlist of
candidate species for IUCN threat assessment was established by eliminating those species known
with certainty to be of Least Concern. Of 1980 bryophyte taxa known to occur in Europe, 242
liverworts and 693 mosses were selected for further consideration.
This report is a summation of these results and gives a methodological overview of how IUCN criteria
could be applied to European bryophytes. The work was financially supported by the Irish National
Parks and Wildlife Service (NPWS) and the Swedish ArtDatabanken (SLU). The report was compiled
by Nick Hodgetts and edited by Neil Lockhart, Tomas Hallingbäck and Edwin Wymer. This work
would not have been possible without the generous help of ECCB country representatives, who have
contributed with current information regarding checklists, Red Lists and bryophyte conservation
generally in each European country. This report is also available in PDF format, and the checklists as
downloadable Excel files, on the ECCB website (http://eccbbryo.nhmus.hu/).
I hope that this report will be useful for all bryologists, especially those interested in the conservation
of bryophytes. I would like to thank all of you who have worked and contributed to this, so please
enjoy this report as the first results of your efforts.
Dr Beáta Papp Chair of ECCB (European Committee for Conservation of Bryophytes)
1
Checklist and country status of European bryophytes
Executive Summary
This report describes the background to the project to revise the Red List for bryophytes in Europe.
Since the launch of the pilot study at the European Committee for the Conservation of Bryophytes
(ECCB) conference in Budapest 2012, steady progress has been made towards updating a checklist of
liverworts, hornworts and mosses for all of Europe. With assistance from the network of Regional Coordinators and Country Contacts, the presence of taxa recorded for each country/territory, with local
Red List status, is presented in this report. This checklist will also be published on the ECCB website
http://eccbbryo.nhmus.hu/node/4.
A workshop meeting held in Ekenäs, Sweden in March 2013 produced an initial list of ‘candidate
species’ from the checklist. These species can proceed forward for IUCN threat assessment. This was
achieved by identifying and then excluding those taxa considered with certainty to be Least Concern.
With the elimination of these Least Concern taxa, there are currently 242 liverwort and hornwort taxa,
and 693 moss taxa, which are candidates for Red List assessment.
This report also reviews and refines IUCN categories and criteria that could be used in the Red List
assessments. Examples of a possible format for data sheets for Red List species are presented.
References to country checklists and Red Lists and an updated list of ECCB Country Contacts are also
provided.
Acknowledgements
I would like to thank all the members of the ECCB and other bryologists Europe-wide, too numerous
to mention individually, who helped me produce this report by generously providing information. I
would particularly like to thank the ECCB Board for constructive discussions and helpful advice, and
Tomas Hallingbäck of the Swedish ArtDatabanken and Neil Lockhart of the Irish National Parks and
Wildlife Service for providing funding for this pilot project.
Nick Hodgetts
2
Checklist and country status of European bryophytes
Introduction
Red Lists are needed to underpin conservation action with the best scientific data available. The
International Union for the Conservation of Nature (IUCN) produces a global Red List to highlight the
most threatened organisms on the planet, but Red Lists are also produced regionally and on a country
basis. Bryophytes, although individually small, are collectively a key part of vegetation communities
and biodiversity. Like other organisms, bryophytes are under increasing threat of decline and
ultimately extinction from anthropogenic pressures, and nowhere more so than in Europe, one of the
most developed and densely-inhabited parts of the globe. New information is needed at a European
level to inform the conservation of bryophytes and their habitats, and a Red List is a key element of
this.
The Global Strategy for Plant Conservation (Secretariat of the Convention on Biological Diversity
2003; http://intranet.iucn.org/webfiles/doc/SSC/SSCwebsite/Plants/global_strategy.pdf) lists as one of
its objectives “understanding and documenting plant diversity”, with a sub-objective to “monitor the
status and trends in global plant diversity and its conservation, and threats to plant diversity, and
identify plant species, plant communities, and associated habitats and ecosystems, at risk, including
consideration of Red Lists”. This proposal for a new European bryophyte Red List therefore clearly
links in with the Strategy.
The European Committee for the Conservation of Bryophytes (ECCB) was inaugurated in 1990 to
address the conservation of bryophytes in Europe. It is composed of a network of bryologists Europewide, with members from universities, museums, conservation agencies and private individuals in
partnership. It has no funding, but has a loose membership and is run principally by a Board that is
re-elected at conferences, which are held every 4 years or so.
The ECCB produced the first Red List of bryophytes in Europe in 1995 (ECCB 1995), using the old
IUCN categories and criteria. Since then, while there have been many positive conservation actions for
bryophytes, the overall picture, both in Europe and globally, is still one of habitat loss and decline.
Habitats sensitive to disturbance such as old-growth forest, fens and bogs and species-rich grassland
continue to become smaller in extent and more vulnerable to pressures from the ever-increasing
human population. It is now accepted by most of the scientific community that climate change is
taking place, with a general warming resulting in a reduction in snow and ice. This is particularly
noticeable in mountain ranges such as the Alps, where the glaciers are clearly shrinking. This is bound
to have a knock-on effect on arctic-alpine plant communities, including the bryophytes. In northern
Europe and Asia, areas of tundra that were once frozen in permafrost are now thawing, again with
profound consequences for the flora and fauna and perhaps even the climate of Europe.
3
Checklist and country status of European bryophytes
Perhaps most seriously, Europe, and the world generally, has fallen on hard times. All over Europe
nature conservation is taking a back seat while governments struggle with their economic priorities,
largely not acknowledging the fact that “the worldwide economy is a wholly owned subsidiary of the
environment, not the other way around” (Gaylord Nelson, US Senator and founder of Earth Day).
Resources for activities such as nature conservation are becoming scarcer, and increasingly
conservation, particularly of less high-profile organisms such as bryophytes, is seen as a luxury that
cannot be afforded. This is, of course, a mistake. Surely now, if ever, it is time to look for new
economic models, to prioritise sustainability over growth, incorporating care for our environment and
the species with which we share the planet into our schemes for economic prosperity.
Our knowledge of the European bryophytes is much better now than it was in 1995, with the
discovery of new species, and the publication of many new checklists and country Red Lists.
Furthermore, the IUCN has refined its criteria and categories and the European Red List needs to be
updated to take this into account. For any Red List to be useful, it needs to use the most up-to-date
information available, and it is now clear that, while ground-breaking, the 1995 Red List was based on
inadequate information and is becoming increasingly outdated. In Zürich in 2011, the ECCB Board
agreed that a new European Red List for bryophytes is both desirable and necessary, and that a pilot
project to assess the feasibility of making a new Red List would be a good first step.
A pilot study was therefore commissioned and published (Hodgetts 2012), and presented at an ECCB
conference in Budapest in 2012. This document explained the importance of bryophytes, collated
existing information about regional and national Red Lists, reviewed the ECCB network of contacts,
and identified gaps in the information resource. The following resolutions were made at the Budapest
conference:
•
It was agreed that a new Red List was both desirable and feasible.
•
Any such project should be led by an ECCB Steering Group composed of Tomas Hallingbäck,
Neil Lockhart (as prime movers in the pilot project) and Beáta Papp (as ECCB chair), plus
whoever else they deem necessary to progress the project. Nick Hodgetts was named as
secretary.
•
The work should be done (under direction from the Steering Group) by a contractor with
some bryological expertise, or a number of contractors with a blend of bryological and
technical expertise.
•
The project will use the latest IUCN categories and criteria.
4
Checklist and country status of European bryophytes
•
Geographical coverage will include all the area covered by the previous Red List document
(ECCB 1995). [A later meeting of the Steering Group clarified this to include the Canary
Islands, Madeira and the Azores, but to exclude Cabo Verde, Asiatic Turkey and Cyprus.]
•
The taxonomy will be based on Hill et al. (2006), plus more recent additions and accepted
amendments for the mosses, and the forthcoming world list of liverworts (Söderström et al., in
prep.) for the liverworts.
•
All taxa will be considered for the Red List, but taxa that are undoubtedly Least Concern can
be eliminated relatively quickly in a workshop meeting. [This was done in 2006 for the
liverworts but still needed to be done for the mosses.]
A meeting of the ECCB Steering Group in Ekenäs, Sweden in March 2013 addressed the final point, by
revising the liverwort list and examining the moss list. It thus produced an initial list of ‘candidate
species’ to be considered for the Red List project. These are included in the tables in Appendix 6 and 7.
These
will
also
be
made
available
as
Excel
spreadsheets
on
the
ECCB
website
http://eccbbryo.nhmus.hu/node/4. These tables comprise a revised and updated checklist of mosses,
liverworts and hornworts for all of Europe. Presence of taxa is recorded for each country/territory,
with local Red List information included where available. There are currently 541 liverworts and
hornwort taxa, of which 242 (45%) are candidates for the Red List, and 1439 mosses, of which 693
(48%) are candidates for the Red List. The final number of species included in a Europe-wide Red List
will be considerably lower than this: the ‘candidate species’ are those which are thought to be
sufficiently rare and/or threatened at least to be run through the IUCN Red List criteria. The
remainder (‘Least Concern’) are those which the Steering Group considered to be so common and/or
widespread that they did not need to be considered further for the Red List.
IUCN categories and criteria
The IUCN system for allocating threat status was originally designed to be applied to large animals on
a global scale, but more recently has been developed so that it is applicable to any organism at any
geographical scale, if interpreted sensibly. The previously approved IUCN threat categories (Extinct,
Endangered, Vulnerable and Rare) were replaced in 1994 by a revised system (World Conservation
Union 1994) that abandoned the Rare category (as it was an expression of frequency rather than of
threat), and introduced the category of Critically Endangered. The sub-Red List category of Near
Threatened was also established, and the category of Extinct in the Wild was distinguished from Extinct.
The IUCN categories and criteria for allocating species to each category were further refined over a
period of several years, with version 3.1 published in 2001, with a second edition in 2012 (IUCN 2012).
5
Checklist and country status of European bryophytes
To accompany the IUCN categories and criteria, a series of guidelines have been produced to assist
with their interpretation and use (IUCN 2014). One of the main innovations in the guidelines is a more
realistic approach to extent of occurrence, using the so-called α-hull method. This is a system for
establishing the range (extent of occurrence) of a taxon while excluding aberrant occurrences well
outside its normal range and is explained more fully elsewhere (IUCN 2014).
The IUCN has also produced Guidelines for Application of IUCN Red List Criteria at Regional Levels
(IUCN 2010). This established a Regionally Extinct category for use in regional Red Lists (as distinct
from Extinct, which is global and final). It also introduced the possibility of changing the threat status
of a taxon on the basis of its status in neighbouring territories: taxa can be downgraded (or upgraded)
depending on conditions outside the region covered and whether or not the population can be
rescued from extra-regional populations.
In applying the IUCN criteria, it will be necessary to choose a cut-off date to represent the threshold
between old and recent records. This has not yet been decided for the European list, but a date of 1970
is suggested. This is essentially a compromise, as data sets in different territories differ widely in
quality and detail.
Any Red List for the whole of Europe will inevitably be a political exercise, in that the territories are
artificial (e.g. only small parts of Kazakhstan and Turkey are within Europe; Macaronesia is within
Europe politically but not geographically, etc). Ideally, Red Lists should be global or at least cover
discrete biogeographical areas (e.g. north-west Atlantic Europe, arctic Europe, etc). Numerous data
sources will be required to draw up a European Red List, and each territory has its own way of doing
things, so it will be necessary to collate large amounts of disparate data and standardise it to some
extent before the IUCN criteria can be applied.
The IUCN criteria will be applied to all those taxa included as ‘candidate species’ for the Red List, as
listed in the moss and liverwort tables (Appendix 6 and 7). These were determined during
‘brainstorming’ meetings of the ECCB Red List Steering Group, during which all taxa that definitely
should not be included in any Red List, on the basis of being too frequent, not undergoing a decline, or
not threatened in at least some parts of Europe, were excluded. A relatively small number of species
were labelled ‘regionally threatened’, as they had declined in some areas but are still frequent and
unthreatened in others. These will also be run through the IUCN Red List criteria to determine their
true status over Europe as a whole.
6
Checklist and country status of European bryophytes
Application of IUCN criteria
The application of IUCN criteria proposed for the European bryophyte flora is essentially a modified
version of that used in the preparation of the recently published Irish Red List for bryophytes
(Lockhart et al. 2012). The Red List consists of taxa in the categories Extinct (or Regionally Extinct),
Critically Endangered, Endangered and Vulnerable. For each of the main threat categories (Critically
Endangered, Endangered and Vulnerable) there is a set of five main criteria A–E, any one of which
qualifies a taxon for listing at that level of threat. The qualifying thresholds within the criteria A–E
differ between threat categories.
Criterion A considers percentage decline, regardless of current range or abundance, and will be used
very sparingly. For bryophytes, it is often difficult to decide what constitutes a mature individual, or
even if an individual can be defined, and so estimates of the size of bryophyte populations are rarely
available. It is also difficult to measure the rate of decline of bryophyte taxa from the available data
that does exist, because records have so often been made at different times, in different areas, and
there has been little systematic monitoring of populations over time. When considering population
decline, the use of generation time is a useful concept for bryophytes as it enables decline over a
longer time period than ten years to be used. Hallingbäck et al. (1998) advise using a maximum of 25
years for one generation (for species that are not known to reproduce sexually), with a sliding scale of
11–25 years for species that reproduce sexually only infrequently, down to 1–5 years for short-lived
ephemeral colonists that reproduce frequently with small, highly mobile spores. In other words, a
system of life strategies, such as that devised by During (1992), needs to be adopted in order to obtain
a broad estimate of generation time.
Subcriterion A1 will not be used, as there are no taxa for which there is certainty about whether their
decline is reversible, understood and ceased. However, subcriteria A2 and A3 allow inferred or
suspected decline, and the inference or suspicion can be based on a decline in habitat. This can
sometimes be seen only too clearly. Thus, the very specific fen habitat of Paludella squarrosa has clearly
declined in some areas, so subcriterion A2c can be used to contribute towards its assessment.
Tomentypnum nitens, although occurring in too many populations to qualify as Vulnerable under other
criteria, is known to have been destroyed at many localities, so A2c might also be appropriate here.
Subcriterion A3c can probably be used for some rare ruderal species, as it seems clear that their habitat
will deteriorate in the future through natural succession. Subcriterion A4 will not be used, as it
requires decline in both the past and the future, a level of detail too specific for bryophytes at our
current state of knowledge.
7
Checklist and country status of European bryophytes
Criterion B is used to categorise taxa that have a restricted distribution and are also declining. Extent
of occurrence has in the past been used sparingly to determine threat category, but the advent of the
α-hull method allow it to be used more extensively. If this shows a significant reduction in extent of
occurrence (i.e. range), when old records are compared with recent records (however these are
defined), then clearly the concept is a useful one for determining threat category. However,
bryophytes tend naturally to have very wide ranges, often with wide disjunctions between
populations, so the concept of extent of occurrence may be less relevant to them than it is to many
other species groups. Consequently, it is suggested that subcriteria B1a and B1b (extent of occurrence)
are used only infrequently and with caution, and subcriterion B1c not at all.
Area of occupancy also presents problems of interpretation. It should be measured, according to the
IUCN Guidelines, on grid squares "which are sufficiently small", and which are of appropriate size for
the biological aspects of the taxon. This criterion is perhaps more applicable to a mobile animal
holding a territory or a home range that can be measured. For a plant it is either much more difficult
to determine the area needed for its survival, or the area might be tiny in comparison. For simplicity,
the area of occupancy can be interpreted in terms of 10 km² squares on the UTM grid, which, while
not always the finest resolution attainable with the existing data, will probably work quite well in
conjunction with the Criterion B subcriteria. It certainly works well at national level (e.g. Hodgetts
2011; Lockhart et al. 2012). However, mapping is done on a 50 km² square basis Europe-wide, so an
initial ‘sift’ of species for which this criterion might be appropriate on the grounds of a restricted area
of occupancy can be carried out using 50 km² square distribution maps, before examining the data
more closely.
The concepts of extreme fluctuation and fragmentation are listed as additional risk factors by IUCN,
but these are seldom applicable to most bryophytes, as many species can fluctuate considerably as
part of their natural population dynamics, and their distribution naturally appears to be fragmented.
One of the few exceptions to this generalisation is found in the specialised bryophyte flora that grows
in the draw-down zone of reservoirs and similar habitats. Subcriterion B2c can be used for these
where their populations could be at risk and subject to extreme fluctuation if water levels are kept
artificially high for too long.
Criterion C requires detailed data on both population size and decline, and so cannot be used because
this level of information is not available for bryophytes Europe-wide.
Criterion D identifies very small or restricted populations and is an expression of rarity, inferring that
a taxon is threatened by human activities or stochastic events simply because it is rare, without
necessarily having declined. It is suggested that Criterion D should be used very sparingly for
8
Checklist and country status of European bryophytes
assigning taxa to the Critically Endangered or Endangered categories, as there is usually no detailed
information on population size. However, it can be used in a few cases where it can reasonably be
inferred that a population consists of fewer than 50 mature individuals or fewer than 250 mature
individuals (depending, of course, on the interpretation of ‘an individual’). Subcriterion D2 can be
used more extensively for assigning taxa with less than 5 localities to the Vulnerable category.
Criterion E uses quantitative analyses to consider the probability of extinction in the wild. This cannot
be used, as there have been no population viability analyses published on bryophytes in Europe, as far
as is known.
Red Data List categories
Extinct (EX). A taxon is Extinct when there is no reasonable doubt that the last individual has died.
Extinct in the Wild (EW). A taxon is Extinct in the Wild when it is known to survive only in cultivation
or as a naturalised population well outside the past range.
Regionally Extinct (RE). A taxon is regarded as Regionally Extinct in Europe if there are no recent
records and all known localities have been visited and surveyed without success. Failure to refind
older records may sometimes just reflect the imprecision of the original locality data, or adverse
weather conditions during recent survey visits, or simply that certain taxa occur sporadically and are
inherently difficult to find. It is possible therefore that some Regionally Extinct taxa may persist, albeit
at a relatively low frequency, and might yet turn up at some future date. An effort should be made to
distinguish between taxa that have probably genuinely disappeared from those that may still occur,
the latter being placed in the Data Deficient category.
Critically Endangered (CR). A taxon is Critically Endangered when the best available evidence indicates
that it meets any of the criteria A to E for Critically Endangered, and it is therefore considered to be
facing an extremely high risk of extinction in the wild.
Endangered (EN). A taxon is Endangered when the best available evidence indicates that it meets any of
the criteria A to E for Endangered, and it is therefore considered to be facing a very high risk of
extinction in the wild.
Vulnerable (VU). A taxon is Vulnerable when the best available evidence indicates that it meets any of
the criteria A to E for Vulnerable, and it is therefore considered to be facing a high risk of extinction in
the wild.
9
Checklist and country status of European bryophytes
Near Threatened (NT). A taxon is Near Threatened when it has been evaluated against the criteria but
does not qualify for Critically Endangered, Endangered or Vulnerable now, but is close to qualifying for or
is likely to qualify for a threatened category in the near future. This is rather subjective and how it
might work across Europe has yet to be discussed.
Data Deficient (DD). A taxon is Data Deficient when there is inadequate information to make a direct,
or indirect assessment of its risk of extinction based on its distribution and/or population status.
Least Concern (LC). A taxon is Least Concern when it has been evaluated against the criteria and does
not qualify for Critically Endangered, Endangered, Vulnerable or Near Threatened. Widespread and
abundant taxa are included in this category. All the taxa eliminated from the ‘candidate list’ during
the preliminary sorting exercises fall into this category.
Not Evaluated (NE). A taxon is Not Evaluated when it has not been assessed against the criteria.
IUCN criteria and categories interpreted for European
bryophytes
Fulfilling any one of these criteria leads to the application of a threat category at the appropriate level.
Criterion A. Rapid decline. A2c can be used for a small number of taxa, on the basis of a decline in
habitat quality; A3c can be used for a small number of taxa, on the basis of a predicted decline and loss
of habitat. > 80% decline (CR); > 50% decline (EN); > 30% decline (VU).
Criterion B. Small range; fragmented, declining or fluctuating. This criterion can only be used for a
small number of taxa, such as specialist reservoir species, as both fragmentation and extreme
fluctuation are common natural features of bryophyte populations.
1. Extent of occurrence (estimated using α-hull method, where α = 2): < 100 km² (CR); < 5000 km² (EN);
< 20,000 km² (VU) AND both a and b:
a.
single location (CR); 5 locations or fewer (EN); 10 locations or fewer (VU).
b. continuing decline observed, inferred or projected, in any of the following:
(i) extent of occurrence
(ii) area of occupancy
(iii) area, extent and/or quality of habitat
(iv) number of locations or subpopulations
(v) number of mature individuals
10
Checklist and country status of European bryophytes
2. Area of occupancy: < 10 km², using 1 hectad (CR); < 500 km², using 5 hectads (EN); < 2000 km², using
20 hectads (VU) AND at least two of a–c:
a. single location (CR); 5 locations or fewer (EN); 10 locations or fewer (VU).
b. continuing decline observed, inferred or projected, in any of the following:
(i) extent of occurrence
(ii) area of occupancy
(iii) area, extent and/or quality of habitat
(iv) number of locations or subpopulations
(v) number of mature individuals
c. extreme fluctuations in any of the following (but only used for reservoir species):
(i) extent of occurrence
(ii) area of occupancy
(iii) number of locations or subpopulations
(iv) number of mature individuals
Criterion D/D1. Very small population. Used very sparingly, and only if there is no reasonable
doubt: < 50 individuals estimated (CR); < 250 individuals estimated (EN); < 1000 individuals estimated
(VU).
Criterion D2. Very small range. Can be used extensively to assign taxa to VU: 5 or fewer locations.
11
Checklist and country status of European bryophytes
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Bryophytes and lichens in a changing environment. Oxford, Clarendon Press.
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Trondheim, ECCB.
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Vána, J. 1998. Guidelines for the application of the revised IUCN threat categories to bryophytes. Lindbergia 23
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Mazimpaka, V., Muñoz, J. & Söderström, L. 2006. An annotated checklist of the mosses of Europe and
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IUCN 2014. Guidelines for using the IUCN Red List categories and criteria. Version 11 (February 2014). Gland,
International Union for the Conservation of Nature.
Lockhart, N., Hodgetts, N. & Holyoak, D. 2012. Ireland Red List No. 8. Bryophytes. Mosses, liverworts & hornworts.
Dublin, National Parks & Wildlife Service.
World Conservation Union 1994. IUCN Red List categories. Gland, International Union for the Conservation of
Nature.
12
Checklist and country status of European bryophytes
Appendix 1. European bryophyte checklists and Red Lists
These references cover Europe as a whole, European regions and individual European countries or
territories. Only the most recent and relevant references are included; there are many older works that
have been superseded. For a more complete list of references for liverworts, see Söderström et al. 2008.
Europe
Düll, R. 1983. Distribution of the European and Mediterranean liverworts (Hepatiocophytina). Bryologische
Beiträge 2: 1–114.
European Committee for Conservation of Bryophytes (ECCB) 1995. Red Data Book of European Bryophytes.
Trondheim, ECCB.
Hill, M.O., Bell, N., Bruggeman-Nannenga, M.A., Brugués, M., Cano, M.J., Enroth, J., Flatberg, K.I., Frahm, J.-P.,
Gallego, M.T., Garilleti, R., Guerra, J., Hedenäs, L., Holyoak, D.T., Hyvönen, J., Ignatov, M.S., Lara, F.,
Mazimpaka, V., Muñoz, J. & Söderström, L. 2006. An annotated checklist of the mosses of Europe and
Macaronesia. Journal of Bryology 28: 198–267.
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
Séneca, A. & Söderström, L. 2009. Sphagnophyta of Europe and Macaronesia: a checklist with distribution data.
Journal of Bryology 31: 243–254.
Söderström, L., Hagborg, A., von Konrat, M. & Renner, M. 2008. Early Land Plants Today: liverwort checklist of
checklists. Fieldiana 47: 105–130.
Söderström, L., Urmi, E. & Váňa, J. 2002. Distribution of Hepaticae and Anthocerotae in Europe and Macaronesia.
Lindbergia 27: 3–47.
Söderström, L., Urmi, E. & Váňa, J. 2007. The distribution of Hepaticae and Anthocerotae in Europe and
Macaronesia – Update 1–427. Cryptogamie, Bryologie 28: 299–350.
Albania
Colacino, C. & Sabovljević, M. 2006. Bryophyte flora of Albania: a preliminary check-list. Cryptogamie, Bryologie 27
(4): 471–498. (for additions see also refs in pers. comm. from Jani Marka, 14.6.11)
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
13
Checklist and country status of European bryophytes
Andorra
Sérgio, C., Brugués, M., Cros, R.M. Casas, C. & Garcia, C. 2006. The 2006 Red List and an updated checklist of
bryophytes of the Iberian Peninsula (Portugal, Spain and Andorra). Lindbergia 31: 109-125.
Austria
Grims, F. & Köckinger, H. 1999. Rote Liste gefährdeter Laubmoose (Musci) Österreichs. 2. Fassung. In: Niklfeld,
H. et al., Rote Listen gefährdeter Pflanzen Österreichs. Wien, Grüne Reihe des Bundesministeriums für Umwelt,
Jugend und Familie, pp. 157–171.
Köckinger, H., Schröck, C., Krisai, R. & Zechmeister, H.G. 2011. Checklist of Austrian bryophytes. Webpage.
http://131.130.59.133/projekte/moose/ (which claims that “the list will be updated continuously”)
Saukel, J. & Köckinger, H. 1999. Rote Liste gefährdeter Lebermoose (Hepaticae) und Hornmoose (Anthocerotae)
Österreichs. 2. Fassung. In: Niklfeld, H. et al., Rote Listen gefährdeter Pflanzen Österreichs. Wien, Grüne Reihe des
Bundesministeriums für Umwelt, Jugend und Familie, pp. 172–179.
Belarus
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Rykovsky, G. & Maslovsky, O. 2005. Bryophytes - In: Belarus Red Book (3 ed.). Minsk, 295–325 pp.
Rykovsky,
G.
&
Maslovsky,
O.
2009.
Minsk. Vol. 1 - 2004, 435 pp., Vol. 2, 213 pp.
Flora
of
Belarus.
Bryophyta.
(in
2
volumes).
Belgium
De Zuttere, P. & Schumacker, R. 1984. Bryophytes nouvelles, méconnues, rares, menacées ou disparues de Belgique.
Minist. Region Wallone, Serv. Conserv. Nature, Trav., 160 pp.
Sotiaux, A. & Vanderpoorten, A. 2001. Check-list of the bryophytes of Belgium. Belgian Journal of Botany 134 (2):
97–120.
Sotiaux, A., Stieperaere, H. & Vanderpoorten, A. 2007. Bryophyte checklist and European Red List of the BrusselsCapital Region, Flanders and Wallonia (Belgium). Belgian Journal of Botany 140 (2): 174–196.
Bosnia-Herzegovina
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
Bulgaria
Ganeva, A. & Natcheva, R. 2003. Check-list of the bryophytes of Bulgaria with data on their distribution. I.
Hepaticae and Anthocerotae. Cryptogamie, Bryologie 24 (3): 229–239. (see also http://www.bryologybg.hit.bg/English/index.htm for updates)
Natcheva, R. & Ganeva, A. 2005. Check-list of the bryophytes of Bulgaria. II. Musci. Cryptogamie, Bryologie 26 (2):
209–232. (see also http://www.bryology-bg.hit.bg/English/index.htm for updates)
Natcheva, R., Ganeva, A. & Spiridonov, G. 2006. Red List of the bryophytes in Bulgaria. Phytologia Balcanica 12(1):
55–62.
Peev, D. (ed.) 2011. Red Data Book of the Republic of Bulgaria. Vol. 1. Plants and Fungi. Sofia, Bulgaria Academy of
Sciences & Ministry of Environment and Water (see also digital edition, http://www.eecodb.bas.bg/rdb/en/vol1/).
14
Checklist and country status of European bryophytes
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207 –244.
Croatia
Sabovljevic, M. 2003. The hepatic check list of Croatia. Archives of Biological Sciences, Belgrade 55(1–2): 59–66 .
Sabovljević, M. 2006. Checklist of mosses of Croatia. Archives of Biological Sciences, Belgrade 58 (1): 45–53.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207 –244.
Cyprus
Frahm J.-P., Lüth M. & van Melick H. 2009. Die Moose Zyperns. Archive For Bryology 46: 1–8.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Czech Republic
Kučera, J. & Váňa, J. 2003. Check- and Red List of bryophytes of the Czech Republic (2003). Preslia 75: 193–222.
Kučera, J. & Váňa, J. 2005. Seznam a červený seznam mechorostů České republiky (2005) Příroda, Praha, 23: 1–104.
Kučera, J., Váňa, J. & Hradílek, Z. 2012. Bryophyte flora of the Czech Republic: updated checklist and Red List
and a brief analysis. Preslia 84: 813–850.
Váňa, J. 1997. Bryophytes of the Czech Republic – an annotated check-list of species (1). Novit. Bot. Univ. Carol.,
Praha 11: 39–89.
Váňa, J. 1998. Bryophytes of the Czech Republic – an annotated check-list of species (2). Novit. Bot. Univ. Carol.,
Praha 12: 7–33.
Denmark
Damsholt, K. 2002. Illustrated flora of Nordic liverworts and hornworts. Lund, Nordic Bryological Society.
Damsholt, K., Goldberg, I. & Øllgaard, H. 2008. Danish and Latin names for Liverworts and Hornworts that occur in
Denmark 2008. Webpage: http://www.bryologkredsen.dk/pdf/danlevermos2008.pdf (accessed 6.6.14).
Damsholt, K., Holmen, K. & Warncke, E. 1969. A list of the bryophytes of Denmark. Botanisk Tidsskift 65: 163–183.
Mogensen, G.S. & Goldberg, I. 2005. Danske og latinske navne for Tørvenmosser, Sortmosser of Bladmosser, der
forekommer i Danmark. Version 4. [Danish and Latin names for Sphagna, Andreaeas and Mosses that occur in
Denmark. Version 4] Botanisk Have & Museum. Københavns Universitet, Danmark. 30 sider. Webpage.
http://www.bryologkredsen.dk/pdf/danmos4.pdf (accessed 6.6.14).
Estonia
Ingerpuu, N. 1998. Sammaltaimed, Bryophyta. In: Lillelecht V (ed.), Eesti punane raamat. Tartu, Eesti Teaduste
Akad. Lood., pp. 37–49. (see also http://www.botany.ut.ee/bryology/ (accessed 10 June 2011)
Ingerpuu, N., Kalda, A., Kannukene, L., Krall, H., Leis, M. & Vellak, K. 2005. Esti Sammalde Nimestik. [List of
Estonian Bryophytes]. http://www.botany.ut.ee/bryology/ (accessed 9 June 2011)
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Anon. 2008. Red Data Book of Estonia. Bryophytes. Webpage. www.zbi.ee/punane/liigid/samblad_e.html (accessed 9
June 2011)
15
Checklist and country status of European bryophytes
More detailed information about Red List species can be found at: http://elurikkus.ut.ee/prmt.php?lang=eng
(Accessed 6.6.14).
Faeroe Islands
Boesen, D.F., Lewinsky, J. & Rasmussen, L. 1975. A check-list of the bryophytes of the Faroes. Lindbergia 3: 69–78.
Finland
Koponen, T., Isoviita, P. & Lammes, T. 1977. The bryophytes of Finland: an annotated checklist. Helsinki, Societas
pro fauna et flora Fennica.
Kotiranta, H., Uotila, P., Sulkava, S. & Peltonen, S.-L. (eds.) 1998. Red Data Book of East Fennoscandia. Helsinki,
Ministr. Envir., 351 pp.
Rassi, P., Alanen, A., Kanerva, T. & Mannerkoski, I. (eds.) 2001. The Red List of Finnish Species. Helsinki, Ministry
of the Environment & Finnish Environment Institute, 432 pp.
Syrjänen, K., Anttila, S., Ulvinen, T., Laaka-Lindberg, S., Huttunen, S., Laitinen, T., Ahonen, I., Fagerstén, R., He,
X., Juslén, A., Korvenpää, T., Korvenpää, T., Parnela, A., Sallantaus, T., Vainio, O., Virtanen, R., Piippo, S.,
Rikkinen, J. 2010. Bryophytes. In: Rassi, P., Hyvärinen, E., Juslén, A. & Mannerkoski, I. (eds.). The 2010 Red List
of Finnish Species. Ympäristöministeriö & Suomen ympäristökeskus, Helsinki. p. 208–230. (See also
http://www.ymparisto.fi/download.asp?contentid=123022&lan=en)
Syrjänen, K. et al. 2010. Bryophytes, 208–230. In: Rassi, P., Hyvärinen, E., Juslén, A. & Mannerkoski I. (eds.) 2010.
The 2010 Red List of Finnish Species, 685 p. Ministry of the Environment and Finnish Environment Institute. (See
also www.environment.fi/redlist)
Anon. 2001. Extinct, Threatened and Near Threatened liverworts. Helsinki, Finnish Environment Institute. Webpage.
www.environment.fi/default.asp?contentid=262104&lan=EN (accessed 9 June 2011)
[Checklist also on website of Markku Savela (accessed 29 August 2011):
http://www.funet.fi/pub/sci/bio/life/plants/bryophyta/bryopsida/
However, this is clearly very incomplete or out-of-date (compare with the 2010 Red List).]
Ulvinen, T. ja Syrjänen, K. 2011: Suomen sammalten levinneisyys eliömaakunnissa 2. 30.5.2011 (Distribution of
Finnish bryophytes in biogeographic provinces).
http://www.ymparisto.fi/download.asp?contentid=127735&lan=fi
France
Augier, J. 1966. Flore des bryophytes. Editions Paul Lechavalier, Paris.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Upper Normandy
Werner, J., Bardat, J., Vanot, M. & Prey, T. 2009. Check-list des bryophytes (Anthocerotae, Hepaticae, Musci) de
Haute-Normandie (France). Cryptogamie, Bryologie 30 (4) : 457–475.
Picardie
Hauguel, J.C. & Wattez, J.R., 2008. Inventaire des bryophytes de Picardie: présence, rareté et menace. Conservatoire
Botanique National de Bailleul. Rapport d'étude. 38 p.
Pyrénées-orientale
Thouvenot, L. 2002. Flore bibliographique des bryophytes du Département des Pyrénées-orientale. Société d’Hostoire
naturelle de Perpignan et des Pyrénées-orientale.
Cantal
De Zuttere, P. 1993. Bryophytes rare ou nouvelles pour le département du Puy-de-Dôme et du Cantal (Auvergne,
France). Une mousse nouvelle pour la France: Tortula lingulata Lindb. var. montenegrina (Brid. & Szygz.) Broth.
Nowellia Bryologica 3–4 : 27–41.
16
Checklist and country status of European bryophytes
Puy-de-Dôme
De Zuttere, P. 1993. Bryophytes rare ou nouvelles pour le département du Puy-de-Dôme et du Cantal (Auvergne,
France). Une mousse nouvelle pour la France: Tortula lingulata Lindb. var. montenegrina (Brid. & Szygz.) Broth.
Nowellia Bryologica 3–4 : 27–41.
Franche-Comté
Bailly, G., Caillet, M., Ferrez, Y. & Vadam, J.C., 2009. Liste rouge des Bryophytes de Franche-Comté, version 2. Les
Nouvelles Archives de la Flore jurassienne, 7 : 61–81.
Lorraine
Werner, J., Schneider, T., Schneider, C. & Mahévas, T. 2005. Les bryophytes de la Lorraine extra-vosgienne. Liste
critique annotée. Cryptogamie, Bryologie 26 (4): 347–402.
Mahévas, T., Werner, J., Schneider, C. & Schneider, T. 2010. Liste rouge des bryophytes de Lorraine (Anthocérotes,
Hépatiques, Mousses). Conservatoire et Jardin Botanique, 61p. & annexe.
Corsica
Sotiaux, A., Pioli, A., Royaud, A., Schumacker, R. & Vanderpoorten, A. 2007. A checklist of the bryophytes of
Corsica (France): new records and a review of the literature. Journal of Bryology 29: 41–53.
Germany
Ludwig, G., Düll, R., Philippi, G., Ahrens, M., Caspari, S., Koperski, M., Lütt, S., Schulz, F. & Schwab, G, 1996.
Rote Liste der Moose (Anthocerophyta et Bryophyta) Deutschlands. Schriftenreihe für Vegetationskunde 28: 189–
306.
[What is effectively a German bryophyte checklist, including conservation status of species, is available at
www.moose-deutschland.de]
Greece (including Crete)
Düll, R. 1995. Moose Griechenlands. Bryologische Beiträge 10: 1–229. (not accessed)
Phitos, D., Constantinidis, T. & Kamari, G. (eds.) 2009. The Red Data Book of rare and threatened plants of Greece.
Patras, Hellenic Botanical Society. [Contains a single species of bryophyte – Buxbaumia viridis]
Preston, C.D. 1981. A check-list of Greek liverworts. Journal of Bryology 11: 537–553.
Preston, C.D. 1984. A check-list of Greek mosses. Journal of Bryology 13: 43–95.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
Hungary
Erzberger, P. & Papp, B. 2004. Annotated checklist of Hungarian bryophytes. Studia bot. hung. 35: 91–149.
Papp, B., Erzberger, P., Ódor, P., Hock, Zs., Szövényi, P., Szurdoki, E. & Tóth, Z. 2010. Updated checklist and Red
List of Hungarian bryophytes. Studia bot. hung. 41: 31–59.
Rajczy, M. 1990. Mohák-Bryophyta. In: Rakonczai, Z. (ed.), Vöros könyv. Budapest, Akademiai Kiadó, pp. 322–325.
Iceland
Ingadóttir, Á. 1996. Válisti 1. Plöntur. Reykjavik, Náttúrufrædistofnun Íslands, 82 pp. (mosses only)
Jóhannsson, B. 2003. Checklist of bryophytes in Iceland.
Fjölrit Náttúrufræðistofnunar 44. Webpage.
http://www.floraislands.is/Annad/moslist.html (accessed 6.6.14).
17
Checklist and country status of European bryophytes
Ireland (incl. Northern Ireland)
Hill, M.O., Blackstock, T.H., Long, D.G. & Rothero, G.P. 2008. A checklist and census catalogue of British and Irish
bryophytes updated 2008. Middlewich, British Bryological Society.
Lockhart, N., Hodgetts, N.G. & Holyoak, D.T. 2012a. Rare and threatened bryophytes of Ireland. Belfast, Ulster
Museum. (includes Red List).
Lockhart, N., Hodgetts, N. & Holyoak, D. 2012b. Ireland Red List No. 8. Bryophytes. Mosses, liverworts & hornworts.
Dublin, National Parks & Wildlife Service.
Italy (including Sicily and Sardinia)
Aleffi, M. 2005. New check-list of the Hepaticae and Anthocerotae of Italy. Flora Mediterranea 15: 485–566.
Aleffi, M. & Schumacker, R. 1995. Check-list and red-list of the liverworts (Marchantiophyta) and hornworts
(Anthocerotophyta) of Italy. Flora Mediterranea 5: 73–161.
Cortini-Pedrotti, C. 1992. Check-list of the mosses of Italy. Flora Mediterranea
http://dbiodbs.univ.trieste.it/global/mosses1#init (accessed 9 June 2011))
2: 119–221. (see also
Cortini Pedrotti, C. & Aleffi, M. 1992. Lista rossa della briofite d'Italia. In: Conti, F., Manzi, A. & Pedrotti, F. (eds.),
Libro rosso delle piante d'Italia. Roma, WWW & Società Botanica Italiana, pp. 559–687.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Kazakhstan (mostly in Asia – small part in Europe)
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Kosovo
No list known (but included in Serbia by Sabovljević et al. 2008)
Latvia
Abolina, A.A. 1994. Latvijas retas un aizsargajamas sunas. Riga, LU Ekologiska Centra Apgads “VIDE”, 24 pp.
Abolina, A.A. 2001. Latvijas sūnu saraksts [List of bryophytes of Latvia]. Latvijas veģetācija 3: 47–87. [Also
available at http://latvijas.daba.lv/scripts/db/saraksti/saraksti.cgi?d=suunas (accessed 13.10.11)]
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Also Latvian Red List available at http://biodiv.lvgma.gov.lv/cooperation/fol288846/fol707680 (accessed 13.10.11).
Liechtenstein
Senn, H.-P. 2000. Die Moose des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft
Liechtenstein-Sargans-Werdenberg 27: 7–248.
Lithuania
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Jukoniene, I. 1996. Rare and threatened bryophyte species in Lithuania. Bot. Lithuanica 2: 327– 342.
Jukoniene I., 2002. Checklist of Lithuanian mosses [Lietuvos lapsamanių sąvadas]. – Botanica Lithuanica 8(4): 303–
322.
Naujalis, J., Kalinauskaite, N. & Grineviciene, M. 1995. Vadovas Lietuvos kerpsamanėms pazinti. Vilnius.
Rašomavičius V.(ed.), 2007. Lietuvos raudonoji knyga. 262–354. Vilnius.
18
Checklist and country status of European bryophytes
Luxembourg
Werner, J. 2003. Liste rouge des bryophytes du Luxembourg. Mesures de conservation et perspectives. Ferrantia
35: 1–71.
Werner, J. 2014. Check-list et liste rouge des bryophytes du Luxembourg. [Checklist and Red List of the bryophytes of
Luxembourg]. Webpage. http://sci.mnhn.lu/colsci/weje/pdf/checkliste2014.pdf (accessed 6.6.14).
Macedonia
Cekova, M. 2005. Pregled na brioflorata na republika Makedonija [Survey of bryophyte flora of R. Macedonia]. PMF,
Institut za biol., 40 pp.
Martinčič, A. 2009. Contributions to the bryophyte flora of Republic of Macedonia. Hacquetia 8/2: 97–114.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
Malta
Frahm, J.-P. & Lüth, M. 2008. The bryophyte flora of the Maltese Islands. Archive for Bryology 29: 1–10.
Moldova
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Monaco
No list known, although there are some liverworts mentioned by Söderström et al. (2002, 2007).
Montenegro
Dragićević, S. & Veljić, M. 2006. Pregled mahovina Crne Gore [Survey of Bryophyta of Montenegro]. Prirodnjački
Muzej Crne Gore [Natural History Museum of Montenegro], Podgorica, 99 pp.
Erzberger, P., Papp, B. & Dragićević, S. 2008. Notes on some newly recorded bryophytes from Montenegro.
Journal of Bryology 30: 167–170.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Sabovljević, M. 2000. Checklist of hepatics of the Federal Republic of Yugoslavia. Lindbergia 25: 37–42. (Data
separated for Serbia and Montenegro)
Sabovljević, M., Cvetić, T. & Stevanović, V. 2004. Bryophyte Red List of Serbia and Montenegro. Biodiversity and
Conservation 13: 1781–1790.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207 –244.
Sabovljević, M. & Stevanovic, V. 1999. Moss conspectus of the Federal Republic of Yugoslavia. Flora Mediterranea
9: 65–95. (Data separated for Serbia and Montenegro)
19
Checklist and country status of European bryophytes
Netherlands
Siebel, H.N., During, H.J. & van Melick, H.M.H. 2005. Veranderingen in de Standaardlijst van de Netherlandse
blad-, lever- en hauwmossen [Checklist of Dutch bryophytes and liverworts]/ Buxbaumiella 73: 26–54.
Siebel, H.N., Bijlsma, R.J. & Bal, D. 2006. Toelichting op de Rode Lijst Mossen. Ministry of Agriculture, Nature and
food quality, report DK2006/034. (see also www.verspreidingsatlas.nl/soortenlijst.aspx?groep=B (accessed 9
June 2011))
Norway
Mainland
Direktoratet for naturforvalning (DN). 1999. Nasjonal rødliste for truete arter 1998 [Norwegian Red List 1998]. DNrapport 1999–3: 1–161. See also www.nhm.uio.no/botanisk/mose/red.htm (accessed 9 June 2011)
Hassel, K., Blom, H.H., Flatberg, K.I., Halvorsen, R. & Johnsen, J.I. 2010. Moser. Anthocerophyta,
Marchantiophyta, Bryophyta. In: Kålås, J.A., Viken, Å., Henriksen, S. and Skjelseth, S. (eds.) 2010. The 2010
Norwegian Red List for Species. Norwegian Biodiversity Information Centre. (See also www.artsdatabanken.no
and http://www.biodiversity.no/Article.aspx?m=279&amid=12279.
Up-to-date checklist available at the home pages of The Norwegian Biodiversity Information Center
http://www2.artsdatabanken.no/artsnavn/Contentpages/Sok.aspx.
Svalbard
Frisvoll, A.A. & Blom, H.H. 1997. Trua moser i Noreg med Svalbard. Førebelse ark. NTNU, Vit.- mus., Bot. Notat
1997–3: 1–170.
Frisvoll, A.A. & Elvebakk, A. 1996. Part 2. Bryophytes. In: Elvebakk, A & Prestrud, P. (eds.) A catalogue of
Svalbard plants, fungi, algae and cyanobacteria, pp. 57–172. Oslo, Norwegian Polar Institute.
Poland
Klama, H. 2006. Red list of the liverworts and hornworts in Poland, Czerwona lista watrobowców i glewików w
Polsce. In: Mirek, Z., Zarzycki, K. Wojewoda, & W. Szelag, Z. Red list of plants and fungi in Poland, Czerwona
lista roslin i grzybów Polski. Kraków, Polish Academy of Sciences, W. Szafer Institute of Botany.
Ochyra, R. 1992. Czerwona lista mchów zagroonych Polsce (Red list of threatened mosses in Poland). Kraków, Polish
Academy of Sciences, pp. 79–85.
Ochyra, R., Zarnowiec, J. & Bednarek-Ochyra, H. 2003. Census Catalogue of Polish mosses. Kraków, Polish Academy
of Sciences, W. Szafer Institute of Botany.
Szweykowski, J. 1992. Czerwona lista watrobowców zagroonych w Polske [List of threatened liverworts in Poland]. In:
Zarzycki K., Wojewoda W. & Heinrich Z. (eds.), List of threatened plants in Poland (2 nd ed.). Kraków, W. Szafer
Institute of Botany, Polish Academy of Sciences, Kraków, pp.75–78.
Szweykowski, J. 2006. An annotated checklist of Polish liverworts and hornworts. Kraków, Polish Academy of
Sciences, W. Szafer Institute of Botany.
Zarnowiec, J., Stebel, A. & Ochyra, R. 2004. Threatened moss species in the Polish Carpathians in the light of a
new Red List of mosses in Poland. In: Stebel, A. & Ochyra, R. (eds.), Bryological studies in the Western
Carpathians. Poznań, Sorus.
Portugal
Mainland
Sérgio, C., Casas, C., Brugués, M. & Cros, R.M. 1994. Lista Vermelha dos Briófitos da Península Ibérica [Red List of
Bryophytes of the Iberian Peninsula]. Lisboa, ICN, 45 pp.
Sérgio, C. & Carvalho, S. 2003. Annotated catalogue of Portuguese bryophytes. Portugaliae Acta Biologica 21: 5–
230. (Sérgio, pers. comm. 13.6.11)
Sérgio, C., Brugués, M., Cros, R.M. Casas, C. & Garcia, C. 2006. The 2006 Red List and an updated checklist of
bryophytes of the Iberian Peninsula (Portugal, Spain and Andorra). Lindbergia 31: 109–125.
Sérgio, C., Garcia, C.A., Sim-Sim, M., Vieira, C., Hespanhol, H. & Stow, S. 2013. Atlas e Livro Vermelho dos Briófitos
Ameaçados de Portugal. Lisboa, Universidade de Lisboa – Museu Nacional de História Natural e da Ciência.
20
Checklist and country status of European bryophytes
Azores
Gabriel, R., Sjögren, E., Schumacker, R., Sérgio, C., Frahm, J.-P. & Sousal, E. 2005. 4.1. Lista dos Briófitos
(Bryophyta) [List of Bryophytes (Bryophyta)]. In: Lista de espécies terrestres dos Açores [List of terrestrial species
from the Azores].
Gabriel, R., Sjögren, E., Schumacker, R., Sérgio, C., Aranda, S.C., Claro, D., Homem, N. & Martins, B. 2010. List of
bryophytes (Anthocerotophyta, Marchantiophyta, Bryophyta). In: PAV. Borges, A. Costa, R. Cunha,
R.Gabriel, V. Gonçalves, A.F. Martins, I. Melo, M. Parente, P. Raposeiro, P. Rodrigues, R.S. Santos, L. Silva, P.
Vieira & V. Vieira (eds.) A list of the terrestrial and marine fungi, flora and fauna from the Azores. pp. 99–115,
Princípia, Cascais. (Gabriel, pers. comm. 15.6.11)
Gabriel, R., Homem, N., Couto, A., Aranda, S.C. & Borges, P.A.V. (submitted). Azorean Bryophytes: a
preliminary review of rarity patterns. Açoreana (Gabriel, pers. comm. 15.6.11)
Madeira
Sérgio, C., Schumacker, R., Fontinha, S. & Sim-Sim, M, 1992. Evaluation of the status of the bryophyte flora of
Madeira with reference to endemic and threatened European species. Biological Conservation 59: 223–231.
Sérgio, C., Sim-Sim, M. & Carvalho, M. 2006. Annotated catalogue of Madeiran bryophytes. Bol. Mus. Munic.
Funchal 10: 5–164.
Sérgio, C., Sim-Sim, M., Fontinha, S. & Figueira, R. 2008. 5.1 Lista dos briófitos (Bryophyta) (List of bryophytes
(Bryophyta)). In: Borges, P.A.V. et al. Listagem dos fungos, flora e fauna terrestres dos arquipélagos da
Madeira e Selvagens (A list of the terrestrial fungi, flora and fauna of Madeira and Selvagens archipelagos).
Funchal, Direcção Regional do Ambiente do Governo Regional da Madeira. (Sérgio, pers. comm. 13.6.11)
Sim-Sim, M., Luis, L., Garcia, C., Fontinha, S., Lobo, C., Martins, S. & Stech, M. 2008. New data on the status of
threatened bryophytes of Madeira Island. Journal of Bryology 30: 226–228.
Romania
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
Ştefănuţ, S. 2008. The hornwort and liverwort atlas of Romania. Bucharest, Edit. Ars Docendi, 510 pp. (see also
www.ibiol.ro/bucegia/hepatice/redlist.htm (accessed 9 June 2011))
Ştefănuţ S. & Goia, I. 2012. Checklist and Red List of Bryophytes of Romania. Nova Hedwigia 95 (1–2): 59–104.
Russia
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Konstantinova, N.A. & Bakalin, V.A. 2009. Checklist of liverworts (Marchantiophyta) of Russia. Arctoa 18: 1–64.
Bardonov, L.V. & Ignatov, M.C. 2005. Bryophytes. In: Red Data Book of the Russian Federation. (see also:
http://en.wikipedia.org/wiki/Red_Data_Book_of_the_Russian_Federation)
Bashkortostan
Baisheva, E.Z. 2002. Bryophyta. In: Red Data Book of the Bashkortostan Republic. Vol. 2. Bryophytes, algae, lichens and
fungi. Ufa, Tabigat: 17–44.
Dagestan
Konstantinova, N.A. 2011. Contribution to the hepatic flora of the Republic of Dagestan (Eastern Caucasus,
Russia). Arctoa 20: 175–182.
Radzhi, A.D. 1998. Bryophytes. In: Red book of Dagestan. Rare and endangered species of animal and plants.
Makhachkala, Dagestanskoe Knizhnoe Izd.: 191–194.
Kabardino-Balkar
Shkhagapsoev, S.H. 2000. Bryophytes. In: Red book of Kabardino-Balkar Republic. Nalchik, Ael'-Fa]: 298.
21
Checklist and country status of European bryophytes
Karelia
Maksimov, A.I., Potemkin, A.D. & Maksimova, T.A. 2007. Bryophytes. In: Ivanter, E.V. & Kuznetsov, O.L. (eds.).
Red Data Book of the Republic of Karelia. Ministerstvo sel’skogo, rybnogo khozyaijstva i ekologii Respublici
Karelia, Karel’skiij nauchnyij tsentr Rossiijskoij akademii hauk, Petrozavodskiij gosudarstvennyij universitet.
Petrozavodsk, “Karelia”:79–98. This is also a checklist.
Komi
Taskaeva, A.I. 1998. Krasnaja kniga Respubliki Komi. Moskva, Syktyvkar: Izdatelstvo DIK.
Krasnodar
Konstantinova, N.A., Akatova, T.V., Ignatova, E.A. & Ignatov, M.S. 2008. Bryophyta. In: Red Data Book of the
Krasnodar Province. (Plants and fungi). 2d ed. Krasnodar, Dizain byuro No. 1: 450–477.
Kursk Province
Popova, N.N. 2001. Bryophyta . In: Red Data Book of Kursk Province. Vol. 2. Rare and endangeded plants and fungi.
Tula, Centralno-Czernozemnyj Gos. Prir. Biospher. Zapovednik & al.: 21–45.
Lipetsk
Popova, N.N & Abramova, L.I. 2005. Bryophytes. In: Novikov, V.S. (ed.). Red Data Book of Lipetsk Province. Vol. 1.
Plants, Fungi, Lichens. Moscow, KMK: 15–70.
Moscow (City)
Ignatov, M.S. 2001. Bryophyta. In: Red Data Book of Moscow City. Moscow Government: 543–567.
Moscow(Region)
Ignatov, M.S. 1998. Bryophytes. In: Zubakin, V.A. & Tikhomirov, V.N. (eds.). Red Data Book of Moscow Province.
Moscow, Argus & Russki Univ.
Ignatov, M.S. 2008. Bryophytes. In: Varlygina, T.I., Zubakin, V.A. & Sobolev, N.A. (eds.). Red Data Book of Moscow
Province. 2d edition. Moscow.
Murmansk
Konstantinova, N.A., Belkina, O.A. & Likhachev, A.Y. 2003. Bryophytes - In: Konstantinova, N.A., Koryakin, A.S.
& Makarova, O.A. (eds.). Red Data book of the Murmansk Province. Murmansk, Murmanskoe Knizhnoe
Izdatelstvo.
Nenetsky
Afonina, O.M. & Konstantinova, N.A. (no date) Bryophytes. In: Matveeva, N.V. (ed.) Red Data Book of Nenetsky
Autonomous District. Official edition. Naryan-Mar: 108–125
Ryazan
Volosnova, L.P. et al. 2003. Bryophyta. In: Red book of Ryazan Province. Ryazan, Uzoroch'e: 27–36.
St. Petersburg
Andreeva, E.N., Afonina, O.M., Kuzmina, E.O. & Kurbaova, L.E. 2004. Bryophyta. In: Red Data Book of St.Petersburg. St.-Petersburg: 325–342.
Tzvelev, N.N. 2000. Red Data Book of nature of the Leningrad region. Vol. 2 - Plants and Fungi. St. Petersburg, 671 pp.
St. Petersburg, Karelia & Murmansk
Kotiranta, H., Uotila, P., Sulkava, S. & Peltonen, S.-L. (eds.) 1998. Red Data Book of East Fennoscandia. Helsinki,
Ministr. Envir., 351 pp.
Tver’
Zykov, I.V., Notov, A.A. & Spirina, U.N. 2002. Bryophyta. – Divisio Bryophytes. In: Red Data Book of the Tver’ Province.
Tver’, Veche Tveri & ANTEK: 10–35.
Several other provinces also have checklists (Lars Söderström, pers. comm., May 2012).
San Marino
Zodda, G. 1930. In: Pampanini, R. (ed.). Flora della Repubblica di San Marino. San Marino pp. 1–68.
22
Checklist and country status of European bryophytes
Serbia
Sabovljević, M. 2000. Checklist of hepatics of the Federal Republic of Yugoslavia. Lindbergia 25: 37–42. (Data
separated for Serbia and Montenegro)
Sabovljević, M. & Stevanovic, V. 1999. Moss conspectus of the Federal Republic of Yugoslavia. Flora Mediterranea
9: 65–95. (Data separated for Serbia and Montenegro)
Sabovljević, M., Cvetić, T. & Stevanović, V. 2004. Bryophyte Red List of Serbia and Montenegro. Biodiversity and
Conservation 13: 1781–1790.
Ros, R.M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T.L., Brugués, M., Cano, M.J., Cros, R.M., Dia,
M.G., Dirkse, G.M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J.M., Herrnstadt, I., Khalil, K.,
Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai. M.S., Rodríguez-Nuñez, S., Sabovljević, M., Sérgio, C.,
Shabbara, H., Sim-Sim, M. & Söderström, L. 2007. Hepatics and Anthocerotes of the Mediterranean, an
annotated checklist, Cryptogamie, Bryologie 28 (4): 351–437.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
Slovakia
Kubinská, A., Janovicová, K. & Peciar, V. 1993. The checklist of bryophytes in Slovakia. Biológia, Bratislava 48: 99–
143.
Kubinská, A., Janovicová, K. & Peciar, V. 1996. The list of extinct, missing and threatened bryophytes (Bryophyta)
of Slovakia (1st version). Biológia, Bratislava 51: 373–380.
Kubinská, A. & Janovicová, K. 1998. Bryophytes, pp. 297–332. In: Marhold, K. & Hindák, F. (eds). Checklist of nonvascular and vascular plants of Slovakia. Veda, Bratislava (see also
http://ibot.sav.sk/checklist/index.php?lang=en).
Kubinská, A., Janovicová, K. & Soltes, R. 2001: Red list of bryophytes of Slovakia (December 2001). In: Baláz, D., et
al. (eds)., Cerveny zoznam rastlín a zivocichov Slovenska. Ochr. Prír. 20 (Suppl.): 48–81.
Kubinská, A., Janovicová, K. & Soltes, R. 2001. Updated checklist of liverworts, hornworts and mosses of
Slovakia. Bryonora, Praha 28: 4–10.
Slovenia
Martinčič, A. 1992. Rdeci seznam ogroženih listnatih mahov (Musci) v Sloveniji. Varstvo Narave 18: 1–190. (Red
List; mosses only)
Martinčič, A. 2003. Seznam listnatih mahov (Bryopsida) Slovenije [Survey of mosses of Slovenia]. Hacquetia 2 (1):
91–166.
Spain
Mainland
Garilleti, R. & Albertos, B. 2012. Atlas y Libro Rojo de los Briófitos Amenazados de España. Ministerio de Agricultura,
Alimentacion y Medio Ambiente, Madrid. 287 pp.
Sérgio, C., Casas, C., Brugués, M. & Cros, R.M. 1994. Lista Vermelha dos Briófitos da Península Ibérica [Red List of
Bryophytes of the Iberian Peninsula]. Lisboa, ICN, 45 pp.
Sérgio, C., Brugués, M., Cros, R.M. Casas, C. & Garcia, C. 2006. The 2006 Red List and an updated checklist of
bryophytes of the Iberian Peninsula (Portugal, Spain and Andorra). Lindbergia 31: 109–125. (see also
http://pagines.uab.cat/briologia/en/content/bryophyte-lists)
Baleares
Cros, R.M., Saez, L. & Brugués, M. 2008. The bryophytes of the Balearic Islands: an annotated checklist. Journal of
Bryology 30: 74–95. (see also http://pagines.uab.cat/briologia/en/content/bryophyte-lists)
23
Checklist and country status of European bryophytes
Canary Isles
Dirkse, G.M., Bouman, A.C. & Losada-Lima, A. 1993. Bryophytes of the Canary Islands, an annotated checklist.
Cryptogamie Bryologie Lichenologie 14 (1): 1–47.
Dirkse, G.M. & Losada-Lima, A. 2011. Additions and amendments to the moss flora of the Canary Islands.
Cryptogamie, Bryologie 32 (1): 37–41.
González-Mancebo, J.M., Romaguera, F., Ros, R.M., Patiño, J. & Werner, O. 2008. Bryophyte flora of the Canary
Islands: an updated compilation of the species list with an analysis of island distribution patterns in the
context of the Macaronesian Region. Cryptogamie, Bryologie 29 (4): 315–357.
Sweden
Gärdenfors, U. (ed.) 2010. Rödlistade arter i Sverige 2005 [The 2010 Red List of Swedish species]. Uppsala,
ArtDatabanken, 590 pp. (See also http://www.artfakta.se, accessed 6.6.14)
Hallingbäck, T., Hedenäs, L. & Weibull, H. 2006. Ny checklista för Sveriges mossor [Checklist of bryophytes
recorded from Sweden]. Svensk Botanisk Tidskrift 100 (2): 96–148. (See also
http://www.sbf.c.se/MV/Moss_checklista_2006.pdf, accessed 6.6.14)
Switzerland
Schnyder, N., Bergamini, A., Hofman, H., Müller, N., Schubiger-Bossard, C. & Urmi, E. 2004. Rote Liste der
gefährdeten Moose der Schweiz. Bern, Hrsg. BUWAL, FUB & NISM. BUWAL-reihe: Vollzug umwelt, 99 pp.
Anon. 2011. National inventory of Swiss bryophytes. Zürich, Institut für Systematische Botanik. Webpage.
www.nism.uzh.ch/index.php?content=einleitung&lang=en (accessed 9 June 2011)
Turkey (mostly in Asia – small part in Europe)
Kürschner, H. & Erdağ, A. 2005. Bryophytes of Turkey: an annotated reference list of the species with synonyms
from the recent literature and an annotated list of Turkish bryological literature. Turkish Journal of Botany 29:
95–154.
Sabovljević, M. & Natcheva, R. 2006. Check list of the liverworts and hornworts of South-Eastern
Europe. Phytologia Balcanica 12 (2): 169–180
Sabovljević, M., Natcheva, R., Dihoru, G., Tsakiri, E., Dragićević, S., Erdağ, A. & Papp, B. 2008. Check-list of the
mosses of SE Europe. Phytologia Balcanica 14 (2): 207–244.
Uyar, G. & Çetin, B. 2004. A new check-list of the mosses of Turkey. Journal of Bryology 26: 203–220.
Ukraine
Ignatov, M.S., Afonina, O.M., Ignatova, E.A. et al. 2006. Check-list of mosses of East Europe and North Asia.
Arctoa 15: 1–130.
Sheljag-Sosonka, J.R. (ed.) 1996. Kniga Ukraïni. Roslinnij svit. Kiev, Vidavnictvo "Ukrains'ka Enciklopedija" imeni
M.P. Bazana.
Váňa, J. & Virchenko, V.M. 1993. A list of Anthocerotes and Hepatics of Ukraine. Ukrajins’kyj Botaničnyj Žurnal 50:
89–93.
United Kingdom
Hill, M.O., Blackstock, T.H., Long, D.G. & Rothero, G.P. 2008. A checklist and census catalogue of British and Irish
bryophytes updated 2008. Middlewich, British Bryological Society.
Great Britain (incl. England, Scotland, Wales & Isle of Man)
Church, J.M., Hodgetts, N.G., Preston, C.D. & Stewart, N.F. 2001. British Red Data Books. Mosses and liverworts.
Peterborough, JNCC, 168 pp.
Hodgetts, N.G. 2011. A revised Red List of bryophytes in Britain. Field Bryology 103: 40–49.
Northern Ireland
See Ireland
24
Checklist and country status of European bryophytes
Channel Islands
Hill, M.O., Blackstock, T.H., Long, D.G. & Rothero, G.P. 2008. A checklist and census catalogue of British and Irish
bryophytes updated 2008. Middlewich, British Bryological Society.
Gibraltar
No list known, although there are some liverworts mentioned by Söderström et al. (2002, 2007).
Vatican
No list known, although there are some liverworts mentioned by Söderström et al. (2002, 2007).
25
Checklist and country status of European bryophytes
Appendix 2. European bryological societies and journals
Societies
British Bryological Society
Area covered: Britain and Ireland
Website: http://rbg-web2.rbge.org.uk/bbs/bbs.htm
Journals/Newsletters: Journal of Bryology; Field Bryology
Bryological Association of South-Eastern Europe
Area covered: SE Europe, principally the Balkans
Website: - (only old address found)
Contact: Dr. Marko Sabovljevic, Ass't. Prof. Insitute of Botany and Garden, Faculty of Biology, University of
Belgrade, Takovska 43, 11000 Belgrade, Serbia ([email protected],)
Journals/Newsletters: Bryologisch-lichenologische Arbeitsgemeinschaft für Mitteleuropa (BLAM) (The Bryological and Lichenological
Working Group of Central Europe)
Area covered: Central Europe
Website: http://www.blam-hp.eu/home_en.html
Journals/Newsletters: Herzogia
Czech Botanical Society – Bryological/Lichenological Section
Area covered: Czech Republic
Website: http://botanika.bf.jcu.cz/BLS/english/index.html
Journals/Newsletters: Bryonora
Dutch Bryological and Lichenological Society (BLWG)
Area covered: Netherlands
Website: http://www.blwg.nl/
Journals/Newsletters: Lindbergia (with Nordic Bryological Society); Buxbaumiella
Finnish Bryological Society
Area covered: Finland
Website: http://pro.tsv.fi/sammalseura/cd.html
Journals/newsletters: Bryobrothera; Bryobrotherella
Mossornas Vänner
Area covered: Sweden, Finland, Denmark, Norway & Iceland
Website: http://www.sbf.c.se/MV/
Journals/Newsletters: Myrinia
Nordic Bryological Society
Area covered: Denmark, Finland, Iceland, Norway, & Sweden
Website: http://www.sbf.c.se/NBS/Root.htm
Journals/Newsletters: Lindbergia (with Dutch Bryological and Lichenological Society)
Spanish Bryological Society (Sociedad Española de Briología)
Area covered: Spain
Website: http://www.uam.es/informacion/asociaciones/SEB/
Journals/Newsletters: Boletín de la Sociedad Española de Briología
Swiss Association of Bryology and Lichenology
Area covered: Switzerland
Website: http://www.bryolich.ch/english/index_engl.html
Journals/Newsletters: Meylania
26
Checklist and country status of European bryophytes
Other Journals or newsletters
Arctoa
Area covered: principally Russia
Website: http://www.arctoa.ru/en/
Cryptogamie, Bryologie
Area covered: international, but based in France
Website: http://www.cryptogamie.com/pagint_en/editeur/revue_bryo.php#
Limprichtia
Area covered: western and central Europe.
Website: http://www.jan-peter-frahm.de/Limprichtia/Limprichtia_vols.htm
Muscillanea
Area covered: Belgium
Website: Contact: Herman Stieperaere, Nationale Plantentuin van Belgie, Domein de Bouchout, B-1860 Meise, Belgium.
Nov. System Plant non Vasc
Website: Nowellia Bryologica
Area covered: Belgium
Website: http://users.skynet.be/fb062663/nowellia.htm
Phytologia Balconica
Area covered: the Balkans; mainly vascular plants but occasional bryological papers.
Website: http://www.bio.bas.bg/~phytolbalcan/
27
Checklist and country status of European bryophytes
Appendix 3. Floras
There are many bryophyte Floras covering various geographical areas in Europe. The more relevant and modern
ones, or the latest known for a territory, are presented here. Only major Floras are listed, not Floras of small
subdivisions of countries, and not publications dealing only with a small taxonomic group of plants.
Atherton, I., Bosanquet, S. & Lawley, M. (eds.) 2010. Mosses and liverworts of Britain and Ireland. A field guide.
Plymouth, British Bryological Society.
Casas, C., Brugués, M., Cros, R.M. & Sérgio, C. 2006. Handbook of the mosses of the Iberian Peninsula and the Balearic
Islands. Barcelona, Institut d’Estudis Catalans.
Casas, C., Brugués, M., Cros, R.M., Sérgio, C. & Infante, M. 2009. Handbook of liverworts and hornworts of the Iberian
Peninsula and the Balearic Islands. Barcelona, Institut d’Estudis Catalans.
Cortini Pedrotti, C. 2001. Flora dei Muscha d’Italia, Parte 1: Sphagnopsida, Andreaeopsida, Bryopsida. Casalini Libri.
Cortini Pedrotti, C. 2005. Flora dei Muscha d’Italia, Parte 2: Bryopsida. Antonio Delfino.
Frey, W., Frahm, J.-P., Fischer, E. & Lobin, W. 2006. The liverworts, mosses and ferns of Europe. English edition
revised and edited by T.L. Blockeel. Colchester, Harley Books.
Grims, F. 1999. Die Laubmoose Osterreichs. Verlag der Osterreichischen Akademie der Wissenschaften.
Guerra, J. (ed.) 2007–2012. Flora Briofitica Ibérica. Vols. 1–6. Sociedad Espanola de Briologia.
Hallingbäck, T., Lonnell, N. & Weibull, H. 2006. Encyclopedia of the Swedish Flora and Fauna: Bladmossor:
Sköldmossor–Blåmossor. Bryophyta: Buxbaumia–Leucobryum. Uppsala, ArtDatabanken (Swedish Threatened
Species Unit).
Hallingbäck, T., Lonnell, N. & Weibull, H. 2008. Encyclopedia of the Swedish Flora and Fauna: Bladmossor:
Kompaktmossor–Kapmossor. Bryophyta: Anoectangium–Orthodontium. Uppsala, ArtDatabanken (Swedish
Threatened Species Unit).
Hedenäs, L. 1992. Flora of Madeiran pleurocarpous mosses (Isobryales, Hypnobryales, Hookeriales). Gebrüder
Borntraeger Verlag.
Ignatov, M.S. & Ignatova, E.A. 2003. Moss Flora of the middle part of European Russia, Vol. 2: Fontinalaceae–
Amblystegiaceae. KMK Scientific Press.
Johannsson, B. 1996-… Icelandic bryophytes (various). Icelandic Museum of Natural History.
Jukoniene, I. 2003. Mosses of Lithuania. Institute of Botany.
Lewinsky, J. 1987. The vegetation and bryophyte flora of the Faroe Islands (Denmark): excursion guide…..?
Nyholm, E. 1993. Illustrated Flora of Nordic mosses. Fasc. 3. Oikos editorial office.
Paton, J.A. 1999. The liverwort flora of the British Isles. Colchester, Harley Books.
Smith, A.J.E. 1990. The liverworts of Britain and Ireland. Cambridge, Cambridge University Press.
Smith, A.J.E. 2004. The moss flora of Britain and Ireland. Second edition (2006 reprint). Cambridge, Cambridge
University Press.
28
Checklist and country status of European bryophytes
Appendix 4. ECCB Country Contacts
This table is updated from versions supplied by ECCB members at the beginning of the pilot project.
Country
Name
Institute
e-mail
Albania
Marka, Jani
Department of Biology
Faculty of Natural Sciences
Tirana University
[email protected]
Andorra
Brugués, Domenech
Montserrat
[email protected]
Austria
Zechmeister, Harald
Azores (Portugal)
Gabriel, Rosalina
Baleares Islands
(Spain)
Brugués, Domenech
Montserrat
Belgium
Raeymaekers, Geert
Botanica, Facultat de
Ciencies, Universitat
Autonoma de Barcelona
University of Vienna,
Faculty of Life
Sciences, Dept. of
Conservation Biology,
Vegetation- and Landscape
Ecology
Secção de Biologia Depto.
Ciências Agrárias, Angra
do Heroísmo
Botanica, Facultat de
Ciencies, Universitat
Autonoma de Barcelona
Belarus
Maslovsky, Oleg
Bosnia-Herzegovina
Sabovljević, Marko
Britain (England,
Scotland, Wales, Isle
of Man)
Bulgaria
Hodgetts, Nick
Canary Island
(Spain)
Dirkse, Gerard M.
Caucasus
Channel Islands
(Britain)
Corsica (France)
Ganeva, Anna
González-Mancebo,
Juana María
29
[email protected]
[email protected]
Institute of Experimental
[email protected]
Botany Academy of
Science, Minsk
Institute of Botany and
[email protected]
Garden, Faculty of Biology,
University of Belgrade
[email protected]
Institute of Botany Acad. ,
Sofia
Natuurmuseum Nijmegen
[email protected]
Universidad de La
Laguna, Dept de
Botánica, Fac. De
Biologia, Tenerife
[email protected]
[email protected]
[email protected]
Hodgetts, Nick
[email protected]
Hugonnot, Vincent
Conservatoire Botanique
National du Massif
Central, ChavaniacLafayette
Muséum nationale
d'Histoire naturelle, Paris
Aristotle University of
Thessaloniki Department
of Botany, Thessaloniki
[email protected]
Alegro, Antun
University of Zagreb
[email protected]
Sabovljević, Marko
Institute of Botany and
[email protected]
Tsakiri, Eudoxia
Caucasus
Croatia
[email protected]
Losada-Lima, Anna
Leblond, Sebastien
Crete (Greek)
[email protected]
[email protected]
[email protected]
Checklist and country status of European bryophytes
Country
Czech Republic
Name
Institute
e-mail
Váňa, Jiři
Garden, Faculty of Biology,
University of Belgrade
Charles University
[email protected]
Department of Botany and
Ecology, Praha
Cyprus
Denmark
Frederiksen, Rasmus Danish Bryological Society
Fuglsang
(Bryologkredsen)
Ingerpuu, Nele
University of Tartu
Institute of Ecology and
Earth Sciences Department
of Botany
Vellak, Kai
University of Tartu
Institute of Ecology and
Earth Sciences Department
of Botany
[email protected]
Faroe Islands
(Denmark)
Finland
Syrjänen, Kimmo
Finnish Environment
Institute Species Protection
Unit, Helsinki
[email protected]
France
Hugonnot, Vincent
Conservatoire Botanique
National du Massif
Central, ChavaniacLafayette
Muséum nationale
d'Histoire naturelle, Paris
[email protected]
Zentrum für
Biodokumentation Am
Bupwerk Reden,
Schiffweiler
[email protected]
Estonia
Leblond, Sébastien
[email protected]
[email protected]
[email protected]
Germany
Caspari, Steffen
Gibraltar (Britain)
Hodgetts, Nick
Greece
Tsakiri, Eudoxia
Aristotle University of
Thessaloniki Department
of Botany, Thessaloniki
Hungary
Papp, Beáta
Hungarian Natural History [email protected]
Museum Botanical
Department, Budapest
Iceland
Kristinsson, Hördur
Náttúrufræðistofnun
Íslands, Reykjavík
Ireland
Lockhart, Neil
Italy
Aleffi, Michele
National Parks and
[email protected]
Wildlife Service,
Department of Arts,
Heritage and the Gaeltacht,
Dublin 2
Dipartimento di Botanica
[email protected]
ed Ecologia, Laboratorio di
Briologia, Università degli
Studi di
Camerino, Camerino
[email protected]
[email protected]
[email protected]
30
Checklist and country status of European bryophytes
Country
Name
Institute
e-mail
Kosovo
Sabovljević, Marko
Latvia
Mežaka, Anna
Institute of Botany and
[email protected]
Garden, Faculty of Biology,
University of Belgrade
Department of Botany and [email protected]
Ecology, Faculty of
Biology, University of
Latvia,
Kaliningrad Area
Kazakhstan
Liechtenstein
Lithuania
Jukonienė, Ilona
Luxembourg
Werner, Jean
Macedonia
Sabovljević, Marko
Madeira (Portugal)
Fontinha, Suzana
Simsim, Manuela
Malta
Institute of Botany,
Laboratory of Flora and
Geobotany, Vilnius
[email protected]
[email protected]
Institute of Botany and
[email protected]
Garden, Faculty of Biology,
University of Belgrade
Jardim Botanico de
[email protected]
Madeira, Funchal
Universidade de Lisboa,
Faculdade de Ciências de
Lisboa,
[email protected]
Lafranco, Edwin
Mifsud, Stephan
[email protected]
Moldova
Monaco
Hugonnot, Vincent
Conservatoire Botanique
National du Massif
Central, ChavaniacLafayette
Leblond, Sebastien
Montenegro
31
[email protected]
[email protected]
Dragicevic, Snezana
Natural History Museum
of Montenegro, Podgorica
Sabovljević, Marko
Institute of Botany and
[email protected]
Garden, Faculty of Biology,
University of Belgrade
[email protected]
Netherlands
Siebel, Henk
Northern Ireland
Lockhart, Neil
Norway
Hassel, Kristian
Poland
Klama, Henryk
[email protected]
National Parks and
[email protected]
Wildlife Service,
Department of Arts,
Heritage and the Gaeltacht,
Dublin 2
NTNU, Department of
[email protected]
Biology, Trondheim
Technichal University of
Łódź, Branch in BielskoBiała, Faculty of Textile
Engineering and
Environmental Protection,
Bielska-Biała
[email protected]
Checklist and country status of European bryophytes
Country
Name
Institute
e-mail
Żarnowiec, Jan
Technichal University of
Łódź, Branch in BielskoBiała, Faculty of Textile
Engineering and
Environmental Protection,
Bielska-Biała
Museu Laboratorio
Botânico, Faculdade de
Ciências, Lisboa
Grădina Botanică “Al.
Borza”, Universitatea
“Babeş-Bolyai”, ClujNapoca
Institute of Biology,
Romanian
Academy, Bucharest
The Polar-Alpine Botanical
Garden, Kola Science
Center of the Russian
Acad. Sci., Murmansk Obl.
Main Botanical Garden,
Academy of Science
Botanicheskaya, Moscow
Dipartimento di Botanica
ed Ecologia, Laboratorio di
Briologia, Università degli
Studi di
Camerino, Camerino
Institute of Botany and
Garden, Faculty of Biology,
University of Belgrade
University - DiSciVA sez.
Botanical and Botanical
Garden, Cagliary
Department of
Environmental Biology and
Biodiversity, University of
Palermo
[email protected]
Research Station of the
Tatra National Park,
Tatranská Lomnica
Biotechnical faculty,
Department of biology
Aškerčeva, Ljubljana
Institute of Botany and
Garden, Faculty of Biology,
University of Belgrade
Departamento de Botánica
Universidad de
Valencia, Burjassot
Botanica, Facultat de
Ciencies, Universitat
Autonoma de
Barcelona, Bellaterra
[email protected]
Portugal
Sérgio, Cecília
Romania
Goia, Irina
Stefanut, Sorin
Russia
Konstantinova,
Nadya
Ignatov, Mikhail
San Marino (Italy)
Aleffi, Michele
Serbia
Sabovljević, Marko
Sardinia (Italy)
Cogoni, Annalena
Sicily (Italy)
Campisi, Patrizia
Giovanna, Maria Dia
Slovakia
Šoltés, Rudolf
Slovenia
Martinčič, Andrej
Sabovljević, Marko
Spain
Albertos, Belén
Brugués, Domenech
Montserrat
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
[email protected]
belen. [email protected]
[email protected]
32
Checklist and country status of European bryophytes
Country
Name
Institute
e-mail
Sweden
Hallingbäck, Tomas
[email protected]
Switzerland
Schnyder, Norbert
Threatened Species Unit,
Swedish Agricultural
University, Uppsala
Institut für Systematische
Botanik, Zürich
Turkey
Kirmaci, Mesut
Adnan Menderes
University, Arts and
Science Faculty, Biology
Department, Aydin
Zonguldak Karaelmas
University, Sciences and
Arts Faculty, Department
of Biology, Zonguldak
Kherson State University,
Kherson
[email protected]
Dipartimento di Botanica
ed Ecologia, Laboratorio di
Briologia, Università degli
Studi di
Camerino, Camerino
[email protected]
Svalbard (Norway)
Uyar, Güray
33
Ukraine
Boiko, Michail
Fedoseevich
Vatican (Italy)
Aleffi, Michele
[email protected]
[email protected]
[email protected]
Checklist and country status of European bryophytes
Appendix 5. Data sheet examples for potential Red List species
Three examples have been prepared to show possible formats for data sheets in a new Red List.
Acaulon fontiquerianum Casas & Sérgio
Fontiqueri’s pygmy-moss
Family: Pottiaceae
Order: Pottiales
Subclass: Bryidae
Class: Bryopsida
Division: Bryophyta
Kingdom: Plantae
Status in Europe: Near Threatened
Criteria: -
Photo: C. Aedo
Description
Tiny acrocarpous moss, 1.5–2(–3) mm, not triquetrous, greenish-yellow to yellowish-brown, with 5–
6(–8) ovate to ovate-lanceolate leaves, not or only very slightly keeled, with nerve excurrent in an
arista 80–300(–400) µm long; margins plane. Leaf cells smooth (A. dertosense and A. casasianum
papillose). Nerve in section with 2–5 large, prominent cells on ventral side (A. muticum and A.
mediterraneum without differentiated cells on ventral side). Sporophytes immersed in concave
perichaetial leaves.
34
Checklist and country status of European bryophytes
This species was described in 1990 from Spain and Portugal (Casas & Sérgio 1990) and has
subsequently been found elsewhere in the region (Lo Giudice 1995, Sérgio et al. 1993, etc).
Ecology and biology
An annual ephemeral species of dry exposed soils, with a slight preference for acidic substrates. Often
on banks, roadsides, or at the edges of xerophytic scrub communities.
Altitudinal range: 90–1550 m
Autoicous.
European distribution
This species is known only from scattered sites in the western Mediterranean and the Canary Islands.
It is considered Vulnerable in Spain and Portugal and Endangered in Sardinia, but has no official
status in mainland France, Corsica, Sicily or the Canary Islands. It is likely to have more sites than
indicated on the distribution map, but such a small plant is very easily overlooked.
35
Checklist and country status of European bryophytes
World distribution
As far as is known, A. fontiquerianum is a European endemic, although it is not improbable that it may
be found in North Africa.
Threats
The fragility of the habitat, and its vulnerability to afforestation and fragmentation.
Conservation
This plant has no legal protection at present. It is recommended that activities such as forestry, roadwidening etc. should not take place without first assessing the impact on known populations, and that
habitat is not destroyed for agriculture and livestock. While undoubtedly rare and with a narrow
range and ecological tolerance, there is no evidence that this species has declined.
Designated sites
None known.
References
Casas, C. & Sérgio, C. 1990. Acaulon fontiquerianum sp. nov. de la Peninsula Ibérica. Cryptogamie, Bryologie,
Lichènologie 11: 57–61.
Lo Giudice, R. 1995. Acaulon fontiquerianum Casas & Sérgio (Pottiaceae, Bryophytina), new to the bryoflora of
Italy. Flora Mediterranea 5: 69–72.
Sérgio, C., Hébrard, J.P. & Casas, C. 1993. Acaulon fontiquerianum Casas & Sérgio (Musci, Pottiaceae) nouveau
pour la bryoflore du Portugal, de France et de Corse. Orsis 8: 11–19.
36
Checklist and country status of European bryophytes
Tayloria rudolphiana (Garov.) Bruch & Schimp.
Rudolph’s gland-moss
syn. Tayloria delavayi (Besch.) Besch.
Family: Splachnaceae
Order: Funariales
Subclass: Bryidae
Class: Bryopsida
Division: Bryophyta
Kingdom: Plantae
Status in Europe: Endangered Criteria: B1a; B2a, bi, bii, biii, biv
Photo: Michael Lüth
Description
Robust acrocarp 4 cm high or more; leaves with a longly excurrent nerve ending in a reddish point;
seta reddish, robust; capsule erect, ellipsoid, with sterile basal portion pyriform and narrower than the
spore-bearing upper portion; calyptra large, extending below capsule.
Ecology and biology
As with other members of the Splachnaceae, T. rudolphiana prefers nitrogen-rich organic subtrates.
However, its ecology is unique in that it is usually epiphytic, generally growing on the remains of bird
droppings on Acer pseudoplanatus, and more rarely Acer campestre, Fagus sylvatica and Pinus sylvestris. It
37
Checklist and country status of European bryophytes
is rarely reported from other substrates, such as dead wood, bones and rock. It is probably restricted
to areas with a high humidity (Kiebacher et al. 2012). Subalpine.
Altitudinal range: 850–1700 m (3800–4400 m in China).
Autoicous, spores presumably dispered by flies (Kiebacher et al. 2012). It has developed brood-cells in
culture (Martinez & Price 2011) but these have not yet been observed in the wild.
European distribution
Confined in Europe to a narrow band running across Germany (Endangered), Austria (‘seriously
threatened with extinction’) and Switzerland (Vulnerable).
World distribution
Outside Europe only in China (Yunnan).
Threats
Threats not known in detail, but it is probably vulnerable to inappropriate forest management
practices such as the felling of old trees.
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Checklist and country status of European bryophytes
Conservation
This plant is listed on Appendix 1 of the Bern Convention and protected under the EU Habitats
Directive. It is also ‘priority 1’ on the Swiss National Priority Species list. There is a statutory
obligation to designate SACs for the best populations of this plant in EC countries. Old forest where it
grows should be protected in sufficiently large areas to guarantee the survival of populations of birds
yielding the substratum for the moss. Research into the unusual ecology of this species is necessary to
determine its requirements in detail.
Designated sites
????
References
Hallingbäck, T. & Hodgetts, N.G. (eds.) 2000. Mosses, liverworts and hornworts. Status survey and conservation Action
Plan for bryophytes. Gland, IUCN.
Kiebacher, T., Bürgi, M., Scheidegger, C. & Bergamini, A. 2012. Bryophyte diversity of sycamore pastures in the
northern Alps with a special emphasis on Tayloria rudolphiana. Swiss Federal Research Institute WSL,
Birmensdorf; unpublished.
Martinez, K. & Price, M. 2011. Brood cells in the rare epiphytic moss Tayloria rudolphiana (Garov.) Bruch &
Schimp. (Splachnaceae). Cryptogamie, Bryologie 32: 1–10.
39
Checklist and country status of European bryophytes
Thamnobryum angustifolium (Holt) Nieuwl.
Derbyshire feather-moss
syn. Porotrichum angustifolium Dixon; Thamnium angustifolium Holt
Family: Neckeraceae
Order: Hypnales
Subclass: Bryidae
Class: Bryopsida
Division: Bryophyta
Kingdom: Plantae
Status in Europe: Endangered
Criterion: D
Photo: Des Callaghan
Description
A medium-sized, dark green, dendroid pleurocarp with shoots up to 4 cm long, often encrusted with
calcareous material below. It can be distinguished from the common Thamnobryum alopecurum by the
structure of the branch leaves, which are narrower, very strongly toothed, parallel-sided and have a
broad and poorly-defined costa (Furness & Gilbert 1980; Hodgetts & Blockeel 1992; Holt 1886). The
leaves of T. cataractarum are less strongly toothed but have an even broader costa.
T. angustifolium was discovered in Derbyshire by G.A. Holt in 1883, where it still occurs, although the
plants are now apparently smaller than the early collections. The Cumbria site was discovered in 2008.
There has been much discussion about the nature of this species. Furness & Gilbert (1980) showed that
it maintains its characters distinct from T. alopecurum in cultivation. Hodgetts & Blockeel (1992)
considered that it is more closely related to T. cataractarum and the Madeiran T. fernandesii than to T.
alopecurum. However, more recent molecular work by Olsson et al. (2009) suggests that, while T.
40
Checklist and country status of European bryophytes
angustifolium is undoubtedly a distinct entity, colonies of this plant (and of other narrowly endemic
Thamnobryum species) may originate from the surrounding populations of T. alopecurum. In this case,
the two populations of T. angustifolium are independently derived from local T. alopecurum, and have
evolved convergently in response to the rheophilous habitat.
Ecology and biology
In the Derbyshire Dales, where it has been known for many years, it grows on a shaded limestone
rock in a calcareous spring emerging from a cave in a wooded ravine, where it is inundated by water
in winter but dry in summer. It is surrounded by a dense sward of T. alopecurum. At its more recentlydiscovered site in Cumbria, it grows on the vertical sides of sandstone slabs on the bank of the River
Eden, where it is submerged in fast-flowing water for most of the year.
Altitudinal range: 50–244 m.
Dioicous, only male plants known.
European distribution
Endemic to England, where it occurs at two sites, one in Derbyshire, the other in Cumbria. Oceanic
temperate (Hill et al. 2007). A specimen supposedly from Ireland is of doubtful provenance and a
record from Madeira is erroneous.
41
Checklist and country status of European bryophytes
World distribution
See above.
Threats
T. angustifolium is at risk because of the extremely restricted extent of the population. Collecting by
botanists is a significant threat, as the plant is conspicuous, the population very small and at least one
of the sites well-known. The size of old herbarium specimens suggests that the shoots are now smaller
than they used to be, perhaps because of over-collecting. Reduction in water flow, due perhaps to
lowering of the water table or climatic changes, is also a potential threat. The overall extent of the
Derbyshire population was reduced during 1996 and 1997 by drought, when the water flow from the
cave was much diminished from the norm. However, it was flowing well again in late 1997 and
regular monitoring is now underway to determine whether the plant is recovering. Initial
observations suggest that it is doing so, with fresh green shoots appearing from areas that had
appeared dead following the drought. Irresponsible activities relating to caving may be a significant
threat to this species. For example, large areas of rock are sometimes excavated in order to make small
caves more accessible, and there is known to be an interest among cavers in finding the source of the
stream that emerges at the Thamnobryum site. There are fewer obvious threats at the Cumbria site,
although over-zealous riverbank management and pollution are both potential threats.
Conservation
A notice has been placed at the entrance to the cave alerting cavers to the proximity of a rare plant and
Natural England is monitoring the situation carefully. Water pollution may also be a potential threat.
T. angustifolium is a prime candidate for taking into cultivation as an insurance against disappearance
from its only known site. A Biodiversity Action Plan has been written for this species, and it is
included on a list of the world’s most threatened bryophytes (Hallingbäck & Hodgetts 2000). It also
receives legal protection under the Wildlife & Countryside Act 1981, and is a candidate for ex situ
conservation at Kew Gardens.
Designated sites
Derbyshire Dales NNR
Peak District National Park
Peak District Dales SAC
Cressbrook Dale SSSI
River Eden and Tributaries SSSI
42
Checklist and country status of European bryophytes
References
Furness, S.B. & Gilbert, O.L. 1980. The status of Thamnobryum angustifolium (Holt) Crundw. Journal of Bryology 11:
139–144.
Hallingbäck, T. & Hodgetts, N.G. (eds.) 2000. Mosses, liverworts and hornworts. Status survey and conservation Action
Plan for bryophytes. Gland, IUCN.
Hodgetts, N.G. & Blockeel, T.L. 1992. Thamnobryum cataractarum, a new species from Yorkshire, with observations
on T. angustifolium and T. fernandesii. Journal of Bryology 17: 251–262.
Hill, M.O., Preston, C.D., Bosanquet, S.D.S. & Roy, D.B. 2007. BRYOATT. Attributes of British and Irish mosses,
liverworts and hornworts. Abbots Ripton, NERC Centre for Ecology and Hydrology & Countryside Council for
Wales.
Holt, G.A. 1886. A British moss new to science. Journal of Botany 24: 65.
Olsson, S., Rumsey, F., Grundmann, M., Russell, S., Enroth, J. & Quandt, D. 2009. The origin of the British and
Macaronesian endemic Thamnobryum species (Neckeraceae). Journal of Bryology 31: 1–10.
43
Checklist and country status of European bryophytes
Appendix 6. Checklist and country status of European
liverworts and hornworts
This table is based upon a working list supplied by Nick Hodgetts (October 2013) and a revision supplied on
26/07/2014. Nomenclature is based on information from the Early Land Plants Today project (Lars Söderström
pers. comm. 2011), which is currently finalising a World Checklist of Liverworts and Hornworts. The subdivision
of Russia is based on the system used by Söderström et al. 2007. Occurrence and Threat data are based on the
latest available checklists and Red Lists from each country/territory, backed up with information from the
network of ECCB Country Contacts. Most countries that have a Red List use IUCN threat categories to assign a
threat status to each taxon. However, where countries use their own system of threat categories, the symbols are
explained in the Legend (see below). Only published records are included in the table. Where no checklist exists
for a country/territory, information has been compiled from miscellaneous published sources and revised by the
local ECCB contact(s).
Legend
Status values italicised in red with a darker background fill have an associated ‘Status note’ (see below)
Common categories
●
■
□
?
RE
CR
EN
VU
NT
DD
DD*
NE
y
Occurrence of species confirmed - either Least Concern or no information about status
Occurrence of infraspecific taxon confirmed
At least some reports of the species presumably refer to this infraspecific taxon, although not positively
confirmed in any source
Some doubt about occurrence
Taxon recorded in some literature but later rejected
Regionally Extinct
Critically Endangered
Endangered
Vulnerable
Near Threatened
Data Deficient
Data Deficient but recently recorded
Not Evaluated
Include in candidate list? - y = yes
Country-specific categories
Germany
0
1
2
3
G
R
V
D
Ausgestorben oder verschollen (Regionally Extinct)
Vom Aussterben bedroht (Critically Endangered)
Stark gefährdet (Endangered)
Gefährdet (Vulnerable)
Gefährdung anzunehmen (Risk assumed)
Extrem selten (Extremely rare)
Zuruckgehend (Near Threatened)
Daten ungenugend (Data deficient)
Poland
Ex
E
V
R
I
Extinct
Endangered
Vulnerable
Rare
Indeterminate
Latvia
0
1
2
3
4
Extinct
Endangered
Vulnerable
Rare
Little known or insufficiently explored species
Ukraine
R
Rare
44
45
y
y
y
●
●
●
●
●
VU
‐
EN
VU
●
NT
VU
?
EN
●
●
● ● ● ● ● ● ●
●
●
●
●
‐
‐
●
VU
y
y
y
y
y
y
y
●
●
‐
●
●
●
●
●
●
●
●
●
EN
●
EN
‐
●
●
●
EN
●
●
●
EN
NT
VU
CR
CR
●
?
● VU
● ● ●
●
●
●
VU
●
●
‐
●
●
●
●
●
●
●
●
●
●
NT
●
●
y
●
●
●
y
●
●
●
●
●
●
●
DD
● ●
●
●
●
●
●
●
DD
● ●
●
●
●
‐
●
‐
●
●
●
●
●
● CR CR ●
●
●
●
●
●
y
y
y
NT
y
■
●
●
●
■
● NT
●
■
RE
● ●
● ●
□
□ □
●
●
■
■
●
●
□
■
CR
●
●
■
■
●
●
●
●
■
□
●
■ ■
RE
●
●
■
■
●
●
●
●
‐
● VU
● ● ●
● ● ●
■ ■
VU
● ●
● ●
●
●
●
●
●
DD
DD ●
EN
‐
‐
‐
●
●
●
●
DD ●
●
●
●
●
NT
●
VU
NT
NT
●
●
●
■
■
■
□
NT
●
CR
NT
●
VU
●
●
‐
●
□
●
CR
●
●
●
●
□
DD ●
NT ●
□ □
●
●
□
‐
●
●
●
●
‐
●
●
●
□
□
□
●
●
NT
●
●
□
●
●
‐
●
□
●
●
□
●
●
■
NT
NT
■
NT
NT
NT
VU
NT
●
●
●
●
●
●
●
●
●
■
■
●
●
NT
●
NT
VU
NT
NT
●
●
●
●
DD
CR
NT ●
●
■
■
●
CR
■
■
CR
‐
●
‐
CR
●
●
■
DD ●
●
DD*
NT
NT
NT
●
●
■
●
DD ●
●
●
●
‐
‐
■
‐
RE
●
●
●
●
●
●
●
□
■
●
●
●
■
■
■
●
DD
●
●
●
●
‐
●
●
VU
●
●
■
■
VU
●
■
y
y
●
‐
●
NT ●
●
●
●
● ‐ ‐
● NT ●
□ □ □
●
●
■
NT
● ●
■ □
●
●
■
●
●
■
●
●
■
●
●
●
●
VU
DD
●
●
■
□
EN
●
●
●
●
●
●
●
●
●
■
■
●
●
●
●
●
●
VU
NT
●
●
■
□
NT
●
y
●
●
■
●
●
●
●
‐
●
●
y
●
■
■
●
●
y
y
●
●
y
y
y
y
●
■
■
‐
●
●
●
●
■
■
‐
RE
●
●
● ● ‐
CR ‐
●
■
●
●
●
●
●
●
●
‐
●
■
□
●
●
●
□
‐
●
●
●
● ●
CR
●
■
□
● ●
‐
●
●
■
●
● EN EN VU
● ● ● ●
● ● ● ●
● ● ● ●
● RE RE
‐
DD
● ● ● ●
■ ■ ■ □
■ ■ ■
● ● ●
● ● ● DD
‐ EN
‐
● ● ●
● VU VU
● ● ● ●
● ● ●
■ ■ ■
■ DD
● VU VU ●
●
●
●
●
●
●
■
■
●
●
●
‐
●
‐
‐
●
□
‐
DD
DD
NT
□
?
‐
●
□
‐
CR
CR
●
●
EN
●
□
□
●
●
●
●
■
■
‐
CR
□
□
DD
●
‐
VU
●
□
‐
DD*
●
VU
‐
VU
●
‐
?
●
□
EN
●
VU
●
EN
‐
VU
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
EN
● NT ●
● EN
● DD
‐
●
● VU
● VU
●
●
?
●
NT
● ● ●
● NT NT ●
● VU VU
‐
‐
‐ ●
‐ ‐ ●
DD
VU
NT ● NT
Monaco
●
●
y
●
●
●
Malta
Madeira
Italy
France
Cyprus
Corsica
Canary Islands
Balearic Islands
●
●
● VU VU
EN VU
● ● ●
NT
VU
Azores
Northern Ireland
Andorra
Ireland
Great Britain
Gibraltar
Channel Islands
Sweden
●
●
y
y
y
y
y
y
y
Svalbard
Taxon
Acanthocoleus aberrans
Acrobolbus wilsonii
Adelanthus lindenbergianus
Anastrepta orcadensis
Anastrophyllum alpinum Anastrophyllum assimile
Anastrophyllum donnianum
Anastrophyllum joergensenii Anastrophyllum michauxii
Aneura maxima Aneura mirabilis
Aneura pinguis Aneura pseudopinguis Anthelia julacea
Anthelia juratzkana Anthoceros agrestis Anthoceros caucasicus Anthoceros neesii Anthoceros punctatus Apomarsupella revoluta Arnellia fennica Asterella africana Asterella lindenbergiana Asterella saccata Asterella tenella Barbilophozia barbata Barbilophozia hatcheri Barbilophozia lycopodioides Barbilophozia rubescens Barbilophozia sudetica Bazzania azorica Bazzania flaccida Bazzania pearsonii Bazzania tricrenata Bazzania trilobata Bazzania trilobata var. depauperata
Bazzania trilobata var. trilobata Biantheridion undulifolium
Blasia pusilla Blepharostoma trichophyllum Blepharostoma trichophyllum subsp. brevirete Blepharostoma trichophyllum subsp. trichophyllum Bucegia romanica Calycularia laxa Calypogeia arguta Calypogeia azorica Calypogeia azurea Calypogeia fissa Calypogeia fissa subsp. fissa Calypogeia fissa subsp. neogaea Calypogeia integristipula Calypogeia muelleriana Calypogeia neesiana Calypogeia sphagnicola Calypogeia suecica Cephalozia affinis Cephalozia ambigua Cephalozia bicuspidata Cephalozia bicuspidata subsp. bicuspidata Cephalozia bicuspidata subsp. lammersiana Cephalozia catenulata Cephalozia connivens Cephalozia crassifolia Cephalozia lacinulata Cephalozia leucantha Cephalozia loitlesbergeri Cephalozia lunulifolia Cephalozia macounii Cephalozia macrostachya Cephalozia macrostachya var. macrostachya
Cephalozia macrostachya var. spiniflora Cephalozia pleniceps Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
●
‐
VU
RE
●
●
●
EN
CR
?
●
VU
●
●
●
●
●
●
EN
NT
●
2
CR
R
V
VU 0
CR R
● V
EN
‐
●
●
●
●
NT
1
2
‐
0
R
●
VU
●
NT VU
I
R
RE
V
●
NT
●
●
●
R
●
E
R
?
R
‐
●
CR
●
CR
Ukraine
Russia South
Russia Northwest
Russia North
Russia East
Novaya Zemlya
Russia Central
Moldova
Lithuania
Latvia
Kaliningrad
Kazakhstan
Franz Josef Land
Estonia
Crimea
Caucasus
Belarus
Turkey
Slovenia
Serbia
Romania
Montenegro
Macedonia
Kosovo
Hungary
Appendix 6 - Liverworts
Greece
Croatia
Crete
Bulgaria
Bosnia‐Herzegovina
Albania
Switzerland
Slovakia
●
Poland
V
Netherlands
Liechtenstein
●
Luxembourg
Germany
●
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
EN
●
●
●
●
CR EN
VU ●
●
●
VU
●
VU
‐
DD* DD
EN CR
‐
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
VU
‐
DD
●
‐
NT
●
‐
?
‐
●
●
●
●
●
●
●
●
●
●
●
●
?
‐
‐
●
●
●
●
NT
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
●
?
?
?
EN
●
●
●
●
?
●
●
VU
CR
●
●
●
●
●
D
1
●
EN
●
●
●
V
3
3
●
●
● EN
CR CR
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‐
●
E
‐
●
●
●
EN ●
EN VU
‐
●
●
●
●
●
●
●
●
●
●
●
●
VU
‐
‐
●
●
●
●
●
●
●
●
●
?
VU
●
●
●
●
●
●
●
●
V
●
●
●
●
●
●
●
●
VU
●
●
VU
●
CR
●
VU
●
●
●
‐
●
□
●
●
‐
●
□
●
●
●
●
●
●
?
●
NT
●
●
□
□
□
■
■
□
□
NT V
● V
■ ■
■ ■
RE 2
● V
● ●
□
‐
■ ■
●
●
■
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●
●
■
■
●
●
□
●
□
●
●
●
●
●
●
●
●
□
?
■
□
□
■
□
Ex RE CR
EN VU ●
● VU
●
‐
●
●
●
■
■ DD*
□
■
■
■
‐
● EN
●
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●
●
●
●
●
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●
●
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●
●
●
●
●
●
●
●
●
●
●
●
EN
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□
□
□
□
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‐
●
●
■
■
●
●
■
■
●
●
□
●
●
CR
■
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●
●
●
●
●
●
●
?
●
●
●
●
●
‐
‐
●
●
●
●
RE
●
●
●
●
●
●
1
●
●
●
●
●
●
‐
●
●
‐
●
●
●
‐
●
●
●
●
●
●
●
●
●
‐
●
‐
●
●
●
●
●
EN
EN
EN
□
●
●
□
?
●
●
■
■
●
●
■
■
●
●
●
NT
●
2
●
●
●
□
□
■
□
■
■
□
●
●
●
●
‐
‐
●
●
●
●
●
●
●
●
●
●
□
□
‐
□
□
□
□
■
□
●
●
□
□
●
●
●
●
■
■
●
●
?
‐
●
●
■
■
●
●
●
□
■
●
●
‐
●
●
●
□
□
■
●
●
●
●
■
●
●
□
●
NT
□
●
●
●
VU
●
●
VU
●
■
■
●
EN
●
●
●
●
●
● ●
● ●
● D
NT G
NT 3
?
‐
D
● ●
■ ■
□ ■
NT V
● V
‐
●
□
●
●
EN
●
VU
●
‐
●
CR
□
●
□
●
●
●
●
□
RE D
NT ●
VU 3
●
3
‐
?
VU 3
□ ■
■
VU 3
●
●
□
●
●
●
●
□
●
●
●
VU
●
NT
●
□
R
●
□
●
I
□
□
●
●
●
●
●
●
DD DD* ●
EN ●
CR
R
●
●
●
●
■
■
■
■
■
■
VU DD* V
EN VU ●
●
●
EN
■
●
NT
RE
I
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R
●
?
●
□
■
●
●
EN
□
●
●
■
●
●
●
●
●
●
DD VU
●
●
●
VU NT
●
●
■
■
■
?
●
●
NT NT
‐
‐
● EN
CR VU
●
●
‐
‐
VU
VU
●
●
●
●
●
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●
●
□
●
●
□
●
EN
□
●
□
DD*
● ●
VU
VU
● VU
●
■
●
●
VU
●
■
■
EN
VU
●
●
□
●
●
●
●
●
□
●
●
DD
NT
DD
●
EN
?
●
□
●
□
□
●
■
●
●
●
■
●
■
■
● DD
●
●
●
□
●
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●
●
● ●
EN
CR
●
●
●
EN
●
●
●
DD
●
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●
●
●
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●
●
□
●
EN
□
●
□
●
●
●
●
■
●
VU
●
NT
NT
■
●
●
‐
●
●
■
■
●
●
VU
●
●
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■
VU
VU ●
VU
● VU VU
●
●
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VU
● ●
●
●
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●
●
●
●
□
●
●
●
●
●
●
●
□
●
●
●
●
●
□
●
●
●
●
?
●
●
●
DD
‐
‐
‐
‐
‐
●
●
●
●
●
●
□
□
CR
●
●
●
●
□
●
●
●
●
●
●
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DD
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●
□
●
□
●
●
NT
●
●
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●
●
●
●
1
□
●
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‐
□
●
●
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‐
●
●
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
■
●
●
●
●
●
●
●
●
●
●
■
■
‐
●
‐
●
●
●
●
●
●
‐
●
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NT NT
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●
●
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●
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●
CR
DD
●
●
‐
●
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Sardinia
San Marino
Portugal
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Spain
●
□
EN
NT
CR
● ● ●
■ ■ ■
‐
NT DD
DD
●
●
●
y
y
Sicily
●
●
■
■
●
‐
●
●
y
y
y
●
●
●
●
□
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y
y
Malta
●
●
Madeira
●
●
‐
●
□
■
Italy
●
NT
DD
● ●
■ □
?
France
Cyprus
Balearic Islands
Azores
Northern Ireland
Andorra
Ireland
Great Britain
Gibraltar
Channel Islands
Sweden
Corsica
y
Canary Islands
y
y
Svalbard
Taxon
Cephaloziella arctogena Cephaloziella aspericaulis Cephaloziella baumgartneri Cephaloziella calyculata Cephaloziella dentata Cephaloziella divaricata Cephaloziella divaricata var. divaricata
Cephaloziella divaricata var. scabra Cephaloziella elachista Cephaloziella elegans Cephaloziella granatensis Cephaloziella grimsulana Cephaloziella hampeana Cephaloziella integerrima Cephaloziella massalongii Cephaloziella nicholsonii Cephaloziella phyllacantha Cephaloziella polystratosa Cephaloziella rubella Cephaloziella spinigera Cephaloziella stellulifera Cephaloziella turneri Cephaloziella uncinata Cephaloziella varians Cheilolejeunea cedercreutzii Chiloscyphus pallescens Chiloscyphus pallescens var. fragilis Chiloscyphus pallescens var. pallescens Chiloscyphus polyanthos Chiloscyphus polyanthos var. polyanthos
Chiloscyphus polyanthos var. rivularis Cladopodiella fluitans Cladopodiella francisci Clevea hyalina Clevea spathysii Cololejeunea azorica Cololejeunea calcarea Cololejeunea madeirensis Cololejeunea microscopica Cololejeunea minutissima Cololejeunea rossettiana Cololejeunea schaeferi Cololejeunea sintenisii Colura calyptrifolia Conocephalum conicum Conocephalum salebrosum Corsinia coriandrina Crossocalyx hellerianus
Cryptocolea imbricata Cyathodium foetidissimum Diplophyllum albicans Diplophyllum obtusatum Diplophyllum obtusifolium Diplophyllum taxifolium Douinia ovata Drepanolejeunea hamatifolia Dumortiera hirsuta Endogemma caespiticia
Eremonotus myriocarpus Exormotheca pustulosa Exormotheca welwitschii Fossombronia angulosa Fossombronia caespitiformis Fossombronia caespitiformis subsp. caespitiformis
Fossombronia caespitiformis subsp. multispira Fossombronia echinata Fossombronia fimbriata Fossombronia fleischeri Fossombronia foveolata Fossombronia incurva Fossombronia maritima Fossombronia mittenii Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
?
●
Ukraine
Russia South
Russia Northwest
Russia North
Russia East
Novaya Zemlya
Russia Central
Moldova
Lithuania
Latvia
Kaliningrad
Kazakhstan
Franz Josef Land
Estonia
Crimea
Caucasus
Belarus
Turkey
Slovenia
Serbia
Romania
Montenegro
Macedonia
Kosovo
Hungary
Appendix 6 - Liverworts
Greece
Croatia
Crete
Bulgaria
Bosnia‐Herzegovina
Albania
Switzerland
Slovakia
Poland
Netherlands
Luxembourg
Liechtenstein
Germany
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
●
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Sardinia
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NT
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Italy
●
●
Monaco
France
●
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Cyprus
●
●
Madeira
Corsica
●
Azores
Northern Ireland
Andorra
Ireland
●
EN
●
y
●
●
VU
●
y
y
●
●
●
●
Spain
?
●
●
Canary Islands
●
EN
● ●
●
●
Sicily
y
y
y
y
y
y
y
y
y
y
VU
● ●
●
●
Balearic Islands
y
y
RE ●
● ●
Gibraltar
●
Channel Islands
●
Sweden
●
Great Britain
●
●
Svalbard
Taxon
Fossombronia pusilla Fossombronia wondraczekii Frullania azorica Frullania bolanderi Frullania dilatata Frullania ericoides Frullania fragilifolia Frullania inflata Frullania jackii Frullania microphylla Frullania oakesiana Frullania parvistipula Frullania polysticta Frullania riparia Frullania sergiae Frullania tamarisci Frullania teneriffae Geocalyx graveolens Gongylanthus ericetorum Gymnocolea borealis Gymnocolea fascinifera Gymnocolea inflata Gymnocolea inflata subsp. acutiloba Gymnocolea inflata subsp. inflata
Gymnomitrion adustum Gymnomitrion alpinum Gymnomitrion brevissimum Gymnomitrion commutatum Gymnomitrion concinnatum Gymnomitrion corallioides Gymnomitrion crenulatum Gymnomitrion obtusum Haplomitrium hookeri Harpalejeunea molleri Harpanthus flotovianus Harpanthus scutatus Herbertus borealis Herbertus hutchinsiae Herbertus cf. juniperoideus
Herbertus norenus Herbertus sendtneri Herbertus stramineus Herbertus sp.
Heterogemma capitata
Heterogemma laxa Heteroscyphus denticulatus Heteroscyphus fissistipus Hygrobiella laxifolia Isopaches alboviridis Isopaches bicrenatus Isopaches decolorans Jubula hutchinsiae Jubula hutchinsiae subsp. caucasica Jubula hutchinsiae subsp. hutchinsiae
Jungermannia atrovirens Jungermannia borealis Jungermannia exsertifolia Jungermannia polaris Jungermannia pumila Kurzia pauciflora Kurzia sylvatica Kurzia trichoclados Leiomylia anomala Lejeunea canariensis Lejeunea cavifolia Lejeunea eckloniana Lejeunea flava Lejeunea hibernica Lejeunea lamacerina Lejeunea mandonii Lejeunea patens Lepidozia cupressina Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
V
0
R
EN
DD
DD
●
●
●
●
●
●
●
●
●
CR
●
DD*
●
VU
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
?
●
CR
CR
VU
VU
●
VU
NT
3
VU
3
●
●
EN
E
EN
‐
V
●
NT
CR
●
●
VU
●
VU
●
●
□
■
RE
EN
RE
●
●
●
●
●
CR
V
□
□
1
‐
NE
‐
●
NE
RE
CR
●
2
EN
●
□
■
R
VU 2
EN V
VU
‐
EN
●
●
●
●
●
DD* ●
●
?
●
RE CR
●
●
■
■
VU
VU
●
●
●
●
●
NT
●
●
VU
●
DD*
?
EN
●
■
EN
EN
EN
EN
●
EN
■
VU
VU
●
VU
●
NT
■
VU
VU
VU
?
EN
□
R
V
●
V
RE VU
● VU
VU
EN VU
● EN
E
I
‐
CR EN
●
●
■
■
●
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CR
VU
CR
●
EN
●
●
●
●
●
●
‐
‐
●
●
CR
●
R
■
I
●
●
●
●
●
●
VU
●
●
●
CR
●
●
●
‐
●
CR
Ukraine
●
?
●
Russia South
●
Russia East
●
Novaya Zemlya
●
Russia Central
4
Moldova
Kaliningrad
Kazakhstan
Franz Josef Land
Estonia
Crimea
Caucasus
Belarus
EN
EN
DD
●
Turkey
●
?
●
●
EN
Slovenia
●
VU
●
Serbia
●
Romania
●
Montenegro
●
Macedonia
●
●
Russia Northwest
● DD ●
VU DD
EN NT
●
●
Russia North
NT RE Ex VU
● VU
● DD
Lithuania
●
●
●
Latvia
●
●
●
Kosovo
●
●
Hungary
3
●
●
●
Greece
CR
EN
●
Croatia
●
●
Crete
VU
VU
●
●
EN VU
NT VU
Appendix 6 - Liverworts
Bulgaria
●
Bosnia‐Herzegovina
3
I
E
Albania
●
R
●
Switzerland
●
NT
VU
Slovakia
●
DD V
● V
Poland
●
●
Netherlands
EN
VU
Luxembourg
Liechtenstein
Belgium
Germany
Austria
Czech Republic
Checklist and country status of European bryophytes
●
●
●
●
●
VU
‐
NT
●
VU
●
●
●
●
□
□
□
□
□
□
?
?
●
2
●
●
4
●
●
●
●
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●
●
●
●
□
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□
●
●
DD
DD
●
●
●
●
●
●
●
□
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●
●
●
□
■
●
●
●
?
●
‐
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●
●
●
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‐
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‐
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●
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‐
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●
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●
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●
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●
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●
●
●
●
●
●
●
?
●
?
‐
‐
●
?
CR
●
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NT
VU
●
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1
●
‐
1
●
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●
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●
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2
1
EN
●
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VU R
VU
●
VU
●
NT V
‐
●
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VU
VU
●
VU
●
VU V
?
? NE
NE
NT 3
VU 3
●
3
EN V
●
3
●
●
●
●
●
●
●
●
●
●
●
V
‐
●
●
‐
?
R
EN VU
●
●
CR
CR
VU
VU
CR
CR
●
●
●
●
●
●
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R
●
RE
●
●
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CR
●
●
●
‐
●
●
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VU
●
VU
EN
DD*
VU
●
●
●
●
●
NT
●
●
●
●
●
● DD
●
●
‐
EN
VU
●
EN
●
●
VU
●
●
●
EN VU
● NT
●
●
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●
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●
VU
●
●
VU
CR
●
●
VU
VU
●
●
R
●
EN
●
●
DD
●
●
?
●
●
●
●
●
NT
●
●
●
●
●
●
■
●
●
●
●
●
‐
●
●
●
●
VU
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
‐
●
●
●
VU
●
?
●
●
●
●
●
●
●
●
●
●
●
2
●
‐
●
●
●
●
●
●
‐
‐
‐
●
●
●
●
●
‐
●
●
●
●
NT
●
●
●
●
●
●
●
●
‐
?
DD
50
51
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
● ●
●
●
●
?
●
●
●
●
NE
?
●
●
y
y
y
y
y
y
‐
VU
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
●
□
■
●
■
●
■
■
■
y
NT
□
●
y
●
●
y
y
EN
CR
● ● ●
VU
● ‐
CR
CR
?
y
y
y
●
■
■
■
●
□
■
□
●
□
■
■
NT ●
● NT ●
●
■
□
□
‐
●
●
●
●
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■
■
●
●
●
NT ●
● ● ●
● RE ●
●
●
●
●
●
‐
● NT ● ●
VU ‐ ●
● DD
● EN ● ● ●
‐
●
VU ● ● ●
● NT ● ● ‐
y
y
y
y
y
y
DD ●
●
NE
●
‐
●
●
?
●
●
?
●
●
●
●
●
‐
‐
●
●
●
‐
‐
●
●
●
●
●
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NE
●
‐
RE ‐
‐
●
NT
‐
●
●
●
●
●
●
●
NT
●
‐
‐
DD
NT
●
●
●
●
●
●
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●
●
‐
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NT
NT
●
●
●
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?
●
●
●
●
●
●
●
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‐
●
●
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●
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●
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NT
NT
NT
●
●
●
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●
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‐
‐
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●
‐
●
●
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□
●
●
●
■
●
●
?
●
● ● ●
DD DD
● ● ●
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●
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■
□
●
●
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●
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●
●
●
●
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●
●
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●
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●
●
● DD
● ●
● ●
● ●
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●
●
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●
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●
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■ ■ ■
■ ■ ■
● ● ●
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● ● ●
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NT
● ● ● ●
● NT NT ●
VU
VU
DD
● VU VU ●
● VU VU ●
●
● NT
● ● ● DD
● NT NT ●
●
●
□
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●
●
●
‐
●
‐
●
‐
EN
VU
■
■
■
●
●
●
●
●
●
●
●
●
●
VU
‐
●
●
●
●
●
●
●
●
VU
●
□
●
□
?
●
□
■
□
●
●
●
□
●
●
?
●
■
■
●
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EN
●
●
●
●
●
●
●
CR
VU
●
●
●
■
□
□
●
● DD
● EN
● NT ●
● NT ●
●
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● CR
●
NT
●
●
●
NT
NT
●
‐
?
●
●
●
●
●
●
□
■
□
●
●
‐
●
●
●
●
●
●
●
●
CR
?
●
●
●
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
Monaco
Malta
●
●
●
●
●
●
□
■
●
y
● ●
‐
● ● ●
●
● ● ●
● ● ●
● ● ●
■
■ ■ ■
● ● ●
? ■
■ □ ■
EN ● EN
EN ● NT
□ ■ □
■
‐ ● ●
●
●
●
●
●
●
VU
●
VU
NT
●
●
● ‐ EN ● ‐
● NT ‐ ● ‐ NT
□
■
?
Madeira
Sweden
y
Italy
CR
France
●
●
●
●
‐
●
NT ‐
‐ ‐
● VU ●
y
Cyprus
●
y
Corsica
●
y
■
●
?
●
●
●
Canary Islands
●
●
Balearic Islands
‐
CR
●
Azores
●
●
‐
●
Andorra
●
●
●
Great Britain
□
Gibraltar
□
Channel Islands
Northern Ireland
●
Ireland
●
□
‐
●
●
■
‐
●
●
□
y
y
Svalbard
Taxon
Lepidozia cupressina subsp. cupressina
Lepidozia cupressina subsp. pinnata Lepidozia pearsonii Lepidozia reptans Lepidozia stuhlmannii Leptoscyphus cuneifolius Leptoscyphus porphyrius Liochlaena lanceolata Liochlaena subulata Lophocolea bidentata Lophocolea bispinosa Lophocolea brookwoodiana Lophocolea coadunata Lophocolea fragrans Lophocolea heterophylla Lophocolea minor Lophocolea semiteres Lophozia ascendens Lophozia ciliata Lophozia groenlandica Lophozia guttulata Lophozia lantratoviae Lophozia savicziae Lophozia schusterana Lophozia silvicola Lophozia silvicoloides Lophozia ventricosa Lophozia wenzelii Lophoziopsis excisa Lophoziopsis excisa var. elegans Lophoziopsis excisa var. excisa Lophoziopsis longidens Lophoziopsis longidens subsp. arctica Lophoziopsis longidens subsp. longidens Lophoziopsis pellucida Lophoziopsis polaris Lophoziopsis polaris var. polaris Lophoziopsis polaris var. sphagnorum Lophoziopsis propagulifera Lophoziopsis rubrigemma Lunularia cruciata Mannia androgyna Mannia californica
Mannia controversa Mannia fragrans Mannia gracilis Mannia pilosa Mannia sibirica Mannia triandra Marchantia paleacea Marchantia polymorpha Marchantia polymorpha subsp. montivagans Marchantia polymorpha subsp. polymorpha Marchantia polymorpha subsp. ruderalis Marchesinia mackaii Marsupella andreaeoides Marsupella apiculata Marsupella aquatica Marsupella arctica Marsupella boeckii Marsupella condensata Marsupella emarginata Marsupella funckii Marsupella profunda Marsupella sparsifolia Marsupella sphacelata Marsupella spiniloba Marsupella sprucei Marsupella stableri Mastigophora woodsii Mesoptychia badensis Mesoptychia bantriensis Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
Ukraine
●
●
●
Russia South
●
RE
●
Russia Northwest
●
●
●
●
Russia North
●
●
●
●
●
Russia East
●
●
●
CR
●
Novaya Zemlya
●
Russia Central
●
●
●
●
●
Moldova
●
●
●
Lithuania
●
●
Latvia
●
●
Kaliningrad
●
●
Kazakhstan
●
●
Franz Josef Land
●
●
Estonia
●
●
Crimea
●
Caucasus
●
Belarus
●
●
Turkey
●
Slovenia
●
Serbia
●
Romania
●
Crete
●
Bulgaria
●
Kosovo
Bosnia‐Herzegovina
●
Albania
Switzerland
Slovakia
Poland
Netherlands
●
Montenegro
●
□
Macedonia
●
Hungary
●
Greece
●
Appendix 6 - Liverworts
Croatia
□
‐
Luxembourg
Liechtenstein
Germany
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
□
●
●
●
●
●
●
●
●
?
●
●
●
●
●
R
●
VU
‐
●
‐
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT NE
CR NE
● ●
●
?
NE
●
V
●
●
●
●
●
●
●
‐
●
●
‐
●
●
●
●
●
?
●
●
VU
●
●
●
●
●
●
NT
●
□
EN
■
●
■
●
□
□
NT LC
□
□
‐
D
●
●
●
●
●
●
●
●
●
● ●
CR NE
● V
□
●
□
●
□ ■
NT NE
□
□
□
●
□
?
●
●
●
●
●
●
●
EN D
●
‐
●
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●
●
● CR
●
●
●
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■
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VU
●
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●
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●
●
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●
●
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●
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●
●
●
VU VU DD*
●
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●
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■ DD
□ DD
R
●
CR VU
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R
R
V
●
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●
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●
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●
●
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NT 3
●
●
●
CR
●
R
3
R
●
VU VU
NT ●
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●
EN
2
●
NT
●
●
●
●
●
VU V
● G
R
●
●
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●
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NT CR
EN CR
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●
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●
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●
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CR ‐
●
●
■
EN
●
■
■
‐
●
□
●
●
●
■
■
■
●
EN
●
●
■
■
■
●
●
●
●
■
■
■
■
●
●
●
■
■
■
DD
●
●
VU
●
●
●
DD
●
●
●
● DD*
●
VU ●
●
‐ VU ‐
● ●
●
■ ■
■
■ ■ VU
■ ■
■
CR
CR
●
● DD
● EN
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
1
●
VU
VU
‐
●
●
□
□
●
●
■
■
■
●
□
●
■
■
■
●
□
●
●
●
■
■
■
□
■
□
●
●
●
□
□
●
●
●
●
■
■
■
●
■
■
■
●
●
●
●
‐
●
●
●
●
●
●
□
●
■
●
□
□
●
●
■
●
●
●
●
●
‐
●
●
●
●
●
●
■
■
■
●
●
●
●
●
■
■
■
●
●
●
●
NT
●
3
●
●
●
●
●
●
●
‐
□
●
■
■
■
●
●
●
●
●
●
●
●
‐
●
●
VU
VU
●
●
●
●
●
●
●
●
CR
CR
● ●
● VU
●
●
●
DD*
VU
● DD
CR
● EN ●
CR
VU
CR
‐
VU
●
●
●
■
■
□
●
●
●
●
●
●
●
‐
■
●
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
52
53
Sicily
Spain
●
●
●
●
NT
VU
●
●
●
CR
NT
●
VU
□
□
■
■
□
□
●
VU
VU
●
●
●
●
NT
NT
●
●
●
●
VU
●
●
●
NT
●
●
●
●
●
●
●
NT
CR
VU
VU
CR
●
●
□
●
●
●
●
●
●
●
□
EN
EN
VU
VU
NT
■
●
‐
●
‐
●
●
NT
NT
NT
Monaco
□
Malta
■
y
●
● NT ●
●
●
●
●
●
●
●
?
●
●
□
■
●
□
●
●
■
■
■
●
●
●
■
□
■
●
■
■
● ● ● ●
NT NT NT
NT VU
●
CR ●
y
y
■
●
●
□
●
‐
■
●
□
●
■
■
■
●
●
●
●
y
●
●
●
●
●
●
■
●
□
□
EN EN
■
■ ■
NT
●
●
●
●
●
●
●
EN
NT
●
●
●
‐
●
□
●
?
●
●
●
●
●
□
●
●
●
●
●
●
●
■
●
DD ● ●
DD DD DDDD*
● ● ●
‐
NT
DD
● ● ● ●
● NT ‐ NT ●
□ □
□ □
‐
●
●
VU
●
‐
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
y
y
●
□
● VU
●
● VU
● NT
NT
CR
● ●
□ □
y
●
●
●
●
●
●
●
●
●
●
●
●
NT
● ●
‐ NT
● ●
NT
y
●
●
●
‐
●
□
●
‐
●
‐
●
●
●
●
‐
‐
●
●
●
●
NT
●
●
●
●
●
●
●
●
y
●
●
‐
●
●
●
●
y
●
● ● ●
● ● ●
● ● ●
● ● ●
● NT NT
●
VU
‐
● ● ●
● ● ●
□
●
‐
VU
□
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
‐
‐
●
● ● ● ● ●
●
VU
● ● ●
●
● NT
●
VU
●
● ● ● ● ‐
● DD*
DD
● EN EN
●
●
●
●
●
●
y
y
●
●
y
●
●
●
y
y
y
● ● ‐ ●
CR
NT ● NT
‐ ● ● ●
●
EN
VU ‐ DD
●
●
●
□
■
●
y
y
●
NT ●
● ●
□
□ ■
● ● ●
● ●
●
‐
‐
●
●
□
□
●
●
RE ● DD
‐ ‐ ‐
RE
‐
●
●
■
■
●
● ●
●
●
■
●
●
NT ●
● EN
● ● ●
● ● ●
● ● ●
□ ‐
■ ■ ■
● ● ●
● ●
NT
● ●
‐
●
●
‐
●
●
●
‐
●
●
‐
□
●
□
‐
●
‐
●
●
●
●
y
●
y
‐
●
y
y
‐
●
?
●
●
NT
y
‐
●
●
●
● ● ●
● ● ●
● ● ●
● EN
● ● ●
● EN
‐
●
‐
●
●
●
●
●
●
●
●
●
●
□
□
●
●
●
●
‐
●
●
●
●
●
■
■
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
□
‐
CR
●
NT
□
●
●
●
NT
□
●
□
DD
●
DD
VU ●
CR
EN ●
CR
DD
●
● VU ‐
● CR
●
●
●
●
●
●
‐
●
●
●
VU
CR
●
●
VU
VU
●
●
‐
VU
●
CR
CR
NT
NT
NT
●
●
●
●
●
●
●
●
●
●
■
NT
CR
NT
□
NT
□
●
●
VU
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
●
●
●
●
●
●
VU
●
●
□
●
‐
●
●
●
●
●
NT
NT
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
NT
EN
‐
●
● DD ●
●
●
●
●
DD
●
●
VU
●
‐
?
●
VU
Vatican City
Sardinia
San Marino
Portugal
●
Madeira
□
Italy
□
France
□
□
Cyprus
□
Corsica
■
Canary Islands
■
Balearic Islands
Northern Ireland
Andorra
■
Azores
Ireland
■
Great Britain
■
Gibraltar
□
Channel Islands
□
Sweden
□
Svalbard
Taxon
Mesoptychia bantriensis subsp. bantriensis
Mesoptychia bantriensis subsp. wallfischii Mesoptychia collaris Mesoptychia fitzgeraldiae Mesoptychia gillmanii Mesoptychia heterocolpos Mesoptychia heterocolpos var. arctica Mesoptychia heterocolpos var. harpanthoides Mesoptychia heterocolpos var. heterocolpos
Mesoptychia rutheana Mesoptychia rutheana var. laxa Mesoptychia rutheana var. rutheana
Mesoptychia sahlbergii Mesoptychia turbinata Metzgeria conjugata Metzgeria consanguinea Metzgeria furcata Metzgeria leptoneura Metzgeria pubescens Metzgeria simplex Metzgeria violacea Microlejeunea ulicina Mnioloma fuscum Moerckia blyttii Moerckia flotoviana Moerckia hibernica Mylia taylorii Nardia breidleri Nardia compressa Nardia geoscyphus Nardia geoscyphus var. geoscyphus
Nardia geoscyphus var. suberecta Nardia insecta Nardia japonica Nardia scalaris Neoorthocaulis attenuatus Neoorthocaulis binsteadii Neoorthocaulis floerkei Neoorthocaulis hyperboreus Notothylas orbicularis Nowellia curvifolia Obtusifolium obtusum Odontoschisma denudatum Odontoschisma elongatum Odontoschisma macounii Odontoschisma prostratum Odontoschisma sphagni Oleolophozia perssonii Orthocaulis atlanticus Orthocaulis cavifolius Oxymitra incrassata Pallavicinia lyellii Pedinophyllum interruptum Pellia endiviifolia Pellia epiphylla Pellia epiphylla subsp. borealis Pellia epiphylla subsp. epiphylla
Pellia neesiana Peltolepis quadrata Petalophyllum ralfsii Phaeoceros carolinianus Phaeoceros laevis Phymatoceros bulbiculosus Plagiochasma appendiculatum Plagiochasma rupestre Plagiochila arctica Plagiochila asplenioides Plagiochila bifaria Plagiochila britannica Plagiochila carringtonii Plagiochila exigua Plagiochila heterophylla Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
■
●
●
●
‐
●
●
●
●
●
●
VU
EN
●
DD
●
●
VU
●
●
□
NE
NE
3
V
● NE
DD NE
● VU 3
● CR 3
●
●
●
●
●
□
EN 2
CR ●
?
2
● V
NE
VU 2
● V
□ □
●
CR
● RE
VU R
●
●
●
CR
●
●
VU
●
●
●
●
●
RE
‐
RE
EN EN
□
□
● DD
3
●
●
●
●
●
●
V
V
●
● EN
NT EN
●
●
●
3
●
RE
●
●
●
EN
VU CR
EN
VU
●
●
●
●
●
CR 2
● V
EN 3
●
3
D
R
EN
RE
3
D
VU
R
RE
CR
RE
●
●
●
□
●
VU
EN
‐
●
●
●
●
■
●
●
●
EN ?
RE V
● ●
● ●
● ●
□
□ ■
● V
D
●
‐
VU
□
□
■
□
●
●
●
●
●
■
●
VU
□
□
NT
■
●
CR
●
VU
●
VU
VU
■
●
●
VU
●
●
●
●
●
●
●
●
□
●
□
VU
□
●
3
‐
VU
●
‐
EN
●
●
●
●
●
●
V
R
●
●
CR
●
●
●
●
●
‐
●
■
■
?
EN
?
VU
●
EN
EN
●
□
■
EN
●
DD*
EN
VU
VU
NT
●
□
●
●
R
●
R
‐
V
●
●
R
●
●
●
●
●
●
V
●
●
●
‐
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
VU
DD
DD
●
●
●
●
EN
□
●
●
●
● DD
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
?
●
●
●
●
□
□
●
●
DD*
DD*
●
●
●
EN
DD
□
●
□
●
VU
●
●
●
‐
● VU
● DD
VU
●
●
●
□
■
●
● VU
● ●
CR
●
●
●
●
●
●
●
●
2
DD
R
E
E
●
●
●
■
■
●
R
CR VU DD
RE ‐
●
●
● ●
●
●
● ●
●
●
●
?
□
■
□
●
●
EN EN
●
EN
●
●
●
●
□
●
●
●
□
●
●
●
□
●
●
●
●
●
●
●
●
□
●
□
●
□
●
●
●
●
DD* ‐
● ●
●
?
●
●
●
VU
●
●
●
●
●
●
?
●
□
●
VU
CR
●
●
●
●
●
●
□
●
□
●
●
●
●
EN
●
●
●
●
●
VU
●
‐
●
●
●
VU
□
●
□
●
VU
●
●
●
V
NE
●
● EN
RE
●
●
●
●
‐
●
□
□
●
●
?
‐
‐
●
●
●
●
‐
●
‐
1
●
●
●
□
●
□
‐
‐
●
□
●
1
●
●
●
●
●
?
●
‐
?
●
3
●
●
‐
●
●
●
‐
●
●
●
●
□
●
■
●
□
‐
●
■
●
●
●
‐
‐
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
□
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
□
●
□
●
●
●
●
●
●
□
□
●
●
●
●
●
●
□
●
■
●
●
●
●
●
●
●
●
●
●
?
●
●
□
●
□
●
□
●
□
●
●
‐
●
●
●
●
●
●
●
●
□
●
‐
●
□
●
●
●
●
●
●
□
●
‐
●
NT
‐
CR
DD
DD
●
●
?
VU
●
●
●
?
●
?
●
‐
●
●
●
?
●
?
‐
‐
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Ukraine
□
●
●
●
■
□
□
●
●
VU
● NT
●
● NT
● ● ●
● DD
Russia South
●
●
●
●
Russia Northwest
●
DD*
●
Russia North
●
●
‐
●
Russia East
Novaya Zemlya
Moldova
Russia Central
●
●
●
●
●
●
VU
●
●
●
●
●
■
□
□
?
● DD* ●
‐
●
●
CR
●
●
?
□
●
□
3
●
●
DD
●
□
‐
CR
DD
●
●
●
‐
VU
●
●
●
●
VU
EN
DD
● ●
DD
●
●
■
?
?
‐
‐
●
●
NT ●
? NT
EN NT
‐ VU
VU EN
‐
●
?
● DD
□ □
VU
●
VU CR
● ●
‐
● ●
●
VU
VU
CR VU
●
?
□
■
V
Lithuania
■
● DD
□
Latvia
Switzerland
■
E
●
●
Kaliningrad
Slovakia
□
Albania
Poland
‐
●
Kazakhstan
□
□
1
VU
●
●
Franz Josef Land
□
NT
■
●
□
Estonia
□
●
■
●
●
Crimea
□
CR
VU
●
□
Caucasus
●
●
Belarus
R
G
●
Turkey
CR
●
Slovenia
●
●
□
CR
VU ●
Serbia
VU
●
●
●
Romania
●
Montenegro
●
Macedonia
V
Kosovo
●
□
Hungary
●
□
Appendix 6 - Liverworts
Greece
●
□
Croatia
□
Crete
Liechtenstein
□
Bulgaria
Germany
□
Bosnia‐Herzegovina
Czech Republic
□
Netherlands
Belgium
□
Luxembourg
Austria
Checklist and country status of European bryophytes
NT
●
●
●
●
‐
●
●
●
‐
●
DD*
VU
54
55
●
●
●
■
■
■
□
●
● ‐
EN
■ □ □
■ ■
●
●
VU ‐
●
□
□
●
●
■
■
‐
●
●
●
●
● VU ●
‐
y
y
●
□
□
●
●
?
●
?
■
‐
●
■
■
□
●
●
●
■
□
EN ●
?
● ●
VU
■ ■
●
●
●
●
NT
■
□
● ●
● ●
●
●
●
‐
●
■
□
□
□
■
■
□
□
□
□
□
●
●
●
●
●
DD
VU
‐
DD
●
●
●
● DD* ‐
●
●
●
DD
VU
●
●
●
■
●
●
●
●
●
‐
● DD ●
●
●
●
‐
●
●
VU
DD
‐
● NT ‐
● DD
●
●
● NT ●
●
● NT ●
●
‐
● NT ‐
●
● NT ‐
■
□
□
■
●
●
●
‐
●
●
●
●
● NT NT ●
‐
●
●
‐
●
●
NT
y
?
EN
y
●
‐
●
■
‐
● NT
NT
● ●
■ DD
■ □
‐ DD
‐
‐
●
●
●
●
■
●
●
■
□
□
‐ EN ‐
‐
●
●
●
‐
●
●
●
●
‐
●
‐
● VU ●
●
y
y
●
●
●
●
‐
●
●
‐ ●
‐
● ●
●
●
□
■
DD
‐
●
‐
●
●
●
●
●
●
y
y
y
y
y
y
●
●
●
●
●
●
●
●
●
●
●
■ ■
‐
VU
● ●
● ●
● RE
VU NT
● ●
● ●
● NT
●
●
●
●
●
●
● ●
RE
‐
● ‐
● ●
●
●
●
‐
●
●
●
●
●
VU
EN
‐
EN
CR
●
●
●
●
?
●
●
●
●
●
‐
●
●
●
●
●
EN
●
●
●
●
●
□
□
□
□
●
VU
VU
VU
●
NT
●
NT
●
●
●
●
●
●
‐
●
●
●
●
‐
●
EN
VU
‐
●
CR
●
VU
●
●
●
●
●
●
● DD ●
□
●
●
□
■
□
□
●
●
‐
●
VU
●
□
●
●
DD*
●
?
NT ●
■
●
●
●
□
■
●
?
●
●
■
□
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
●
●
NT
●
●
□
?
■
■
□
■
■
■
NT
●
VU
●
●
●
●
VU
●
?
●
□
●
●
□
●
●
■
NT
NT
● ●
●
●
● ● ●
● DD
●
●
●
●
●
CR
CR
CR
NT
VU
●
●
●
●
●
●
●
●
EN
EN
NT
NT
CR
EN
■
■
□
■
□
□
□
□
□
□
●
NT
●
DD
■
NT
NT
●
●
●
●
●
●
■
□
●
●
●
●
●
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
●
●
□
●
●
□
●
●
□
□
●
●
●
●
●
●
●
■
●
●
□
●
●
y
y
●
● EN ●
●
●
y
EN ‐ EN
●
VU
EN
RE ● VU
RE
EN
●
●
VU
EN
EN
●
EN
●
y
●
●
VU
● ●
‐
‐
●
‐
●
●
‐
?
‐
‐
●
● EN
NE
‐
●
●
‐
y
y
y
y
●
● DD ●
NT
● DD ●
■
●
●
VU
□
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
CR
■
■
●
●
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
●
‐
●
‐
●
RE
●
●
●
●
?
●
●
●
●
‐
DD
●
●
●
●
●
‐
●
DD
●
●
‐
●
‐
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
‐
●
●
●
●
●
CR
●
●
●
Vatican City
Spain
●
VU
●
□
y
San Marino
●
Monaco
●
Malta
●
Madeira
NT
Italy
●
France
●
Cyprus
EN
Corsica
●
●
y
y
y
y
y
y
y
y
y
Portugal
Canary Islands
Balearic Islands
Sicily
y
Sardinia
y
y
y
Azores
Northern Ireland
Andorra
Ireland
Great Britain
Gibraltar
Channel Islands
Sweden
Svalbard
Taxon
Plagiochila longispina Plagiochila maderensis Plagiochila papillifolia Plagiochila porelloides Plagiochila porelloides var. norvegica Plagiochila porelloides var. porelloides Plagiochila porelloides var. subarctica Plagiochila punctata Plagiochila retrorsa Plagiochila spinulosa Plagiochila stricta Plagiochila virginica Pleurocladula albescens Pleurocladula albescens var. albescens Pleurocladula albescens var. islandica Pleurozia purpurea Porella arboris‐vitae Porella baueri Porella canariensis Porella cordaeana Porella inaequalis Porella obtusata Porella pinnata Porella platyphylla Prasanthus suecicus Preissia quadrata Preissia quadrata subsp. hyperborea Preissia quadrata subsp. quadrata
Protolophozia elongata Protolophozia herzogiana Pseudomarsupidium decipiens Pseudotritomaria heterophylla Ptilidium ciliare Ptilidium pulcherrimum Radula aquilegia Radula carringtonii Radula complanata Radula holtii Radula jonesii Radula lindenbergiana Radula nudicaulis Radula visianica Radula voluta Radula wichurae Reboulia hemisphaerica Reboulia hemisphaerica subsp. australis Reboulia hemisphaerica subsp. dioica Reboulia hemisphaerica subsp. hemisphaerica
Reboulia hemisphaerica subsp. paradoxa Riccardia chamedryfolia Riccardia incurvata Riccardia latifrons Riccardia latifrons subsp. arctica Riccardia latifrons subsp. latifrons Riccardia multifida Riccardia palmata Riccia atlantica Riccia atromarginata Riccia beyrichiana Riccia bicarinata Riccia bifurca Riccia breidleri Riccia bullosa Riccia canaliculata Riccia cavernosa Riccia ciliata Riccia ciliifera Riccia crinita Riccia crozalsii Riccia crustata Riccia crystallina Riccia duplex Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
Luxembourg
Netherlands
Poland
Slovakia
Switzerland
Albania
Bosnia‐Herzegovina
Bulgaria
Croatia
Greece
Hungary
Kosovo
Macedonia
Montenegro
Romania
Serbia
Slovenia
Turkey
Belarus
Caucasus
Crimea
Estonia
Franz Josef Land
Kaliningrad
Latvia
Lithuania
Russia Central
Russia East
Russia North
Russia Northwest
Russia South
Ukraine
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
□
■
□
□
□
□
■
■
■
■
■
□
□
■
■
■
□
■
■
■
■
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
●
●
●
●
□
R
□
■
□
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
NE
●
CR
‐
?
CR
R
□
‐
●
□
●
■
VU
‐
●
●
●
●
NT V
●
?
●
●
●
NT V
●
●
■
■
●
NT
DD
E
E
DD VU
DD
●
●
DD* ●
NT EN
●
●
‐
NT RE
●
?
●
EN
■
■
●
●
‐
●
●
●
●
EN
VU
●
●
●
●
VU
●
●
●
●
●
‐
●
□
?
?
●
●
● ●
●
● DD DD
●
●
‐
●
●
●
●
●
●
NT
●
‐
VU
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
□
□
□
□
□
■
●
□
●
●
●
●
●
●
‐
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
1
‐
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
□
‐
□
■
■
■
□
■
□
■
■
□
□
■
■
■
●
□
□
□
□
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
R
R
●
VU
●
VU
0
●
●
●
●
●
□
□
□
□
□
□
■
□
□
□
□
□
□
EN
●
●
R
E
V
●
RE VU
CR VU
●
●
●
1
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
□
●
●
□
DD
●
□
●
3
□
●
●
□
●
●
□
●
●
□
●
●
□
●
●
□
●
●
●
●
●
●
●
3
●
■
VU
□
■
□
?
NE
□
●
●
●
●
●
●
V
●
●
●
●
●
●
3
●
●
●
VU G
●
●
●
NT
3
NT
□
□
□
□
VU
VU
●
●
●
●
□
●
●
□
●
●
●
●
VU VU
□
□
●
●
□
●
0
G
E
●
□
●
DD
●
‐
●
‐
‐
●
●
■
?
DD
CR
● ●
●
●
●
VU
●
Novaya Zemlya
Liechtenstein
●
Moldova
Germany
●
Kazakhstan
Czech Republic
●
Crete
Belgium
Appendix 6 - Liverworts
Austria
Checklist and country status of European bryophytes
●
●
●
●
‐
?
VU V
VU 3
● V
□
●
●
□
3
3
?
2
CR VU
● EN
●
●
●
NT
□
●
●
□
■
NT CR
‐
□
●
●
●
●
●
●
●
‐
NT
●
●
●
●
VU
●
●
● DD
●
●
●
●
●
NT
□
□
□
□
□
□
●
●
●
●
●
VU
●
●
● DD
DD
● EN
□
●
●
□
VU
●
□
●
●
□
‐
●
●
●
●
●
●
●
● DD
●
●
●
●
●
●
●
● DD
NT
● VU
● NT
● VU
● CR
●
●
●
●
●
EN
VU
●
●
RE
EN
EN
EN
CR
VU
● DD G
● NT 3
● NT G
● NT V
NT ?
NE
‐
●
G
?
G
●
R
DD*
EN
●
●
VU CR
VU
NT
●
●
‐
●
●
●
E
R
RE
NT
EN
●
DD
CR
EN RE
‐
‐
?
●
‐
●
●
RE
CR
VU
VU
VU
VU
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
CR
EN
●
□
EN
NT
● DD
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
□
□
■
□
□
□
?
●
●
EN
●
●
●
□
●
●
■
●
●
□
●
●
●
●
●
●
●
?
●
●
●
●
VU
●
□
●
●
□
●
●
□
●
●
□
●
●
●
●
●
●
●
EN
●
●
DD
● DD*
● VU
● ● ●
● CR ●
●
DD ●
‐
EN
DD
●
●
□
●
●
●
‐
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
1
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
‐
●
●
●
●
●
●
‐
56
57
■
‐
●
□
■
y
●
■
y
●
EN
VU
EN
● DD
●
?
■
●
□
●
●
y
●
●
● ●
DD
● ●
●
●
●
■
■ ■ □
●
● ● ●
■ DD
■ NT □
● VU
● CR ●
● VU ●
● EN ●
●
□
□
●
●
■
■
EN
●
●
‐
‐
●
●
●
●
●
●
?
●
● NT
● NT ●
● NT
● DD
●
□
‐
?
●
□
●
□
NE NE
□ □
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
□
‐
■
●
□
□
●
□
●
□
●
□
□
‐
□
□
●
□
VU ‐
‐
‐
‐
□
●
●
‐
■
●
●
●
NT
■
●
■
NT
y
y
●
‐
●
●
■
●
●
●
□
‐
● NT
●
●
□
DD
NT ●
■
NT
□
●
●
●
●
□
□
●
■
■
VU
●
‐
●
●
□
■
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
●
■
NT
●
●
□
□
□
■
?
□
EN
DD
●
● DD
●
y
●
●
y
●
y
●
●
■
y
●
y
y
●
□
y
y
y
y
y
●
●
●
●
●
●
●
●
‐ ●
● ● ● ● ●
EN
●
●
CR
VU
EN
●
●
● EN
VU
VU ● ● ● ●
CR
EN
EN
■
? RE
EN
● ● ●
□ ■ ■
■
●
●
●
■
■
● ●
VU
● ●
■ ■
VU ● ● ● ●
‐ ● ●
●
‐
EN
DD
●
●
VU ●
NT ● ● ● ●
‐ ‐
● ● ● ● ●
● ● ● ● ● ●
□ □ ■ □ □ ■
NT ‐ □ ■ ■
VU
EN ● DD
‐
●
● ● ● ● ‐ ●
□ ■ ■ ■ ‐ □
●
‐
●
?
●
●
●
●
●
●
●
●
●
‐
●
●
●
‐
●
●
●
●
DD
● ●
●
●
●
‐
DD
□
● ● ●
‐
NT VU
□ □
● VU
●
●
●
●
●
●
□
VU
□
●
□
DD
● ● ●
●
NT VU ● VU
●
□
●
□
●
□
●
●
□
‐
●
● VU ●
● EN
● NT
● DD
● NT
‐
‐
● NT
CR
■
■
● NT ●
EN
● NT ●
■ ■ □
?
NT
NT
●
●
●
●
VU
CR
CR
●
●
□
●
□
■
NT
■
‐
‐
●
□
EN
■
●
NT
●
VU
VU
●
●
?
□
●
‐
●
●
●
●
●
‐
□
●
■
■
VU
●
‐
●
●
●
●
●
●
NT
□
●
●
●
□
EN
‐
●
EN
EN
● VU
DD
NT
●
●
● DD DD
□ □ □
‐
●
●
●
●
●
□
DD
‐
●
DD
□
●
●
‐
●
●
●
‐
●
●
●
●
□
DD
□
●
?
●
DD
‐
●
‐
Vatican City
Sardinia
San Marino
Portugal
●
●
●
●
VU
DD
●
■
■
NT
y
y
y
y
y
y
y
y
y
y
y
Monaco
□
NT
CR
VU
Malta
□
● DD ●
■
■ ■ ■
Madeira
□
●
Italy
□
●
France
●
Cyprus
●
●
Corsica
●
Canary Islands
●
Balearic Islands
●
Azores
●
Andorra
Gibraltar
Channel Islands
Sweden
VU ●
□
□ □
●
●
Spain
●
■
□
●
Sicily
●
VU
Northern Ireland
●
Ireland
●
Great Britain
y
y
Svalbard
Taxon
Riccia fluitans Riccia frostii Riccia glauca Riccia glauca var. ciliaris Riccia glauca var. glauca Riccia gothica Riccia gougetiana Riccia gougetiana var. armatissima Riccia gougetiana var. gougetiana
Riccia huebeneriana Riccia lamellosa Riccia ligula Riccia macrocarpa Riccia melitensis Riccia michelii Riccia nigrella Riccia papillosa Riccia perennis Riccia rhenana Riccia rhenana var. rhenana Riccia rhenana var. violacea Riccia sommieri Riccia sorocarpa Riccia sorocarpa subsp. arctica Riccia sorocarpa subsp. erythrophora Riccia sorocarpa subsp. sorocarpa
Riccia subbifurca Riccia trabutiana Riccia warnstorfii Riccia warnstorfii var. ciliaris Riccia warnstorfii var. warnstorfii Ricciocarpos natans Riella affinis Riella bialata Riella cossoniana Riella helicophylla Riella notarisii Riella parisii Saccobasis polita Saccobasis polymorpha Saccogyna viticulosa Sauteria alpina Scapania aequiloba Scapania apiculata Scapania aspera Scapania brevicaulis Scapania calcicola Scapania carinthiaca Scapania carinthiaca var. carinthiaca
Scapania carinthiaca var. massalongii Scapania compacta Scapania crassiretis Scapania curta Scapania curta var. curta
Scapania curta var. grandiretis Scapania curta var. isoloba Scapania cuspiduligera Scapania degenii Scapania glaucocephala Scapania gracilis Scapania gymnostomophila Scapania helvetica Scapania hyperborea Scapania irrigua Scapania irrigua subsp. irrigua Scapania irrigua subsp. rufescens Scapania kaurinii Scapania ligulifolia
Scapania lingulata Scapania lingulata var. lingulata
Scapania lingulata var. microphylla Scapania mucronata Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
●
CR
?
●
□
DD
‐
□
□
□
●
●
■
■
●
●
□
■
EN G
EN
●
I
□
□
□
■
EN
□
□
□
●
●
CR
●
●
□
□
■
□
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
●
●
□
●
●
?
‐
●
‐
●
●
3
●
●
‐
●
Ukraine
□
□
●
Russia South
●
●
●
●
●
Russia East
●
Novaya Zemlya
●
●
Moldova
●
●
Kazakhstan
●
●
Russia Central
Franz Josef Land
●
Estonia
Crimea
Caucasus
Belarus
Turkey
Slovenia
Serbia
Romania
Kosovo
□
DD
Hungary
NT
●
●
●
●
Russia Northwest
□
VU
Russia North
□
●
●
●
Lithuania
●
●
CR
●
■
■
Latvia
●
EN VU
●
■
■
●
●
●
●
●
●
‐
EN
Kaliningrad
□
●
Greece
□
●
Croatia
□
●
‐
■
●
DD
● NT
●
□ □
Crete
□
●
Bulgaria
□
●
Bosnia‐Herzegovina
●
●
Montenegro
●
●
NT VU
Appendix 6 - Liverworts
Macedonia
RE
□
V
?
□ ■
NE
DD ?
●
Albania
●
●
‐
●
Switzerland
●
VU
Slovakia
●
Poland
Germany
●
Netherlands
Czech Republic
●
Luxembourg
Belgium
VU
RE
●
■
■
Liechtenstein
Austria
Checklist and country status of European bryophytes
●
●
●
●
■
●
●
●
●
●
●
■
●
●
VU
?
RE
‐
CR
●
□
VU
VU
CR
NT D
□ □
CR
□
●
□
●
□
●
●
●
●
●
●
●
EN
EN DD
□
□
EN
EN
□
●
●
EN
‐
DD
□
●
●
●
●
●
●
●
■
●
■
●
●
●
●
●
●
□
□
D
□
VU
□
●
□
EN
□
●
EN
●
VU
3
EN RE
●
□
EN
□
●
□
NT
■
3
□
CR
□
●
□
●
●
●
□
■
DD VU
‐
●
●
●
□
●
●
□
□
●
●
□
●
●
VU
●
■
●
●
■
■
VU VU
●
DD
●
●
●
□ □
‐ DD
●
DD
●
□
NT
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
●
□
●
●
●
■
□
□
□
□
□
□
■
□
□
■
□
■
□
□
■
■
□
DD
□
●
□
●
●
1
●
●
●
●
●
●
●
●
VU
●
CR
●
‐
□
●
‐
RE
●
R
‐
●
EN
●
●
E
R
‐
●
●
□
NT VU
●
●
● VU
●
●
‐
NT VU
CR VU
●
●
●
●
VU
DD
●
●
?
●
●
VU
●
●
●
VU
●
RE
VU
‐
●
●
□
●
●
VU
●
●
●
□
?
●
NT V
CR ●
VU V
●
●
●
●
EN G
RE R
□
■ ■
DD 2
●
●
□
●
□
D
■
CR
●
NT CR
●
□
VU CR
□
□
●
●
●
□
■
■
EN VU
DD VU
● EN
□
■
●
●
●
●
CR
●
●
●
●
●
●
●
EN
DD
VU
●
●
●
●
●
VU
●
●
●
●
■
EN
‐
●
□
●
●
●
●
□
●
■
●
■
●
●
CR VU
●
● ● EN ●
●
?
● ● ●
●
DD
● ● VU DD
●
EN
□
CR
CR
●
■
●
■
VU
DD
●
■
●
●
●
●
●
●
●
●
●
●
□
●
‐
NT
●
●
NT
●
●
?
●
●
‐
●
●
‐
‐
●
■
●
□
●
■
●
■
●
●
●
‐
●
●
●
●
●
□
●
●
●
●
□
●
■
●
□
‐
●
●
■
■
‐
●
●
■
‐
●
■
■
VU
●
●
■
●
□
VU V
NE
R
NE
EN 3
CR D
●
□
V
■
■
CR
●
●
‐
RE
R
●
●
□
●
□
EN
■
●
□
NT
EN
●
‐
‐
‐
‐
VU CR
VU VU
‐
●
●
□
■
?
●
●
VU
●
□
●
□
●
●
□
NT
□
●
□
●
□
CR
‐
‐
DD
CR
VU
●
■
●
●
●
□
●
□
●
□
●
□
●
●
‐
●
□
●
VU
●
■
VU
●
□
EN V
□ □
● DD V
NT RE
□
□
●
VU
R
■
●
EN
●
NT
●
■
NT
CR EN
□
■
DD* ●
●
?
●
●
●
●
●
●
□
●
●
□
‐
●
●
R
●
□
●
□
●
□
?
VU
●
●
□
●
●
□
●
□
?
NT
■
NT
●
‐
●
□
1
●
‐
□
●
●
●
●
□
●
□
●
□
●
●
●
□
●
□
●
58
59
EN ●
EN ●
y
y
●
●
■
●
□
●
□
● NT ●
●
□
●
●
●
●
□
●
●
‐
■
‐
●
■
●
●
DD
VU
● ●
●
●
□
y
●
y
y
y
y
●
■
●
■
●
EN
● ● ●
● ●
●
●
DD ●
● ●
●
■
■
●
■
● ●
● DD
● ●
● ●
●
? ●
● NT
●
●
● ●
● ‐
●
‐
●
●
●
●
●
●
‐
●
●
‐
●
●
●
●
●
‐
●
●
●
●
●
■
Spain
□
□
□
■
EN
□
□
●
●
NT
●
NT
●
●
●
■
■
●
NT
NT
NT
●
●
‐
●
■
VU
■
□
●
DD
●
■
NT
■
VU
□
■
□
●
■
●
■
■
●
□
●
DD ●
□
■
●
□
DD ●
□
□
DD
VU
● ●
□
●
●
CR
VU
NT RE
NT
NT
●
●
CR
CR
VU
‐
●
●
DD
EN
● ●
● ● ● ●
● ● ● ●
● NT NT
●
● NT NT ●
● NT NT
VU
●
VU CR
●
●
●
DD ●
●
●
CR
NT
●
●
●
?
●
●
●
●
●
NT
NT
CR
●
●
●
●
●
VU
●
●
?
●
●
●
●
●
‐
●
●
●
●
NT
NT
NT
NT
NT
●
?
●
●
●
●
CR
●
●
●
●
●
●
●
?
●
●
●
●
VU
●
VU
●
●
●
●
●
●
VU
●
●
●
● NT ●
‐ DD
● CR ‐
●
●
y
●
●
y
●
?
?
●
●
● ●
● ●
VU ‐
●
●
●
●
●
●
● NT NT
● ● ● ●
NT
●
●
●
●
y
‐
y
y
y
●
●
● RE RE
EN NT
VU
‐ ‐
●
●
‐
●
●
●
●
●
‐
‐
●
●
●
●
EN
VU
●
●
●
●
●
●
●
●
●
● ●
‐
NT
●
●
●
●
●
●
●
EN
● ●
●
● ●
NE ‐
● ●
● VU
● ●
●
●
‐
●
●
●
●
●
●
●
●
●
NT
CR
●
●
●
VU
●
?
‐
●
VU
●
●
●
●
●
●
NT
NT
NT
● DD ●
●
‐
●
VU
‐
‐
‐
‐
● DD
● NT
‐
●
●
●
EN
●
CR
●
●
●
‐
●
DD*
●
●
●
‐
●
●
●
‐
y
● ●
VU
□
RE
NT
y
y
y
CR
● VU
●
●
●
□
●
●
●
●
●
VU
EN
‐
●
●
●
●
Vatican City
Sicily
Sardinia
San Marino
Portugal
Monaco
DD
DD
● DD DD
●
□
●
●
Malta
‐
DD
●
●
Madeira
CR
NT
■
●
□
Italy
●
●
■
●
France
●
■
●
●
‐
‐
Cyprus
DD
● DD ●
□
●
EN
● ●
■
■
● ●
Corsica
DD
□
EN ●
●
NT
● EN ‐
● ● ●
●
●
●
‐
●
●
□
●
●
●
●
●
■
‐
●
DD
□
●
●
□
‐
NT
NT
■
‐
●
‐
EN
EN ● NT
● ● ● ● ● ●
EN ● ●
● NT ● ● ● ●
●
●
●
● ● ● ● ‐ ● ●
□
●
●
■
□
y
●
CR
■
■
●
● VU
●
□
●
□
Balearic Islands
●
Canary Islands
● ● ●
● EN
Azores
●
●
Northern Ireland
●
Andorra
●
Ireland
●
Great Britain
‐
Gibraltar
●
EN
●
●
●
●
■
●
y
y
y
y
y
y
y
y
NT ●
● ●
□
Channel Islands
NT ‐
Sweden
●
y
Svalbard
Taxon
Scapania nemorea Scapania nimbosa Scapania obcordata Scapania obscura Scapania ornithopodoides Scapania paludicola Scapania paludicola var. paludicola Scapania paludicola var. rotundiloba Scapania paludosa Scapania parvifolia Scapania praetervisa Scapania scandica Scapania scandica var. argutedentata Scapania scandica var. grandiretis Scapania scandica var. scandica
Scapania scapanioides Scapania simmonsii Scapania sphaerifera Scapania spitsbergensis Scapania subalpina Scapania tundrae Scapania uliginosa Scapania umbrosa Scapania undulata Scapania undulata var. minor Scapania undulata var. undulata
Scapania verrucosa Scapania zemliae Schistochilopsis grandiretis Schistochilopsis hyperarctica Schistochilopsis incisa Schistochilopsis incisa var. incisa Schistochilopsis incisa var. inermis Schistochilopsis opacifolia Schizophyllum sphenoloboides Schljakovia kunzeana Schljakovianthus quadrilobus Solenostoma callithrix Solenostoma caucasicum Solenostoma confertissimum Solenostoma gracillimum Solenostoma handelii Solenostoma hyalinum Solenostoma obovatum Solenostoma paroicum Solenostoma pusillum Solenostoma sphaerocarpum Solenostoma subellipticum Southbya nigrella Southbya tophacea Sphaerocarpos michelii Sphaerocarpos stipitatus Sphaerocarpos texanus Sphenolobopsis pearsonii Sphenolobus minutus Sphenolobus saxicola Syzygiella autumnalis Syzygiella rubricaulis Targionia hypophylla Targionia lorbeeriana Telaranea azorica Telaranea europaea Telaranea murphyae Telaranea nematodes Telaranea sejuncta Tetralophozia filiformis Tetralophozia setiformis Trichocolea tomentella Tricholepidozia tetradactyla Trilophozia quinquedentata Tritomaria exsecta Tritomaria exsectiformis Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
VU
I
□
‐
CR
□
VU ‐ VU 3
DD*
CR
VU ● VU D
VU ● DD D
I
●
R
●
VU ●
VU
VU VU
●
●
●
□
□
VU
□
■
□
●
EN
□
□
■
I
■
●
■
VU
□
EN
□
?
●
□
VU VU
●
□
●
■
●
□
1
■
●
□
●
□
●
●
●
●
●
□
VU
□
VU
●
●
■
●
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
RE
NT
3
●
●
●
1
V
V
■
?
□
EN
VU EN
□
VU D
●
■
NE
■
●
■
●
EN 2
EN D
●
VU NE
● NE
□
●
NT NE
● NE
R
‐
‐
● NE
VU NE
VU RE
□
□
CR EN
●
●
VU
R
□
VU
VU
●
●
●
●
●
●
●
●
●
●
●
●
□
‐
■
‐
■
VU
●
3
●
RE
●
●
● V
VU G
VU NE
●
●
CR
2
VU
●
●
●
VU
●
2
3
●
●
●
●
●
●
●
●
●
V
V
V
R
EN
●
■
●
●
NT CR
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
■
●
■
■
□
●
●
●
●
●
●
● DD
● VU
●
●
●
●
■
EN
□
●
□
□
□
■
●
●
●
?
●
●
●
●
●
●
●
●
●
●
■
●
●
●
●
VU
●
●
●
●
●
●
●
●
VU
VU
●
□
□
●
□
●
□
□
■
VU
□
□
□
■
●
●
EN
●
V
R
DD ●
NT NT
VU
●
●
VU NT
●
●
VU
VU
●
●
●
●
●
●
‐
‐
●
●
VU VU
●
□
●
□
●
●
●
●
●
■
■
●
●
VU
●
●
●
CR
●
●
●
●
●
●
●
RE
‐
CR
●
●
●
●
●
●
●
VU NT
●
‐
●
●
●
●
●
‐
●
NT
EN
●
●
● DD
●
●
EN
●
●
‐
● ●
DD* ●
●
●
●
●
EN
●
?
●
●
●
●
●
●
●
‐
●
●
EN
●
●
●
●
●
□
●
□
●
■
■
DD* EN
●
□
EN
EN
●
●
●
●
●
RE ‐
● EN
●
● DD
□ □
● DD* ●
DD* ●
DD* ●
EN
VU
●
□
●
●
RE
‐
●
●
●
■
RE
‐
●
● DD* CR
DD* VU
●
■
●
R
●
Ukraine
●
Russia South
●
□
●
□
●
EN
●
■
■
●
R
●
□
●
●
EN
●
●
●
●
3
●
□
‐
●
CR VU
●
●
●
‐
●
RE
DD
●
●
●
●
●
●
●
●
●
‐
DD
3
□
□
VU
1
●
NE
VU
□
●
Russia Northwest
‐
Russia North
VU
Russia East
●
Novaya Zemlya
●
Russia Central
●
Moldova
●
Lithuania
●
Latvia
●
Kaliningrad
●
Kazakhstan
VU
Franz Josef Land
●
Estonia
●
Crimea
●
Caucasus
Bulgaria
●
Belarus
Bosnia‐Herzegovina
●
Turkey
Albania
●
Slovenia
Switzerland
●
Serbia
Slovakia
●
Romania
Poland
EN
Montenegro
Netherlands
●
Macedonia
Luxembourg
●
Kosovo
Liechtenstein
V
Hungary
Germany
●
Greece
Czech Republic
●
Appendix 6 - Liverworts
Croatia
Belgium
●
Crete
Austria
Checklist and country status of European bryophytes
●
●
EN
●
■
●
■
●
●
NT
●
●
●
●
?
●
DD
●
DD* ●
●
●
●
●
●
●
●
●
‐
‐
‐
●
EN
●
□
●
●
●
●
●
□
●
●
●
●
●
EN
RE
●
●
VU
●
●
VU
●
●
●
□
●
□
●
□
●
●
●
●
●
■
●
□
●
?
●
●
●
●
●
●
●
●
●
●
□
●
●
●
●
●
‐
●
‐
●
●
●
●
●
●
●
●
■
■
●
●
●
●
●
●
●
●
●
●
●
■
■
●
●
■
●
□
●
‐
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
‐
●
●
●
●
●
‐
●
‐
‐
‐
DD* ●
CR
● ●
NT
●
●
●
●
●
●
● DD
●
EN
●
‐
●
EN
●
●
NT
VU
DD
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
VU
●
●
‐
‐
●
●
3
●
●
?
●
‐
●
●
●
‐
●
●
●
●
‐
●
●
●
●
●
●
●
‐
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
1
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
60
61
y
‐
●
□
DD
‐
●
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
Monaco
■
NT
Malta
■
●
Madeira
Italy
Cyprus
Corsica
□
France
□
DD
Canary Islands
□
Balearic Islands
□
Azores
Northern Ireland
□
Andorra
Ireland
■
●
Gibraltar
?
□
●
Channel Islands
●
■
●
Great Britain
□
●
Sweden
■
Svalbard
Taxon
Tritomaria exsectiformis subsp. arctica Tritomaria exsectiformis subsp. exsectiformis
Tritomaria scitula Tylimanthus anisodontus Tylimanthus laxus Norway
Red/italic status values with a darker fill indicate note present
Iceland
‐ Literature record but later rejected
Finland
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
Faeroe Islands
■ Infraspecific taxon confirmed
Include in candidate list?
● Species conrmed; LC/status unknown
Appendix 6 - Liverworts
Denmark
Checklist and country status of European bryophytes
□
□
□
R
□
CR
■
●
□
Greece
Hungary
□
□
□
■
■
VU
■
□
□
●
Lithuania
□
□
Russia North
□
‐
□
●
□
●
□
□
Ukraine
Russia South
Russia East
‐
Russia Northwest
Novaya Zemlya
Moldova
Latvia
□
Russia Central
‐
Kaliningrad
□
Kazakhstan
Franz Josef Land
Estonia
Crimea
Caucasus
Belarus
Turkey
Slovenia
Serbia
Romania
Montenegro
Macedonia
Kosovo
Croatia
□
●
Crete
Bulgaria
Bosnia‐Herzegovina
Albania
Switzerland
□
Slovakia
□
Poland
Netherlands
□
0
Luxembourg
□
?
Liechtenstein
Germany
Belgium
■
●
Czech Republic
Austria
Checklist and country status of European bryophytes
Appendix 6 - Liverworts
□
●
62
Checklist and country status of European bryophytes
Status notes (Status values italicised in red with a darker background fill)
63
Taxon
Country
Note
Anastrophyllum assimile
Aneura maxima
Barbilophozia hatcheri
Barbilophozia hatcheri
Calypogeia integristipula
Calypogeia sphagnicola
Cephalozia catenulata
Cephalozia catenulata
Cephalozia loitlesbergeri
Cephalozia pleniceps
Cephaloziella calyculata
Cephaloziella calyculata
Cephaloziella elachista
Cephaloziella grimsulana
Cephaloziella integerrima
Cephaloziella massalongi
Cephaloziella phyllacantha
Cephaloziella spinigera
Cephaloziella varians
Clevea spathysii
Cololejeunea rossettiana
Cololejeunea rossettiana
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Conocephalum conicum
Douinia ovata
Drepanolejeunea hamatifolia
Dumortiera hirsuta
Eremonotus myriocarpus
Exormotheca pustulosa
Fossombronia foveolata
Fossombronia maritima
FRANCE
SLOVAKIA
MONTENEGRO
SERBIA
FRANCE
FRANCE
FRANCE
SPAIN
FRANCE
FRANCE
FRANCE
SERBIA
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
UKRAINE
MONTENEGRO
UKRAINE
BELARUS
CORSICA
CRETE
DENMARK
ESTONIA
LATVIA
LITHUANIA
MACEDONIA
RUSSIA NW
SARDINIA
SERBIA
SICILY
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
SWITZERLAND
TURKEY
Fossombronia mittenii
Gymnomitrion brevissimum
Gymnomitrion commutatum
Gymnomitrion obtusum
Harpalejeunea molleri
Harpanthus flotovianus
Hygrobiella laxifolia
Jungermannia borealis
Jungermannia polaris
Kurzia sylvatica
Kurzia trichoclados
Lejeunea patens
Lepidozia cupressina
Lophozolea fragrans
Lophozia ascendens
Lophozia guttulata
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
AUSTRIA
● (at risk)
● (DD suggested)
Low risk
Low risk
● (at risk)
● (at risk)
● (at risk)
DD-vanished
● (at risk)
● (at risk)
● (at risk)
Low risk
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
Rare
Low risk
Rare
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
s.l.?
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
RE (but refound since)
Included on basis of information received
(Mesud Kirmaci via Papp, March 2014), but
seems unlikely
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
VU according to Christian Schröck, as L.
longiflora
Checklist and country status of European bryophytes
Taxon
Country
Note
Lophoziopsis propagulifera (=Lophozia
propagulifera)
Mannia controversa
Mannia triandra
Marsupella boeckii
Marsupella condensata
Mesoptychia gillmanii
Moerckia blyttii
Moerckia flotoviana
Moerckia flotoviana
Moerckia hibernica
Moerckia hibernica
Moerckia hibernica
Moerckia hibernica
Mylia taylorii
Nardia insecta
Neoorthocaulis floerkei
Neoorthocaulis floerkei
Odontoschisma elongatum
Oleolophozia perssonii
Orthocaulis atlanticus
Oxymitra incrassata
Peltolepis quadrata
Plagiochasma rupestre
Plagiochila bifaria
Plagiochila exigua
Plagiochila spinulosa
Pleurocladula albescens
Pleurocladula albescens
Prasanthus suecicus
Radula lindenbergiana
Radula visianica
SWITZERLAND
? (name not known to NS)
FRANCE
SERBIA
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
GERMANY
ANDORRA
FINLAND
LATVIA
SLOVAKIA
FRANCE
FRANCE
FRANCE
SERBIA
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
ANDORRA
FRANCE
UKRAINE
FRANCE
FRANCE
ITALY
Riccardia incurvata
Riccia cavernosa
Riccia lamellosa
Riccia macrocarpa
Riccia papillosa
Riccia trabutiana
Sauteria alpina
Scapania crassiretis
FRANCE
ROMANIA
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
BULGARIA
Scapania helvetica
Scapania paludicola
Scapania paludosa
Scapania parvifolia
UKRAINE
FRANCE
FRANCE
BULGARIA
Schljakovia kunzeana
Solenostoma subellipticum
Sphenolobus minutus
Telaranea europaea
Tritomaria scitula
FRANCE
SPAIN
SERBIA
FRANCE
FRANCE
● (at risk)
Low risk
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
s.l.
s.l.?
VU (s.l. or s.s.?)
s.l.?
s.l.?
● (at risk)
● (at risk)
● (at risk)
Low risk
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
DD-vanished
● (at risk)
Rare
● (at risk)
● (at risk)
after Köckinger only an aberrant form of male
R. lindenbergiana (N. Schnyder, 13.2.14). Ref?
● (at risk)
DD (now confirmed)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
There is one record by Duell (1999), det. Duda
(comm. Natcheva March 2014)
Rare
● (at risk)
● (at risk)
There is one record by Duell (1999), det. Duda
(comm. Natcheva March 2014)
● (at risk)
DD-vanished
Low risk
● (at risk)
● (at risk)
64
Checklist and country status of European bryophytes
Appendix 7. Checklist and country status of European mosses
This table is based upon a working list supplied by Nick Hodgetts (October 2013) and a revision supplied on
26/07/2014. Nomenclature is based on the current European moss checklist (Hill et al. 2006), updated with
additions and nomenclatural changes from more recent literature. A list of taxa and references for nomenclatural
changes from Hill et al. 2006 is provided. The subdivisions of Russia are based on the system used by Ignatov et al.
2006. Occurrence and Threat data are based on the latest available checklists from each country/territory, backed
up with information from the network of ECCB Country Contacts. Most countries that have a Red List use IUCN
threat categories to assign a threat status to each taxon. However, where countries use their own system of threat
categories, the symbols are explained in the Legend (see below). Only published records are included in the table.
Where no checklist exists for a country/territory, information has been compiled from miscellaneous published
sources and revised by the local ECCB contact(s).
Legend
Status values italicised in red with a darker background fill have an associated ‘Status note’ (see below)
Common categories
●
Occurrence of species confirmed - either Least Concern or no information about status
■
Occurrence of infraspecific taxon confirmed
□
At least some reports of the species presumably refer to this infraspecific taxon, although not
positively confirmed in any source
?
Some doubt about occurrence
Taxon recorded in some literature but later rejected
RE
Regionally Extinct
CR
Critically Endangered
EN
Endangered
VU
Vulnerable
NT
Near Threatened
DD
Data Deficient
DD*
Data Deficient but recently recorded
NE
Not Evaluated
y
Include in candidate list? - y = yes
Country-specific categories
Netherlands
EB
BE
KW
GE
Ernstig bedreigd (Highly Endangered)
Bedreigd (Endangered)
Kwetsbaar (Vulnerable)
Gevoulig (Susceptible)
Belgium
Mn
Menacées (Threatened)
Germany
0
1
2
3
G
R
V
D
nb
nm
Poland
Ex
E
V
R
I
65
Ausgestorben oder verschollen (Regionally Extinct)
Vom Aussterben bedroht (Critically Endangered)
Stark gefährdet (Endangered)
Gefährdet (Vulnerable)
Gefährdung anzunehmen (Risk assumed)
Extrem selten (Extremely rare)
Zuruckgehend (Near Threatened)
Daten ungenugend (Data deficient)
Nicht bewertet (subspecies/variety not evaluated because too little knowledge about this taxon)
Not mentioned (the name does not occur in the Red List at all - mostly varieties not recognised in
Germany)
Extinct
Endangered
Vulnerable
Rare
Indeterminate
Checklist and country status of European bryophytes
Austria
0
1
2
3
4
Vollständig vernichtet (Completely destroyed - i.e. Extinct)
Von vollständiger vernichtung bedroht (Seriously threatened with extinction)
Stark gefährdet (Highly endangered)
Gefährdet (Endangered)
Gefährdung anzunehmen (Risk assumed)
Latvia
0
1
2
3
4
Extinct
Endangered
Vulnerable
Rare
Little known or insufficiently explored species
Lithuania
0
1
2
3
4
Extinct
Critically Endangered
Endangered
Vulnerable
Rare
Belarus
0
Extinct
Ukraine
R
Rare
66
DD
Acaulon casasianum
y
Acaulon dertosense
y
Acaulon fontiquerianum
y
Acaulon mediterraneum
y
Acaulon muticum
y
Acaulon piligerum
y
Acaulon triquetrum
y
●
●
EN
VU
RE
●
●
●
Achrophyllum dentatum
Alleniella complanata
●
●
Aloina aloides
●
●
VU
● RE RE
● DD ●
DD ●
●
●
DD
●
●
DD
●
●
DD ●
● ●
EN
●
●
●
●
DD
●
DD ?
● DD ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
Aloina bifrons
y
Aloina brevirostris
y
Aloina humilis
y
Aloina obliquifolia
y
Aloina rigida
●
●
CR
●
● ●
EN ●
●
●
EN ●
● EN EN
●
●
●
●
●
●
●
NT
●
●
●
●
DD
● NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Amphidium lapponicum
● RE RE
●
● DD
y
Anacolia webbii
y
Andoa berthelotiana
y
VU ●
●
●
●
●
●
●
●
●
●
Andreaea blyttii
● VU
● ● ●
Andreaea frigida
y
Andreaea heinemannii
y
●
DD
●
NT ●
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
‐
●
Andreaea mutabilis
y
Andreaea nivalis
y
●
●
NT
●
●
●
?
●
●
●
●
●
●
?
DD
?
VU
●
●
●
●
●
●
□
□
●
●
Andreaea rothii subsp. rothii
VU
●
●
●
●
■
●
●
●
●
●
□
Anoectangium aestivum
y
Anoectangium handelii
y
Anomobryum bavaricum
y
□
□
●
●
□
●
NT
●
● NT ●
Anomobryum julaceum
●
●
●
●
●
■
■
■
■
■
●
□
■
■
■
■
■
●
●
●
●
●
●
■
■
■
■
■
■
●
●
y
y
Anomodon tristis
y
Anomodon viticulosus
■
■
□
□
●
●
●
●
VU ● DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
Aongstroemia longipes
y
●
Archidium alternifolium
●
●
●
●
●
EN ●
●
●
●
●
Arctoa fulvella
●
NT
VU ●
● EN ● NT
●
●
● VU ● VU
●
y
Aschisma carniolicum
y
●
●
y
Arctoa hyperborea
●
●
●
●
VU
●
●
●
●
NT
●
●
●
●
■
●
●
●
■
VU
VU
VU
●
VU
VU
●
VU
●
●
■
DD
●
●
●
●
■
■
■
□
■
■
■
■
■
●
●
●
●
●
■
■
□
■
■
□
□
●
●
●
●
●
●
●
●
EN
VU
●
●
●
●
● DD ●
● NT ●
Antitrichia californica
Antitrichia curtipendula
●
■
■
●
●
●
●
DD
●
●
●
DD NT
●
●
●
Anomodon rugelii
●
‐
y
Anomodon rostratus
●
●
?
●
●
Anomodon longifolius
●
VU
Anomobryum concinnatum
Anomodon attenuatus
●
●
●
■
●
■
●
■
Andreaea rupestris var. papillosa
y
●
●
● VU VU
●
Andreaea rothii subsp. falcata
Andreaea sinuosa
●
●
●
●
Andreaea rothii
Andreaea rupestris var. rupestris
●
●
Andreaea obovata
Andreaea rupestris
●
VU
●
VU
y
●
VU
VU
Andreaea megistospora
●
●
EN
●
Andreaea heinemannii subsp. heinemannii
67
●
●
Andreaea heinemannii subsp. crassifolia
Arctoa anderssonii
●
DD
●
Andreaea alpina
Andreaea flexuosa ●
●
●
●
y
●
●
●
●
Andreaea alpestris
Andreaea crassinervia
●
●
●
●
●
y
y
●
●
DD
●
●
Anacamptodon splachnoides
DD ●
● DD
●
●
Amblystegium serpens
Aplodon wormskioldii
●
●
DD
y
Anacolia menziesii
●
●
●
Amphidium mougeotii
●
●
●
Amblyodon dealbatus
Anomobryum lusitanicum
EN
●
VU
●
NT
●
Aloina ambigua
Amphidium tortuosum
VU
●
●
●
NT
Alleniella besseri
Alophosia azorica
■
NT
●
RE
●
●
●
●
●
● NT NT
●
●
●
●
NT CR
●
CR
● ●
●
●
●
●
●
●
VU
●
●
●
●
●
VU
●
●
VU
●
●
‐
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT ●
● LC
VU
● VU
VU
●
CR
●
●
●
●
●
●
●
DD
●
VU
Vatican City
■
Spain
■
■ NT NT
Abietinella abietina var. hystricosa
Sicily
●
■
Sardinia
●
■
San Marino
●
■
Portugal
●
Monaco
●
Malta
Italy
●
Madeira
France
●
■ EN EN □
Cyprus
Andorra
●
■
Corsica
Northern Ireland
●
□
Canary Islands
Ireland
●
□
Balearic Islands
Great Britain
●
□
Azores
Svalbard
●
■
Gibraltar
Norway
●
□
Channel Islands
Iceland
●
Sweden
Finland
Abietinella abietina var. abietina
Faeroe Islands
Taxon
Abietinella abietina
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
●
●
●
□
■
□
□
□
□
DD D
■
EN
■
NT BE
R
□
Ukraine
●
□
Sub‐polar & North Urals
●
□
SE Russia
Central Russia
●
■
NW Russia
Caucasus (in Europe)
●
■
NE Russia
Belarus
●
■
Moldova
Arctic Russia
●
□
Middle and South Urals
Slovenia
●
□
Lithuania
Serbia
●
□
Latvia
Romania
●
■
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
□
EN
Kazakhstan (in Europe)
Montenegro
●
□
Kaliningrad
Macedonia
●
□
Estonia
Albania
●
■
Crimea
Switzerland
●
□
Turkey
Slovakia
EB
■
Kosovo
Poland
●
□
Hungary
Netherlands
●
V
Greece
Luxembourg
nb
■
Croatia
Liechtenstein
●
□
Crete
Germany
●
■
Bulgaria
Czech Republic
●
Bosnia‐Herzegovina
Belgium
Appendix 7 - Mosses
Austria
Checklist and country status of European bryophytes
■
2
2
2
●
●
3
●
VU
●
3
●
●
DD
●
●
●
DD
●
3
●
2
●
EN
V
●
V
V
VU CR
DD* ●
● DD* ●
●
●
VU VU
?
●
●
●
●
●
●
●
NT
●
CR EN
NT
DD
VU
●
DD
●
●
0
●
DD*
●
●
●
R
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
BE
●
●
R
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
DD
●
●
●
NT
●
I
RE DD* ●
●
NT
●
●
●
NT
●
CR
●
VU
●
DD*
CR
NE
3
●
●
●
●
●
G
●
R
●
V
2
●
●
●
●
●
EN
●
1
●
EN
4
CR
●
CR
GE
I
NT GE
●
VU
●
DD
NT
●
●
●
●
NT
RE
E
●
●
●
●
R
●
EN
●
●
●
3
E
EN RE
nb
●
●
●
●
0
●
EN VU
DD
●
●
●
●
●
●
DD*
●
●
●
●
●
●
VU
●
●
●
●
RE
●
●
●
●
●
?
●
VU
●
VU
EN
●
●
DD*
●
●
●
●
0
●
●
●
●
●
●
●
●
●
●
CR DD DD
● VU ●
VU DD
●
●
●
●
●
●
●
NT
●
●
RE
R
■
G
□
■
4
□
EN
3
●
●
●
●
■
nb
EN
R
CR
●
CR CR DD
●
EN
●
■
□
VU
●
●
□
□
DD*
VU
●
●
3
●
3
●
●
□
●
●
●
VU
●
●
●
■
□
□
●
VU
●
●
●
DD*
nm
I
NT NT
DD
1
VU
3
●
●
3
●
●
3
●
●
■
■
■
●
●
●
●
BE
●
●
I
●
●
V
VU
●
CR
V
RE nm
●
EN
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
?
□
CR
●
NT
1
●
●
●
●
●
●
●
●
□
□
●
●
●
●
■
■
■
■
■
■
□
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
NT VU
●
●
BE
●
●
●
BE
E
2
●
●
●
DD* NT
●
●
●
●
●
●
●
●
●
●
●
NT
●
CR
●
NT
EN VU
●
EN
●
●
CR
●
DD
●
DD
●
●
●
DD
●
●
NT VU
●
EN
●
●
VU
●
●
●
●
●
●
R
EN CR
●
EN VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
VU
EN
●
●
EN
●
●
●
●
●
●
NT
●
2
1
●
●
0
●
●
■
■
●
3
●
DD*
3
●
●
VU
□
●
■
VU
●
●
□
■
□
●
EN
CR
nm
●
●
●
DD*
●
●
RE
●
●
CR
EN NT
CR KW ●
nb
●
●
●
CR VU
nb
CR
●
●
R
●
●
●
●
●
●
I
●
●
●
□
●
●
CR VU
■
●
●
●
CR
nm
□
2
R
■
■
●
R
●
■
●
●
●
VU
4
●
●
●
R
●
●
●
●
●
4
●
●
CR
●
CR
DD
●
●
●
2
DD*
●
●
?
1
●
●
●
●
●
DD
VU VU
●
CR
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
1
●
●
●
CR
●
DD
68
CR RE RE
●
●
●
●
●
● NT NT
●
●
●
●
●
●
●
●
●
● ● ● ●
● VU VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Aulacomnium androgynum
Aulacomnium palustre
●
Aulacomnium turgidum
●
●
●
?
●
●
■
●
Barbula convoluta var. sardoa
Barbula crocea
DD
●
DD
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
●
■
■
■
■
■
□
■
□
□
□
■
■
□
■
■
■
■
■
■
■
■
■
■
■
■
■
NT
●
●
■
CR
y
■
VU
DD
●
●
●
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD
Bartramia halleriana
●
Bartramia ithyphylla
●
●
●
□
Barbula unguiculata
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
● RE RE
●
●
●
●
● VU VU ●
●
●
●
DD
●
●
●
NT
●
y
Bartramia pomiformis
●
●
Bartramia rosamrosiae ●
●
●
CR
●
●
● DD
●
NT ●
●
●
●
●
y
Blindia acuta
●
Blindia caespiticia
y
Brachydontium trichodes
y
●
●
●
CR
●
●
●
●
●
●
VU
DD
●
●
●
●
●
● EN EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
EN
Brachymenium commutatum
‐
Brachymenium notarisii
y
Brachymenium paradoxum
y
Brachymenium philonotula
y
Brachytheciastrum collinum
y
Brachytheciastrum dieckei
y
Brachytheciastrum fendleri
y
●
VU ●
●
●
●
●
●
●
?
‐
y
Brachytheciastrum vanekii
y
Brachytheciastrum velutinum
●
NT
●
●
●
●
●
●
●
●
●
●
VU
●
● EN EN ●
Brachytheciastrum velutinum var. salicinum
●
●
●
●
●
●
●
●
Brachytheciastrum olympicum
Brachytheciastrum trachypodium
●
●
●
■
●
●
●
●
●
●
●
●
●
■
■
●
●
●
DD
●
‐
●
●
●
●
■
■
DD
■
Brachytheciastrum velutinum var. vagans
Brachytheciastrum velutinum var. velutinum
□
Brachythecium albicans
●
Brachythecium buchananii ●
□
□
■
□
□
□
□
●
●
●
●
●
●
●
●
DD
●
■
□
■
□
●
□
□
●
■
■
□
●
●
●
□
■
■
■
■
●
●
DD
●
DD
●
●
●
●
CR
y
NT
Brachythecium campestre
Brachythecium capillaceum
□
y
Brachythecium cirrosum
RE ●
Brachythecium coruscum
VU
●
Brachythecium erythrorrhizon
●
● VU
DD ●
●
VU
●
●
●
DD
●
●
●
●
DD
●
●
●
●
□
Brachythecium erythrorrhizon subsp. asiaticum
Brachythecium erythrorrhizon subsp. erythrorrhizon
Brachythecium geheebii
■
y
Brachythecium glareosum
Brachythecium laetum
●
●
y
●
□
□
□
■
●
EN
●
●
●
●
●
●
●
●
●
●
‐
VU
■
□
□
●
●
●
●
●
●
●
DD*
Brachythecium mildeanum
●
Brachythecium mildeanum var. mildeanum
□
●
Brachythecium mildeanum var. udum
●
●
●
●
●
●
●
●
■
□
□
■
□
□
□
□
■
‐
●
□
■
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Brachythecium rutabulum var. rutabulum
□
□
■
□
□
□
□
□
□
□
□
□
□
□
□
■
Brachythecium salebrosum
●
●
NT
●
●
●
● DD ●
●
●
●
●
●
Brachythecium rutabulum var. atlanticum
●
Braunia alopecura
y
Braunia imberbis y
●
Breidleria pratensis
Breutelia azorica
69
●
■
Brachythecium turgidum
y
●
●
●
●
●
VU
● ●
●
●
■
Brachythecium rutabulum
Brachythecium tommasinii
●
EN
Brachythecium rivulare
● ● ●
EN
NT
●
●
● VU VU
●
●
●
●
●
●
●
●
●
●
●
●
□
■
□
■
□
●
●
●
●
●
●
■
●
●
●
●
●
●
DD
●
?
□
●
VU
DD
Vatican City
●
Spain
●
Sicily
●
●
Sardinia
●
●
San Marino
●
Portugal
●
●
Monaco
?
●
Malta
●
Madeira
Cyprus
Corsica
Canary Islands
●
y
●
Bartramia subulata
VU ●
NT ●
DD ●
Barbula convoluta var. convoluta
Bartramia laevisphaera ●
●
Barbula convoluta
Bartramia breviseta
VU ●
●
Barbula bolleana
Barbula enderesii
Balearic Islands
●
●
●
Atrichum undulatum
Barbula consanguinea Azores
Andorra
Northern Ireland
Ireland
Great Britain
CR
●
●
●
y
Italy
●
Atrichum tenellum
y
France
●
Atrichum flavisetum
Barbula bicolor
●
VU
EN
Atrichum crispum
Barbula amplexifolia Gibraltar
Channel Islands
Sweden
y
Svalbard
y
Atrichum angustatum
Norway
Atrichum androgynum Iceland
y
Finland
y
Atractylocarpus alpinus
Faeroe Islands
Taxon
Aschisma cuynetii
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
NT
3
●
●
●
●
●
●
●
●
●
●
●
●
●
VU KW ●
●
●
V
●
VU
●
4
●
●
DD
●
●
●
●
VU
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
R
EN
DD*
R
VU
VU
●
Ukraine
Sub‐polar & North Urals
SE Russia
●
NW Russia
3
NE Russia
●
Moldova
Middle and South Urals
●
Lithuania
●
Kazakhstan (in Europe)
●
Kaliningrad
●
Estonia
●
Crimea
●
●
●
●
●
●
●
□
□
■
■
□
■
□
DD
■
●
I
●
●
●
●
●
●
●
●
NT
3
●
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
?
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
●
●
●
3
●
3
●
●
●
●
●
●
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
■
■
□
■
■
□
□
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
■
■
■
■
VU
●
●
■
●
●
●
●
■
●
●
VU
●
DD*
●
●
●
●
●
●
V
●
Latvia
●
4
D
●
Central Russia
DD
●
RE
●
Caucasus (in Europe)
●
●
●
3
CR
●
Belarus
GE
DD D
3
●
RE
3
●
●
Arctic Russia
●
Turkey
●
Slovenia
●
Serbia
NT
Romania
●
Montenegro
●
Macedonia
DD
Kosovo
Hungary
●
Crete
Albania
Switzerland
Slovakia
Poland
Netherlands
Luxembourg
Liechtenstein
Greece
●
Croatia
RE EN
Bulgaria
●
R
Appendix 7 - Mosses
Bosnia‐Herzegovina
1
Germany
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
●
●
EN VU
●
●
●
CR
●
●
●
NT RE
●
●
●
●
●
●
VU
●
●
●
NT DD
‐
●
●
●
●
●
●
●
●
●
●
●
● DD* ●
●
CR VU
NT CR
●
●
●
VU DD
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
DD
●
●
●
VU
EN
●
VU
●
EN
DD
●
●
●
●
●
●
●
●
RE
●
●
0
VU
●
2
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
NE
●
●
●
●
●
●
nm
V
●
EB
2
4
VU
●
●
●
V
R
4
●
●
●
V
R
VU
●
●
●
●
●
●
●
DD
CR
●
●
●
●
R
I
●
●
●
DD*
DD
●
●
●
●
DD
●
●
●
●
●
D
●
●
●
●
●
D
□
□
■
●
●
●
●
3
●
2
●
●
NT VU
I
EN
●
●
●
●
●
●
●
●
■
■
■
●
●
●
■
●
●
DD
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
□
■
□
□
□
■
□
□
□
■
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
DD
EN ●
●
●
DD NT
VU
●
●
DD
●
●
●
●
●
NT
NT
?
●
EN
DD
●
●
●
●
VU
●
●
●
●
●
DD*
?
●
?
●
●
●
●
●
●
NT
●
●
●
●
●
■
□
nm
4
●
3
●
EN
3
●
●
Mn EN
●
■
□
D
nb
●
■
□
DD D
●
4
●
●
●
■
D
□
●
□
●
V
□
●
BE
□
V
●
●
●
●
CR
□
CR DD* ●
●
●
EN
●
●
□
□
□
□
●
●
DD
●
●
CR
●
NT
●
●
NT
●
●
●
●
●
□
□
DD*
●
●
●
●
?
●
?
●
●
●
□
●
●
●
■
■
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
● KW ●
●
●
●
● DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
nm
□
□
□
□
□
□
□
□
□
□
□
□
□
■
□
■
■
□
□
■
□
□
■
□
■
■
■
■
□
■
■
□
●
●
●
□
●
□
●
●
?
●
3
□
● DD*
□
●
●
●
■
●
●
●
●
●
●
● KW ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
■
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
■
nm
□
□
□
□
□
□
□
□
□
●
●
●
Mn
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
I
VU
●
4
R
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
1
RE VU
2
Mn
●
2
●
CR
●
E
●
●
CR
●
●
●
NT
●
●
●
●
●
4
●
●
●
70
y
Bruchia flexuosa
y
Bruchia vogesiaca
y
Bryhnia scabrida
y
Bryobrittonia longipes
y
Bryoerythrophyllum alpigenum
y
Bryoerythrophyllum caledonicum
y
Bryoerythrophyllum campylocarpum
y
●
Bryoxiphium madeirense
y
Bryoxiphium norvegicum
y
Bryum apiculatum
y
Bryum argenteum
Bryum austriacum y
Bryum blindii
y
Bryum calophyllum
y
Bryum cellulare
y
Bryum cryophilum
y
Bryum demaretianum
y
Bryum dichotomum
y
Bryum dyffrynense
y
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
EN
●
●
●
●
●
●
●
Bryum marratii
y
●
NT
VU ●
Bryum miniatum
y
Bryum minii
y
Bryum muehlenbeckii
y
Bryum oblongum
y
EN ● DD
●
●
VU EN
NT
Bryum kunzei
●
●
●
●
●
NT
●
●
DD
NT
VU
●
DD ●
●
●
● DD
●
●
● DD ● NT
●
Bryum turbinatum
y
●
NT
Bryum valparaisense
y
Bryum versicolor y
Bryum violaceum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
DD ●
●
●
●
●
●
●
●
●
DD
?
●
●
●
●
●
VU
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
VU
DD
●
DD
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
VU
●
●
●
VU
●
●
EN
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
EN CR
DD ●
● ●
CR
●
●
●
y
●
●
●
●
●
●
●
● DD
●
VU
VU
●
●
●
● ● ●
EN ● DD
●
y
●
EN
●
● ●
NT
DD ●
●
●
DD
●
●
●
●
● ● ●
NT EN
●
y
●
●
●
●
NT RE
●
CR
●
●
NT
●
●
NT
●
●
●
●
●
●
●
●
●
●
y
Calliergon richardsonii
●
Calliergonella cuspidata
●
Calliergonella lindbergii
●
●
DD
●
●
?
DD
●
●
●
DD
●
●
●
y
Callicladium haldanianum
●
●
DD
VU
●
DD
‐
●
●
●
NT ● VU
Buxbaumia aphylla
Calliergon giganteum
●
RE RE
Buxbaumia viridis
Calliergon cordifolium
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
● ●
●
●
●
●
●
●
●
●
Calomnion complanatum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD ●
?
●
●
EN
●
●
CR
●
VU
‐
●
●
●
●
VU
EN
●
●
●
●
●
●
●
●
DD
●
●
NT
●
●
●
y
Calyptrochaeta apiculata
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
Monaco
Malta
Madeira
Italy
France
●
●
● DD
●
Bryum weigelii
71
●
y
●
Campyliadelphus chrysophyllus
●
● EN EN
DD
●
y
Calymperes erosum
●
NT
●
y
Bryum tenuisetum
Calliergon megalophyllum
DD
●
●
Bryum subapiculatum
Bryum wrightii
●
●
DD
●
Bryum warneum
●
NT NT NT
● VU
●
●
Bryum sibiricum ●
●
●
●
y
●
EN
●
●
●
●
Bryum sauteri
●
DD
●
● NT ●
Bryum schleicheri
●
●
●
y
●
DD
EN ●
●
Bryum salinum
●
VU EN
●
Bryum ruderale
●
NT
● NT ● DD
y
Bryum riparium
●
DD
●
●
Bryum radiculosum
VU
●
● DD ●
●
●
EN VU
DD EN EN
Bryum knowltonii
EN
●
NT
Bryum klinggraeffii
●
●
●
y
●
●
●
Bryum intermedium
●
●
DD
Bryum gemmilucens
y
Cyprus
●
Bryum gemmiferum
Bryum gemmiparum
●
●
●
Bryum elegans
●
NT
●
Bryum dixonii
Corsica
DD
●
y
EN
●
Bryum canariense
Bryum caucasicum
VU
VU
DD
●
y
CR
●
y
Bryoerythrophyllum rubrum
●
●
DD*
DD
● EN ●
Bryoerythrophyllum recurvirostrum
Canary Islands
●
Balearic Islands
●
Azores
Northern Ireland
●
Andorra
Ireland
Gibraltar
Great Britain
●
CR
Bryoerythrophyllum ferruginascens
Bryum funckii
Channel Islands
Sweden
Svalbard
Norway
Iceland
Finland
●
Brotherella lorentziana
Bryoerythrophyllum inaequalifolium
Faeroe Islands
Taxon
Breutelia chrysocoma
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
RE
3
0
●
●
VU
●
Ukraine
Sub‐polar & North Urals
SE Russia
NW Russia
RE
0
●
●
●
●
●
nb
4
●
●
●
●
●
4
●
NE Russia
Moldova
Middle and South Urals
Lithuania
Latvia
Kazakhstan (in Europe)
Kaliningrad
Estonia
Crimea
Central Russia
Caucasus (in Europe)
Belarus
Arctic Russia
Turkey
VU
0
0
Slovenia
Serbia
Romania
Montenegro
Macedonia
Kosovo
Hungary
Appendix 7 - Mosses
Greece
Croatia
Crete
Bulgaria
Bosnia‐Herzegovina
Albania
Switzerland
Slovakia
Poland
Netherlands
Luxembourg
Liechtenstein
Germany
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
●
VU
●
●
●
●
VU
CR
R
●
CR
●
D
●
●
●
●
●
2
nb
●
●
●
●
●
●
●
●
●
●
●
CR EN
●
●
●
●
●
GE
V
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
DD
●
VU
●
●
●
●
●
●
●
DD
CR
VU
0
●
●
●
●
DD
CR
0
RE
●
●
●
●
●
●
●
●
●
●
?
●
4
●
●
●
●
●
●
●
●
NT
●
●
3
●
DD
3
●
●
●
3
D
●
●
VU
●
NT
D
VU
3
●
●
CR
2
V
●
●
●
●
1
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
● DD* ●
●
●
●
●
●
BE
●
●
EN
CR
EN VU
●
KW Ex CR
R
●
●
●
●
DD
●
●
●
NT
●
●
●
●
●
NT EN
DD VU
NT
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
EN
●
●
NT
●
●
●
VU
VU DD
1
D
*
●
VU
DD*
EN KW ●
?
3
DD D
RE NT
●
DD VU
●
●
●
●
●
●
●
VU
VU
●
●
DD*
●
VU
●
●
●
●
●
●
●
●
●
●
DD
●
●
VU
●
●
●
●
●
VU
●
DD
●
●
●
●
●
●
VU
●
●
●
●
DD*
●
●
●
●
●
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR VU
DD
●
●
●
●
●
●
●
●
●
CR
●
●
●
●
EN
DD
CR
EN
●
DD
DD
CR
EN
●
CR
●
●
●
●
●
●
●
EN
●
VU
●
●
●
CR
●
VU
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
4
1
2
●
●
DD nb
CR nb
●
VU
●
I
●
●
●
●
● VU
VU ●
●
●
●
?
●
DD
●
●
●
NT
●
●
●
VU
EN
●
●
●
●
●
●
●
1
●
●
●
●
●
●
●
●
●
●
●
●
4
●
DD
●
VU
●
R
●
RE EN
2
RE
I
VU
●
●
NT
●
DD*
●
CR
●
●
●
●
CR
●
●
●
●
●
●
● DD* ●
●
●
●
●
●
DD
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
RE
●
●
●
●
●
●
●
●
●
●
●
●
●
0
●
●
●
?
●
2
3
RE
●
2
●
NT
●
●
KW V
●
RE
●
V
NT
●
●
●
VU
●
●
DD
●
0
DD
●
NT
●
DD
●
CR
3
RE NT
3
2
3
VU
VU
2
●
●
VU
●
3
RE
0
1
3
3
●
●
3
●
●
●
●
●
●
●
NT EN DD
●
EN
VU
●
EN
VU NT DD
VU VU
●
NT
VU
●
●
EN EN
BE
E
●
●
●
CR EN CR NT
EN ● EN ●
●
●
VU BE
●
●
● VU
NT ●
●
●
●
EN
●
●
NT
CR
RE
E
E
VU
●
●
●
●
●
●
●
●
V
●
●
●
●
●
●
●
●
●
● KW ●
●
●
NT EB
●
●
●
●
●
●
●
1
●
●
●
●
●
●
●
VU EB
●
●
●
?
●
● DD* ●
VU ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
3
●
1
●
3
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
●
1
●
1
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
3
●
●
VU
●
●
●
VU NT
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
VU
NT
DD
●
●
●
●
●
●
●
●
●
●
DD
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
72
y
Campylidium calcareum ●
●
●
Campylidium sommerfeltii Campylium laxifolium
y
y
●
●
EN ● DD
Campylium protensum
●
●
●
●
●
●
Campylium stellatum
●
●
●
●
●
Campylophyllum halleri
NT
Campylopus atrovirens
●
Campylopus brevipilus
●
Campylopus cygneus
y
Campylopus flaccidus
y
●
Campylopus fragilis
●
Campylopus gracilis
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● VU
●
●
●
●
●
●
●
y
Campylopus setifolius
y
●
Campylopus shawii
y
● NT
Campylopus subporodictyon
y
●
● DD
●
●
y
Catoscopium nigritum
y
Ceratodon conicus
y
●
● VU
●
●
●
●
●
●
●
●
●
●
●
●
RE
●
●
●
●
Ceratodon purpureus subsp. purpureus
□
□
■
□
□
□
■
VU ●
●
●
Cinclidium subrotundum
Cinclidotus aquaticus
y
Cinclidotus confertus
y
Cinclidotus danubicus
y
Cinclidotus fontinaloides
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
VU
RE
● VU VU
DD
●
●
●
CR
●
●
NT
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
VU
●
●
DD
EN
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
■
■
■
■
■
■
□
□
■
■
■
■
■
?
■
DD
□
■
■
■
■
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD ●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
y
VU DD
NT
y
Cirriphyllum crassinervium
●
Cirriphyllum piliferum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
y
Cleistocarpidium palustre
y
Climacium dendroides
DD
●
●
Cnestrum alpestre
●
●
Cnestrum glaucescens
y
VU
Cnestrum schisti
y
●
Conardia compacta
y
●
Conostomum tetragonum
Coscinodon cribrosus
y
Coscinodon humilis
y
Cratoneuron curvicaule
y
Cratoneuron filicinum
y
Crossidium crassinerve
y
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
● RE
●
●
●
●
VU ●
RE
●
●
●
CR ● DD
● ●
● ●
●
●
●
●
EN
●
●
●
DD ● DD
EN
●
Crossidium aberrans
●
●
●
Claopodium whippleanum
Clasmatodon parvulus ●
CR
●
●
●
●
●
●
●
●
●
●
●
● EN ●
●
DD
●
Crossidium davidai
●
●
VU
●
●
●
●
●
●
NT
●
●
?
DD
●
●
●
●
●
●
●
DD
●
●
●
Crossidium geheebii
y
Crossidium laevipilum
y
Crossidium laxefilamentosum
y
●
●
●
●
●
Crossidium squamiferum
●
●
●
●
●
Crossidium squamiferum var. pottioideum
Crossidium squamiferum var. squamiferum
●
□
●
●
● DD ●
■
Cryphaea heteromalla
●
Ctenidium molluscum
●
EN
● NT ●
●
CR ●
●
●
●
●
●
●
●
●
□
●
□
●
□
●
●
■
■
●
●
●
●
■
●
●
●
■
●
□
□
●
■
●
■
●
●
●
?
●
●
●
●
■
●
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
Monaco
Malta
Madeira
●
● VU VU
Cinclidotus riparius
73
●
●
VU
y
●
●
●
CR
Cheilothela chloropus
y
●
●
●
Ceratodon purpureus subsp. stenocarpus
Cinclidium stygium
●
●
● NT NT
●
●
Cinclidium latifolium
‐
●
VU ●
●
Cinclidotus vivesii
EN
●
● EN EN
●
Cinclidotus pachylomoides
●
●
●
DD
●
● VU
●
y
●
● RE
VU ●
Ceratodon purpureus
Cinclidium arcticum
●
●
●
NT
Campylopus subulatus
y
Italy
NT
●
●
Campylopus schimperi
Campylostelium strictum
France
●
●
●
Campylostelium saxicola
Cyprus
RE
Campylopus pilifer
y
Corsica
●
●
●
y
Campylostelium pitardii
Canary Islands
●
●
?
●
●
Campylopus pyriformis
Balearic Islands
●
●
y
Campylopus introflexus
Azores
Andorra
Northern Ireland
Ireland
Great Britain
Gibraltar
Channel Islands
Sweden
● NT NT
●
●
●
Campylopus oerstedianus
●
●
●
VU
Campylopus flexuosus
Campylopus incrassatus
Svalbard
Norway
Iceland
VU ● NT
●
Campylium longicuspis
Finland
Faeroe Islands
Taxon
Campyliadelphus elodes
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
●
●
NT
3
●
VU KW ●
●
●
●
●
●
●
EN
V
●
●
●
●
●
NT
●
●
●
●
●
●
NE
0
BE
NT
●
●
●
●
●
●
I
VU
●
3
●
●
3
●
●
GE
R
DD
●
DD
●
●
●
VU
4
R
●
DD RE
2
●
●
●
●
●
●
●
●
●
●
●
●
D
●
VU
3
EV GE
RE NT
V
●
RE
2
●
●
●
4
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD*
4
●
●
●
Ukraine
●
Middle and South Urals
Kazakhstan (in Europe)
Kaliningrad
●
●
Estonia
Belarus
Arctic Russia
Turkey
Slovenia
Serbia
Romania
Kosovo
●
●
●
●
●
VU
●
E
RE VU
CR ●
E
DD DD
●
CR EN
●
●
I
CR EN
RE
E
NT VU
VU
●
●
●
VU NT
●
DD
NT
●
●
●
NT
●
●
●
●
●
RE
●
DD*
●
●
●
●
●
DD
●
NT
RE
VU
DD
●
●
?
EN
●
4
●
?
DD
●
3
●
●
NT
Mn
●
●
EN
●
Sub‐polar & North Urals
●
●
EN VU
●
SE Russia
VU
●
●
●
NW Russia
●
●
●
●
NE Russia
●
●
CR
Moldova
●
Lithuania
●
●
Latvia
●
●
Crimea
●
?
Hungary
●
Greece
?
●
?
Central Russia
●
Croatia
EN VU DD* ● DD*
DD
DD
Crete
Bulgaria
Bosnia‐Herzegovina
Albania
Switzerland
R
Slovakia
BE
● KW ●
I
Caucasus (in Europe)
●
Montenegro
●
nb
Poland
Luxembourg
Liechtenstein
2
●
●
Appendix 7 - Mosses
Macedonia
●
Germany
Czech Republic
Mn DD
Netherlands
2
●
Belgium
Austria
Checklist and country status of European bryophytes
●
●
●
●
CR
CR
DD
CR
●
R
●
VU
●
DD VU
DD
●
0
●
NT
●
1
●
●
3
●
?
0
DD
?
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
■
■
□
□
□
□
□
□
□
■
□
□
□
■
?
□
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
●
●
VU
●
●
●
DD
DD*
VU
●
R
●
●
●
2
Mn
?
2
RE
E
CR
●
●
EN
●
EN
●
●
2
●
●
●
3
●
1
●
●
●
RE
●
VU EN
●
CR
●
●
●
NT
●
E
EN
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
?
DD
●
●
nm
●
●
VU
●
●
VU
●
●
NT
●
●
NT
●
●
●
●
●
EN
●
●
●
●
●
●
●
CR
●
1
●
●
●
●
●
●
●
●
●
E
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
0
2
●
●
Mn VU
●
V
●
●
0
?
R
EN
G
EN GE
●
R
EN VU
● KW ●
I
●
●
CR VU
I
VU
●
VU VU
EN NT
R
CR
●
●
●
CR
●
VU
●
●
●
●
●
0
3
●
●
●
●
●
●
●
●
VU GE
●
EV
●
●
DD
●
VU
●
●
●
●
●
DD*
●
●
●
DD
EN
DD
DD
EN
●
NT
●
NT
●
●
CR
●
●
●
●
●
VU
●
?
●
●
●
●
●
●
●
● DD*
●
●
●
●
●
DD
●
●
●
●
R
●
RE CR
G
VU
●
●
●
D
■
□
●
□
●
●
●
●
●
●
CR
●
EN
●
■
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
EN
●
●
●
●
●
●
●
●
VU
3
■
●
●
DD*
R
3
DD
●
●
●
●
●
BE
●
●
□
VU
□
●
□
●
●
●
□
●
□
●
□
●
■
●
●
●
●
□
●
●
□
●
□
CR
□
●
●
●
●
DD
●
□
■
□
DD
●
●
●
■
3
●
●
●
●
●
■
■
■
●
●
●
●
3
●
●
74
●
●
y
●
NT
●
NT
●
RE
●
Cynodontium gracilescens
y
●
EN
●
Cynodontium jenneri
y
●
●
●
● RE ●
●
NT
●
●
●
●
VU
●
Cynodontium strumiferum
●
●
●
●
●
●
VU
Cynodontium suecicum
y
●
Cynodontium tenellum
NT
●
●
●
●
●
●
Cyrtomnium hymenophylloides
NT ●
●
●
●
VU
●
●
●
Cyrtomnium hymenophyllum
Daltonia splachnoides
y
y
Dichelyma capillaceum
y
Dichelyma falcatum
y
Dichodontium palustre
●
●
●
●
●
●
●
●
●
●
●
EN
● ●
●
●
●
●
●
●
●
●
Dicranella cerviculata
●
●
●
●
●
Dicranella crispa
●
●
●
●
●
●
●
●
●
●
●
●
●
Dichodontium pellucidum
●
●
Dicranella heteromalla
●
●
●
●
●
●
●
●
●
DD
●
●
●
DD
DD
EN
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD DD
●
●
●
●
● EN EN
VU NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● VU VU
● ● ● ●
●
●
●
●
● VU
●
●
●
●
●
●
●
NT
●
●
CR
●
●
●
●
VU
DD
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
y
NT
DD
NT
●
●
●
●
●
●
●
●
Dicranella schreberiana
●
●
●
●
●
●
●
●
●
Dicranella staphylina
●
●
●
●
●
●
●
●
●
Dicranella subulata
●
●
●
●
●
●
●
●
●
●
Dicranella varia
●
●
●
●
●
●
●
●
●
●
y
●
●
CR
Dicranodontium denudatum
y
Dicranum acutifolium
y
Dicranum angustum
y
Dicranum brevifolium
y
Dicranum canariense
y
●
●
NT ●
●
Dicranum bonjeanii
●
●
●
Dicranoweisia cirrata
●
●
●
●
●
●
●
●
● VU ●
● ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
DD
DD ●
●
‐
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
Dicranum drummondii
●
Dicranum elongatum
●
●
●
●
Dicranum fulvum
●
Dicranum fuscescens
●
Dicranum laevidens
y
Dicranum leioneuron
y
Dicranum majus
Dicranum montanum
●
●
●
●
●
Dicranum flexicaule
y
●
●
DD
●
DD
DD
●
●
●
●
●
DD
●
●
●
●
CR
●
DD
CR
●
●
●
●
●
●
●
RE
●
RE
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Dicranum polysetum
●
Dicranum scoparium
●
●
RE
●
●
●
NT
●
●
VU ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
DD
● ●
●
●
●
RE
●
●
● ●
VU ●
●
●
y
Dicranum scottianum
●
●
y
Dicranum groenlandicum
●
●
●
DD ●
DD
●
Dicranum fragilifolium
●
●
●
Dicranum crassifolium
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● VU ●
●
●
●
●
●
●
DD
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
EN
‐
●
‐
●
●
EN
●
●
RE
●
DD
●
●
●
●
●
●
NT
●
VU
y
Dicranum spadiceum
NT ●
Dicranum spurium
●
Dicranum tauricum
●
Dicranum transsylvanicum
y
Dicranum undulatum
y
Dicranum viride
y
Didymodon acutus
●
●
NT ●
●
y
Didymodon australasiae
y
●
●
VU
●
●
●
●
●
EN
NT
EN
●
Didymodon asperifolius
●
●
●
●
●
● DD ●
●
● EN
●
●
●
VU RE
● EN EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
RE
●
●
VU
●
●
●
DD
●
●
VU
EN
●
●
●
●
●
●
●
●
VU
●
●
DD
●
Vatican City
Spain
Sicily
Sardinia
Portugal
Monaco
Malta
Madeira
San Marino
●
RE
●
●
75
●
‐
●
Dicranella rufescens
Dicranum septentrionale ●
DD
●
Dicranella howei
Dicranum muehlenbeckii
●
y
Dicranella grevilleana
Dicranum flagellare
●
‐
NT
●
●
EN
DD
VU
●
●
Dicranum dispersum
Italy
France
Cyprus
Corsica
Canary Islands
Balearic Islands
●
NT
Dichodontium flavescens
Dicranodontium uncinatum
●
VU
CR
● VU
y
Dicranodontium asperulum
●
●
●
Dialytrichia saxicola Dicranella humilis
?
●
VU ●
Dialytrichia mucronata
Dicranella campylophylla
CR
● VU
Daltonia stenophylla
Dendrocryphaea lamyana
●
‐
Cynodontium polycarpon
●
Azores
Andorra
Northern Ireland
EN NT
Cynodontium fallax
Cynodontium bruntonii
Ireland
Great Britain
Gibraltar
Channel Islands
Sweden
Svalbard
Norway
Iceland
y
Finland
y
Cynodontium asperifolium
Faeroe Islands
Taxon
Cyclodictyon laetevirens
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
VU
●
VU
CR
●
VU
●
EN
?
●
●
●
?
?
Moldova
●
●
SE Russia
Middle and South Urals
Lithuania
Latvia
●
Kazakhstan (in Europe)
●
Kaliningrad
Central Russia
Caucasus (in Europe)
Belarus
Arctic Russia
Turkey
Slovenia
Serbia
CR VU DD
Ukraine
●
Romania
Montenegro
Macedonia
Kosovo
Hungary
Greece
Croatia
●
●
●
●
?
●
●
R
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
EV
●
●
●
●
●
?
Sub‐polar & North Urals
DD EN
●
Estonia
I
●
Crimea
●
Crete
NT
Bulgaria
R
Bosnia‐Herzegovina
R
Albania
Switzerland
DD VU
NW Russia
?
R
Appendix 7 - Mosses
NE Russia
4
●
Slovakia
●
Poland
Germany
●
Netherlands
Czech Republic
●
Luxembourg
Belgium
3
Liechtenstein
Austria
Checklist and country status of European bryophytes
VU
4
●
●
●
●
VU
2
R
CR VU
● DD*
●
I
VU EN
●
●
●
●
EN
●
●
●
●
VU
●
●
EN
●
●
DD EN DD
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
DD
●
●
●
nm
VU
●
1
E
E
VU
NT
I
EN DD*
●
GE
●
●
DD
4
●
?
D
●
●
●
●
●
●
3
●
●
●
4
●
●
●
●
4
●
●
●
V
DD
2
●
●
●
●
● KW ●
VU
●
R
●
●
●
●
●
●
●
●
CR
●
●
2
●
VU
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
VU DD
EN EN
●
●
VU
●
VU EN
●
●
●
NT
3
●
3
●
2
DD
●
●
NT
CR
EN
●
DD* VU
●
●
●
●
●
?
●
DD
NT
●
●
CR
●
●
●
DD
●
●
VU
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
DD
●
●
NT VU
●
●
●
NT
● DD* VU
●
●
●
DD*
●
●
DD
●
●
CR
●
●
●
●
●
●
●
●
●
●
EN
●
●
CR
●
●
●
●
●
EN
●
DD
●
●
●
●
●
●
●
●
●
●
I
●
●
●
●
●
NT
DD*
●
EN CR
●
●
●
VU
●
●
●
VU
●
●
●
DD* VU
● DD* ●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
0
●
●
●
●
●
●
3
Mn
●
EN
2
●
●
●
●
V
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
DD
VU
VU EN
●
●
VU DD
●
●
●
●
VU
●
I
●
●
●
VU
●
●
●
DD
●
●
●
●
●
●
CR
DD
●
DD
DD*
●
●
●
●
●
●
●
●
●
●
3
●
●
●
DD*
●
●
●
●
●
●
●
●
VU
nm
V
●
CR
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
3
●
Mn
CR
●
1
●
●
R
VU KW ●
●
●
●
●
●
●
nm
CR
●
●
●
●
●
3
4
●
●
●
●
●
●
EV BE
●
●
●
●
●
●
VU
●
●
●
●
DD
●
●
●
EN DD
●
VU VU
●
DD
●
DD
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
NT
●
●
1
1
3
3
●
●
Mn NT
●
●
●
2
●
●
●
●
●
●
●
R
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
DD
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
?
DD*
●
●
DD
●
●
●
●
●
●
●
●
?
VU
●
●
DD EN
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
3
●
●
●
●
CR
E
EN NT
●
EN
●
NT
R
EN
●
●
EN
●
●
VU RE
●
●
●
●
●
I
●
●
●
●
V
‐
●
GE
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
VU
VU
●
●
EN
DD
● ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN DD
3
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
R
●
VU
CR
●
1
●
RE
●
●
●
●
●
●
●
EN
DD DD* ●
●
●
●
●
●
‐
●
DD
NT
●
DD DD
CR
●
●
● DD* ●
EN VU DD
●
DD
●
● DD* ●
●
●
●
●
●
●
●
●
●
●
EN
VU
●
●
●
●
●
●
●
●
VU
CC
EN
NT
●
●
DD*
●
●
●
DD
●
● DD* ●
●
●
●
●
●
EN KW V
●
●
I
●
●
●
●
●
●
●
●
NE
3
●
●
DD*
● DD*
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
?
RE
●
●
4
●
I
●
●
CR
●
●
●
●
VU
●
VU VU
●
VU EN VU
●
●
VU
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
NT
●
3
1
●
●
●
●
●
●
‐
●
●
‐
●
●
●
●
●
●
76
●
●
VU
●
●
●
DD
●
●
●
Didymodon fallax
●
●
●
●
Didymodon ferrugineus
●
● NT ●
●
●
●
●
●
●
●
●
●
●
●
●
● DD ●
●
Didymodon giganteus
●
●
●
●
●
●
DD
●
●
●
●
EN
VU
●
●
●
●
●
●
Didymodon glaucus
y
VU
CR
CR
●
Didymodon icmadophilus
y
● VU
DD
NT RE
●
●
●
●
●
DD
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
DD
●
●
●
●
●
●
●
NT
●
Didymodon insulanus
Didymodon johansenii
RE ●
●
y
●
●
●
●
●
●
●
●
●
●
●
●
RE ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Didymodon lamyanus ?
Didymodon luridus
●
Didymodon maschalogenus y
Didymodon maximus
y
VU
NT
Didymodon nicholsonii
Didymodon rigidulus
●
●
●
●
●
●
●
●
●
●
●
●
EN ●
●
●
●
VU
●
●
●
●
● DD
●
Didymodon sicculus
Didymodon sinuosus
●
Didymodon spadiceus
●
Didymodon subandreaeoides
y
Didymodon tomaculosus
y
Didymodon tophaceus
y
Didymodon validus y
Didymodon vinealis
Discelium nudum
y
Distichium capillaceum
Distichium hagenii
● VU ● DD ●
●
●
● NT ●
●
●
NT
●
●
Ditrichum cornubicum
y
●
Ditrichum flexicaule
●
Ditrichum gracile
Ditrichum heteromallum
●
●
Ditrichum lineare
y
Ditrichum punctulatum
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● NT NT
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
●
EN ●
●
●
●
●
●
●
●
●
●
●
NE ●
●
●
RE
●
●
●
Drepanocladus angustifolius ● ●
DD ●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● CR CR
●
●
●
●
●
●
●
●
●
●
●
●
●
VU ● NT ●
●
●
VU
●
● EN EN
● ● ● ●
●
●
●
●
Drepanocladus polygamus
y
●
●
NT NT NT
● ● ●
●
●
DD
●
Drepanocladus sendtneri
y
●
●
●
y
●
●
● NT NT
Drepanocladus sordidus
EN
EN
● ● VU
Drepanocladus trifarius y
Drepanocladus turgescens y
●
●
●
●
Echinodium renauldii
y
Echinodium setigerum
y
Echinodium spinosum
y
Encalypta affinis
y
●
●
●
● ●
CR ●
●
●
●
●
EN
‐
●
●
●
VU
●
NT
●
DD ●
‐
●
●
EN
●
●
●
●
●
●
●
● VU
VU
●
VU
DD
●
CR
●
●
●
●
●
VU
■
□
□
■
■
□
□
●
●
●
●
●
●
Encalypta alpina
NT ●
Encalypta brevicolla
●
y
●
●
●
EN
77
●
●
●
Encalypta affinis subsp. macounii
y
DD
●
●
Encalypta affinis subsp. affinis
y
●
●
● DD DD
VU ● EN
● ● ● ● ●
Encalypta obovatifolia
●
●
●
●
●
Encalypta pilifera ●
●
●
y
y
●
●
Drepanocladus lycopodioides Encalypta mutica
DD
●
●
y
●
●
●
y
y
●
RE
y
Encalypta microstoma
●
●
NT
Drepanocladus longifolius
Encalypta longicolla
●
●
Drepanocladus brevifolius y
●
●
y
Encalypta intermedia
●
DD
●
Drepanocladus arcticus
Encalypta brevipes
●
●
DD
●
y
Encalypta ciliata
●
●
●
EN CR
● DD DD ●
●
●
●
●
●
Drepanocladus aduncus
●
● DD
●
Ditrichum subulatum
Ditrichum zonatum
●
● EN
●
Ditrichum pusillum
●
●
●
●
VU ●
●
●
Ditrichum pallidum
Ditrichum plumbicola
●
RE
EN ● VU
NT ● ● ● ●
●
y
●
●
●
y
Distichophyllum carinatum
●
● VU
●
Distichium inclinatum
●
●
● VU
●
●
Diphyscium foliosum
● DD
DD* ● ●
●
●
Didymodon umbrosus
●
●
CR ●
● ●
■
●
●
●
●
●
●
●
●
● DD
●
● VU
RE
● CR CR ●
●
●
DD NT
●
●
CR
● VU
VU
EN
●
● NT
●
●
●
●
●
VU
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
DD
●
EN
●
Monaco
Malta
Madeira
Italy
France
Cyprus
Corsica
Canary Islands
Balearic Islands
Azores
Andorra
Northern Ireland
Ireland
Great Britain
Gibraltar
y
Channel Islands
Didymodon erosus
Sweden
y
Svalbard
y
Didymodon eckeliae Norway
Didymodon cordatus
Iceland
y
Finland
y
Didymodon brachyphyllus
Faeroe Islands
Taxon
Didymodon bistratosus
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
NE Russia
NW Russia
SE Russia
Sub‐polar & North Urals
Ukraine
●
Moldova
Middle and South Urals
Lithuania
Latvia
Kazakhstan (in Europe)
Kaliningrad
D
Estonia
VU
●
Crimea
3
Central Russia
●
●
Caucasus (in Europe)
NT
●
●
Belarus
●
●
VU
EN
Arctic Russia
●
●
●
●
Turkey
●
●
Slovenia
?
●
Serbia
●
●
Romania
●
‐
Montenegro
●
●
VU
Macedonia
●
●
●
Kosovo
●
●
Hungary
●
●
Greece
●
VU
Croatia
●
●
Crete
●
●
Bulgaria
●
●
EN VU
Appendix 7 - Mosses
Bosnia‐Herzegovina
●
●
●
Albania
●
●
●
Switzerland
●
CR
Slovakia
‐
Poland
Germany
VU
Netherlands
Czech Republic
●
Luxembourg
Belgium
3
Liechtenstein
Austria
Checklist and country status of European bryophytes
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
VU
●
●
●
●
●
DD
●
DD
●
‐
●
CR VU
D
●
EN
●
●
●
●
EN
3
●
NT
●
●
●
VU
●
●
●
RE VU
CR VU
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
NT
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD* ●
CR
●
●
VU
●
VU
●
●
●
DD
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
4
●
●
●
●
2
●
VU
●
●
●
NT
●
4
●
●
NE
EN
D
D
V
V
3
●
●
●
EN
●
●
VU
●
●
●
●
●
●
EN
1
●
●
●
●
●
●
●
nm
●
●
●
V
●
●
Mn VU
●
●
DD VU
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
EN
●
DD
●
●
EN
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
DD*
●
VU
●
●
●
NT NT
●
●
3
●
●
●
●
●
DD*
●
●
●
●
EB
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
GE
●
V
●
●
NT
●
●
VU
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
4
●
●
●
VU
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
V
GE
●
●
● DD*
NT CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
RE
●
●
●
●
●
●
NT
●
RE
●
NT KW
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
NT
●
NT KW ●
●
CR
●
●
●
●
●
VU VU
DD EN
●
VU
EN
VU KW ●
NT EN
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
DD* NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
VU
●
●
●
●
●
●
●
●
R
●
3
●
●
●
3
●
CR
●
1
●
●
●
●
●
●
●
D
●
4
●
●
●
●
●
NE
●
●
EN
●
R
V
●
●
●
●
●
●
DD NT DD*
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
DD
1
Mn RE
● VU
1
3
EB
CR ●
E
●
CR VU
EN EN DD*
CR
RE
1
D
EB
R
V
EN CR
DD
3
2
CR
2
1
●
E CR NT
Ex
CR
4
RE
R
●
●
●
●
●
□
□
nm
□
□
CR
□
□
●
●
●
2
2
1
●
●
E
●
●
NE
DD*
EN
NT
VU
●
VU
VU
●
NT VU VU
EN
DD
●
●
EN ● VU
VU VU VU VU
● CR
VU
●
DD CR DD
NT
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
DD*
●
●
●
●
●
●
●
●
2
●
1
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
1
2
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
●
VU
●
●
DD
●
NT
●
DD
●
●
●
●
●
●
0
●
Mn VU
2
●
4
3
●
●
R
CR
●
EN
●
I
DD
●
●
●
●
●
●
VU
●
VU
●
VU
●
●
●
●
●
●
●
●
3
●
EN
1
●
●
●
●
●
DD
CR
●
EN
VU
?
●
● DD*
●
●
EN
●
●
●
●
VU
●
D
●
●
●
●
●
●
78
□
■
Encalypta streptocarpa
●
●
●
●
Entodon challengeri
y
Entodon cladorrhizans
y
●
Entodon concinnus
Entodon schleicheri
y
Entosthodon abramovae y
Entosthodon attenuatus
Entosthodon commutatus □
y
Encalypta vulgaris
●
●
●
●
□
□
□
■
EN
EN
□
●
●
●
●
● NT
●
●
●
●
●
●
●
●
●
●
?
● VU ●
●
●
●
●
●
●
●
●
●
●
y
Entosthodon handelii y
Entosthodon hungaricus
y
DD
Entosthodon kroonkurk y
●
Entosthodon mouretii
y
Entosthodon muhlenbergii
y
●
RE
NT ●
CR
●
●
●
NT
●
● NT NT
●
● RE
VU ●
y
?
● ● ●
NT EN
Entosthodon stenophyllus y
Ephemerum cohaerens
y
Ephemerum crassinervium y
RE
DD
●
●
■
■
y
■
■
■
■
■
●
●
●
●
DD
●
●
●
EN VU
● NT NT
Ephemerum crassinervium subsp. rutheanum ●
Ephemerum minutissimum
y
RE
●
●
● VU
y
●
Eucladium verticillatum var. verticillatum
□
Eurhynchiastrum pulchellum
●
VU
●
●
□
●
●
●
●
●
■
□
●
□
Eurhynchium angustirete
●
●
□
□
□
●
●
VU ●
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
?
DD
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
NT
●
DD*
●
●
NT
●
●
●
●
●
DD
●
●
DD
●
●
●
●
●
●
●
●
DD
●
VU
●
●
●
●
●
●
●
DD
●
●
■
■
●
●
●
●
●
●
■
■
■
■
●
VU
●
●
DD
DD
●
●
■
■
■
■
■
DD
●
●
DD
●
●
●
●
●
●
●
●
●
●
DD
●
●
CR
●
●
●
●
●
●
●
●
●
■
●
■
●
●
●
●
■
●
●
●
■
□
□
□
□
■
□
□
□
□
□
□
■
●
●
●
■
■
■
■
DD
■
■
●
●
●
●
■
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
■
□
●
●
●
●
DD
■
●
DD
DD
□
■
■
y
Fabronia pusilla
y
Fissidens adianthoides
●
●
●
●
●
●
●
●
y
Fissidens arnoldii
y
Fissidens asplenioides
y
Fissidens azoricus
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD ●
□
□
Fissidens crassipes
Fissidens crassipes subsp. crassipes
●
DD
●
NT
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
□
■
■
■
■
■
■
DD
●
●
●
●
●
●
●
●
●
?
●
●
■
■
■
■
■
■
■
■
■
‐
■
■
□
■
■
□
■
■
■
■
●
●
●
●
■
●
VU
DD
●
EN
●
●
●
?
‐ NT ●
●
●
■
□
□
□
?
‐
■
■
?
●
‐
●
■
●
■
●
●
DD
●
●
●
●
●
●
y
● DD
EN RE
● ● ●
NT
NT
●
●
●
●
●
NT
●
● VU
●
RE ● VU
●
●
NT
●
●
● ● ● ‐
● VU VU
●
●
□
■
●
●
●
●
● ●
DD
●
●
●
‐
●
●
●
●
□
●
●
□
●
y
●
●
● DD*
CR
Fissidens crispus
y
●
DD ●
●
y
y
●
●
●
Fissidens celticus
Fissidens grandifrons
●
□
Fissidens bryoides var. caespitans
Fissidens gymnandrus
●
●
●
●
y
●
●
□
Fissidens gracilifolius
● DD ●
‐ DD
EN
Fissidens bryoides
Fissidens exilis
●
●
Fissidens bryoides var. bryoides
Fissidens dubius
?
●
●
●
Fissidens arcticus
■
VU
VU
●
Fabronia ciliaris
CR
●
DD
●
□
●
●
●
●
RE ●
●
●
●
●
●
■
●
●
●
●
●
●
■
■
□
■
■
●
●
NT
●
■
●
●
●
● DD
DD
●
●
●
●
●
●
● DD*
‐
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
DD
●
●
Vatican City
Sardinia
San Marino
Monaco
Portugal
NT
●
●
●
●
●
DD
●
●
●
DD
DD
●
●
●
●
VU
●
■
●
Malta
Italy
●
●
□
●
●
■
●
Madeira
France
●
●
●
●
●
Exsertotheca intermedia
79
■
y
Exsertotheca crispa
Fissidens fontanus
■
●
●
●
■
Eurhynchiastrum pulchellum var. pulchellum
Fissidens curvatus
■
●
●
■
Eurhynchiastrum pulchellum var. praecox
Fissidens crassipes subsp. warnstorfii
■
●
●
EN
Eurhynchiastrum pulchellum var. diversifolium
Fissidens coacervatus
●
●
●
Eucladium verticillatum var. angustifolium
Exsertotheca baetica
●
DD
DD
● EN EN
Epipterygium tozeri
Eurhynchium striatum
●
■
●
●
VU ?
Entosthodon schimperi
Eucladium verticillatum
●
DD
●
● DD ●
DD ●
y
Entosthodon fascicularis
Ephemerum spinulosum
●
●
Entosthodon duriaei Ephemerum serratum
●
●
y
Ephemerum recurvifolium
●
●
■
NT
Entosthodon dagestanicus Ephemerum crassinervium subsp. sessile ●
●
Entosthodon convexus
Entosthodon pulchellus
Cyprus
□
●
VU
●
□
y
Entosthodon obtusus
Corsica
Canary Islands
Balearic Islands
Azores
Andorra
Northern Ireland
● CR CR ●
■
Spain
DD
Ireland
●
●
Great Britain
●
●
Gibraltar
●
●
Sicily
Encalypta rhaptocarpa var. rhaptocarpa
Encalypta spathulata
NT ●
● ●
Channel Islands
■
Sweden
Encalypta rhaptocarpa var. leptodon
Svalbard
●
Norway
●
y
Iceland
Encalypta rhaptocarpa
Taxon
Encalypta procera
Finland
Faeroe Islands
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
□
□
□
■
□
■
EN DD
□
■
?
●
□
■
■
■
■
■
●
‐
●
□
CR
●
●
□
Ukraine
□
●
●
Sub‐polar & North Urals
●
●
●
SE Russia
■
NW Russia
■
●
●
NE Russia
■
■
Moldova
●
Middle and South Urals
●
Lithuania
●
Latvia
●
Kazakhstan (in Europe)
Estonia
●
●
Kaliningrad
Crimea
●
Belarus
●
Arctic Russia
●
Turkey
●
Romania
●
Montenegro
?
Macedonia
●
Kosovo
Hungary
Bulgaria
Greece
Bosnia‐Herzegovina
●
Croatia
Albania
●
Crete
Switzerland
●
Central Russia
1
●
Caucasus (in Europe)
R
DD
NT
Slovenia
■
2
●
Appendix 7 - Mosses
Serbia
■
Slovakia
DD G
Poland
■
Netherlands
●
Luxembourg
Liechtenstein
EN nb
Belgium
●
Austria
Germany
Czech Republic
Checklist and country status of European bryophytes
●
●
●
■
■
■
■
■
■
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
V
●
●
BE
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
RE
●
3
●
●
●
DD
V
●
●
BE
●
NT
EN
●
●
CR VU
VU
●
●
CR
●
●
NT
●
●
DD
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
?
●
●
●
●
●
●
●
●
●
3
●
VU
2
3
●
VU BE
R
R
NT EN
●
●
●
CR
0
Mn CR
2
1
2
EB
●
I
1
Mn
● EN
2
●
●
●
E
●
R
■
■
●
●
R
VU CR DD
●
●
CR
CR DD* ●
NT
●
NT
NT
●
VU
●
●
●
●
●
●
●
●
●
NT
●
●
DD
DD
VU
■
■
■
■
NT
●
●
●
EN
●
DD
●
●
●
●
●
●
●
●
●
VU
●
EN
●
●
●
●
●
● DD*
●
●
●
●
●
●
●
●
●
●
●
●
0
0
●
●
DD
VU CR
‐
2
■
■
3
●
●
●
0
Mn
●
3
●
●
DD
V
●
‐
●
●
DD*
NT KW E
VU CR DD*
CR
●
●
VU
●
R
NT VU DD*
DD
●
●
●
GE
●
□
●
CR
DD
●
■
DD
VU CR
DD
VU EN VU
3
●
3
●
●
●
●
●
●
VU
●
●
●
R
VU
●
■
●
●
V
●
■
□
□
nm
□
□
●
●
●
3
●
VU
●
■
‐
■
□
■
3
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
●
V
●
EN
D
□
□
□
□
□
●
●
●
●
■
■
□
□
□
GE
●
●
●
●
●
●
●
●
●
EB
●
●
●
● KW ●
●
●
●
●
●
NT
●
□
□
■
□
□
□
■
NT
●
●
●
●
●
■
■
■
■
●
□
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
VE
●
3
4
□
■
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
NT
● DD* ●
DD*
●
●
●
●
VU
VU
VU
■
EN
●
CR
DD CR
VU
□
□
□
■
□
●
□
●
□
□
●
●
?
●
□
●
3
□
■
●
RE
●
■
●
●
□
●
■
●
●
●
●
■
□
Ex CR
●
□
□
●
●
●
●
□
●
DD
NT
□
■
RE VU DD*
●
●
●
■
■
●
●
■
□
■
●
●
●
R
●
●
□
3
●
●
●
●
3
■
●
●
■
?
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
VU
●
●
2
1
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
□
□
■
□
□
□
□
●
D
3
●
DD nb
□
□
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
■
EN
□
□
□
□
□
■
■
DD
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
VU
■
●
●
E
EN
●
●
VU
●
●
●
●
●
EN
●
●
●
1
●
●
□
□
□
□
□
□
□
□
□
■
■
□
□
□
□
□
□
□
□
□
● DD* ●
■
●
■
●
●
EN
EN
DD*
●
●
●
DD
●
DD
●
●
●
●
●
NT
●
●
●
VU
●
●
●
●
●
●
●
DD
3
DD
?
●
NT
DD DD
CR
●
3
●
●
●
●
●
V
●
● KW ●
●
●
●
4
●
NT
●
VU
●
VU
●
●
●
●
●
●
●
1
●
●
G
DD*
●
●
NT NT
●
●
●
VU
●
●
●
●
●
DD NT
CR
●
VU
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
DD
VU
●
●
NT
VU
●
●
●
3
4
●
●
●
●
●
●
1
3
●
●
●
●
●
●
●
●
●
EN
●
I
●
●
●
●
DD*
●
●
●
80
Fissidens osmundoides
y
Fissidens polyphyllus
y
Fissidens pusillus
y
Fissidens rufulus
y
Fissidens serratus
y
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
● VU
● VU
●
●
●
RE
●
●
●
●
●
●
DD
●
● VU
●
●
●
●
●
●
●
●
●
●
● DD ●
●
●
●
●
●
●
●
●
EN
●
●
y
●
●
●
●
●
□
●
□
■
●
□
●
●
■
□
□
Fissidens viridulus var. incurvus
●
●
●
●
■
■
■
■
●
●
●
■
●
●
●
■
■
■
●
●
DD
●
●
●
●
DD
●
●
●
●
■
■
?
■
□
●
□
●
■
●
□
●
□
●
■
●
■
●
■
●
□
●
■
●
■
●
■
●
■
●
■
■
■
■
■
□
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
□
■
■
■
■
■
■
■
■
■
■
■
■
■
●
●
●
●
■
■
■
●
●
●
●
●
●
●
● DD ●
●
●
●
Fontinalis antipyretica subsp. antipyretica
□
□
■
□
■
■
■
■
■
■
□
□
□
■
■
■
■
■
□
■
Fontinalis antipyretica subsp. gracilis
■
Fontinalis antipyretica subsp. kindbergii
■
Fontinalis dalecarlica
●
Fontinalis dichelymoides
y
Fontinalis hypnoides
y
■ NT NT
■
●
●
Fontinalis hypnoides var. hypnoides
●
□
●
□
●
■
DD
●
●
●
●
●
●
●
●
●
■
■
■
■
DD
■
NT
■
●
■
●
■
●
●
■
●
●
‐
●
□
□
□
●
●
●
●
■
●
□
■
y
Funaria arctica
y
Funaria hygrometrica
●
■
■
●
●
●
■
■
■
■
□
□
●
●
●
●
●
●
●
●
●
●
●
Funariella curviseta
Glyphomitrium daviesii
y
Goniomitrium seroi
y
■
●
●
y
■
● DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
y
Gigaspermum mouretii
●
● NT
●
●
●
●
●
DD NT
EN
CR ●
●
●
● VU
NT
CR
● DD
VU
●
●
●
VU
NT EN
●
●
●
Grimmia anomala
Grimmia arenaria
y
EN
DD
Grimmia atrata
y
DD*
●
Grimmia caespiticia
y
Grimmia capillata
y
Grimmia crinita
y
Grimmia curviseta
y
DD
●
●
?
NT ●
●
●
●
DD
●
VU ●
●
Grimmia elatior
Grimmia elongata
NT ●
Grimmia funalis
●
y
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
CR
VU
●
●
●
DD
●
●
VU
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
VU
NT ●
Grimmia dissimulata
Grimmia donniana
?
●
● DD
●
●
● NT
●
●
●
●
●
●
●
●
VU
●
●
VU
●
●
●
●
●
●
VU
●
●
● VU
● NT NT
●
●
●
●
● NT NT ●
●
●
●
DD ●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
●
●
DD
?
●
●
●
y
●
●
●
NT
●
Grimmia incurva
●
●
●
●
VU
●
?
NT ●
●
● RE RE ●
?
●
●
●
●
●
● ●
● RE RE ●
●
?
●
y
●
●
●
●
●
y
Grimmia mollis
EN
●
●
Grimmia montana
NT ●
●
●
●
●
●
●
●
●
DD RE
●
EN
Grimmia anodon
Grimmia muehlenbeckii
●
●
VU
Grimmia longirostris
●
●
●
Grimmia laevigata
●
●
DD ●
Grimmia hartmanii
●
●
Grimmia alpestris
Grimmia decipiens
‐
■
■
Funaria anomala ■
●
■
y
■
■
Fontinalis squamosa var. dixonii
Funaria aequidens
■
DD
Fontinalis squamosa var. curnowii
Fontinalis squamosa var. squamosa
■
●
Fontinalis hypnoides var. duriaei
Fontinalis squamosa
■
■
●
NT
●
■
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
● ●
DD ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT DD DD ●
y
● EN
●
DD
VU
●
●
● VU ●
●
●
NT
●
●
VU
●
●
●
●
DD
●
●
●
Vatican City
Spain
Sicily
Sardinia
Portugal
Monaco
San Marino
●
●
●
●
81
DD
●
■
●
●
Grimmia orbicularis
●
■
●
●
Grimmia nutans
●
■
●
●
Grimmia meridionalis DD
■
●
●
Grimmia lisae
●
●
●
■
●
Fontinalis antipyretica
Grimmia horrida ●
●
●
□
Grimmia fuscolutea
●
●
●
●
Fissidens viridulus var. viridulus
Fontinalis antipyretica subsp. bryhnii
●
●
●
●
●
Fissidens taxifolius subsp. pallidicaulis
Funaria microstoma
DD
●
●
●
Fissidens taxifolius subsp. taxifolius
DD
●
●
Fissidens sublimbatus
Fissidens viridulus
Malta
?
●
● EN EN
VU VU
Fissidens taxifolius
Madeira
Italy
France
Cyprus
Corsica
●
●
Fissidens serrulatus
Fissidens sublineaefolius
Canary Islands
Balearic Islands
Azores
Andorra
Northern Ireland
Ireland
Great Britain
Gibraltar
NT NT NT
●
Fissidens rivularis
Channel Islands
DD
●
●
Fissidens ovatifolius
Sweden
y
Svalbard
y
Fissidens nobreganus
Norway
Fissidens monguillonii
Iceland
y
Finland
y
Fissidens microstictus
Faeroe Islands
Taxon
Fissidens jansenii
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
●
●
●
Ukraine
●
Sub‐polar & North Urals
●
●
SE Russia
●
NW Russia
●
NE Russia
●
Moldova
●
Middle and South Urals
●
Lithuania
Latvia
●
Kazakhstan (in Europe)
Central Russia
●
Kaliningrad
Caucasus (in Europe)
●
Estonia
Belarus
●
Crimea
Arctic Russia
Turkey
Slovenia
Serbia
●
Romania
●
Montenegro
●
Macedonia
●
●
Kosovo
Greece
●
Hungary
Croatia
●
Appendix 7 - Mosses
Crete
NT
Bulgaria
R
Bosnia‐Herzegovina
EB
Albania
Luxembourg
Liechtenstein
Switzerland
●
Slovakia
Mn
Poland
●
R
Netherlands
●
Germany
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
●
EN
2
VU
VU
●
●
●
●
●
Mn
3
●
■
NE
●
●
□
●
3
●
●
R
NT
●
■
●
■
●
●
●
GE
●
●
●
EN
●
□
●
DD
●
VU
●
●
●
●
VU
●
DD* VU EN
●
●
□
●
□
●
□
●
●
□
●
●
■
●
■
■
■
■
■
DD DD
4
□
■
■
■
■
■
VU DD
●
●
●
●
●
●
●
●
●
□
□
□
■
□
□
□
■
□
D
●
□
●
●
●
DD
□
□
●
●
●
■
●
■
●
■
●
●
■
■
●
●
●
■
■
●
■
□
●
□
●
●
■
?
● DD* ●
●
■
■
●
DD
■
●
●
●
●
●
■
●
■
●
●
●
●
CR
●
?
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
●
■
●
■
●
□
□
□
□
●
□
□
□
□
●
□
●
□
●
□
●
□
●
□
□
●
□
□
●
●
●
●
R
■
■
NT
■
EN
■
■
■
DD
■
■
■
■
NT
■
■
■
■
EN
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
■
□
□
□
□
□
□
□
■
■
■
■
■
□
■
□
□
■
■
■
■
■
■
■
■
■
■
□
■
■
□
□
■
VU
■
■
■
■
■
■
■
DD
E
EN
?
●
D
1
●
CR
■
□
●
●
G
E
●
CR VU
●
●
●
■
■
■
■
□
●
□
?
□
■
0
3
●
■
●
2
V
VU
□
R
□
□
□
nm
□
□
□
□
CR VU
□
□
●
●
DD
●
VU CR
●
●
●
●
VU
3
1
●
●
●
●
●
VU
3
2
●
●
●
●
?
□
□
EN
NT
RE
□
□
■
■
■
EN
□
□
●
■
●
■
●
■
●
■
●
□
□
■
■
■
■
●
●
●
?
●
●
●
●
●
■
●
■
■
●
●
2
●
●
●
nb
●
●
●
●
I
●
●
CR
●
●
●
nb
●
VU
●
●
●
EN
VU
3
R
●
●
GE
●
1
●
NT
R
I
NT ●
DD NT
VU nb
●
●
DD
R
0
●
EN
3
EN GE
R
1
Mn
?
R
●
●
●
●
V
VU
DD D
●
V
VU
●
CR
G
●
NT
R
●
2
●
●
nb
●
●
3
●
Mn
●
3
●
●
Mn
●
D
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU CR VU
VU
●
●
●
DD*
●
●
●
●
DD
●
●
●
EN GE
●
VU
VU
VU
●
●
●
EN DD
VU
VU DD*
VU
DD
NT ●
DD
●
●
4
●
●
●
●
●
●
●
●
V
VU
3
●
●
G
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
● DD* CR
●
●
● VU
● DD* CR
●
●
DD VU
DD
VU
VU
●
DD VU
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
EN
●
●
●
DD*
EN
●
●
●
●
●
VU
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
RE
EN
●
●
●
●
DD
●
VU
●
DD
●
●
●
●
●
●
●
●
CR
●
●
NT
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
VU
EN
●
●
●
●
●
CR
●
●
●
VU
●
VU
●
●
●
●
●
●
●
●
●
●
VU
VU CR VU
EN
NT
●
DD DD*
CR ● DD*
●
CR
VU
●
●
●
●
●
●
●
GE
●
EN
●
D
●
●
EN
●
●
●
●
●
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
CR
●
●
●
NT
●
●
● DD* VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
●
●
●
●
●
R
VU
●
●
●
●
●
●
●
●
●
●
●
82
Grimmia reflexidens
y
Grimmia teretinervis
y
●
●
●
●
VU ●
●
●
●
●
●
●
●
y
Grimmia unicolor
y
●
●
●
●
●
●
Corsica
Cyprus
France
Italy
Madeira
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● ●
DD ●
NT
● NT ●
●
●
●
●
Gymnostomum aeruginosum var. aeruginosum
□
□
□
■
□
EN
●
■
□
VU
● ●
●
●
●
●
□
□
■
●
●
●
●
●
●
●
●
●
●
●
●
●
Monaco
●
Malta
NT
DD
●
●
● DD ●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
■
●
●
●
●
●
●
DD
●
●
●
●
●
DD
Gymnostomum aeruginosum
●
●
●
● NT NT ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
DD
●
■
□
■
□
y
CR
CR
Gymnostomum calcareum
●
●
●
●
●
DD ●
□
□
●
●
●
●
■
■
Gymnostomum viridulum
●
y
Gyroweisia tenuis
□
□
●
●
□
●
RE ● NT
VU
●
VU
Hageniella micans
y
Hamatocaulis lapponicus
y
Hamatocaulis vernicosus
y
Haplocladium angustifolium
y
Haplocladium microphyllum
y
Haplocladium virginianum
y
Hedenasiastrum percurrens y
●
■
EN
●
●
●
EN
EN
● VU
VU ● NT
DD
DD
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● NT
●
● NT NT
●
□
●
□
●
■
●
●
●
NT DD
●
EN
●
RE
●
●
●
●
●
●
● VU
●
●
●
Hedwigia ciliata var. ciliata
□
□
■
□
■
■
■
□
□
■
■
Hedwigia ciliata var. leucophaea
■
●
Helicodontium capillare
y
Helodium blandowii
y
●
Hennediella heimii
y
●
DD ●
●
●
●
●
■
■
●
●
●
●
●
●
●
●
●
■
■
□
■
■
■
●
●
●
●
●
●
NT
●
■
□
■
■
■
DD
■
DD
●
NT
●
DD
●
●
●
●
●
●
●
●
●
Hennediella heimii var. arctica
●
●
RE
●
●
●
●
●
DD
RE
□
□
□
□
□
□
□
●
●
●
DD
●
EN
●
■
Hennediella heimii var. heimii
□
□
□
□
□
□
■
Hennediella macrophylla
●
Hennediella stanfordensis
●
Herzogiella seligeri
Herzogiella striatella
y
Herzogiella turfacea
y
●
●
●
●
●
●
VU
Heterocladium dimorphum
●
●
●
●
NT
●
●
●
Heterocladium flaccidum
●
Heterocladium wulfsbergii
y
Heterophyllium affine
y
Hilpertia velenovskyi
y
●
●
VU
●
●
●
●
●
●
●
●
●
?
NT
DD ●
Heterocladium heteropterum
●
●
●
●
● ● ●
● NT NT
●
●
?
Homalia lusitanica
Homalia trichomanoides
●
●
●
●
●
●
●
●
y
□
●
●
●
●
●
●
●
●
●
●
● DD ●
□
□
■
□
□
□
□
□
□
□
●
●
Hookeria lucens
●
●
●
Hygroamblystegium fluviatile
●
● ● NT
NT
EN
●
●
●
Hygroamblystegium humile
●
●
Hygroamblystegium tenax
●
EN ● EN
●
Hygroamblystegium varium
●
●
●
●
●
●
●
●
●
●
●
●
●
Hygrohypnum alpinum
y
VU ●
●
Hygrohypnum cochlearifolium
y
VU
●
●
●
●
●
●
●
●
●
VU
NT ●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
RE
●
●
●
DD
●
●
●
●
■
●
■
●
●
DD
●
●
■
●
■
□
■
■
■
●
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Homomallium incurvatum
●
●
●
■
■
●
●
●
Homalothecium sericeum
Hygrohypnum alpestre
●
●
DD
NT
Homalothecium philippeanum
●
●
●
NT ●
Homalothecium lutescens var. fallax
Homalothecium lutescens var. lutescens
●
● VU ●
●
●
●
●
VU
●
Homalothecium lutescens
●
●
●
Homalothecium aureum
83
●
■
●
●
Hygrohypnum duriusculum
●
DD
□
●
Hedwigia ciliata
Hedwigia stellata
●
DD
□
● VU ●
●
■
●
□
●
■
□
□
NT
●
□
■
●
●
●
■
□
RE
Habrodon perpusillus
Homalia webbiana
●
■
Gymnostomum lanceolatum
Gyroweisia reflexa
■
■
VU
Gymnostomum calcareum var. atlanticum Gymnostomum calcareum var. calcareum
●
●
Gymnostomum aeruginosum var. obscurum
Gymnostomum boreale
●
●
●
●
DD
■
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD*
●
●
● NT NT
● EN EN
●
?
●
● NT NT
●
DD
●
●
●
DD
●
● NT NT
●
● DD ●
●
●
●
●
●
?
DD
●
●
DD
●
●
RE
●
●
●
● VU
●
NT CR
●
●
●
NT
●
●
●
DD
NT
●
Vatican City
Canary Islands
Azores
Andorra
Northern Ireland
Ireland
Great Britain
●
●
NT
●
●
● ● ● ●
● NT NT ●
RE
VU
y
●
Spain
●
●
●
Grimmia ungeri
●
●
Grimmia torquata
Grimmia triformis
●
EN
Grimmia tergestina
Grimmia trichophylla
Gibraltar
●
Balearic Islands
●
● EN
●
●
Grimmia ramondii
Channel Islands
Norway
●
Sweden
Iceland
●
Svalbard
Finland
●
Sicily
y
Grimmia pulvinata
●
Sardinia
Grimmia poecilostoma
●
San Marino
y
Portugal
Grimmia plagiopodia
Faeroe Islands
Taxon
Grimmia ovalis
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
Mn
●
●
VU
R
●
3
●
CR G
● nb
●
Mn VU
●
●
RE
GE
2
VU
●
●
●
●
RE
●
R
V
●
VU
■
■
□
■
nm
□
●
●
DD
●
●
●
●
EN
●
NT
DD*
●
●
●
●
● KW ●
●
●
●
●
●
●
VU
●
VU
VU
● DD*
VU
●
●
●
●
●
●
DD NT
●
●
●
●
VU
●
DD
●
●
●
●
EN
●
●
● DD* EN
●
●
●
●
●
EN
●
●
DD* EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
NT
●
●
EN
●
●
●
●
●
NT
●
●
●
●
VU
●
●
●
●
NT
●
●
CR
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
□
■
□
□
□
□
□
□
●
●
●
CR
VU
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
●
□
●
□
□
□
□
□
□
□
VU
●
●
VU
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
Ukraine
●
Sub‐polar & North Urals
●
NE Russia
●
Moldova
●
Middle and South Urals
NE
Lithuania
●
Latvia
●
Kazakhstan (in Europe)
Estonia
●
●
DD
Kaliningrad
Crimea
●
●
●
●
Caucasus (in Europe)
●
Belarus
●
SE Russia
●
3
●
●
CR
●
●
●
●
●
●
NW Russia
●
●
Central Russia
4
●
EN
Arctic Russia
●
Turkey
●
Slovenia
●
Serbia
Greece
●
Romania
Croatia
●
Montenegro
Crete
●
Macedonia
Bulgaria
●
Kosovo
Bosnia‐Herzegovina
●
VU
Hungary
Albania
●
Switzerland
NT GE
Appendix 7 - Mosses
Slovakia
●
2
Poland
3
Netherlands
●
RE
Luxembourg
Liechtenstein
●
●
Germany
Belgium
●
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Austria
Checklist and country status of European bryophytes
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●
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84
Hygrohypnum montanum
y
Hygrohypnum norvegicum
y
Hygrohypnum polare
y
Hygrohypnum smithii
y
Hygrohypnum styriacum
y
Hygrohypnum subeugyrium
y
●
●
●
●
●
●
●
●
●
Hymenoloma compactum
●
Hymenoloma crispulum
y
Hymenostylium gracillimum y
VU
●
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EN
●
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NT
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●
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● NT NT ●
● ● ● ●
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● DD
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y
Hyocomium armoricum
●
Hyophila involuta
y
Hypnum aemulans y
Hypnum andoi
●
●
Hypnum bambergeri
Hypnum callichroum
VU
●
●
NT ●
●
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NT
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●
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●
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Hypnum cupressiforme
●
●
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●
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■
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■
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■
●
●
●
●
Hypnum cupressiforme var. filiforme
Hypnum cupressiforme var. heseleri
■
Hypnum cupressiforme var. lacunosum
Hypnum cupressiforme var. resupinatum
■
■
■
■
Hypnum cupressiforme var. subjulaceum
Hypnum fertile
■
Hypnum hamulosum
y
● VU ●
y
Hypnum imponens
y
NT
●
●
●
Hypnum jutlandicum
●
●
●
Hypnum pallescens
Hypnum plicatulum
y
Hypnum procerrimum
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Hypnum revolutum
●
Hypnum revolutum var. revolutum
□
●
y
y
Indusiella thianschanica
y
Isopterygiopsis alpicola
y
●
●
●
Isopterygiopsis pulchella
y
Isothecium algarvicum
y
Isothecium alopecuroides
●
●
●
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●
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●
●
●
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●
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■
Isothecium myosuroides var. brevinerve
●
■
■
Isothecium myosuroides var. myosuroides
□
□
■
□
□
■
■
□
y
Iwatsukiella leucotricha
y
Jaffueliobryum latifolium
y
■
□
■
■
●
Kiaeria blyttii
●
●
●
●
●
●
Kiaeria falcata
● EN ●
●
●
●
● CR
Kiaeria glacialis
●
●
●
●
●
85
●
●
●
VU
Isothecium myosuroides var. brachythecioides
Isothecium prolixum ●
EN
■
●
DD
Isothecium holtii
Isothecium myosuroides
?
●
Isopterygiopsis muelleriana
Isopterygium tenerum
DD
●
NT
●
y
●
●
y
Imbribryum mildeanum
●
RE
VU
●
Imbribryum alpinum
●
●
EN
Hypnum vaucheri
Hypopterygium tamarisci
□
y
Hypnum subimponens
●
VU
●
Hypnum revolutum var. dolomiticum
Hypnum uncinulatum
●
VU
Hypnum recurvatum
Hypnum sauteri
●
y
Hypnum holmenii
VU
●
●
●
●
●
‐
□
□
□
□
■
●
●
●
VU
EN
●
●
?
‐
Vatican City
Spain
●
Sicily
●
Sardinia
●
San Marino
●
Portugal
●
●
Monaco
●
●
Malta
●
Madeira
Cyprus
●
Corsica
●
Canary Islands
Balearic Islands
NT
Azores
VU
Andorra
Northern Ireland
●
●
NT ●
Hymenostylium recurvirostrum var. recurvirostrum
Hymenostylium xerophilum ●
●
●
Hymenostylium recurvirostrum
Hymenostylium recurvirostrum var. insigne
●
VU
Hylocomiastrum umbratum
Hymenoloma mulahaceni
●
●
● EN
Hylocomiastrum pyrenaicum
Hylocomium splendens
●
●
VU
VU
VU
● ● ● ● ● ●
Hygrohypnum ochraceum
Ireland
CR
Great Britain
● NT ●
●
●
Italy
y
NT
●
France
Hygrohypnum molle
●
Gibraltar
●
Channel Islands
Norway
●
●
Sweden
Iceland
●
●
●
Hygrohypnum luridum
Svalbard
Finland
Faeroe Islands
Taxon
Hygrohypnum eugyrium
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
●
●
RE
●
R
4
●
●
●
●
●
●
●
DD VU
4
●
●
VU
●
●
●
●
●
●
●
●
VU
VU
●
Ukraine
●
Sub‐polar & North Urals
●
SE Russia
●
NW Russia
●
NE Russia
●
Moldova
●
Middle and South Urals
●
Lithuania
EN
Kazakhstan (in Europe)
●
Slovenia
●
EN
?
Serbia
Macedonia
Kosovo
Hungary
Crete
●
Latvia
CR VU
●
Kaliningrad
EN EN
R
VU
●
Estonia
●
●
Crimea
CR
●
Central Russia
●
●
Caucasus (in Europe)
VU
NT VU
●
?
●
Belarus
R
●
●
Arctic Russia
VU VU
●
●
●
Turkey
●
Greece
●
Romania
DD
VU
NT
Croatia
R
Bulgaria
●
Bosnia‐Herzegovina
●
Albania
●
Appendix 7 - Mosses
Montenegro
●
●
Switzerland
●
Slovakia
3
●
Poland
?
●
Netherlands
Mn
Luxembourg
2
●
Liechtenstein
Belgium
Germany
Austria
Czech Republic
Checklist and country status of European bryophytes
CR
●
●
●
●
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DD
●
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●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
3
●
●
●
CR 3
CR nb
●
●
●
●
● KW ●
VU
NT
●
●
●
●
●
●
●
●
?
3
VU
●
●
●
nb
●
●
●
●
■
□
■
□
□
■
□
●
●
VU
VU
DD
●
●
●
●
●
●
●
CR
□
●
NT
●
●
●
●
●
●
●
●
●
□
□
●
●
●
●
●
●
●
EN
●
●
●
●
●
□
□
■
□
□
□
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
1
●
●
●
□
□
□
□
□
□
□
●
NT DD* ●
D
■
□
■
□
●
4
●
●
●
●
●
VU
●
EN
R
●
NT
●
DD
RE
NT
●
VU
●
●
●
●
●
●
●
●
RE
●
●
●
●
●
●
●
●
●
●
●
86
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU ●
●
●
●
●
●
●
● VU ●
●
●
Leptodontium gemmascens
y
●
EN
● EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
DD
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
y
DD
Leptotheca gaudichaudii
●
Lescuraea incurvata NT ●
●
●
●
●
●
VU
●
●
●
● RE ●
●
Lescuraea plicata RE ●
● ●
●
●
Lescuraea radicosa ●
DD
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
DD
●
DD
●
●
●
Lescuraea saviana ●
Lescuraea saxicola
●
●
●
●
●
●
●
●
●
●
●
Lescuraea mutabilis
Lescuraea patens ●
RE
●
●
NT
●
DD
●
●
●
●
●
●
DD
●
DD
●
y
Leskea polycarpa
●
●
●
●
●
●
y
●
Leucobryum glaucum
●
Leucobryum juniperoideum
●
Leucodon canariensis
y
Leucodon flagellaris
y
Leucodon immersus
y
Leucodon pendulus
y
Leucodon sciuroides
●
●
●
●
DD
●
●
Leucodon sciuroides var. morensis
●
●
●
●
■
Leucodon sciuroides var. sciuroides
□
Leucodon treleasei
y
Lindbergia dagestanica y
Lindbergia grandiretis y
Loeskeobryum brevirostre
■
□
■
■
●
●
●
●
●
●
VU ●
●
●
●
●
●
●
DD
●
●
●
●
?
NT
●
●
●
■
■
■
●
●
y
●
EN
Meesia triquetra
y
Meesia uliginosa
y
●
●
●
●
●
●
CR
CR
VU
NT
● ●
● RE
● ● ●
●
RE
VU
NT
●
Microbryum davallianum
Microbryum floerkeanum
y
Microbryum fosbergii
y
Microbryum longipes
y
Microbryum rectum
□
●
y
Microbryum curvicollum
●
●
●
●
●
□
■
□
□
□
●
●
●
●
■
■
■
■
VU
●
●
●
□
□
DD
●
●
●
■
■
□
■
■
□
■
●
●
●
●
●
●
●
NT
●
●
RE
● EN
●
● DD ●
●
●
●
DD
●
●
●
●
●
●
DD
●
●
●
CR
CR
●
DD
●
●
●
●
DD
●
●
DD
VU
●
Microbryum starckeanum
●
RE ●
● ●
● RE RE
CR ●
●
●
●
y
Microeurhynchium pumilum ●
●
●
■
Meesia longiseta
●
●
●
●
●
Meesia hexasticha
●
●
RE
●
● ● ● ● ●
EN
EN ● VU
VU
●
●
?
Loeskypnum badium
DD DD DD
●
■
□
●
●
EN
Micromitrium tenerum
y
Mielichhoferia elongata
y
CR
Mielichhoferia mielichhoferiana
y
CR
NT ●
●
Mnium blyttii
●
RE
● VU
EN
VU
●
EN
RE
●
●
●
●
●
●
●
VU
●
● NT ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
□
■
■
●
●
●
●
●
●
●
DD
●
●
EN
VU
●
●
NT
●
●
y
Mnium hornum
●
Mnium lycopodioides
Mnium marginatum
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
■
■
●
●
Mnium marginatum var. dioicum
Mnium marginatum var. marginatum
□
□
Mnium spinosum
●
●
□
□
□
■
■
NT ●
●
●
●
NT
■
●
●
●
●
■
●
●
Mnium thomsonii
Molendoa hornschuchiana
y
Molendoa schliephackei
y
●
●
●
●
●
●
●
●
●
●
●
□
■
Mnium spinulosum
Mnium stellare
●
●
●
●
●
● NT NT ●
●
●
●
●
●
?
●
■
■
●
●
●
●
●
●
●
NT
●
●
●
●
DD
●
y
●
Molendoa taeniatifolia
y
●
Molendoa tenuinervis
y
●
●
□
Molendoa sendtneriana
●
●
●
DD
●
●
DD
Vatican City
●
Spain
DD
●
●
87
Portugal
●
NT
Leptophascum leptophyllum
Mnium heterophyllum
●
● NT NT
Leptodontium proliferum Microcampylopus laevigatus
●
●
●
y
y
Leucobryum albidum
Sicily
●
●
Leptodontium flexifolium
Lescuraea secunda
Sardinia
●
●
●
Leptodon smithii
Leptodontium styriacum
San Marino
●
Leptodon corsicus Leptodon longisetus ●
●
?
Monaco
●
●
Malta
●
●
DD
Madeira
●
●
Italy
●
●
France
●
● DD
Leptodictyum riparium
●
NT
●
Leptobarbula berica
Leptobryum pyriforme
●
Cyprus
●
Andorra
Great Britain
Gibraltar
Channel Islands
●
Corsica
●
Canary Islands
● VU ●
Balearic Islands
●
Sweden
Svalbard
Norway
●
Azores
●
●
Northern Ireland
●
Kindbergia praelonga
Iceland
DD* ● VU
Kiaeria starkei
Ireland
y
Finland
Faeroe Islands
Taxon
Kiaeria riparia
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
Sub‐polar & North Urals
Ukraine
SE Russia
NW Russia
NE Russia
Moldova
Middle and South Urals
Lithuania
Latvia
Kazakhstan (in Europe)
Kaliningrad
Estonia
Crimea
Central Russia
Caucasus (in Europe)
Belarus
●
Arctic Russia
●
Turkey
●
Slovenia
Romania
DD*
Serbia
Montenegro
Macedonia
●
Kosovo
●
Hungary
●
Appendix 7 - Mosses
Greece
●
VU
●
Croatia
VU
●
●
Crete
●
●
Bulgaria
●
●
Bosnia‐Herzegovina
●
●
Albania
●
Switzerland
●
Slovakia
●
Poland
Liechtenstein
●
Netherlands
Germany
●
Luxembourg
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
●
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
0
●
GE
RE VU
●
RE VU
3
CR
G
NT GE
4
R
‐
●
*
DD*
●
V
●
●
EN
●
●
●
●
●
●
EN
●
●
●
●
EN
●
●
EN
R
EN
EN
●
R
●
●
nb
DD R
●
●
●
●
●
●
VU
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
4
●
VU
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
VU
NT
●
●
●
VU
●
●
CR
●
●
●
●
●
●
●
VU
●
●
●
●
●
?
●
●
●
VU
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU CR VU ●
● VU ● NT
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
?
●
●
●
●
●
R
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
2
●
●
●
●
●
●
●
●
●
●
●
●
●
●
3
●
●
●
●
●
●
●
●
nb
●
●
BE
●
●
●
D
□
□
■
●
●
●
●
■
□
□
□
□
□
□
■
●
●
■
□
□
□
●
●
■
□
□
●
●
●
●
■
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
■
□
□
□
■
●
3
3
●
NT
G
●
●
BE
V
RE
●
●
VU
●
?
DD
●
EN
●
●
●
●
●
0
Ex RE
0
RE
0
E
EN RE
● DD*
?
2
●
Mn CR
CR
1
3
V
E
EN NT
VU ●
●
●
DD
●
●
3
3
●
●
VU
●
●
●
●
●
●
●
●
V
V
VU BE R
● KW ●
VU EN
NT DD
DD
●
0
Mn VU
V
NT GE
●
EN CR
DD*
0
●
Mn DD
2
1
GE
CR RE
R
DD
VU EN
EN
●
●
●
●
●
●
●
EN
●
●
●
NT
●
●
DD
●
●
●
2
●
EN VU
●
●
●
0
● CR EN
VU NT
●
●
●
EN
●
●
●
NT
GE
EN
VU
●
●
CR
●
●
●
CR
●
CR
●
RE
●
0
4
●
●
●
●
RE
●
●
●
●
NT
EN
●
●
●
●
2
4
●
●
●
●
●
●
●
●
EN
VU
●
●
NT
●
●
●
DD
●
●
CR
R
VU
VU
DD
●
●
●
●
●
●
●
●
CR
●
●
●
●
●
DD*
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
NT
●
VU
●
?
NT
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
■
□
□
□
□
□
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
V
●
● KW ●
●
●
●
●
●
●
□
□
■
nm
□
□
□
□
□
■
□
□
□
●
V
●
●
●
●
●
●
●
●
●
●
●
●
●
● KW ●
DD
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
CR
●
R
●
●
R
●
●
EN
●
●
●
●
●
VU
●
?
2
●
●
3
□
●
●
●
CR CR
VU DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
●
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
□
■
□
●
●
●
●
●
3
●
●
●
DD
VU
●
●
3
●
●
●
4
●
●
VU
●
?
●
●
●
DD
●
●
●
4
4
●
●
●
●
DD*
0
RE
EN
●
VU
0
●
●
RE
RE
CR
RE
●
NT CR EN
●
●
●
●
4
4
R
NT DD*
CR
88
NT ●
Myurium hochstetteri
y
Myuroclada maximowiczii
y
Neckera cephalonica
y
Neckera menziesii
y
y
Neckera pumila
y
Oedipodium griffithianum
y
Oligotrichum hercynicum
y
Oncophorus elongatus
y
●
●
●
NT
●
EN
●
NT ●
●
●
●
VU
●
●
VU ●
●
●
DD
●
●
●
●
Oreoweisia torquescens
y
Orthodontium gracile
y
●
VU ●
VU
●
●
●
RE
●
VU
●
●
●
●
●
●
DD
●
Orthodontium pellucens
y
Orthothecium chryseon
y
●
●
●
●
●
●
●
●
●
●
VU
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
□
■
□
□
□
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
CR
y
●
●
●
●
●
●
●
●
□
●
VU
CR
● ● ●
EN ●
●
NT ●
●
●
●
●
y
y
Orthotrichum consimile
y
Orthotrichum consobrinum y
Orthotrichum crenulatum
y
Orthotrichum cupulatum
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
RE
●
● NT NT
●
●
●
●
●
●
●
●
DD
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
●
●
DD ●
●
●
●
●
●
●
●
●
●
● DD ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
CR
DD
●
●
DD
●
CR
●
●
VU ●
●
●
●
●
●
● NT ●
●
Orthotrichum cupulatum var. bistratosum
●
●
●
●
DD
●
●
■
■
□
□
■
■
■
■
DD
■
DD
●
●
■
Orthotrichum cupulatum var. cupulatum
●
●
●
Orthotrichum casasianum
■
●
Orthotrichum alpestre
Orthotrichum cambrense ●
CR
●
y
y
■
●
●
●
DD
Orthotrichum callistomum
●
DD
EN ● VU
Orthotrichum acuminatum
Orthotrichum anomalum
●
DD
□
■
VU
● ●
● NT ●
Orthothecium rufescens
Orthotrichum affine var. affine
VU
●
●
y
Orthotrichum affine
●
□
■
□
□
■
■
■
■
■
□
■
□
y
Orthotrichum cupulatum var. riparium
■
■
■
■
●
●
●
■
■
●
■
■
■
■
y
Orthotrichum diaphanum
●
Orthotrichum handiense
y
Orthotrichum hispanicum
y
Orthotrichum holmenii y
Orthotrichum ibericum
y
Orthotrichum laevigatum
y
Orthotrichum lyellii
y
Orthotrichum microcarpum
y
Orthotrichum moravicum y
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
EN
●
●
●
CR ● VU
●
Orthotrichum macrocephalum
●
CR
●
●
Orthotrichum patens
y
CR
VU
Orthotrichum pellucidum
y
CR
●
Orthotrichum philibertii
y
Orthotrichum pulchellum
y
Orthotrichum pallens
●
●
●
●
●
●
●
●
●
EN EN
EN
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
DD DD
CR
● VU
●
●
●
●
●
●
● NT
●
●
●
●
DD
●
VU
●
●
●
●
●
EN
●
●
●
●
●
●
●
DD
●
● VU ●
●
●
●
DD
DD
●
NT VU
● VU
●
●
Vatican City
Spain
Sicily
San Marino
Portugal
Sardinia
●
DD
● CR
DD
●
Orthothecium intricatum
89
VU
●
●
●
●
●
●
Orthodontium lineare
Orthotrichum pylaisii
●
●
●
RE
■
y
Orthotrichum pumilum
Monaco
●
●
y
Oreas martiana
Orthotrichum dagestanicum Malta
●
VU
Oncophorus wahlenbergii var. wahlenbergii
Orthotrichum cupulatum var. fuscum
Madeira
Cyprus
Corsica
Canary Islands
Balearic Islands
Azores
●
●
●
Oncophorus wahlenbergii var. compactus
Orthotrichum affine var. bohemicum Andorra
EN
●
Oncophorus virens
Orthothecium strictum
●
VU
●
Oncophorus dendrophilus Orthothecium lapponicum
●
●
●
y
Oedipodiella australis
Oncophorus wahlenbergii
●
●
●
y
Nyholmiella obtusifolia
Ochyraea tatrensis
DD
●
y
Nogopterium gracile
Nyholmiella gymnostoma
●
● RE
Neckera oligocarpa
Nobregaea latinervis
●
●
● VU
NT
● EN EN ●
y
Myurella tenerrima
Neckera pennata
Italy
Myurella sibirica
France
●
Myurella julacea
● VU
Northern Ireland
NT
DD
VU
● ● ● ● ● ●
Ireland
●
y
Great Britain
y
Myrinia pulvinata
Gibraltar
Taxon
Molendoa warburgii
Channel Islands
Sweden
Svalbard
Norway
Iceland
Finland
Faeroe Islands
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
●
NW Russia
SE Russia
Sub‐polar & North Urals
Ukraine
●
●
NE Russia
Moldova
●
●
1
3
Middle and South Urals
●
●
Lithuania
●
DD
Latvia
EN
●
●
Kazakhstan (in Europe)
Estonia
●
●
Kaliningrad
Crimea
●
EN
VU
Belarus
●
●
Arctic Russia
Central Russia
EN
VU NT
Caucasus (in Europe)
4
●
●
●
●
BE
●
Turkey
●
NT
Slovenia
●
●
Serbia
R
●
Romania
●
Montenegro
Bulgaria
●
Macedonia
Bosnia‐Herzegovina
●
Kosovo
Albania
●
Hungary
Switzerland
●
Greece
Slovakia
●
Appendix 7 - Mosses
Croatia
Poland
●
Crete
Liechtenstein
V
Netherlands
Germany
EN
Luxembourg
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
2
CR
2
DD
CR CR ●
CR
●
●
●
●
●
●
●
●
CR
●
●
4
●
1
VU
1
RE
2
●
●
3
●
0
●
?
3
●
RE
●
1
●
0
●
●
●
E
EN CR
EB
E
EN VU
●
R
●
CR
●
RE VU
●
VU
●
●
EN
●
VU VU VU
●
●
●
EN
●
●
DD
●
DD VU DD
●
●
●
●
●
●
DD
●
●
VU
●
●
DD
NT
●
●
DD*
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
2
2
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
DD
● VU ●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
●
●
●
●
●
●
●
?
●
?
R
●
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
VU
●
VU
EN
●
●
●
●
4
RE
●
0
■
nm
●
●
I
NT ●
VU VU
●
●
‐
‐
●
●
●
●
RE
●
□
□
■
□
4
4
●
●
●
□
R
●
●
●
●
D
●
●
V
●
Mn RE
□
VU
□
■
■
3
CR VU
●
4
□
●
●
●
●
●
●
EN
●
D
●
●
VU CR
●
●
●
●
●
●
●
●
●
●
DD
●
VU
●
●
NT
●
●
●
●
●
DD EN
R
●
●
●
CR
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
3
●
●
■
CR
R
●
●
●
●
●
CR EN DD* ●
●
●
●
●
●
DD*
●
●
DD*
●
●
RE
●
R
●
●
●
■
□
■
4
RE
●
●
●
●
●
●
●
●
EN GE
nb
●
VU
●
●
V
□
■
□
□
D
2
●
●
●
●
●
●
●
●
■
■
□
□
□
□
□
VU DD
D
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
□
□
■
□
□
■
■
■
■
□
□
■
■
■
■
■
■
■
□
■
■
■
■
■
■
■
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
VU
●
2
1
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
R
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
nb
‐
●
3
●
●
RE NT
CR
●
●
DD
●
●
NT
●
●
VU GE
●
●
NT
R
CR NT
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
?
VU
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Mn
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
CR EN
EN ● DD
●
●
●
NT
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
90
Orthotrichum scanicum
●
●
●
●
CR
●
CR
●
RE
●
y
Orthotrichum sordidum
y
Orthotrichum speciosum
●
●
●
●
●
●
●
●
RE
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
y
Orthotrichum stellatum
y
●
●
●
●
■
□
□
□
NT VU
●
●
●
●
●
●
Orthotrichum striatum
●
EN ●
●
●
Orthotrichum tenellum
●
y
Orthotrichum vittii
y
Orthotrichum vladikavkanum
y
●
●
●
DD
●
●
●
●
VU
●
DD
●
NT
●
●
DD
●
●
NT VU
●
EN
●
●
Oxyrrhynchium speciosum
●
●
●
●
●
RE ●
●
●
●
●
●
●
●
●
●
●
VU
●
●
● CR
●
NT ●
● NT NT
y
● NT
Oxystegus minor y
●
Oxystegus recurvifolius y
●
●
● NT ●
●
●
Oxystegus tenuirostris var. tenuirostris
●
●
●
●
●
● VU ●
●
NT ●
●
NT ●
●
Palustriella decipiens
Palustriella falcata
●
●
●
●
●
●
●
●
●
●
●
● DD ●
●
●
●
●
●
●
●
●
●
●
●
●
● NT ●
●
●
●
●
●
Paraleucobryum longifolium
●
Paraleucobryum sauteri
y
Pelekium atlanticum
y
Pelekium minutulum
y
●
Philonotis capillaris ●
y
Philonotis fontana
y
Philonotis marchica
y
■
■
■
■
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● EN
NT ●
●
●
●
NT
■
□
□
●
●
●
●
●
●
●
●
●
●
●
Physcomitrium eurystomum
y
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
■
□
□
□
□
□
□
●
EN
●
?
●
NT
●
DD NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
VU
●
●
●
DD
●
●
●
●
●
EN ●
●
●
●
●
●
●
●
●
●
●
NT
●
●
● VU ●
●
●
●
●
●
●
●
NT
NT
●
●
●
●
●
● EN EN
●
●
● NT NT ●
DD NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
● VU VU ●
●
●
●
●
●
●
●
● NT ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD ●
●
●
●
●
●
●
●
●
NT VU
●
●
●
y
●
●
●
y
Physcomitrium arenicola
DD
●
●
●
●
●
y
DD
●
●
●
●
●
●
●
Philonotis seriata
Physcomitridium readeri ●
■
NT
●
Physcomitrella patens
●
RE CR
●
●
Philonotis rigida
Philonotis tomentella
●
●
RE CR
●
Philonotis hastata
●
●
●
●
Philonotis falcata ●
VU
Philonotis calcarea
y
●
●
●
‐
VU
Philonotis caespitosa
Philonotis cernua
DD
NT
●
y
Paraleucobryum enerve
●
■
VU
●
●
DD ●
● NT
● ● ●
●
Oxystegus tenuirostris var. holtii y
●
●
●
Oxystegus hibernicus
Palustriella commutata
●
■
□
DD
●
●
NT
y
Paludella squarrosa
□
■
●
● VU VU
Oxystegus daldinianus y
■
●
■
●
Oxyrrhynchium schleicheri
Palamocladium euchloron
■
NT
●
Oxyrrhynchium hians
Oxystegus tenuirostris
□
CR
VU ●
Orthotrichum urnigerum
□
● VU VU
●
y
□
■
Orthotrichum stramineum
Orthotrichum tortidontium
EN
NT
NT
●
●
●
●
‐
●
●
●
●
●
EN
●
●
□
□
●
●
●
●
● DD ●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
Physcomitrium eurystomum subsp. acuminatum
Physcomitrium eurystomum subsp. eurystomum
□
Physcomitrium pyriforme
●
Physcomitrium sphaericum
y
Plagiomnium affine
Plagiomnium confertidens
y
Plagiomnium curvatulum
y
Plagiomnium cuspidatum
Plagiomnium drummondii
●
●
●
●
●
●
●
NT
y
●
Plagiomnium ellipticum
Plagiomnium medium
●
●
Plagiomnium rostratum
●
Plagiomnium undulatum
●
91
●
RE
Plagiomnium elatum
Plagiomnium undulatum var. madeirense
□
●
●
●
●
DD ●
●
●
VU
● ●
●
●
●
●
●
NT ●
●
●
●
●
●
●
●
●
RE
●
●
●
●
●
●
● DD ●
VU
●
●
●
●
DD
●
DD
●
●
●
DD
●
●
●
● NT NT
●
●
●
●
●
●
●
● ●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD ●
●
●
●
●
DD
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
DD
●
■
■
■
●
■
●
Vatican City
Spain
●
■
Orthotrichum sprucei
Sicily
Sardinia
San Marino
Portugal
Monaco
Malta
Madeira
Italy
●
●
●
●
Orthotrichum speciosum var. speciosum
Philonotis uncinata
France
●
Orthotrichum speciosum var. brevisetum
Palustriella pluristratosa
Cyprus
Corsica
Canary Islands
Balearic Islands
Azores
Andorra
Northern Ireland
Ireland
Great Britain
Gibraltar
Channel Islands
● NT NT
Orthotrichum schimperi
Orthotrichum shawii
Sweden
y
●
●
Orthotrichum rupestre
Svalbard
●
Norway
y
Iceland
y
Orthotrichum rogeri
Finland
Taxon
Orthotrichum rivulare
Faeroe Islands
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
?
nb
DD
●
●
NT
DD
NT GE
CR ●
R
E
●
●
CR CR
DD*
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
Mn
●
●
●
●
■
□
■
nm
□
□
●
1
R
Ukraine
Sub‐polar & North Urals
●
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
SE Russia
NW Russia
NE Russia
Moldova
Middle and South Urals
Lithuania
Latvia
●
Kazakhstan (in Europe)
●
CR
CR
●
Kaliningrad
●
Central Russia
●
Caucasus (in Europe)
NE
Belarus
●
●
Arctic Russia
●
●
EN CR
Turkey
Romania
Montenegro
Macedonia
●
Estonia
?
Crimea
●
●
Kosovo
Hungary
Greece
Croatia
Crete
Bulgaria
Bosnia‐Herzegovina
DD
GE DD* RE VU
Slovenia
●
●
Appendix 7 - Mosses
Serbia
NE
Albania
3
3
Switzerland
VU
CR
Slovakia
●
●
EN
Poland
●
0
1
Netherlands
D
Luxembourg
VU
●
Liechtenstein
Germany
●
Belgium
1
Austria
Czech Republic
Checklist and country status of European bryophytes
●
●
NT
●
●
●
●
●
●
●
□
□
□
■
□
□
□
□
□
■
GE
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
DD
VU
●
3
4
CR
CR
1
●
CR
●
EN
●
CR
●
3
Mn
●
●
●
CR
●
●
V
DD
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
V
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
2
●
DD
●
●
●
●
R
DD VU
●
●
●
●
●
●
DD
DD* NT
EN
NT
● DD*
●
●
●
●
●
●
●
●
●
4
VU
2
R
CR CR
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
CR
3
●
●
3
NT
●
●
●
●
●
●
●
●
DD*
●
DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
VU
●
●
●
DD
●
●
●
EN
●
●
●
●
●
●
●
●
●
V
●
●
●
NT
●
●
●
●
□
□
■
□
□
□
□
□
□
□
□
□
●
VU
●
NT
●
●
●
●
●
□
□
□
□
□
□
□
□
□
●
2
EN
1
●
V
●
●
V
●
●
●
D
●
●
●
●
●
●
●
RE
●
●
●
●
●
●
●
●
DD
RE
E
NT BE
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
VU
●
RE
0
●
●
●
V
●
●
●
3
2
●
EN
3
●
●
●
V
●
NT
●
●
●
●
I
●
EN VU
●
●
●
●
●
EN
●
?
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
VU
●
●
DD
●
●
●
●
●
●
●
VU
●
●
EN VU DD* ●
VU
●
●
EN
●
DD
●
●
●
DD
●
●
●
●
NT
●
●
VU
●
●
●
DD
●
●
DD
VU DD
VU
R
VU
●
●
R
CR GE
●
NT RE
V
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
VU
●
●
●
●
●
●
EN
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
DD*
●
DD VU
●
●
VU
●
●
●
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
DD EN DD NT
●
EN
●
2
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
2
●
●
●
□
●
EN
●
□
NT
●
●
●
?
●
VU
●
●
●
●
●
●
●
●
0
3
●
●
●
●
NT
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
2
RE CR
●
RE
●
●
●
●
●
3
3
●
●
●
●
●
3
EV EB
V
●
3
RE
●
●
●
VU
G
I
●
●
Mn VU
●
3
VU CR DD
VU
●
●
VU VU
GE
●
VU VU
●
●
●
NT
●
●
●
●
●
NT
●
●
VU
NT
●
●
●
VU
●
●
●
●
●
DD
●
●
● NT
VU CR
EN
●
E
□
□
■
nm
●
●
●
●
2
●
Mn VU
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
■
□
□
●
●
●
●
●
VU
●
R
●
●
●
□
●
RE
●
●
●
●
●
●
●
●
●
□
■
●
●
DD
●
●
●
●
●
●
●
●
VU
●
●
VU
●
●
●
●
●
1
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
?
●
□
□
■
■
●
●
CR
●
3
3
●
●
●
●
●
●
Mn NT
●
3
●
●
●
●
‐
●
●
NT BE
●
●
●
VU KW ●
EN ●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
NT
DD
●
●
●
●
VU
●
●
●
●
●
DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
●
●
●
●
●
●
0
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
●
●
●
●
●
●
■
●
●
●
●
■
●
●
●
■
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
92
Plagiothecium berggrenianum
□
●
■
●
DD
●
□
■
□
□
■
□
□
□
■
■
□
□
Monaco
●
Malta
●
Cyprus
● CR
Corsica
●
■
●
●
●
●
●
Plagiothecium denticulatum
●
●
●
●
●
Plagiothecium denticulatum var. denticulatum
■
□
■
□
□
●
Plagiothecium denticulatum var. obtusifolium
●
●
●
● VU
●
●
●
●
●
●
● VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
□
■
■
■
■
■
□
□
■
■
□
□
■
■
■
■
■
■
■ NT NT
Plagiothecium denticulatum var. undulatum
■
■
■
■
■
■
Plagiothecium laetum
●
●
●
NT
●
●
●
●
● VU VU ●
● VU
●
●
●
●
●
NT
●
●
DD ●
●
●
●
●
●
●
●
●
EN
●
NT
●
●
●
DD ●
●
●
●
●
●
●
■
■
■
●
●
●
y
Plagiothecium nemorale
Plagiothecium piliferum
y
Plagiothecium succulentum
●
●
●
●
●
y
●
●
●
●
●
RE
● VU
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
DD
●
●
VU
●
●
●
●
●
●
●
Plagiothecium undulatum
●
●
●
●
Plasteurhynchium meridionale
VU
Plasteurhynchium striatulum
NT
VU
●
● NT
●
●
●
●
●
● DD ●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
NT
●
Platydictya jungermannioides
●
●
Platygyrium repens
●
●
●
Pleuridium acuminatum
●
VU
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Pleuridium subulatum
●
NT ●
●
●
●
●
●
●
?
● DD ●
●
●
●
●
●
●
●
Pleurozium schreberi
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● EN EN
?
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
DD ●
●
●
●
●
●
●
DD
●
●
●
●
●
●
VU ●
●
●
●
DD
Platyhypnidium grolleanum
●
●
Pogonatum dentatum
Pogonatum nanum
Pogonatum urnigerum
y
Pohlia annotina
VU
●
Pohlia atropurpurea
y
Pohlia beringiensis y
Pohlia bolanderi
y
Pohlia bulbifera
●
●
●
●
●
●
EN
● NT NT
●
●
●
●
●
●
●
●
●
●
Pohlia camptotrachela
●
●
Pohlia cruda
●
●
●
●
●
Pohlia crudoides
●
●
●
●
●
●
□
□
■
Pohlia elongata var. acuminata
Pohlia elongata var. greenii
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
RE
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
□
□
□
■
■ NT NT □
y
□
■
■ EN
EN
NT
DD
● ● ● ● ●
●
●
y
DD
Pohlia flexuosa var. pseudomuyldermansii
●
□
●
●
●
●
●
● NT ●
●
●
●
●
Pohlia lescuriana
Pohlia longicolla
y
●
●
●
●
●
●
●
DD ●
●
●
●
●
?
●
●
VU
●
■
□
●
■
■
■
■
■
■
● NT NT
●
●
DD
●
●
●
EN
■
□
●
□
■
●
■ □ □
● DD
●
●
●
●
●
●
●
●
●
DD
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Pohlia nutans subsp. nutans
□
□
■
□
■
□
■
□
□
□
□
□
□
□
□
■
□
□
●
●
●
●
●
●
●
●
Pohlia saprophila
y
Pohlia scotica
y
Pohlia sphagnicola
y
Pohlia tundrae
y
Pohlia vexans
y
Pohlia wahlenbergii
●
●
●
●
●
●
●
93
●
●
DD
●
●
DD NT
□
●
●
□
□
●
VU
●
●
●
■
■
■
●
●
●
●
EN
●
RE
VU
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
■ DD DD
■
●
●
●
●
●
■
■
■
■
■
■
●
Pohlia wahlenbergii var. calcarea
Pohlia wahlenbergii var. glacialis
●
VU
VU
●
●
●
●
■
VU
●
●
●
■
?
VU
●
●
DD
DD
●
●
■
●
●
●
●
●
■
□
●
y
●
■
□
●
Pohlia proligera
VU
VU ●
Pohlia nutans
■
‐
DD
Pohlia melanodon
Pohlia nutans subsp. schimperi
●
●
●
●
DD
DD DD
DD
●
Pohlia lutescens
NT
●
VU
●
Pohlia flexuosa var. flexuosa
Pohlia obtusifolia
●
●
●
■
■
Pohlia filum
●
●
■
Pohlia elongata var. elongata
●
●
●
●
NT
Pohlia elongata
● ● ●
NT EN
●
●
Pohlia ludwigii
●
●
●
NT ● VU
●
●
Pohlia drummondii
●
●
●
●
Pohlia flexuosa
●
●
VU ●
●
●
●
Pohlia erecta
●
y
Pohlia andalusica
Pohlia andrewsii
●
y
Pogonatum aloides
Pogonatum neesii
●
■
■ RE
●
● VU ●
●
●
■
Vatican City
Spain
□
?
●
●
Plagiothecium svalbardense
□
■
●
Plagiothecium curvifolium
Plagiothecium platyphyllum
■
□
●
Plagiothecium cavifolium
Plagiothecium neckeroideum
■
□
●
■
y
Plagiothecium latebricola
Sicily
□
Sardinia
□
San Marino
Balearic Islands
□
Portugal
Azores
□
Madeira
Andorra
□
Italy
Northern Ireland
□
Canary Islands
Ireland
□
●
Plagiopus oederianus var. alpinus
Plagiopus oederianus var. oederianus
■
Great Britain
□
Gibraltar
□
Channel Islands
Norway
■
Sweden
Iceland
□
Svalbard
Finland
□
France
Plagiopus oederianus
Faeroe Islands
Taxon
Plagiomnium undulatum var. undulatum
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
Sub‐polar & North Urals
Middle and South Urals
□
□
□
□
□
●
●
●
●
VU
3
●
●
●
●
●
■
□
□
nm
□
□
□
□
□
□
□
□
□
□
■
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
□
■
■
■
■
■
■
■
NT
■
■
VU VU
●
VU
●
V
●
●
EV
●
●
●
●
CR
●
●
●
●
●
VU
3
●
2
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
● DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
□
□
□
□
■
■
□
□
EN
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
■
■
●
●
EN
R
●
●
●
V
CR
●
●
●
●
●
●
●
●
●
●
■
■
●
●
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
RE CR
●
●
●
V
VU
●
●
GE
EN
●
●
●
●
●
●
●
●
●
●
VU
●
DD
●
●
NT
●
●
●
● DD*
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
VU
●
EN
●
●
●
●
●
●
●
●
VU
●
●
NT
●
●
●
●
●
●
●
●
● DD*
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
■
■
●
■
DD*
■
■
●
●
●
●
●
NT
●
●
●
2
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
● DD* ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
● DD* ●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
4
●
VU
3
●
●
●
VU
●
●
●
DD CR
●
●
3
?
?
●
?
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
4
●
●
●
●
●
●
●
●
●
●
● KW ●
●
●
●
●
I
●
●
●
●
● KW
?
●
●
●
■
●
●
●
●
■
■
●
●
●
●
●
■
●
●
●
●
●
NT
●
●
□
●
●
●
●
●
DD*
●
●
●
■
●
●
●
●
DD
●
●
●
■
●
●
●
●
●
●
●
?
●
●
□
■
●
●
●
□
●
●
●
●
□
EN
●
DD
●
●
□
■
■
EN
●
●
□
■
●
VU
●
●
CR
●
●
■
VU
●
□
Ukraine
□
SE Russia
□
●
NW Russia
□
●
NE Russia
□
●
■
Moldova
□
●
Lithuania
■
●
Latvia
Kaliningrad
Kazakhstan (in Europe)
Estonia
□
VU
Crimea
□
●
Central Russia
□
●
Belarus
Turkey
□
●
Arctic Russia
Slovenia
□
●
Serbia
□
●
Romania
□
●
Montenegro
□
●
Macedonia
□
I
Kosovo
□
Hungary
Crete
□
RE
Greece
Bulgaria
□
●
Croatia
Poland
□
V
Albania
Netherlands
nm
Switzerland
Luxembourg
■
VU
Slovakia
Liechtenstein
□
●
Belgium
□
●
■
Austria
Germany
Caucasus (in Europe)
Appendix 7 - Mosses
Czech Republic
Bosnia‐Herzegovina
Checklist and country status of European bryophytes
●
●
●
●
●
●
●
●
EN
●
●
RE
●
EN
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
D
●
●
●
●
●
●
● KW ●
EB ●
R
●
EN
●
●
●
●
●
●
●
NT
●
●
●
●
●
DD
●
●
●
DD
●
●
NT
●
●
●
DD NT
NT
●
●
●
nb
●
●
●
● DD* ●
●
DD
●
NE
●
●
●
●
●
DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
4
●
●
●
●
●
V
●
●
●
●
●
●
CR
NT
●
●
●
●
●
●
●
DD VU
DD VU
●
●
●
●
●
●
●
●
●
CR
●
DD
●
●
●
●
●
●
● EN
DD
●
●
●
●
●
●
●
●
●
●
●
●
VU
NE
●
●
●
●
●
●
●
●
3
●
●
●
●
●
□
■
●
nb
●
●
CR BE
●
●
0
■
■
●
2
●
●
●
VU
NE
●
4
4
□
●
●
●
●
●
●
●
DD
□
■
□
■
DD
●
●
●
●
NT
●
●
●
□
□
□
□
VU
□
□
□
□
■
D
■
●
●
G
●
VU
R
●
nm
□
●
●
●
●
EN
3
●
●
●
VU
DD
R
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
□
□
■
□
■
NT
D
■
●
●
RE
●
R
V
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
■
□
□
■
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
DD
■
■
●
●
●
■
DD VU
●
VU
■
■
●
●
●
?
●
●
●
●
CR
■
NE
●
●
●
?
EN NT
●
?
●
●
●
●
VU
●
●
?
●
●
●
●
●
●
●
DD
NT ●
RE NT
VU NT
■
■
DD CR
VU
●
□
DD
CR
●
●
●
●
●
●
●
●
NE
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
■
■
□
□
□
□
■
■
■
■
■
□
■
□
□
■
■
■
■
■
■
EN
CR
■
DD
●
VU
NT
●
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
●
□
□
□
VU ●
EN NT
●
VU
VU
●
?
●
●
●
■
EN
VU
●
●
■
●
●
●
?
0
?
2
4
CR
R
●
nb
R
VU
3
●
VU
●
●
●
●
●
D
■
EN
D
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
■
■
■
●
●
●
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
94
Polytrichastrum sphaerothecium
●
●
●
●
●
●
●
●
●
●
●
□
□
■
■
□
□
■
■
■
■ ■ ■
■ DD
□
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Polytrichum commune var. perigoniale Polytrichum hyperboreum
Polytrichum jensenii
●
Polytrichum longisetum
Polytrichum pallidisetum
Polytrichum strictum
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
Pseudobryum cinclidioides
y
●
●
Pseudocampylium radicale
y
●
NT
Pseudocrossidium hornschuchianum
●
●
Pseudocrossidium obtusulum
y
Pseudocrossidium replicatum
y
Pseudocrossidium revolutum
■
DD
●
●
●
●
●
●
●
VU ●
●
●
●
●
●
NT
●
□
■
□
■
■
■
□
□
■
■
■
□
■
□
VU ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
DD DD
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
?
●
●
●
●
●
DD
EN
●
NT
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
Pseudoleskeella papillosa
y
NT
y
NT ●
Pseudoleskeella tectorum
●
●
●
●
●
EN
●
●
CR
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
NT
●
y
Pseudoscleropodium purum
VU
●
Pseudoleskeella rupestris
●
●
●
●
● NT ●
●
●
●
●
●
●
●
●
● NT ●
●
●
●
●
●
●
●
●
●
●
EN ●
?
DD
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
DD
●
DD
EN
●
●
●
●
●
y
Pseudotaxiphyllum elegans
Pseudotaxiphyllum laetevirens
■
●
y
Pseudoleskeella nervosa
Pseudosymblepharis bombayensis ■
●
DD
●
NT
Pseudoleskeella catenulata
Pseudorhynchostegiella duriaei
●
●
●
●
Pseudoamblystegium subtile
□
●
DD
●
●
y
Pseudephemerum nitidum
Pseudoleskea artariae
●
●
DD
●
Polytrichum swartzii
Pottiopsis caespitosa ●
●
y
Polytrichum piliferum
■
●
●
Polytrichum commune var. commune Polytrichum juniperinum
■
●
●
Polytrichum commune
Polytrichum formosum
●
●
y
●
Psilopilum cavifolium
CR ● VU ● NT
Psilopilum laevigatum
CR ● VU ●
●
Pterigynandrum filiforme
●
●
●
●
●
●
●
● RE RE ●
Pterigynandrum filiforme var. filiforme
□
□
■
□
□
□
■
■
Pterigynandrum filiforme var. majus
●
VU
■
□
□
VU ●
□
□
●
□
■
■
●
●
●
●
●
●
●
■
■
□
□
■
■
■
■
■
DD DD
Pterygoneurum compactum ■
■
Pterygoneurum crossidioides Pterygoneurum kozlovii
y
Pterygoneurum lamellatum
y
●
Pterygoneurum ovatum
Pterygoneurum papillosum
y
Pterygoneurum sampaianum
y
NT
EN
RE RE RE
● RE
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Pterygoneurum squamosum ●
Pterygoneurum subsessile
●
Ptilium crista‐castrensis
●
Ptychomitrium incurvum
y
Ptychomitrium nigrescens
y
●
●
●
Ptychomitrium polyphyllum
●
●
Ptychostomum archangelicum
●
●
●
●
●
●
●
y
●
●
●
●
●
●
●
●
●
●
●
●
●
y
●
Ptychostomum arcticum
Ptychostomum boreale Ptychostomum bornholmense
Ptychostomum capillare
Ptychostomum cernuum y
Ptychostomum compactum
●
●
●
● CR
RE ●
●
DD
DD
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
VU
●
●
●
●
●
DD ●
● NT NT
●
●
●
●
●
●
●
●
●
●
Ptychostomum compactum var. rutheanum
Ptychostomum creberrimum
Ptychostomum cyclophyllum
y
Ptychostomum demissum
y
Ptychostomum moravicum
Ptychostomum pallens
95
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
RE
●
CR
●
●
?
●
DD
●
DD
●
●
DD
●
●
RE EN EN
● ● ●
●
□
●
□
■ ■ ■
NT DD
□
●
■
●
EN
●
EN
● ●
●
● NT ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
EN ●
●
●
□
●
●
■
●
■
●
● DD
●
●
●
y
●
●
●
●
EN ● NT
●
●
●
●
●
●
●
●
●
●
●
● VU VU ●
●
●
●
●
●
●
NT
DD
DD
● ● ● ● ● ● ●
□
●
●
RE
●
●
□
●
●
●
●
■
●
●
EN
●
Ptychostomum donianum
Ptychostomum imbricatulum
●
●
●
Ptychostomum compactum var. compactum
Ptychostomum longisetum
●
●
●
●
●
●
●
●
■
■
●
●
●
□
□
□
●
NT
‐
●
□
●
●
VU
DD DD ●
DD
EN
‐
●
●
‐
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD
● ● ●
●
●
DD
EN ●
Vatican City
Spain
●
Sicily
●
Sardinia
●
San Marino
●
Portugal
●
□
Monaco
●
y
□
Malta
●
Polytrichastrum sexangulare
□
Madeira
●
Italy
□
●
France
■
●
Cyprus
■
●
Corsica
■
●
Canary Islands
■
●
Balearic Islands
Northern Ireland
□
●
Azores
Ireland
□
●
Andorra
Great Britain
□
●
Gibraltar
Sweden
■
Channel Islands
Svalbard
□
●
Norway
□
●
Iceland
Finland
Polytrichastrum alpinum
Faeroe Islands
Taxon
Pohlia wahlenbergii var. wahlenbergii
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
Crimea
Estonia
Kaliningrad
□
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
□
■
■
■
□
■
■
■
■
■
□
■
□
■
□
□
■
■
■
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Kosovo
●
□
?
Hungary
●
■
■
●
Ukraine
Central Russia
□
●
■
■
Sub‐polar & North Urals
Caucasus (in Europe)
□
●
●
■
■
SE Russia
Belarus
□
●
●
●
NW Russia
Arctic Russia
□
●
□
NE Russia
Turkey
□
●
●
●
Moldova
Slovenia
□
●
□
Middle and South Urals
Serbia
□
●
●
●
Lithuania
Romania
□
●
●
■
■
Latvia
Montenegro
□
●
■
●
●
Kazakhstan (in Europe)
Macedonia
□
●
□
●
■
■
□
□
●
□
●
●
□
■
EN
□
●
□
□
■
●
□
●
●
□
■
●
■
■
■
□
□
□
GE
●
□
Greece
●
□
Croatia
G
□
Crete
RE
●
□
□
Bulgaria
□
●
Bosnia‐Herzegovina
Luxembourg
□
●
Albania
Liechtenstein
■
●
Switzerland
Germany
■
●
Slovakia
Czech Republic
□
●
Poland
Belgium
■
Appendix 7 - Mosses
Netherlands
Austria
Checklist and country status of European bryophytes
●
●
●
●
●
4
●
●
●
●
●
●
●
G
●
●
●
●
●
3
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
VU NT
●
VU VU
●
●
●
●
●
NT
●
●
●
●
VU
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
□
□
□
□
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
1
●
CR
3
3
●
●
●
●
●
V
2
●
EN
2
3
●
●
G
3
●
●
●
●
●
2
DD DD*
●
●
●
R
VU
●
EN GE
E
CR VU
CR
R
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
R
●
EN
DD
●
●
●
●
●
●
●
●
●
VU EN
●
EN
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
VU
●
●
EN VU DD
●
CR EN
●
●
●
NT
●
●
●
●
●
●
CR
●
●
●
●
●
●
NT
●
NT
●
CR
●
●
●
●
●
●
4
VU
R
4
CR
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
DD* ●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
RE
●
●
●
DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
V
●
■
□
□
nm
□
VU GE
□
□
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
R
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
DD
●
●
●
●
●
VU
●
●
●
●
●
●
DD
●
●
●
●
●
DD* NT
NT
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
1
1
●
●
●
●
●
□
□
■
□
□
■
■
□
■
□
■
■
□
□
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
■
■
nm
■
●
●
●
●
●
Mn
●
●
●
●
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●
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VU DD*
●
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●
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●
●
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●
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nm
●
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●
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●
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●
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?
●
■
■
■
●
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■
CR CR
VU
CR
1
3
RE EN
●
●
3
V
RE
NT EB
●
CR
CR CR
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●
●
●
●
?
●
EN
●
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CR CR
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●
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●
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●
●
●
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●
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3
●
●
●
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3
●
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3
●
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I
●
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●
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●
●
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●
●
●
CR
NT
DD*
EN
●
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●
●
?
EN
●
?
DD
●
EN DD
●
●
NT
●
●
●
●
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●
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●
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●
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●
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4
●
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●
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CR GE
●
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●
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VU KW ●
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●
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●
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●
●
●
●
RE
EN ●
●
●
●
●
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●
●
EN VU
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
●
●
●
●
NT
●
□
□
□
NT DD* ●
□
DD
□
●
EN RE
DD VU
●
■
□
●
nm
EN D
0
●
EN
●
□
DD
DD
●
●
●
●
●
●
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●
●
●
●
●
●
DD
●
●
●
●
●
●
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●
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R
●
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DD
VU
●
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RE
0
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
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EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
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●
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●
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●
□
●
□
●
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
●
□
●
●
●
●
●
●
□
●
□
□
□
●
□
●
●
1
4
●
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
DD
DD*
●
●
VU
■
CR DD*
●
●
EN ● DD
● DD ●
R
●
□
●
●
●
●
2
●
□
●
●
□
?
□
DD EN DD
RE
●
●
●
●
EN
■
■
●
●
DD
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
96
●
Ptychostomum torquescens
●
Ptychostomum zieri
Pylaisia polyantha
□
y
Pyramidula tetragona
y
Racomitrium aciculare
■
□
□
□
● NT ●
●
●
●
●
●
RE
●
●
●
●
RE
●
●
●
RE
● VU VU
●
● NT NT
●
■
■
■
●
●
●
●
●
●
●
●
●
●
EN
●
●
VU
●
●
VU ●
●
●
●
●
●
●
DD
●
●
●
●
●
DD
● VU ●
●
●
●
●
●
●
■
■
■
■
■
●
●
●
●
●
●
●
●
●
●
VU
●
VU
NT
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● VU VU ●
●
●
●
NT
●
Racomitrium canescens subsp. canescens
□
□
■
□
■
□
■
□
□
□
■
□
□
□
●
■
●
●
●
●
●
Racomitrium ellipticum
●
●
□
Racomitrium canescens
●
●
●
□
Racomitrium aquaticum
●
●
●
□
■
Racomitrium affine
●
●
●
□
□
?
EN ●
□
■
■
●
● ● ●
● VU VU ●
●
Racomitrium elongatum
●
●
●
●
●
●
●
●
Racomitrium ericoides
●
●
●
●
●
●
●
●
●
●
●
●
Racomitrium fasciculare
●
●
●
●
●
●
●
●
●
●
●
●
●
CR
●
●
●
EN
?
Racomitrium hespericum
●
●
Racomitrium heterostichum
●
Racomitrium himalayanum
y
Racomitrium lamprocarpum
y
Racomitrium lanuginosum
Racomitrium lusitanicum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
VU
●
●
●
●
●
●
●
●
●
DD
●
y
●
VU
Racomitrium macounii
RE ●
●
●
● VU
●
●
●
●
●
●
Racomitrium macounii subsp. alpinum
■
□
■
■
■
■
■
■
■
■
■
■
■
●
●
●
■
●
■
●
■
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
■
Racomitrium macounii subsp. macounii
●
Racomitrium microcarpon
Racomitrium nivale ●
y
●
Racomitrium sudeticum
Rhabdoweisia crenulata
y
Rhabdoweisia crispata
y
●
●
●
● VU
●
Rhabdoweisia fugax
Rhamphidium purpuratum
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
● ● ●
● NT NT
●
●
● VU VU ●
Rhizomnium andrewsianum
y
CR ● EN ● NT
y
CR
Rhizomnium magnifolium
Rhizomnium pseudopunctatum
●
●
Rhizomnium punctatum
●
●
Rhodobryum ontariense
Rhodobryum roseum
●
●
●
●
●
●
●
●
●
● NT NT
● ●
EN
●
●
●
●
●
●
●
DD
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
● NT NT
y
●
●
Rhynchostegiella litorea
● RE RE
NT
DD
□
y
Rhynchostegiella tenuicaulis
y
Rhynchostegiella trichophylla
y
●
□
●
●
●
●
□
□
□
□
EN ●
●
●
●
■
CR
y
RE
●
●
●
● NT
EN ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Rhynchostegium riparioides ●
● NT ●
●
●
●
●
●
●
Rhynchostegium strongylense
y
Rhytidiadelphus loreus
Rhytidiadelphus squarrosus
EN ●
DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● NT ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
●
●
●
●
□
●
●
●
■
□
□
■
■
□
● NT ●
●
●
●
●
●
●
□
□
□
■
●
●
DD
□
●
DD
●
●
●
● NT NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Rhytidium rugosum
●
●
●
●
●
Saelania glaucescens
●
●
●
●
●
VU
NT ●
●
●
●
●
●
●
y
●
●
●
●
●
●
●
● VU ●
●
?
●
●
●
●
●
●
●
●
●
●
●
NT RE
●
●
●
●
●
●
●
●
●
●
DD
CR
Rhytidiadelphus subpinnatus
Rhytidiadelphus triquetrus
NT
●
●
●
●
●
●
DD
●
●
●
●
y
●
●
●
●
●
Rhynchostegium rotundifolium
●
●
y
●
97
●
●
●
Rhynchostegium megapolitanum
Sanionia orthothecioides
●
●
● NT NT
Rhynchostegium murale
Sanionia georgicouncinata
●
●
●
VU
Rhynchostegium alopecuroides Rhynchostegium confertum
●
VU
VU
■
Rhynchostegiella teneriffae
Rhynchostegium confusum ●
NT
●
Rhynchostegiella tenella var. tenella
●
●
Rhynchostegiella tenella var. meridionalis
Rhynchostegium arcticum
●
VU
y
Rhynchostegiella tenella
●
●
●
●
Rhynchostegiella curviseta
Rhynchostegiella macilenta
●
y
Rhizomnium gracile
Rhynchostegiella bourgaeana
VU
DD
y
Racomitrium obtusum
Racomitrium panschii
●
●
VU ●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
DD
●
VU
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
Vatican City
●
■
●
●
●
●
■
■
●
●
●
■
□
●
●
Spain
DD
Sicily
●
Sardinia
●
San Marino
●
Portugal
●
●
●
Racomitrium canescens subsp. latifolium
●
●
● VU ●
Monaco
●
●
Malta
●
●
Madeira
●
●
□
Italy
■
France
■
Cyprus
■
■
Corsica
●
Canary Islands
●
Balearic Islands
●
■
Ireland
●
■
Great Britain
●
■
Gibraltar
●
■
Channel Islands
●
■
Sweden
Svalbard
●
●
●
Pylaisia selwynii
●
Azores
Ptychostomum rubens
●
Andorra
■
●
■
Northern Ireland
Ptychostomum pseudotriquetrum var. pseudotriquetrum
Norway
●
■
Iceland
●
Ptychostomum pseudotriquetrum var. bimum
Finland
Faeroe Islands
Taxon
Ptychostomum pseudotriquetrum
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
■
D
□
□
□
■
●
●
●
●
●
●
●
●
●
Mn EN
●
0
●
●
DD G
●
●
●
□
BE
V
●
●
●
●
●
●
●
●
●
CR
1
V
●
●
●
●
● KW ●
VU
●
●
NT
●
■
●
●
●
■
■
●
■
□
■
■
●
?
CR
●
●
●
●
●
●
VU
●
●
VU
●
●
NT
●
DD
●
NT DD* ●
VU
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
CR DD
EN
●
●
●
VU
●
●
●
DD
DD
●
●
●
●
●
●
●
DD* ●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
□
■
■
■
■
■
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
CR EN
●
●
■
●
●
●
●
VU
●
●
●
●
●
●
●
Ukraine
■
■
●
Sub‐polar & North Urals
□
VU
■
SE Russia
■
●
□
●
NW Russia
■
●
■
●
NE Russia
■
●
□
Moldova
■
●
■
● DD* ●
Middle and South Urals
■
□
●
Lithuania
■
Latvia
●
□
Kazakhstan (in Europe)
●
■
Kaliningrad
●
■
Estonia
●
■
Crimea
●
■
●
●
Central Russia
●
■
■
●
Caucasus (in Europe)
●
■
Belarus
●
Slovenia
●
□
Serbia
Romania
●
■
●
●
□
●
■
Montenegro
●
Macedonia
●
Kosovo
●
■
Arctic Russia
□
●
●
Turkey
■
●
●
■
Hungary
●
Appendix 7 - Mosses
Greece
●
■
Croatia
●
Crete
Netherlands
●
Bulgaria
Luxembourg
●
D
Bosnia‐Herzegovina
Liechtenstein
nb
■
Albania
Germany
●
Switzerland
Czech Republic
●
3
Slovakia
Belgium
●
Poland
Austria
Checklist and country status of European bryophytes
VU
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
●
●
●
NT
●
●
●
●
●
CR
DD*
●
●
NT
●
●
V
●
● KW ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
□
■
nm
□
□
□
■
□
□
□
□
■
□
■
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
□
●
●
●
●
●
?
●
●
●
VU
●
●
●
●
●
●
1
□
●
●
●
□
●
●
●
●
●
■
●
● KW ●
●
●
●
DD*
3
VU KW ●
DD
●
DD
‐
●
V
EV KW ●
NT VU
●
●
●
●
● KW ●
●
●
●
●
V
●
●
●
●
NT
●
●
R
●
EN
●
VU
■
■
■
■
■
□
■
●
EN
●
V
I
●
●
●
●
●
●
●
●
●
●
BE
●
●
●
DD NE
●
●
EN
●
●
EN
●
EN
●
●
●
●
DD*
●
?
●
●
DD
●
●
●
●
●
□
■
VU VU
●
●
EN
●
DD
●
●
●
●
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
●
■
■
VU
DD*
●
●
■
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
2
●
RE EN
● NT
●
●
V
EN
●
●
●
3
V
●
V
R
●
●
●
DD
●
●
●
VU
DD EN
●
●
●
CR
●
●
●
CR
●
VU
●
●
●
●
NT
●
●
●
VU
?
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
3
●
Mn EN
●
●
3
●
KW ●
DD
●
VU
●
NT
●
● DD* ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
VU
●
DD
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
●
●
●
●
●
●
●
V
●
●
CR
●
●
●
●
NT
●
●
●
●
●
V
●
●
BE
●
●
●
3
●
●
●
●
●
CR CR
D
3
●
NT
●
●
●
●
●
NT
●
●
●
DD*
●
●
●
●
VU
□
■
■
4
●
EN
●
●
CR
D
□
□
□
□
□
□
NT
●
□
□
□
□
□
■
□
□
□
□
□
GE
E
CR VU DD* ● DD* ●
●
●
VU
●
CR
●
VU
E
EN VU DD*
●
DD
●
VU
●
●
●
●
●
●
●
●
●
4
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
D
●
●
●
●
■
●
DD
●
●
●
●
■
□
VU VU
●
●
●
●
●
0
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
V
●
●
●
●
●
●
●
V
●
●
●
●
●
DD
NT
●
●
VU NT
●
●
●
●
●
●
EN GE
●
●
●
●
●
NT
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
2
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN NT
●
●
●
●
●
●
●
●
GE
●
●
DD
●
●
●
VU
●
●
●
●
●
●
DD* EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
V
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
V
●
● KW ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
EN
2
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
EN
●
●
●
●
●
NT EN
VU
●
●
●
●
● DD*
●
●
●
●
●
●
●
●
●
1
●
●
●
●
●
●
●
●
1
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
R
●
●
98
●
●
●
●
●
●
●
●
● VU
●
●
●
●
●
●
VU
y
Schistidium agassizii
●
Schistidium apocarpum
●
Schistidium atrofuscum
Schistidium boreale
● EN
●
Schistidium brunnescens
●
Schistidium brunnescens subsp. brunnescens
□
●
●
y
Schistidium canadense y
●
●
?
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
●
●
●
●
NT ●
Schistidium crenatum
●
Schistidium dupretii
●
●
●
VU
●
●
●
●
●
●
●
●
DD DD DD ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
VU
DD
y
●
VU
●
●
● DD DD
●
●
●
□
□
■
■
□
■
■
■
■
■
VU
●
●
VU
Schistidium elegantulum subsp. wilsonii
●
●
NT ●
●
Schistidium flaccidum
Schistidium flexipile
□
■
●
Schistidium frigidum
●
●
●
●
●
NT
●
DD
Schistidium frigidum var. frigidum
■
□
□
□
■
■
□
■
Schistidium frigidum var. havaasii
●
●
●
■
●
■
●
●
■
●
●
●
DD
●
●
DD
DD
■
■
Schistidium frisvollianum
y
Schistidium grande
y
Schistidium grandirete
●
● DD
y
●
● DD
Schistidium helveticum
y
EN
Schistidium holmenianum
y
●
●
DD
DD
●
●
●
●
Schistidium lancifolium
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Schistidium maritimum subsp. maritimum
□
□
■
□
■
□
■
□
■
□
□
□
Schistidium maritimum subsp. piliferum
■
Schistidium obscurum y
Schistidium occidentale
y
Schistidium papillosum
y
■
●
y
Schistidium pulchrum
●
●
●
●
y
●
Schistidium robustum
Schistidium sibiricum y
Schistidium sinensiapocarpum
y
Schistidium sordidum
y
Schistidium spinosum
y
●
DD* ●
●
●
NT
● ●
Schistidium subjulaceum
y
●
●
●
●
●
● VU
●
VU
●
●
●
DD
●
●
DD
●
DD
●
●
●
?
●
●
●
●
●
●
●
●
DD ●
●
●
●
●
● NT NT
●
?
●
●
DD
●
VU
●
VU
DD
●
EN
■
■
●
●
●
●
●
●
□
■
■
□
■
■
Schistidium succulentum y
Schistidium tenerum
y
VU ● NT ●
Schistidium trichodon
y
NT
Schistidium trichodon var. nutans
●
●
■
Schistidium trichodon var. trichodon
y
DD DD
●
●
Schistidium submuticum subsp. arcticum
Schistidium venetum
●
●
DD
Schistidium submuticum subsp. submuticum
Schistostega pennata
?
●
●
●
●
●
y
NT
●
● VU VU
●
Schistidium submuticum
Schistidium umbrosum
■
●
●
●
●
y
●
●
●
●
Schistidium strictum
Schistidium subflaccidum ●
●
VU
●
y
■
CR
●
Schistidium poeltii
Schistidium scandicum
●
●
Schistidium platyphyllum
Schistidium rivulare
●
●
Schistidium maritimum
●
■
□
●
●
●
VU ●
●
●
●
NT
●
y
Sciuro‐hypnum curtum y
●
Sciuro‐hypnum dovrense y
EN
Sciuro‐hypnum flotowianum
y
● VU VU
●
●
□
■
■
●
●
■
NT
Schizymenium pontevedrense
99
■
●
□
Sciuro‐hypnum latifolium
●
●
Schistidium elegantulum
Sciuro‐hypnum glaciale
●
●
● DD ●
Schistidium elegantulum subsp. elegantulum
Schistidium recurvum
●
●
VU
Schistidium confusum
Schistidium pruinosum
●
EN
●
●
VU ●
●
Schistidium confertum
Schistidium crassipilum
●
●
●
?
●
■
Schistidium bryhnii
●
●
Schistidium brunnescens subsp. griseum
Schistidium echinatum ●
●
●
■
●
●
□
□
●
●
●
●
●
EN
VU ●
NT ●
●
●
DD
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
DD
VU
Vatican City
●
Spain
●
Sicily
●
Sardinia
●
●
San Marino
●
Portugal
●
Monaco
Madeira
●
Malta
Italy
●
Cyprus
France
●
Corsica
Andorra
●
●
Canary Islands
Northern Ireland
●
●
Balearic Islands
Ireland
●
●
Azores
Great Britain
●
●
Gibraltar
●
●
●
Channel Islands
●
●
Sweden
●
●
Svalbard
DD ●
●
Norway
●
●
Iceland
●
●
Finland
●
●
●
Sarmentypnum sarmentosum
Schistidium abrupticostatum ●
●
●
●
Sarmentypnum exannulatum
●
Faeroe Islands
Taxon
Sanionia uncinata
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
3
●
●
●
●
●
●
D
4
0
●
●
●
●
I
●
●
●
EN
●
?
●
●
●
DD VU DD
●
DD
●
VU
●
●
NT DD DD* ●
●
●
●
●
●
NT
●
●
●
●
VU EN VU
●
●
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
?
●
CR
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
nb
●
●
DD
●
●
NT
●
●
●
●
●
●
EN
●
●
●
●
■
■
3
■
□
□
■
□
□
■
□
□
■
■
■
□
□
□
■
■
D
■
■
■
■
■
■
NT
■
●
●
●
●
●
●
●
●
■
■
DD*
●
●
●
Ukraine
●
●
Sub‐polar & North Urals
●
●
SE Russia
●
VU
NW Russia
●
●
NE Russia
●
●
Moldova
●
●
Middle and South Urals
●
●
Lithuania
●
CR
Latvia
NT
●
Kazakhstan (in Europe)
●
●
Kaliningrad
●
●
Estonia
●
●
Crimea
●
Central Russia
●
Caucasus (in Europe)
●
Belarus
R
Arctic Russia
NT
●
Turkey
●
●
Slovenia
●
Serbia
●
Romania
●
●
Montenegro
Switzerland
●
Macedonia
Slovakia
●
Kosovo
Poland
●
CR KW ●
Hungary
Netherlands
NT
●
Greece
Luxembourg
●
3
Croatia
Liechtenstein
●
●
Crete
Germany
●
●
Bulgaria
Czech Republic
●
●
Bosnia‐Herzegovina
Belgium
●
Appendix 7 - Mosses
Albania
Austria
Checklist and country status of European bryophytes
■
CR
●
●
●
4
●
●
VU
2
●
●
●
●
EN
●
●
●
●
DD
●
●
DD*
●
● DD* ●
●
NT
●
?
DD
●
●
NT
●
●
●
EN
● DD*
●
●
●
●
●
●
●
●
NE
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
D
●
●
DD*
●
VU
● DD* NE
●
●
●
●
●
●
●
●
●
●
□
□
□
●
●
●
●
●
●
●
●
●
●
DD*
●
● DD*
●
●
●
●
●
NE
●
●
●
■
□
■
■
□
□
□
□
□
□
□
□
□
□
□
□
■
□
4
●
EN
2
●
EN
●
CR
●
●
E
DD VU DD* ●
□
NT
●
●
I
●
●
●
●
●
●
●
●
●
DD*
●
●
□
□
●
4
3
R
●
4
DD*
●
G
VU
I
● DD* DD* ● DD* ●
●
●
●
D
0
GE
●
VU
●
●
●
0
□
■
□
■
■
●
NT
●
NT
●
●
NE
●
●
●
●
●
●
●
■
●
4
●
●
●
4
●
●
●
●
●
●
G
EN
●
●
●
I
4
●
●
●
●
●
●
●
CR
GE
DD DD*
DD
DD*
DD* DD*
DD*
I
VU DD
●
DD*
■
●
●
●
EN
DD
● DD* ●
CR
●
NT
● DD* EN
●
?
●
●
●
●
●
●
●
●
●
●
●
●
?
DD*
●
●
●
NT
CR
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
R
DD*
●
G
DD*
●
●
DD*
●
NT
●
● DD*
●
●
●
●
●
■
VU
●
■
●
●
●
●
●
■
■
■
■
■
●
●
VU
●
●
■
■
□
●
V
■
D
■
nm
●
CR
I
EN
●
●
●
NT
●
●
■
□
□
■
●
■
□
4
□
DD*
□
●
□
□
3
■
●
●
●
●
●
●
?
VU
●
●
EN VU
DD*
‐
●
?
?
●
●
●
●
0
●
?
●
4
●
■
□
DD*
●
●
●
●
DD
V
●
R
●
VU RE
R
DD VU
●
VU
● DD
VU
●
●
●
●
●
●
3
1
●
●
●
●
●
●
●
●
VU
●
VU
VU
●
VU
DD*
●
●
EN
DD
●
●
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
100
●
●
●
●
●
●
●
●
●
Sciuro‐hypnum populeum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Sciuro‐hypnum starkei
Sciuro‐hypnum tromsoeense y
Scleropodium cespitans
y
NT
●
●
Scleropodium touretii
Scopelophila cataractae
y
Scopelophila ligulata
y
●
●
DD
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
DD
●
●
●
●
●
● NT NT
●
● EN EN
VU ●
●
●
●
●
●
●
●
Scorpidium revolvens
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
DD
●
●
●
DD
●
●
●
●
CR
●
●
●
Scorpiurium deflexifolium
● ● ●
● EN ●
y
●
●
VU
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
DD
EN
●
●
●
?
●
●
DD
●
●
●
●
●
●
●
●
DD
●
●
y
Seligeria brevifolia
y
Seligeria calcarea
y
Seligeria calycina
y
Seligeria campylopoda
y
Seligeria carniolica
y
Seligeria diversifolia
y
VU ●
●
●
●
VU
EN
●
VU
●
DD
EN
VU
●
NT
●
●
CR
EN
CR
●
VU
●
●
RE
VU
Seligeria donniana
NT
●
NT
●
●
●
●
VU
DD
●
●
●
●
●
DD
●
●
●
y
Seligeria irrigata
y
Seligeria oelandica
y
VU ● VU
VU VU VU
Seligeria patula
y
VU
DD NT ?
Seligeria polaris
y
● ●
VU
●
EN
●
Seligeria pusilla
RE
●
●
●
●
●
●
Seligeria recurvata
RE
●
●
●
●
●
●
●
VU
EN
VU
?
●
●
VU
●
●
●
●
●
●
●
●
●
Seligeria subimmersa
y
Seligeria trifaria
y
Seligeria tristichoides
y
Sematophyllum adnatum
y
Sematophyllum demissum
y
DD
●
●
●
●
●
VU
● VU
●
VU NT
NT VU
Sematophyllum substrumulosum
Serpoleskea confervoides Sphagnum affine
●
CR
●
●
● EN ●
●
●
●
●
● NT NT
● VU VU
■
■
●
●
●
?
●
Sphagnum affine var. affine
●
●
●
●
●
NT
NT
?
Sphagnum affine var. flagellare ■
■
■
■
■
■
■
■
■
●
●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
y
●
Sphagnum angustifolium
●
Sphagnum annulatum
Sphagnum aongstroemii
y
●
●
●
●
●
●
Sphagnum austinii
●
Sphagnum balticum
●
●
●
●
Sphagnum capillifolium
●
●
●
●
●
●
● DD DD ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● ●
● NT ●
●
●
●
●
●
●
●
●
●
●
●
DD ●
Sphagnum contortum
●
●
●
●
Sphagnum cuspidatum
●
●
●
●
●
●
●
●
●
●
●
●
Sphagnum fallax
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Sphagnum fimbriatum
●
●
●
●
●
●
●
●
●
●
●
■
●
□ □ □
● VU
□
□
●
□
●
Sphagnum centrale
y
Sphagnum compactum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
Sphagnum fimbriatum subsp. concinnum
□
●
Sphagnum flexuosum
y
Sphagnum girgensohnii
Sphagnum inundatum
□
●
■
●
●
●
●
●
●
● ● ●
● NT ●
●
●
●
●
●
Sphagnum jensenii
Sphagnum lenense
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
●
●
●
‐
●
EN
●
●
VU
●
■
●
●
●
●
●
●
● NT NT ●
● ● ●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
●
●
CR
●
●
?
●
●
?
●
y
Sphagnum lindbergii
●
Sphagnum magellanicum
●
Sphagnum majus
●
Sphagnum majus subsp. majus
Sphagnum majus subsp. norvegicum
101
●
●
■
Sphagnum fimbriatum subsp. fimbriatum
Sphagnum fuscum
●
●
●
Sphagnum auriculatum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
RE VU
●
●
●
●
●
●
EN
■
□
■
■
□
■
□
□
■
■
■
■
■
●
●
●
?
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
Monaco
‐
●
Sphagnum arcticum
Malta
●
●
●
Sphagnum angermanicum
Madeira
●
●
●
●
VU
Italy
●
?
●
Seligeria acutifolia
France
Cyprus
?
●
●
●
Seligeria galinae
Corsica
● VU ●
●
●
Seligeria austriaca
Canary Islands
●
●
‐
●
Scorpiurium sendtneri
Balearic Islands
Azores
●
●
EN
●
?
?
●
●
y
●
DD
●
VU VU
Scorpiurium circinatum
●
●
● ●
NT
Scorpidium cossonii
Scorpidium scorpioides
Andorra
Northern Ireland
Ireland
Great Britain
●
Sciuro‐hypnum plumosum
Sciuro‐hypnum reflexum
Gibraltar
Channel Islands
Sweden
Svalbard
Norway
Iceland
y
Finland
y
Sciuro‐hypnum ornellanum
Faeroe Islands
Taxon
Sciuro‐hypnum oedipodium
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
RE
0
RE
?
●
●
●
●
●
●
VU
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
VU
●
●
●
●
●
●
●
DD
●
●
●
●
●
VU
●
●
●
●
?
●
DD NT
3
3
●
EN
G
●
EN
2
RE
●
?
●
●
EN
●
●
R
4
R
●
EN
●
●
●
●
● DD*
●
RE
●
●
●
?
VU
●
RE EB
E
CR VU
?
VU
VU
●
●
EN
●
DD* DD
VU
D
●
VU DD*
VU
●
●
●
EN
EN
●
●
CR
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
DD*
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN VU
●
●
●
NE
●
●
DD* DD
●
●
●
●
●
●
EN
CR
GE
R
NT VU
NT
R
CR
0
●
●
●
●
DD
VU
DD
●
VU
●
VU
●
NT
●
EN
●
●
NT GE
R
●
R
4
DD D
●
●
●
●
●
●
●
●
●
●
●
DD* CR DD
RE
●
●
●
●
DD
●
●
VU
●
●
EN
DD* VU
●
●
●
4
●
●
●
●
●
●
DD*
●
Ukraine
●
Sub‐polar & North Urals
●
Moldova
●
SE Russia
Middle and South Urals
NW Russia
Lithuania
NE Russia
Latvia
Kaliningrad
Kazakhstan (in Europe)
Estonia
Crimea
Central Russia
Caucasus (in Europe)
●
VU
●
EN
●
●
2
●
VU
●
●
●
DD
●
G
4
●
●
●
CR EB
2
VU
●
●
●
●
DD*
●
●
●
VU
NT
●
●
●
DD*
CR
●
Belarus
DD
Arctic Russia
Slovenia
●
Turkey
Serbia
Kosovo
Hungary
Greece
Croatia
Crete
Bulgaria
Bosnia‐Herzegovina
VU
●
●
R
●
4
?
●
4
●
?
?
●
CR
2
●
2
?
?
Romania
DD DD
DD DD*
Montenegro
●
●
Appendix 7 - Mosses
Macedonia
●
Albania
●
0
Switzerland
Liechtenstein
●
Slovakia
Germany
‐
Poland
Czech Republic
●
4
Netherlands
Belgium
●
Luxembourg
Austria
Checklist and country status of European bryophytes
●
●
●
●
CR
CR VU
R
DD VU
NE
?
●
VU
●
●
●
VU
●
●
NT
●
●
●
●
VU
●
●
●
●
●
VU GE
●
●
● DD* ●
NT
●
●
?
VU
●
●
●
●
EN
●
●
VU
●
●
●
●
●
●
NT
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
V
●
R
●
●
● DD*
●
DD
VU
●
●
●
VU
RE
●
R
VU
VU
●
●
1
●
●
●
VU
●
2
■
■
■
●
●
●
●
●
●
●
DD*
EB
●
I
nm
■
■
V
GE
●
●
●
EN VU
●
●
●
●
●
●
●
VU
●
DD
● DD ●
NE
DD
●
DD*
●
●
VU
●
?
●
●
□
■
■
●
●
●
●
●
●
■
□
□
■
● DD NT
●
●
●
●
NT
DD*
●
■
VU
●
●
●
●
DD
3
●
0
●
●
RE
2
NT
●
●
EN
● DD* ●
●
●
●
●
EN
●
●
2
●
●
●
RE
3
Mn
●
D
●
2
●
●
●
NT
3
2
3
●
●
3
EV
●
●
3
●
●
●
NT
●
●
3
●
●
●
●
●
●
●
V
VU KW ●
●
●
●
KW ●
KW ●
●
CR DD*
VU VU VU
NT
VU
●
●
●
RE
●
DD
●
●
●
●
1
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
R
●
●
●
?
VU
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
VU
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
VU
VU ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
4
●
●
●
●
●
●
●
●
●
EN
VU
●
●
●
●
●
EN
●
●
EN
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
NT
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD*
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
□
●
□
●
□
●
□
●
□
●
□
●
□
●
□
●
□
●
□
●
●
●
●
●
●
●
● DD*
NT
VU VU
EN
●
NT
VU NT
●
●
●
●
●
EN
●
●
●
●
1
●
NT
●
■
□
3
3
●
3
□
●
■
●
Mn NT
●
●
●
DD
nm
V
□
2
V
V
●
□
VU
□
●
□
●
□
●
□
NT
KW V
●
VU
●
●
●
VU EB
EN ●
●
●
□
●
□
●
□
●
□
VU
‐
●
●
EN
EN
●
●
●
●
●
●
●
●
V
1
●
1
□
●
VU
●
□
●
●
●
●
EN
●
●
□
●
●
●
VU VU
●
●
●
●
●
●
●
●
●
●
●
● VU ●
VU VU EN
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
DD
●
●
●
●
●
●
●
●
VU
●
NT
1
●
●
●
●
●
V
●
●
●
VU
●
●
●
●
●
●
●
3
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
3
●
●
2
BE
●
CR
●
VU
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
■
nm
□
□
□
□
□
■
□
□
□
□
□
□
■
□
□
□
□
□
102
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● ● ● ●
● NT NT
●
●
●
●
●
y
Sphagnum riparium
y
Sphagnum rubellum
●
●
●
●
●
y
●
y
Sphagnum subfulvum
●
●
●
●
●
DD*
●
●
y
●
●
●
Sphagnum subfulvum subsp. purpureum
Sphagnum subfulvum subsp. subfulvum
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
Sphagnum subnitens subsp. subnitens
□
Sphagnum subsecundum
●
Sphagnum tenellum
●
Sphagnum teres
●
y
y
Sphagnum warnstorfii
□
●
●
●
●
●
●
● DD DD
●
●
■
■
■
■
●
●
●
●
●
●
●
●
●
y
Straminergon stramineum
Syntrichia caninervis var. abranchesii
y
●
●
●
●
●
●
●
●
VU
VU
■
■
■
■
■
■
●
●
●
●
□
□
□
■
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
VU
■
●
● NT
●
●
●
●
●
●
●
●
●
●
●
● NT NT ●
●
● VU VU ●
●
■
●
● DD*
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
●
●
EN
EN
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
VU ●
●
●
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
□
DD
?
?
VU
●
●
●
●
●
●
●
●
●
●
●
NT
●
■
■
Syntrichia caninervis var. gypsophila
■
Syntrichia echinata
y
Syntrichia fragilis
y
Syntrichia glabra
y
Syntrichia handelii
y
Syntrichia laevipila
EN ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Syntrichia montana var. calva
●
●
●
Syntrichia norvegica
●
●
□
●
●
●
■
●
□
●
●
EN
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
■
■
□
●
■
●
●
●
●
●
●
□
●
●
□
●
●
DD ●
■
□
Syntrichia papillosissima
NT RE RE ●
NT
Syntrichia princeps
Syntrichia ruralis var. ruraliformis
■
Syntrichia ruralis var. ruralis
■
●
●
●
●
●
DD* ■
□
■
■
□
□
●
●
●
●
●
●
■
■
■
■
■
■
■
■
■
■
■
■
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
□
■
□
■
■
■
□
●
●
●
□
y
Syntrichia subpapillosissima
103
■
●
●
●
●
■
□
□
■
●
■
●
●
●
●
●
●
●
●
●
●
■
DD
■
■
■
●
●
●
●
Taxiphyllum wissgrillii
■
y
Syntrichia ruralis
Taxiphyllum densifolium
VU
■
■ □
EN
VU
●
■
Syntrichia montana var. montana
●
●
DD
VU
●
●
●
VU
EN
y
●
Syntrichia virescens
■
●
●
●
Syntrichia montana
Syntrichia papillosa
■
●
●
●
Syntrichia latifolia
Syntrichia sinensis
●
RE
Syntrichia caninervis var. caninervis
Syntrichia rigescens
●
●
Syntrichia caninervis var. astrakhanica
Syntrichia minor
●
VU
●
● DD
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
■
VU
●
●
●
●
●
y
●
●
●
●
●
●
●
Vatican City
Spain
Sicily
Sardinia
San Marino
●
●
□
●
y
Portugal
●
y
Syntrichia caninervis
Monaco
●
●
●
●
●
Syntrichia calcicola
Malta
●
●
●
●
●
Stegonia latifolia
●
●
●
Splachnum sphaericum
y
●
●
■
Splachnum rubrum
Splachnum vasculosum
●
●
■
●
●
Splachnum luteum
y
●
●
●
●
y
●
●
?
●
y
Splachnum pensylvanicum
●
●
●
●
●
■
Sphagnum wulfianum
Splachnum melanocaulon
●
●
●
EN
●
Splachnum ampullaceum
●
●
● NT NT ●
■
Sphagnum troendelagicum
●
●
●
Sphagnum subnitens subsp. ferrugineum
Sphagnum tundrae
Madeira
Cyprus
●
●
●
■
■
Sphagnum subnitens
Syntrichia bogotensis
●
NT DD
●
Sphagnum strictum
Splachnobryum obtusum
●
y
Sphagnum squarrosum
●
VU
●
●
Sphagnum russowii
●
VU
●
●
Sphagnum recurvum Sphagnum skyense
●
●
y
Sphagnum quinquefarium
Sphagnum rubiginosum
Corsica
RE
Sphagnum palustre
Sphagnum pylaesii
●
●
Sphagnum papillosum
Sphagnum pulchrum
●
?
y
Sphagnum platyphyllum
Canary Islands
●
Balearic Islands
●
Azores
Northern Ireland
●
Andorra
Ireland
●
Gibraltar
Channel Islands
Sweden
Svalbard
Norway
Finland
Iceland
●
Italy
Sphagnum obtusum
Sphagnum olafii
● EN
Great Britain
●
y
France
Sphagnum nitidulum Faeroe Islands
Taxon
Sphagnum molle
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
3
I
R
BE
●
●
GE
●
●
DD
VU NT
CR ●
EN
●
●
●
●
●
●
●
●
●
●
●
NT
VU
●
●
●
DD
●
●
●
●
●
●
VU
●
●
●
●
●
●
DD
VU
●
●
●
●
Sub‐polar & North Urals
Middle and South Urals
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
2
●
●
●
●
●
●
●
●
●
●
4
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
2
●
Ukraine
●
SE Russia
●
NW Russia
●
NE Russia
2
Moldova
1
Crimea
2
●
●
CR VU VU
NT VU VU
Kazakhstan (in Europe)
Central Russia
Caucasus (in Europe)
Belarus
Arctic Russia
Turkey
Slovenia
Serbia
VU
VU
VU
EN VU
●
Lithuania
2
●
●
Latvia
●
CR RE
DD
Kaliningrad
●
●
CR
DD VU
Estonia
●
●
●
Romania
●
Montenegro
●
3
2
Macedonia
●
●
Mn CR
DD VU
Kosovo
●
Hungary
●
3
3
Appendix 7 - Mosses
Greece
●
Croatia
●
Crete
2
Bulgaria
NT
Bosnia‐Herzegovina
●
Albania
2
Switzerland
KW V
Slovakia
2
Poland
Germany
RE
Netherlands
Czech Republic
●
Luxembourg
Belgium
1
Liechtenstein
Austria
Checklist and country status of European bryophytes
VU
●
●
●
●
●
●
●
●
●
●
●
‐
VU DD*
2
Mn
●
V
BE
●
3
●
●
G
VU
●
●
?
NT
●
●
●
●
●
EN
●
●
●
NT
●
●
●
●
●
●
G
NT
●
●
●
●
●
●
●
●
CR
CR
●
●
●
●
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
VU NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
EN
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
VU
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
●
●
●
0
●
R
VU
VU
EN
●
■
□
■
2
●
●
3
NT KW ●
□
DD
■
□
● DD*
□
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●
●
DD
●
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●
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●
□
□
●
●
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●
●
2
●
EN
■
■
□
■
nm
3
●
●
3
□
●
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□
■
EN EB
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●
●
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●
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●
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●
●
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●
2
●
●
3
EV
3
●
●
3
NT KW ●
●
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●
3
●
●
2
CR
●
●
●
●
EN NT
EN
●
V
□
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●
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●
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●
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●
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●
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●
●
●
●
●
●
VU VU
●
●
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●
●
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●
●
●
●
●
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●
2
●
●
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●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
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●
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●
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EN
●
●
●
●
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DD
2
RE NT
2
RE
V
CR NT
CR NT
EN
●
●
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1
●
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2
●
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CR
●
0
●
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4
●
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G
●
●
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●
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●
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●
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●
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●
●
?
VU
●
●
●
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●
CR
NT CR EN
●
●
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●
NT
●
●
●
●
NE
●
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●
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●
●
EN
RE
1
●
RE
●
●
●
●
●
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■
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●
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CR
EN
●
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●
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■
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●
0
●
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●
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●
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3
3
●
●
DD
NT
●
●
● KW I
●
●
R
DD VU DD*
EN DD
●
VU
CR
●
●
●
●
●
●
DD
NT
●
●
●
●
●
CR
VU
●
●
●
●
VU
●
●
●
●
●
□
●
□
●
?
?
1
●
●
●
●
■
■
□
●
■
●
●
●
●
3
■
□
3
□
CR
●
●
●
VU
■
■
■
●
□
●
■
●
●
●
nm
R
●
●
■
■
■
■
?
nb
●
●
●
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●
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●
●
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●
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■
●
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●
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●
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●
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●
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NT
R
VU
●
●
●
EN
●
●
●
■
■
■
■
■
■
●
□
●
●
●
■
EN
□
●
VU
●
●
●
●
●
●
●
DD
■
■ DD* ■
■
■
■
■
□
■
●
■
CR
●
●
●
■
□
■
■
●
■
DD
■
●
□
●
□
VU
■
●
●
●
●
●
●
●
●
NT
●
●
●
●
□
●
□
●
VU
□
●
●
●
■
●
●
●
●
□
●
●
●
●
□
●
●
●
●
●
CR
●
●
●
●
●
●
●
■
DD
■
■
VU
■
■
■
■
■
■
■
●
EN
●
●
●
●
■
■
■
●
●
■
■
■
DD
■
●
●
●
●
■
■
■
■
■
■
■
■
●
●
●
●
●
●
●
●
●
●
□
□
■
■
■
■
■
■
■
■
●
●
●
D
●
●
●
●
●
?
●
●
●
●
●
●
●
GE
R
●
R
VU
●
●
●
●
●
●
●
VU
VU
●
●
●
●
●
●
VU
●
●
● DD* ●
DD*
●
EN
●
●
●
DD
●
●
●
●
●
●
●
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
104
●
●
●
●
●
●
VU
●
EN
●
●
NT
●
●
●
DD
VU
●
●
●
●
●
●
●
●
DD
Tayloria serrata
y
EN
Tayloria splachnoides
y
EN
NT
Tayloria tenuis
y
NT
●
NT
CR
●
●
●
●
●
●
●
● DD
●
Tetraplodon angustatus
y
●
●
Tetraplodon pallidus
●
●
NT ●
VU ● EN
● ● ●
●
●
●
RE
●
●
●
●
●
●
DD
●
Tetraplodon paradoxus
y
Tetraplodon urceolatus
y
Tetrastichium fontanum
y
●
VU
●
CR
Tetrastichium virens
y
●
CR
●
Tetrodontium brownianum
y
DD
●
CR
VU
Tetrodontium ovatum
y
DD
●
VU
Tetrodontium repandum
y
NT
●
●
●
●
Thamnobryum alopecurum
EN
●
●
●
CR ●
●
● NT NT
●
RE
●
●
Thamnobryum angustifolium
y
EN
Thamnobryum cataractarum
y
EN
Thamnobryum fernandesii
y
Thamnobryum maderense
y
Thamnobryum neckeroides
y
Thamnobryum rudolphianum
y
Thamnobryum subserratum y
Thuidiopsis sparsa
y
Thuidium tamariscinum
●
●
●
Timmia austriaca
Timmia bavarica
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● VU VU
●
●
●
●
●
●
●
●
●
VU ●
●
●
●
VU ●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
y
y
Tortella alpicola
y
Tortella arctica
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Tortella flavovirens var. flavovirens
■
□
□
Tortella flavovirens var. glareicola
■
●
●
●
●
DD
● ●
●
VU VU VU
DD
●
●
●
●
●
●
●
●
●
●
□
■
□
□
□
■
□
□
■
●
■
■
●
VU
●
●
■
■ NT NT
■
■ EN EN □
●
■
●
●
●
●
●
‐
●
●
NT
●
Tortella inclinata var. densa
□
■
Tortella inflexa
●
●
●
●
Tortella limbata
y
Tortella limosella
y
●
CR
●
●
●
●
●
●
●
EN
●
●
●
●
●
●
DD
●
●
●
VU
●
●
CR
●
VU
EN
●
●
●
●
●
●
■
■
□
□
●
■
●
●
●
□
■
■
●
●
■
●
●
●
●
●
■
■
■
●
Tortella inclinata var. inclinata
●
●
NT ●
Tortella humilis
■
●
■
●
●
●
●
●
●
■
■
■
■
□
□
■
□
■
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
DD
●
●
●
DD
●
●
●
●
●
■
●
RE
Tortella nitida
●
y
●
●
●
●
●
●
●
●
●
●
●
●
Tortella squarrosa RE ●
●
●
●
●
●
●
●
●
DD
?
●
DD
●
●
●
●
●
●
●
●
■
●
●
■
●
●
●
■
●
■
●
■
●
●
●
DD
●
●
NT
●
■
■
□
□
□
□
■
□
■
□
□
■
■
□
□
□
■
■
■
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
●
■
■
●
□
□
■
■
□
■
■
■
●
●
●
●
●
●
●
Tortella tortuosa var. tortuosa
□
□
■
□
□
□
Tortula acaulon ●
●
●
Tortula acaulon var. acaulon □
■
□
● NT
Tortella tortuosa var. fragilifolia
Tortula acaulon var. papillosa ■
■
■
■
Tortula acaulon var. pilifera ■
■
■
■ DD
Tortula acaulon var. retortifolia 105
●
VU
Tortella flavovirens var. papillosissima
Tortula amplexa
●
●
VU
Tortula acaulon var. schreberiana ●
●
VU
●
Tortella inclinata
●
●
DD
●
●
Tortella tortuosa
DD
●
NT VU
Tortella bambergeri
Tortella fragilis
●
● DD
Tortella flavovirens
Tortella rigens
●
●
●
Tomentypnum nitens
DD
●
●
EN
Timmiella barbuloides
Timmiella flexiseta
●
●
VU
EN ●
y
DD
●
●
RE
Timmia norvegica
y
●
●
●
●
Timmiella anomala
●
●
Thuidium recognitum
Timmia sibirica
● VU ●
●
●
y
●
●
Thuidium delicatulum
Timmia megapolitana
●
●
●
y
●
●
●
Thuidium assimile
Timmia comata
●
■
■
●
DD
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
Vatican City
Spain
Sicily
Sardinia
San Marino
Portugal
EN
●
y
Tetraplodon mnioides
●
DD
Tayloria rudolphiana
Tetraphis pellucida
Monaco
DD
Malta
●
●
Madeira
Italy
Cyprus
Corsica
Canary Islands
Balearic Islands
Azores
Andorra
●
y
Tayloria lingulata
Tetraplodon blyttii
Northern Ireland
Ireland
Great Britain
Gibraltar
Channel Islands
Sweden
Svalbard
Norway
Iceland
● VU ●
VU
France
y
Tayloria froelichiana
Tayloria hornschuchii
Finland
Faeroe Islands
Taxon
Tayloria acuminata
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
EN VU
?
R
1
2
●
EN
●
4
RE
1
●
●
●
●
SE Russia
NW Russia
NE Russia
Moldova
Middle and South Urals
Lithuania
Latvia
Kazakhstan (in Europe)
Kaliningrad
Estonia
Crimea
Central Russia
Caucasus (in Europe)
Belarus
Arctic Russia
Turkey
●
●
●
●
EN
●
●
●
●
●
4
VU
3
●
4
RE VU
2
●
●
GE
I
EN
V
CR EN
I
CR DD
●
●
●
CR
DD
●
EN
●
●
●
●
CR
DD
●
●
●
VU
●
●
●
●
●
●
VU
●
●
VU EN
VU
R
EN VU
VU EN
●
VU
●
●
●
●
EN
●
EN
●
●
●
●
●
●
3
3
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
NT
●
DD
0
●
●
●
●
●
●
●
●
4
NT
R
●
●
EN
D
●
●
●
●
CR
NT
I
●
RE
NE
?
DD
●
VU
I
●
●
●
●
●
CR VU
●
DD*
●
●
●
●
0
●
R
EN
VU
DD*
●
4
●
●
VU
●
3
Ukraine
I
VU
RE VU
3
VU EN
CR
●
Sub‐polar & North Urals
EN
Romania
Montenegro
Macedonia
Kosovo
Hungary
Greece
Croatia
Crete
Bulgaria
Bosnia‐Herzegovina
VU
R
4
Albania
Switzerland
Slovakia
Poland
Netherlands
Luxembourg
I
●
Slovenia
0
2
Appendix 7 - Mosses
Serbia
0
●
Liechtenstein
Germany
Czech Republic
Belgium
Austria
Checklist and country status of European bryophytes
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
NT
●
1
3
●
●
DD*
●
●
●
●
●
●
●
●
●
EB
●
●
●
●
●
●
●
●
●
●
V
●
●
EB
●
●
●
●
●
NT
●
●
●
●
V
●
●
EB
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
RE
2
●
NT
●
●
●
●
●
EN
●
●
●
●
●
EN
3
●
●
●
DD
●
●
●
EN
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
●
●
●
GE Ex
●
●
●
VU VU
VU
DD
?
CR
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
NT
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
CR
●
●
●
●
CR
0
●
●
●
●
●
●
EN
●
●
●
EN
●
?
3
Mn NT
2
●
CR EB
V
●
●
●
●
EN
EN
●
●
EN NT
●
●
●
●
VU
●
●
●
●
●
DD
●
R
●
●
●
●
●
●
●
●
●
CR
●
●
●
●
□
nm
□
□
□
□
□
□
□
■
□
□
□
□
□
□
□
3
RE
V
●
VU
●
●
●
●
VU DD* ●
CR
●
●
●
NT
●
●
●
●
●
NT
●
●
■
■
●
●
●
■
□
□
●
●
●
■
■
■
■
NT GE
□
R
□
●
●
■
NT
■
■
■
■
●
□
GE
RE
nm
●
DD*
●
●
●
□
●
□
●
NT
●
●
●
CR
□
□
■
□
□
● DD* ●
●
●
□
●
●
●
●
□
●
●
●
●
NT
3
●
?
●
●
■
●
●
□
□
■
□
●
●
●
nb
■
□
■
■
●
●
●
●
●
●
●
●
2
3
●
●
●
●
■
●
●
●
●
●
●
●
3
●
●
●
●
□
□
□
□
□
●
●
●
■
□
■
●
VU
□
EN
●
●
●
●
●
□
□
VU
●
●
CR
●
●
●
NT VU
●
●
●
●
●
●
●
●
●
NT
●
●
●
●
■
●
■
●
●
■
●
■
●
●
■
●
●
●
●
■
●
●
□
●
●
CR
●
●
●
●
□
VU
GE
R
●
●
●
□
□
□
□
■
■
□
■
□
□
■
■
■
□
□
□
□
□
□
□
□
□
□
□
□
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
□
■
□
□
□
■
□
■
■
□
■
■
■
■
■
■
■
■
■
□
□
□
□
□
□
□
□
□
■
■
■
■
■
■
■
■
■
■
■
●
●
●
VU
3
●
●
R
?
●
●
●
D
■
●
DD*
●
●
●
●
●
●
●
●
●
□
□
□
□
●
●
●
●
□
●
□
●
□
□
□
□
□
D
■
■
D
NT
■
□
□
■
DD*
■
■
■
106
Tortula cernua
y
NT ● ●
● ●
CR ● DD ● CR
Tortula cuneifolia
y
●
Tortula freibergii
y
Tortula guepinii
y
●
● VU VU
EN
EN CR CR
●
●
●
●
●
y
Tortula lindbergii ●
●
●
●
●
●
●
●
●
●
●
●
?
●
● DD
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
EN ●
EN
●
●
DD
●
●
●
●
●
●
●
●
EN
VU
●
EN
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
DD ● VU
●
●
●
EN
●
● CR
●
?
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
y
Tortula marginata
●
Tortula mucronifolia
EN ●
●
● NT NT
● DD
●
●
●
●
Tortula muralis
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
Tortula muralis subsp. muralis
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
□
■
● DD ●
■
●
■
●
■
●
●
●
●
DD ●
●
●
●
Tortula muralis subsp. obtusifolia Tortula pallida
■ CR
y
Tortula protobryoides ●
Tortula randii
y
Tortula revolvens
y
Tortula rhodonia y NT
EN
DD
?
●
●
y
Tortula truncata
●
●
RE
●
●
●
●
●
●
Tortula vahliana
y
Tortula viridifolia
y
Tortula vlassovii y
Tortula wilsonii
y
Trachycystis ussuriensis
y
Trematodon ambiguus
y
●
●
●
Trematodon brevicollis
y
CR ●
●
VU
Trematodon laetevirens
y
EN
EN
VU
Trematodon longicollis
y
Trematodon perssoniorum
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
y
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
● DD
VU RE
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
?
DD*
●
●
●
●
●
●
●
●
●
●
●
RE
VU
●
●
●
●
●
● EN
●
DD ●
●
Ulota calvescens
y
Ulota coarctata
y
●
●
●
Ulota crispa
Ulota curvifolia
y
Ulota drummondii
y
●
Ulota hutchinsiae
y
●
Ulota macrospora
y
Ulota phyllantha
y
Vesicularia reimersiana
y
Voitia hyperborea
y
Voitia nivalis
y
Warnstorfia fluitans
●
●
●
DD ●
● RE
●
●
●
●
●
●
●
●
●
DD
●
DD
●
●
VU
●
●
VU
●
DD
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
‐
●
VU ●
●
●
●
●
●
●
●
VU
●
●
●
NT
●
●
●
●
●
●
●
●
RE
●
●
● DD DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
VU
●
●
● ● ●
● EN
●
●
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
VU
VU
VU
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
Ulota rehmannii
●
●
VU
●
y
Ulota bruchii
●
●
●
●
●
●
●
●
●
●
●
●
●
●
RE
●
●
●
●
Warnstorfia procera
y
Warnstorfia pseudostraminea
y
●
●
Warnstorfia trichophylla
y
●
●
Warnstorfia tundrae
●
NT
●
●
●
●
●
DD ●
●
●
●
●
●
●
●
DD ●
●
●
●
●
●
DD
●
●
DD
DD
DD
●
●
●
●
●
●
NT DD
●
●
●
●
●
Weissia condensa var. armata
107
●
●
●
Weissia condensa var. condensa
DD
DD
●
●
Weissia brachycarpa
●
VU RE
Trichostomum crispulum
Weissia condensa
●
●
Trichostomum brachydontium
Triquetrella arapilensis
DD
VU
●
y
Trichostomum arcticum
●
NT ● NT
Tortula ucrainica
Trichodon cylindricus
● NT
●
?
y
Tortula subulata
Tortula systylia
● DD
●
Tortula schimperi
Tortula solmsii
● RE RE
●
●
●
●
●
●
●
●
●
●
DD*
●
DD
●
●
●
●
DD
●
●
■
□
□
□
■
■
□
□
□
□
□
□
□
■
●
●
■
□
□
■
Vatican City
San Marino
Portugal
Monaco
Madeira
Malta
Italy
●
●
●
DD
Tortula leucostoma
●
●
●
Tortula inermis
y
●
●
●
Tortula hoppeana
Tortula laureri
●
Spain
●
y
●
●
●
DD ●
● DD
Tortula israelis
●
Sicily
●
Tortula canescens
Tortula caucasica ●
Sardinia
y
y
France
Canary Islands
● NT NT
y
Tortula bolanderi
Cyprus
Balearic Islands
●
Corsica
Azores
Andorra
Northern Ireland
Ireland
Great Britain
Gibraltar
●
●
●
Tortula brevissima
Tortula lingulata
●
y
Tortula atrovirens
Tortula bogosica
Channel Islands
Sweden
Svalbard
Norway
Iceland
Finland
Faeroe Islands
Taxon
Tortula ampliretis
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
●
●
EN
D
●
CR
3
●
●
R
●
●
●
CR
●
●
CR
●
●
CR EN
●
●
DD
●
Ukraine
Sub‐polar & North Urals
SE Russia
NW Russia
NE Russia
●
DD
Moldova
●
●
●
●
●
●
●
●
CR
Middle and South Urals
●
●
●
●
Lithuania
●
●
●
Latvia
●
NT
Kazakhstan (in Europe)
●
VU
●
Kaliningrad
●
DD
●
DD
●
Estonia
●
EN
Crimea
●
●
Central Russia
●
VU
Caucasus (in Europe)
●
●
Belarus
●
●
Arctic Russia
●
Turkey
●
Slovenia
EN
EN
Serbia
●
●
Romania
●
CR
Montenegro
●
●
VU
VU
Macedonia
DD
●
Kosovo
Hungary
R
Greece
RE
Croatia
●
1
●
Crete
●
●
RE RE
●
DD*
VU
G
Appendix 7 - Mosses
Bulgaria
●
2
Bosnia‐Herzegovina
G
Albania
RE
Switzerland
0
Slovakia
3
Poland
Germany
CR
Netherlands
Czech Republic
●
Luxembourg
Belgium
2
Liechtenstein
Austria
Checklist and country status of European bryophytes
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
●
●
●
●
●
●
DD
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
4
2
4
2
3
●
●
V
CR
R
●
RE
●
●
BE
●
●
VU
VU
●
●
●
●
●
●
VU KW
●
●
CR
3
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
3
DD
●
●
VU
●
●
●
●
●
●
■
■
■
■
■
●
●
●
●
●
●
●
●
■
■
■
■
DD
●
●
●
●
●
■
VU CR DD
●
R
NT CR
■
●
●
BE
R
VU
R
DD
DD D
●
●
EN
●
●
4
●
●
●
● KW ●
●
2
●
●
●
●
●
●
●
●
●
●
?
●
●
●
●
●
DD
●
3
3
?
EN
●
●
●
●
●
●
●
■
■
■
■
■
■
■
■
EN
●
●
●
NT
●
NT
●
●
NT
●
DD
■
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
■
■
■
■
■
■
■
■
■
■
■
■
■
■
■
3
●
●
●
●
●
■
■
■
■
■
■
●
●
●
●
●
●
●
●
?
●
●
●
●
●
NT
●
●
●
3
●
3
●
●
●
●
●
VU
RE
●
●
●
●
R
●
●
●
●
●
●
●
●
●
●
CR
●
CR
●
EN
●
●
●
●
●
●
DD*
DD
●
●
EN
●
3
●
DD
R
4
VU
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
2
RE
●
DD*
●
nb
●
●
R
nm
2
Mn CR
●
●
●
●
2
GE
R
VU
4
CR
●
CR
DD* CR
CR
VU
●
●
●
●
NT
●
●
3
●
EN
●
●
●
●
●
●
●
●
●
3
●
DD
V
●
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
BE
●
VU
●
●
V
VU
●
EN
●
●
VU
●
●
VU
●
●
●
●
●
● DD*
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
DD* DD VU
●
●
●
●
●
●
●
●
VU
●
NT
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
VU
1
●
Mn CR
●
●
4
●
3
●
●
●
●
●
E
VU CR
V
NT
●
RE
0
●
RE EN
1
●
Mn
●
●
●
●
EN
●
DD
●
NT
NT
●
●
●
●
●
●
●
●
●
DD
nb
3
0
EN GE
GE
E
CR DD
E
CR VU
CR
●
R
●
●
●
●
VU
EN
●
RE
NT
●
●
●
●
●
VU
●
●
●
●
●
DD*
VU
●
●
●
●
●
●
CR
●
E
Ex RE
●
●
●
2
3
0
RE
●
●
●
EN
DD
V
EN
●
●
D
●
nb
E
●
NT DD
●
NT
●
?
●
●
VU
●
EN
●
RE
CR
●
●
V
●
3
●
●
3
●
□
□
■
nm
□
● KW ●
EN
●
●
●
●
●
●
●
●
●
NT
CR
●
●
●
VU
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
VU
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
□
□
□
□
□
■
□
□
□
□
□
□
■
●
●
●
●
●
1
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
■
108
y
Weissia perssonii
y
Weissia rostellata
y
●
Weissia rutilans
y
●
Weissia squarrosa
y
●
Weissia sterilis
y
VU
y
Zygodon conoideus
y
Zygodon conoideus var. conoideus
●
●
●
●
DD
●
●
●
■
■
□
■
□
■
■
■
■
■
■
■
■
■
■
■
■
●
■
DD
DD
●
●
●
●
●
●
●
DD
■
●
● VU VU
●
●
●
●
●
RE
NT
VU ●
●
EN
VU
● NT NT
●
VU ●
EN
NT
●
●
●
●
●
● DD*
● VU VU
●
NT
●
●
VU
●
●
●
●
DD
●
DD
●
●
□
■
□
NT ●
■
■
Zygodon conoideus var. lingulatus
●
●
●
●
●
?
●
●
VU
●
●
●
●
●
DD
●
●
VU
DD
■
■
■
□
□
□
□
□
□
●
VU
DD
●
●
●
●
●
●
CR
●
■
y
Zygodon forsteri
y
EN
●
●
●
Zygodon gracilis
y
EN
●
EN
●
●
●
●
●
●
DD
●
●
Zygodon rupestris
Zygodon sibiricus
y
Zygodon stirtonii
y
109
●
DD
Zygodon dentatus
Zygodon viridissimus
VU
●
●
EN
●
●
NT
●
●
EN
●
●
●
●
?
●
●
●
DD*
●
●
●
●
●
●
●
● VU ●
●
●
●
●
●
●
●
●
●
●
●
●
●
Vatican City
●
Monaco
●
CR
Weissia wimmeriana
Zygodon catarinoi
●
Spain
Weissia multicapsularis
VU
Sicily
●
Sardinia
Weissia longifolia
San Marino
■ ●
EN
y
■
Portugal
Weissia controversa var. densifolia
■
Malta
■ DD
■
Madeira
■
Italy
●
□
France
Corsica
●
Cyprus
Canary Islands
●
■
Azores
●
■
Andorra
●
■
Ireland
●
■
Great Britain
●
■
Gibraltar
●
■
Weissia controversa var. crispata
Weissia levieri
Northern Ireland
Channel Islands
●
■
Sweden
●
□
Svalbard
●
□
Norway
●
■
Iceland
●
□
Finland
Balearic Islands
Weissia controversa var. controversa
Faeroe Islands
Taxon
Weissia controversa
Appendix 7 - Mosses
Denmark
● Species conrmed; LC/status unknown
■ Infraspecific taxon confirmed
□ Unconrmed infraspecic taxon
? Some doubt about occurrence
‐ Literature record but later rejected
Red/italic status values with a darker fill indicate note present
Include in candidate list?
Checklist and country status of European bryophytes
Hungary
Macedonia
Montenegro
Romania
Serbia
Slovenia
Turkey
Belarus
Caucasus (in Europe)
Central Russia
Crimea
Estonia
Kaliningrad
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
●
□
□
□
□
■
■
■
□
□
□
■
□
■
■
■
■
□
■
□
□
■
■
■
■
DD
●
■
NT
■
■
NT NT
●
■
V
■
D
●
●
V
■
●
VU KW ■
●
●
●
0
●
DD
●
VU KW R
2
●
EN
V
NT
R
DD CR
1
●
VU
2
●
EN RE
R
EN VU
D
●
I
DD
DD*
●
●
●
●
●
●
●
●
●
●
●
NT
VU
■
●
■
●
CR
●
DD
●
●
●
●
●
●
●
●
●
●
●
●
●
DD
●
CR
●
●
EN
●
■
■
■
●
●
●
●
●
●
●
●
●
●
●
CR VU DD
NT
3
Ukraine
Greece
Sub‐polar & North Urals
Croatia
●
□
SE Russia
Crete
●
■
NW Russia
Bulgaria
●
□
NE Russia
Bosnia‐Herzegovina
●
■ KW ■
Moldova
Albania
Middle and South Urals
Switzerland
● KW ●
□
Lithuania
Slovakia
●
■
Latvia
Poland
nb
■
Arctic Russia
Liechtenstein
●
□
Kosovo
Germany
●
■
Netherlands
Czech Republic
●
Luxembourg
Belgium
Kazakhstan (in Europe)
Appendix 7 - Mosses
Austria
Checklist and country status of European bryophytes
●
●
?
●
VU
●
●
VU
●
●
CR
●
●
●
?
Mn
●
VU
□
nm
□
3
NT
●
4
2
●
CR
NT
●
EN
●
●
●
V
EN
V
●
VU
?
□
●
R
●
●
●
●
0
●
●
?
E
CR NT
DD
E
CR VU
E
VU
CR
DD
●
EN
●
NE
●
DD*
●
●
VU
●
●
DD
●
●
●
●
●
3
●
● DD*
●
●
EN
E
●
●
?
●
●
●
?
VU
?
●
●
●
NE
CR
3
●
●
NT
●
●
●
●
●
110
Checklist and country status of European bryophytes
Status notes (Status values italicised in red with a darker background fill)
Taxon
Acaulon fontiquerianum
Acaulon muticum
Acaulon muticum
Acaulon triquetrum
Alleniella besseri
Aloina aloides
Aloina ambigua
Aloina bifrons
Aloina obliquifolia
Aloina obliquifolia
Amphidium mougeotii
Anacamptodon splachnoides
Anacamptodon splachnoides
Anacolia webbii
Andreaea alpestris
Anomobryum bavaricum
Anomodon longifolius
Antitrichia californica
Antitrichia californica
Archidium alternifolium
Atrichum angustatum
Atrichum flavisetum
Atrichum flavisetum
Aulacomnium androgynum
Aulacomnium palustre
Barbula bolleana
Barbula bolleana
Brachydontium trichodes
Brachytheciastrum collinum
Brachythecium campestre
Brachythecium campestre
Brachythecium cirrosum
Brachythecium erythrorrhizon
Brachythecium erythrorrhizon
Brachythecium tommasinii
Brachythecium turgidum
Breidleria pratensis
Breutelia chrysocoma
Bryoerythrophyllum ferruginascens
Bryum cellulare
Bryum elegans
Bryum elegans
Bryum funckii
Bryum intermedium
Bryum intermedium
Bryum klinggraeffii
Bryum kunzei
Bryum marratii
Bryum radiculosum
Bryum schleicheri
Bryum subapiculatum
Bryum tenuisetum
Bryum turbinatum
Bryum versicolor
Bryum warneum
Callicladium haldanianum
Callicladium haldanianum
Calliergon megalophyllum
Calliergon richardsonii
Calliergon richardsonii
111
Country
SICILY
MONTENEGRO
SICILY
SPAIN
SICILY
AUSTRIA
ALBANIA
FRANCE
AUSTRIA
FRANCE
SICILY
FRANCE
SWITZERLAND
SICILY
FRANCE
FRANCE
ALBANIA
FRANCE
SICILY
AUSTRIA
BELGIUM
GB
SWEDEN
SICILY
SICILY
FRANCE
SWITZERLAND
SPNU_RUSSIA
SICILY
FRANCE
SICILY
SICILY
FRANCE
SPAIN
SICILY
FRANCE
FRANCE
FRANCE
FRANCE
SICILY
BULGARIA
SICILY
SICILY
FINLAND
SICILY
SICILY
AUSTRIA
LATVIA
ALBANIA
SICILY
SICILY
SICILY
LATVIA
FRANCE
LATVIA
FRANCE
MONTENEGRO
FRANCE
FRANCE
LATVIA
Note
● (at risk)
CR (one old record)
● (at risk)
● (healthy population - Belén Albertos, 11.2.14)
● (at risk)
DD (probably no correct reports)
● (old, no voucher)
● (at risk)
NE (formerly = A. rigida)
● (at risk)
● (at risk)
● (at risk)
RE (but refound in 2010 in one site)
● (at risk)
● (at risk)
● (at risk)
● (old, no voucher)
● (at risk)
● (at risk)
0 (recently refound)
RE?
● (NE)
● (but not recognised in Se)
● (at risk)
● (at risk)
● (at risk)
VU (erroneously recorded)
deleted
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (one locality - Belén Albertos, 11.2.14) (status??)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
VU (as B. stirtonii)
● (at risk)
● (at risk)
● ('subsp. nitidulum' regarded as EN)
● (at risk)
● (at risk)
● (doubtful taxon, although type from Austria)
● (probably disappeared)
● (old, no voucher)
● (at risk)
● (at risk)
● (at risk)
● (probably disappeared)
● (at risk)
● (probably disappeared)
● (at risk)
EN (old data)
● (at risk)
● (at risk)
● (v. rare)
Checklist and country status of European bryophytes
Taxon
Calymperes erosum
Campyliadelphus elodes
Campylidium sommerfeltii
Campylium protensum
Campylopus flexuosus
Campylopus fragilis
Campylopus oerstedianus
Campylopus pyriformis
Campylopus pyriformis
Campylostelium pitardii
Ceratodon purpureus subsp. stenocarpus
Cheilothela chloropus
Cinclidotus aquaticus
Cinclidotus danubicus
Cnestrum alpestre
Conostomum tetragonum
Cratoneuron curvicaule
Cryphaea heteromalla
Cynodontium bruntonii
Cynodontium tenellum
Cynodontium tenellum
Cyrtomnium hymenophylloides
Dendrocryphaea lamyana
Dialytrichia mucronata
Dialytrichia mucronata
Dichelyma falcatum
Dichodontium pellucidum
Dicranella cerviculata
Dicranella grevilleana
Dicranella humilis
Dicranella rufescens
Dicranella schreberiana
Dicranodontium denudatum
Dicranodontium uncinatum
Country
SICILY
LATVIA
ALBANIA
AUSTRIA
MONTENEGRO
MONTENEGRO
SICILY
SICILY
SLOVENIA
SICILY
FRANCE
FRANCE
SICILY
ITALY
FRANCE
FRANCE
SICILY
SICILY
SICILY
FRANCE
LATVIA
FRANCE
SWITZERLAND
AUSTRIA
IRELAND
FRANCE
SICILY
FRANCE
FRANCE
SICILY
SICILY
SICILY
LATVIA
SLOVENIA
Dicranoweisia cirrata
Dicranoweisia cirrata
Dicranum acutifolium
Dicranum bonjeanii
Dicranum dispersum
Dicranum majus
Dicranum polysetum
Dicranum scottianum
Didymodon bistratosus
Didymodon erosus
Didymodon johansenii
Didymodon sicculus
Didymodon validus
Diphyscium foliosum
Distichium inclinatum
Distichophyllum carinatum
Ditrichum flexicaule
Ditrichum flexicaule
Ditrichum flexicaule
Ditrichum flexicaule
Ditrichum flexicaule
Ditrichum pallidum
Ditrichum pusillum
Ditrichum subulatum
Drepanocladus aduncus
Drepanocladus polygamus
Drepanocladus sordidus
Drepanocladus turgescens
ALBANIA
SICILY
AUSTRIA
SICILY
FRANCE
FRANCE
ALBANIA
FRANCE
FRANCE
FRANCE
FRANCE
FRANCE
SWEDEN
SICILY
SICILY
SWITZERLAND
DENMARK
ESTONIA
FAEROE ISLANDS
LATVIA
NL
SICILY
SICILY
FRANCE
SICILY
SICILY
LATVIA
MONTENEGRO
Note
● (at risk)
● (v. rare)
? (old, no voucher)
● (not recognised as valid taxon in Austria)
VU (one, old data)
VU (one, old data)
● (at risk)
● (at risk)
RE (if correctly reported)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (declined - Belén Albertos, 11.2.14)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (v. rare)
● (at risk)
DD (erroneously recorded)
DD (probably wrongly reported)
VU New to Ireland since Red List 2010
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (v. rare)
Listed in Ros et al. 2013 but subsequently deleted by
Martincic (pers. comm. via B. Papp, 7.3.14).
● (old, no voucher)
● (at risk)
NE (previously included in D. brevifolium)
● (at risk; if record correct)
● (at risk)
● (at risk)
● (old, no voucher)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (but not recognised in Se)
● (at risk)
● (at risk)
RE (but refound at one site in 2005)
● (should this be gracile?)
● (should this be gracile?)
● (should this be gracile?)
● (D. gracile not published from Latvia)
KW (gracile?)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (v. rare)
VU (one, old data)
112
Checklist and country status of European bryophytes
113
Taxon
Encalypta ciliata
Country
IRELAND
Encalypta intermedia
Encalypta rhaptocarpa
Encalypta rhaptocarpa
Encalypta rhaptocarpa var. leptodon
Entodon cladorrhizans
Entodon concinnus
Entosthodon fascicularis
Entosthodon hungaricus
Entosthodon muhlenbergii
Entosthodon muhlenbergii
Entosthodon pulchellus
Ephemerum crassinervium
Ephemerum crassinervium subsp. rutheanum
Ephemerum crassinervium subsp. rutheanum
Ephemerum crassinervium subsp. sessile
Ephemerum minutissimum
Ephemerum recurvifolium
Ephemerum recurvifolium
Ephemerum serratum
Ephemerum serratum
Eucladium verticillatum var. angustifolium
Fissidens bryoides var. caespitans
Fissidens crispus
Fissidens exilis
Fissidens fontanus
Fissidens gracilifolius
Fissidens ovatifolius
Fontinalis squamosa
Funaria microstoma
Funaria microstoma
Funariella curviseta
Funariella curviseta
Gigaspermum mouretii
Grimmia anomala
Grimmia donniana
Grimmia elatior
Grimmia hartmanii
Grimmia longirostris
Grimmia mollis
Grimmia montana
Grimmia ovalis
Grimmia ramondii
Grimmia ramondii
Grimmia tergestina
Grimmia torquata
Gymnostomum aeruginosum
Gymnostomum aeruginosum var. obscurum
Gymnostomum calcareum
Gymnostomum lanceolatum
Helodium blandowii
Herzogiella striatella
Heterocladium wulfsbergii
Homalia lusitanica
Homalothecium lutescens var. fallax
Hygroamblystegium humile
Hygrohypnum eugyrium
Hygrohypnum luridum
Hygrohypnum ochraceum
Hygrohypnum polare
Hygrohypnum polare
Hylocomium splendens
AUSTRIA
ALBANIA
SICILY
AUSTRIA
AUSTRIA
SICILY
LATVIA
SICILY
AUSTRIA
IRELAND
IRELAND
MONTENEGRO
FRANCE
ITALY
SICILY
LATVIA
MONTENEGRO
SICILY
LATVIA
MONTENEGRO
AUSTRIA
SICILY
SICILY
FRANCE
SICILY
SICILY
SICILY
SLOVENIA
MONTENEGRO
SICILY
FRANCE
MONTENEGRO
SICILY
FRANCE
SICILY
SICILY
LATVIA
SICILY
FRANCE
SICILY
LATVIA
LATVIA
SICILY
SICILY
SICILY
LATVIA
AUSTRIA
LATVIA
AUSTRIA
FRANCE
FRANCE
IRELAND
SICILY
AUSTRIA
LATVIA
FRANCE
SICILY
ITALY
FRANCE
ITALY
SICILY
Note
CR (EN) - Second record for Ireland in 2011, status
would now be EN
DD (doubtfully present)
● (old, no voucher)
● (at risk)
■ (but not recognised as valid taxon in Austria)
DD (doubtfully present)
● (at risk)
● (v. rare)
● (at risk)
0 (recently refound)
RE (CR) - refound in 2012, status would now be CR
EN New to Ireland since Red List 2010
VU (one, old data)
■ (at risk)
?■ 'serratum var. rutheanum'
■ (at risk)
3? (E. Eur checklist says this sp.)
VU (one, old data)
● (at risk)
3? (Latvian Red List says this species)
VU (one, old data)
■ (not recognised as valid taxon in Austria)
■ (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
RE (if correctly reported)
VU (one, old data)
● (at risk)
● (at risk)
VU (one, old data)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (v. rare)
● (at risk)
● (at risk)
● (at risk)
● (probably disappeared)
● (v. rare)
● (at risk)
● (at risk)
● (at risk)
● (v. rare)
■ (doubtful taxon in Austria)
● (v. rare)
● (doubtful taxon)
● (at risk)
● (at risk)
NT(●)
● (at risk)
■ (if recognised, the dominant var. in Austria)
● (v. rare)
● (at risk)
● (at risk)
● (declined - Belen Albertos, 11.2.14)
● (at risk)
● (declined - Belén Albertos, 11.2.14)
● (at risk)
Checklist and country status of European bryophytes
Taxon
Hymenoloma crispulum
Hymenoloma mulahaceni
Country
SICILY
SPAIN
Hymenostylium gracillimum
Hypnum cupressiforme var. filiforme
Hypnum imponens
Hypnum jutlandicum
Hypnum jutlandicum
Hypnum pallescens
Hypnum pallescens
Hypnum uncinulatum
Hypnum vaucheri
Hypopterygium tamarisci
Isopterygiopsis pulchella
Isopterygiopsis pulchella
Kiaeria blyttii
Leptodon smithii
Leptodontium flexifolium
Leptodontium flexifolium
Lescuraea radicosa
Lescuraea saxicola
Meesia hexasticha
Meesia longiseta
Meesia uliginosa
Meesia uliginosa
Microbryum davallianum
Microbryum longipes
Microbryum starckeanum
Mielichhoferia elongata
Mielichhoferia mielichhoferiana
Mnium stellare
Molendoa hornschuchiana
Myurella julacea
Myurella tenerrima
Neckera menziesii
Nyholmiella obtusifolia
Oncophorus wahlenbergii
Oreas martiana
Orthodontium lineare
Orthodontium pellucens
Orthothecium intricatum
Orthotrichum alpestre
Orthotrichum moravicum
Orthotrichum pallens
Orthotrichum pulchellum
Orthotrichum rivulare
Orthotrichum scanicum
Orthotrichum scanicum
Orthotrichum schimperi
Orthotrichum schimperi
Orthotrichum schimperi
Oxystegus tenuirostris
Oxystegus tenuirostris
Palustriella pluristratosa
Palustriella pluristratosa
Pelekium minutulum
Philonotis caespitosa
Philonotis rigida
Plagiomnium elatum
Plagiomnium medium
Plagiopus oederianus var. alpinus
Plagiothecium cavifolium
Plagiothecium latebricola
FRANCE
AUSTRIA
FRANCE
LATVIA
SICILY
AUSTRIA
FRANCE
FRANCE
SICILY
IRELAND
LATVIA
SICILY
LATVIA
GERMANY
FRANCE
SWITZERLAND
SICILY
SICILY
LATVIA
SLOVENIA
LATVIA
MONTENEGRO
LATVIA
PORTUGAL
AUSTRIA
SICILY
SICILY
SICILY
MONTENEGRO
ALBANIA
FRANCE
PORTUGAL
SICILY
LATVIA
SLOVAKIA
SWITZERLAND
FRANCE
SICILY
SICILY
SLOVAKIA
SICILY
SICILY
SWITZERLAND
AUSTRIA
SICILY
AUSTRIA
FINLAND
GERMANY
LATVIA
SICILY
AUSTRIA
FRANCE
AUSTRIA
SICILY
FRANCE
SICILY
SICILY
AUSTRIA
SICILY
SWITZERLAND
Note
● (at risk)
● (not been evaluated but only one locality and
clearly CR - Belén Albertos 11.2.14)
● (at risk)
■ (taxon probably without any value)
● (at risk)
● (v. rare)
● (at risk)
● (should be divided into 2 spp.)
● (at risk)
● (at risk)
● (at risk)
● LC, if the same as immigrans
● (v. rare)
● (at risk)
● (v. rare)
0 (but refound)
● (at risk)
VU (erroneously recorded)
● (at risk)
● (at risk)
● (probably disappeared)
RE (if correctly reported)
● (probably disappeared)
VU (two, old)
● (probably disappeared)
VU (if correctly reported)
0 (recently refound)
● (at risk)
● (at risk)
● (at risk)
CR (one, old data)
● (1960, no voucher)
● (at risk)
● (RE)
● (at risk)
● (v. rare)
Newly recorded. CR suggested.
● (not yet found in Ch)
● (at risk)
● (at risk)
● (at risk)
● (DD suggested)
● (at risk)
● (at risk)
DD (erroneously recorded)
0 (recently refound)
● (at risk)
NE (but not hitherto recognised as a taxon)
● (but not separated from O. pumilum)
not accepted
● (v. rare)
● (at risk)
● (recently found but without taxonomic value)
● (at risk)
DD (doubtfully recorded)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
■ (dubious taxon)
● (at risk)
● (erroneously recorded)
114
Checklist and country status of European bryophytes
115
Taxon
Plagiothecium undulatum
Platyhypnidium grolleanum
Pogonatum aloides
Pogonatum dentatum
Pogonatum nanum
Pohlia andalusica
Pohlia camptotrachela
Pohlia elongata
Pohlia elongata var. acuminata
Pohlia filum
Pohlia lescuriana
Pohlia lutescens
Pohlia melanodon
Pohlia proligera
Polytrichum commune var. perigoniale
Polytrichum longisetum
Polytrichum strictum
Polytrichum swartzii
Pseudoamblystegium subtile
Pseudoamblystegium subtile
Pseudoleskeella nervosa
Pseudotaxiphyllum elegans
Pterygoneurum ovatum
Ptychomitrium nigrescens
Ptychostomum compactum
Ptychostomum creberrimum
Ptychostomum cyclophyllum
Ptychostomum donianum
Ptychostomum pallens
Ptychostomum pallens
Pyramidula tetragona
Racomitrium affine
Racomitrium elongatum
Racomitrium ericoides
Racomitrium ericoides
Racomitrium microcarpon
Racomitrium nivale
Country
SICILY
CZECH REPUBLIC
LATVIA
LATVIA
LATVIA
FRANCE
LATVIA
SICILY
AUSTRIA
LATVIA
LATVIA
SICILY
LATVIA
SICILY
SICILY
SICILY
SICILY
LATVIA
ALBANIA
SICILY
SICILY
SICILY
LATVIA
SICILY
ALBANIA
SICILY
PORTUGAL
SWITZERLAND
ALBANIA
SICILY
SICILY
SICILY
SLOVAKIA
BULGARIA
SICILY
FRANCE
AUSTRIA
Racomitrium obtusum
Racomitrium sudeticum
Rhabdoweisia crispata
Rhabdoweisia fugax
Rhabdoweisia fugax
Rhamphidium purpuratum
Rhodobryum ontariense
Rhynchostegium alopecuroides
Rhynchostegium arcticum
Rhynchostegium strongylense
Rhytidiadelphus squarrosus
Sanionia uncinata
Sarmentypnum exannulatum
Schistidium agassizii
Schistidium apocarpum
Schistidium atrofuscum
Schistidium confertum
Schistidium flaccidum
Schistidium maritimum
Schistidium obscurum
Schistidium platyphyllum
Schistidium rivulare
Sciuro-hypnum curtum
Sciuro-hypnum dovrense
FRANCE
SICILY
LATVIA
LATVIA
SICILY
PORTUGAL
LATVIA
ALBANIA
AUSTRIA
SICILY
SICILY
SICILY
SICILY
FRANCE
CORSICA
FAEROE ISLANDS
SICILY
SICILY
FRANCE
SWITZERLAND
SICILY
SICILY
BULGARIA
SWITZERLAND
Note
● (at risk)
Problematic taxon (Jiri Váňa Feb 2014)
● (v. rare)
● (v. rare)
● (v. rare)
● (at risk)
● (v. rare)
● (at risk)
■ (but not recognised as valid taxon)
● (probably disappeared)
● (probably disappeared)
● (at risk)
● (probably disappeared)
● (at risk)
■ (at risk)
● (at risk)
● (at risk)
● (v. rare)
● (old, no voucher)
● (at risk)
● (at risk)
● (at risk)
● (v. rare)
● (at risk)
● (old, no voucher)
NT (doubtful report)
EN (if correctly reported)
DD (erroneously recorded)
● (old, no voucher)
● (at risk)
● (at risk)
● (at risk)
Newly recorded. NT suggested.
All specimens are R. elongata
● (at risk)
● (at risk)
● validly combined in Racomitrium in Köckinger et
al. 2008 on p. 288 as Racomitrium nivale (Köckinger,
Bednarek-Ochyra & Ochyra) Köckinger
● (at risk)
● (at risk)
● (v. rare)
● (v. rare)
● (at risk)
RE (but refound) CR?
● (v. rare)
● (1960, no voucher)
● (present - a modification of R. murale?)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
●s.l.
●s.l.
● (at risk)
● (at risk)
● (at risk)
● (not yet recorded in Ch)
● (at risk)
● (at risk, if correctly recorded)
Deleted by Natcheva March 2014
? (not yet recorded in Ch)
Checklist and country status of European bryophytes
Taxon
Sciuro-hypnum flotowianum
Sciuro-hypnum latifolium
Sciuro-hypnum oedipodium
Sciuro-hypnum ornellanum
Sciuro-hypnum reflexum
Sciuro-hypnum starkei
Sciuro-hypnum starkei
Scleropodium cespitans
Scleropodium touretii
Scorpiurium deflexifolium
Scorpiurium sendtneri
Seligeria campylopoda
Seligeria pusilla
Seligeria recurvata
Sematophyllum substrumulosum
Sematophyllum substrumulosum
Sphagnum affine
Sphagnum affine
Sphagnum affine
Sphagnum affine
Sphagnum affine var. flagellare
Sphagnum angustifolium
Sphagnum auriculatum
Sphagnum auriculatum
Sphagnum austinii
Sphagnum austinii
Sphagnum balticum
Country
SICILY
SLOVENIA
FRANCE
FRANCE
SICILY
LATVIA
SICILY
SICILY
AUSTRIA
SICILY
SICILY
LATVIA
LATVIA
LATVIA
IRELAND
SICILY
AUSTRIA
FRANCE
LITHUANIA
SLOVAKIA
SWITZERLAND
SLOVAKIA
SICILY
SLOVAKIA
AUSTRIA
SLOVAKIA
AUSTRIA
Sphagnum contortum
Sphagnum flexuosum
SICILY
MONTENEGRO
Sphagnum inundatum
Sphagnum inundatum
Sphagnum lindbergii
Sphagnum magellanicum
AUSTRIA
SICILY
FRANCE
SICILY
Sphagnum majus
Sphagnum molle
Sphagnum platyphyllum
Sphagnum recurvum
Sphagnum riparium
Sphagnum riparium
Splachnum ampullaceum
Splachnum ampullaceum
Splachnum sphaericum
Splachnum sphaericum
Splachnum sphaericum
Syntrichia fragilis
Syntrichia fragilis
Syntrichia handelii
Syntrichia latifolia
Syntrichia norvegica
Syntrichia papillosa
FRANCE
FRANCE
FRANCE
MONTENEGRO
FRANCE
SLOVAKIA
FRANCE
ITALY
FRANCE
ITALY
LATVIA
AUSTRIA
FRANCE
SICILY
SICILY
SICILY
SICILY
Note
● (at risk)
(Grom 1969: wrong det.; Glow. 1910: locality in Italy)
● (at risk)
● (at risk)
● (at risk)
● (v. rare)
● (at risk)
● (at risk)
0 (very doubtful old report)
● (at risk)
● (at risk)
● (v. rare)
● (v. rare)
● (v. rare)
VU (NT?) - new records since 2010, status NT?
● (at risk)
1 (as part of S. imbricatum)
● (at risk)
? (old record of S. imbricatum)
?EN
■ (but NS does not know which var. occurs in Ch)
?DD
● (at risk)
?EN
1 (as part of S. imbricatum)
?EN
0 (old reports wrong but correctly determined
material found recently)
● (at risk)
"I check all papers with Sphagnum spp. from
Montenegro. In Dragicevic & Veljic (2006) there is
data - S. flexuosum from Barno jezero (Birks & Walers
1972/3). Also in the same paper, we have data for S.
recurvum. I guess that we put S. recurvum as
synonym of S. flexuosum. Why, I don't know to
explain. Therefore it is necessary to delete S.
flexuosum from the list to Montenegro and put S.
recurvum." (Papp, pers. comm 9.4.14). Both spp. Left
blank, as S. fallax is recorded for Montenegro.
3 (included in S. subsecundum)
● (at risk)
● (at risk)
RE (based on herbarium specimen revision, the only
report of Sphagnum magellanicum for Sicily is by
Bottini (1919) from Madonie Mountains, without a
precise locality. It has never been found again, even
by Raimondo & Dia (1978), who led field research
● (at risk)
● (at risk)
● (at risk)
See S. flexuosum
● (at risk)
?VU
● (at risk)
● (declined - Belén Albertos, 11.2.14)
● (at risk)
● (declined - Belén Albertos, 11.2.14)
● (v. rare)
0 (recently refound)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
● (at risk)
116
Checklist and country status of European bryophytes
117
Taxon
Syntrichia princeps
Tetraplodon mnioides
Tetrodontium brownianum
Thamnobryum subserratum
Timmiella barbuloides
Timmiella barbuloides
Timmiella flexiseta
Tomentypnum nitens
Tortella alpicola
Tortella bambergeri
Tortella tortuosa var. fragilifolia
Country
IRELAND
FRANCE
FINLAND
LATVIA
FRANCE
SWITZERLAND
SICILY
PORTUGAL
FRANCE
FRANCE
AUSTRIA
Tortula bolanderi
Tortula brevissima
Tortula cernua
Tortula cernua
Tortula hoppeana
Tortula pallida
Tortula pallida
Tortula randii
Tortula revolvens
Tortula rhodonia
Tortula solmsii
Tortula subulata
Tortula wilsonii
Trematodon ambiguus
Trematodon ambiguus
Trematodon longicollis
Trichodon cylindricus
Ulota bruchii
Ulota coarctata
Ulota crispa
Ulota crispa
Ulota curvifolia
Ulota rehmannii
Warnstorfia pseudostraminea
Warnstorfia pseudostraminea
Warnstorfia trichophylla
Warnstorfia trichophylla
Warnstorfia tundrae
Weissia brachycarpa
Weissia controversa
Weissia levieri
Zygodon forsteri
Zygodon sibiricus
Zygodon sibiricus
SICILY
SICILY
LATVIA
SLOVAKIA
SICILY
FRANCE
SICILY
LATVIA
SICILY
SWITZERLAND
SICILY
SICILY
FAEROE ISLANDS
FRANCE
LATVIA
SICILY
SICILY
LATVIA
LATVIA
ALBANIA
SICILY
ALBANIA
FRANCE
AUSTRIA
SLOVENIA
AUSTRIA
LATVIA
LATVIA
LATVIA
LATVIA
SICILY
MONTENEGRO
NL
SWEDEN
Note
RE (CR) - refound 2012, status would now be CR
● (at risk)
DD (= T. ovatum in Fi)
● Abolina et al. 2011
● (at risk)
?DD (erroneously recorded)
● (at risk)
VU (if correctly reported)
● (at risk)
● (at risk)
Not recognised in Austria; could be an erroneous
combination as all Tortella tortuosa can have fragile
leaves
● (at risk)
● (at risk)
● (v. rare)
Newly recorded. CR suggested.
● (at risk)
● (at risk)
● (at risk)
● (probably disappeared)
● (at risk)
DD (probably never occurred in Ch)
● (at risk)
● (at risk)
● (sic)
● (at risk)
● (v. rare)
● (at risk)
● (at risk)
● (v. rare)
● (v. rare)
● (old, no voucher)
● (at risk)
● (old, no voucher)
● (at risk)
0 (recently refound)
RE (if correctly reported)
DD (doubtful for Austria)
● (v. rare)
● (v. rare)
● (v. rare)
● (v. rare)
● (at risk)
EN (if report correct)
● (where did I receive this record from??
● (where did I receive this record from??)
Checklist and country status of European bryophytes
Appendix 8. Taxon references for additions and nomenclatural
changes to the European moss checklist of Hill et al. 2006
Genus
Specific/Subspecific
epithets
Authority
References for additions and changes
to Hill et al. 2006)
Alleniella
besseri
Olsson et al. 2011
Alleniella
complanata
Andreaea
flexuosa
(Lobarz.) S.Olsson, Enroth &
D.Quandt
(Hedw.) S.Olsson, Enroth &
D.Quandt
R.Br. bis
Anomobryum
bavaricum
Atrichum
Barbula
androgynum
amplexifolia
Barbula
Bartramia
Bartramia
consanguinea
laevisphaera
rosamrosiae
Brachythecium
Braunia
Bryum
Bryum
Bryum
Bryum
buchananii
imberbis
austriacum
muehlenbeckii
sibiricum
versicolor
(Thwaites & Mitt.) A.Jaeger
(Taylor) Müll.Hal.
Damayanti, J.Muñoz, J.-P.Frahm &
D.Quandt
(Hook.) A.Jaeger
(Sm.) N.Dalton & D.G. Long
Köckinger, Holyoak & Suanjak
Bruch & Schimp.
Lindb. & Arnell
A.Braun ex Bruch & Schimp.
Campylidium
Campylidium
Campylopus
Clasmatodon
Dialytrichia
Dicranum
Didymodon
Didymodon
calcareum
sommerfeltii
subporodictyon
parvulus
saxicola
septentrionale
eckeliae
lamyanus
(Crundw. & Nyholm) Ochyra
(Myrin) Ochyra
(Broth.) B.H.Allen & Ireland
(Hampe) Sull.
(Lamy) M.J.Cano
Tubanova & Ignatova
R.H.Zander
(Schimp.) Thér.
Didymodon
Didymodon
Drepanocladus
Drepanocladus
Drepanocladus
maschalogenus
validus
angustifolius
brevifolius
lycopodioides
(Renauld & Cardot) Broth.
Limpr.
(Hedenäs) Hedenäs & C.Rosborg
(Lindb.) Warnst.
(Brid.) Warnst.
Drepanocladus
trifarius
Drepanocladus
turgescens
(F.Weber & D.Mohr) Broth. ex
Paris
(T.Jensen) Broth.
Encalypta
Entosthodon
Entosthodon
Entosthodon
Entosthodon
Entosthodon
Entosthodon
Entosthodon
Ephemerum
Ephemerum
pilifera
abramovae
commutatus
dagestanicus
duriaei
handelii
kroonkurk
stenophyllus
crassinervium
crassinervium subsp.
rutheanum
crassinervium subsp. sessile
Ephemerum
(Warnst. in Hamm.) Holyoak &
Köckinger
(Müll.Hal.) A.Jaeger
(Mitt.) A.Jaeger
Funck
Fedosov & Ignatova
Durieu & Mont.
Fedosov & Ignatova
Mont.
(Schiffn.) Laz.
Dirkse & Brugués
Fedosov & Ignatova
(Schwägr.) Hampe
(Schimp. in Ruthe) Holyoak
(Bruch) Holyoak
Olsson et al. 2011
[subsp. luisieri Sérgio & Sim-Sim];
Sérgio & Sim-Sim 2012
Holyoak, D.T. & Köckinger, H. 2010;
Ros et al. 2013
Sérgio et al. 2010; Ros et al. 2013
Köckinger, H. & Kučera, J. 2007; Ros et
al. 2013
Köckinger et al. 2012; Ros et al. 2013
Damayanti et al. 2012
Damayanti et al. 2012
Ignatov & Milyutina 2010
Dalton et al. 2012; Ros et al. 2013
Köckinger et al. 2013
[=Imbribryum alpinum]
Ignatov et al. 2010
[=Bryum dichotomum? Not included by
Hill et al. (2006) but retained for now
on the recommendation of
representatives of several countries,
pending further molecular work at
RBGE]
Ochyra et al. 2003; Ros et al. 2013
Ochyra et al. 2003; Ros et al. 2013
Allen & Ireland 2002
Müller 2007
Cano 2007; Ros et al. 2013
Tubanova et al. 2010
Puche et al. 2006; Ros et al 2013
Considered by Afonina et al. (2010) to
be synonymous with D. brachyphyllus
Köckinger & van Melick 2007
Jiménez 2006; Ros et al. 2013
Hedenäs & Rosborg 2008
Hedenäs & Rosborg 2008
Hedenäs & Rosborg 2008; Ros et al.
2013
Hedenäs & Rosborg 2008; Ros et al.
2013
Hedenäs & Rosborg 2008; Ros et al.
2013
Fedosov 2012
Fedosov et al. 2010
Brugués & Sérgio 2010; Ros et al. 2013
Fedosov et al. 2010
Ros et al. 2013
Ros et al. 2013
Ros et al. 2013
Fedosov et al. 2010
Holyoak 2010; Ros et al. 2013
[= E. hibernicum] Holyoak 2010; Ros et
al. 2013
Holyoak 2010; Ros et al. 2013
118
Checklist and country status of European bryophytes
119
Genus
Specific/Subspecific
epithets
Authority
References for additions and changes
to Hill et al. 2006)
Exsertotheca
baetica
Guerra et al. 2010; Olsson et al. 2011
Exsertotheca
crispa
Exsertotheca
intermedia
Funaria
Grimmia
Grimmia
Grimmia
Gymnostomum
Hedenasiastrum
Hymenoloma
Hymenoloma
Hymenoloma
Hymenostylium
anomala
horrida
laevigata
meridionalis
calcareum var. atlanticum
percurrens
compactum
crispulum
mulahaceni
gracillimum
Hymenostylium
Hypnum
xerophilum
aemulans
González-Mancebo, O.Werner,
J.Patiño & Ros
(Hedw.) S.Olsson, Enroth &
D.Quandt
(Brid.) S.Olsson, Enroth &
D.Quandt
Jur.
Muñoz & H.Hespanhol
(Brid.) Brid.
(Müll.Hall.) E.Maier
Sérgio
(Hedenäs) Ignatov & Vanderp.
(Schleich. ex Schwägr.) Ochyra
(Hedw.) Ochyra
(Höh.) Ochyra
(Nees & Hornsch.) Köckinger &
Kučera
Köckinger & Kučera
Breidl.
Imbribryum
Imbribryum
Isothecium
alpinum
mildeanum
prolixum
Leptodon
corsicus
Leptodon
Leptodontium
Lescuraea
Lescuraea
Lescuraea
longisetus
proliferum
incurvata
patens
plicata
Lescuraea
Lescuraea
Lindbergia
Lindbergia
radicosa
saviana
dagestanica
grandiretis
Microeurhynchium
Nogopterium
Nyholmiella
pumilum
gracile
gymnostoma
Nyholmiella
Oncophorus
obtusifolia
dendrophilus
Orthotrichum
Orthotrichum
Orthotrichum
Orthotrichum
Orthotrichum
Orthotrichum
Oxystegus
affine var. bohemicum
cambrense
consobrinum
dagestanicum
holmenii
moravicum
daldinianus
Oxystegus
Oxystegus
Oxystegus
Philonotis
minor
recurvifolius
tenuirostris var. holtii
capillaris
Plášek & Sawicki
Bosanquet & F.Lara
Cardot
Fedosov & Ignatova
Lewinsky-Haapasaari
Plášek & Sawicki
(De Not.) Köckinger, O.Werner &
Ros
Köckinger, O.Werner & Ros
(Taylor) R. H. Zander
(Braithw.) A. J. E. Sm.
Lindb.
Philonotis
falcata
(Hook.) Mitt.
(Huds. ex With.) N.Pedersen
(Jur.) J.R.Spence
(Mitt.) M.Stech, Sim-Sim, Tangney
& D.Quandt
Enroth. Sotiaux, D.Quandt &
Vanderp.
(Mont.) Enroth
Herzog
(Hedw.) E.Lawton
Lindb.
(Schleich. ex F.Weber & D.Mohr)
Broth.
(Mitt.) Mönk.
(De Not.) E.Lawton
Ignatova & Ignatov
(Lindb. ex Broth.) Ignatov &
Ignatova
(Wilson) Ignatov & Vanderp.
(Hedw.) Crosby & W.R.Buck
(Bruch ex Brid.) Holmen &
Warncke
(Brid.) Holmen & Warncke
Hedd. & Blockeel
Olsson et al. 2011
Olsson et al. 2011
Ros et al. 2013
Muñoz et al. 2009; Ros et al. 2013
Muñoz et al. 2009; Ros et al. 2013
Maier 2010; Ros et al. 2013
Sérgio 2006
Aigoin et al. 2009; Ros et al. 2013
Werner et al. 2013
Werner et al. 2013
Werner et al. 2013
Köckinger & Kucera 2011
Köckinger & Kucera 2011
Species described from Austria
omitted by Hill et al. (2006) but
inserted into list on the advice of
Harald Zechmeister, Heribert
Köckinger and Christian Schröck
(pers. comm. January 2013)
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Stech et al. 2008; Ros et al. 2013
Sotiaux et al. 2009; Ros et al. 2013
Olsson et al. 2011; Ros et al. 2013
Porley & Edwards 2010
Ros et al. 2013
Ros et al. 2013
Ros et al. 2013
Ros et al. 2013
Ros et al. 2013
Ignatova et al. 2010
Ignatova et al. 2010
Aigoin et al. 2009; Ros et al. 2013
Crosby & Buck 2011
Goffinet et al. 2004
Goffinet et al. 2004
Hedderson & Blockeel 2006; Ros et al.
2013
Plášek et al. 2011
Bosanquet & Lara 2012
Lara et al. 2009; Ros et al. 2013
Fedosov & Ignatova 2010
Fedosov et al. 2010
Plášek et al. 2009
Köckinger et al. 2010
Köckinger et al. 2010
Köckinger et al. 2010; Ros et al. 2013
Köckinger et al. 2010; Ros et al. 2013
Koponen & Isoviita 2010; Ros et al.
2013
Ignatov et al. 2010
Checklist and country status of European bryophytes
Genus
Specific/Subspecific
epithets
Authority
References for additions and changes
to Hill et al. 2006)
Physcomitridium
Pohlia
Polytrichum
readeri
beringiensis
commune var. commune
(Müll.Hal.) G.Roth
A.J.Shaw
Polytrichum
commune var. perigoniale
(Michx.) Hampe
Polytrichum
Polytrichum
Polytrichum
Pottiopsis
formosum
longisetum
pallidisetum
caespitosa
Hedw.
Sw. ex Brid.
Funck
(Brid.) Blockeel & A.J.E.Sm.
Pseudoamblystegium
Pseudocampylium
Pseudorhynchostegiella
Pseudosymblepharis
Pterygoneurum
subtile
radicale
duriaei
bombayensis
compactum
(Hedw.) Vanderp. & Hedenäs
(P. Beauv.) Vanderp. & Hedenäs
(Mont.) Ignatov & Vanderp.
(Müll.Hal.) P.Sollman
M.J.Cano, J.Guerra & Ros
Pterygoneurum
crossidioides
W.Frey, Herrnst. & Kürschner
Pterygoneurum
squamosum
Segarra & Kürschner
Ptychostomum
Ptychostomum
archangelicum
arcticum
Ptychostomum
boreale
Ptychostomum
bornholmense
Ptychostomum
Ptychostomum
capillare
cernuum
(Bruch & Schimp.) J.R.Spence
(R.Br.) J.R.Spence ex Holyoak &
N.Pedersen
(F.Weber & D.Mohr) Ochyra &
Bednarek-Ochyra
(Wink. & R.Ruthe) Holyoak &
N.Pedersen
(Hedw.) Holyoak & N.Pedersen
(Hedw.) Hornsch.
Hooper et al. 2010; Ellis et al. 2013
Ignatov et al. 2006
Included as P. uliginosum (Wallr.)
Schriebl by Hill et al. 2006, but in the
opinion of Hill et al. 2008, the
taxonomy and nomenclature of P.
commune Hedw. are still not clear, so
the approach taken by the latter is
retained
Included as P. commune Hedw. by Hill
et al. 2006, but in the opinion of Hill et
al. 2008, the taxonomy and
nomenclature of P. commune Hedw.
are still not clear, so the approach
taken by the latter is retained
Bell & Hyvönen 2010
Bell & Hyvönen 2010
Bell & Hyvönen 2010
This taxon includes Trichostomum
triumphans & Weissia tyrrhena,
according to Ros & Werner 2007
Vanderpoorten & Hedenäs 2009
Vanderpoorten & Hedenäs 2009
Aigoin et al. 2009
Ignatova et al. 2012
Synonymised with P. ovatum in
Spanish Flora (Guerra 2004–2006)
Synonymised with P. ovatum in
Spanish Flora (Guerra 2004–2006)
Synonymised with P. ovatum in
Spanish Flora (Guerra 2004–2006)
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
Ptychostomum
compactum
compactum var. compactum
compactum var. rutheanum
creberrimum
cyclophyllum
demissum
donianum
imbricatulum
longisetum
moravicum
pallens
pseudotriquetrum
pseudotriquetrum var.
bimum
pseudotriquetrum var.
pseudotriquetrum
rubens
torquescens
Ptychostomum
Racomitrium
zieri
nivale
Ptychostomum
Hornsch.
(Warnst.) Holyoak & N.Pedersen
(Taylor) J.R.Spence & H.P.Ramsay
(Schwägr.) J.R.Spence
(Hook.) Holyoak & N.Pedersen
(Grev.) Holyoak & N.Pedersen
(Müll.Hal.) Holyoak & N.Pedersen
(Blandow ex Schwägr.) J.R.Spence
(Podp.) Ros & Mazimpaka
(Sw.) J.R.Spence
(Hedw.) J.R.Spence & H.P.Ramsay
(Schreb.) Holyoak & N.Pedersen
[=Bryum pallescens] Holyoak &
Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
[=Bryum uliginosum] Holyoak &
Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
(Mitt.) Holyoak & N.Pedersen
(Bruch & Schimp.) Ros &
Mazimpaka
(Hedw.) Holyoak & N.Pedersen
(Köckinger, Bednarek-Ochyra &
Ochyra) Köckinger
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
Holyoak & Pedersen 2007
[=Bucklandiella nivalis] Köckinger et al.
2007; Köckinger 2008
120
Checklist and country status of European bryophytes
121
Genus
Specific/Subspecific
epithets
Authority
References for additions and changes
to Hill et al. 2006)
Rhynchostegium
alopecuroides
(Brid.) A.J.E.Sm.
Rhynchostegium
confusum
Rhynchostegium
riparioides
K.Cezón, J.Muñoz, Hedenäs &
Huttunen
(Hedw.) Cardot
Huttunen & Ignatov 2010; Ros et al.
2013
Cezón et al. 2010; Ros et al. 2013
Sarmentypnum
Sarmentypnum
Schistidium
Schistidium
Schistidium
Schistidium
Schistidium
Schistidium
Schistidium
Sciuro-hypnum
exannulatum
sarmentosum
abrupticostatum
canadense
echinatum
obscurum
sibiricum
subflaccidum
succulentum
curtum
(Schimp.) Hedenäs
(Wahlenb.) Tuom. & T.J.Kop.
(Bryhn) Ignatova & H.H.Blom
(Dupr.) Ignatova & H.H.Blom
Ignatova & H.H.Blom
H.H.Blom, Köckinger & Ignatova
Ignatova & H.H.Blom
(Kindb.) H.H.Blom
Ignatova & H.H.Blom
(Lindb.) Ignatov
Sciuro-hypnum
Sciuro-hypnum
dovrense
tromsoeense
Serpoleskea
confervoides
(Limpr.) Draper & Hedenäs
(Kaurin & Arnell) Draper &
Hedenäs
(Brid.) Schimp.
Sphagnum
affine var. flagellare
Sphagnum
Sphagnum
nitidulum
recurvum
(Schlieph. ex Röll) L.Söderstr. &
Hedenäs
Warnst.
P.Beauv.
Thamnobryum
subserratum
(Hook. ex Harv.) Nog. & Z.Iwats.
Tortella
Tortella
arctica
squarrosa
(Arnold) Crundw. & Nyholm
(Brid.) Limpr.
Tortula
acaulon
(With.) R.H.Zander
Tortula
acaulon var. acaulon
Tortula
acaulon var. papillosa
(Lindb.) R.H.Zander
Tortula
acaulon var. pilifera
(Hedw.) R.H.Zander
Tortula
acaulon var. retortifolia
(J.Guerra & Ros) R.H.Zander
Tortula
acaulon var. schreberiana
(Dicks.) R.H.Zander
Tortula
caucasica
Broth.
Tortula
lindbergii
Broth.
Tortula
Tortula
muralis subsp. obtusifolia
protobryoides
(Schwägr.) Culm.
R.H.Zander
Tortula
Zygodon
vlassovii
catarinoi
(Laz.) Ros & Herrnst.
C. Garcia, F. Lara, Sérgio & SimSim
Werner et al. 2007; Huttunen &
Ignatov 2010; Ros et al. 2013
Hedenäs 2006
Hedenäs 2006
Ignatova et al. 2009
Ignatova et al. 2009
Ignatova et al. 2009
Ignatova et al. 2009
Ignatova et al. 2009
Blom et al. 2006
Ignatova et al. 2009
Ignatov & Milyutina 2007; Ros et al.
2013
Draper & Hedenäs 2009
Draper & Hedenäs 2008; Ros et al. 2013
Vanderpoorten & Hedenäs 2009; Ros
et al. 2013
Séneca & Söderström 2009
Séneca & Söderström 2009
Séneca & Söderström 2009; Ros et al.
2013
Ignatova & Ignatov 2011; Abolina
2011; Köckinger et al. 2008
Ellis et al. 2011
Werner et al. 2005; Grundmann et al.
2006; Ros et al. 2013
Zander 1993; Werner et al. 2002, 2004;
Ros et al. 2013
Zander 1993; Werner et al. 2002, 2004;
Ros et al. 2013
Zander 1993; Werner et al. 2002, 2004;
Ros et al. 2013
Zander 1993; Werner et al. 2002, 2004;
Ros et al. 2013
Zander 1993; Werner et al. 2002, 2004;
Ros et al. 2013
Zander 1993; Werner et al. 2002, 2004;
Ros et al. 2013
[=T. modica] Ros et al. 2008; Ros &
Herrnstadt 2010; Ros et al. 2013
[=T. lanceola] Ros et al. 2008; Ros et al.
2013
Košnar, J. & Kolář 2009; Ros et al. 2013
Zander 1993; Werner et al. 2002, 2004;
Ros et al. 2013
Ros & Herrnstadt 2010; Ros et al. 2013
Garcia et al. 2006
Checklist and country status of European bryophytes
References for additions and nomenclatural changes to the moss checklist of Hill et al. 2006
Abolina, A.A. 2011. New moss records from Republic of Adygeya. 3. Arctoa 20: 256.
Afonina, O.M., Czernyadjeva, I.V., Ignatova, E.A. & Kučera, J. 2010. Five species of Didymodon (Pottiaceae,
Bryophyta) new for Russia. Arctoa 19: 51–62.
Aigoin, D.A., Huttunen, S., Ignatov, M.S. Dirkse, G.M. & Vanderpoorten, A. 2009. Rhynchostegiella
(Brachytheciaceae): molecular re-circumscription of a convenient taxonomic repository. Journal of Bryology 31:
213–221.
Allen, B.H. & Ireland, R.R. 2002. The taxonomic status of Dicranodontium subporodictyon in western North
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