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“Callianassa” aff. “Callianassa” sismondae A. MilneEdwards, 1860 “Callianassa”almerai Müller, 1993 Galathea. weinfuerteri Backmeyer, 1950 Petrolisthes haydni Müller, 1984b Pisidia viai Müller, 1984 Pisidia? subnodosa sp. nov. Dardarnus hungaricus LĘrenthey, 1929 Anapagurus miocenicus Müller, 1977 Pagurus retznensis sp nov. Dromia evae sp. nov. Kerepesia viai Müller, 1976 Kromtitis koberi (Bachmayer &Tollman, 1954) Calappa praelata LĘrenthey 1929 Lobocarcinus sp. aff. L. sismondai (von Meyer, 1843) Carpilius antiquus Glaessner, 1928 Daira speciosa (Reuss, 1871) Hyastenus sp. Majidae gen. et sp. indet. Glabropilumnus nitidus sp. nov. Pilumnus aff mediterraneus (LĘrenthey, 1877) Portunus muelleri sp. nov. Liocarcinus aff rakosensis LĘrenthey, 1929 Charybdis fragilis (Müller, 1984b) Eomaldivia friebei sp. nov. Panopeus wroni Müller, 1984b Chlorodiella juglans Müller 1984b Chlorodiella loczyi Müller, 1984b Chlorodiella mediterranea (LĘrenthey, 1929) Chlorodiella tetenyensis Müller, 1984b Pilodius vulgaris (Glaessner, 1928) Xantho moldavicus (Yanakevich, 1977) Haydnella steiningeri Müller, 1984b Xantho moldavicus (Yanakevich, 1977) A Messinian, Italy; P Messinian, Spain; G Messinian Greece; R Badenian, Russia; 6 ? + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + Other localities Units Poland Retznei Hungary Crabs from the Badenian of Retznei, Austria, with distribution of firm and/or tentative recordings, and occurrences elsewhere. Austria – other localities Table 1. + + + + + + + + + + + + I U OGP + + O + + + + + + + + O NROU + A O + + + + + + + + + + + + + + + + + + + + + N Messinian, Oran; I Oligocene, Italy; + + + + + + + + + + + + U U O O AOU AO AOU O Langhian, Spain; U Badenian, Ukraine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