Journal of Hydrology 237 (2000) 74–85 www.elsevier.com/locate/jhydrol Simple equations to represent the volume–area–depth relations of shallow wetlands in small topographic depressions M. Hayashi a,*, G. van der Kamp b a Department of Geology and Geophysics, University of Calgary, Calgary, Alberta, Canada T2N 1N4 b National Water Research Institute, Saskatoon, Saskatchewan, Canada S7N 3H5 Received 28 September 1999; revised 5 April 2000; accepted 14 July 2000 Abstract Small topographic depressions have important functions in hydrology and ecology because they store water in the form of shallow lakes, wetlands or ephemeral ponds. The relations between the area A, the volume V, and the depth h of water in depressions are important for evaluating water and dissolved-mass balances of the system. The A–h and V–h relations are usually determined from fine-resolution elevation maps based on detailed survey data. Simple equations are presented in this paper, which can be used to: (1) interpolate A–h and V–h data points obtained by a detailed survey; (2) approximate unknown A–h and V–h relations of a depression from a minimal set of field data without a time-consuming elevation survey; and (3) serve as a geometric model of depressions in simulation studies. The equations are simple power functions having two constants. The first constant s is related to the size of the depression, and the second constant p is related to the geometry of the depression. The power functions adequately represent A–h and V–h relations of all 27 wetlands and ephemeral ponds examined in this paper, which are situated in the northern prairie region of North America. Assuming that the power functions are applicable for other similar topographic depressions, an observer only needs to measure A and h twice to determine the two constants in the equation. The equations will be useful in field studies requiring approximate A–h and V–h relations and in theoretical and modeling studies. 䉷 2000 Elsevier Science B.V. All rights reserved. Keywords: Wetlands; Ephemeral lakes; Ponds; Water balance; Bathymetry; Water storage 1. Introduction Topographic depressions that hold water in the form of small lakes, wetlands or ephemeral ponds have important hydrological and ecological functions. They store snowmelt and storm water to attenuate flood peaks, and provide habitats for birds and animals that are dependent on aquatic plants and invertebrates. To study these functions hydrologists need to evaluate * Corresponding author. Tel: ⫹1-403-220-2794; fax: ⫹1-403284-0074. E-mail address: [email protected] (M. Hayashi). water balance and dissolved-mass balance in the depressions. For example, after a runoff event the flux of water and nutrients into a wetland can be estimated from the change of water volume in the wetland and the change of concentration of dissolved species. A practical approach for determining water volume V and area A is to measure the depth of water (h) and estimate A and V from predetermined area–depth (A– h) and volume–depth (V–h) relations. These relations are specific to each depression, and are usually derived from a detailed bathymetry map. Because of this site-specific nature, most hydrological research articles report A–h and V–h relations merely as a 0022-1694/00/$ - see front matter 䉷 2000 Elsevier Science B.V. All rights reserved. PII: S0022-1694(00 )00 300-0 M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 75 and one can be derived from another. For example, suppose that the water level in a lake rises by a small amount Dh. The resulting volume change DV in the lake is equal to ADh. Therefore, V at any h is given by V h Fig. 1. Slope profile of symmetric basins with y=y0 r=r0 p : For example, p 2 indicates a parabolic slope. tool for mass balance calculations, and rarely treat them as a significant topic of study. However, it is useful to look for common factors in the A–h and V–h relations for a variety of topographic depressions because this can lead to a more general understanding of their shapes and storage characteristics. Generalized forms of V–h and A–h relations have been used by some investigators in the mathematical modeling of lakes. For example, Gates and Diessendorf (1977) assumed V is proportional to A to model lake level fluctuation in response to stochastic forcing, Bengtsson and Malm (1997) assumed A is proportional to h 2 to study the sensitivity of lake level to climatic condition, and O’Connor (1989) assumed V is proportional to h m and A is proportional to hm⫺1 to simulate the variation of dissolved solids in lakes and reservoirs. The scope of this paper is limited to ephemeral ponds and wetlands in small natural depressions, for example, “pothole” wetlands in glaciated plains. Simple power functions with two parameters will be proposed to represent A–h and V–h relations, and tested using data for 27 shallow lakes, wetlands and ponds in the northern prairie region of North America. The power functions presented in this study are expected to be widely applicable to small wetlands and ponds in isolated and smoothly sloped depressions, even though they have only been tested in the northern prairie environment. 2. Theory A–h and V–h relations are dependent on each other, Zh 0 A h dh 1 where h is a dummy variable of integration and h is the depth measured at the deepest point of the lake. This relationship between V and A is fundamental and applies to all lakes and wetlands that have a horizontal water surface. Experimentally determined V–h and A–h relations must satisfy Eq. (1), a failure of which indicates that the V–h and A–h relations are inconsistent. The A–h and V–h relations of a basin can often be approximated by simple analytical expressions such as polynomials or power functions. In this paper power functions are proposed that are based on the shape of simple symmetric basins formed by rotating a slope profile around the central axis (Fig. 1). The slope profiles are given by y=y0 r=r0 p 2 where y [L] is the relative elevation of the land surface at a distance r [L] from the center, y0 [L] is the unit elevation, for example 1 m in SI units, r0 is the radius corresponding to y0, and p is a dimensionless constant. It follows from Eq. (2) that the area of the water surface corresponding to a depth of water h measured at the center of the basin r 0 is given by A pr02 h h0 2=p 2=p h s h0 3 where h0 [L] is the unit depth, s [L 2] is a scaling constant, which is equal to the area of water surface when h h0 : The constant p provides the link between the shape of the basin (Fig. 1) and A–h relation. A small value, for example p 2; corresponds to a paraboloid basin that has smooth slopes extending from the center to the edge, and a large value corresponds to a basin that has a flat bottom. In an extreme case, we can set p ! ∞: This corresponds to a cylinder, for which A s regardless of h. It follows from Eq. (1) that the volume of water corresponding to h is 76 M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 Fig. 2. Location of the present and previous study sites. (1) St. Denis; (2) Fort Qu’Appelle; (3) Melfort; (4) Saskatoon; (5) Swift Current; (6) Wilkie; (7) Ward; (8) Dickey; (9) Stutsman. Shaded area indicates the extent of the prairie wetland region (Winter, 1989). given by V s h1⫹ 2=p 1 ⫹ 2=p h2=p 0 4 Unlike the hypothetical basins represented in Fig. 1, real wetlands have more complex, asymmetric shapes, and they commonly occur in the lowest part of catchments where slopes are concave. With suitable p and r0, Eq. (2) represents the concave portion of most slope profiles reasonably well. A natural depression is made up by many slope profiles each having different values of p and r0, and no single slope profile represents the entire depression. Therefore, one might expect that Eqs. (3) and (4) cannot adequately represent natural depressions. However, as will be shown, the field data suggest that A–h relations of all natural depressions examined in this study are well approximated by Eq. (3). In this case p and s in Eq. (3) represent the shape and the size of depressions in some average sense. 3. Field sites and methods 3.1. Present study The present field study was conducted in the St. Denis National Wildlife Area (NWA) in Saskatchewan, Canada as part of a multidisciplinary research project to understand the hydrology and the ecology of prairie wetlands. The site is located at 106⬚06 0 W and 52⬚02 0 N, which is approximately 40 km east of Saskatoon (Fig. 2). The topography of the site is described as moderately rolling knob and kettle moraine with slopes varying from 10 to 15% (Miller et al., 1985). The area is underlain by glacial tills, which have a high clay content of 20–30%. The mean annual precipitation in Saskatoon is 360 mm, of which 84 mm occurs as snow (Atmospheric Environment Service, 1997). Air temperature frequently becomes lower than ⫺30⬚C in winter during which the soil frost penetrates as deep as 2 m. The uplands around the wetlands have been under cultivation for 50–100 years. Within the St. Denis NWA, four wetlands identified as S92, S109, S120, and S125S were selected for detailed elevation surveys. The extent of the wetlands is loosely defined by the growth of aquatic vegetation such as sedge and spike rush and by the presence of soft organic-rich soil, but the water-covered areas of the wetlands drastically change during a year. The wetlands become entirely or partially inundated in spring after snowmelt runoff. Runoff rarely occurs in summer and water levels in the wetlands gradually decline (Hayashi et al., 1998). Topographical maps of the four wetlands are shown in Fig. 3. Surveyed areas did not completely cover the catchments of S92, S109, and S125S, and part of drainage divides are drawn along the limit of the surveyed area. The missing area is small compared to the areas included in the maps. In addition to the four wetlands, a detailed elevation survey was conducted for four small depressions on the cultivated uplands. These small depressions hold ephemeral ponds only for a week to a few weeks in early spring, and are not considered wetlands in usual sense. However, they are hydrologically important because they store snowmelt water and recharge local groundwater. Three such depressions (D1, D2, and D3 in Fig. 3b) are located in and adjacent to the catchment of S109. The fourth one (S104) is located 200 m northeast of S109. The catchments of the wetlands and depressions were surveyed in 1994, 1998, and 1999 using total stations. For the wetlands, survey points were spaced horizontally at 10–15 m intervals in the uplands, and M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 77 Fig. 3. Topographical maps showing the elevation above the mean sea level of the four catchments in the St. Denis NWA. Principal contour interval is 1 m. Scale bars indicates 50 m. Wetlands are indicated by shades and drainage divides are indicated by thick lines. The location of small depressions D1, D2, and D3 are indicated in (b). 5–10 m intervals in the wetlands. For the small depressions, survey points were spaced horizontally at 2–5 m intervals. Estimated measurement error is within a few centimeters for elevation and within a few tens of centimeters for horizontal location. The software package Surfer (Golden Software, Golden, CO, USA) was used to estimate the elevation on regularly spaced grids by interpolation and to construct digital elevation models (DEMs). The maps shown in Fig. 3 are based on the DEMs. From the DEMs, V–h and A–h relations were calculated using the volume and area integration tool of Surfer. The depth of water h is defined as the elevation difference between the water surface and the lowest point in the depression, which means that h 0 when the wetland becomes completely dry. The kriging method (Davis, 78 h (m) A (m 2) V (m 3) A (m 2) V (m 3) 0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 S92 170 520 750 970 1180 1380 1590 1810 2050 2350 2730 3200 5.2 41 100 190 300 430 570 740 940 1160 1410 1700 S109 190 500 830 1130 1410 1690 1970 2250 2630 3120 3660 4120 6.5 41 110 210 330 490 670 880 1120 1410 1750 2140 A (m 2) V (m 3) S120 520 900 1190 1430 1660 1880 2100 2300 2570 2830 3150 21 93 200 330 480 660 860 1080 1330 1600 1890 A (m 2) V (m 3) S125S 360 790 1210 1610 1990 2360 2740 3120 3510 3850 17 74 170 320 500 710 970 1260 1590 1970 A (m 2) V (m 3) S104 180 340 500 660 820 1010 1210 9.5 36 78 140 210 300 410 A (m 2) V (m 3) D1 150 360 610 7.2 32 81 A (m 2) V (m 3) D2 63 130 210 310 460 3.2 13 30 56 94 A (m 2) V (m 3) D3 88 160 270 370 470 600 760 4.6 18 40 72 110 170 230 M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 Table 1 Area A and volume V of water corresponding to depth h in the wetlands and small depressions in the St. Denis NWA M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 79 Table 2 Upper depth limit hmax, scaling constant s, power constant p, root-mean-squared error of area Aerr and volume Verr of the wetlands, the relative magnitude of Aerr with respect to A1m, and Verr with respect to V1m. Asterisks indicate that the relative magnitudes of Aerr and Verr are evaluated against A and V at hmax s (m 2) p Aerr (m 2) hmax (m) St. Denis S92 S109 S120 S125S S104 D1 D2 D3 1.2 1.2 1.1 1.0 0.7 0.3 0.5 0.7 2450 3180 2820 3840 1720 2880 1160 1130 1.80 1.61 2.66 2.10 1.95 1.55 1.45 1.66 Shjeflo (1968) Pothole 1 Pothole 2 Pothole 4 Pothole 5 Pothole 5A Pothole 6 Pothole 7 Pothole 8 Pothole C1 1.4 1.7 1.5 2.3 1.6 1.5 1.5 1.1 1.9 51 900 109 700 93 400 78 200 9600 33 300 86 800 123 100 162 900 5.12 3.64 4.31 5.48 2.49 6.19 3.52 3.33 5.33 1360 8110 4610 1390 239 200 1820 6260 4540 2.6 6.9 5.2 1.7 2.6 0.6 2.1 5.3 2.8 685 2010 814 961 100 359 801 1120 3870 1.7 2.4 1.5 1.5 2.1 1.5 1.7 1.3 3.4 Lakshman (1971) Ft. Qu’Appelle 1 Ft. Qu’Appelle 2 Ft. Qu’Appelle 17 Ft. Qu’Appelle 19 Ft. Qu’Appelle 20 Melfort 7 Saskatoon 16 Swift Current 1 Wilkie 6 Wilkie 12 1.4 1.5 1.5 1.2 0.9 1.1 0.9 2.0 0.6 1.2 7160 8570 5790 6720 4310 8990 1960 34 100 11 800 2150 3.22 3.00 2.79 3.75 3.11 2.44 4.06 3.26 4.72 3.28 57 342 83 142 98 392 31 2610 242 46 0.8 4.7 1.6 1.5 1.4 ⴱ 4.1 1.7 ⴱ 7.4 2.6 ⴱ 2.1 101 125 37 90 54 236 44 1240 118 13 2.0 3.1 1.4 1.6 1.4 ⴱ 4.3 5.1 ⴱ 7.1 2.9 ⴱ 0.9 1986, p. 239) was used for interpolation. Preliminary analysis showed that the calculated V–h and A–h relations were essentially independent of the choice of semivariogram and grid spacing. The relations presented in this paper were calculated using linear semivariograms with no drift and 5 m grids for the wetlands and 1 m grids for the small depressions. 3.2. Previous studies Two sets of published data are used in this paper; the first data set from North Dakota, USA (Shjeflo, 1968) and the second data set from Saskatchewan, Canada (Lakshman, 1971). The Shjeflo wetlands are located in three locations; Potholes 1, 2, and 4 in Ward 97 101 55 20 17 2.4 11 15 Aerr/A1m (%) Verr (m 3) Wetland ID 4.1 3.2 1.9 0.5 1.4 ⴱ 0.4 ⴱ 2.5 ⴱ 2.0 ⴱ 17 20 8.6 5.3 2.6 0.9 1.6 2.6 Verr/V1m (%) 1.5 1.4 0.5 0.3 0.6 ⴱ 1.1 ⴱ 1.7 ⴱ 1.1 ⴱ County, Potholes 5, 5A, 6, 7, and 8 in Dickey County, and Pothole C1 in Stutsman County (Fig. 2). The Lakshman wetlands are located near Fort Qu’Appelle, Melfort, Saskatoon, Swift Current and Wilkie (Fig. 2). Survey methods and the density of surveyed points were not clearly described in the original articles, which only included tables of the elevation of water surface with respect to an arbitrary datum and the area and the volume corresponding to each elevation. Shjeflo (1968) provided the elevation of the lowest point in each basin so that we could calculate the depth of water corresponding to each elevation and determine A–h and V–h relations. Lakshman (1971) did not provide such data, and we needed to estimate the lowest elevation by inspecting the water level record. 80 M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 Fig. 4. A–h relation of S92 and S109. Solid circles indicate data points and curves indicate the power function (Eq. (3)) with the values of s and p listed in Table 2. The estimation was only possible for those wetlands that dried up frequently in the reported study period of 1964–1970. In addition the elevation-area-volume data were incomplete for some wetlands. Therefore, out of 25 wetlands in Lakshman (1971), V–h and A–h relations were determined for only 15 of them. The accuracy of estimating h is expected to be in the order of 0.05 m. 4. Results 4.1. St. Denis wetlands The A–h and V–h relations of the wetlands and small depressions are listed in Table 1. The data are listed for a depth range between 0 and hmax. The hmax is defined by the overflow point for S120, S125 and all micro-depressions, and by the highest water level recorded in 1968–1997 for S92 and S109 (van der Kamp et al., 1999). The values of hmax are listed in Table 2. Fig. 4 shows the A–h relation of S92, which represents an irregularly-shaped end member, and for S109, which represents a reasonably regularly-shaped end member. Solid circles indicate data points calculated from the DEM and curves show Eq. (3) with the best-fit values of s and p determined by the leastsquares method. Table 2 lists the values of s and p. The power function (Eq. (3)) approximates the A–h relation of all wetlands reasonably well. To evaluate the goodness of fit between the data points and the power function, root-mean-squared (RMS) error Aerr is defined by v u X u1 m A ⫺ APF 2 5 Aerr t m i1 DEM Fig. 5. V–h relation of S92 and S109. Solid circles indicate data points and curves indicate the power function (Eq. (4)) with the values of s and p listed in Table 2. M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 Fig. 6. Relationship between s and p. where ADEM is the area calculated from DEM, and APF is the area given by the power function, and m is the number of data points. The magnitude of Aerr is comparable to the values of A in a range 0 ⬍ h ⱕ 0:1 m; and APF does not give a meaningful estimate in this depth range. The relative magnitude of Aerr 81 becomes less significant at greater depth range as A increases. The ratio Aerr =A is generally smaller than 10% in a range 0:3 m ⬍ h ⱕ hmax : Table 2 lists Aerr as well as Aerr =A1m ; where A1m is the data point closest to h 1 m: Once s and p are determined by fitting Eq. (3) to A– h data points, the same s and p can be used in Eq. (4) to approximate V–h relation. Fig. 5 shows the V–h relation of S92 and S109, in which solid circles indicate data points calculated from DEM and curves show Eq. (4). The goodness of fit between the data points and the power function (Eq. (4)) is expressed by RMS error, Verr, defined similarly to Eq. (5). The relative magnitude of Verr with respect to V is generally smaller than 10% in a range 0:3 m ⬍ h ⱕ hmax : Table 2 lists Verr and Verr =V1m ; where V1m is the data point closest to h 1 m: Eqs. (3) and (4) were similarly applied to the A–h and V–h relations of wetlands S120 and S125, and small depressions D1, D2, D3, and S104. The leastsquares-fit values of s and p, and RMS errors Aerr and Verr are listed in Table 2. The match between the power functions and the data points for the wetlands and small depressions is similar to that S92 and S109. The ratio Aerr =A and Verr =V are generally smaller than 10% in a depth range 0:3 m ⬍ h ⱕ hmax for wetlands and 0:1 m ⬍ h ⱕ hmax for small depressions. The relationship between s and p is shown in Fig. 6. St. Denis wetlands and depressions have relatively small sizes, which is reflected in the range of the scaling constant s. The values of p fall in a relatively narrow region around 2, which indicates that the depressions have a reasonably smooth shape that resembles a paraboloid (Fig. 1). 4.2. Previously studied wetlands Fig. 7. A–h and V–h relations of Pothole 4 (Shjeflo, 1968). Solid circles indicate data points and curves indicate the power functions with values of s and p listed in Table 2. A–h and V–h data were available for 10 wetlands in Shjeflo (1968) and 15 wetlands in Lakshman (1971). The data were examined with respect to the consistency condition given by Eq. (1). At each reported value of h, a value of V was calculated from A–h relation using Eq. (1), and compared to the reported value of V. If the relative difference was greater than 15% at any h greater than 0.2 m, the data set was considered inconsistent. The source of inconsistency may be inappropriate methods used to calculate A and V from the survey data. Nine wetlands in Shjeflo 82 M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 (1968) and ten in Lakshman (1971) satisfied the condition, and will be used in the following analysis. The upper depth limit hmax, listed in Table 2, is arbitrarily set to be the highest water level recorded in the study period (1960–1964 in Shjeflo (1968) and 1964– 1970 in Lakshman (1971)) plus 0.5 m. If A–h and V–h data do not cover hmax, the highest value of h within the data set is considered to be hmax. Eq. (3) was fitted to the A–h relation of the nine Shjeflo wetlands to determine s and p (Table 2). The least-squares method was applied to each data set within a range 0:1 m ⱕ h ⬍ hmax : The data points having h less than 0.1 m were excluded from the analysis because the accuracy of h at such a small value is questionable. As an example Fig. 7a shows the A–h relation of a wetland identified as Pothole 4, and Fig. 7b shows the V–h relation of the same wetland. Solid circles indicate data points and curves show the power functions (Eqs. (3) and (4)). The power functions agree with the data points reasonably well for both A–h and V–h relations. Similarly the power functions adequately represent A–h and V–h relations of all other Shjeflo wetlands. The RMS errors Aerr and Verr and the relative magnitudes Aerr =A1m and Verr =V1m are listed in Table 2. For all nine Shjeflo wetlands, Aerr =A and Verr =V are generally smaller than 10% in a range 0:3 m ⬍ h ⱕ hmax : The power function (Eq. (3)) was also applied to the Lakshman wetlands to determine the least-squares-fit values of s and p (Table 2). The agreement between data points and the power function is reasonably good for all ten wetlands. The RMS errors and their relative magnitude with respect to A1m and V1m are listed in Table 2. For all ten Lakshman wetlands, Aerr =A and Verr =V are generally smaller than 10% in a range 0:3 m ⬍ h ⱕ hmax : The relationship between s and p is shown in Fig. 6. The positive correlation indicates that Shjeflo and Lakshman wetlands, which are much larger in size than St. Denis wetlands, have higher values of p. The implication of the size-shape relationship will be discussed later. 5. Simple field methods for determining approximate A–h and V–h relations In general, accurate determination of A–h and V–h relations requires a detailed elevation survey over an entire wetland, which is labor intensive and time consuming. Based on the above results, it is likely that Eqs. (3) and (4) approximately represent A–h and V–h relations of most small topographic depressions. Therefore, at least for the first approximation, one only needs to measure A and h at a few different times to determine s and p. For a given wetland, let z be the elevation of the water surface with respect to an arbitrary datum, for example a staff gauge, and zmin be the elevation of the lowest point in the wetland. Eq. (3) can be written as A s z ⫺ zmin =h0 2=p 6 In principle, three unknown constants s, p, and zmin can be determined from three independent measurements of A and z. However, it is easy to measure zmin for most wetlands in small depressions. For example, an observer can locate the lowest point in the wetland just before it becomes completely dry, or an observer on a boat can probe around the central part of the wetland to find the deepest point. Therefore, in most cases only s and p need to be determined from two independent measurements of A and z. There are a number of ground-based and airborne methods to estimate A of many wetlands relatively easily. If time and resources are limited, an observer may chose to estimate p from the size–shape relationship shown in Fig. 6. For example, Fig. 6 indicates that p is likely close to 2 in small seasonal wetlands and ephemeral ponds. In this case, an observer may assume p 2 and determine s from a single measurement of A and h. The accuracy of A–h and V–h relations determined this way may not be high, but some field studies can benefit from the simplicity and the practicality of the method. We used this method in the St. Denis NWA in 1998 to estimate the volume of snowmelt runoff collected in several depressions that did not have detailed survey data. This measurement showed that significant portion of snowmelt runoff is stored in depressions without draining to the main wetland in the catchment, and gave us an important step forward in understanding the hydrology of prairie wetlands. M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 83 Fig. 8. Hypsometric curves of the catchments of S92 and S109. Shaded regions indicate wetlands. 6. Discussion 6.1. Hypsometry: statistical meaning of the shape parameter p The field data indicate that the A–h relation of many wetlands is represented by a well-defined value of p that reflects the basin shape in some average sense. This finding is not trivial and warrants some discussion. To this end it is informative to reexamine how the A–h relation is determined from detailed survey data. The A–h relation of a basin is determined from a high-resolution DEM of the basin. On the DEM, the basin is hypothetically filled with water to a series of values of h, and for each h the grid points that are under the water surface are counted. If there are n grid points under the water surface located at h, then the area of the water surface A is given by A h n hDa 7 where Da is the area associated with each grid point. The above procedure involves “counting” the number of grid points, which suggests that A can also be expressed in terms of the frequency distribution of grid points within the basin. The cumulative frequency distribution F h is defined as F h n h=N 8 where N is the total number of grid points in the basin. Since the total area of the basin Atot is given by Atot NDa 9 Fig. 9. North–south (NS) and east–west (EW) slope profiles of Pothole 4 and S109. Open squares indicate EW profiles and solid circles indicate NS profiles. Curves show the best fit power functions having power p. (a) Pothole 4. (b) S109. \tflt="PS6F00" \tfnm="PS6F00" A h and F h are related by A h Atot F h 10 Eq. (10) clearly demonstrates the equivalence between A–h relation and frequency distribution. Similar ideas have been used to study the distribution of landmass within a drainage basin by geomorphologists, who use the term hypsometric analysis for studying how F h changes in relation to landform evolution (Strahler, 1952; Willgoose and Hancock, 1998). The F h of the entire catchments of S92 and S109 are shown in Fig. 8. The graphs, called hypsometric curves, have F h on the horizontal axis and h on the vertical axis following the geomorphological convention (Strahler, 1952). The values of A corresponding to F are also plotted on the top horizontal axis. Each wetland occupies only a small portion of the catchment, which is indicated by a shaded region at the toe of the hypsometric curves. The wetland 84 M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 portion of the hypsometric curves have relatively simple shape that is adequately represented by the power function (Eq. (3)) even though the overall shape of the hypsometric curves is more complex. It is clear from the above discussion that the A–h relation of a depression should be regarded as the frequency distribution of land elevation within the depression. Therefore, two given depressions may have an identical A–h relation even though their actual shape is significantly different. The shape parameter p in Eq. (3) represents the slope profile of a hypothetical basin (Fig. 1) that is hypsometrically equivalent to the actual depression. 6.2. Size–shape relationship and landform evolution Fig. 6 shows that larger wetlands tend to have higher values of p. In general large prairie wetlands have water for a long period of time in a given year, in fact many of them are semi-permanent lakes, while small prairie wetlands have water only for a few months after snowmelt. As a result, large wetlands have a flat bottom formed by sedimentation. For example Fig. 9a shows north–south (solid circles) and east–west (open squares) profiles of Pothole 4 in Shjeflo (1968) and Fig. 9b shows north–south and east–west profiles of S109 in St. Denis. Markers in Fig. 9 show the data points and lines show Eq. (2) using the least-squares-fit values of p and r0. As indicated in Fig. 9, the profiles in Pothole 4 have large p reflecting the flat bottom, while those in S109 have small p reflecting relatively smooth slope from the center to the edge. In Fig. 9 p takes a range of values because the wetlands are asymmetric and each individual slope has different curvature. The A–h relation integrates all slope profiles within the wetland and defines a single value of p in an average sense. For example, a representative p of Pothole 4 (Fig. 9a) may be given by a harmonic average pav of the north, east, south, and west profiles; pav 4:85: This is comparable to p 4:31; which was obtained by fitting Eq. (3) to the A–h data set. Similarly, a harmonic average of the four profiles of S109 (Fig. 9b) is pav 1:78; which is comparable to p 1:61; which was obtained by fitting Eq. (3) to the A–h data set. The wetlands examined in this paper occur in the northern prairie region that is characterized by semi- arid climate having warm summer and cold winter, and by thick glacial tills and hummocky topography. Many prairie wetlands become partially or completely dry in late summer and fall, and the land surface is subjected to gravity-driven soil creep induced by freezing and thawing, drying and wetting (Kirkby, 1967), cultivation (de Jong et al., 1998), and animal burrow activities (Black and Montgomery, 1991). Geomorphological literatures suggest that soil creep tends to dissipates the irregularity of the landform by diffusion-like processes (Culling, 1960). It is reasonable to expect that the dissipation of irregularity at a local scale results in the smooth frequency distribution of land elevation at a basin-wide scale, and hence to simple A–h and V–h relations. If the landform evolution is solely driven by soil creep, slope profiles are expected to approach curves having a low value of p (Fig. 1). In contrast, if the landform evolution is strongly influenced by other mechanisms like underwater sedimentation, slope profiles may take a higher value of p that reflects the balance between several driving forces. It is not clear if similar smoothing mechanisms exist in different environments, for example deep lakes on a rocky terrain or playas on tropical savanna. Therefore, the applicability of Eqs. (3) and (4) is so far limited to the prairie region where the equations have been tested. However, it will be interesting to examine whether the equations are useful for the lakes and wetlands in different environments. 7. Conclusions The main objective of this paper is to examine the relation between the volume V, area A, and depth h of wetlands in isolated depressions. For the 27 wetlands and ephemeral ponds examined in this paper, A is proportional to h2=p to a good approximation, and V is proportional to h1⫹2=p ; where p is a dimensionless constant. In other words, A–h and V–h relations are expressed as power functions. The constant p is related to the shape of the depression, more specifically the functional form of slope profiles. For example, a paraboloid-shaped depression has p 2; and a cylinder-shaped depression has p ! ∞: Natural depressions have more complex and asymmetric shape. In general, low values of p occur in depressions M. Hayashi, G. van der Kamp / Journal of Hydrology 237 (2000) 74–85 having smooth slopes from the center to the margin, and high values occur in depressions having a flat bottom. For the 27 depressions, p ranges approximately between 2 and 6. Large wetlands have high p and small wetlands have low p reflecting the basin shape. By definition the A–h relation is closely related to the frequency distribution of land elevation within the depression, and it reflects the shape of the depression in a statistical sense. Therefore, A–h curves are similar to the hypsometric curves that are used by geomorphologists to study landform evolution. Assuming that the power functions approximate the unknown A–h and V–h relations of a given wetland, an observer only needs to determine two constants in the function; the scaling constant s and the shape constant p. This can be achieved by two independent measurements of A and h, and offers a less laborintensive alternative to the standard method of determining A–h and V–h relations from detailed elevation data. This simple geometric model can be used in field studies requiring approximate values of A and V. It also provides a valuable tool for theoretical and modeling studies because the parameters s and p sensitively reflect the size and the geometry of small lakes and wetlands. The applicability of the equations have only been tested in the northern prairie region of North America, and future studies are required to examine if the equations adequately represent lakes and wetlands in other environments. Acknowledgements We gratefully acknowledge the Canadian Wildlife Service for the use of the St. Denis NWA. We thank Randy Schmidt, Vijay Tumber, Catheryne Staveley, David Parsons, Trevor Dusik, Herman Wan, and Geoff Webb for their assistance in the collection and analysis of survey data; and Yvonne Martin for the discussion on landform evolution. We also thank Lars Bengtsson and an anonymous reviewer for constructive comments. 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