NOTE BRÈVE
ASSEMBLAGES OF MILLIPEDES AND CENTIPEDES ACCORDING
TO THE TYPE OF PASTURE MANAGEMENT: PRELIMINARY RESULTS
FROM THE CZECH REPUBLIC
Jan MIKULA1 !"#$%&#'!ŠARAPATKA1 !()*$%+,!KOPECKÝ2 & Ivan H. TUF1,*
RÉSUMÉ. — Les peuplements de millipèdes et de centipèdes résultant du type de gestion des pâturages:
résultats préliminaires en République tchèque-!.!/0!123!*0!+0330!432*0!435%3!*0!+#675808!)5!847583%3%#9!03!
)0:!:382+3280:!*0:!;8#2706093:!*0!+093%7<*0:!03!*0!6%))%7<*0:!*59:!)0:!7=3285;0:!09!>#9+3%#9!*0!*%>>48093:!
3?70:!*0!;0:3%#9-!/0:!3?70:!*0!;0:3%#9!4&5)24:!435%093!)0!7=3285;0!*0!6#23#9: !)0!>52+,5;0!03!)0!75%))5;0!*59:!
*%&08:0:!+#61%95%:#9:-!@%>>48093:!6#*0:!*0!>#9*53%#9:!*0!7=3285;0:!#93!4;5)06093!434!4&5)24:!A!64)59;0!
+#6608+%5)! *0:! +2)3280:! >#2885;<80: ! :0609+0:! )#+5)0:! 64)59;40:! 5&0+! *0:! 78#7#83%#9:! *%>>480930:! *0!
7)5930:!*%+#3?)4*#90: !03!290!'5+,<80!:7#935940-!B5);84!)5!858034!*0:!+093%7<*0:!03!*0:!6%))%7<*0: !)C0>>03!
)0!7)2:!7#:%3%>!7#28!)0:!6%))%7<*0:!5!434!#1:08&4!*59:!)0!+5*80!*2!75%))5;0!+58!+0)2%A+%!#>>80!*0!)5!653%<80!
#8;59%D20! 6#830-! /5! 60%))0280! >5E#9! *0! ;4808! )0! 7=3285;0! 0:3! *C23%)%:08! 29! 64)59;0! )#+5)! *0! :0609+0:! F!
>#830!78#7#83%#9!*0!7)5930:!*%+#3?)4*#90:!+#660!9#288%3280!*0!D25)%34!:2748%0280!09!+#67585%:#9!*2!;5G#9SUMMARY. — H,0!5%6!#>!3,%:!:32*?!I5:!3#!+#67580!*%:38%123%#9!59*!5::061)5;0!:382+3280:!%9!+093%A
70*0:!59*!6%))%70*0:!%9!75:3280:!I%3,!*%>>08093!6595;06093!58859;06093:-!H,0!0&5)2530*!3?70:!#>!6595;0A
6093!%9+)2*0*!:,007!;85G%9; !6#I%9; !62)+,%9;!59*!&58%#2:!+#61%953%#9:!#>!3,0:0!380536093:-!@%>>08093!
>#29*53%#9!6#*0:!#>!75:3280!I080!5):#!0&5)2530*!J!+#6608+%5)!6%K3280:!#>!>#**08!+8#7: !)#+5)!6%K3280:!#>!
:00*:!I%3,!*%>>08093!78#7#83%#9:!#>!*%+#3?)0*#9#2:!,081: !59*!:7#93590#2:!>5))#I!)59*-!@0:7%30!)#I!5129A
*59+0!#>!1#3,!+093%70*0:!59*!6%))%70*0: !3,0!6#:3!7#:%3%&0!0>>0+3!#1:08&0*!%9!6%))%70*0:!I5:!%9!62)+,%9;!
*20!3#!3,0!:287)2:!#>!*05*!#8;59%+!653308-!H,0!10:3!I5?!3#!6595;0!75:3280!%:!3#!2:0!5!6%K3280!#>!)#+5)!:00*:!
I%3,!5!,%;,!78#7#83%#9!#>!*%+#3?)0*#9#2:!,081:!5:!>##*!#>!,%;,08!:21:3593%5)%3?!+#67580*!3#!;85::-
L#%)!%9&083018530:!7)5?!59!%67#83593!8#)0!%9!0+#:?:306!>29+3%#9!5:!3,0?!%678#&0!3,0!9238%093!
+?+)0! 3,8#2;,! *0+#67#:%3%#9! 78#+0::0:! 59*! %3:! 80;2)53%#9! 1?! 780*53%#9! 59*! 3,0?! 6#*%>?! 3,0!
7,?:%+5)!78#7083%0:!#>!:#%)!MNO:+,6%33!P!Q8%>R3,: !STTUV!W#)308:,!XYYSV!/5&0))0!et al., 2006).
Z)3,#2;,! 3,0%8! 8#)0! %:! 29*080:3%6530* ! 3,0?! %8807)5+051)?! 7583%+%7530! %9! 5! :0)>A#8;59%G%9;! :#%)!
:?:306!59*!78#&%*0!&5)251)0!0+#:?:306!:08&%+0:!M/5&0))0!et al- !XYY[\-!H,2: !>#8!%9:359+0 !3,0?!
7)5?!59!%67#83593!8#)0!*28%9;!0+#)#;%+5)!80:3#853%#9!#>!*%:32810*!5805:!MH5'#&:O] !XYYSV!L9?*08!
P!^09*8%K !XYYU\-!_#8!3,0:0!805:#9: !%3!:006:!3#!10!%67#83593!3#!6595;0!:#%)!65+8#>5295!%9!
1! @075836093! #>! N+#)#;?! 59*! N9&%8#960935)! L+%09+0: ! _5+2)3?! #>! L+%09+0 ! `5)5+O?! a9%&08:%3? ! L&#1#*?! X[ !
bcAddSe[!()#6#2+ !bG0+,!f0721)%+
2! @075836093!#>!c##)#;?!59*!_%:,!_586%9; !_5+2)3?!#>!Z;8#1%#)#;? !_##*!59*!g53285)!f0:#28+0: !bG0+,!a9%&08A
:%3?!#>!/%>0!L+%09+0:!%9!`85;20 !h56]+Oi!SXT !bcAS[jXS!`85,5![!A!L2+,*#) !bG0+,!f0721)%+
k!+#880:7#9*%9;!523,#8l!%&59-32>m27#)-+G
Rev. Écol. (Terre Vie), vol. 69, 2014.
– 167 –
#8*08!3#!65%935%9!:#%)!>083%)%3? !0:70+%5))?!%9!5;8#0+#:?:306:!M/5&0))0!et al- !STTe\-!^09+0 !59?!
6595;06093!80;%60!6#*%>?%9;!3,0!+#6629%3?!:382+3280!#>!:#%)!65+8#A%9&083018530:!%:!)%O0)?!3#!
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65+8#>5295!%9!b09385)!N28#7059!+#9*%3%#9:-!b093%70*0:!580!780*53#8:!80;2)53%9;!7#72)53%#9:!
#>!:65))08!%9&083018530:!M0-;-!:78%9;35%): !7#3I#86: !0583,I#86: !#8%153%*!6%30: !03+-\ !*0+#6A
7#:%9;!#8;59%+!653308!M`#:08,!STUU\V!6%))%70*0:!580!*038%3%&#80:!*0:38#?%9;!)%3308!59*!>283,08%9;!
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3859:>#8608:!M/5&0))0!et al- !STTd\-!"#3,!6#*0)!;8#27:!#>!:#%)!%9&083018530:!,5&0!1009!2:0*!
10>#80!>#8!0&5)253%#9!#>!D25)%3?!#>!75:3280:!I%3,!#90!65%9!80:2)3!J!3,0!65%9!90;53%&0!>5+3#8!
#>!+#9&093%#95)!5;8%+2)3280!6595;06093!%:!3,0!2:0!#>!70:3%+%*0:!Mh)%9;08 !STTXV!")5+O1289!P!
Z83,28 !XYYS\-!b#67)030!5159*#96093!#>!6595;06093!%:!2:0>2)!>#8!6?8%57#*:!%9!75:3280:!5:!
I0))!5:!605*#I:!ML00108!et al- !XYYj\-!(28!:32*?!I5:!5%60*!3#!0&5)2530!3,0!%675+3!#>!*%>>08093!
3?70:!#>!75:3280!6595;06093!#9!5::061)5;0:!#>!+093%70*0:!59*!6%))%70*0:MATERIAL AND METHODS
H,0! R0)*! %9&0:3%;53%#9! I5:! 708>#860*! #9! 3,800! )#+5)%3%0:! %9! N5:3! B#85&%5 ! %9! 3,0! bG0+,! f0721)%+-! H,0! R8:3!
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*%>>08093! 3?70:! #>! 6595;06093-! @%>>08093! +#61%953%#9:! #>! :0&085)! *%&08:0! 380536093:! I080! 708>#860*! #9! %9*%&%*25)!
7)#3:!M:%G0!#>!7)#3!jqSY!6\-!H,0!380536093!&58%593:!I080l!S\!708659093!:,007!;85G%9; !X\!6#I%9; !p\!5)3089530!;85G%9;A
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#>!c218%!Mg!eToXUC !N!SUo[C\ !:%32530*!#9!5!R0)*!I%3,!5!6#:5%+!#>!7)#3:!M:%G0!#>!7)#3!jqj!6\!:#I9!I%3,!*%>>08093!3?70:!
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R0)*!,5:!1009!+#938#))0*-!H,0!:0+#9*!)#+5)%3?!Mc218%A?#29;\!I5:!+80530*!%9!XYY[-!H,0!3,%8*!)#+5)%3?!Mc218%A#)*\!,5:!1009!
380530*!:%9+0!STTd !I,09!3,0!0K708%6093!10;59L#%)!%9&083018530:!%9+)2*%9;!6?8%57#*:!I080!+#))0+30*!>8#6!je!7%3>5))!>#865)*0,?*0!3857:!M5!;)5::!'58!I%3,!5!6035)!
+#&08!59*!59!%9:0830*!7)5:3%+!7#3!I%3,!X!*)!#>!e!r!>#865)*0,?*0 !p!3857:!708!7)#3!58859;0*!%9!5!)%90!I%3,!p!6!%9308:75+%9; !
0K7#:0*! >8#6! B5?! 3#! g#&06108! XYYd ! +,0+O0*! 6#93,)?\-! s2593%353%&0! *535! >8#6! 7%3>5))! 3857:! I5:! 0&5)2530*! 1?!
bZg(b(!78#;85660!>#8!W%9*#I:!e-jt!MH08!"855O!P!u6%)5208 !STTU\-!H,0!0>>0+3!#>!*%>>08093!3?70:!#>!6595;06093!
#9! 6?8%57#*! *%:38%123%#9! I5:! 0&5)2530*! 1?! 29%6#*5)! +59#9%+5)! +#880:7#9*09+0! 595)?:%:! MbbZ\ ! 59*! B#930Ab58)#!
70862353%#9!30:3:-
RESULTS
Z)3#;03,08 !SYe!%9*%&%*25):!#>!+093%70*0:!59*!6%))%70*0:!I080!385770* !80780:093%9;!j!:70A
+%0:!#>!6%))%70*0:!>8#6!3,0!`#)?*0:6%*50 !g065:#653%*50!59*!v2)%*50!>56%)%0:!59*!e!:70+%0:!
#>!+093%70*0:!>8#6!3,0!^09%+#7%*50!59*!/%3,#1%%*50!>56%)%0:!80:70+3%&0)?!MH51-!w\-!H,0!,%;,0:3!
926108:!#>!%9*%&%*25):!I080!80+#8*0*!>8#6!3I#!7)#3:!I%3,!*%>>08093!6595;06093!380536093: !
#90!80780:0930*!1?!5)3089530!;85G%9;A6#I%9;!I%3,!>083%)%G53%#9!59*!3,0!#3,08!1?!5!6%K3280!#>!
)#+5)!:00*:!I%3,!5!,%;,!78#7#83%#9!#>!,081:!MH51-!w\-!H,0!L,599#9AW05&08!*%&08:%3?!%9*0K!I5:!
,%;,0:3!#9!75:3280!6#I0*!3I%+0!5!?058-!
@20!3#!3,0!)#I!926108!#>!%9*%&%*25): !#9)?!65308%5)!>8#6!3,0!B#8O#&!)#+5)%3?!I5:!595A
)?:0*!1?!bZg(b(-!bbZ!1%7)#3!:26658?!M_%;-!S\!:,#I:!3,0!*%:38%123%#9!#>!6?8%57#*: !3,0!
R8:3!5K%:!*0:+8%10:!XS-e!r!59*!3,0!I,#)0!6#*0)!*0:+8%10:!jT-X!r!&58%51%)%3?!#>!:70+%0:!*535-!
H,0!I,#)0!6#*0)!%:!:%;9%R+593!M_!x!X-Up[ !7!x!Y-YYXY\ !3,0!:%;9%R+593!0>>0+3!#9!*%:38%123%#9!#>!
6?8%57#*!:70+%0:!%:!0&%*093!%9!5)3089530!;85G%9;A6#I%9;!M_!x![-XT !7!x!Y-YYX !*0:+8%1%9;!ee!r!
&58%51%)%3?\!59*!62)+,%9;!M_!x!X-Sj !7!x!Y-YeU !*0:+8%1%9;!XU-U!r!&58%51%)%3?\-!(90!#>!3,0!6#:3!
%67#83593!80:2)3:!80>08:!3#!3,0!,%;,!780>0809+0!#>!3,0!Unciger foetidus!6%))%70*0!>#8!:%30:!I%3,!
62)+,%9;-!H,%:!7533089!I5:!5):#!0&%*093!>#8!5))!6%))%70*0:!%9!595)?:%:"0:%*0:! 3,0! )#I! 926108! #>! 385770*! %9*%&%*25):! #9! :65))! 7)#3:! 53! 3,0! c218%A#)*! 59*!
c218%A?#29;!)#+5)%3%0:!%3!%:!0&%*093!3,53!:7#93590#2:!>5))#I!)59*!%:!59!29+#6>#8351)0!:%30!>#8!
6?8%57#*:– 168 –
TABLE I
Cylindroiulus latestriatus!Mb283%: !SUej\
Unciger foetidus Mb-/-h#+, !SUpU\
Unciger transsilvanicus!My08,#0>> !SUTT\
Lamyctes emarginatus g0I7#83 !SUee
!"#$%!&'()$*+,-"&'(/%99502: !SdjU
Lithobius muticus b-/-h#+, !SUed
Lithobius microps B0%9083 !SU[U
g26108!#>!%9*%&%*25):
:,007!;85G%9;!
A
A
A
+
A
+
+
A
10
[j
j
p
L,599#9AW05&08!*%&08:%3?!%9*0K
Nemasoma varicorne!b-/-h#+, !SUed
A
Morkov
g26108!#>!:70+%0:
Polydesmus inconstans!/53G0) !SUUe
Recorded species of myriapods (ind./3 traps/5 months) on pastures with different types of management and calculation
of Shannon-Weaver diversity index of assemblages
0.46
:,007!;85G%9;!I%3,!>083%)%G53%#9
A
A
A
+
+
A
A
A
A
6
2
Y-XU
5)3089530!;85G%9;A6#I%9;!
A
A
A
A
A
+
+
A
A
p
2
Y-XU
5)3089530!;85G%9;A6#I%9;!I%3,!
>083%)%G53%#9
A
A
A
A
A
+
+
+
A
12
p
0.40
X!6#I%9;:!708!?058
A
A
A
+
+
A
+
+
A
10
4
Y-jp
p!6#I%9;:!708!?058
A
A
A
+
A
A
+
A
A
4
2
Y-pY
X!6#I%9;:!708!?058!I%3,!62)+,%9;
A
A
A
+
+
A
A
+
A
11
p
Y-pd
p!6#I%9;:!708!?058!I%3,!62)+,%9;
A
A
A
+
+
A
A
A
A
U
2
0.29
X!6#I%9;:!708!?058!I%3,!
>083%)%G53%#9
A
A
A
A
A
A
+
A
A
1
1
0.00
p!6#I%9;:!708!?058!I%3,!
>083%)%G53%#9
A
A
A
A
A
A
A
A
A
A
A
A!
pp
d
Zubri-young
)#+5)!:00*!6%K3280!I%3,!)#I!
78#7#83%#9!#>!,081:
+
A
+
A
A
A
+
A
+
U
4
Y-ed
)#+5)!:00*!6%K3280!I%3,!,%;,!
78#7#83%#9!#>!,081:
+
+
+
A
A
A
+
+
A
12
j
Y-eU
+#6608+%5)!6%K3280!#>!>#**08!
+8#7:
+
A
+
A
A
+
A
A
A
d
p
Y-pj
:7#93590#2:!>5))#I!)59*
A
A
+
A
A
A
+
A
A
6
2
0.20
6
2
Zubri-old
)#+5)!6%K3280!#>!:00*:!I%3,!,081:!
MXY!O;z,5\
A
A
A
A
+
A
A
A
A
1
1
0.00
)#+5)!6%K3280!#>!:00*:!I%3,!,081:!
MeY!O;z,5\
A
A
A
A
+
A
+
A
A
p
2
Y-XU
+#6608+%5)!6%K3280!#>!>#**08!
+8#7:
A
A
A
A
+
A
A
A
A
2
1
0.00
:7#93590#2:!>5))#I!)59*
A
A
A
A
A
A
A
A
A
A
A
A
– 169 –
_%;280! S-.! bbZ! 1%7)#3! #>! *%:38%123%#9! #>! 6?8%57#*:! 53! :%30:! 5++#8*%9;! 3#! 3?70! #>! 75:3280! 6595;06093! M)#+5)%3?!
B#8O#&\-
DISCUSSION
Z))!80+#8*0*!:70+%0:!580!+#66#9!606108:!#>!+#6629%3%0:!#9!:%30:!*%:32810*!1?!,2659:!
:2+,!5:!R0)*: !,057:!#8!28159!5805:!Mh)%9;08 !STTXV!/0{9%0I:O5 !STT[V!W?3I08 !STT[V!/59;,A
3#9 !XYYS\V!0-;-!%9!b09380 !w)0A*0A_859+0!59*!`#%3#2Ab,580930:!80;%#9:!M_859+0\ !w#8%#!MXYYe\!
#1:08&0*!3,53!L. emarginatus!%:!,%;,)?!80780:0930*!%9!5;8#0+#:?:306: !10+52:0!*0:7%30!+57A
3280:!%9!&58%#2:!,51%353: !5)6#:3!5))!,%:!*535!#9!3,%:!:70+%0:!+560!>8#6!5;8#0+#:?:306:!M+#89!
R0)*:\-!Z))!:70+%0:!I080!+)5::%R0*!5:!028?3#7%+!#8!5*57351)0!3#!5!b09385)!N28#7059!09&%8#96093!
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