Botanical Journal of the Linnean Society, 2010, 162, 130–422. With 12 figures
Patterns and traits of the endemic plants of Greece
KYRIACOS GEORGHIOU* and PINELOPI DELIPETROU
Department of Botany, Faculty of Biology, National and Kapodistrian University of Athens,
Panepistimiopolis, Athens 15784, Greece
Received 25 August 2009; accepted for publication 14 December 2009
Greece is characterized by high plant diversity (5800 species) and endemism (15.6%). This study attempts a first
overall assessment of the taxonomy, distribution, traits and conservation status of the Greek endemic plants. The
endemic species belong to 56 families and 242 genera. Most of the endemic plants have a narrow geographical and
altitudinal distribution range. The southern floristic regions are richer in endemic species. The species area
relationships for endemics (EARs) for island and continental floristic regions explain over 50% of the variation in
number of species and are characterized by steep curves. Analysis of the distributional pattern of the endemics by
similarity coefficients offers useful insights into the palaeogeography and biogeography of Greece. The endemic
species occur at all altitudes, but the altitudinal distribution shows a predominance of local endemics at 0–600 m
in the island regions and in higher zones in the continental regions. The life form spectra show a predominance
of hemicryptophytes and chamaephytes. This trait seems indicative of their habitat and adaptive strategy and may
be related to speciation processes. The overview of the conservation status of the Greek endemics indicates that
over 40% of the taxa are threatened or near threatened. © 2010 The Linnean Society of London, Botanical
Journal of the Linnean Society, 2010, 162, 130–422.
ADDITIONAL KEYWORDS: altitudinal distribution – conservation – distribution – endemism – floristic
analysis – life form – phytogeography – plant diversity.
INTRODUCTION
Greece is phytogeographically a borderline and a crossroads, an eastern Mediterranean and a Balkan
country, the south-easternmost edge of Europe in
immediate contact with South-West Asia and close
contact with North Africa. Its high mountains and
islands have acted through geological history as
stepping stones and refuges for taxa of the surrounding floristic regions and as isolation centres.
Mediterranean-type ecosystems cover more than 90%
of Greece, the vegetation belts ranging from
xerothermo- to cryo-Mediterranean (Hobbs, Richardson & Davis, 1995). In broad terms, the complicated
geological history, dissected topography and varied
bioclimate are the main reasons for the floristic diversity and increased endemism in Greece (Iatrou, 1986;
Thompson, 2005).
Despite active botanical exploration and study, the
general reference literature for the flora of Greece is
segmental and a great part of it not updated. The
*Corresponding author. E-mail: [email protected]
130
early 20th century floras, Conspectus Florae Graecae
(Halácsy 1901–1904, 1908, 1912), Prodromus Florae
Peninsulae Balcanicae (Hayek 1924–1933) and Flora
Aegaea (Rechinger, 1943a) include only parts of
Greece. The more recent Med-Checklist (Greuter,
Burdet & Long, 1984, 1986, 1989) covers a number of
families and the Mountain Flora of Greece (Strid,
1986; Strid & Tan, 1991) covers the mountain–alpine
region. The flora of the Cretan area is exceptionally
well investigated (Rechinger, 1943b; Greuter, 1973;
Greuter, Matthäs & Risse, 1984a, 1984b, 1985; Bergmeier & Matthäs, 1995) and there are recent floras
and checklists (Turland, Chilton & Press, 1993; Jahn
& Schönfelder, 1995; Chilton & Turland, 1997). Flora
Europaea (Tutin et al., 1968–1980, 1993) excluded the
East Aegean islands which were included in the Flora
of Turkey and the East Aegean Islands (Davis, 1965–
1986). Otherwise, the data are scattered across
13 276 published papers, monographs and theses
(Strid, 2006) and in unpublished herbarium material.
Recent floristic and taxonomic activity in Greece has
resulted in significant additions, exclusions and
changes in taxonomic status (Strid & Papanikolaou,
1985; Tan & Iatrou, 2001). The Flora Hellenica
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE
project will eventually bridge the gaps: the first two
volumes have been published (Strid & Tan, 1997,
2002), but the completion of the project will take
several more years.
In the meantime, conservationists lack an overall
assessment of Greek endemic species. The most recent
complete catalogue of Greek endemic plants listed 752
species (IUCN, 1982), a number later corrected to 742
(WWF & IUCN, 1994) which has since been quoted in
the literature (Phitos et al., 1995; Strid & Tan, 1997).
Apart from that, there is a recent endemic flora (Tan &
Iatrou, 2001) of the Peloponnese (Peloponnisos) and a
report of the disparately greater figure of a total of
1275 endemic taxa in Greece (Iatrou, 1996).
Phytogeography and endemism have been studied in
the Aegean (Rechinger, 1949; Rechinger & RechingerMoser, 1951; Snogerup, 1967a; Runemark, 1969, 1970,
1971a, b, Strid, 1970, 1972, 1996; Greuter, 1972, 1975;
Cardona & Contandriopoulos, 1978; Carlström, 1987;
Christodoulakis, 1996; Panitsa et al., 2006; Panitsa,
Tzanoudakis & Sfenthourakis, 2008), Peloponnisos
(Iatrou, 1986) and Evvoia (Trigas, 2003). The origin
and distribution patterns of the Greek flora and the
endemics have been connected to palaeogeographical
patterns (e.g. Turrill, 1929; Rechinger, 1965). The
palaeogeographical evolution of the area of Greece, the
Hellenic orogen, a composite segment of the Alpine
orogen, is relatively recent and complex. The Hellenic
arc lies in an active geological region created by the
late Mesozoic and Cenozoic collision of the European
and African plates and is the result of processes
including south(west)ward rollback of subducting
African lithosphere and associated backarc extension,
gravitational collapse and westward extrusion of Anatolia (e.g. Doutsos et al., 1993; van Hinsbergen, 2004;
Mountrakis, 2006). The mountain range of the Hellenides with a continental mass the Aegais connected the
Balkans with Asia Minor until the Miocene (13 Mya)
when the fragmentation of Aegais started (Creutzburg,
1966; Dermitzakis, 1990). Crete (Kriti) in the South
Aegean Arc was first separated 10 Mya (Creutzburg,
1966; Sissingh, 1976; Dermitzakis, 1990) and 9 Mya
the mid-Aegean Trench separated the eastern Aegean
islands and Asia Minor from the Cyclades and mainland Greece (Dermitzakis & Papanikolaou, 1981).
During the Messinian desiccation of the Mediterranean, connection among certain areas was
re-established (Hsü et al., 1978; Andersen et al., 2001),
although deep canyons with unfavourable climatic
conditions must have acted as dispersal barriers. By
early Pliocene the South Aegean Arc split once more
and Kriti was permanently isolated (Meulenkamp,
1985; Beerli, Hotz & Uzzell, 1996), while the North
Anatolian fault propagated into the north Aegean
region in late Pliocene time (Dinter, 1998). The onset of
a Mediterranean-type climate is placed in the mid-
131
Pliocene (Suc, 1984; Pons & Quézel, 1985; Bertoldi, Rio
& Thunell, 1989; Suc et al., 1994). During the Pliocene
and Pleistocene, certain areas were submerged and
then re-emerged and connections among islands and
mainland landmasses were established or broken in a
complex sequence of events affecting the organisms
inhabiting the area (Dermitzakis & Papanikolaou,
1981; Dermitzakis, 1990; Schúle, 1993; Sfenthourakis,
1996; Hausdorf & Hennig, 2005; Kasapidis et al., 2005;
Poulakakis et al., 2005; Simaiakis & Mylonas, 2006;
Comes, Tribsch & Bittkau, 2008).
The extant Greek flora is typically Mediterranean
with ‘Palaeo-Mediterranean’ tropical and subtropical
taxa which evolved during the pre-Pliocene and
Pliocene (remnants of a flora that became extinct
during the glaciations) and ‘Neo-Mediterranean’ nontropical taxa which appeared after the establishment
of Mediterranean type climate (for a review see
Blondel & Aronson, 1995). The flora has been influenced by east-to-west migration during the Miocene
and by a north-to-south migration in consecutive
waves (Iatrou, 1986). A part of the endemic flora
represents species the ancestors of which had a
broader distribution during the Tertiary (Turrill,
1929). The South Aegean element of the flora correlated to the South Aegean Arc reflects the situation
during the Miocene and the Cardaegean element (the
Cretan area and the South and Central Cyclades)
reflects the situation during the Pleistocene (Greuter,
1971, 1975). In the Cyclades there has been active
differentiation of endemics inside numerous islands
(Cardona & Contandriopoulos, 1978). The endemic
flora of Kriti has a relict character, but with a nonnegligible neoendemic element (Greuter, 1972;
Cardona & Contandriopoulos, 1978). Recent molecular
studies (Bittkau & Comes, 2005; Edh, Widén & Ceplitis, 2007) support the theory that plant diversification
and speciation in the Aegean region is driven mainly
by random (genetic drift) rather than adaptive differentiation among isolated populations (Runemark,
1969; Strid, 1970; Snogerup, Gustafsson & von
Bothmer, 1990; Thompson, 2005). The flora of Peloponnisos and the mountain flora are characterized by a
large proportion of neoendemics, indicating active speciation processes (Strid & Papanikolaou, 1985; Iatrou,
1986) and, on the whole, it is assumed that there are
more neo-endemics than palaeondemics in the Greek
flora (Tan & Iatrou, 2001; Thompson, 2005). According
to the above sources, it seems that two processes, the
evolution of habitat specialists (e.g. serpentine endemics) and of vicariant taxa (e.g. islet specialists) have
been important in endemic species formation.
The aim of this paper is to present, for the first
time, quantitative data on the taxonomy, geographical
and altitudinal distribution, ecology (life form) and
conservation status of all the Greek endemic plants
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
132
K. GEORGHIOU and P. DELIPETROU
and investigate their traits and possible relationships. The study of the above aspects, besides revealing general patterns of endemism in Greece, can also
be used for conservation purposes. The pattern of
distribution of species in genera and families is useful
in making decisions targeting conservation effort
(Fenner, Lee & Bastow Wilson, 1997). The same is
true for geographical and altitudinal distribution (e.g.
Gómez Campo & Malato Beliz, 1985; Cowling &
Samways, 1995a; McDonald & Cowling, 1995; Médail
& Verlaque, 1997; Roos et al., 2004) and species–area
relations can be used to predict extinction (Reid,
1992; Whittaker & Fernández-Palacios, 2007). Life
form, a functional or adaptive plant attribute (Box,
1987; Médail, Roche & Tatoni, 1998), seems to have a
predictive value for conservation in the Mediterranean (Verlaque, Médail & Aboucaya, 2001) and for the
impacts of global climate change (Broennimann et al.,
2006; Thuiller et al., 2006). Inherently, this work is
intended as an incentive for further research into and
cooperation on the knowledge, understanding and
hence the conservation of the endemic plants.
MATERIAL AND METHODS
The data set used in this study was derived from the
database Chloris (Georghiou & Delipetrou, 1990–
2009). Chloris is a relational MS Access database with
coded data on the taxonomy, distribution, biology,
ecology, conservation and protection status of the
endemic, subendemic, rare, threatened and protected
plants of Greece. It is regularly updated and currently
contains data on 3222 taxa of Pteridophyta and Spermatophyta based on all the relevant floras, another
1130 published literature sources and 180 not published (including Internet) valid sources.
The term ‘Greek endemic’ denotes vascular plant
taxa that occur only within the political borders of
Greece. This is quite straightforward for the islands,
but there are at least eight stenoendemic mountain
taxa occurring just on both sides of the borders with
Balkan countries. Such taxa were not included in
data analysis. However, it must be noted that the
inclusion of these taxa in the data sets used for the
calculation of indices or in statistical analysis had no
significant effect on the results (data not shown).
Distribution data are coded using the 13 floristic
regions of Greece (Fig. 1) in the Flora Hellenica
project (Strid & Tan, 1997) in which the division of
the Aegean was mainly based on Rechinger (1943a,
1949), confirmed by floristic data from Strid (1996).
The continental regions are: Peloponnisos (Pe), Sterea
Ellada (StE), South Pindos mountain range (SPi),
North Pindos mountain range (NPi), East Central
Greece (EC), North Central Greece (NC), North-East
Greece (NE). The island regions are: West Aegean
islands, including the large island of Evvoia (WAe);
Ionian Islands (IoI), North Aegean islands (NAe), the
island complex of the Cyclades (Kik), Cretan area
including the large island of Kriti and its surrounding
islets and the islands of Kasos and Karpathos (KK),
East Aegean islands (EAe). Life form data are coded
according to Raunkiaer (Ellenberg & Müller-Dombois,
1966; Pignatti, 1982).
The vegetation zones used in the altitudinal distribution of plants correspond to the altitude range
classes in Table 1. This is a rough correspondence, as
the exact range of each zone in Greece varies, not only
depending on latitude but also locally (e.g. southern
and northern slopes).
Species–area relationships for all species (SARs)
and for endemic (i.e. endemic to one floristic region)
species (EARs) within the Greek floristic regions were
investigated assuming the ‘power equation’ with logarithmic transformation:
log S = c + zlogA (S = species number; A = area;
c, z = constants)
Hierarchical cluster analysis was carried out using
the STATISTICA software and was based on the
presence/absence of species or taxa in each floristic
region. A preliminary analysis was made with the use
of three linkage methods (single, complete and
unweighted pair-group average), each applied using
four measures of similarity: Dice (Chekanowski or
Sørensen, matches weigh double, joint absences
excluded), Jaccard (equal weight to matches and nonmatches, joint absences excluded), Lance–Williams
(Bray–Curtis, complementary to Dice) (eqn. 1) and
the Preston index (eqn. 2), which is not greatly
affected by large differences in the number of taxa (or
the area) of the regions compared (Preston, 1962).
Indices that include joint absences (e.g. Hamman)
were deemed inappropriate because a large number of
taxa occur in only one floristic region and are absent
from all others and this would result in artefactual
relationships between non-similar regions.
Lance − Williams index = ( b + c ) ( 2a + b + c )
(1)
where a = number of taxa present in both of the pair
of regions compared; b and c = taxa present on one
region but absent from the other.
n
n
( F1 F1 + 2 ) + ( F2 F1 + 2 ) = 1
(2)
Preston index z = 1/n; F1 and F2 = number of species
present in each of the pair of regions compared;
F1+2 = total number of species present in the pair of
regions compared. The equation was solved for each
pair of regions by the MSExcel solution tool.
The available data on the conservation status of the
Greek endemic plants (see references in the relevant
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE
133
Figure 1. The floristic regions of Greece (Strid & Tan, 1997).
Table 1. Correspondence of vegetation zones to altitude range classes
Zone
A
B
C
D
E
Altitude
range (m)
0–600
600–1000
1000–1500
1500–2000
2000–3000
Description
Thermo-Mediterranean: Eu-Mediterranean matorral and thermophilous conifer zone (coastal,
hilly, submountainous)
Meso-Mediterranean: Sub-Mediterranean shrub and forest (hilly, submountainous)
Supra-Mediterranean: Fagus and Abies forest zone (including sub-Mediterranean conifers)
Mountain Mediterranean: Boreal conifer (e.g. Picea) zone
Oro-Mediterranean/Alpine. No forests
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
134
K. GEORGHIOU and P. DELIPETROU
Table 2. Species diversity and endemism in regions of the Mediterranean, Europe, Africa and South America
Region
Area km2
Native
species
sp/logarea
logsp/
logarea
Endemic
species
%
Endemic
Source*
Greece
Cyprus
Bulgaria
Italy (territory)
France (territory)
Peninsular Spain (+ Balearics)
Canary Islands
UK
Turkey
Morocco
Southern Africa
Madagascar
Chile
California (MTE)
Argentina
Panama
Venezuela
132 700
9 251
111 000
301 049
551 700
501 000
14 879
244 754
779 500
710 850
2 570 000
587 000
462 820
242 519
2 770 000
75 648
1 148 154
5 855
1 612
3 600
5 300
4 650
7 920
1 800
1 800
8 650
3 675
20 372
10 000
4 669
5 050
9 370
9 900
21 070
1143
406
714
967
810
1390
431
334
1468
628
3178
1734
824
938
1454
2029
3477
0.74
0.81
0.70
0.68
0.64
0.68
0.78
0.60
0.67
0.61
0.67
0.69
0.65
0.69
0.62
0.82
0.71
913
110
320
712
133
550
550
16
2 675
625
16 298
8 000
2 700
1 515
1 100
1 222
8 000
15.6
6.8
8.9
13.4
2.9
6.9
30.6
0.9
30.9
17.0
80.0
80.0
57.8
30.0
11.7
12.3
38.0
1, 2, 3
5, 4
6
7
6
8
8
6, 9
10
6
11
6
6, 12
13
6
6
6
*Sources: 1, Strid & Tan, 1997; 2, Tan & Iatrou, 2001; 3, Chloris database; 4, database CyprusFlora; 5, Hadjikyriakou,
1997; 6, WWF & IUCN, 1994; 7, Pignatti, 1995; 8, Bueno et al., 1995; 9, WWF & IUCN, 1988; 10, Davis, 1965–1986; 11,
Cowling & Hilton-Taylor, 1994; 12, Fuentes et al., 1995; 13, Keeley & Swift, 1995.
section of the results) include classification in: (1) the
old, pre-1994 IUCN categories (Lucas & Synge, 1978);
(2) the new IUCN categories, according to the 1994
version (Mace & Stuart, 1994); and (3) the new IUCN
categories, according to the revised 2001 version
(IUCN, 2001). The conservation status of the plants
that have been classified only according to the pre1994 IUCN categories [i.e. extinct (Ex), endangered
(E), vulnerable (V), rare (R), not threatened (nt),
indeterminate (I), inadequate data (?)] was not transferred to the new categories because there is no
reliable correspondence, except from the old category
inadequate data (?), which was transferred to the new
category Data Deficient (DD) and from the old category not threatened (nt) which was transferred to
the new category Least Concern (LC). The conservation status of the plants classified according to the
1994 version was transfered to the 2001 version on a
case-by-case reassessment.
RESULTS
BIODIVERSITY
AND ENDEMISM
Taking species richness in relation to area size as a
measure of biodiversity, Greece, with an estimated
6437 native plant taxa or c. 5800 species (Strid &
Tan, 1997, 2002; Tan & Iatrou, 2001), ranges among
the highest in European and Mediterranean or
Mediterranean-type climate areas (Table 2).
Table 3. Greek endemic species and taxa
Endemic Endemic
Doubtful
sexual
apomictic ? Endemic taxa
Total
Species*
913
Subspecies 345
Varieties
24
Taxa
1226
65
1
66
27
28
2
52
56
37
5
98
1061
411
31
1442
*Fifty species with infraspecific subdivisions.
The vascular plant species recorded in the database
Chloris as endemic to Greece number 1061 (1442
taxa) (Table 3). Even excluding apomictic species and
species doubtfully endemic or of doubtful taxonomic
status, this number becomes 913 species (1226 taxa).
Species endemism reaches 15.6% of the total native
flora which, taking into account the small area of the
country, is quite high in comparison with other
Mediterranean-type climate and European countries
(Table 2). Using Bykov’s index (le = endemism factual/
endemism normal) (Fig. 2) as a quantitative measure
of endemism (Cowling & Samways, 1995b), Greece
has 2.1 times the normal endemism for its area,
which is lower than South Africa (le = 3.6) or the
exceptionally high-ranking Canary Islands (le = 9.3),
but comparable with Turkey and Cyprus (le approximately 2) and higher than Bulgaria, Italy, Peninsular
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE
135
Figure 2. Bykov’s nomogram. The diagonal line is defined by 100% endemism for the earth and by 1% endemism in
625 km2 and depicts normal endemism (En). Points represent the factual endemism (Ef). Areas above the diagonal line
have less than normal endemism and areas below the line have more. Data for Greece and the Greek floristic regions do
not include doubtful or apomictic species.
Spain and the Balearics and Morocco (le = 0.7–1.2).
Within the floristic regions of Greece, the Cretan area
(le = 3.1) is indicated as the most important centre of
endemism, a well-known fact commented upon by
Greuter (1972), Rechinger (1949, 1965) and Strid,
1986. Peloponnisos and the West Aegean (le = 1.2) are
the only other regions exceeding the expected endemism. The East Aegean islands and the Cyclades
have close to normal levels of endemism (le = 0.8).
FLORISTIC
ANALYSIS
The endemic species (apomictic and doubtful species
included) belong to 53 families and 239 genera. There
are no endemic families and there are six monotypic
endemic genera: Horstrissea Greuter, P.Gerstberger &
B.Egli, Jankaea Boiss., Peltaria Jacq., Petromarula
Vent. ex R.Hedw., Phitosia Kamari & Greuter and
Thamnosciadium Hartvig. The families richer in
endemic species in absolute numbers (Fig. 3) are Asteraceae, Caryophyllaceae, Brassicaceae and Lamiaceae.
Their degree of endemism (27, 30, 24 and 22%, respectively) is higher than that of the general flora (15.6%).
However, the highest degree of endemism is seen in
Campanulaceae (49%). At infraspecific level, the
endemic taxa belong to 58 families and 281 genera.
The genera of the endemic species are presented in
Figure 4. The total species/genus ratio in the endemics is 4.4, and 45% of the genera have only one
endemic species. However, certain genera, notably
Centaurea L., Campanula L., Silene L., Allium L.,
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
136
K. GEORGHIOU and P. DELIPETROU
Figure 3. The families of the Greek endemic species: sexual species (striped bar), apomictic species (open bar) and
doubtful species (black bar).
Verbascum L., Ophrys L., Viola L. and Stachys L.,
show a high degree of species richness and endemism
(endemic/all species: 61/122, 42/77, 38/119, 39/79,
23/72, 40/90 and 17/52, respectively). [Species concepts and the number of species recognized in Ophrys
vary considerably between authors. Here, a relatively
narrow concept is used, but it should be noted that
this may result in an over-representation of this
genus in these analyses. For example, in the recent
treatment of Pedersen & Faurholdt (2007), no Ophrys
spp. are endemic to Greece, although some infraspecific taxa are.]
DISTRIBUTION
IN FLORISTIC REGIONS
The majority of the endemic species are restricted to
only one floristic region (Fig. 5) and none occurs in all
13 regions. The distribution across regions (Fig. 6)
shows that southern and eastern Greece is richer in
absolute numbers of endemics. Peloponnisos, the
Cretan area and Sterea Ellada are the regions with
the highest numbers of endemic species, in descend-
ing order, and the Cretan area has the highest
number of local endemics (173 species, 235 taxa,
excluding apomictic and doubtful taxa).
The species–area relationships for all species (SARs)
and for endemic species (EARs) are shown in Figure 7.
The SAR for the 13 floristic regions (species numbers
according to Tan & Iatrou, 2001) was highly significant
with a z-value (the regression coefficient) of 0.20 and
explained a considerable amount of species number
variation (76%). The analysis for continental and
island regions separately resulted in a significant SAR
with a steeper slope (z = 0.38) and a high R2 (0.87) for
the continental regions and, in SAR significant only at
1% error level, with a shallower slope (z = 0.14) and
lower R2 (0.54) for the island regions. The EAR for all
the floristic regions was not significant and explained
only a low percentage of species number variation. The
EAR for the island regions was significant with a steep
curve (z = 1.15) and 82% of the variation in the number
of endemics was explained by the total size of the
islands per region. The EAR for the continental regions
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE
137
Figure 4. The genera of the Greek endemic species: sexual species (striped bar), apomictic species (open bar) and
doubtful species (black bar).
resulted in an equally steep curve (z = 1.49), which
explained 46% of the variation in the number of
endemics but was significant only at 1% error level.
The respective c-values (island c = -2.5, continental
c = -4.7) indicate that continental regions with a lower
initial slope have fewer endemics per unit area. All
types of EARs gave similar results when the doubtful
and apomictic species were excluded or for the endemic
taxa, i.e. with subspecies and varieties included. The
performance of the semi-log model was worse in all
cases (data not shown).
Analysis of the distributional pattern of the endemics and of the relations among floristic regions, based
on their common endemic taxa, was made by hierarchical clustering (Fig. 8). The preliminary analysis
resulted in two distinct classification patterns for
each linkage method, which were essentially similar
for the species and the taxa data sets, regardless of
whether apomictic and doubtful taxa were included:
one using the Lance–Williams dissimilarity coefficient, which was identical to the classification with
Dice and Jaccard, and one using the Preston z dis-
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
138
K. GEORGHIOU and P. DELIPETROU
Figure 5. Greek endemic species occurring in 1–10 floristic regions: sexual species (open bar), apomictic species
(black bar) and doubtful species (grey bar). The graph
includes 1059 species with adequate data.
Figure 6. The distribution of the Greek endemic and local
endemic species in floristic regions: Greek endemics,
sexual (striped bar); Greek endemics, apomictic or doubtful (grey bar); local endemics, sexual (open bar); local
endemics, apomictic or doubtful (black bar). The graph
includes 1059 species with adequate data.
similarity coefficient. The differences in the resulting
clustering were more marked among the two indices
than among the three linkage methods, and the
unweighted pair-group average (UPGMA) was finally
selected as it is based on information about all intercluster pairs (Garson, 2008).
The analysis for all taxa (including subspecies and
varieties) seems advantageous because the taxonomic
position of several taxa is not clear and exclusion of
endemics of infraspecific rank might screen cases of
actual endemism. Nevertheless, the results were
essentially similar when the analysis was performed
at species level. Table 4 (unshaded part) shows the
Figure 7. SARs: regression lines for all floristic regions
(S); insular regions (Si); and continental regions (Sc).
EARs: regression lines for all regions (E); insular regions
(Ei); and continental regions (Ec). The analysis included
1059 endemic species with adequate data (doubtful and
apomictic species included).
S: y = 0.2009x – 2.5687 (n = 13, R2 = 0.76353, Fs =
35.518, P = 0.0001)
Si: y = 1.1607x – 2.5005 (n = 6, R2 = 0.83705, Fs = 20.548,
P = 0.01055)
Sc: y = 1.5600x – 4.9890 (n = 7, R2 = 0.47862, Fs = 4.590,
P = 0.08506)
E: y = 0.4188x – 0.0857 (n = 13, R2 = 0.16086, Fs = 2.109,
ns)
Ei: y = 1.1536x – 2.4915 (n = 6, R2 = 0.81493, Fs = 17.613,
P = 0.01373)
Ec: y = 1.4853x – 4.6609 (n = 7, R2 = 0.46330, Fs = 4.316,
P = 0.09235)
actual numbers of common Greek endemic taxa
between regions and Table 5 demonstrates the similarity matrices used in clustering, the Preston index
(unshaded part) and the Lance–Williams coefficient
(shaded part). Both indices gave rather low similarity
measures, apparently because of the presence of a
large number of taxa restricted to only one region.
The two distributional patterns (Fig. 8) are quite
similar with certain features in common.
The most prominent of these features are: (1) the
Central, South and East Aegean islands (Kik, KK,
EAe) comprise a unit showing greater similarity
among themselves than with any other region; (2) the
Western Aegean (WAe) and the Northern Aegean
(NAe) have greater affinity to the adjacent mainland
areas rather than to the remainder of the Aegean
islands; and (3) each continental region has greater
similarity with its adjacent regions, the highest similarities shown in two pairs, Pe and StE in Southern
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE
139
is connected to EAe and Pe. There is also a closer
connection of EAe with Kik than with the KK.
The endemic taxa occurring in only two regions
were also examined (Table 4), as the existence of
biregional endemics is a good indication of phytogeographical connections between regions. Table 4 shows
a clearer picture of the relations between regions and
underlines the above-noted affinities, such as the
Pe–KK connection (22 biregional endemics, 12 of
them from the islands of Kythira and Antikythira)
and the EAe–Kik connection. Of the eight taxa
endemic to Kik and EAe, two occur only on Ikaria in
EAe, whereas the distribution of the other six in EAe
includes at least one of the islands of Ikaria, Samos
and Chios. Besides, all three WAe–EAe biregional
endemics occur in the Ikaria–Samos area. The connection of NAe to NE is indicated by seven biregional
endemics, all of which occur on the islands of Thasos
and/or Samothraki.
ALTITUDINAL
Figure 8. Dendrogram of the hierarchical cluster analysis of the distribution of Greek endemic taxa in floristic
regions. The analysis included 1328 taxa with adequate
data (doubtful taxa excluded). A, analysis with the
Preston coefficient. B, analysis with the Lance–Williams
coefficient.
Greece and SPi and NPi in Pindos. The main difference between the two distributional patterns is that
the Preston index grouped NAe with the adjacent NE
mainland and IoI with the adjacent regions of
Western Greece rather than separate from all other
regions; also, it grouped EC with the Southern Greece
regions rather than with the more distant NE.
The similarity matrices (Table 5) highlight certain
connections not apparent in the classification. The
Ionian Islands (IoI) are closest to SPi and the other
two regions with west-facing coastal parts, Pe and
StE. Because of the proximity of the large island of
Evvoia to the mainland, WAe firstly shares endemic
species with StE and, secondly, with the other two
regions with Aegean-facing coasts and islets, EC and
Pe, but maintains a relation with Kik in the central
Aegean. In the first instance, KK is closer to Kik, with
which it comprises the ‘Cardaegean’, and, secondly, it
DISTRIBUTION
The occurrence of the Greek endemic species at each
altitudinal zone (zones A to E, Table 1) shows that
they can be found at all altitudes up to c. 3000 m (Mt
Olympos summit at 2917 m). There is a rise at altitudes from 0 to 600 m and a gradual fall at higher
altitudes, more marked above 2000 m (Fig. 9). It
should, however, be kept in mind that such high
altitudes represent a small percentage of the territory
of Greece. This general altitudinal distribution
pattern is observed in the local endemics of the floristic regions of Pe and KK (Fig. 9A). In the other
island regions, except NAe (Fig. 9A), the fall is
steeper at altitudes above 600 m, apparently because
of the lack of high altitudes on most islands. However,
in continental Greece, including StE, NE and EC
(Fig. 9B), the endemics are concentrated at altitudes
above 600 m. This pattern is more pronounced in the
high-altitude mountainous areas of SPi, NPi and NC
(Fig. 9B). On the one hand, although the endemics
occur at all altitudes, the altitudinal range (Fig. 10) of
each of them is not as wide: nearly half of them have
a range of 0–600 m and a quarter of them are
restricted to only one altitude or within a range of
200 m. On the other hand, a significant 31% has an
altitudinal range of over 1000 m. The detailed distribution of endemics to altitudinal zones (Fig. 11A,
main graph) shows that 35% of them occur in only one
zone, 23% only at the thermo-Mediterranean zone
(0–600 m) and 15% only at the mountain- and/or
oro-Mediterranean zones (above 1500 m). However, a
considerable number of species (23%) occur in three to
five altitudinal zones. The altitudinal zone pattern is
differentiated for local endemics in the floristic
regions (Fig. 11B). In the island regions, there is a
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
140
K. GEORGHIOU and P. DELIPETROU
Table 4. Common endemic taxa between floristic regions in absolute numbers (unshaded area) and distribution of
biregional endemics (shaded area)
Pe
StE
WAe
IoI
SPi
NPi
EC
NC
NE
NAe
Kik
KK
EAe
Pe
StE
WAe
IoI
SPi
NPi
EC
NC
NE
NAe
Kik
KK
EAe
423
47
3
8
0
0
2
1
0
0
3
22
0
188
352
14
6
8
2
2
7
0
0
2
1
0
85
112
207
0
1
0
3
0
3
1
11
0
3
52
53
17
90
1
0
0
0
0
0
0
1
0
49
82
26
30
121
7
0
4
1
0
0
0
0
35
58
17
19
64
121
0
7
0
0
0
0
0
32
46
33
11
26
19
74
7
1
0
0
0
0
30
57
23
15
47
43
32
158
2
0
0
0
0
22
32
27
8
15
12
21
28
106
7
0
0
0
10
12
14
6
5
4
8
5
14
46
1
0
3
46
43
49
13
6
3
6
3
9
8
157
23
8
72
40
31
14
8
5
7
3
6
6
66
371
11
36
33
32
11
7
3
8
8
7
11
50
51
148
The bold type indicates the Greek endemic taxa in each region. The analysis included 1328 taxa with adequate data
(doubtful taxa were excluded).
Table 5. Dissimilarity coefficients between pairs of floristic regions: Preston z (unshaded area); Lance–Williams coefficient (shaded area)
Pe
StE
WAe
IoI
SPi
NPi
EC
NC
NE
NAe
Kik
KK
EAe
Pe
StE
WAe
IoI
SPi
NPi
EC
NC
NE
NAe
Kik
KK
EAe
0.00
0.51
0.73
0.80
0.82
0.87
0.87
0.90
0.92
0.96
0.88
0.82
0.87
0.60
0.00
0.60
0.76
0.65
0.75
0.78
0.78
0.86
0.94
0.83
0.84
0.87
0.76
0.65
0.00
0.89
0.84
0.90
0.77
0.87
0.83
0.89
0.73
0.89
0.82
0.74
0.72
0.91
0.00
0.72
0.82
0.87
0.88
0.92
0.91
0.89
0.94
0.91
0.81
0.63
0.87
0.77
0.00
0.47
0.73
0.66
0.87
0.94
0.96
0.97
0.95
0.87
0.76
0.91
0.86
0.56
0.00
0.81
0.69
0.89
0.95
0.98
0.98
0.98
0.83
0.70
0.77
0.90
0.78
0.84
0.00
0.72
0.77
0.87
0.95
0.97
0.93
0.91
0.81
0.91
0.90
0.73
0.75
0.76
0.00
0.79
0.95
0.98
0.99
0.95
0.91
0.86
0.86
0.94
0.91
0.92
0.82
0.84
0.00
0.82
0.93
0.97
0.94
0.92
0.90
0.87
0.93
0.95
0.95
0.90
0.95
0.84
0.00
0.92
0.97
0.89
0.86
0.86
0.79
0.92
0.97
0.99
0.96
0.99
0.95
0.92
0.00
0.75
0.67
0.87
0.92
0.92
0.94
0.97
0.98
0.96
0.99
0.97
0.95
0.77
0.00
0.80
0.89
0.89
0.86
0.93
0.97
0.99
0.94
0.96
0.96
0.89
0.74
0.83
0.00
The analysis included 1328 taxa with adequate data (doubtful taxa were excluded).
predominance of local endemics restricted to or occurring at the thermo-Mediterranean zone, which
decreases as the coverage in high mountains
increases (e.g. KK and NAe). Conversely, most of the
local endemics of the continental regions are
restricted to zones above 1000 or 1500 m.
BIOLOGY
Life form is the functional attribute investigated in
the endemic species (Table 6). The hemicryptophytes
(93% scapose, caespitose or rosette-forming perennials and 7% biennials) are by far the most frequent,
followed by the chamaephytes (83% subshrubs or
dwarf shrubs, 15% mat-forming or prostrate and 1%
succulent). The proportions of geophytes and annuals
(therophytes) are significantly lower and only 1% of
the endemic species are phanerophytes (mostly nanophanerophytes).
In Greece, as shown by the floras of various areas
(Table 6, Voliotis, 1982; Vogiatzakis, Griffiths &
Mannion, 2003), the life form spectra typically shift
from a predominance of annuals at lower altitudes and
southern regions with higher temperatures and
extended drought periods to a predominance of hemicryptophytes and chamaephytes at higher altitudes
and northern regions with lower temperatures and
higher rainfall. Compared with this trend for the total
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE
141
Figure 10. Frequency classes of the altitude range of the
Greek endemic species. The analysis included 860 species
with adequate data (doubtful and apomictic species
excluded).
Figure 9. Occurrence of the Greek endemic species in
altitudinal zones (all species occurring at the indicated
range are included) in Greece (bars, secondary y axis) and
of local endemics in each floristic region (lines, primary y
axis). A, insular regions. B, continental regions. The analysis included 860 species with adequate data (doubtful and
apomictic species excluded). Altitudinal zones are defined
in Table 1.
flora, the life form spectrum of the endemics, with a
predominance of perennial hemicryptophytes and
chamaephytes and low proportions of annuals, is closer
to the spectra for high mountains, with the exception of
the usually high proportions of geophytes and chamaephytes among the endemics. A similar trend is
observed in Cyprus where, for the total flora, the
proportion of annuals is higher and the proportion of
hemicryptophytes is lower than for the endemic flora
(Table 6, Hadjikyriakou, 1997). The obligate chasmophytes (75% endemic) of Kriti are mainly chamaephytes and hemicryptophytes as opposed to the
general chasmophytic flora which has an increased
proportion of annuals (Table 6, Kypriotakis, 1997). In
the regional life form spectra of the endemics
(Fig. 11B) it is shown that southern warmer regions
(e.g. EAe) have higher proportions of annuals and
northern regions (e.g. NC) have higher proportions of
hemicryptophytes. This indicates that the trend of the
total flora is repeated in the endemic flora, although
the percentages differ. Pe and StE, with high mountains, extensive coastlines and offshore islets, are close
to the general pattern for the endemics. In order to
investigate whether the life form of the endemics is
related to their altitude range, and therefore to bioclimate, the life form spectra for plants of each altitudinal zone were plotted (Fig. 11A, inserted graph). There
is a trend for a decrease of annuals and geophytes and
an increase of hemicryptophytes in plants occurring at
or restricted to the higher altitudes, but the life form
spectra of the endemics still maintain their differences
from the total flora spectra. The other life forms
maintain more or less the same pattern.
CONSERVATION
STATUS
The main sources for the conservation status of the
Greek endemics are the Red Data Book of Rare and
Threatened Plants of Greece (Phitos et al., 1995) and
the IUCN-WCMC database (IUCN-WCMC, 1997), in
which 873 taxa in total were classified according to
old IUCN categories. A preliminary classification in
new IUCN categories (1994 version) was attempted
for the plants of Southern Greece, including 189
endemic taxa (Georghiou & Delipetrou, 2001), and for
50 taxa by the Mediterranean Island Plant Specialist
Group (De Montmollin & Strahm, 2005). Valid literature sources (e.g. Iatrou, 1986; Trigas, 2003) and
recent unpublished field work provide additional
information on the conservation status or rarity of
321 endemic taxa.
The compiled information on the conservation
status of the endemics is presented in Figure 12, with
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
142
K. GEORGHIOU and P. DELIPETROU
Figure 11. A, altitudinal distribution in zones (only species restricted at the indicated zone are included) of the Greek
endemic species and their life forms in counts (main graph) and percentages (inserted graph). T, annuals (Therophytes);
G, geophytes; H, hemicryptophytes; C, chamaephytes; P, phanerophytes. B, altitudinal distribution in zones of the local
endemics in each floristic region (A, B, C, D, E represent species restricted to each zone; A–E, B–E, C–E, D–E represent
species occurring in more than one zone in the indicated range). The analysis included 860 species with adequate data
(doubtful and apomictic species excluded). Altitudinal zones are defined in Table 1.
the plants characterized according to the old (Lucas &
Synge, 1978) or the new (IUCN, 2001) IUCN threat
categories, depending on the available data. There are
253 (18% of the total) threatened (critically endangered, endangered or vulnerable) and seven extinct or
doubtfully extinct Greek endemic taxa. Another 369
taxa (26% of the total) have been characterized as
indeterminate or rare and, in our experience, these
usually turn out to be either vulnerable (VU) or
near-threatened (NT). However, a significant 19% of
the plants are apparently of least concern (LC).
There are few known cases of extinction in the
endemic Greek flora. Two species are in all probability
extinct. Ammi topalii Beauverd, known only from the
type specimen collected in 1935 (Rechinger, 1961) was
not found in a recent search of its location and may
have disappeared as a result of the change in land use
(Iatrou & Trigas, 1996; Trigas, 2003). Geocaryum
bornmuelleri (Wolff) Engstr., also known from a single
gathering on the island of Thasos in 1891 (Greuter,
1995), has also not been refound (Strid & Tan, 1997).
The doubtfully extinct taxa are Brassica cadmea
Heldr. ex O.E.Schulz, Centaurea tuntasia Heldr. ex
Halácsy, Cerastium illyricum Ard. ssp. crinitum
(Lonsing) P.D.Sell & Whitehead, Geocaryum divaricatum (Boiss. & Orph.) Engstr. and Tetragonolobus
wiedemannii Boiss. Most of them are inadequately
known, and the last three are of doubtful taxonomic
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE
143
Table 6. Life form spectra in floras of Greek areas
Life form (%)†
Area
T
G
H
Ch
P
Greek endemics*
NC
SPi
NE
NPi
WAe
Pe
StE
EC
KK
IoI
NAe
Kik
EAe
Vrontous Mt (1849 m, NE)
Voras Mt (2524 m, NC)
Vermion Mt (2052 m, NC)
Vermion 500–1000 m
Vermion 1000–1650 m
Vermion 1650–2052 m
Menoikio Mt (1963 m, NE)
Kyllini Mt (2376 m, Pe)
Kyllini 0–600 m
Kyllini 2000–2376 m
Cholomon Mt (1165 m, NE)
Central Evvoia Island (WAe)
Vertiskos Mt (1103 m, NE)
Sithonia peninsula (NE)
Samos Island (EAe)
Cretan area (KK)
Kriti – chasmophytes
Kriti – obligate chasmophytes
Geraneia Mt (1269 m, StE)
Skiathos Island (WAe)
Skopelos Island (WAe)
Kythira Island (Pe)
Nisyros Island (EAe)
Arkoi–Leipsoi islet complex (EAe)
Cyprus
Cyprus endemics
11
7
8
9
11
12
12
12
16
17
18
19
19
21
13
14
15
35
18
4
20
24
39
10
26
29
30
42
42
42
31
0
43
45
47
52
55
59
49
11
17
14
24
16
15
18
14
17
14
19
25
14
27
27
12
11
14
16
16
9
6
13
12
9
15
8
10
9
15
15
11
3
14
12
13
12
9
15
17
23
44
57
45
52
55
43
44
44
52
31
34
49
34
30
54
57
51
26
37
66
54
43
31
55
32
43
42
30
24
21
22
30
26
25
19
17
17
15
21
40
27
21
23
22
20
27
28
24
18
31
21
16
17
19
8
7
10
6
9
17
10
10
5
23
12
10
5
8
10
14
26
60
11
7
10
11
9
6
5
22
1
1
1
1
0
1
1
2
0
3
2
3
3
4
12
10
10
17
20
4
10
10
13
3
15
4
11
9
9
6
10
7
6
8
9
8
10
5
7
4
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Chloris database
Voliotis, 1977
Voliotis, 1979
Chochliouros, 2005
Chochliouros, 2005
Chochliouros, 2005
Chochliouros, 2005
Karagiannakidou, 1991
Dimopoulos, 1993
Dimopoulos, 1993
Dimopoulos, 1993
Voliotis, 1967
Phitos, 1960
Pavlides, 1982
Pavlides, 1976
Christodoulakis, 1986
Jahn & Schönfelder, 1995
Kypriotakis, 1997
Kypriotakis, 1997
Constantinidis, 1997
Economidou, 1969
Economidou, 1973
Yannitsaros, 1969
Papatsou, 1975
Panitsa, 1997
Hadjikyriakou, 1997; CyprusFlora database
Hadjikyriakou, 1997; CyprusFlora database
*The analysis included 935 species with adequate data (doubtful and apomictic species excluded).
†T, annuals (Therophytes); G, geophytes; H, hemicryptophytes; C, chamaephytes; P, phanerophytes.
status (Routsi, 1993, Greuter, 1995; Strid & Tan,
1997, 2002). Some other allegedly extinct endemics
(Phitos et al., 1995; Tan & Strid, 1997), Alkanna
sartoriana Boiss. & Heldr. (Vassiliadis, 1996) and
Astragalus maniaticus Kit Tan & Strid (Tan & Iatrou,
2001), have been rediscovered. According to the
above, at the worst, only 0.5% of the endemic taxa are
extinct. However, 35% of the endemics have not been
evaluated at all (NE) or are insufficiently known
(DD), including at least 35 taxa that have not been
collected or looked for, for the last 60 to > 100 years.
DISCUSSION
The floristic analysis of the endemic taxa highlighted
certain families which are important at Mediterra-
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
144
K. GEORGHIOU and P. DELIPETROU
Figure 12. Conservation status of the Greek endemic
taxa (including doubtful and apomictic taxa). New IUCN
categories (IUCN, 2001): EX, extinct; ?EX, doubtfully
extinct; CR, critically endangered; EN, endangered; VU,
vulnerable; NT, near threatened; LC, least concern; DD,
data deficient; NE, not evaluated. Old IUCN categories
(Lucas & Synge, 1978): E, endangered; V, vulnerable; R,
rare; I, indeterminate.
nean level. Asteraceae have an excess of endemic
genera in East Asia and North America (Qian, 2001).
Caryophyllaceae show active endemism in Greece
(Trigas, Iatrou & Karetsos, 2007) and, with Brassicaceae, have large concentrations of genera in the
Mediterranean basin (Heywood et al., 2007). The
Mediterranean basin is a centre of diversification for
Lamiaceae (Heywood et al., 2007) and the evolutionary centre of all the Greek genera of Campanulaceae
(Eddie et al., 2003). The 10–12 families with the
highest numbers of endemic species more or less fall
within the richest families of the total Greek flora (as
indicated by the studied floras of several areas), a
trend also observed in other Mediterranean type
climate provinces (Beard, Chapman & Gioia, 2000). A
striking difference between the total and endemic
Greek flora is seen in Poaceae and Fabaceae which,
along with Asteraceae, invariably figure among the
three richest and most diverse families in Greece and
in the Mediterranean (Blondel & Aronson, 1995;
Maxted & Bennett, 2001), but rank lower in both
absolute number of endemics and degree of endemism
(6 and 10%, respectively) in Greece. Also, the 10–12
families richest in endemics in Greece coincide with
those in neighbouring Cyprus (Hadjikyriakou, 1997)
and Turkey (Davis, 1965–1986). The main differences
between the three countries are noted in Fabaceae
(higher rank in Turkey), Campanulaceae (higher rank
in Greece) and Crassulaceae (higher rank in Cyprus).
The total species/genus ratio (4.4) is somewhat
lower than the ratio for the total flora (5.5; Strid &
Tan, 1997) and medium compared with the 3.0–9.0 for
the floras of other Mediterranean-type climate
regions (Blondel & Aronson, 1995; Fenner et al.,
1997). The total Greek flora is characterized by a
majority of small-sized genera, with at least 473
genera represented by only one species (Strid & Tan,
1997), and this trend is repeated in the endemic flora.
The high endemism of certain genera, such as Centaurea, which comprises both relict and neoendemic
taxa (Georgiadis, 1980; Routsi, 1993), and Allium
may indicate recent speciation (Fenner et al., 1997).
The distribution of the endemics is characterized by
a high frequency of narrowly distributed species
(Fig. 5) and by a north-to-south and a west-to-east
increase of local endemism (Fig. 6). The first result
supports the notion that increased diversity is related
to high incidence of local combined with regional
endemism in Mediterranean basin floras (Greuter,
1991, 1995). The second result confirms previous
reports, based on subtotals of the endemic taxa, that
endemism in the mountain flora increases southwards, with taxa tending to be more narrowly distributed and reproductively isolated (Strid, 1986, 1993),
and that in the genus Centaurea there is an east-towest reduction of the number of endemics, with the
western taxa having evolved more recently (Georgiadis, 1980; Georgiadis, Iatrou & Georgiou, 1986). Both
Bykov’s indices for floristic regions (Fig. 2) and the
counts of endemic species as a function of area (Fig. 7)
are consistent with the above and show the island
regions to be centres of endemism.
The slope of the SAR for the total flora (z = 0.20) of
the floristic regions was at the margin of the range
(0.1–0.2) of z expected for the flora of mainland intraprovincial (nested sampling, sensu Rosenzweig, 1995,
2004) SARs or SARs expected for mainland areas
larger than 10 km2 (Crawley & Harral, 2001) or of
non-isolated islands (McArthur & Wilson, 1967). Such
SARs are interpreted as the result of accumulation of
habitat heterogeneity within a region/province
(Rosenzweig, 1995), implying that the floristic regions
of Greece constitute parts of the same province.
However, as the total flora SAR has not been constructed from a nested design, and as some floristic
regions are continental and some islands (i.e. with
different underlying processes for biodiversity), any
such theoretical interpretation is precarious. Indeed,
the SAR slope and explanatory value change when
the model is applied to continental and island regions
separately and has lower explanatory value for island
regions, indicating the greater involvement of other
factors; for example, habitat diversity and isolation
(Yeakley & Weishampel, 2000; Triantis et al., 2003;
Field et al., 2008). For instance, the longer isolated
KK has a relatively low total species number despite
the size and habitat diversity of the island of Kriti.
In all cases, EARs had slopes steeper than those of
the SARs for the total flora as predicted for log–log
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PATTERNS OF PLANT ENDEMISM IN GREECE
models by Harte & Kinzig (1997) and also confirmed
in several cases either for islands (Roos et al., 2004;
Triantis, Mylonas & Whittaker, 2008) or continental
nested areas (Ulrich, 2005), and even in the contiguous US (Harte & Kinzig, 1997). The steep curves
indicate a larger increase in the number of endemics
than the total species richness for the same areas
and may partly reflect the large number of smallrange species (Reid, 1992). The steep single-island
endemic curves have been interpreted as an evolutionary dynamics metric of speciation (Triantis et al.,
2008; Whittaker, Triantis & Ladle, 2008), but our
study does not have the detail needed for such conclusions. On the whole, the results of the EARs indicate that area is a good predictor of endemism in the
floristic regions, especially in the island regions, but
is not adequate alone, and other variables such as
history and isolation are expected to play a significant role (Yeakley & Weishampel, 2000; Whittaker,
Willis & Field, 2001). The higher concentration of
endemics in the island regions is a trait also
observed at the scale of the Med-Checklist territories
(Greuter, 1995).
The analysis of the distribution of the endemics by
similarity coefficients and hierarchical classification
can be a tool for the study of palaeogeography and
biogeography in Greece. The pattern of affinities of
WAe to the other regions is similar to the phytogeographical connections of Evvoia that arose by comparisons of the total floras (Trigas, Iatrou & Panitsa, 2008)
and the similarly of Pe to StE was also found for
mountain taxa (Strid, 1993). The KK–Pe affinity, commented upon by Greuter (1975), may reflect the connection of West Kriti with Peloponnisos until the late
Miocene (Dermitzakis & Papanikolaou, 1981), possible
Pliocene connections of West Kriti with the island of
Antikythira (Vardinoyannis, 1994) and possible Kriti
to Kythira colonization later than 4 Mya (Parmakelis
et al., 2006). The Kik–EAe affinity and biregional
endemics agree with: the connection of the EAe islands
of Ikaria and Chios to the Cyclades with which they
most probably formed a continuous landmass until the
lower Pliocene (map in Simaiakis & Mylonas, 2006);
the possible Evvoia (WAe)–Attiki (StE)–Cyclades (Kik)
connection to Asia Minor through the islands of Chios
and Samos during the Tortonian and Messinian (Dermitzakis & Papanikolaou, 1981); and the Kik–EAe
bridge during the Pleistocene (map in Poulakakis
et al., 2005). The NAe–NE affinity and biregional
endemics agree with the connection of Thasos and
Samothraki to the mainland until the Middle Pleistocene and their early separation from the other
Aegean islands (Dermitzakis & Papanikolaou, 1981;
Durand et al., 1999). A coupling of the detailed distribution with phylogenetic relationships of the endemics
would prove enlightening.
145
On the whole, although the Lance–Williams coefficient resulted in slightly higher similarity measures,
the Preston index performed better in terms of interpretation. The Lance–Williams coefficient is more
sensitive to the large disparity in the total number of
endemics in each region and the Preston index was
also considered preferable to the Jaccard index in the
numerical analysis of the floras of Aegean Islands and
the adjacent Asia regions (Carlström, 1987).
The altitudinal distribution of the endemics reveals
that they occur at all altitudes, but there is a concentration in the thermo-Mediterranean zone below
600 m, particularly in the island regions. There is,
however, a clear positive relationship of the number of
species with altitude, such as found in endemics of the
southern Iberian peninsula (Giménez et al., 2004),
Bolivia (Kessler, 2000) and South African fynbos
(McDonald & Cowling, 1995), in the continental floristic regions, where mountain isolation has promoted
endemism (Strid & Papanikolaou, 1985). A significant
number of species transgresses the altitudinal zones,
and hence bioclimates, apparently being adapted to a
wider range of environments. A similar trend has been
found in Corsica, in contrast to south-east France, and
has been considered as a pattern of widening ecological
niche in island environments (Médail & Verlaque,
1997). In KK, where both island and mountain isolation have played a role in plant endemism (Legakis &
Kypriotakis, 1994, also see Introduction), 43% of the
local endemics occur in the zones from below 600 m to
above 1000 m, but there is also a significant proportion
(20%) restricted to the zones above 1000 m.
Life form, the fundamental characteristic of a plant
(Barboni et al., 2004), is an attribute related to bioclimate and altitudinal distribution (Raunkiaer, 1934;
Gentry, 1988; Danin & Orshan, 1990; Verlaque et al.,
2001). The life form spectra of the Greek endemics,
although following the trends of the total Greek flora,
maintain at all altitudinal zones and in all floristic
regions the high percentages of hemicryptophytes
typical of mountainous regions. These spectra,
although reminiscent of the theory that Aegean
endemics are mainly mountain endemics with distribution areas older than the current land/sea distribution (Rechinger, 1965), are related to but not
adequately explained by the bioclimate or altitudinal
zone of the endemic plants. High proportions of hemicryptophytes among the Greek endemics have been
observed even in the floras of small islets (Vallianatou, 2005). In New Zealand, the endemic flora life
form spectra also resemble those of the alpine flora
(McGlone, Duncan & Heenan, 2001) and the prominence of woody endemics (including shrubs and subshrubs) was related to dispersal ability and habitat
and hence to the trend towards reduced dispersability
and the shift from herbaceous annuals to woody
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146
K. GEORGHIOU and P. DELIPETROU
perennials found in isolated island populations (Carlquist, 1974, 2009, Böhle, Hilger & Martin, 1996;
Cody & McOverton, 1996). This trend might apply to
island or mountain top endemics.
The difference in the life form spectra between the
endemic and the total flora is partly related to the
differences in family composition, although again
the endemic flora follows the total flora trend. For
example, Fabaceae have a high proportion of annuals
in both in the total (c. 40%, see literature in Table 6)
and the endemic flora (27%), but they are represented
by many fewer species in the endemic flora than
Asteraceae, which in contrast have a high proportion of
hemicryptophytes in both the total (see literature in
Table 6) and the endemic flora (80%). The flora of
Cataluña, which has a high proportion of endemic
members of Fabaceae, also has an increased proportion
of chamaephytic members of Fabaceae compared with
other Mediterranean area floras (Blondel & Aronson,
1995).
In Mediterranean-type climate regions, high proportions of herbaceous perennials and soft shrubs characterize the endemic floras of Sardinia (Bacchetta, Iiriti
& Pontecorvo, 2004), south-eastern Spain (Melendo
et al., 2003), California and Italy (Beard et al., 2000),
whereas woody shrubs and herbaceous perennials
characterize the endemic floras south-west Australia
and the Cape region of South Africa (Beard et al., 2000;
Ojeda et al., 2001). These life form spectra are at least
partly interpreted as a result of habitat specialization
to infertile and rocky substrata. Life form is expected
to be an attribute related to the habitat of the endemics
and is indicative of their ecological strategy (Grime,
1979; Grime, Hodgson & Hunt, 1988) and of the degree
of disturbance (Leishman & Westoby, 1992; McIntyre,
Lavorel & Tremont, 1995; Médail et al., 1998).
Endemic species are known to be stress tolerant,
occupying more infertile, steeper and rockier habitats
with reduced competition than more common plants
(Médail & Verlaque, 1997; Desmet & Cowling, 1999;
Ojeda et al., 2001; Lavergne et al., 2004) and predominance of stress-tolerance strategies was found in the
endemics of Evvoia (Trigas et al., 2008). Regarding the
habitat of the Greek endemics, a preliminary investigation (data not shown) and the literature (Rechinger
& Rechinger-Moser, 1951; Snogerup, 1985; Strid &
Papanikolaou, 1985; Strid & Tan, 1997; Trigas et al.,
2008) indicate that the majority of the endemics are
inhabitants of rocks, cliffs and screes (70–80% hemicryptophytes and chamaephytes) or, less often, of dry,
poor soils with phrygana (40% geophytes and 40%
hemicryptophytes and chamaephytes), garrigue or
grassland. These habitats are most often characterized
by low productivity and intense drought stress combined with low- to high-intensity unpredictable yet
recurrent or even continuous, anthropogenic (e.g.
grazing) or natural (e.g. wind) disturbance (sensu
Noble & Slatyer, 1980). This is the case in small islets,
cliffs and rocks, grazed phrygana and grassland, fireprone habitats affected by wind and salt spray. In fact,
habitats such as coastal rocks may promote speciation
processes (Crespo, De la Torre & Costa, 2003). Perennation by roots or stress-tolerant leaves and vegetative
reproduction are characters associated with a stresstolerant strategy (Grime et al., 1988). The taproots and
ligneous underground or slightly above ground parts of
hemicryptophytes and chamaephytes may effectively
withstand and help in the recovery from wind blowdowns. Indeed, the cliff floras are characterized by
slow-growing, desiccation tolerant or avoidant, perennial plants (Larson, Matthes & Kelly, 2000). In contrast, medium grazing, frequent in phrygana habitats,
favours geophytes and hemicryptophytes (McIntyre
et al., 1995; Noy-Meir & Oron, 2001).
Do the predominant life forms of the endemics
represent adaptations to the present ecological conditions or are they related to phylogenetic restrictions,
evolved in plants before the onset of the Mediterranean climate? The Mediterranean islands have functioned as museums for elements of former vegetation
that have been replaced elsewhere, but in the Aegean
islands there are many cases of post-Pleistocene
neoendemics (Snogerup, 1967a, b, Strid, 1970;
Bothmer, 1974; Thompson, 2005), and neoendemics
are also prominent in the continental flora (seed
introduction). The life form pattern of the endemics
cannot be explained adequately by altitudinal distribution or family composition, but it seems indicative
of their habitat and adaptive strategy. It could also be
surmised that it is related to speciation processes.
Chasmophytism is supposed to be a secondary development for certain elements that found refuge in the
Aegean cliffs (Snogerup, 1985) and lignification can
develop by island isolation (review in Whittaker &
Fernández-Palacios, 2007). Moreover, longevity is a
trait connected to a local persistence syndrome considered as a key feature for the ecology and long-term
viability of rare endemics (García & Regino, 2003;
Lavergne et al., 2004).
The overview of the conservation status of the
Greek endemics indicates that more than 40% of the
taxa are threatened or near threatened but only 0.5%
may be extinct. In many cases, the criteria for the
threat are based on the small area of occurrence or
extent of occupancy which is an intrinsic characteristic rather than a result of distribution area reduction caused by anthropogenic threats. A similar
situation occurs in Cyprus, where 32% of the endemics are threatened and none extinct and the restricted
distribution is the threat criterion for 60% of the
threatened endemics (Tsintides et al., 2007). Rarity
can mean vulnerability, but when, as in the case of
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PATTERNS OF PLANT ENDEMISM IN GREECE
Greece, either relic species or newly evolving neoendemics are involved, rarity can also be related to
endemism (Ricklefs, 1995). This is characteristic of
the floras of the Mediterranean basin, where species
diversity is positively correlated to endemism and
rarity and, despite the large number of rare species,
the confirmed cases of extinction amount to a mere
0.11% of the endemic Mediterranean element
(Greuter, 1995). In Greece, many rare endemics also
inhabit inaccessible habitats and are minimally
affected by anthropogenic threats. However, grazing,
a human activity widespread in Greece and most
often included in the main threats of rare species,
may have actually pushed certain species to inaccesible cliffs. Nevertheless, some endemics seem to
tolerate grazing (Bergmeier, 1998). Moreover, controlled grazing by maintaining a diverse landscape
may increase plant diversity and favour certain
plants linked to disturbance (Verdú, Crespo &
Galante, 2000).
However hopeful the above facts may seem, the
ever-increasing land-use changes in coastal areas
pose a significant threat, especially to island endemics and to coastal and wetland species. Sand dune
inhabitants with limited distributions such as Silene
ammophila Boiss. & Heldr. (endemic to KK) are
under immediate threat. Finally, global climate
change is an impending threat, especially for the
mountain endemics with restricted altitudinal
distribution.
ACKNOWLEDGEMENTS
The database Chloris was supported by the General
Secretariat of Research and Technology, Ministry of
Development, Greece (Research projects: Conservation and protection of endemic, rare and threatened
plants of Greece, 1993–1995; Database of endemic,
sub-endemic, rare, threatened and protected plants of
the Greek flora – CHLORIS, 1998–2000). We wish to
thank Professor Arne Strid for his useful comments
on the manuscript. We also thank Dr Ioannis Bazos
for his valuable advice and contribution in data collection. We are indebted to two anonymous reviewers
for useful comments on the manuscript.
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154 K. GEORGHIOU and P. DELIPETROU
APPENDIX
LIST OF THE ENDEMIC PLANTS OF GREECE
CONTENTS
Explanation of symbols (page 154)
The floristic regions of Greece (page 155)
Checklist of the endemic plants (pages 156)
Alphabetic synonym list (page 369)
Floras and checklists (page 395)
Literature (page 396)
Taxa currently considered synonyms or non-existent or mistaken names cited as greek endemics (page 420)
EXPLANATION OF SYMBOLS
! (In front of a plant name): doubtfully endemic
? (In front of a plant name): doubtful taxon
A (In front of a plant name): apomictic taxon
U (In front of a plant name): possibly apomictic taxon
S (In front of a plant name): sexual taxon (only for Limonium spp. and Taraxacum spp.)
+ (In the table of the floristic regions): present
? (In the table of the floristic regions): doubtfully present
* (In the table of the floristic regions): imperfectly known distribution
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PATTERNS OF PLANT ENDEMISM IN GREECE 155
THE FLORISTIC REGIONS OF GREECE
Pe
Peloponnese
StE
Sterea Ellada
WAe
West Aegean islands
IoI
Ionian islands
SPi
South Pindos
NPi
North Pindos
EC
East Central Greece
NC
North Central Greece
NE
North-East Greece
NAe
North Aegean islands
Kik
Cyclades
KK
Crete, Karpathos
EAe
East Aegean islands
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
?
3
Acanthus greuterianus Snogerup, B.Snogerup & Strid
Acanthaceae
L/1926
Acer hyrcanum Fisch. & C.A.Mey. subsp. reginae amaliae (Orph. ex
Boiss.) E.Murray
Sapindaceae
Acer opalus Mill. subsp. hyrcanum (Fischer & Meyer) var. reginaeamaliae (Orph. ex Boiss.) Aldén
Acer amaliae Boiss. (nom. illeg.)
Acer hyrcanum Fischer & Meyer subsp. reginae-amaliae (Boiss.)
A.E.Murray
Acer reginae-amaliae Orph. ex Boiss.
FE 2:239 FGC 1:289 FGS 1:24 MCL 1:40 MFG 1:582 PFPB 1:604
L/202 L/391 L/395 L/512 L/607 L/843 L/1871 L/1963 L/2013
4
5
Acantholimon androsaceum (Jaub. & Spach) Boiss.
Plumbaginaceae
Acantholimon echinus (L.) Boiss. subsp. creticum (Boiss.) Papan. &
Kokkini
Acantholimon echinus L. var. creticum Boiss.
Acantholimon ulicinum sensu auct. cret.
Statice androsacea Jaub. & Spach
EFK :224 FAe :426 FCA :125 FCSI :70 FE 3:30 FT 7:498
MCL 4:307 MFG 1:748
L/393 L/518 L/1111
Acantholimon aegaeum F.K.Mey.
Plumbaginaceae
Acantholimon androsaceum sensu auct. Samos
Acantholimon echinus sensu auct. Samos
FAe :426 FT 10:212 FT 7:498 MCL 4:307
L/393 L/796 L/975 L/1957
?
2
Taxon
Abies cephalonica Loudon
Pinaceae
Abies apollinis Link
Abies cephalonica Loudon f. arcadiaca Henkel & W.Hochst
Abies cephalonica Loudon f. parnassica Henkel & W.Hochst
Abies cephalonica Loudon var. apollinis (Link) Beissner
Abies cephalonica Loudon var. graeca (Fraas) Liu
Abies panachaica Heldr.
Abies reginae-amaliae Heldr.
AFE 2:12 FE 1:38 FGC 3:450 FGS 1:115 FGS 2:91 FHe 1:2 MCL 1:30 MFG 1:39
L/159 L/512 L/1200 L/1467 L/1844 L/1846 L/1871
L/1963 L/1973 L/2013
Taxon
Qualifiers
1
Taxon
Number
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+
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+
StE
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+
Pe
+
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WAe
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SPi
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+
IoI
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EC
NPi
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NC
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
156 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Achillea occulta Constantin. & Kalpoutz.
Asteraceae
L/1892
Achillea pindicola Hausskn. subsp. integrifolia (Halácsy) Franzen
Asteraceae
Achillea clavennae L. var. integrifolia Halácsy
Achillea pauciloba (Heldr. ex Heimerl) Heimerl var. Halacsyana Heimerl
Achillea pauciloba (Heldr. ex Heimerl) Heimerl var. integrifolia
(Halácsy) Heimerl
MFG 2:437
Achillea pindicola Hausskn. subsp. pindicola
Asteraceae
Achillea pauciloba (Heldr. ex Heimerl) Heimerl
Achillea umbellata Sm. subsp. pauciloba Heldr. ex Heimerl
Achillea umbellata Sm. var. pauciloba (Heldr. ex Heimerl) Heimerl
MFG 2:436
L/121 L/2200
11
12
Achillea barbeyana Heldr. & Heimerl
Asteraceae
Ptarmica barbeyana (Heldr. & Heimerl) Nyman
Ptarmica vulgaris DC.
FE 4:161 FGC 2:43 MFG 2:437 RDB :4
L/330
9
10
Achillea ambrosiaca (Boiss. & Heldr.) Boiss.
Asteraceae
Achillea erba-rotta All. subsp. olympica (Heimerl) I.B.K.Richardson
Achillea moschata Wulfen subsp. olympica Heimerl
Achillea olympica (Heimerl) Halácsy
FE 4:161 FGC 2:43 FGC 2:44 MFG 2:438 RDB :2
L/953 L/1180
8
Achillea aegyptiaca L.
Asteraceae
Achillea tournefortii DC. (nom. illeg.)
FE 4:164 FGC 2:50 FSPS 2:114 MFG 2:449
L/111 L/121 L/145 L/169 L/377 L/861
7
!
Achillea absinthoides Halácsy
Asteraceae
FE 4:164 FGC 2:48 MFG 2:448
L/477
6
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*
PATTERNS OF PLANT ENDEMISM IN GREECE 157
18
!
Adonis cyllenea Boiss., Heldr. & Orph.
Ranunculaceae
Chrysocyathus cylleneus (Boiss., Heldr. & Orph.) Chrtek & Slavikova
AFE 8:111 FE 1:268 FGC 1:6 FT 1:143 MCL 4:392 MFG 1:209 RDB :6
L/485 L/824 L/1120 L/1232 L/1706 L/1769 L/1963 L/2201
Adonis cretica (Hunth.) Runemark
Ranunculaceae
Adonis aestivalis L. subsp. cretica (Hunth.) C.Steinb.
Adonis dentata Delile subsp. cretica (Hunth.) Riedl
Adonis microcarpa DC. subsp. cretica (Huth.) Vierh.
EFK :98 FAe :195 FCA :130 FHe 2:37
L/268 L/453
Acinos alpinus (L.) Moench subsp. nomismophyllus (Rech.f.)
Leblebici
Lamiaceae
Calamintha alpina (L.) Lam. subsp. nomismophylla Rech.f.
FE 3:165 MCL 3:323 MFG 2:130
L/1941 L/1972 L/1973
16
17
Achnatherum fallacinum H.Scholz & Raus
Poaceae
FCSII :
L/1922
15
?
Achillea umbellata Sm.
Asteraceae
Achillea umbellata Sm. subsp. monocephala (Heldr. ex Heimerl)
Tzanoud. & Iatroú
Achillea umbellata Sm. subsp. vulgaris Heimerl nom. illeg.
Achillea umbellata Sm. subsp. vulgaris Heimerl var. monocephala
Heldr. ex Heimerl
Achillea umbellata Sm. subsp. vulgaris Heimerl var. polycephala
Heimerl
Ptarmica umbellata (Sm.) DC.
FE 4:161 FGC 2:42 FGS 2:49 MFG 2:435
L/160 L/512 L/530 L/607 L/953 L/1193 L/1844 L/1871 L/1963 L/2013
14
Taxon
Achillea taygetea Boiss. & Heldr.
Asteraceae
Achillea aegyptiaca L. subsp. taygetea (Boiss. & Heldr.) Hayek
Achillea aegyptiaca L. var. taygetea (Boiss. & Heldr.) Halácsy
FE 4:164 FGC 2:50 MFG 2:449
L/512 L/607 L/1963
Taxon
Qualifiers
13
Taxon
Number
.
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+
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+
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+
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WAe
StE
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Pe
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SPi
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IoI
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
158 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Aegonychon goulandriorum (Rech.f.) Holub
Boraginaceae
Lithospermum goulandriorum Rech.f.
MCL 1:94 MFG 2:23
L/477 L/1149 L/1929
Aegonychon goulandriorum (Rech.f.) Holub subsp. goulandriorum
Boraginaceae
Lithospermum goulandriorum Rech.f. subsp. goulandriorum
MCL 1:94 MFG 2:23
L/477 L/1149 L/1929
Aegonychon goulandriorum (Rech.f.) Holub subsp. thessalicum
(Aldén) Valdés
Boraginaceae
Lithospermum goulandriorum Rech.f. subsp. thessalicum Aldén
MCL 1:94 MFG 2:23
L/477 L/1929
Aethionema carlsbergii Strid & Papan.
Brassicaceae
Aethionema orbiculatum auct. non (Boiss.) Hayek
AFE 11:164 FE 1:389 FHe 2:262 MCL 3:35 MFG 1:327 RDB :8
L/389 L/404a L/607 L/1963
Aethionema orbiculatum (Boiss.) Hayek
Brassicaceae
Crenularia orbiculata Boiss.
Eunomia orbiculata Griseb.
AFE 11:162 FE 1:388 FHe 2:262 MCL 3:36 MFG 1:327
L/607 L/1972 L/1973
21
22
23
24
25
Aegilops comosa Sm. subsp. heldreichii (Boiss.) Eig
Poaceae
Aegilops comosa Sm. var. subventricosa Boiss.
Triticum comosum (Sm.) K.Richt. subsp. heldreichii (Boiss.) Greuter
Triticum heldreichii (Boiss.) K.Richt.
EFK :390 FCA :176 FCSI :94 FE 5:201 FT 9:239
B/3060s L/430 L/607 L/913 L/1052 L/1111 L/1957
20
?
Aegilops biuncialis Vis. subsp. archipelagica (Eig) Raus
Poaceae
Aegilops biuncialis Vis. var. archipelagica Eig
FCSII :
L/551 L/1917 L/1918
19
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 159
31
Alchemilla aroanica (Buser) Rothm.
Rosaceae
Alchemilla acutiloba Steven subsp. aroanica Buser
FE 2:62 MFG 1:429 RDB :12
L/607 L/1963
Ajuga orientalis L. subsp. aenesia (Heldr.) Phitos & Damboldt
Lamiaceae
FGC 2:466
L/1123
30
!
Agrostemma githago L. subsp. thessalum (Bornm.) Greuter
Caryophyllaceae
Agrostemma gracile auct.
Agrostemma gracile L. var. thessalum Bornm.
Agrostemma thessalum (Formánek) Hayek
Githago thessala Formánek
FHe 1:238
L/309b L/350
29
Aethionema samium T.R.Dudley & Christod.
Brassicaceae
FT 11:37
Aethionema retsina Phitos & Snogerup
Brassicaceae
AFE 11:164 FE 1:389 FHe 2:262 MCL 3:36 MFG 1:328 RDB :10
L/918
Taxon
Aethionema saxatile (L.) R.Br. subsp. creticum (Boiss. & Heldr.)
I.A.Andersson
Brassicaceae
Aethionema ovalifolium (DC.) Boiss.
Aethionema polygaloides DC.
Aethionema creticum Boiss. & Heldr.
Aethionema graecum Boiss. & Spruner var. creticum (Boiss. & Heldr.)
Halácsy
Aethionema saxatile (L.) R.Br. var. creticum (Boiss. & Heldr.) Halácsy
AFE 11:167 EFK :122 FAe :241 FCA :78 FE 1:389 FGS 1:12 FHe 2:264 FT 1:320 FT 10:37 MCL 3:36 MFG 1:328 MFG 1:331
L/196 L/197 L/278 L/321 L/321 L/539 L/540 L/607 L/776 L/856 L/869 L/930 L/1111 L/1957 L/1963
?
Taxon
Qualifiers
28
27
26
Taxon
Number
.
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+
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+
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WAe
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StE
Pe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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+
+
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EAe
Distribution
Qualifier
160 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Alkanna calliensis Heldr. ex Boiss.
Boraginaceae
FE 3:96 FGC 2:342 FGS 1:76 MCL 1:66
L/2013
Alkanna graeca Boiss. & Spruner
Boraginaceae
FE 3:96 FGC 2:344 FGS 1:76 MCL 1:67 MFG 2:40
L/121 L/447 L/512 L/530 L/607 L/1244 L/1960 L/1963 L/1972 L/1973 L/2013 L/2200
Alkanna graeca Boiss. & Spruner subsp. baeotica (DC.) Nyman
Boraginaceae
Alkanna baeotica DC.
FE 3:96 FGC 2:344 MCL 1:67 MFG 2:40
B/1316 B/2423 L/489 L/530 L/607 L/1871 L/1960 L/1963
Alkanna graeca Boiss. & Spruner subsp. graeca
Boraginaceae
FE 3:96 MCL 1:67 MFG 2:40
L/447 L/607 L/902 L/1871 L/1963 L/2013
Alkanna methanaea Hausskn.
Boraginaceae
FE 3:96 FGC 2:343 FGS 1:76 FGS 2:63 MCL 1:67
L/395 L/607 L/1844 L/1963
Alkanna pelia (Halácsy) Rech.f.
Boraginaceae
Alkanna stribrnyi var. pelia Halácsy
FE 3:96 FGC 2:345 MCL 1:68
Alkanna sartoriana Boiss. & Heldr.
Boraginaceae
FE 3:97 FGC 2:345 MCL 1:68 RDB :14
L/606 L/607 L/1147 L/1963
Alkanna sieberi DC.
Boraginaceae
EFK :248 FAe :451 FCA :38 FE 3:97 FGC 2:346 MCL 1:68
L/518 L/554 L/707 L/1111 L/2093
32
33
34
35
36
37
38
39
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+
+
+
+
+
+
+
+
+
+
+
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 161
Allium aegilicum Tzanoud.
Amaryllidaceae
L/1111 L/1691 L/1917
Allium aeginiense Brullo, Giusso & Terrasi
Amaryllidaceae
L/2222
Allium apolloniensis B.Biel, Kit Tan & Tzanoud.
Amaryllidaceae
L/1921
Allium archeotrichon Brullo, Pavone & Salmeri
Amaryllidaceae
FT 11:227
L/1111 L/1482
Allium bourgeaui Rech.f. subsp. creticum Bothmer
Amaryllidaceae
Allium bourgeaui subsp. bourgeaui sensu Bothmer 1974 p.p.
EFK :362 FCA :180 FCSI :105 FE 5:64
L/430 L/682 L/1111
Allium brachyspathum Brullo, Pavone & Salmeri
Amaryllidaceae
FCSII :
L/1111 L/2093
Allium brulloi Salmeri
Amaryllidaceae
L/1471
Allium brussalisii Tzanoud. & Kypr.
Amaryllidaceae
L/2221
Allium calamarophilon Phitos & Tzanoud.
Amaryllidaceae
RDB :16
L/471 L/1136 L/1871
41
42
43
44
45
46
47
48
49
Taxon
Allium achaium Boiss. & Orph.
Amaryllidaceae
FE 5:59 FGC 3:251 MFG 2:709
L/121 L/682 L/1467 L/1871 L/1963 L/2013 L/2200
Taxon
Qualifiers
40
Taxon
Number
.
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+
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+
StE
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+
+
Pe
+
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WAe
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IoI
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+
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+
SPi
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+
NPi
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EC
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NC
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NE
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NAe
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+
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+
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Kik
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+
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KK
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+
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EAe
Distribution
Qualifier
162 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Allium candargyi Karavok. & Tzanoud.
Amaryllidaceae
FT 11:228
L/1080 L/1192
Allium chalkii Tzanoud. & Kollmann
Amaryllidaceae
FT 11:225
L/478 L/1192
Allium circinnatum Sieber subsp. circinnatum
Amaryllidaceae
Allium circinnatum Sieber, Rechinger 1943; Tutin & al. 1980; Stearn
1978
EFK :359 FCA :180 FCSI :105 FE 5:57
L/430 L/471 L/682 L/726 L/727 L/1111
Allium circinnatum Sieber subsp. peloponnesiacum Tzanoud.
Amaryllidaceae
FCA :180
L/471 L/607 L/726 L/727 L/1111 L/1917 L/1917 L/1963
Allium dirphianum Brullo, Guglielmo, Pavone, Salmeri & Terrasi
Amaryllidaceae
L/1879
53
54
55
56
57
Allium callimischon Link subsp. haemostictum Stearn
Amaryllidaceae
Allium callimischon sensu Rechinger 1949
Allium cruentatum Sieber nom. nud.
EFK :361 FCA :180 FE 5:59 FT 8:130
L/430 L/682 L/1055 L/1111 L/1194
!
52
Allium callimischon Link
Amaryllidaceae
EFK :361 FCA :180 FE 5:59 FT 8:130
L/471 L/472 L/646 L/682 L/1055 L/1111 L/1123 L/1192 L/1194 L/1830 L/1846 L/1963
Allium callimischon Link subsp. callimischon
Amaryllidaceae
Allium boryanum Kunth
FE 5:59
L/471 L/607 L/682 L/1055 L/1111 L/1123 L/1194 L/1830 L/1963
!
51
50
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 163
Allium goulimyi Tzanoud.
Amaryllidaceae
L/471 L/472 L/726 L/1192 L/1211 L/2201
Allium guttatum Steven subsp. dilatatum Zahar.
Amaryllidaceae
Allium dilatatum Zahar.
EFK :361 FCA :181 FCSI :106 FE 5:67
L/430 L/471 L/682 L/728 L/729 L/1221
Allium heldreichii Boiss.
Amaryllidaceae
FE 5:68 FGC 3:248 MFG 2:713
L/471 L/477 L/682 L/1055 L/1246 L/2201
64
65
66
Allium frigidum Boiss. & Heldr.
Amaryllidaceae
FE 5:59 FGC 3:251 FGS 2:83 FT 8:135 MFG 2:706
L/471 L/472 L/682 L/1192 L/1963
62
Allium gomphrenoides Boiss. & Heldr.
Amaryllidaceae
FE 5:68 FGC 3:248
L/119 L/471 L/607 L/682 L/729 L/765 L/1052 L/1111 L/1192 L/1878 L/1917 L/1963
Allium favosum Zahar.
Amaryllidaceae
Allium parnassicum auct. non Halácsy
Allium stamineum Orphanides, non Boiss.
FE 5:61
L/471 L/566 L/682 L/1192 L/1226
61
63
Allium euboicum Rech.f.
Amaryllidaceae
Allium paniculatum L. subsp. euboicum (Rechf.) Stearn
FE 5:60
L/323b L/530 L/608 L/682 L/1192 L/1871
60
!
Allium erythraeum Griseb.
Amaryllidaceae
Allium obtusiflorum DC. (including taxon)
FE 5:60 FGC 3:251
L/315 L/682 L/1192
59
Taxon
Allium dodecanesi Karavok. & Tzanoud.
Amaryllidaceae
FT 11:229
L/913 L/1080 L/1192
Taxon
Qualifiers
58
Taxon
Number
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StE
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WAe
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IoI
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SPi
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NPi
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EC
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NC
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NAe
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KK
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EAe
Distribution
Qualifier
164 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Allium hymettium Boiss. & Heldr.
Amaryllidaceae
Allium stamineum Boiss. var. hymettium Boiss.
FE 5:63 FGC 3:254
L/471 L/530 L/607 L/682 L/1055 L/1192 L/1871 L/1871 L/2013
Allium integerrimum Zahar.
Amaryllidaceae
FE 5:68
L/682 L/1053 L/1192 L/1221
Allium ionicum Brullo & Tzanoud.
Amaryllidaceae
L/1192 L/1692 L/1800 L/1843a
Allium karistanum Brullo, Pavone & Salmeri
Amaryllidaceae
L/933 L/1192 L/1683 L/1871
Allium lagarophyllum Brullo, Pavone & Tzanoud.
Amaryllidaceae
L/315 L/1192
Allium luteolum Halácsy
Amaryllidaceae
FE 5:62 FGC 3:256 FGS 2:84
L/145 L/169 L/270 L/270 L/471 L/682 L/776 L/869 L/1192
Allium macedonicum Zahar.
Amaryllidaceae
FE 5:61 MFG 2:708
L/682 L/1107 L/1108 L/1192 L/1226
Allium maniaticum Brullo & Tzanoud.
Amaryllidaceae
Allium obtusiflorum DC. (including taxon)
L/262 L/315 L/682 L/1192 L/1963
Allium parnassicum (Boiss.) Halácsy
Amaryllidaceae
Allium achaium Boiss. & Orph. var. parnassicum Boiss.
FE 5:62 FGC 3:255 FGS 1:105 MFG 2:709
L/471 L/477 L/682 L/1226 L/1267a L/2194
67
68
69
70
71
72
73
74
75
.
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+
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+
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 165
Allium pilosum Sm.
Amaryllidaceae
FAe :719 FE 5:62 FGC 3:257 FT 8:148
L/270 L/471 L/682 L/796 L/1079 L/1192 L/1957
Allium platakisii Tzanoud. & Kypr.
Amaryllidaceae
L/317 L/1078 L/1111 L/1192 L/1583 L/1692
Allium rausii Brullo, Guglielmo, Pavone, Salmeri & Terrasi
Amaryllidaceae
L/1879
Allium rhodiacum Brullo, Pavone & Salmeri
Amaryllidaceae
FT 11:225
L/289 L/1192
Allium ritsii Iatroú & Tzanoud.
Amaryllidaceae
L/243 L/1192 L/1963
Allium runemarkii Trigas & Tzanoud.
Amaryllidaceae
L/1683 L/1871
Allium sphaerocephalon L. subsp. aegaeum (Heldr. & Halácsy)
Karavok. & Tzanoud.
Amaryllidaceae
FAe :716 FGC 3:716 FT 11:229
L/169 L/1079
Allium sphaerocephalon L. subsp. trachypus (Boiss. & Spruner)
Stearn
Amaryllidaceae
Allium sphaerocephalon L. var. aegaeum Heldr. & Halácsy
Allium arvense Guss. var. trachypus Boiss.
Allium sphaerocephalon L. subsp. sphaerocephalon var. trachypus Boiss.
Allium trachypus Boiss. & Spruner
FAe :716 FE 5:66 FGC 3:246
L/682 L/1111 L/1973
77
78
79
80
81
82
83
84
Taxon
Allium phitosianum Brullo, Guglielmo, Pavone, Salmeri & Terrasi
Amaryllidaceae
L/1879
Taxon
Qualifiers
76
Taxon
Number
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WAe
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StE
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Pe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
166 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Allium tardans Greuter & Zahar.
Amaryllidaceae
Allium paniculatum sensu auct. karp., non L. 1759
Allium staticiforme sensu auct. cret., non Sm?
EFK :360 FCA :181 FCSI :107 FE 5:62
L/201 L/268 L/270 L/430 L/471 L/473 L/551 L/646
L/682 L/932 L/1111 L/1226 L/1692 L/1766
Allium thessalicum Brullo, Pavone, Salmeri & Tzanoud.
Amaryllidaceae
Allium erythraeum sensu auct. non Griseb. p.p.
Allium obtusiflorum sensu auct. non DC. p.p.
L/315 L/682 L/1192 L/1871
Alyssum baldaccii Vierh. ex E.I.Nyár.
Brassicaceae
Alyssum argenteum sensu Boiss. Fl. Orient.
Alyssum fallacinum sensu auct. cret. non Hausskn.
AFE 11:51 EFK :117 FCA :78 FCSI :38 FCSII : FE 1:367 FHe 2:219 MCL 3:42 RDB :20
L/518 L/524 L/831
Alyssum degenianum Nyár.
Brassicaceae
Alyssum murale Waldst. & Kit. subsp. stojanoffi (Nyár.) Dudley in
J. Arnold Arbor. 1964 non sensu orig.
AFE 11:49 FE 1:367 FHe 2:217 MCL 3:45 MFG 1:297
L/178 L/895 L/1252
Alyssum densistellatum T.R.Dudley
Brassicaceae
Alyssum praecox auct. graec. non Boiss.
AFE 11:48 FAe :224 FE 1:366 FHe 2:212 MCL 3:41
L/323a L/831 L/1699 L/1871
Alyssum euboeum Halácsy
Brassicaceae
AFE 11:54 FAe :226 FE 1:367 FHe 2:219 MCL 3:42
L/99 L/323a L/608 L/831 L/1699 L/1871
85
86
87
88
89
90
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 167
Alyssum lesbiacum (Candargy) Rech.f.
Brassicaceae
Odontarrhena lesbiaca P.Candargy
FAe :226 FHe 2:220 FT 1:409 MCL 3:43
B/2447 L/111 L/986 L/1699 L/1880
97
Alyssum idaeum Boiss. & Heldr.
Brassicaceae
Alyssum curetum Gand.
Alyssum idaeum var. curetum (Gand.) Hayek
AFE 11:47 EFK :118 FCA :78 FCSI :38 FE 1:366 FHe 2:216 MCL 3:43 MFG 1:292
L/831 L/856 L/1111 L/1699 L/2248
95
Alyssum lassiticum Halácsy
Brassicaceae
AFE 11:47 EFK :118 FCA :78 FCSI :38 FE 1:366 FHe 2:215 MCL 3:43 MFG 1:292
L/518 L/831 L/1111 L/1583 L/2255
Alyssum heldreichii Hausskn.
Brassicaceae
AFE 11:51 FE 1:367 FHe 2:218 MCL 3:43 MFG 1:299
L/159 L/178 L/178 L/1699 L/1729 L/1901 L/2201
94
96
Alyssum handelii Hayek
Brassicaceae
AFE 11:47 FE 1:366 FHe 2:215 MCL 3:42 MFG 1:293 RDB :24
L/832 L/1180 L/1699
93
!
Alyssum gustavssonii Hartvig
Brassicaceae
AFE 11:44 FE 1:365 FHe 2:212 MCL 3:42 MFG 1:293
L/1963 L/2013
92
Taxon
Alyssum fragillimum (Bald.) Rech.f.
Brassicaceae
Alyssum alspestre L. var. fragillimum Bald.
Alyssum nebrodense Tineo var. fragillimum (Bald.) Halácsy
AFE 11:59 EFK :117 FAe :226 FCA :78 FCSI :38 FE 1:369 FHe 2:221 MCL 3:42 MFG 1:303 RDB :22
B/3060f L/518 L/519 L/522 L/554 L/753 L/831 L/1111 L/1766
Taxon
Qualifiers
91
Taxon
Number
.
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StE
Pe
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WAe
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IoI
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SPi
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+
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NPi
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EC
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+
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NC
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NE
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NAe
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Kik
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+
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KK
+
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EAe
Distribution
Qualifier
168 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
102
!
101
?
?
!
100
?
?
!
99
98
Alyssum nebrodense Tineo subsp. tenuicaule Hartvig
Brassicaceae
AFE 11:59 FE 1:368 FHe 2:221 MCL 3:46 MFG 1:300
L/1963
Alyssum montanum L. subsp. repens (Baumg.) Schmal. var.
trichostachyum (Rupr.) Baumg.
Brassicaceae
Alyssum spathulaefolium Formánek
Alyssum montanum L. subsp. epiroticum (Baumg.) Hayek
Alyssum montanum L. subsp. montanum proles epiroticum Baumg.
Alyssum virescens Halácsy
Alyssum repens var. stenophyllum Halácsy
Alyssum repens Rupr.
Alyssum repens subsp. trichostachyum (Rupr.) Hayek
FHe 2:211 MFG 1:289
L/178 L/2200
Alyssum montanum L. subsp. repens (Baumg.) Schmal. var.
brachyphyllum Halácsy
Brassicaceae
Alyssum montanum L. subsp. montanum proles laxum Baumg.
Alyssum montanum L. subsp. montanum proles humile Baumg.
FHe 2:211
L/2200
Alyssum montanum L. subsp. montanum var. hymettium Boiss
Brassicaceae
Alyssum montanum L. var. graecum Halácsy p.p. minor
Alyssum montanum L. var. angustifolium Heldr.
Alyssum spruneri Jordan & Fourr.
FHe 2:210 MFG 1:288
L/828
Alyssum montanum L. subsp. montanum var. graecum Halácsy
Brassicaceae
Alyssum vourinonense T.R.Dudley & Rech.f.
Alyssum montanum L. subsp. montanum proles thessalum (Halácsy)
Baumg.
Alyssum thessalum Halácsy
Alyssum olympicum Halácsy
Alyssum montanum L. subsp. graecum (Halácsy) Hayek
FHe 2:209 MFG 1:287
L/1901
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PATTERNS OF PLANT ENDEMISM IN GREECE 169
Alyssum simulans Runemark
Brassicaceae
Alyssum foliosum sensu Halácsy Consp. Fl. Graec. ‘Pentelikon’ p.p.
FGC 1:98 FHe 2:207
L/1871
Alyssum sphacioticum Boiss. & Heldr.
Brassicaceae
Alyssum atlanticum sensu Sm. Fl. Graec. Prodr.
AFE 11:47 EFK :118 FAe :223 FCA :79 FCSI :39 FE
1:367 FHe 2:215 MCL 3:49 MFG 1:291 RDB :26
L/519 L/519 L/522 L/753 L/831 L/1111 L/1699
Alyssum taygeteum Heldr.
Brassicaceae
Alyssum doerfleri var. taygeteum (Heldr.) Bornm.
AFE 11:48 FE 1:366 FHe 2:214 MCL 3:41 MFG 1:295
L/512 L/607 L/831 L/1915 L/1963
Alyssum tenium Halácsy
Brassicaceae
AFE 11:50 FAe :226 FE 1:367 FGC 1:93 FHe 2:218 MCL 3:50 RDB :28
L/169 L/831 L/1699
Alyssum xifocarpum P.Candargy
Brassicaceae
Alyssum umbellatum Desv. sensu T.R.Dudley & al. in Fl. Turkey
FAe :224 FHe 2:205 MCL 3:50
L/1880
Amelanchier parviflora Boiss. subsp. chelmea (Halácsy) Ziel.
Rosaceae
Amelanchier chelmea (Halácsy) Browicz
Amelanchier cretica var. chelmea Halácsy
FT 4:168-172 MFG 1:437 RDB :30
L/391 L/395 L/443 L/767 L/967 L/1772 L/1871 L/1901 L/1963
104
105
106
107
108
109
Taxon
Alyssum samium Dudley & Christod.
Brassicaceae
Alyssum condensatum Boiss. & Hausskn. subsp. flexibile (Nyár) sensu
T.R.Dudley 1964
FHe 2:221
L/753 L/796 L/975
Taxon
Qualifiers
103
Taxon
Number
.
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+
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+
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+
+
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StE
Pe
+
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WAe
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SPi
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IoI
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NPi
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EC
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NC
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NE
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NAe
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Kik
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+
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KK
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+
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EAe
Distribution
Qualifier
170 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Anchusa leucantha Selvi & Bigazzi
Boraginaceae
L/2216
Anchusa samothracica Bigazzi & Selvi
Boraginaceae
L/1684 L/2216
Anchusa stylosa M.Bieb. subsp. spruneri (Boiss.) Selvi & Bigazzi
Boraginaceae
Anchusa spruneri Boiss.
Anchusa stylosa M.Bieb. var. major DC.
Anchusa stylosa M.Bieb. subsp. spruneri (Boiss.) Nyman
FAe :449 FE 3:108 FGC 2:328 FGS 1:75 MCL 1:71 PFPB 2:64
L/607 L/1252 L/1267b L/1963 L/2216
Anchusa undulata L. subsp. sartorii (Guşul.) Selvi & Bigazzi
Boraginaceae
Anchusa macrosyrinx Rech.f.
Anchusa sartorii Guşul.
Anchusa hybrida Ten. var. setigera Halácsy
Anchusa undulata Sm. var. maritima Rech.f.
FAe :448 FE 3:107, 108 FGC 2:325 MCL 1:71, 72 PFPB 2:60
L/111 L/169 L/451 L/561 L/776 L/869 L/1146 L/1288 L/2216
112
113
114
115
Ammi topalii Beauverd
Apiaceae
FAe :404 FE 2:353
L/99 L/323a L/608 L/707 L/1871
Anchusa cespitosa Lam.
Boraginaceae
EFK :250 FAe :447 FCA :38 FCSI :13 FCSII : FE 3:107
FT 6:402 MCL 1:70 MFG 2:45 RDB :32
B/3060f L/518 L/554 L/1048 L/1111 L/1766 L/2216
?
111
110
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+
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*
PATTERNS OF PLANT ENDEMISM IN GREECE 171
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Andrachne telephioides L. subsp. oreocretensis Aldén
Euphorbiaceae
EFK :180 FCA :88 MCL 3:205 MFG 1:565
L/253 L/899 L/1111
Androcymbium rechingeri Greuter
Colchicaceae
Androcymbium gramineum (Cav.) J.F.Macbr. var. puniceum (Maire)
Maire
EFK :350 FCA :182 FCSI :107 FE 5:21 RDB :34
L/255 L/1111 L/1247
Anemone hortensis L. subsp. heldreichii (Boiss.) Rech.f.
Ranunculaceae
Anemone stellata var. heldreichii Boiss.
AFE 8:85 EFK :97 FAe :183 FCA :130 FCSI :74 FE 1:264
FGC 1:6 FHe 2:28 MCL 4:394
L/273 L/518 L/554 L/707 L/1111
117
118
119
Taxon
Anchusella variegata (L.) Bigazzi, Nardi & Selvi
Boraginaceae
Borragine murale variegata con flore odorato, di Candia Zanoni
Borago cretica minima Contareni non descripta Bellucci
Borrago verrucosa minor flore variegato Muntig
Lycopsis aegyptiaca auct non L., Gand
Borago variegata cretica Wheler
Buglossum creticum culicolisb procumbentibus, flore variegato odorato,
echii foliis verrucosis Hermann
Buglossoides cretica Rvinus
Buglossum creticum verrucosum Strisser
Lycopsis foliis drepano-dentatis Lineé
Anchusa ovata auct non Lehm., Mazziari
Buglossum creticum minimum odoratum flore varion eleganti Jonqcuet
Anchusa perlata Lam. nom. illeg.
Anchusa variegata (L.) Lehm.
Anchusa variegata (L.) Lehm. subvar. cephalonica (Bornm. & Gusul.)
Gusul.
Anchusa variegata (L.) Lehm. var. cephalonica Bornm. & Gusul.
Lycopsis variegata L.
EFK :250 FAe :449 FCA :39 FCSI :14 FCSII : FE 3:109
FGC 2:329 FGS 1:76 FGS 2:63 FT 6:400 MCL 1:72
L/150 L/408 L/451 L/518 L/537 L/554 L/607 L/700
L/767 L/913 L/1107 L/1111 L/1112 L/1244 L/1871 L/1878 L/1913 L/1917 L/1963 L/2181
Taxon
Qualifiers
116
Taxon
Number
.
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+
StE
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+
Pe
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+
WAe
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+
IoI
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NPi
SPi
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+
EC
.
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?
NC
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NE
.
.
.
+
NAe
.
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+
Kik
+
+
+
+
KK
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.
+
EAe
Distribution
Qualifier
172 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Anthemis abrotanifolia (Willd.) Guss.
Asteraceae
Anthemis boissieri Stoj. & Acht.
Anthemis cretica auct., non L.
Cotula abrotanifolia Willd.
EFK :314 FAe :625 FCA :57 FCSI :25 FE 4:150 MFG 2:427
L/253 L/522 L/554 L/1014
Anthemis ammanthus Greuter
Asteraceae
EFK :313 FAe :632 FCA :57 FCSI :25 FE 4:158
L/255 L/338 L/495 L/518 L/554 L/773 L/932 L/1049 L/1111 L/1710 L/2093
Anthemis ammanthus Greuter subsp. ammanthus
Asteraceae
Ammanthus ageratifolius auct. cret.
Ammanthus intermedius Gand. (non Anthemis intermedia Guss.)
Ammanthus maritimus Boiss. & Heldr. (non Anthemis maritima L.
Chrysanthemum maritimum)
EFK :313 FAe :632 FCA :57 FCSI :25 FE 4:158
L/201 L/255 L/268 L/273 L/338 L/495 L/518 L/537 L/554 L/932 L/1049 L/1111
Anthemis ammanthus Greuter subsp. paleacea Greuter
Asteraceae
Ammanthus maritimus auct. p.p. [non Boiss. & Heldr.]
Ammanthus tomentellus p.p.
EFK :313 FCA :57 FE 4:158
L/255 L/273 L/338 L/495 L/518 L/554 L/1049 L/1111
Anthemis brachmannii Boiss. & Heldr.
Asteraceae
FE 4:158 FGC 2:55
L/607 L/1963
Anthemis cretica L. subsp. panachaica (Halácsy) Oberprieler &
Greuter
Asteraceae
Anthemis panachaica Halácsy
Anthemis montana L. var. linnaeana Hal.
FE 4:152 FGC 2:57 MCL 2:26 MFG 2:422
L/1932 L/2237
120
121
122
123
124
125
.
.
.
.
.
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
+
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+
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+
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.
PATTERNS OF PLANT ENDEMISM IN GREECE 173
Anthemis cretica L. subsp. spruneri (Boiss. & Heldr.) Govaerts
Asteraceae
Anthemis spruneri Boiss. & Heldr.
Anthemis montana L. var. incana (Boiss.) Hayek
Anthemis montana L. var. spruneri (Boiss. & Heldr.) Halácsy
Anthemis orientalis (L.) Degen subsp. incana (Boiss.) Hayek var.
spruneri (Boiss. & Heldr.) Hayek
FE 4:152 MFG 2:427
L/121 L/499 L/530 L/607 L/1014 L/1973 L/2200
Anthemis filicaulis (Boiss. & Heldr.) Greuter
Asteraceae
Ammanthus filicaulis Boiss. & Heldr.
Anthemis filicaulis (Boiss. & Heldr.) Greuter f. discoidea Greuter
EFK :313 FAe :632 FCA :58 FCSI :26 FE 4:158
L/338 L/518 L/554 L/1111
Anthemis glaberrima (Rech.f.) Greuter
Asteraceae
Ammanthus glaberrimus Rech.f.
EFK :314 FCA :58 FE 4:59 RDB :36
L/138 L/338 L/537 L/554 L/932 L/1111 L/1710
Anthemis laconica Franzén
Asteraceae
MFG 2:425
L/499 L/607 L/1014 L/1963
Anthemis parnesia Boiss. & Heldr.
Asteraceae
FAe :626 FE 4:153 FGC 2:60 FGS 2:49
L/607
127
128
129
130
131
Taxon
Anthemis cretica L. subsp. sibthorpii (Griseb.) Govaerts
Asteraceae
Anthemis sibthorpii Griseb.
Anthemis orientalis (L.) Degen subsp. sibthorpii (Griseb.) Hayek
Santolina montana Sm.
FE 4:149 MFG 2:425 RDB :42
L/404a L/1014 L/1972 L/1973
Taxon
Qualifiers
126
Taxon
Number
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+
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+
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WAe
.
StE
Pe
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SPi
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IoI
.
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NPi
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EC
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.
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NC
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NE
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NAe
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Kik
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+
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KK
.
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EAe
Distribution
Qualifier
174 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Anthemis peregrina L. subsp. heracleotica (Boiss. & Heldr.)
Georgiou
Asteraceae
Anthemis peregrina L. subsp. guicciardii (Heldr. ex Sart.) Zangeri
Anthemis guicciardii Heldr. ex Sart.
Anthemis peregrina L. var. guicciardii (Heldr. ex Sart.) Boiss.
Anthemis peregrina L. var. heracleotica Boiss & Heldr.
Anthemis tomentosa L. subsp. heracleotica (Boiss. & Heldr.)
R.Fernandes
FAe :627 FE 4:154 MCL 2:32
L/119 L/128 L/516 L/530 L/773 L/1871 L/1932
Anthemis rhodensis Boiss.
Asteraceae
FAe :625 FT 5:191 RDB :38
L/197 L/1111
Anthemis rigida Boiss. ex Heldr. subsp. liguliflora (Halácsy) Greuter
Asteraceae
Anthemis cretica (L.) Nyman var. liguliflora Halácsy
Anthemis heracleotica sensu Rechinger Fl. Aeg. p.p.
Anthemis pusilla Greuter var. liguliflora (Halácsy) Greuter
EFK :314 FAe :627 FAe :628 FCA :58 FCSI :26
L/119 L/551 L/773 L/1111
Anthemis rosea Sm. subsp. rosea
Asteraceae
FAe :626 FT 5:200 RDB :40
L/796 L/975
Anthemis samariensis Turland
Asteraceae
L/2255
Anthemis scopulorum Rech.f.
Asteraceae
Anthemis tomentosa L. subsp. scopulorum (Rech.f.) Grierson
EFK :315 FAe :627 FCA :58 FE 4:152 FT 5:204
B/3060c L/169 L/197 L/206 L/446 L/495 L/773 L/776 L/861 L/913 L/1049 L/1078 L/1104 L/1111 L/1710 L/1878 L/1917 L/1924
132
133
134
135
136
137
+
.
+
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.
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+
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+
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+
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+
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+
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+
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 175
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Anthemis scopulorum Rech.f. var. discoidea Georgiou
Asteraceae
L/773 L/913 L/1924
Anthemis scopulorum Rech.f. var. ofidousa Georgiou
Asteraceae
Anthemis pusilla subsp. ammanthiformis sensu Greuter & Rechinger,
1967 p.p.
L/119 L/773 L/1924
Anthemis scopulorum Rech.f. var. scopulorum
Asteraceae
L/773 L/913 L/1924
Anthemis tomentella Greuter
Asteraceae
Ammanthus maritimus Fl. Aeg. p.p.; Halácsy 1912
Ammanthus tomentellus sensu Gandoger 1915, 1916, 1920, p.p.;
Rechinger 1943
EFK :313 FAe :632 FCA :58 FCSI :26 FE 4:158
L/255 L/338 L/518 L/554 L/1111
Anthemis werneri Stoj. & Acht.
Asteraceae
Anthemis flexicaulis Rech.f.
FAe :626 FAe :627 FE 4:153 FE 4:154
L/323b L/773 L/776 L/928 L/1018 L/1627 L/1710 L/1871 L/2181
Anthemis werneri Stoj. & Acht. subsp. insularis Georgiou
Asteraceae
FE 4:154
L/773 L/1018 L/1627 L/2181
Anthemis werneri Stoj. & Acht. subsp. werneri
Asteraceae
Anthemis guicciardii sensu Rechinger Fl. Aeg. p.p.; Hansen 1971
Anthemis heracleotica sensu Rechinger Fl. Aeg. p.p.
Anthemis tomentosa sensu Rechinger 1960 p.p.
FAe :626 FAe :627 FE 4:153
L/128 L/323b L/773 L/776 L/783 L/1018 L/2181
139
140
141
142
143
144
145
Taxon
Anthemis scopulorum Rech.f. var. conica Georgiou
Asteraceae
L/773 L/1924
Taxon
Qualifiers
138
Taxon
Number
.
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StE
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+
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Pe
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+
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WAe
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IoI
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SPi
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NPi
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.
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EC
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NC
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NE
+
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+
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NAe
+
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+
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+
+
+
+
Kik
.
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+
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+
.
KK
.
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.
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+
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+
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EAe
Distribution
Qualifier
176 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Aquilegia ottonis Orph. ex Boiss. subsp. taygetea (Orph.) Strid
Ranunculaceae
Aquilegia amaliae var. taygetea (Orph.) Hayek
Aquilegia taygetea Orph.
AFE 8:231 FE 1:288 FHe 2:73 MCL 4:396 MFG 1:228 RDB :46
L/1963
Arabis cretica Boiss. & Heldr.
Brassicaceae
Arabis muralis Bertol var. cretica (Boiss. & Heldr.) B.M.G.Jones
Arabis serpillifolia Vill. subsp. cretica (Boiss. & Heldr.) B.M.G.Jones
AFE 10:198 EFK :116 EFK :116 FCA :79 FCSI :39 FE 1:355 FHe 2:188 MCL 3:54 MFG 1:264
L/856 L/1111 L/1766
Arabis subflava B.M.G.Jones
Brassicaceae
Arabis ochroleuca Boiss. & Heldr. non (Lam.) Lam.
AFE 10:207 FE 1:356 FHe 2:190 MCL 3:50 MFG 1:267
L/1963
Arenaria aegaea Rech.f.
Caryophyllaceae
Arenaria serpyllifolia L. subsp. aegaea (Rech.f.) Akeroyd
AFE 6:28 EFK :78 FAe :141 FCA :45 FCSI :17 FCSII : FE 1:147 FHe 1:167 FT 11:45 MCL 1:167
L/150 L/169 L/196 L/197 L/201 L/206 L/255 L/268 L/537 L/759a L/776 L/896 L/913 L/932 L/1049 L/1111 L/1710 L/1917
148
149
150
151
Aquilegia ottonis Orph. ex Boiss. subsp. ottonis
Ranunculaceae
Aquilegia amaliae Heldr. ex Boiss. subsp. meridionalis Quézel &
Contandr.
AFE 8:231 FE 1:288 FHe 2:73 MCL 4:395 MFG 1:227 RDB :46
L/828 L/1963
147
!
Anthyllis splendens Willd.
Fabaceae
Anthyllis aegaea Turrill
EFK :170 FAe :376 FCA :102 FCSI :56 FE 2:178 MCL 4:6 RDB :44
L/145 L/206 L/255 L/707 L/1111 L/1201
146
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 177
Arenaria fragillima Rech.f.
Caryophyllaceae
Arenaria oxypetala Sm. sensu Halácsy
Arenaria pamphylica sensu Hayek.
AFE 6:24 EFK :77 FAe :139 FCA :45 FCSI :17 FE 1:146 FHe 1:163 MCL 1:163 MFG 1:87
L/518 L/554 L/759a L/1111
Arenaria gionae Gustavsson
Caryophyllaceae
AFE 6:22 FE 1:145 FHe 1:160 MCL 1:162 MFG 1:89 RDB :48
L/1032
Arenaria guicciardii Heldr. ex Boiss.
Caryophyllaceae
AFE 6:29 EFK :78 FAe :141 FCA :45 FE 1:147 FHe 1:167 FT 2:30 MCL 1:165 MFG 1:91
L/121 L/197 L/252 L/530 L/587 L/607 L/796 L/1111 L/1123 L/1880 L/1957 L/1963 L/2200
Arenaria leucadia Phitos & Strid
Caryophyllaceae
FHe 1:165 RDB :50
L/1135
Arenaria peloponnesiaca Rech.f.
Caryophyllaceae
AFE 6:28 FHe 1:167 MCL 1:168
L/607 L/607 L/1800
Arenaria phitosiana Greuter & Burdet
Caryophyllaceae
Arenaria litoralis Phitos
AFE 6:23 FE 1:145 FHe 1:162 MCL 1:164 RDB :52
L/443 L/715 L/767 L/928
Arenaria runemarkii Phitos
Caryophyllaceae
FHe 1:166 FT 11:44
L/539 L/540 L/759a
153
154
155
156
157
158
159
Taxon
Arenaria filicaulis Fenzl subsp. teddii (Turrill) Strid
Caryophyllaceae
Arenaria teddii Turrill
AFE 6:23 FAe :140 FE 1:145 FHe 1:161 MCL 1:164 MFG 1:86
L/1107
Taxon
Qualifiers
152
Taxon
Number
.
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+
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StE
Pe
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+
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+
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IoI
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WAe
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SPi
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NPi
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EC
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NC
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NE
.
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NAe
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Kik
.
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+
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+
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KK
+
.
.
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+
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EAe
Distribution
Qualifier
178 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Arum idaeum Gand.
Araceae
Arum maculatum sensu Greuter 1973
EFK :409 FCA :159 FE 5:270
L/212 L/255 L/430 L/559 L/1111
Arum purpureospathum P.C.Boyce
Araceae
EFK :410 FCA :159 FCSI :91 RDB :58
L/138 L/212 L/397 L/559 L/1111 L/2176
167
Armeria maritima Willd. subsp. smolikana Babalonas
Plumbaginaceae
MCL 4:313 MFG 1:750
L/950
164
166
Armeria johnsenii Papan. & Kokkini
Plumbaginaceae
Armeria undulata sensu Rechinger 1960, eub.
MCL 4:315
L/323a L/788 L/1871
163
Armeria sancta Janka
Plumbaginaceae
Armeria majellensis Boiss. var. brachyphylla Boiss.
FAe :431 FE 3:33 MCL 4:318 PFPB 2:13
L/1973
Armeria icarica J.R.Edmonson
Plumbaginaceae
FT 7:504 MCL 4:315
L/539 L/540
162
165
Aristolochia microstoma Boiss. & Spruner
Aristolochiaceae
AFE 3:120 FAe :174 FE 1:89 FGC 3:90 FGS 2:75 FHe 1:74 MCL 1:53
L/415 L/451 L/530 L/607 L/781 L/1963 L/1973 L/1981
161
?
Aristolochia cretica Lam.
Aristolochiaceae
AFE 3:116 EFK :62 FAe :175 FCA :37 FCSI :12 FE 1:88 FHe 1:71 MCL 1:53
L/518 L/554 L/709 L/1111
160
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 179
Asperula arcadiensis Sims
Rubiaceae
FE 4:12 FGC 1:735 MFG 2:296
L/511 L/512 L/607 L/1199 L/1844 L/1963
Asperula aristata L. subsp. thessala (Boiss. & Heldr.) Hayek
Rubiaceae
Asperula aristata L. subsp. peristeraea Halácsy ex Hayek
Asperula longiflora Waldst. & Kit var. thessala (Boiss. & Heldr.)
Halácsy
Asperula thessala Boiss. & Heldr.
FE 4:6 FGC 1:741 MFG 2:286
L/178 L/196 L/565 L/1107 L/1217 L/1246 L/1963 L/1973 L/2200 L/2201
Asperula baenitzii Heldr. ex Boiss.
Rubiaceae
FE 4:13 FGC 1:734 MFG 2:297,298 RDB :60
L/530 L/736 L/781 L/1062 L/1103
Asperula boissieri Heldr. ex Boiss.
Rubiaceae
Asperula gionae Quézel & Contandr.
Asperula lutea Sm. var. pulvinaris Boiss.
FE 4:9 FGC 1:736 MFG 2:292
L/121 L/160 L/512 L/607 L/1963 L/2013 L/2200
Asperula boryana (Walpers) Ehrend.
Rubiaceae
Galium boryanum Walp.
Sherardia pusilla Chaub. & Bory
FE 4:14 FGC 1:717 MFG 2:298
L/511 L/512 L/607 L/1963
Asperula brachyphylla Trigas & Iatroú
Rubiaceae
L/1871 L/2223
Asperula crassula Greuter & Zaffran
Rubiaceae
Asperula crassifolia sensu Gandoger 1916, Rechinger 1943
EFK :236 FAe :568 FCA :137 FCSI :78 RDB :62
L/253 L/518 L/1111 L/1583
169
170
171
172
173
174
175
Taxon
Asperula abbreviata (Halácsy) Rech.f.
Rubiaceae
Asperula lutea var. abbreviata Halácsy
FAe :566 FE 4:8 MFG 2:294
L/196 L/869 L/1711
Taxon
Qualifiers
168
Taxon
Number
.
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+
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IoI
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+
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WAe
StE
Pe
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+
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SPi
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+
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NPi
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+
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Distribution
Qualifier
180 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Asperula elonea Iatroú & T.Georgiadis
Rubiaceae
FGC 1:737 RDB :64
L/607 L/917 L/1963
Asperula euboea (Ehrend.) Trigas
Rubiaceae
MFG 2:290
L/323b L/1871 L/1963
Asperula icarica Ehrend. & Schönb.-Tem.
Rubiaceae
FT 7:742
L/539 L/540
Asperula idaea Halácsy
Rubiaceae
Asperula idaea Halácsy var. fallax Rech.f.
Asperula idaea Halácsy var. lassitica Rech.f.
EFK :235 FCA :137 FE 4:7 FGC 1:737 MFG 2:289
B/3060f L/518 L/856 L/1111 L/1766
Asperula lilaciflora Boiss. subsp. coa (Rech.f.) Ehrend.
Rubiaceae
Asperula coa Rech.f.
Asperula lutea var. rigida sensu FAe rhodos
FAe :566 FT 7:748
L/150 L/150 L/197 L/288 L/1111
Asperula lilaciflora Boiss. subsp. runemarkii Ehrend. &
Schönb.-Tem.
Rubiaceae
FT 7:747
L/539 L/540 L/1957
Asperula lutea Sm.
Rubiaceae
FE 4:8 FGC 1:737 MFG 2:290
L/323b L/512 L/607 L/1830 L/1963 L/1973 L/1981 L/2013 L/2200
Asperula malevonensis Ehrend. & Schönb.-Tem.
Rubiaceae
Asperula boissieri Heldr. ex Boiss. var. transiens Halácsy
FGC 1:736 MFG 2:294
L/911 L/1963
176
177
178
179
180
181
182
183
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 181
Asperula muscosa Boiss. & Heldr.
Rubiaceae
FE 4:13 FGC 1:734 MFG 2:297 RDB :66
Asperula naufraga Ehrend. & Guterm.
Rubiaceae
L/1689 L/1800
Asperula nitida Sm. subsp. mytilinica Ehrend.
Rubiaceae
Asperula nitida Sm. var. hirtella Boiss. p.p.
FT 7:745 MFG 2:294
L/321 L/1957
Asperula oetaea (Boiss.) Heldr. ex Halácsy
Rubiaceae
Asperula lutea Sm. var. oetaea Boiss.
Asperula lutea subsp. oetaea (Boiss.) Nyman
FE 4:8 FGC 1:739 MFG 2:292
L/121 L/512 L/1963 L/2013 L/2200
Asperula ophiolithica Ehrend.
Rubiaceae
Asperula stricta auct. eur., non Boiss.
Asperula stricta auct. eur., non Boiss. subsp. pruinosa Ehrend.
Asperula stricta auct. eur., non Boiss. subsp. stricta
FE 4:7 MFG 2:287
L/323b L/1006 L/1871
Asperula pinifolia (Boiss.) Heldr. ex Ehrend. & Schönb.-Tem.
Rubiaceae
Asperula lutea var. pinifolia Boiss.
Asperula nitida sensu auct. graec.
FE 2:8 FGC 1:734 FGC 1:738 FT 7:744 MFG 2:293
L/121 L/512 L/2200
Asperula pubescens (Willd.) Ehrend. & Schönb.-Tem.
Rubiaceae
Asperula incana Sm.
Crucianella pubescens Willd.
EFK :236 FCA :137 FCSI :78 FE 4:11 MFG 2:295
L/856 L/1111 L/1766
185
186
187
188
189
190
191
Taxon
Asperula mungieri Boiss. & Heldr.
Rubiaceae
Asperula lutea Sm. subsp. mungieri (Boiss. & Heldr.) Ehrend. & Krendl
FGC 1:739 MFG 2:291
L/512 L/607 L/1915 L/1963
Taxon
Qualifiers
184
Taxon
Number
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Distribution
Qualifier
182 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Asperula pulvinaris (Boiss.) Heldr. ex Boiss.
Rubiaceae
FE 4:8 FGC 1:736 MFG 2:295
L/530 L/781
Asperula rigida Sm.
Rubiaceae
Asperula kritensis Coust. & Gand.
Galium suberosum sensu auct. cret., non Sm.
EFK :236 FCA :137 FCSI :78 FE 4:13 MFG 2:298
L/268 L/537 L/762 L/917 L/1111 L/1766
Asperula rigidula Halácsy
Rubiaceae
Asperula lutea Sm. subsp. rigidula (Halácsy) Ehrend.
Asperula lutea Sm. var. rigida Boiss.
Asperula lutea Sm. var. rigidula Halácsy
Asperula stricta Boiss. var. stricta Boiss.
Asperula stricta Boiss. var. tomentosa Boiss.
FE 2:8 FGC 1:738 MFG 2:291
L/323b L/512 L/530 L/607 L/781 L/1467 L/1871 L/1915
Asperula samia Christod. & T.Georgiadis
Rubiaceae
FT 10:215 MFG 2:289
L/182 L/796 L/975
Asperula saxicola Ehrend.
Rubiaceae
FE 4:14 MFG 2:299 RDB :68
L/410c L/607 L/1963
Asperula suberosa Sm.
Rubiaceae
Asperula athoa Boiss.
FAe :568 FE 4:7 MFG 2:289
L/1972 L/1973
Asperula suffruticosa Boiss. & Heldr.
Rubiaceae
FAe :567 FE 4:7 FGC 1:738 MFG 2:287
L/323b L/1871
192
193
194
195
196
197
198
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 183
Asplenium creticum Lovis, Reichst. & Zaffran
Aspleniaceae
EFK :41 FCA :30 FE 1:22 FT 10:4 MCL 1:10 MFG 1:24
L/255 L/419 L/518 L/945 L/1111 L/1153
Astragalus agraniotii Orph. ex Boiss.
Fabaceae
FE 2:121 FGC 1:434 MCL 4:33 MFG 1:472 RDB :70
L/607 L/911 L/1963 L/2235
Astragalus apollineus Boiss. & Heldr.
Fabaceae
FE 2:122 FGC 1:431 MCL 4:61 MFG 1:476
L/512 L/843 L/2235
Astragalus austroaegaeus Rech.f.
Fabaceae
Astragalus insulae-carpathi Eig
Astragalus tauricolus auct. non Boiss. Hayek Prodr. fl. Balc.; Rechinger
fl. Aeg.
Astragalus tauricolus Boiss. subsp. austoaegaeus (Rech.f.) Ponert
Astragalus tauricolus Boiss. var. niveus Barbey
EFK :147 FAe :318 FCA :103 FE 2:114 FT 3:92 MCL 4:36
L/197 L/581 L/707 L/1111 L/2235
Astragalus creticus Lam.
Fabaceae
EFK :145 FAe :318 FCA :103 FCSI :56 FE 2:117 FE 2:117 FT 3:122 MCL 4:21 MCL 4:21 MFG 1:465
MFG 1:465
L/518 L/522 L/587 L/594 L/796 L/843 L/847 L/856 L/876 L/1111 L/1137 L/1600 L/2013 L/2200 L/2235
201
202
203
204
205
!
Asperula tournefortii Sieber ex Sprengel
Rubiaceae
Asperula majori Barbey (included in taxon)
Asperula tournefortii Sieber ex Sprengel var. revoluta Rech.f.
EFK :236 FAe :565 FCA :137 FCSI :78 FE 4:13 FT 7:764
L/196 L/197 L/206 L/268 L/537 L/1111
200
Taxon
Asperula taygetea Boiss. & Heldr.
Rubiaceae
EFK :236 FAe :565 FCA :137 FE 4:11 FGC 1:735 MFG 2:296
L/119 L/607 L/765 L/1111 L/1588 L/1917 L/1963
Taxon
Qualifiers
199
Taxon
Number
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Distribution
Qualifier
184 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Astragalus dolinicola (Brullo & Giusso) Brullo & Giusso
Fabaceae
Astracantha dolinicola Lam.
FCSII :
L/522 L/594 L/2235
Astragalus drupaceus Orph. ex Boiss.
Fabaceae
FE 2:117 FGC 1:439 MCL 4:33 RDB :72
L/395 L/607 L/674 L/1963 L/2235
Astragalus graecus Boiss. & Spruner var. rhodensis Pamp.
Fabaceae
Astragalus gilvus sensu Agerer-Kirchhof (1976)
FAe :318 FT 3:100
L/197 L/913
Astragalus ictericus Dingler
Fabaceae
FE 2:116 MCL 4:35
L/2235
209
210
211
212
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Astragalus cylleneus Fisch.
Fabaceae
Astracantha thracica (Griseb.) Podlech subsp. cyllenea (Fisch.) Greuter
Astragalus calavrytensis Beauverd & Topali
Astragalus parnassi Boiss. subsp. cylleneus (Fisch.) Hayek
Astragalus thracicus Griseb. subsp. cylleneus (Boiss. & Heldr.) Strid
FE 2:117 FT 3:166 MCL 4:30 MFG 1:467
L/160 L/395 L/512 L/876 L/942 L/1152 L/2235
208
Astragalus creticus Lam. subsp. creticus
Fabaceae
Astracantha cretica (Lam.) Podlech
Astragalus creticus Lam. f. glaberrimus Greuter & Zaffran
EFK :145 FAe :318 FCA :103 FCSI :56 FE 2:117 FT 3:122 MCL 4:21 MFG 1:465
L/518 L/796 L/856 L/876 L/1137 L/1600 L/2235
Astragalus creticus Lam. subsp. minoicus Brullo & Giusso
Fabaceae
L/2235
!
207
206
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 185
Astragalus laconicus Iatroú & Kit Tan
Fabaceae
L/1210b L/1600a L/1963 L/2235
Astragalus lacteus Heldr. & Sart. ex Boiss.
Fabaceae
Astragalus baldaccii Degen
FE 2:121 MCL 4:43 MFG 1:473
L/512 L/607 L/843 L/1112 L/1832d L/2013 L/2235
Astragalus maniaticus Kit Tan & Strid
Fabaceae
L/702 L/1963 L/2235
Astragalus nummularius Lam.
Fabaceae
EFK :147 FAe :318 FCA :103 FCSI :56 FE 2:116 MCL 4:56
L/1111 L/2235
Astragalus thracicus Griseb. subsp. monachorum (Širj.) Strid
Fabaceae
Astracantha thracica (Griseb.) Podlech subsp. monachorum (Širj.)
Greuter
Astracantha monachorum (Širj.) Podlech
Astragalus monachorum Širj.
FE 2:118 MCL 4:29 MFG 1:467
L/785 L/876 L/942 L/1973 L/2235
Asyneuma giganteum (Boiss.) Bornm.
Campanulaceae
Podanthum giganteum Boiss.
EFK :297 FAe :605 FCA :42 FT 6:70 MCL 1:121 RDB :76
L/138 L/197 L/758 L/1111
Athamanta arachnoidea Boiss. & Orph.
Apiaceae
Athamanta macedonica (L.) Sprengel subsp. arachnoidea (Boiss. &
Orph.) Tutin
Athamanta macedonica (L.) Sprengel var. arachnoidea (Boiss. & Orph.)
Halácsy
FE 2:340 MFG 1:687
L/511 L/607 L/1963
214
215
216
217
218
219
220
Taxon
Astragalus idaeus Bunge
Fabaceae
Astragalus hirsutus Raul, non Vahl.
EFK :147 FAe :320 FCA :103 FCSII 0: FE 2:121 MCL 4:50 MFG 1:472 RDB :74
L/365 L/750 L/1111 L/1861 L/2235
Taxon
Qualifiers
213
Taxon
Number
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EAe
Distribution
Qualifier
186 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Atriplex recurva D’Urv.
Amaranthaceae
Atriplex tatarica auct..
Atriplex tatarica L. subsp. recurva (D’Urv.) Rech.f.
Atriplex tatarica L. var. recurva Halácsy
Atriplex tatarica L. var. sibthorpii Halácsy
AFE 5:40 EFK :69 FAe :122 FCA :54 FCSI :24 FE 1:116 FHe 1:124 MCL 1:294
L/111 L/128 L/163 L/196 L/206 L/268 L/447 L/537 L/554 L/759a L/776 L/783 L/861 L/928 L/1111 L/1627 L/1710 L/2181
Aubrieta erubescens Griseb.
Brassicaceae
AFE 10:214 FE 1:357 FHe 2:193 MCL 3:58 MFG 1:272 RDB :78
L/1119 L/1972 L/1973
Aubrieta glabrescens Turrill
Brassicaceae
Aubrieta gracilis Spruner subsp. glabrescens (Turrill) Akeroyd
Aubrieta deltoidea (L.) DC. var. degeniana Bald.
Aubrieta gracilis Spruner f. degeniana (Bald.) Hayek
Aubrieta gracilis Spruner subsp. scardica (Wettst.) Phitos var. degeniana
(Bald.) Phitos
AFE 10:214 FE 1:358 FHe 2:195 MCL 3:58 MFG 1:272 RDB :80
L/1119
Aubrieta gracilis Spruner
Brassicaceae
AFE 10:214 FE 1:357 FHe 2:194 MCL 3:58 MFG 1:274
L/121 L/1119 L/2200
Aubrieta scyria Halácsy
Brassicaceae
AFE 10:214 FE 1:357 FHe 2:193 MCL 3:58 RDB :82
L/404b L/1119 L/1871
Aubrieta thessala H.Boissieu
Brassicaceae
Aubrieta gracilis Spruner ex Boiss. subsp. thessala (Boissieu) Maire &
Petitm.
Aubrieta gracilis Spruner var. thessala (Boissieu) Hayek
Aubrieta intermedia auct. non Heldr. & Orph.
AFE 10:215 FE 1:357 FHe 2:195 MCL 3:58 MFG 1:274
L/1119 L/1217 L/1832d
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222
223
224
225
226
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PATTERNS OF PLANT ENDEMISM IN GREECE 187
Aurinia moreana Tzanoud. & Iatroú
Brassicaceae
Alyssum orientale Ard. var. alpinum Halácsy
Alyssum saxatile L. var. alpinum (Halácsy) Halácsy
AFE 11:70 FE 1:370 FHe 2:230 MCL 3:59 MFG 1:280
L/499 L/607 L/831 L/1193 L/1844 L/1963
Ballota pseudodictamnus (L.) Benth. subsp. pseudodictamnus
Lamiaceae
FAe :513 FCA :95 FE 3:150 FGC 2:533 FGC 2:533 FGS 1:87 FT 7:157 MCL 3:281
L/119 L/554 L/932 L/1052 L/1096 L/1111
Bellevalia brevipedicellata Turrill
Asparagaceae
EFK :357 FCA :182 FCSI :108 FE 5:45 RDB :84
L/255 L/707 L/762 L/1111 L/1194 L/1574 L/1583 L/2224
Bellevalia hyacinthoides K.Perss. & Wendelbo
Asparagaceae
Puschkinia dubia Kunth
Foxia spicata (Boiss.) Parl.
Bellevalia spicata Boiss.
Hyacinthus spicatus Sm.
Strangweia hyacinthoides Sm.
Strangweia spicata (Sm.) Boiss.
FE 5:44 FGC 3:263 FGS 1:84
L/63 L/282 L/530 L/776 L/1111 L/1176 L/1244 L/1846 L/1963 L/2224
Bellevalia sitiaca Kypr. & Tzanoud.
Asparagaceae
Bellevalia brevipedicellata sensu auct. cret. orient., non Turrill
Muscari macrocarpum sensu auct. Siteia, non Sweet
FCSII :
L/336 L/554 L/1111 L/1574 L/2093 L/2224
Bellis longifolia Boiss. & Heldr.
Asteraceae
EFK :305 FAe :607 FCA :59 FCSI :27 FE 4:112 MFG 2:405
L/518 L/522 L/551 L/554 L/1111
228
229
230
231
232
233
Taxon
Aurinia gionae (Quézel & Contadriopoulos) Greuter & Burdet
Brassicaceae
Alyssum corymbosum sensu Halácsy Consp. Fl. Graec. p.p. minor
Alyssum gionae Quézel & Contadriopoulos
AFE 11:66 FE 1:370 FHe 2:229 MCL 3:58 MFG 1:279
L/121 L/607 L/828 L/830 L/1963 L/2200
Taxon
Qualifiers
227
Taxon
Number
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IoI
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WAe
StE
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NPi
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Distribution
Qualifier
188 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Berteroa obliqua (Sm.) DC. subsp. pindicola Kit Tan
Brassicaceae
Berteroa obliqua (Sm.) DC. var. pindicola (Halácsy) Quezél & Contadr.
Berteroa mutabilis (Vemt.) DC. subsp. obliqua (Sm.) Bald. sensu Bald.
1897
Berteroa obliqua sensu auct., Halácsy 1894
Berteroa stricta Boiss. & Heldr. var. pindicola Halácsy
AFE 11:62 FE 1:369 FHe 2:226 MFG 1:303
L/1143
Beta nana Boiss. & Heldr.
Amaranthaceae
AFE 5:14 FE 1:111 FHe 1:112 MCL 1:297 MFG 1:78 RDB :86
L/455 L/512 L/513 L/587 L/607 L/1180 L/1246 L/1963 L/1973
Biarum davisii Turrill
Araceae
FCA :159 FE 5:271 RDB :88
L/346 L/906 L/1111 L/1236 L/1766 L/2236
Biarum fraasianum (Schott) Nyman
Araceae
Biarum fraasianum (Schott) N.E.Br. nom. superfl.
Ischarum fraasianum Schott
FE 5:271 FGC 3:294 RDB :90
L/906 L/1120 L/2236
Biarum rhopalospadix C.Koch
Araceae
Cyllenium spruneri (Boiss.) Schott
Ischarum spruneri (Boiss.) Schott
Biarum spruneri Boiss.
FE 5:271 FGC 3:293
L/308c L/607 L/906 L/1830 L/1871 L/1963 L/2236
Biarum tenuifolium (L.) Schott subsp. abbreviatum (Schott) Richter
Araceae
Biarum tenuifolium (L.) Schott var. abbreviatum (Schott) Engler
Biarum abbreviatum Schott
B/3060c L/276 L/906 L/1111 L/1917 L/1963 L/2236 L/2236
234
235
236
237
238
239
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 189
Bolanthus chelmicus Phitos
Caryophyllaceae
AFE 7:135 FE 1:222 FHe 1:327 MCL 1:171
L/607 L/609 L/1963
Bolanthus chelmicus Phitos subsp. chelmicus
Caryophyllaceae
FE 1:222 FHe 1:328 MCL 1:171
L/609 L/1963
Bolanthus chelmicus Phitos subsp. meteoricus Phitos
Caryophyllaceae
Bolanthus thymifolius sensu Phitos 1981
FHe 1:328
L/609 L/1963
Bolanthus creutzburgii Greuter
Caryophyllaceae
AFE 7:135 FCA :45 FE 1:223 FHe 1:329 MCL 1:170 MFG 1:173
L/408 L/518 L/609 L/1111 L/1766
Bolanthus fruticulosus (Bory & Chaub.) Barkoudah
Caryophyllaceae
Gypsophila fruticulosa (Bory & Chaub.) Boiss.
Petrorhagia suffruticosa Rech.f. & Phitos
Saponaria fruticulosa Bory & Chaub.
AFE 1:134 FE 1:222 FHe 1:326 MCL 1:171
L/119 L/267 L/607 L/609 L/765 L/1118 L/1963
Bolanthus graecus (Schreber) Barkoudah
Caryophyllaceae
Cucubalus polygonoides Willd.
Gypsophila graeca (Schreber) Britten
Gypsophila ocellata Sm.
Gypsophila polygonoides (Willd.) Halácsy
Gypsophila polygonoides subsp. ocellata (Sm.) Hayek
Saponaria graeca Schreber
Saponaria polygonoides (Willd.) Jaub & Spach
Saponaria thymifolia var. ocellata (Sm.) Boiss.
Silene polygonoides (Willd.) Poiret
AFE 7:134 FE 1:222 FHe 1:327 MCL 1:171 MFG 1:172
L/119 L/530 L/609 L/765 L/869 L/1871 L/2201
241
242
243
244
245
246
Taxon
Biarum tenuifolium (L.) Schott subsp. idomenaeum P.C.Boyce &
Athanasiou
Araceae
Biarum tenuifolium auct. kret., Rechinger 1949
Biarum zelebori cret. sensu Greuter 1973, p.p.
EFK :410 FCA :159
L/255 L/276 L/906 L/2236
Taxon
Qualifiers
240
Taxon
Number
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EAe
Distribution
Qualifier
190 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Bolanthus intermedius Phitos
Caryophyllaceae
AFE 7:134 FE 1:223 FHe 1:328 MCL 1:171
L/608 L/609 L/1871
Bolanthus laconicus (Boiss.) Barkoudah
Caryophyllaceae
Gypsophila fasciculata Marg. & Reut. var. laconica Boiss.
Gypsophila laconica (Boiss.) Boiss.
AFE 7:134 FE 1:222 FHe 1:326 MCL 1:171
L/512 L/607 L/609 L/1963
Bolanthus thymifolius (Sm.) Phitos
Caryophyllaceae
Bolanthus graecus var. thymifolius (Sm.) Barkoudah
Bolanthus thessalus (Jaub. & Spach) Barkoudah
Gypsophila polygonoides (Willd.) Halácsy subsp. thessala (Jaub. &
Spach) Hayek
Gypsophila polygonoides (Willd.) Halácsy subsp. thymifolia (Sm.) Hayek
Gypsophila polygonoides (Willd.) Halácsy var. thymifolia (Sm.) Hayek
Gypsophila thessala (Jaub. & Spach) Halácsy
Gypsophila thymifolia Sm.
Saponaria thymifolia (Sm.) Boiss.
AFE 7:135 FE 1:223 FHe 1:328 MCL 1:171
L/609 L/1871
Bornmuellera baldaccii (Degen) Heywood subsp. rechingeri Greuter
Brassicaceae
AFE 11:72 FE 1:371 FHe 2:234 MCL 3:67 MFG 1:307
L/854
Bornmuellera tymphaea (Hausskn.) Hausskn.
Brassicaceae
Alyssum tymphaeum (Hausskn.) Halácsy
Ptilotrichon tymphaeum (Hausskn.) Halácsy
Vesicaria tymphaea Hausskn.
AFE 11:73 FE 1:371 FHe 2:234 MCL 3:67 MFG 1:307
L/159 L/513 L/831 L/854 L/1851 L/1901
Brachypodium sylvaticum (Huds.) P.Beauv. subsp. creticum
H.Scholz & Greuter
Poaceae
Brachypodium pinnatum sensu Zaffran 1976
EFK :388 FCA :165 FE 5:189 MFG 2:793
L/430 L/1111 L/1766 L/2200
247
248
249
250
251
252
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PATTERNS OF PLANT ENDEMISM IN GREECE 191
?
257
258
Bromus elidis H.Scholz
Poaceae
L/586 L/2237
?
256
Bufonia stricta (Sm.) Gürke
Caryophyllaceae
Bufonia brachyphylla Boiss. & Heldr. nom. illeg.
Moehringia stricta Sm.
AFE 6:69 EFK :79 FAe :134 FCA :45 FCSI :18 FE 1:161
FGC 1:245 FHe 1:193 MCL 1:173 MFG 1:107
L/408 L/518 L/530 L/554 L/707 L/736 L/781 L/1111 L/1963
Bromus pindicus Hausskn.
Poaceae
FE 5:185 FGC 3:389 MFG 2:791
L/159 L/528
Brassica cretica Lam. subsp. laconica M.Gustafsson & Snogerup
Brassicaceae
AFE 11:239 FE 1:407 FHe 2:282
L/274 L/607 L/1026 L/1877 L/1963
255
Brassica cretica Lam. subsp. cretica
Brassicaceae
Brassica cretica Lam. subsp. nivea (Boiss. & Spruner) Gustafsson. &
Snogerup
Brassica cretica Lam. var. nivea (Boiss. & Spruner) O.E.Schulz
(included in taxon)
Brassica montana Pourret subsp. cretica (Lam.) O.E.Schulz
Brassica nivea Boiss. & Spruner
Brassica nivea Boiss. & Spruner subsp. cretica (Lam.) Onno
AFE 11:239 EFK :125 FCA :80 FCSI :39 FE 1:407 FHe 2:281 MCL 3:68
L/122 L/197 L/274 L/443 L/928 L/1111 L/1627 L/1973
!
254
Taxon
Brassica cadmea Heldr. ex O.E.Schulz
Brassicaceae
Brassica fruticulosa Cyr.
Sinapis radicata S. & Sm.
AFE 11:249 FE 1:408 FGC 1:79 FHe 2:285 MCL 3:68 PFPB 1:453
Taxon
Qualifiers
253
Taxon
Number
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EAe
Distribution
Qualifier
192 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Bufonia stricta (Sm.) Gürke subsp. cecconiana (Bald.) Rech.f.
Caryophyllaceae
Bufonia brachyphylla Boiss & Heldr. var. cecconiana Bald.
Bufonia brachyphylla sensu auct. p.p.
Bufonia stricta (Sm.) Gurke var. cecconiana (Bald.) Gurke
Bufonia stricta var. typica sensu Halácsy Consp. Fl. Graec. p.p.
AFE 6:69 EFK :79 FAe :134 FCA :45 FCSI :18 FHe 1:193 MCL 1:173 MFG 1:107
L/408 L/518 L/554 L/707 L/1111 L/1963
Bufonia stricta (Sm.) Gürke subsp. stricta
Caryophyllaceae
Bufonia stricta (Sm.) Gurke var. hymettia Halácsy
Bufonia stricta (Sm.) Gurke var. typica Halácsy p.p.
Bufonia stricta var. cecconiana sensu auct. p.p.
AFE 6:69 EFK :79 FAe :134 FCA :45 FCSI :18 FGC 1:245 FHe 1:193 MCL 1:173 MFG 1:107
L/408 L/518 L/530 L/607 L/736 L/1111 L/1871 L/1963
Bufonia stricta (Sm.) Gürke subsp. stricta var. euboica (Phitos &
Kamari) Constantin.
Caryophyllaceae
FHe 1:193 RDB :96
L/297 L/1871
Bunium tenerum Hausskn.
Apiaceae
FGC 1:672 MFG 1:672
Bupleurum aira Snogerup
Apiaceae
FE 2:347 RDB :98
L/152 L/869 L/2098
Bupleurum capillare Boiss. & Heldr.
Apiaceae
FE 2:347 FGC 1:691 RDB :100
L/138 L/2098
Bupleurum gaudianum Snogerup
Apiaceae
EFK :215 FCA :148 FCSI :85 FCSII : RDB :102
L/253 L/294 L/762 L/1111 L/2093 L/2098
259
260
261
262
263
264
265
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 193
Campanula aizoon Boiss. & Spruner subsp. aizoides (Zaffran) Fed.
Campanulaceae
Campanula aizoides Zaffran ex Greuter
EFK :295 FCA :42 MCL 1:123 MFG 2:373 RDB :110
L/160 L/178 L/255 L/518 L/519 L/607 L/1111 L/1963
Campanula aizoon Boiss. & Spruner subsp. aizoon
Campanulaceae
MCL 1:123 MFG 2:372 RDB :110
L/184
Campanula albanica Witasek subsp. sancta (Hayek) Podlech
Campanulaceae
Campanula rotundifolia L. subsp. sancta Hayek
FE 4:90 MCL 1:136 MFG 2:387
L/2200 L/2237
?
?
270
271
272
273
Campanula amorgina Rech.f.
Campanulaceae
FAe :600 FE 4:84 MCL 1:129
L/169 L/196 L/2237
Campanula aizoon Boiss. & Spruner
Campanulaceae
EFK :295 FCA :42 MCL 1:123 MFG 2:372 RDB :110
L/160 L/184 L/255 L/512 L/518 L/607 L/1963
269
Calamintha hirta (Briq.) Hayek
Lamiaceae
Calamintha sylvatica Bromf. subsp. ascendens (Jordan) P.W.Ball
Satureja calamintha (L.) Scheele var. hirta Briq.
Satureja sancta Greuter & Burdet
FE 3:166 MCL 3:327 MFG 2:131
L/940 L/1941
268
Bupleurum greuteri Snogerup
Apiaceae
Bupleurum glumaceum Greuter & Rechinger 1967
RDB :104
L/119 L/305 L/1111 L/1878 L/1963 L/2098
Taxon
Bupleurum kakiskalae Greuter
Apiaceae
EFK :215 FCA :148 FCSI :85 RDB :106
L/138 L/255 L/337 L/1111 L/1237 L/1583
?
Taxon
Qualifiers
267
266
Taxon
Number
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EAe
Distribution
Qualifier
194 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Campanula anchusiflora Sm.
Campanulaceae
Campanula rupestris subsp. anchusiflora (Sm.) Hayek
FAe :599 FE 4:90 FGC 2:254 MCL 1:139
L/607 L/925 L/1963
Campanula andrewsii A.DC.
Campanulaceae
Campanula laciniata Andr., non L.
Campanula rupestris Sibth. & Sm. subsp. eu-rupestris Hayek var.
andrewsii (DC.) Hayek
Campanula tomentosa sensu auct. non Vent., Boiss. Fl. Or., p.p.
Campanula tomentosa var. bracteosa Heldr., p.p.
FAe :598 FE 4:81 FGC 2:255 MCL 1:139
L/512 L/607 L/925 L/1264 L/1963 L/1973
Campanula andrewsii A.DC. subsp. andrewsii
Campanulaceae
FE 4:81 MCL 1:139
L/607 L/925 L/1264 L/1963
Campanula andrewsii A.DC. subsp. hirsutula Phitos
Campanulaceae
FE 4:81 MCL 1:140
L/607 L/925 L/1264 L/1963
Campanula asperuloides (Boiss. & Orph.) Engler
Campanulaceae
Campanula asperuloides (Boiss. & Orph.) Engler subsp. taygetea
(Quézel & Contandr.) Greuter & Burdet
Diosphaera asperuloides (Boiss. & Orph.) Buser
Trachelium asperuloides Boiss. & Orph.
FE 4:95 MCL 1:124 RDB :112
L/138 L/512 L/607 L/709 L/980 L/1963 L/2237
Campanula calaminthifolia Lam.
Campanulaceae
Campanula calaminthifolia Lam. var. olivieri (A.DC.) Halácsy
Campanula olivieri A.DC.
FAe :600 FE 4:84 FT 6:34 MCL 1:129
L/111 L/539 L/540 L/869
274
275
276
277
278
279
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 195
Campanula celsii A.DC.
Campanulaceae
Campanula eriantha Hampe
Campanula lyrata auct. non Lam
Campanula rupestris subsp. celsii (A.DC.) Hayek
Campanula eriantha Vent.
Campanula eriantha Vent. var. typica Heldr.
FE 4:81 FGC 2:254 MCL 1:140
L/323b L/516 L/530 L/925 L/1264 L/1713 L/1871 L/1963
Campanula celsii A.DC. subsp. carystea Phitos
Campanulaceae
FE 4:81 MCL 1:140
L/323b L/925 L/1871 L/1963
Campanula celsii A.DC. subsp. celsii
Campanulaceae
FE 4:81 MCL 1:40
L/516 L/530 L/925 L/1713 L/1963
Campanula celsii A.DC. subsp. parnesia Phitos
Campanulaceae
Campanula anchusiflora auct. non Sibth. & Sm.
FE 4:81 MCL 1:140
L/925 L/1963
Campanula celsii A.DC. subsp. spathulifolia (Turrill) Phitos
Campanulaceae
Campanula rupestris Sm. var. gracilis Heldr.
Campanula rupestris Sm. var. spathulifolia Turrill.
FE 4:81 MCL 1:140
L/530 L/785 L/925 L/1264 L/1963
Campanula columnaris Contandr.
Campanulaceae
MCL 1:123 MFG 2:373 RDB :114
L/184 L/587
281
282
283
284
285
286
Taxon
Campanula carpatha Halácsy
Campanulaceae
Campanula tubulosa Lam. var. oreophila Phitos
Campanula saxatilis auct. non L. Boiss. Fl. Or. Suppl.
Campanula tubulosa auct. non Lam. Karp.
Campanula tubulosa Lam. var. carpatha (Halácsy) Hayek
EFK :296 FAe :597 FCA :42 MCL 1:134
L/119 L/273 L/925 L/1049 L/1111
Taxon
Qualifiers
280
Taxon
Number
+
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+
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EAe
Distribution
Qualifier
196 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Campanula constantini Beauverd & Topali
Campanulaceae
Campanula rupestris sensu auct. Euboea, Rechinger 1960 p.p.
FAe :598 FE 4:82 MCL 1:140
L/323b L/608 L/926 L/927 L/1871
Campanula creutzburgii Greuter
Campanulaceae
Campanula drabifolia subsp. creutzburgii (Greuter) Fed.
EFK :295 FCA :42 FCSI :15 FE 4:88 MCL 1:126
L/119 L/255 L/518 L/1008 L/1111
Campanula cymaea Phitos
Campanulaceae
Campanula celsii sensu auct. Euboea, Rechinger 1960 p.p.
Campanula tomentosa Vent., non Lam. var. calycina Heldr.
FE 4:82 MCL 1:140 RDB :116
L/323b L/926 L/1871
Campanula drabifolia Sm.
Campanulaceae
Campanula attica Boiss. & Heldr.
Campanula drabifolia Sm. var. major Boiss.
FAe :601 FCA :42 FCY 2:1049 FE 4:88 FT 6:53 MCL 1:126
L/119 L/150 L/192 L/516 L/530 L/696 L/796 L/856 L/986 L/1111 L/1846 L/1864 L/1878 L/1981
Campanula euboica Phitos
Campanulaceae
Campanula rupestris sensu auct. Euboea, Rechinger 1960 p.p.
FE 4:81 MCL 1:140
L/323b L/925 L/1871
Campanula garganica Ten. subsp. acarnanica (Damboldt) Damboldt
Campanulaceae
Campanula acarnanica Damboldt
FE 4:88 MCL 1:127
L/978 L/1123 L/1846
Campanula garganica Ten. subsp. cephallenica (Feer) Hayek
Campanulaceae
Campanula cephallenica Feer
FE 4:88 MCL 1:127
L/946 L/1123 L/1244 L/1800
287
288
289
290
291
292
293
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 197
Campanula heterophylla L.
Campanulaceae
FAe :600 FE 4:85 MCL 1:129
L/169 L/206 L/776 L/869
Campanula hierapetrae Rech.f.
Campanulaceae
FAe :600 FCA :42 FE 4:84 EFK :295 MCL 1:129 RDB :120
L/554 L/1111
Campanula incurva Aucher ex A.DC.
Campanulaceae
Campanula leutweinii Heldr.
FE 4:84 FGC 2:256 FT 6:14 MCL 1:129 RDB :122
L/539 L/540 L/861 L/928 L/1871
Campanula jacquinii (Sieber) A.DC. subsp. jacquinii
Campanulaceae
Trachelium jacquinii (Sieber) Boiss. subsp. jacquinii
Diosphaera jacquinii (Sieber) Buser
Phyteuma jacquinii Sieber
Trachelium jacquinii (Sieber) Boiss. p.p.
EFK :294 FAe :604 FCA :42 FCSI :16 FE 4:94 FT 11:175 MCL 1:129 MFG 2:389
L/300 L/518 L/1963
Campanula laciniata L.
Campanulaceae
Campanula erucifolia Feer
EFK :295 FAe :598 FCA :43 FCSII : FE 4:82 MCL 1:130 RDB :124
L/196 L/206 L/255 L/707 L/925 L/1111 L/1111 L/1583
Campanula lavrensis (Tocl & Rohlena) Phitos
Campanulaceae
Campanula andrewsii A.DC. subsp. breueri Tocl & Rohlena
Campanula andrewsii A.DC. var. lavrensis Tocl & Rohlena
Campanula lavrensis A.DC. var. breueri (Tocl & Rohlena) Phitos
Campanula rupestris sensu auct. Athos, Fl. Aeg. p.p.
Campanula rupestris Sm. var. andrewsii (A.DC.) Hayek f. breueri
(Tocl & Rohlena) Hayek
FAe :598 FE 4:81 MCL 1:140
L/447 L/925 L/1832d L/1972 L/1973
295
296
297
298
299
300
Taxon
Campanula goulimyi Turrill
Campanulaceae
FE 4:81 MCL 1:140
L/323b L/785 L/925 L/1871
Taxon
Qualifiers
294
Taxon
Number
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StE
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WAe
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IoI
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SPi
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NPi
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EAe
Distribution
Qualifier
198 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Campanula merxmuelleri Phitos
Campanulaceae
FE 4:82 MCL 1:140 RDB :126
L/915 L/925 L/1896
Campanula nisyria Papatsou & Phitos
Campanulaceae
FT 6:13 MCL 1:131
L/191
Campanula oreadum Boiss. & Heldr.
Campanulaceae
MCL 1:133 MFG 2:378
L/485 L/695 L/1246
Campanula pangaea Hartvig
Campanulaceae
L/1472
Campanula papillosa Halácsy
Campanulaceae
MCL 1:133 MFG 2:379
L/512 L/607 L/1963
Campanula pelviformis Lam.
Campanulaceae
Campanula corymbosa Desf.
Campanula pelviformis Lam. var. micrantha DC.
EFK :296 FAe :597 FCA :43 FCSI :16 FE 4:82 MCL 1:134
L/518 L/554 L/707 L/856 L/925 L/1111
Campanula pinatzii Greuter & Phitos
Campanulaceae
Campanula drabifolia subsp. pinatzii (Greuter & Phitos) Fed.
EFK :294 FCA :43 FCSI :16 FE 4:88 MCL 1:126
L/119 L/201 L/1008 L/1111
Campanula pindicola Aldén
Campanulaceae
MCL 1:138 MFG 2:386
L/477
302
303
304
305
306
307
308
309
?
Campanula lyrata Lam. subsp. icarica Runemark ex Phitos
Campanulaceae
FT 6:12 MCL 1:131
L/150 L/197 L/539 L/540 L/796
301
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 199
Campanula rechingeri Phitos
Campanulaceae
MCL 1:140
L/925 L/928 L/1189
Campanula reiseri Halácsy
Campanulaceae
Campanula reiseri var. leonis Halácsy
Campanula reiseri var. pelagia Phitos
FAe :598 MCL 1:140 RDB :128
L/423 L/443 L/550 L/925 L/928 L/2181
Campanula rhodensis A.DC.
Campanulaceae
Campanula drabifolia Sibth. & Sm.
FAe :601 FT 6:53 MCL 1:126
L/119 L/197 L/581 L/1111
Campanula rupestris Sm.
Campanulaceae
Campanula tomentosa Vent. subsp. eu-rupestris Hayek
Campanula tomentosa Vent. var. brachyantha Boiss.
FAe :598 FE 4:82 FGC 2:255 FGS 2:170 MCL 1:140 MFG 2:377 RDB :130
L/99 L/323b L/512 L/530 L/861 L/926 L/927 L/1259a L/1844 L/1973 L/2013
Campanula rupicola Boiss. & Spruner
Campanulaceae
FE 4:85 MCL 1:140 MFG 2:380
L/512 L/587
Campanula sartorii Boiss. & Heldr.
Campanulaceae
FE 4:87 MCL 1:140 RDB :132
L/111 L/169 L/978 L/2181
311
312
313
314
315
316
Taxon
Campanula radicosa Bory & Chaub.
Campanulaceae
FE 4:86 MCL 1:135 MFG 2:381
L/512 L/607 L/1963 L/2200
Taxon
Qualifiers
310
Taxon
Number
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StE
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IoI
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WAe
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SPi
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NPi
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NAe
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Kik
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EAe
Distribution
Qualifier
200 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Campanula saxatilis L.
Campanulaceae
Campanula mollis auct. non L.
Campanula saxatilis L. var. simonelli Bald.
EFK :295 FAe :598 FCA :43 FCSI :16 FE 4:82 MCL 1:141
L/119 L/518 L/554 L/707 L/765 L/925 L/1963
Campanula saxatilis L. subsp. cytherea Rech.f. & Phitos
Campanulaceae
Campanula carpatha auct. cyth. non Halácsy
Campanula saxatilis L. var. cythaerensis Beauverd & Top., nom. nud.
Campanula tubulosa auct. cyth. non Lam.
FAe :597 FE 4:82 MCL 1:141 RDB :134
L/119 L/765 L/925 L/1111 L/1917 L/1963
Campanula saxatilis L. subsp. saxatilis
Campanulaceae
EFK :295 FAe :598 FCA :43 FCSI :16 FE 4:82 MCL 1:141
RDB :134
L/518 L/554 L/707 L/925 L/1111
Campanula sciathia Phitos
Campanulaceae
Campanula andrewsii var. pelia Halácsy
Campanula pelia Hausskn.
FE 4:83 MCL 1:140
L/122 L/926 L/928
Campanula scopelia Phitos
Campanulaceae
Campanula celsii sensu auct. Skop., Fl. Aeg. p.p.
Campanula reiseri sensu auct. Skop., Fl. Aeg. p.p.
FAe :598 FE 4:83 MCL 1:140
L/439 L/926 L/928
Campanula spatulata Sm. subsp. filicaulis (Halácsy) Phitos
Campanulaceae
Campanula pauciflora Desf.
Campanula sibthorpiana Halácsy var. filicaulis Halácsy
Campanula spatulata Sm. subsp. spruneriana sensu auct. Kret.,
Rechinger 1943 p.p.
Campanula spatulata subsp. sibthorpiana var. filicaulis (Halácsy)
Hayek
EFK :295 FAe :602 FCA :43 FCSI :16 FE 4:80 MCL 1:142
MFG 2:374
L/268 L/518 L/554 L/695 L/707 L/930 L/1111
317
318
319
320
321
322
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PATTERNS OF PLANT ENDEMISM IN GREECE 201
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Campanula thessala Maire
Campanulaceae
Campanula patenticalyx Bocquet?
FE 4:83 FGC 2:170 MCL 1:140 MFG 2:377
L/926 L/927 L/1261b
Campanula topaliana Beauverd
Campanulaceae
FE 4:81 MCL 1:140
L/530 L/607 L/925 L/1199 L/1963 L/1973
Campanula topaliana Beauverd subsp. cordifolia Phitos
Campanulaceae
Campanula calavrytana Beauverd & Top.
Campanula rupestris Sibth. & Sm. subsp. eu-rupestris Hayek var.
virescens (Halácsy) Hayek
Campanula rupestris Sibth. & Sm. var. virescens Halácsy
FE 4:81, MCL 1:140
L/607 L/925 L/926 L/1199
Campanula topaliana Beauverd subsp. delphica Phitos
Campanulaceae
FE 4:81 MCL 1:140
L/925
Campanula topaliana Beauverd subsp. topaliana
Campanulaceae
FE 4:81, MCL 1:140
L/530 L/607 L/925 L/1963
Campanula tubulosa Lam.
Campanulaceae
Campanula subidaea Gand.
EFK :296 FAe :597 FCA :43 FCSI :16 FE 4:82 MCL 1:134
L/518 L/554 L/707 L/925 L/1111
324
325
326
327
328
329
Taxon
Campanula stenosiphon Boiss. & Heldr.
Campanulaceae
Campanula glomerata L. var. stenosiphon Boiss.
FE 4:85 FGC 2:260 MCL 1:142
L/159 L/607 L/1963
Taxon
Qualifiers
323
Taxon
Number
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StE
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IoI
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WAe
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SPi
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NPi
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EC
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NC
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
202 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Carex cretica Gradst. & Kern
Cyperaceae
EFK :418 FCA :160 FE 5:308 FT 9:130
L/255 L/257 L/539 L/540 L/1111
Carlina barnebiana B.L.Burtt & P.H.Davis
Asteraceae
EFK :320 FCA :60 FCSI :27 FE 4:210
L/273 L/1111 L/2093
Carlina corymbosa L. subsp. curetum (Heldr. ex Halácsy) Rech.f.
Asteraceae
Carlina curetum Heldr. ex Halácsy
Carlina curetum Heldr. ex Halácsy subsp. curetum
EFK :320 FAe :644 FCA :60 FCSI :27 FE 4:209
L/273 L/518 L/554 L/1111 L/1766
Carlina diae (Rech.f.) Meusel & Kastner
Asteraceae
Lyrolepis diae Rech f.
Lyrolepis piae Nord.
EFK :319 FCA :60 FE 4:209 RDB :136
L/138 L/255 L/268 L/291 L/537 L/554 L/1111 L/1583
Carlina sitiensis Rech.f.
Asteraceae
EFK :319 FCA :60 FCSI :28 FE 4:209
L/201 L/518 L/551 L/554 L/1111 L/2093
Carthamus leucocaulos Sm.
Asteraceae
Carthamus leucocaulos Sm. var. arachnoideus Davis
Kentrophyllum leucocaulon DC.
EFK :330 FAe :670 FCA :61 FCSI :28 FE 4:302 FT 5:594
L/128 L/150 L/197 L/273 L/346 L/447 L/539 L/540
L/554 L/762 L/776 L/783 L/913 L/1111 L/1871 L/2093 L/2181
332
333
334
335
336
337
Cardamine amara L. subsp. barbaraeoides (Halácsy) Maire & Petitm.
Brassicaceae
Cardamine raphanifolia Pourret subsp. barbareoides (Halácsy) Strid
Cardamine barbaraeoides Halácsy
AFE 10:157 FE 1:349 FHe 2:179 MCL 3:80 MFG 1:257
L/121 L/513 L/2200
Carduus euboicus Franco
Asteraceae
FE 4:225
L/456d
!
331
330
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 203
Centaurea acarnanica (Matthäs) Greuter
Asteraceae
Centaurea subciliaris Boiss. & Heldr. subsp. acarnanica Matthäs
L/1846 L/1916
Catapodium borgesii H.Scholz
Poaceae
FCSII :
L/1111 L/1693
343
344
Carum heldreichii Boiss.
Apiaceae
FE 2:354 MFG 1:700
L/121 L/160 L/178 L/512 L/587 L/2200
342
?
Carum depressum Hartvig & Kit Tan
Apiaceae
L/1963
341
Carum appuanum (Viv.) Grande subsp. palmatum (Hartvig) Bechi &
Garbari
Apiaceae
Carum rigidulum (Viv.) W.D.J.Koch ex DC. subsp. palmatum Hartvig
MFG 1:699
L/178 L/1107 L/2237
340
!
Carthamus tenuis (Boiss. & Bl.) Bornm. subsp. gracillimus (Rech.f.)
Hanelt
Asteraceae
Carthamus gracilimus Rech.f.
FT 5:594
L/197 L/1111
339
Taxon
Carthamus rechingeri P.H.Davis
Asteraceae
EFK :330 FCA :61 FE 4:303
L/1111
Taxon
Qualifiers
338
Taxon
Number
.
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Pe
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StE
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WAe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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+
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KK
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EAe
Distribution
Qualifier
204 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea achaia Boiss. & Heldr.
Asteraceae
Centaurea achaia Boiss. & Heldr. var. ferox Boiss. & Heldr.
Centaurea achaia Boiss. & Heldr. var. intermedia Boiss. & Heldr.
Centaurea achaia Boiss. & Heldr. var. submutica Boiss. & Heldr.
Centaurea sibthorpii (Halácsy) Boiss. & Heldr. p.p.
Colymbada sibthorpii (Halácsy) Holub
FE 4:267
L/284 L/530 L/607 L/610 L/736 L/1058 L/1215 L/1963
Centaurea acicularis Sm. var. acicularis
Asteraceae
Colymbada acicularis (Sm.) Holub
FAe :664 FT 5:548
L/1058
Centaurea aetolica Phitos & T.Georgiadis
Asteraceae
RDB :138
L/284 L/610 L/798 L/1058 L/1259a L/1833
Centaurea affinis Friv. subsp. laconiae Prodan
Asteraceae
Centaurea affinis Friv. subsp. peloponnesiaca (Halácsy) Dostál
Centaurea affinis Friv. var. peloponnesiaca Halácsy
Centaurea dissecta Boiss. subsp. affinis var. parlatoris f. peloponnesiaca
(Halácsy) Dostal
Centaurea dissecta Boiss. var. peloponnesiaca
FE 4:272 FGS 2:164 MFG 2:497
L/310a L/470 L/530 L/607 L/888 L/1963
Centaurea affinis Friv. subsp. pallidior (Halácsy) Hayek
Asteraceae
Acosta pallidior (Halácsy) Holub
Centaurea affinis Friv. var. pallidior (Halácsy) Halácsy
Centaurea grisebachii (Nyman) Heldr. subsp. paucijuga (Halácsy) Dostál
Centaurea pallidior Halácsy
Centaurea pallidior Halácsy var. pallidissima Halácsy
Centaurea paucijuga Halácsy
FE 4:273 MFG 2:498
L/607 L/888 L/1107 L/1211 L/1467 L/2013 L/2201
Centaurea albanica Halácsy
Asteraceae
Centaurea alba L. subsp. albanica (Halácsy) Dostál
Centaurea deusta Bald.
Centaurea ustulata Hal.
FE 4:288 FGC 2:134
L/1242
345
346
347
348
349
350
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 205
Centaurea argentea L. subsp. argentea
Asteraceae
FCSII :
L/765 L/1111 L/1690 L/1963 L/2237
Centaurea argentea L. subsp. chionantha (Turland & L.Chilton)
Greuter
Asteraceae
Centaurea argentea L. var. chionantha Turland & L.Chilton
Centaurea argentea L. subsp. macrothysana (Rech.f.) Turland &
L.Chilton var. chionantha Turland & L.Chilton
FCSII :
L/1690 L/1916 L/2237
Centaurea argentea L. subsp. macrothysana (Rech.f.) Turland &
L.Chilton
Asteraceae
Centaurea argentea L. var. macrothysana Rech.f.
FCSII :
L/1111 L/1690 L/1916 L/2237
Centaurea athoa DC. subsp. parnonia E.Gamal-Eldin & Wagenitz
Asteraceae
Centaurea rupestris L. subsp. parnonia (Halácsy) Routsi & T.Georgiadis
Centaurea finazzeri sensu auct. from Taygetos.
Centaurea macedonica Boiss. subsp. parnonia (Halácsy) Dostál
Centaurea parnonia Halácsy
Centaurea rupestris L. var. minor Boiss.
Colymbada parnonia (Halácsy) Holub
FE 4:264 MFG 2:492
L/284 L/470 L/607 L/1058 L/1203 L/1588 L/1916 L/1963
Centaurea atropurpurea Olivier
Asteraceae
Centaurea oliveriana DC.
Centaurea oliveriana DC. var. amorgina Boiss.
Colymbada oliveriana (DC.) Holub
Hyalea oliveriana Jaub. & Spach
FAe :663 FE 4:266 FGC 2:152 MFG 2:493
L/111 L/145 L/196 L/206 L/284 L/776 L/869 L/1058
352
353
354
355
356
Taxon
Centaurea argentea L.
Asteraceae
Centaurea argentea L. var. brachythysana Rech.f.
Centaurea argentea L. var. macrothysana Rech.f.
Centaurea cineraria Sieb.
Centaurea raquisina Sibth. & Sm.
EFK :329 FAe :666 FCA :61 FCSII : FE 4:271
L/518 L/554 L/707 L/888 L/1052 L/ 1111 L/1690 L/1916 L/1963 L/2237
Taxon
Qualifiers
351
Taxon
Number
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EAe
Distribution
Qualifier
206 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea attica Nyman
Asteraceae
FE 4:275 FGC 2:144 FGS 1:54
L/323b L/427 L/530 L/888 L/1980
Centaurea attica Nyman subsp. attica
Asteraceae
Centaurea boissieri Walp., non DC.
Centaurea graeca sensu auct., Boiss. & Spruner; Fl. Aeg., non Griseb.
Centaurea paniculata sensu auct., Sibth. & Sm. Fl. Graec. Prodr. subsp.
auct. sensu Sibth. & Sm. Fl. Graec. Prodr.
Centaurea parlatoris Heldr var. boissieri Heldr. & Sart.
FAe :663 FAe :667 FE 4:275 FGC 2:144
L/323b L/888 L/1871
Centaurea attica Nyman subsp. megarensis (Halácsy & Hayek)
Dostál
Asteraceae
Centaurea megarensis Halácsy & Hayek
FE 4:275 FGS 1:54
L/138 L/530 L/781 L/888
Centaurea attica Nyman subsp. pateraea (Halácsy) T.Georgiadis
Asteraceae
Centaurea attica Nyman var. pateraea Halácsy
FGC 2:144 FGS 1:54
L/427 L/530 L/888
Centaurea attica Nyman subsp. pentelica (Hausskn.) Dostál
Asteraceae
Centaurea pentelica Hausskn.
FE 4:275 FGC 2:143
L/323b L/888 L/1871
Centaurea brunnea Halácsy
Asteraceae
Centaurea alba Halácsy subsp. brunnea (Halácsy) Dostál
Centaurea deusta Ten. var. brunnea Halácsy
FE 4:288 FGC 2:135
L/1242
Centaurea carystea Trigas & Constantin.
Asteraceae
L/2226
357
358
359
360
361
362
363
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 207
Centaurea chrysocephala Phitos & T.Georgiadis
Asteraceae
MFG 2:505 RDB :146
L/563 L/1017 L/1259a
Centaurea cithaeronea Phitos & Constantin.
Asteraceae
RDB :148
L/314 L/529 L/530
Centaurea corinthiaca Boiss. & Heldr.
Asteraceae
Centaurea achaia Boiss. & Heldr. subsp. corinthiaca (Boiss. & Heldr.)
Phitos & Georgiadis
Centaurea achaia Boiss. & Heldr. var. corinthiaca (Boiss. & Heldr.)
Boiss.
FE 4:267
L/284 L/530 L/610 L/1058 L/1203
Centaurea ebenoides Heldr. ex S.Moore
Asteraceae
Colymbada ebenoides (Heldr.) Holub
FE 4:267
L/99 L/99 L/284 L/323b L/608 L/1058 L/1833 L/1871
Centaurea euboica Rech.f.
Asteraceae
Centaurea sibthorpii Halácsy p.p.
FE 4:267
L/99 L/284 L/323b L/607 L/608 L/610 L/1058 L/1833 L/1871
Centaurea euboica Rech.f. subsp. euboica
Asteraceae
L/99 L/284 L/323b L/608 L/610 L/1058 L/1871
366
367
368
369
370
371
Centaurea chalcidicaea Hayek
Asteraceae
Acosta chalcidicaea (Hayek) Holub
Centaurea affinis Friv. subsp. chalcidicaea (Hayek) T.Georgiadis
Centaurea sancta-annae J.Wagner
FAe :667 FE 4:274 MFG 2:499
L/888 L/1972 L/1973 L/1975
Taxon
Centaurea charrelii Halácsy & Dörfler
Asteraceae
FE 4:270 RDB :144
L/995
Taxon
Qualifiers
365
364
Taxon
Number
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StE
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WAe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
208 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea euboica Rech.f. subsp. intermedia Phitos & T.Georgiadis
Asteraceae
L/284 L/323b L/608 L/610 L/1058 L/1871
Centaurea finazzeri Adamović subsp. kozanii (Routsi &
T.Georgiadis) Greuter
Asteraceae
Centaurea rupestris L. subsp. kozanii Routsi & T.Georgiadis
L/284 L/891 L/1058 L/1916 L/2201
Centaurea greuteri E.Gamal-Eldin & Wagenitz
Asteraceae
Centaurea grisebachii (Nyman) Heldr. subsp. confusa (Halácsy)
Dostál
Asteraceae
Centaurea affinis Friv. var. microcaphala Halácsy
Centaurea confusa Halácsy
Centaurea grisebachii (Nyman) Formánek var. confusa (Halácsy) Hayek
Centaurea macedonica Formánek
FE 4:274 MFG 2:500
L/470 L/543 L/888
Centaurea grisebachii (Nyman) Heldr. subsp. occidentalis
E.Gamal-Eldin & Wagenitz
Asteraceae
MFG 2:500
L/888 L/2201
Centaurea heldreichii Halácsy
Asteraceae
Centaurea alba L. subsp. heldreichii (Halácsy) Dostál
FE 4:289 FGC 2:133 RDB :150
L/138 L/1202
Centaurea huljakii Wagner
Asteraceae
FAe :668 FE 4:289 MFG 2:499
L/447 L/1204 L/1832d L/1973
Centaurea iberica Trev. ex Sprengel subsp. holzmanniana (Boiss.)
Dostál
Asteraceae
Centaurea iberica Trev. ex Sprengel var. holzmanniana Boiss.
FE 4:282 FGC 2:163
372
373
374
375
376
377
378
379
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 209
Centaurea incompleta Halácsy
Asteraceae
FE 5:264 RDB :152
L/775 L/1202 L/1246
Centaurea johnseniana Kit Tan & Strid
Asteraceae
L/1894
Centaurea kalambakensis Freyn & Sint.
Asteraceae
FE 4:274 RDB :154
L/138 L/888
Centaurea lacerata (Hausskn.) Halácsy
Asteraceae
Centaurea affinis Friv. subsp. lacerata (Hausskn.) Maire & Petitm.
FE 4:272 MFG 2:498
L/888 L/1846
Centaurea laconica Boiss.
Asteraceae
FAe :663 FE 4:267
L/284 L/607 L/776 L/930 L/1058 L/1963 L/2181
Centaurea laconica Boiss. subsp. laconica
Asteraceae
Centaurea eryngioides Lam. var. subarachnoidea Boiss & Heldr.
Centaurea subarachnoidea Boiss & Heldr.
Colymbada laconica (Boiss.) Holub
FE 4:267
L/284 L/607 L/1058 L/1963
381
382
383
384
385
386
Taxon
Centaurea idaea Boiss. & Heldr.
Asteraceae
Centaurea solstialis subsp. idaea (Boiss. & Heldr.) Gugler
Centaurea solstialis var. idaea (Boiss. & Heldr.) Bald.
EFK :327 FAe :668 FCA :61 FCSI :29 FE 4:284 MFG 2:503
L/518 L/522 L/554 L/1111 L/1766
Taxon
Qualifiers
380
Taxon
Number
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WAe
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IoI
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+
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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EAe
Distribution
Qualifier
210 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea laconica Boiss. subsp. lineariloba (Halácsy) Wagenitz &
E.Gamal-Eldin
Asteraceae
Centaurea guicciardi Boiss. var. lineariloba Halácsy & Dörfler
Centaurea spruneri Boiss. subsp. guicciardi (Boiss.) Hayek var.
lineariloba (Halácsy & Dörfler ) Hayek
Centaurea spruneri Boiss. & Heldr. subsp. lineariloba (Halácsy &
Dörfler) Dostál
Centaurea spruneri Boiss. & Heldr. var. lineariloba (Halácsy & Dörfler)
Hay.
FAe :663 FE 4:267 FGC 2:156
L/284 L/554 L/776 L/1058 L/1203 L/2181
Centaurea lactiflora Halácsy
Asteraceae
Centaurea ovina Formánek
FE 4:281 FGC 2:150
L/888
Centaurea lactucifolia Boiss.
Asteraceae
Centaurea chalkensis Fors.-Major & Barbey
Centaurea lactucifolia Boiss. subsp. chalkensis (Fors.-Major & Barbey)
Rech.f.
FT 5:549 RDB :156
L/197 L/284 L/288 L/1058 L/1111
Centaurea lancifolia Sieber ex Sprengel
Asteraceae
Aberboa lancifolia (Sprengel) DC.
Chartolepis lancifolia (Sprengel) Fenzl
Wagenitzia lancifolia (Sieber ex Sprengel) Dostál
EFK :326 FAe :669 FCA :61 FCSI :29 FE 4:301 MFG 2:523 RDB :158
L/518 L/519 L/1111 L/1583 L/1973
Centaurea laureotica Heldr. ex Halácsy
Asteraceae
Centaurea asperula Halácsy
Centaurea attica Nyman subsp. asperula (Halácsy) Dostal
Centaurea laureotica Heldr. ex Halácsy
FE 4:275 FE 4:281 FGC 2:144 FGS 2:54
L/516 L/888
387
388
389
390
391
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 211
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea ossaea Halácsy
Asteraceae
Centaurea attica Nyman subsp. ossaea (Halácsy) Dosral
Centaurea dissecta var. alpina Heldr., pl. exs. A 1884, non Heldr in
Boiss. Fl. Or.
FE 4:275 MFG 2:502
L/888
401
Centaurea nervosa Willd. subsp. promota E.Gamal-Eldin & Wagenitz
Asteraceae
MFG 2:513
L/2013
398
Centaurea orphanidea Heldr. & Sart. ex Boiss.
Asteraceae
FAe :667 FE 4:282 FGC 2:148 FGS 2:164
L/99 L/323b L/530 L/888
Centaurea musarum Boiss. & Orph.
Asteraceae
FE 4:290 MFG 2:509 RDB :160
L/529 L/1694
397
400
Centaurea musakii T.Georgiadis
Asteraceae
L/888
396
Centaurea niederi Heldr.
Asteraceae
FE 4:270 RDB :162
L/138 L/339 L/607 L/888 L/1016 L/1202 L/1963
Centaurea messenicolasiana T.Georgiadis, G.Dimitrellos & Routsi
Asteraceae
L/563 L/2251
395
399
Centaurea mantoudii T.Georgiadis
Asteraceae
L/608 L/888 L/1871 L/1960
394
!
Centaurea litochorea T.Georgiadis & Phitos
Asteraceae
MFG 2:503
L/1217 L/1246
393
Taxon
Centaurea leonidia Kalpouz. & Constantin.
Asteraceae
L/1877
Taxon
Qualifiers
392
Taxon
Number
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+
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StE
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WAe
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IoI
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+
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+
+
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SPi
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+
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NPi
+
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EC
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NC
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Kik
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EAe
Distribution
Qualifier
212 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea pangaea Greuter & Papan.
Asteraceae
MFG 2:514
L/907 L/1107
Centaurea pannosa DC.
Asteraceae
FE 4:271
L/888 L/1972
Centaurea paxorum Phitos & T.Georgiadis
Asteraceae
L/1017
Centaurea pelia DC.
Asteraceae
Centaurea pelia DC. f. hamata (Hausskn.) Hayek
Centaurea pelia DC. var. refracta Hausskn.
Centaurea xanthina Boiss. & Heldr. non Spreng.
FE 4:281 FGC 2:149 FGS 1:60
L/499 L/512 L/530 L/888 L/1467 L/1832d L/2013
Centaurea peucedanifolia Boiss. & Orph.
Asteraceae
FE 4:279
L/888 L/1973
Centaurea poculatoris Greuter
Asteraceae
EFK :329 FCA :61 FCSI :29 FE 4:275 RDB :164
L/336 L/337 L/752 L/888 L/1111 L/1583
Centaurea prespana Rech.f.
Asteraceae
L/888 L/1112 L/1151 L/2205
Centaurea princeps Boiss. & Heldr.
Asteraceae
Centaurea alba L. subsp. princeps (Boiss. & Heldr.) Gugler
Centaurea alba var. princeps (Boiss. & Heldr.) Boiss.
FE 4:289 MFG 2:504 RDB :166
L/121 L/138
Centaurea pseudocadmea Wagenitz
Asteraceae
Centaurea deustiformia Adamović subsp. pseudocadmea (Wagenitz)
Dostál
FE 4:289 MFG 2:502
L/310a L/530 L/1120 L/1915
402
403
404
405
406
407
408
409
410
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 213
Centaurea ptarmicifolia Halácsy ex Hayek
Asteraceae
Centaurea deustiformia Adamović subsp. ptarmicifolia (Halácsy &
Hayek) Dostál
Centaurea epirotica (Bald.) Halácsy
FE 4:289 MFG 2:506
L/563
Centaurea raphanina Sm.
Asteraceae
Colymbada raphanina (Sm.) Holub
FAe :664 FAe :666 FCA :62 FE 4:268 FT 5:549 MCL 2:134 MFG 2:494
L/111 L/128 L/206 L/443 L/512 L/776 L/783 L/856 L/869 L/928 L/1049 L/1058 L/1111 L/1288 L/1627 L/1766 L/1833 L/1844 L/1963 L/1981 L/2181 L/2237
Centaurea raphanina Sm. subsp. mixta (DC.) Runemark
Asteraceae
Centaurea eriopoda Rech.f.
Centaurea halacsyi Dörfler
Centaurea hellenica Boiss. & Spruner
Centaurea hellenica Boiss. & Spruner f. intermedia Heldr.
Centaurea mixta DC.
Centaurea mixta Sm. subsp. eu-mixta Gugler
Centaurea mixta Sm. var. eriopoda Rech.f.
Centaurea mixta Sm. var. eu-mixta (Gugler) Rech.f.
Centaurea mixta Sm. var. halacsyi (Dörfler) Rech.f.
Centaurea mixta Sm. var. myconia (Boiss. & Sart.) Hayek
Centaurea mixta Sm. var. nigrotriangulata Rech.f.
Centaurea myconia Boiss. & Sart.
Centaurea nigrotriangulata Rech.f.
Colymbada mixta DC.
FAe :664 FE 4:269 FT 5:549 MCL 2:134 MFG 2:495
L/111 L/122 L/128 L/169 L/177 L/439 L/443 L/512
L/516 L/522 L/530 L/539 L/540 L/607 L/707 L/776 L/783 L/861 L/869 L/928 L/1058 L/1111
L/1288 L/1627 L/1844 L/1871 L/1878 L/1963 L/1981 L/2181
412
413
414
Taxon
Centaurea psilacantha Boiss. & Heldr.
Asteraceae
Colymbada psilacantha (Boiss. & Heldr.) Holub
FE 4:267 FGC 2:154
L/284 L/1058
Taxon
Qualifiers
411
Taxon
Number
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StE
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IoI
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SPi
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NPi
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NC
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NAe
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Kik
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EAe
Distribution
Qualifier
214 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea rechingeri Phitos
Asteraceae
Centaurea scyria Runemark
FE 4:266 FT 10:168
L/284 L/423 L/443 L/767 L/796 L/915 L/928 L/1058
Centaurea redempta Heldr.
Asteraceae
EFK :328 FAe :663 FCA :62 FCSI :29 FE 4:267
L/119 L/284 L/518 L/765 L/1058 L/1111 L/1878 L/1963
Centaurea redempta Heldr. subsp. cytherea (Rech.f.) Routsi &
T.Georgiadis
Asteraceae
Centaurea cytherea Rech.f.
FE 4:267
B/3060c L/119 L/284 L/765 L/1058 L/1111 L/1878 L/1963
Centaurea redempta Heldr. subsp. redempta
Asteraceae
EFK :328 FAe :663 FCA :62 FCSI :29 FE 4:267
L/284 L/518 L/1058 L/1111
Centaurea solstitialis L. subsp. erythracantha (Halácsy) Dostál
Asteraceae
Centaurea erythracantha Halácsy
FE 4:284 FGS 1:61
L/1246
417
418
419
420
421
Centaurea raphanina Sm. subsp. saxatilis (K.Koch) Greuter
Asteraceae
Phaeopappus saxatilis (K.Koch) Boiss.
Centaurea saxatilis (K.Koch) B.D.Jacks.
Centaurium saxatile K.Koch
MCL 2:134
L/1916 L/2237
416
?
Centaurea raphanina Sm. subsp. raphanina
Asteraceae
EFK :328 FAe :666 FCA :62 FCSI :29 FE 4:269 MCL 2:134 MFG 2:494
L/273 L/518 L/554 L/856 L/1049 L/1058 L/1111 L/1766 L/1833
415
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 215
?
424
Centaurea subsericans Halácsy
Asteraceae
Acosta subsericans (Halácsy) Holub
FE 4:275 FGS 2:54 MFG 2:501
L/530 L/781 L/1915
Centaurea thasia Hayek
Asteraceae
Centaurea ipsaria Stoj. & Kitanov
FAe :666 FE 4:272
L/554 L/888 L/1015 L/1267b L/1349 L/1916
Centaurea thessala Hausskn.
Asteraceae
FE 4:275 FE 4:282 FGC 2:148 FGC 2:149
L/888
Centaurea thessala Hausskn. subsp. drakiensis (Freyn. & Sint.)
T.Georgiadis
Asteraceae
Centaurea attica Nyman subsp. drakiensis (Freyn & Sint.) Dostál
Centaurea drakiensis Freyn & Sint.
FE 4:275 FGC 2:149
L/888
425
426
427
428
Centaurea spruneri Boiss. & Heldr. subsp. spruneri
Asteraceae
FE 4:267
L/284 L/1058 L/1259b L/1448
Centaurea spruneri Boiss. & Heldr. subsp. minoa Heldr. ex Boiss
Asteraceae
Centaurea guicciardi Boiss. subsp. minoa Heldr. ex Boiss.
Centaurea minoa Heldr. ex Boiss.
Centaurea spruneri Boiss. & Heldr. var. minoa (Boiss.) Hayek
Centaurea spruneri Boiss. & Heldr. var. minoa (Boiss.) Rech.f.
Colymbada guicciardii (Boiss.) Holub
Colymbada spruneri (Boiss. & Heldr.) Holub
EFK :328 FAe :663 FCA :62 FCSI :29 FE 4:267
L/284 L/554 L/1058
?
423
Taxon
Centaurea spinosa L. var. tragacanthoides (Rech.f.) Wagenitz
Asteraceae
Centaurea tragacanthoides Rech.f.
FAe :668 FT 5:500
L/796 L/888 L/2207
Taxon
Qualifiers
422
Taxon
Number
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StE
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Pe
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IoI
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WAe
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SPi
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NPi
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EC
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NC
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
216 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Centaurea tuntasia Heldr. ex Halácsy
Asteraceae
FE 4:264 FGC 2:159 RDB :168
L/1058 L/1203
Centaurea tymphaea Hausskn. subsp. brevispina (Hausskn.) Dostál
Asteraceae
Centaurea brevispina Hausskn.
Centaurea macedonica Heldr., non Griseb., non C. diffusa var.
brevispina Boiss.
Centaurea tymphaea Hausskn. var. brevispina (Hausskn.) Hayek
FE 4:282 FGC 2:148
L/888
Centaurea vermia Rech.f.
Asteraceae
FE 4:274
L/888
432
433
434
Centaurea vlachorum Hartvig
Asteraceae
MFG 2:510 RDB :170
L/995 L/1040 L/1165
Centaurea triamularia Aldén
Asteraceae
MFG 2:510
L/477
431
435
Centaurea transiens Halácsy
Asteraceae
Centaurea grisebachii (Nyman) Formánek subsp. transiens (Halácsy)
T.Georgiadis
FE 4:274 FGC 2:143
L/888 L/890b L/1246
430
?
Centaurea thessala Hausskn. subsp. thessala
Asteraceae
Centaurea orphanidea Heldr. & Sart. var. thessala Hausskn.
Centaurea thessala Hausskn.
FE 4:282 FGC 2:148
L/888
429
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PATTERNS OF PLANT ENDEMISM IN GREECE 217
Centaurium erythraea Rafn subsp. limoniiforme (Greuter) Greuter
Gentianaceae
Centaurium limoniiforme Greuter
FE 3:58 MCL 3:235
L/119 L/392 L/765 L/1052 L/1111
Centranthus junceus Boiss. & Heldr.
Caprifoliaceae
Centranthus longiflorus Steven subsp. junceus (Boiss. & Heldr.)
I.B.K.Richardson
Centranthus ruber (L.) DC. subsp. junceus (Boiss. & Heldr.) Maire &
Petitm.
FE 4:56 MFG 2:349
L/469 L/512 L/1973 L/2013 L/2200
Centranthus sieberi Heldr.
Caprifoliaceae
Centranthus nevadensis Boiss. subsp. sieberi (Heldr.) I.B.K.Richardson
EFK :291 FCA :155 FE 4:56 MFG 2:348
L/469 L/1111 L/1766
Cephalanthera cucullata Boiss. & Heldr.
Orchidaceae
Cephalanthera rubra sensu Teschner 1975, non (L.) Rich.?
Epipactis cucullata (Boiss. & Heldr. ex Rchb.f.) Wettstein
EFK :422 FAe :842 FCA :187 FCSI :111 FE 5:328 RDB :174
L/138 L/526 L/611 L/791 L/867 L/1098 L/1111 L/1395 L/1463 L/1766 L/2185
Cephalaria fanourii Perdetzoglou & Kit Tan
Caprifoliaceae
L/995 L/1801
Cephalaria flava (Sm.) Szabó subsp. setulifera (Boiss. & Heldr.)
Kokkini
Caprifoliaceae
Cephalaria setulifera Boiss. & Heldr.
FE 4:58 MCL 3:177 MFG 2:352
L/476 L/530 L/736 L/749 L/781 L/1871 L/1915 L/2013
437
438
439
440
441
442
Taxon
Centaurea xylobasis Rech.f.
Asteraceae
FT 5:513 RDB :172
L/707 L/796 L/975
Taxon
Qualifiers
436
Taxon
Number
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Pe
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StE
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WAe
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SPi
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IoI
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
218 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Cephalaria tenuiloba Strid
Caprifoliaceae
MCL 3:177 MFG 2:352 RDB :178
L/121 L/1181 L/2200 L/2201 L/2201
Cerastium brachypetalum Pers. subsp. atheniense (Lonsing) P.D.Sell
& Whitehead
Caryophyllaceae
Cerastium atheniense Lonsing
Cerastium brachypetalum sensu Halácsy non Pers. s. stricto
AFE 6:112 FE 1:173 FHe 1:210 MCL 1:176
L/323a L/530 L/1164 L/1871
Cerastium brachypetalum Pers. subsp. corcyrense (Lonsing) P.D.Sell
& Whitehead
Caryophyllaceae
Cerastium corcyrense Moschl
Cerastium litigiosum sensu Halácsy
AFE 6:110 FE 1:173 FGC 1:223 FHe 1:209 MCL 1:181 MFG 1:120
L/441 L/532 L/601 L/1164
Cerastium brachypetalum Pers. subsp. doerfleri (Halácsy ex Hayek)
P.D.Sell
Caryophyllaceae
Cerastium doerfleri Halácsy ex Hayek
AFE 6:110 EFK :81 FAe :146 FCA :45 FCSI :18 FE 1:173 FHe 1:209 MCL 1:182 MFG 1:121
L/518 L/1111 L/1164
444
445
446
447
?
Cephalaria squamiflora (Sieber) Greuter subsp. squamiflora
Caprifoliaceae
Cephalaria ?ambrosioides sensu Consp. Fl. Graec.
Cephalaria balearica sensu Barbey
Cephalaria centauroides var. cretacea sensu Raulin 1878
Cephalaria cretacea sensu Rchb. & Schultes 1827
Scabiosa centauroides sensu Sieber & auct. cret.
Scabiosa cretacea auct.
Scabiosa uralensis auct. var. cretacea sensu Rchb. 1826
EFK :291 FAe :590 FCA :86 FE 4:57 FGC 1:769 FT 4:547 MCL 3:178
L/145 L/255 L/273 L/423 L/539 L/540 L/767 L/1111 L/1247
443
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PATTERNS OF PLANT ENDEMISM IN GREECE 219
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Cerastium candidissimum Correns
Caryophyllaceae
Cerastium tomentosum auct. non L.
AFE 6:88 FE 1:167 FHe 1:201 MCL 1:176 MFG 1:112
L/160 L/512 L/530 L/587 L/607 L/1846 L/1871 L/1871 L/1963 L/2013 L/2200
Cerastium illyricum Ard.
Caryophyllaceae
Cerastium pilosum Sibth. & Sm. nom. illeg.
AFE 6:108 FE 1:172 FHe 1:205 MCL 1:180 MFG 1:116
L/122 L/159 L/447 L/607 L/1054 L/1244 L/1769 L/1800 L/1846 L/1963 L/2201
Cerastium illyricum Ard. subsp. brachiatum (Lonsing) Jalas
Caryophyllaceae
Cerastium brachiatum Lonsing
Cerastium brachiatum Lonsing subsp. decrescens Lonsing
Cerastium brachiatum subsp. prolixum Lonsing
Cerastium illyricum Ard. subsp. decrescens (Lonsing) P.D.Sell &
Whitehead
Cerastium illyricum Ard. subsp. prolixum (Lonsing) P.D.Sell &
Whitehead
Cerastium illyricum Ard. var. macropetalum Boiss.
Cerastium illyricum Bory non Ard.
Cerastium pedunculare sensu Hayek p.p., non Bory & Chaub.
AFE 6:108 FE 1:172 FHe 1:206 MCL 1:180 MFG 1:118
L/607 L/785 L/1054 L/1800 L/1963
Cerastium illyricum Ard. subsp. crinitum (Lonsing) P.D.Sell &
Whitehead
Caryophyllaceae
Cerastium crinitum Lonsing
AFE 6:109 FE 1:172 FHe 1:206 MCL 1:180
Cerastium illyricum Ard. subsp. Illyricum
Caryophyllaceae
Cerastium pedunculare sensu Halácsy [non Bory & Chaub.]
Cerastium pedunculare sensu Hayek p.p. [non Bory & Chaub.]
Cerastium pelligerum Bornm. & Hayek
Cerastium pilosum Sibth & Sm.
AFE 6:109 FE 1:172 FHe 1:205
L/607 L/1123 L/1244
449
450
451
452
453
Taxon
Cerastium brachypetalum Pers. subsp. pindigenum (Lonsing)
P.D.Sell & Whitehead
Caryophyllaceae
Cerastium litigiosum sensu Halácsy
Cerastium pindigenum Lonsing
AFE 6:110 FE 1:173 FHe 1:209 MCL 1:182 MFG 1:120
L/121 L/601 L/1164 L/2200
Taxon
Qualifiers
448
Taxon
Number
.
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+
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+
+
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WAe
.
+
+
StE
+
+
+
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Pe
+
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+
+
+
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IoI
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+
+
SPi
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+
+
NPi
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+
+
EC
.
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+
NC
.
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NE
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NAe
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Kik
.
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.
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KK
.
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.
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EAe
Distribution
Qualifier
220 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Cerastium moesiacum Friv. subsp. glutinosum (Halácsy) Strid
Caryophyllaceae
Cerastium alpinum L. var. glutinosum Halácsy
Cerastium moesiacum Friv. f. halacsyi Borza
Cerastium moesiacum Friv. f. thasium Hayek
FHe 1:202
L/1267a L/1349
Cerastium pedunculare Bory & Chaub.
Caryophyllaceae
Cerastium laxum Boiss. & Heldr.
AFE 6:109 FE 1:172 FHe 1:207 FT 2:85 MCL 1:182
L/159 L/607 L/1963
Cerastium runemarkii Möschl & Rech.f.
Caryophyllaceae
Cerastium coronense Runemark
AFE 6:100 FE 1:170 FHe 1:203 MCL 1:182
L/196 L/361 L/869 L/1871
Cerastium scaposum Boiss. & Heldr.
Caryophyllaceae
AFE 6:109 EFK :80 FAe :147 FCA :46 FCSI :18 FE 1:172 FHe 1:207 MCL 1:182 MFG 1:118
L/518 L/554 L/1111 L/1766 L/2094
Cerastium scaposum Boiss. & Heldr. subsp. deschatresii Greuter,
N.Böhling & R.L.Jahn
Caryophyllaceae
L/1111 L/2094
Cerastium scaposum Boiss. & Heldr. subsp. peninsularum Greuter,
N.Böhling & R.L.Jahn
Caryophyllaceae
L/1111 L/2094
Cerastium scaposum Boiss. & Heldr. subsp. scaposum
Caryophyllaceae
L/1111 L/2094
Cerastium smolikanum Hartvig
Caryophyllaceae
AFE 6:115 FE 1:174 FHe 1:208 MCL 1:182 MFG 1:118
L/1391
454
455
456
457
458
459
460
461
.
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 221
468
Chamaecytisus creticus (Boiss. & Heldr.) Rothm.
Fabaceae
Chamaecytisus subidaeus (Gand.) Rothm.
Cytisus creticus Boiss. & Heldr.
Cytisus subidaeus Gand.
EFK :144 FCA :104 FE 2:91 MCL 4:73
L/97 L/1111 L/1963
Chaerophyllum heldreichii Orph. ex Boiss.
Apiaceae
FE 2:325 MFG 1:667
L/395 L/607 L/1963 L/2013
Chaerophyllum euboeum Halácsy
Apiaceae
FAe :421
L/323a L/1871
466
!
Chaerophyllum creticum Boiss. & Heldr.
Apiaceae
FCA :149 FCSI :86 FE 2:325 RDB :182
L/1111
465
467
Chaenorhinum idaeum Rech.f.
Plantaginaceae
Chaenorhinum minus L. subsp. idaeum (Rech.f.) R.Fernandes
Microrrhinum idaeum (Rech.f.) Speta
EFK :276 FCA :142 FE 3:226
L/255 L/522 L/1010 L/1111
464
?
Cerastium vourinense Moschl & Rech.f.
Caryophyllaceae
AFE 6:87 FE 1:167 FHe 1:207 MCL 1:185 MFG 1:118
L/1391
463
Taxon
Cerastium theophrasti Merxm. & Strid
Caryophyllaceae
Cerastium alpinum auct. non L.
Cerastium alpinum var. glanduliferum sensu Stoj. & Stefanov
Cerastium uniflorum auct. non Clairv.
Cerastium uniflorum var. mitkaense Wagner
AFE 6:101 FE 1:170 FHe 1:204 MCL 1:174 MFG 1:115 RDB :180
L/404a L/485 L/1094
Taxon
Qualifiers
462
Taxon
Number
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WAe
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StE
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IoI
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SPi
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+
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NPi
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EC
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+
+
NC
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NE
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NAe
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Kik
+
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+
+
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KK
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EAe
Distribution
Qualifier
222 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Chondrilla ramosissima Sm.
Asteraceae
EFK :339 FAe :690 FCA :62 FE 4:343
L/516 L/530 L/543 L/607 L/765 L/1111 L/1696 L/1871 L/1963 L/2013
Cicer graecum Orph. ex Boiss.
Fabaceae
FE 2:129 MCL 4:78 MFG 1:481 RDB :184
L/395 L/607 L/1963
Cirsium creticum (Lam.) d’Urv subsp. dictaeum Greuter, Matthäs &
Risse
Asteraceae
EFK :322 FCA :63
L/253
Cirsium epiroticum Petr.
Asteraceae
FE 4:240 MFG 2:486
?
?
470
471
472
473
474
Cirsium hypopsilum Boiss. & Heldr.
Asteraceae
Cirsium cylleneum Halácsy
Cirsium lobelii Ten. var. hypopsilum (Boiss. & Heldr.) Halácsy
FE 2:235 MFG 2:484
L/160 L/512 L/607 L/1830 L/1871 L/1963
Cirsium mairei Halácsy
Asteraceae
FE 4:240 MFG 2:486
L/121 L/2013 L/2200 L/2201
475
476
Cirsium heldreichii Halácsy
Asteraceae
Cirsium decussatum Boiss.
Cirsium dolopicum Formánek
FE 4:235 MFG 2:483
L/121 L/323b L/1871 L/1963 L/2200
Chamaecytisus dorycnioides (Davidov) Frodin & Heywood
Fabaceae
Cytisus dorycnioides Davidoff
FE 2:93 MCL 4:74
469
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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*
PATTERNS OF PLANT ENDEMISM IN GREECE 223
Clematis elisabethae-carolae Greuter
Ranunculaceae
AFE 8:102 EFK :97 FCA :131 FE 1:267 FHe 2:31 MCL 4:398 MFG 1:208 RDB :190
L/255 L/346 L/408 L/1766
Coincya nivalis (Boiss. & Heldr.) Greuter & Burdet
Brassicaceae
Brassica nivalis Boiss. & Heldr.
Brassicella nivalis (Boiss. & Heldr.) O.E.Schultz
Hutera nivalis (Boiss. & Heldr.) Gomez-Campo
Rhynchosinapis nivalis (Boiss. & Heldr.) Heywood
AFE 11:238 FE 1:406 FGC 1:78 FHe 2:289 MCL 3:87 MFG 1:335 RDB :192
L/1180
Colchicum asteranthum Vassiliad. & K.Perss.
Colchicaceae
L/1824 L/2234
Colchicum chimonanthum K.Perss.
Colchicaceae
L/1576 L/2234
Colchicum confusum K.Perss.
Colchicaceae
L/1576 L/2234
Colchicum cretense Greuter
Colchicaceae
Colchicum creticum Davis 1949; Goulimis 1956
Colchicum pusillum sensu Gandoger 1916; Rechinger 1935
EFK :351 FCA :183 FE 5:22 MFG 2:654
L/518 L/785 L/1111 L/1247 L/2234
Colchicum euboeum (Boiss.) K.Perss.
Colchicaceae
Colchicum bivonae Guss. subsp. euboeum (Orph.) Nyman
Colchicum euboeum Orph.
Colchicum latifolium Sm. var. euboeum Boiss.
Colchicum parnassicum auct. non Sart., Orph. & Heldr.
FAe :709 FE 5:24
L/323b L/1585 L/1871 L/2234
478
479
480
481
482
483
484
Taxon
Cirsium morinifolium Boiss. & Heldr.
Asteraceae
Cirsium hypopsilum auct., non Boiss. & Heldr.
Eriolepis morinaefolia Gand.
EFK :322 FAe :652 FCA :63 FCSI :30 FE 4:235 MFG 2:484
L/522 L/554 L/1111
Taxon
Qualifiers
477
Taxon
Number
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+
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Pe
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+
+
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WAe
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StE
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IoI
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SPi
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+
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NPi
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EC
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NC
+
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
224 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Colchicum graecum K.Perss.
Colchicaceae
Colchicum parnassicum auct., non Sart., Orph. & Heldr. (1859)
Colchicum taygeteum Heldr. in sched. nom. nud.
MFG 2:658
L/121 L/299 L/1262a L/1963 L/2013 L/2200 L/2234
Colchicum lingulatum Boiss. & Spruner subsp. lingulatum
Colchicaceae
FE 5:24 FT 8:344
L/530 L/543 L/1585 L/1871 L/2234
Colchicum parlatoris Orph.
Colchicaceae
Colchicum rhenium Heldr., Epet. Syllog. Parnassou 1901: 252 (1901)
Colchicum neapolitanum (Ten.) Ten. subsp. parlatoris (Orph.) K.Richt.
FE 5:23
L/339 L/607 L/1963 L/2234
Colchicum parnassicum Sart., Orph. & Heldr. ex Boiss.
Colchicaceae
Colchicum lingulatum Boiss. & Spruner var. parnassicum (Sartori,
Orph. & Heldr.) Stef. nom. illegit.
FAe :709 FE 5:24 MFG 2:660
L/299 L/323b L/512 L/1973 L/2013 L/2234
Colchicum peloponnesiacum Rech.f. & P.H.Davis
Colchicaceae
FE 5:22 FT 8:366
L/607 L/1247 L/1844 L/1963 L/2234
Colchicum psaridis Heldr. ex Halácsy
Colchicaceae
Colchicum zahnii Heldr.
Colchicum boissieri Orph. var. gracile Orph., in sched., nom. nud.
Colchicum psaridis Heldr. ex Nyman, nom. nud.
FE 5:22 FT 11:250 FT 8:335
L/1351 L/1963 L/2234
Colchicum pulchellum K.Perss.
Colchicaceae
MFG 2:654
L/299 L/1963 L/2234
Colchicum rausii K.Perss.
Colchicaceae
L/1576 L/2234
485
486
487
488
489
490
491
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
492
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+
+
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+
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+
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 225
Consolida arenaria A.Carlström
Ranunculaceae
Consolida tomentosa sensu Rechinger 1943, partim?
FHe 2:22 FT 10:18 MCL 4:399 RDB :196
L/197 L/279 L/1111 L/2091
Consolida samia P.H.Davis
Ranunculaceae
FHe 2:22 FT 1:133 MCL 4:403 RDB :198
L/138 L/796 L/1005
Consolida tenuissima (Sm.) Soó
Ranunculaceae
Delphinium tenuissimum Sm.
AFE 8:73 FAe :181 FE 1:262 FGC 1:34 FHe 2:25 FT 1:134 MCL 4:403
L/323a L/530 L/736 L/1871 L/1981
Consolida tuntasiana (Halácsy) Soó
Ranunculaceae
Delphinium tuntasianum Halácsy
AFE 8:71 FE 1:261 FHe 2:24 MCL 4:404 RDB :200
L/530 L/736 L/1963
496
497
498
499
Conium divaricatum Boiss. & Orph.
Apiaceae
Conium maculatum L. var. divaricatum Boiss.
EFK :214 FAe :424 FCA :149 FE 2:342 FGC 1:656 FGS 1:44 FT 4:380
L/530
495
?
Colutea insularis Browicz
Fabaceae
Colutea arborescens sensu Rechinger 1943
FT 3:44 MCL 4:79
L/197 L/580 L/1111
494
Taxon
Colchicum sfikasianum Kit Tan & Iatroú
Colchicaceae
Colchicum amabile auct. non Heldr.
Colchicum kochii auct. non Parl.
Colchicum parnassicum auct. non Sart., Orph. & Heldr.
Colchicum polymorphum Orph., nom. nud.
L/1123 L/1261c L/1351 L/1585 L/1963 L/2234
Taxon
Qualifiers
493
Taxon
Number
+
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Pe
+
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StE
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IoI
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WAe
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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+
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EAe
Distribution
Qualifier
226 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Convolvulus argyrothamnos Greuter
Convolvulaceae
EFK :244 FCA :74 FE 3:80 MCL 3:4 RDB :202
L/336 L/337 L/1111 L/1583
Convolvulus mairei Halácsy
Convolvulaceae
FE 3:82 FGS 1:74 MCL 3:6
L/482 L/607 L/1963 L/2176
Corydalis blanda Schott subsp. olympica Lidén
Papaveraceae
FHe 2:102
L/577 L/1963
Corydalis blanda Schott subsp. oxelmannii Lidén
Papaveraceae
FHe 2:102
L/577 L/1963
Corydalis blanda Schott subsp. parnassica (Orph. & Heldr.) Lidén
Papaveraceae
Corydalis parnassica Orph. & Heldr.
AFE 9:70 FE 1:304 FHe 2:102 MCL 4:271 MFG 1:232
L/512 L/577 L/1963
Corydalis thasia (Stoj. & Kit.) Stoj. & Kit.
Papaveraceae
Corydalis solida (L.) Clairv. subsp. longicarpa Lidén
Corydalis tenella Ledeb. ex Nordm. var. thasia Stoj. & Kit.
AFE 9:74 FE 1:304 FHe 2:103
L/554 L/577 L/1267a L/2181
Corydalis uniflora (Sieber) Nyman
Papaveraceae
Corydalis rutifolia (Sm.) DC. var. subuniflora Boiss. & Heldr.
Corydalis rutifolia (Sm.) DC. subsp. uniflora (Sieber) Cullen &
P.H.Davis
Fumaria uniflora Sieber
Pistolochia uniflora (Sieber) Sojak
AFE 9:66 EFK :106 FCA :122 FE 1:304 FHe 2:100 MCL 4:272 MFG 1:232
L/577 L/856 L/1111 L/1766
500
501
502
503
504
505
506
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+
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+
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+
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 227
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Crataegus xkillinica K.I.Chr.
Rosaceae
L/293
Crataegus xpeloponnesiaca Byatt
Rosaceae
L/293 L/1963
Crepis arcuata Kamari & Strid
Asteraceae
MFG 2:583 RDB :206
L/264
Crepis athoa Boiss.
Asteraceae
Hieracioides athoum (Boiss.) O.Kuntze
FE 4:352 MFG 2:591
L/1973
Crepis auriculifolia Sieber ex Sprengel
Asteraceae
Crepis raulinii Boiss.
Hieracioides auriculaefolium (Sieber.) O.Kuntze
EFK :341 FAe :699 FCA :63 FCSI :30 FE 4:351 MFG 2:585 RDB :208
L/499 L/518 L/522 L/554 L/707 L/1070 L/1111 L/1130
Crepis cretica Boiss.
Asteraceae
Crepis neglecta L. f. cretica (Boiss.) Vierh.
Crepis neglecta L. subsp. cretica (Boiss.) Hay.
Crepis neglecta sensu Barb. karp.; Halácsy Conspect. Fl. Graec. p.p.
Crepis neglecta var. fuliginosa sensu auct., Rechinger Fl. Aeg.
Crepis neglecta var. graeca sensu auct. Kret., Rechinger Fl. Aeg.; 1943
Crepis neglecta var. stricta sensu auct., Rechinger Fl. Aeg.
Crepis neglecta var. typica sensu auct., Rechinger Fl. Aeg.
EFK :340 FAe :700 FCA :64 FCSI :30 FE 4:356 MFG 2:595
L/268 L/518 L/522 L/554 L/707 L/762 L/821 L/1111 L/1766 L/2093
508
509
510
511
512
513
Taxon
Crataegus pycnoloba Boiss. & Heldr.
Rosaceae
FE 2:77 MFG 1:441
L/159 L/160 L/512 L/607 L/1003 L/1963
Taxon
Qualifiers
507
Taxon
Number
.
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+
+
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Pe
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StE
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WAe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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+
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KK
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EAe
Distribution
Qualifier
228 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Crepis hellenica Kamari
Asteraceae
Crepis fuliginosa auct. plur., non Crepis fuliginosa Sm.
Crepis fuliginosa Sm. var. graeca (Vierh.) Greuter
Crepis neglecta f. fuliginosa auct. plur., non Crepis fuliginosa Sm.
Crepis neglecta L. f. graeca Vierh.
Crepis neglecta L. subsp. graeca (Vierh.) Rech.f.
Crepis neglecta L. var. graeca (Vierh.) Hayek
Crepis neglecta var. fuliginosa auct. plur., non Crepis fuliginosa Sm.
FAe :700 FT 5:838 MCL 2:186 MFG 2:595
L/163 L/177 L/423 L/443 L/447 L/449 L/530 L/539 L/540 L/707 L/776 L/821 L/869 L/928 L/1101 L/1288 L/1963 L/2013 L/2181 L/2247
!
!
517
518
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Crepis hellenica Kamari subsp. insularis Kamari
Asteraceae
MCL 2:186 MFG 2:595
L/539 L/540 L/776 L/821 L/1963 L/2237
Crepis incana Sm.
Asteraceae
Hieracioides incanum (Sm.) O.Kuntze
FE 4:392 MFG 2:589
L/259 L/512 L/530 L/607 L/1104 L/1871 L/1963
519
520
Crepis hellenica Kamari subsp. hellenica
Asteraceae
FAe :700 FT 5:838 MCL 2:186 MFG 2:595
L/163 L/447 L/449 L/516 L/530 L/607 L/776 L/821 L/1871 L/1963 L/2013 L/2181
Crepis heldreichiana (O.Kuntze) Greuter
Asteraceae
Crepis taygetea Babcock
Hieracioides heldreichianum O.Kuntze
MFG 2:589 RDB :212
L/259 L/607 L/1963
516
Crepis fraasii Sch. Bip. var. mungieri (Boiss. & Heldr.) Hayek
Asteraceae
Crepis fraasii Sch. Bip. subsp. mungieri (Boiss. & Heldr.) P.D.Sell
Crepis mungieri Boiss. & Heldr.
Crepis sieberi. var. mungieri (Boiss. & Heldr.) Boiss.
EFK :339 FCA :64 FE 4:348 FT 5:821 MFG 2:582
L/308d L/987 L/1111
Crepis guioliana Babcock
Asteraceae
FE 4:352 MFG 2:590
L/307a L/1104
!
515
514
+
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+
+
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+
+
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+
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+
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+
PATTERNS OF PLANT ENDEMISM IN GREECE 229
Crepis sibthorpiana Boiss. & Heldr.
Asteraceae
Crepis divaricata Boiss. & Heldr. subsp. sibthorpiana (Boiss. & Heldr.)
Hayek
Hieracioides sibthorpianum (Boiss. & Heldr.) O.Kuntze
Hieracium foetidum Sm.
EFK :341 FCA :64 FCSI :31 FE 4:352 MFG 2:587 RDB :216
B/3060f L/259 L/518 L/519 L/522 L/554 L/1111
Crepis tybakiensis Vierh.
Asteraceae
Crepis flexiscapa Rech.f.
Crepis foetida L subsp. maritima var. tybakensis (Vierh.) Hayek
EFK :341 FAe :702 FCA :64 FCSI :31 FE 4:357
L/253 L/518 L/522 L/554 L/762 L/780 L/1111 L/2093
Crepis xcytherea Kamari
Asteraceae
L/1052 L/1104 L/1878
Crocus biflorus Miller subsp. melantherus B.Mathew
Iridaceae
Crocus melantherus Boiss. & Orph.
L/607 L/1089 L/1588 L/1708 L/1963
Crocus biflorus Miller subsp. stridii (Papan. & Zacharof) B.Mathew
Iridaceae
Crocus stridii Papan. & Zacharof
RDB :218
L/399 L/566 L/607 L/1089 L/1287
Crocus boryi Gay
Iridaceae
Crocus boryanus var. graius Herbert
Crocus cretensis Körnicke
Crocus marathonicus Heldr.
EFK :367 FCA :177 FE 5:98 MFG 2:729
L/607 L/913 L/1089 L/1111 L/1131 L/1244 L/1708 L/1843a L/1844 L/1881 L/1963
522
523
524
525
526
527
Taxon
Crepis merxmuelleri Kamari & Hartvig
Asteraceae
MFG 2:587 RDB :214
L/995 L/1076
Taxon
Qualifiers
521
Taxon
Number
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SPi
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NPi
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EAe
Distribution
Qualifier
230 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Crocus goulimyi Turrill
Iridaceae
Crocus goulimyi Turrill subsp. leucanthus (B.Mathew) B.Mathew
FE 5:97 RDB :220
L/399 L/607 L/785 L/1089 L/1090 L/1708 L/1881 L/1963 L/2237
Crocus hadriaticus Herb.
Iridaceae
Crocus hadriaticus Herb. subsp. parnonicus B.Mathew
Crocus hadriaticus Herb. subsp. parnassicus (B.Mathew) B.Mathew
Crocus hadriaticus Herb. var. chrysovelonicus Herb.
Crocus hadriaticus Herb. var. saundersianus Herb.
Crocus peloponnesiacus L.
FE 5:97 MFG 2:729
L/607 L/1089 L/1123 L/1131 L/1708 L/1846 L/1963 L/2237
Crocus laevigatus Bory & Chaub.
Iridaceae
Crocus boryi Bory & Chaub. var. laevigatus (Bory & Chaub.) Baker
Crocus fontenayi Reut.
Crocus laevigatus Bory & Chaub. var. fontenayi (Reuter) Halácsy
Crocus veneris Maw, non Tappeiner ex Poech
EFK :366 FAe :738 FCA :178 FE 5:98 MFG 2:729
L/128 L/323b L/346 L/430 L/516 L/518 L/530 L/554 L/607 L/707 L/750 L/776 L/778 L/869 L/938 L/1052 L/1089 L/1111 L/1210d L/1708 L/1882 L/1917 L/1963 L/1973 L/2181
Crocus niveus Bowles
Iridaceae
FE 5:97
L/607 L/785 L/1089 L/1708 L/1881 L/1963
Crocus oreocreticus B.L.Burtt
Iridaceae
EFK :366 FCA :178 FE 5:97 MFG 2:728
L/785 L/1089 L/1111
529
530
532
533
531
Crocus cartwrightianus Herb.
Iridaceae
Crocus graecus Chapp.
Crocus graecus Chapp. var. cartwrightianus (Herb.) Hayek
Crocus pallasii Spruner ex Boiss., non Goldb.
Crocus sativus L. var. cartwrightianus (Herb.) Maw
EFK :367 FAe :738 FCA :177 FE 5:97 FGC 3:200 FGS 1:103 FGS 2:82
L/177 L/530 L/707 L/776 L/869 L/1089 L/1111 L/2181
528
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 231
Crocus sieberi Gay subsp. atticus (Boiss. & Orph.) B.Mathew
Iridaceae
Crocus atticus (Boiss. & Orph.) Orph.
Crocus sieberi Gay var. atticus Boiss. & Orph.
FE 5:95 MFG 2:725
L/399 L/443 L/530 L/1089 L/1871 L/2107 L/2181
Crocus sieberi Gay subsp. nivalis (Bory & Chaub.) B.Mathew
Iridaceae
Crocus nivalis Bory & Chaub.
Crocus sieberi f. nivalis (Bory & Chaub.) Heldr.
FE 5:95 MFG 2:726
L/399 L/607 L/1089 L/1708 L/1963 L/2107
Crocus sieberi Gay subsp. sieberi
Iridaceae
Crocus sibthorpianus Herbert var. latifolius Herbert
Crocus sieberi var. heterochromus Halácsy
Crocus sieberi var. versicolor Boiss. & Heldr.
Crocus sieberianus Herbert
FCA :178 FE 5:95 MFG 2:725
L/1089 L/1111 L/1766
Crocus tournefortii Gay
Iridaceae
Crocus boryi Gay subsp. tournefortii (Gay) Greuter, Matthäs & Risse
Crocus boryi Gay var. orphanidis (Hook.f.) Baker
Crocus fontenayi Nyman non Reut.
Crocus orphanidis Hook.f.
Crocus pholegandrius Orph.
EFK :367 FAe :737 FCA :178 FE 5:98 FT 8:437
L/197 L/201 L/268 L/430 L/869 L/1049 L/1089 L/1111 L/1210d
Cruciata taurica (Pall. ex Willd.) Ehrend. subsp. euboea (Ehrend.)
Ehrend.
Rubiaceae
Cruciata coronata (Sm.) Ehrend. subsp. euboea (Ehrend.) Ehrend.
Galium coronatum Sm. var. euboeum Ehrend.
FE 4:37 MFG 2:330
L/323b L/1871
535
536
537
538
539
Taxon
Crocus robertianus C.D.Brickell
Iridaceae
FE 5:98 RDB :224
L/138 L/399 L/1089
Taxon
Qualifiers
534
Taxon
Number
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+
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WAe
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StE
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IoI
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SPi
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NPi
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NC
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Kik
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KK
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EAe
Distribution
Qualifier
232 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Cuscuta rausii M.A.García
Convolvulaceae
FCSII :
L/1111 L/2252
Cyanus baldaccii (Degen ex Halácsy) Holub
Asteraceae
Centaurea baldaccii Degen ex Halácsy
EFK :327 FCA :61 FCSII 0: FE 4:298 FGC 2:140 MFG 2:516 RDB :142
L/1111
Cyclamen creticum Hildebr.
Primulaceae
Cyclamen creticum Hildebr. var. pallide-roseum Grey-Wilson. nom.
inval.
Cyclamen repandum Sm. subsp. creticum (Dörfler) Debussche &
Thompson
Cyclamen repandum Sm. var. creticum Dörfler
Cyclamen hederifolium Aiton subsp. creticum (Hildebr.) O.Schwarz
EFK :222 FCA :129 FCSI :73 FCSII : FE 3:25 MCL 4:376
L/257 L/541 L/856 L/1111 L/1807 L/1809 L/1810 L/1811 L/1848 L/1884 L/1885 L/1886
Cyclamen graecum Link subsp. Graecum
Primulaceae
Cyclaminus graeca (Link) Asch.
Cyclaminos miliakiaresii Heldr.
Cyclaminos graeca (Link) Heldr.
Cyclamen velutinum Jord.
Cyclamen persicum sensu Sibth. & Sm.
Cyclamen penteliki Hildebr.
Cyclamen gaydurowryssii (orthographical variant) Glasau
Cyclamen gaidurowryssii Glasau
Cyclamen aeginiticum Hildebr.
Cyclamen miliarakesii Heldr. ex Halácsy
Cyclamen miliarakesii Heldr. ex Hildebr.
FCSII :
L/1111 L/1807 L/1808 L/1809 L/1848
541
542
543
544
?
Cuscuta atrans Feinbrun
Convolvulaceae
Cuscuta epithymum var. micrantha sensu Davis 1953
EFK :242 FCA :75 FCSI :36 FE 3:76 MCL 3:9 MFG 2:14
L/346 L/1111 L/1766
540
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PATTERNS OF PLANT ENDEMISM IN GREECE 233
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Cyclamen rhodium Gorer f. albiflorum (B.Mathew) J.Compton &
Culham
Primulaceae
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan f. albiflorum
B.Mathew
Cyclamen repandum Sm. subsp. peloponnesiacum Grey-Wilson f.
albiflorum B.Mathew
L/1749 L/1884
Cyclamen rhodium Gorer subsp. peloponnesiacum (Grey-Wilson)
J.Compton & Culham
Primulaceae
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan
Cyclamen repandum Sm. subsp. peloponnesiacum Grey-Wilson
EFK :222 MCL 4:377
L/159 L/395 L/541 L/833 L/1749 L/1750 L/1807 L/1809 L/1810 L/1848 L/1847 L/1884 L/1963 L/2181
Cyclamen rhodium Gorer subsp. rhodium
Primulaceae
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan subsp. rhodense
(Meikle) Kit Tan
Cyclamen repandum Sm. var. rhodense Meikle
Cyclamen repandum Sm. subsp. rhodense (Meikle) Grey-Wilson
FT 6:131 MCL 4:377
L/197 L/541 L/833 L/1111 L/1755 L/1807 L/1809 L/1810 L/1848 L/1884 L/1963
547
548
549
Cyclamen graecum Link subsp. mindleri A.P.Davis & Govaerts
Primulaceae
Cyclamen persicum Miller subsp. mindleri (Heldr.) Knuth
Cyclamen graecum Link subsp. candicum Ietsw. nom. illeg.
Cyclamen mindleri Heldr.
Cyclamen pseudograecum Hildebr.
FCSII :
L/1807 L/1808 L/1809
Taxon
Cyclamen rhodium Gorer
Primulaceae
EFK :222 FT 6:131 MCL 4:377
L/159 L/197 L/395 L/541 L/833 L/1749 L/1750 L/1755 L/1807 L/1809 L/1810 L/1811 L/1847 L/1848 L/1884 L/1885 L/1963
?
Taxon
Qualifiers
546
545
Taxon
Number
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+
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Pe
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WAe
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StE
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SPi
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NPi
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EC
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NAe
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Distribution
Qualifier
234 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Cyclamen rhodium Gorer subsp. vividum (Grey-Wilson) J.Compton
& Culham
Primulaceae
Cyclamen repandum Sm. var. vividum (Grey-Wilson) Kit Tan
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan subsp. vividum
(Grey-Wilson) Kit Tan
Cyclamen repandum Sm. f. vividum Grey-Wilson
L/541 L/1807 L/1809 L/1810 L/1848 L/1847 L/1884 L/1963
Cymbalaria microcalyx (Boiss.) Wettst. subsp. alba (Voliotis) Kit Tan
Plantaginaceae
Cymbalaria microcalyx (Boiss.) Wettst. subsp. microcalyx var. alba
Voliotis
L/1199 L/1963
Cymbalaria microcalyx (Boiss.) Wettst. subsp. dodekanesi (Cuf.)
Greuter
Plantaginaceae
Cymbalaria acutiloba (Boiss. & Heldr.) Speta subsp. dodekanesi
(Greuter) Speta
Cymbalaria acutiloba (Boiss. & Heldr.) Speta subsp. microcalyx
(Greuter) Speta
EFK :277 FCA :142 FE 3:238 FT 6:673 MFG 2:207
L/119 L/191 L/192 L/197 L/253 L/561 L/607 L/750
L/1049 L/1111
Cymbalaria microcalyx (Boiss.) Wettst. subsp. microcalyx
Plantaginaceae
Cymbalaria microcalyx (Boiss.) Wettst. var. heterosepala (Cuf.) Cuf.
Linaria microcalyx (Boiss.) Wettst. var. heterosepala Cuf.
Linaria microcalyx (Boiss.) Wettst. var. orphanidiana Cuf.
EFK :277 FCA :142 FE 3:238 MFG 2:206
L/196 L/253 L/554 L/561 L/607 L/750 L/932 L/1111 L/1174 L/1199 L/1963 L/2093
Cymbalaria microcalyx (Boiss.) Wettst. subsp. minor (Cuf.) Greuter
Plantaginaceae
Linaria microcalyx (Boiss.) Wettst. var. minor Cuf.
FE 3:238 MFG 2:207
L/119 L/607 L/930 L/1123 L/1174 L/1846 L/1963
Cynoglossum sphacioticum Boiss. & Heldr.
Boraginaceae
EFK :252 FAe :445 FCA :39 FCSI :14 FE 3:120 MCL 1:78 MFG 2:59 RDB :230
L/518 L/707 L/1111
550
551
552
553
554
555
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 235
Delphinium hirschfeldianum Heldr. & Holzm.
Ranunculaceae
AFE 8:64 FE 1:259 FHe 2:19 MCL 4:407
563
?
Delphinium hellenicum Pawl.
Ranunculaceae
Delphinium junceum auct. non DC.
Delphinium longipes sensu Hayek, non Moris
Delphinium peregrinum sensu Halácsy, non L.
AFE 8:66 FE 1:260 FHe 2:20 MCL 4:407
L/585 L/1111 L/1713 L/1800 L/1846 L/1878 L/1963
562
Daphne euboica Rech.f.
Thymelaeaceae
FE 2:257 MFG 1:593
L/323a L/391 L/1871
560
Daucus conchitae Greuter
Apiaceae
FT 11:146
L/192
Dactylorhiza smolikana B.Willing & E.Willing
Orchidaceae
L/611 L/2185
559
561
Dactylorhiza pythagorae Gölz & Reinchard
Orchidaceae
Dactylorhiza kalopissii E.Nelson subsp. pythagorae (Gölz & Reinchard)
E.Nelson
FT 11:304
L/611 L/802 L/2185
558
?
Dactylorhiza graeca H.Baumann
Orchidaceae
Dactylorhiza cordigera (Fries) Soó var. rhodopeia H.Presser
Dactylorhiza cordigera (Fries) Soó var. graeca (H.Baumann) H.Presser
Dactylorhiza cordigera (Fries) Soó subsp. graeca (H.Baumann) Kreutz
MFG 2:876 RDB :234
L/526 L/611 L/956 L/1184 L/2185
557
Taxon
Dactylis glomerata L. subsp. rigida (Boiss. & Heldr.) Hayek
Poaceae
Dactylis rigida Boiss. & Heldr.
EFK :382 FAe :776 FCA :168 FE 5:171 MFG 2:776
L/1111
Taxon
Qualifiers
556
Taxon
Number
+
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Pe
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StE
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IoI
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NPi
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NC
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NAe
+
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Kik
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KK
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+
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EAe
Distribution
Qualifier
236 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Dianthus cinnamomeus Sm.
Caryophyllaceae
AFE 7:187 EFK :93 FCA :46 FCY 1:218 FE 1:244 FHe 1:352 FT 2:114 MCL 1:198
L/169 L/869 L/1111
!
!
566
567
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Dianthus desideratus Strid
Caryophyllaceae
FHe 1:371
Dianthus corymbosus Sm.
Caryophyllaceae
Dianthus chalcidicus Halácsy
AFE 7:194 FE 1:246 FGC 1:206 FHe 1:358 MCL
1:191 MFG 1:192
B/1316 L/111 L/121 L/447 L/554 L/566 L/1107 L/1112 L/1832d L/1973 L/2200
569
570
Dianthus cinnamomeus Sm. subsp. naxensis Runemark
Caryophyllaceae
FHe 1:352
L/759a
568
!
Dianthus biflorus Sm.
Caryophyllaceae
Dianthus biflorus Sm. subsp. samaritani (Heldr.) Maire & Pet.
Dianthus cinnabarinus Spruner.
Dianthus cinnabarinus Spruner. var. alpinus Boiss.
Dianthus samaritani Heldr.
AFE 7:192 FE 1:245 FGC 1:213 FHe 1:357 MCL 1:187 MFG 1:190
L/99 L/323b L/512 L/587 L/607 L/1467 L/1846 L/1871 L/1871 L/1963 L/2013 L/2200
565
Dianthus cinnamomeus Sm. subsp. cinnamomeus
Caryophyllaceae
AFE 7:187 EFK :93 FCA :46 FCY 1:218 FE 1:244 FHe 1:352 FT 2:114 MCL 1:198
L/759a L/1111
Dianthus androsaceus (Boiss. & Heldr.) Hayek
Caryophyllaceae
Dianthus lilacinus Boiss. & Heldr. var. androsaceus Boiss. & Heldr.
AFE 7:205 FE 1:235 FHe 1:369 MCL 1:186 MFG 1:196 RDB :242
L/1830 L/1938b L/1963
564
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PATTERNS OF PLANT ENDEMISM IN GREECE 237
Dianthus fruticosus L.
Caryophyllaceae
Dianthus arboreus auct. plur.
AFE 7:201 EFK :92 FAe :155 FAe :156 FCA :46 FE 1:234 FGC 1:201 FHe 1:363 FT 10:72 FT 2:121 MCL 1:193
B/3060c L/196 L/206 L/252 L/518 L/554 L/707 L/776 L/1111 L/1244 L/1690 L/1800 L/1917 L/1963 L/2093 L/2181
Dianthus fruticosus L. subsp. amorginus Runemark
Caryophyllaceae
AFE 7:201 EFK :92 FCA :46 FHe 1:364 MCL 1:193
L/252 L/268 L/414 L/776 L/1111
Dianthus fruticosus L. subsp. carpathus Runemark
Caryophyllaceae
AFE 7:201 EFK :92 FCA :46 FCSI :18 FHe 1:365 MCL 1:193
L/414 L/1111
Dianthus fruticosus L. subsp. creticus (Tausch) Runemark
Caryophyllaceae
Dianthus creticus Tausch
AFE 7:201 EFK :92 FAe :156 FCA :46 FCSI :18 FHe 1:365 FT 2:121 MCL 1:193
L/336 L/414 L/518 L/554 L/1096 L/1111
Dianthus fruticosus L. subsp. fruticosus
Caryophyllaceae
AFE 7:201 FHe 1:364 MCL 1:193
L/414 L/1111 L/2181
Dianthus fruticosus L. subsp. karavious Runemark
Caryophyllaceae
FHe 1:365 FT 10:72 MCL 1:193
L/414 L/1111
572
573
574
575
576
577
Taxon
Dianthus diffusus Sm.
Caryophyllaceae
Dianthus cylleneus Boiss. & Heldr.
Dianthus diffusus Sm. var. cylleneus (Boiss. & Heldr.) Halácsy
Dianthus glandulosopubescens Halácsy
Dianthus pubescens Sm.
Dianthus pubescens Sm. var. glabratus Boiss.
AEE 7:195 FAe :158 FHe 1:360 FT 2:126 MCL 1:191 MFG 1:192
L/447 L/530 L/759a L/1865 L/1871
Taxon
Qualifiers
571
Taxon
Number
.
.
.
.
.
.
.
.
.
+
+
StE
.
+
+
Pe
.
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+
WAe
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+
+
IoI
.
.
.
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.
.
SPi
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.
.
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.
.
NPi
.
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.
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.
EC
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.
NC
.
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.
.
NE
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.
.
.
.
.
.
NAe
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+
.
.
+
+
+
Kik
.
.
+
+
+
+
.
KK
+
.
.
.
.
+
.
EAe
Distribution
Qualifier
238 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Dianthus haematocalyx Boiss. & Heldr. subsp. pruinosus (Boiss. &
Orph.) Hayek
Caryophyllaceae
Dianthus haematocalyx Boiss. & Heldr. var. pruinosus (Boiss. & Orph.)
Boiss.
Dianthus pruinosus Boiss. & Orph.
AFE 7:191 FE 1:245 FHe 1:356 MCL 1:196 MFG 1:190
L/1011
Dianthus haematocalyx Boiss. & Heldr. subsp. ventricosus (Heldr.)
Maire & Petitm.
Caryophyllaceae
Dianthus alpinus Sibth. sensu Smith (1809)
Dianthus haematocalyx Boiss. & Heldr. subsp. sibthorpii (Vieh.) Hayek
Dianthus haematocalyx Boiss. & Heldr. var. alpinus Boiss.
Dianthus sibthorpii Vierh.
Dianthus ventricosus Heldr. ex Halácsy
AFE 7:191 FE 1:245 FHe 1:356 MCL 1:196 MFG 1:189
L/512 L/1011 L/2013
583
584
Dianthus gracilis Sm. subsp. xanthianus (Davidov) Tutin
Caryophyllaceae
Dianthus xanthianus Davidov
AFE 7:190 FE 1:245 FHe 1:354 MFG 1:187
L/1349
581
Dianthus haematocalyx Boiss. & Heldr. subsp. phitosianus
Constantin.
Caryophyllaceae
L/1011
Dianthus fruticosus L. subsp. sitiacus Runemark
Caryophyllaceae
AFE 7:201 FCA :47 FCSI :18 FHe 1:365 MCL 1:193
L/252 L/336 L/414 L/1111
580
582
Dianthus fruticosus L. subsp. rhodius (Rech.f.) Runemark
Caryophyllaceae
Dianthus rhodius Rech.f.
AFE 7:201 FHe 1:365 FT 2:121 MCL 1:193
L/191 L/197 L/414 L/445 L/580 L/707 L/1052 L/1111
579
!
Dianthus fruticosus L. subsp. occidentalis Runemark
Caryophyllaceae
AFE 7:201 FCA :46 FCSI :18 FHe 1:364 MCL 1:193
L/108 L/119 L/252 L/414 L/574 L/607 L/765 L/1111 L/1123 L/1244 L/1690 L/1800 L/1917 L/1963
578
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PATTERNS OF PLANT ENDEMISM IN GREECE 239
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Dianthus juniperinus Sm. subsp. aciphyllus (Sieber ex Ser.) Turland
Caryophyllaceae
Dianthus aciphyllus Ser. var. aciphyllus
Dianthus arboreus L., nomen ambig.
Dianthus juniperinus Sm. var. aciphyllus (Sieber ex Ser.) Halácsy
Dianthus juniperinus Sm. var. sieberi Boiss.
AFE 7:200 EFK :93 FAe :157 FCA :47 FCSI :18 FE 1:234 FHe 1:363 MCL 1:196 MFG 1:194
L/408 L/426 L/518 L/707 L/1111
Dianthus juniperinus Sm. subsp. bauhinorum (Greuter) Turland
Caryophyllaceae
Betonica coronariaarboreacretica J. Bauhin
Caryophyllus arborescenscreticus C. Bauhin
Caryophyllus sylvestrisarboreus Alpini
Dianthus aciphyllus var. bauhinorum Greuter
Dianthus arboreus L., non Sm. & auct. plur., nomen ambiguum
EFK :93 FAe :157 FCA :47 FCSI :18 FHe 1:363
L/408 L/426 L/1111
Dianthus juniperinus Sm. subsp. heldreichii Greuter
Caryophyllaceae
Dianthus juniperinus sensu auct. Raulin 1869 p.p., non Sm.
Dianthus juniperinus Sm var. sieberi sensu auct. non Boiss.
AFE 7:199 EFK :92 FCA :47 FE 1:234 FHe 1:362 MCL 1:196 MFG 1:194
L/255 L/408 L/426 L/1111
Dianthus juniperinus Sm. subsp. idaeus Turland
Caryophyllaceae
EFK :93 FCA :47 FCSI :18 FHe 1:363
L/426 L/1111
Dianthus juniperinus Sm. subsp. juniperinus
Caryophyllaceae
AFE 7:199 EFK :93 FCA :47 FCSI :18 FE 1:234 FHe 1:362 MCL 1:196
L/408 L/426 L/518 L/1111
586
587
588
589
590
Taxon
Dianthus juniperinus Sm.
Caryophyllaceae
Caryophyllus creticus, arboreus Tournefort
Dianthus arboreus sensu auct. p.p., non s. str.
AFE 7:199 AFE 7:200 EFK :92 FAe :157 FCA :46 FCSI :18 FE 1:234 FHe 1:362 MCL 1:196
L/255 L/408 L/426 L/1096 L/1111
Taxon
Qualifiers
585
Taxon
Number
.
.
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.
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.
.
.
StE
.
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Pe
.
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WAe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
+
+
+
+
+
+
KK
.
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.
.
EAe
Distribution
Qualifier
240 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Dianthus serratifolius Sm. subsp. abbreviatus (Heldr. ex Halácsy)
Strid
Caryophyllaceae
Dianthus serratifolius Sm. var. abbreviatus Heldr. & Halácsy
Dianthus taygeteus Quézel & Contandr.
AFE 7:169 FE 1:241 FHe 1:347 MCL 1:203 MFG 1:179
L/530 L/607 L/1963
Dianthus serratifolius Sm. subsp. serratifolius
Caryophyllaceae
AFE 7:169 FE 1:241 FHe 1:347 MCL 1:203 MFG 1:180
L/310a L/530 L/607 L/1830 L/1938 L/1963
596
597
Dianthus myrtinervius Griseb. subsp. caespitosus Strid & Papan.
Caryophyllaceae
Dianthus kajmaktzalanicus Micevski
Dianthus myrtinervius Griseb. var. oxylepis Boiss.
Dianthus oxylepis (Boiss.) Kummerle & Jav.
AFE 7:185 FHe 1:351
L/1129
594
Dianthus serratifolius Sm.
Caryophyllaceae
AFE 7:169 FE 1:241 FHe 1:347 MCL 1:203
L/121 L/310a L/377 L/530 L/607 L/1830 L/1938b L/1963 L/1981 L/2200
Dianthus mercurii Heldr.
Caryophyllaceae
AFE 7:192 FE 1:245 FHe 1:357 MCL 1:187 MFG 1:191
L/607 L/1963
593
595
Dianthus juniperinus Sm. subsp. pulviniformis (Greuter) Turland
Caryophyllaceae
Dianthus pulviniformis Greuter
AFE 7:199 EFK :93 FCA :47 FCSI :18 FE 1:234 FHe 1:362 MCL 1:196 MFG 1:194
L/255 L/408 L/426 L/1111
592
!
Dianthus juniperinus Sm. subsp. kavusicus Turland
Caryophyllaceae
Dianthus juniperinus var. sieberi sensu Rechinger (Phyton, 1943) p.p.
EFK :93 FCA :47 FCSI :18 FHe 1:363
L/426 L/1111
591
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 241
Digitalis leucophaea Sm. subsp. ikarica P.H.Davis
Plantaginaceae
FT 6:687
L/539 L/540
Digitalis leucophaea Sm. subsp. leucophaea
Plantaginaceae
Digitalis lanata Ehrh. subsp. leucophaea (Sibth. & Sm.) H.Werner
FAe :485 FE 3:241 FT 6:687
L/543 L/1267b L/1973
606
Digitalis laevigata Waldst. & Kit. subsp. graeca (Ivanina) Werner
Plantaginaceae
FE 3:241 MFG 2:209
L/121 L/532 L/566 L/1246 L/1830 L/2200
603
605
Dianthus xylorrhizus Boiss. & Heldr.
Caryophyllaceae
AFE 7:163 EFK :94 FAe :159 FCA :47 FCSI :19 FE 1:239 FHe 1:346 MCL 1:189 RDB :246
L/252 L/1111 L/1583
602
Digitalis leucophaea Sm.
Plantaginaceae
FAe :485 FE 3:241 FT 6:687
L/489 L/539 L/543 L/1267b L/1973
Dianthus tymphresteus (Boiss. & Spruner) Heldr. & Sart.
Caryophyllaceae
Dianthus viscidus Bory & Chaub. var. tymphresteus Boiss. & Spruner
AFE 7:195 FE 1:246 FHe 1:359 MCL 1:191 MFG 1:193
L/121 L/160 L/512 L/1467 L/1588 L/1963 L/2013 L/2200
601
604
Dianthus strymonis Rech.f.
Caryophyllaceae
FE 1:236 FHe 1:370 MCL 1:195 MFG 1:198
L/785
600
!
Dianthus stamatiadae Rech.f.
Caryophyllaceae
AFE 7:203 FE 1:235 FHe 1:366 MCL 1:204
L/1442
599
Taxon
Dianthus sphacioticus Boiss. & Heldr.
Caryophyllaceae
AFE 7:185 EFK :93 FAe :159 FCA :47 FCSI :19 FE 1:244 FHe 1:352 MCL 1:204 MFG 1:185
L/518 L/707 L/1111
Taxon
Qualifiers
598
Taxon
Number
+
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+
+
.
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.
StE
+
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Pe
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+
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IoI
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WAe
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.
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SPi
.
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+
.
.
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.
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NPi
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EC
.
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+
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+
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NC
+
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+
+
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+
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NE
+
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+
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NAe
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Kik
.
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+
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KK
.
+
+
.
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.
.
EAe
Distribution
Qualifier
242 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Draba cretica Boiss. & Heldr.
Brassicaceae
Draba cretica Boiss. & Heldr. var. lassitica Halácsy
Draba cretica Boiss. & Heldr. var. sphaciotica Halácsy
AFE 11:87 EFK :120 FCA :81 FCSI :41 FE 1:374 FHe 2:241 MCL 3:96 MFG 1:312
L/522 L/831 L/1111 L/1766
Draba lacaitae Boiss.
Brassicaceae
Draba aizoides L. subsp. lacaitae (Boiss.) Maire & Petitm.
Draba aizoides L. var. brevistyla Boiss. nom. illeg.
Draba erostra Halácsy
AFE 11:85 FE 1:374 FHe 2:240 MCL 3:98 MFG 1:311
L/99 L/121 L/160 L/323b L/344 L/512 L/512 L/530
L/607 L/831 L/1830 L/1963 L/2013 L/2200
Draba laconica Stefanović & Kit Tan
Brassicaceae
FHe 2:239
Draba parnassica Boiss. & Heldr.
Brassicaceae
Draba boissieri Bald.
Draba hispanica Boiss. subsp. parnassica (Boiss. & Heldr.) Nyman
AFE 11:86 FE 1:374 FHe 2:240 MCL 3:99 MFG 1:312
L/323a L/512 L/831 L/1871 L/1963
Draba strasseri Greuter
Brassicaceae
AFE 11:85 FHe 2:239 MCL 3:99
L/942 L/1915 L/1963
Drymocallis halacsyana (Degen) Kurto & Strid
Rosaceae
Potentilla rupestris L. var. baeniczkyi (Friv.) T.Wolf subvar. halacsyana
(Degen) Hayek
Potentilla geoides M.Bieb. subsp. halacsyana (Degen) Strid
Potentilla halacsyana Degen
Potentilla rupestris L. var. halacsyana (Degen) T.Wolf
AFE 13:162 MFG 1:407
L/533 L/895 L/1252 L/1349 L/1940
607
608
609
610
611
612
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PATTERNS OF PLANT ENDEMISM IN GREECE 243
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Drymocallis longisepala (Strid) Kurtto & Strid subsp. epirotica
(Soják) Kurtto & Strid
Rosaceae
Potentilla geoides Bieb. subsp. geoides sensu Strid p.p.
Potentilla longisepala (Strid) Soják subsp. epirotica Soják
AFE 13:162
L/1940
Drymocallis longisepala (Strid) Kurtto & Strid subsp. longisepala
Rosaceae
AFE 13:162
L/1940
Ebenus cretica L.
Fabaceae
EFK :173 FCA :105 FCSI :57 FE 2:191 MCL 4:89
L/752 L/856 L/887 L/1096 L/1111 L/1766 L/2093
Ebenus sibthorpii DC.
Fabaceae
FAe :390 FE 2:191 MCL 4:90
L/99 L/197 L/323a L/530 L/581 L/607 L/1111 L/1871 L/1963
Echinops graecus Miller
Asteraceae
FAe :642 FE 4:213 FGC 2:88
L/323b L/530 L/869 L/1467 L/1871 L/1980 L/2181
Echinops ritro L. subsp. sartorianus (Boiss. & Heldr.) Kozuharov
Asteraceae
Echinops ritro L. var. sartorianus (Boiss. & Heldr.) Boiss.
Echinops sartorianus Boiss. & Heldr.
FE 4:214 FGC 2:91 MFG 2:476
L/456c L/607
Echinops sphaerocephalus L. subsp. taygeteus (Boiss. & Heldr.)
Kozuharov
Asteraceae
Echinops sphaerocephalus L. var. taygeteus (Boiss. & Heldr.) Boiss.
Echinops taygeteus Boiss. & Heldr.
FE 4:214 FGC 2:89
L/456c L/512 L/607
614
615
616
617
618
619
620
Taxon
Drymocallis longisepala (Strid) Kurtto & Strid
Rosaceae
Potentilla longisepala (Strid) Soják
Potentilla geoides Bieb. subsp. longisepala Strid
AFE 13:162 MFG 1:408
L/1901 L/1940
Taxon
Qualifiers
613
Taxon
Number
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+
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SPi
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IoI
+
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WAe
+
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StE
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Pe
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+
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NPi
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+
+
+
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EC
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+
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NC
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NE
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NAe
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+
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Kik
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+
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KK
.
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+
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.
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EAe
Distribution
Qualifier
244 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Epilobium vernonicum Snogerup
Onagraceae
MCL 4:252 MFG 1:652 RDB :248
L/1168
Epipactis cretica Kalopissis & Robatsch
Orchidaceae
Epipactis helleborine sensu auct. cret. non (L.) Crantz, Künkele 1979?
EFK :421 FCA :187 FCSI :111 RDB :250
L/430 L/526 L/611 L/734 L/791 L/1057 L/1111 L/1463 L/1766 L/1821 L/2185
Epipactis halácsyi Robatsch
Orchidaceae
Epipactis purpurata auct. non J.E. Smith
Epipactis graeca Halácsy nom nud.
L/611 L/2185
Epipactis olympica Robatsch
Orchidaceae
L/611 L/2185
Epipactis subclausa Robatsch
Orchidaceae
Epipactis atrorubens (Hoffm.) Besser subsp. subclausa (Robatsch) Kreutz
Epipactis atrorubens auct. non (Hoffm. ex Bernh.) Besser
Epipactis thessala B.Baumann & H.Baumann
MFG 2:867
L/526 L/611 L/2185
Eragrostis minor Host subsp. angusta H. Scholz & Raus
Poaceae
L/1896 L/2237
622
623
624
625
626
627
?
Edraianthus parnassicus (Boiss. & Spruner) Halácsy
Campanulaceae
Campanula parnassica Boiss. & Spruner
Halacsyella parnassica (Boiss. & Spruner) Janchen
FE 4:99 MCL 1:146 MFG 2:394
L/512 L/607 L/1467 L/1963 L/2013 L/2200
621
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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*
PATTERNS OF PLANT ENDEMISM IN GREECE 245
Erysimum atticum Boiss.
Brassicaceae
Erysimum pusillum Bory & Chaub subsp. atticum (Heldr. & Sart. ex
Boiss.) P.W. Ball
Erysimum parnassi Bory & Chaub var. atticum (Heldr. & Sart. ex
Boiss.) Hayek
Erysimum pusillum Bory & Chaub var. atticum (Heldr. & Sart. ex
Boiss.) Hayek
AFE 10:61 FE 1:331 FGC 1:65 FHe 2:132 MCL 3:108
L/261 L/530 L/830
Erysimum boryanum Boiss. & Spruner
Brassicaceae
AFE 10:59 FE 1:330 FGC 1:64 FHe 2:133 FT 1:472 MCL 3:111
L/261 L/607 L/1963 L/1981
634
635
Erysimum asperulum Boiss. & Heldr.
Brassicaceae
AFE 10:73 FE 1:333 FHe 2:147 MCL 3:108 MFG 1:246
L/261 L/830 L/2013 L/2200
!
633
+
+
+
+
.
.
.
Eryngium ternatum Poiret
Apiaceae
EFK :208 FCA :150 FE 2:323 RDB :258
L/1111
632
.
Eryngium amorginum Rech.f.
Apiaceae
EFK :208 FAe :399 FCA :150 FE 2:323 RDB :256
L/111 L/145 L/1111 L/1583
631
.
.
.
Erodium sibthorpianum Boiss. subsp. vetteri (Barley & Major) Davis
Geraniaceae
Erodium vetteri Barbey & Major
FT 2:483 MCL 3:250 MFG 1:552 RDB :172
L/796 L/975
630
.
.
.
Erodium hartvigianum Strid & Kit Tan
Geraniaceae
L/1894
StE
629
Pe
+
Taxon
+
Erodium chrysanthum L’Her. ex DC.
Geraniaceae
Erodium absinthoides Willd. subsp. chrysanthum (L’Herit. ex DC.) Maire
& Pettimengin
FE 2:201 MCL 3:250 MFG 1:551
L/138 L/512 L/530 L/607 L/736 L/759i L/1963
Taxon
Qualifiers
628
Taxon
Number
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.
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.
.
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IoI
WAe
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SPi
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NPi
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+
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EC
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+
.
NC
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NE
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NAe
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Kik
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+
+
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KK
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+
.
.
EAe
Distribution
Qualifier
246 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Erysimum creticum Boiss. & Heldr.
Brassicaceae
AFE 10:72 EFK :114 FCA :82 FCSI :42 FE 1:333 FHe 2:146 FT 1:477 MCL 3:109
L/1111 L/1138
641
Erysimum cephalonicum Polatschek
Brassicaceae
Erysimum pusillum Bory & Chaub. subsp. cephalonicum (Polatschek)
P.W. Ball
AFE 10:61 FE 1:331 FHe 2:135 MCL 3:111 MFG 1:244
L/121 L/261 L/382 L/530 L/607 L/1123 L/1139 L/1711 L/1871 L/1963 L/1981 L/2013 L/2200
639
Erysimum corinthium (Boiss.) Wettst.
Brassicaceae
Erysimum cheiri (L.) Crantz var. corinthium (Boiss.) Polatschek
Cheiranthus cheiri auct. p.p., non L.
Cheiranthus cheiri L. var. corinthius (Boiss.) Hausskn.
Cheiranthus corinthius Boiss.
Erysimum bisaccatum Formánek
AFE 10:50 FAe :215 FE 1:328 FHe 2:139 MCL 3:108
L/129 L/530 L/607 L/1111 L/1264 L/1690 L/1800 L/1877 L/1963
Erysimum candicum Snogerup subsp. carpathum Snogerup
Brassicaceae
Cheiranthus cheiri L. var. corinthius (Boiss.) Hausskn.
AFE 10:50 EFK :113 FCA :82 FCSI :41 FE 1:328 FHe 2:142 MCL 3:108
L/129 L/1111
638
640
Erysimum candicum Snogerup subsp. candicum
Brassicaceae
AFE 10:50 EFK :113 FCA :82 FCSI :41 FE 1:328 FHe 2:142 MCL 3:108
L/129 L/255 L/336 L/856 L/1111 L/1138
637
!
Erysimum candicum Snogerup
Brassicaceae
Cheiranthus cheiri L., pro parte
AFE 10:50 EFK :113 FCA :82 FCSI :41 FE 1:328 FHe 2:142 MCL 3:108
L/111 L/129 L/856 L/1111
636
.
.
+
.
+
.
+
.
.
+
.
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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+
+
+
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.
+
+
+
.
.
.
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.
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PATTERNS OF PLANT ENDEMISM IN GREECE 247
Erysimum mutabile Boiss. & Heldr.
Brassicaceae
AFE 10:64 EFK :113 FCA :83 FE 1:331 FHe 2:138 MCL 3:112 MFG 1:245
L/518 L/522 L/1102 L/1111 L/1138 L/1766
Erysimum naxense Snogerup
Brassicaceae
AFE 10:52 FE 1:329 FHe 2:142 MCL 3:109 RDB :260
L/129 L/869 L/1167
Erysimum olympicum Boiss.
Brassicaceae
AFE 10:72 FE 1:333 FHe 2:145 MCL 3:113 MFG 1:245
L/261
647
648
Erysimum microstylum Hausskn.
Brassicaceae
Erysimum pusillum Bory & Chaub. subsp. microstylum (Hausskn.)
Hayek
AFE 10:61 FE 1:331 FHe 2:138 MCL 3:111 MFG 1:243
L/532 L/1711 L/1846 L/1852 L/2013 L/2201
645
646
Erysimum krendlii Polatschek
Brassicaceae
Erysimum smyrnaeum sensu auct. Samothraki, non Boiss. & Balansa
FHe 2:148
L/1349
644
!
Erysimum hayekii (Jav. & Rech.f.) Polatschek
Brassicaceae
Erysimum aciphyllum sensu Boiss Fl. Orient.; Meikle 1954 Chios
Erysimum pusillum Bory & Chaub. subsp. hayekii Jav. & Rech.f.
AFE 10:61 FAe :209 FE 1:331 FHe 2:137 FT 11:40 MCL 3:110 MFG 1:243
L/169 L/197 L/261 L/321 L/575 L/776 L/869 L/1140 L/1957
643
Taxon
Erysimum graecum Boiss. & Heldr.
Brassicaceae
AFE 10:70 EFK :114 FAe :209 FCA :82 FE 1:333 FGC 1:66 FHe 2:144 FT 1:477 MCL 3:110
L/121 L/261 L/516 L/530 L/561 L/607 L/696 L/856 L/1138 L/1288 L/1349 L/1713 L/1830 L/1871 L/1963 L/2013 L/2200
Taxon
Qualifiers
642
Taxon
Number
+
.
.
.
.
.
.
.
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.
.
+
StE
.
+
Pe
.
.
.
.
.
.
+
WAe
.
.
.
+
.
.
.
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+
.
.
.
.
.
SPi
IoI
.
.
.
+
.
.
.
NPi
.
.
.
+
.
.
+
EC
+
.
.
+
.
.
.
NC
.
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NE
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+
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NAe
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+
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+
.
Kik
.
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+
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KK
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.
.
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+
.
EAe
Distribution
Qualifier
248 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Erysimum parnassi (Boiss. & Heldr.) Hausskn.
Brassicaceae
Cheirantus parnassi Boiss. & Heldr.
Erysimum boryanum Boiss. & Spruner subsp. parnassi (Boiss. & Heldr.)
Nyman
Erysimum pusillum Bory & Chaub. subsp. parnassi (Boiss. & Heldr.)
Hayek
AFE 10:61 FAe :209 FE 1:331 FHe 2:136 MCL 3:113 MFG 1:243
L/261 L/323a L/382 L/512 L/2013
Erysimum pectinatum Bory & Chaub.
Brassicaceae
AFE 10:79 FE 1:334 FHe 2:150 MCL 3:113 MFG 1:246
L/607 L/1963
Erysimum pusillum Bory & Chaub.
Brassicaceae
Erysimum pusillum Bory & Chaub. subsp. trichophyllum (Halácsy)
Hayek
Erysimum trichophyllum Halácsy
AFE 10:61 FAe :209 FE 1:331 FHe 2:133 MCL 3:111 MFG 1:242
L/121 L/261 L/382 L/607 L/1846
Erysimum raulinii Boiss.
Brassicaceae
AFE 10:64 EFK :113 FCA :83 FE 1:331 FHe 2:150 MCL 3:113 MFG 1:242
L/1102 L/1111 L/1138 L/1766
Erysimum rechingeri Jav.
Brassicaceae
Erysimum pusillum Bory & Chaub. subsp. rechingeri (Jáv.) P.W.Ball
AFE 10:61 FAe :209 FE 1:331 FHe 2:136 MCL 3:113
L/111 L/261
Erysimum senoneri (Heldr. & Sart.) Wettst.
Brassicaceae
Cheiranthus cheiri auct non L.
Cheiranthus senoneri Helsr. & Sart.
AFE 10:52 FE 1:329 FHe 2:140 FT 10:57 MCL 3:109
L/129 L/169 L/443 L/539 L/783 L/938
649
650
651
652
653
654
+
.
.
.
.
.
.
.
+
+
+
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
+
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+
.
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+
.
.
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.
+
.
.
.
+
.
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 249
Euphorbia rechingeri Greuter
Euphorbiaceae
Euphorbia myrsinites L. subsp. rechingeri (Greuter) Aldén
Euphorbia myrsinites sensu auct. kret. Rechinger 1943
EFK :183 FCA :90 FCSI :47 FE 2:221 MCL 3:209 MFG 1:570
L/255 L/408 L/518 L/554 L/1111
Euphorbia sultan-hassei Strid
Euphorbiaceae
Euphorbia bivonae sensu Turland 1989, non Steud
Euphorbia myrsinites sensu Rechinger 1944
EFK :182 FCA :90 FCSI :47
L/265 L/336 L/1111 L/1583
661
662
Euphorbia deflexa Sm.
Euphorbiaceae
FCA :87 FE 2:223 MCL 3:210 MFG 1:570
L/121 L/159 L/477 L/499 L/512 L/530 L/587 L/607
L/895 L/1111 L/1246 L/1871 L/1972 L/1973 L/2013 L/2200
659
Euphorbia orphanidis Boiss.
Euphorbiaceae
FE 2:224 MCL 3:218 MFG 1:571
L/512
Euphorbia anthula Lavrent. & Papan.
Euphorbiaceae
MCL 3:206
L/409 L/1176
658
660
Erysimum senoneri (Heldr. & Sart.) Wettst. subsp. senoneri
Brassicaceae
AFE 10:52 FE 1:329 FHe 2:141 FT 10:57 MCL 3:109
L/128 L/129 L/196 L/443 L/915 L/938 L/1871 L/2181
657
!
Erysimum senoneri (Heldr. & Sart.) Wettst. subsp. icaricum
Snogerup
Brassicaceae
AFE 10:52 FE 1:329 FHe 2:141 FT 10:57 MCL 3:109 RDB :264
L/129 L/539 L/540 L/1167
656
Taxon
Erysimum senoneri (Heldr. & Sart.) Wettst. subsp. amorginum
Snogerup
Brassicaceae
AFE 10:52 FE 1:329 FHe 2:141 FT 10:57 MCL 3:109 RDB :262
L/129 L/404b L/1167
Taxon
Qualifiers
655
Taxon
Number
+
+
.
.
.
.
.
.
+
.
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.
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.
.
StE
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Pe
.
.
.
+
.
+
.
.
WAe
.
.
.
.
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+
.
.
.
.
.
.
SPi
.
.
.
.
IoI
.
.
.
+
.
.
.
.
NPi
.
.
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+
.
.
.
.
EC
.
.
.
+
.
.
.
.
NC
.
.
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+
+
.
.
.
NE
.
.
.
+
.
.
.
.
NAe
.
.
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+
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+
Kik
+
+
.
+
.
.
.
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KK
.
.
.
.
.
.
+
.
EAe
Distribution
Qualifier
250 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Festuca cyllenica Boiss. & Heldr. subsp. cyllenica
Poaceae
FE 5:137 FT 9:411 MFG 2:754
L/607
?
?
?
?
?
667
668
669
670
671
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Festuca jeanpertii (St-Yves) Markgr. subsp. achaica (Markgr.-Dann.)
Markgr.-Dann.
Poaceae
FCA :170 FE 5:144 FT 9:425 MFG 2:759
L/607 L/1091 L/1830
Festuca grandiaristata Markgr.-Dann.
Poaceae
Festuca hirtovaginata (Acht.) Markgr.-Dann. (included in taxon)
Festuca thracica Markgr.-Dann. (included in taxon)
FE 5:149 MFG 2:760
L/269 L/528
Festuca cyllenica Boiss. & Heldr. subsp. thasia Markgr.-Dann.
Poaceae
FE 5:137 FT 9:411 MFG 2:754
Festuca cyllenica Boiss. & Heldr. subsp. pangaei Markgr.-Dann.
Poaceae
FE 5:137 FT 9:411 MFG 2:754
L/1107
Ferulago thyrsiflora (Sm.) Koch
Apiaceae
EFK :217 FCA :151 FCSI :86 FE 2:359
L/1111
Ferulago serpentinica Rech.f.
Apiaceae
FE 2:360
L/323a L/1871 L/1871 L/1901
665
666
Ferulago sartorii Boiss.
Apiaceae
Ferulago serpentinica Rech.f.
FAe :412 FE 2:360 FGC 1:645
L/169 L/323a L/2181 L/2181
664
?
Evax perpusilla Boiss. & Heldr.
Asteraceae
FE 4:125 FGC 2:38 FGS 2:49
663
.
.
.
+
.
.
.
.
+
.
.
+
+
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+
+
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+
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+
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+
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+
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+
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.
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+
+
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+
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+
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.
.
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.
.
.
.
.
.
*
PATTERNS OF PLANT ENDEMISM IN GREECE 251
Festuca pseudosupina Vetter
Poaceae
FT 9:438 MFG 2:760 RDB :268
L/269 L/1957
Festuca rubra L. subsp. thessalica Markgr.-Dann.
Poaceae
FE 5:141 MFG 2:752
L/565 L/1211 L/2201
Festuca stygia H. Scholz & Strid
Poaceae
L/301
678
679
680
Festuca polita (Halácsy) Tzvelev var. euboica Markgr.-Dann.
Poaceae
Festuca callieri sensu Krause 1956
MFG 2:761
L/724
677
?
Festuca polita (Halácsy) Tzvelev subsp. cretica (Markgr.-Dann.)
Foggi & H. Scholz
Poaceae
Festuca polita (Halácsy) Tzvelev var. cretica Markgr.-Dann.
EFK :378 FCA :170 FCSII 0: MFG 2:761
L/430 L/518 L/724 L/1939
676
Festuca pindica (Markgr.-Dann.) Markgr.-Dann.
Poaceae
FE 5:136 MFG 2:753
L/269
675
?
Festuca oviniformis Vetter
Poaceae
FE 5:148
L/269
674
Festuca macedonica Vetter
Poaceae
FE 5:149 MFG 2:760
L/269 L/528 L/1107 L/1832d L/2201
Taxon
Festuca olympica Vetter
Poaceae
FE 5:143 MFG 2:757
L/269 L/512 L/528
?
Taxon
Qualifiers
673
672
Taxon
Number
.
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StE
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Pe
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+
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WAe
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IoI
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SPi
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+
.
.
.
NPi
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+
.
.
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EC
.
+
.
.
.
.
.
+
+
NC
.
+
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+
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+
NE
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NAe
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Kik
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KK
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EAe
*
Distribution
Qualifier
252 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Festuca valida (Uechtr.) Penzes subsp. leilaensis Markgr.-Dann.
Poaceae
FE 5:134
Festuca violacea Schleicher ex Gaudin subsp. handelii
Markgr.-Dann.
Poaceae
FE 5:139
L/269 L/528
Fibigia lunarioides (Willd.) Sm.
Brassicaceae
Alyssum lunarioides Willd.
Farsetia lunaroides (Willd.) R.Br.
Fibigia lunaroides (Willd.) Sm. (sphalm.)
Pevalekia lunarioides (Willd.) Trinajstic
AFE 11:60 EFK :119 FCA :83 FE 1:369 FHe 2:225 FT 1:358 MCL 3:116
L/145 L/169 L/197 L/206 L/268 L/537 L/831 L/913
L/1111 L/1710
Filago aegaea Wagenitz subsp. aegaea
Asteraceae
EFK :307 FCA :65 FCY 2:880 FE 4:122
L/255 L/268 L/518 L/1111 L/2093 L/2181
Filago cretensis Gand.
Asteraceae
Filago spathulata C. Presl var. syria Hal.
EFK :308 FAe :610 FCA :66 FCSII : FE 4:122 FT 5:106
L/128 L/150 L/163 L/177 L/443 L/531 L/540 L/762
L/776 L/783 L/869 L/913 L/928 L/1049 L/1111
L/1627 L/1713 L/1917 L/1957 L/2089 L/2093 L/2181 L/2227
Filago cretensis Gand. subsp. cretensis
Asteraceae
Evax cretensis (Gand.) Gand.
EFK :308 FCA :66 FE 4:122
L/163 L/201 L/303 L/423 L/443 L/531 L/539 L/762
L/778 L/913 L/928 L/1111 L/1713 L/1917 L/1957 L/2181
681
682
683
684
685
686
.
.
.
.
+
+
+
.
.
+
.
.
+
+
.
.
.
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+
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+
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+
+
+
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+
+
+
+
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+
+
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 253
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Fritillaria conica Boiss.
Liliaceae
FE 5:33 FGC 3:221 RDB :270
L/138 L/494 L/607 L/1073 L/1158 L/1229 L/1963
Fritillaria davisii Turrill
Liliaceae
FE 5:32
L/607 L/1073 L/1229 L/1963
Fritillaria epirotica Turrill ex Rix
Liliaceae
FE 5:32 MFG 2:676 RDB :272
L/404a L/961 L/1073 L/1229
Fritillaria erhartii Boiss. & Orph.
Liliaceae
Fritillaria erhartii Boiss. & Orph. var. octavii Gand.
Fritillaria regis georgii Heldr. & Holzm. ex Nyman
Fritillaria macrandra Baker
FAe :721 FE 5:34 MFG 2:674
L/111 L/323b L/685 L/1073 L/1229 L/1871
Fritillaria euboeica Rix
Liliaceae
Fritillaria rixii Zaharof
Fritillaria sibthorpiana auct. balkan., non (Sm.) Baker
FE 5:33 MFG 2:674 RDB :274
L/494 L/965 L/1073 L/1158 L/1227 L/1229 L/1871
Fritillaria graeca Boiss. & Spruner
Liliaceae
Fritillaria graeca Boiss. & Spruner subsp. graeca var. graeca
FAe :721 FCA :183 FCSI :108 FE 5:32 FGC 3:220 MFG 2:677
L/160 L/512 L/516 L/530 L/607 L/856 L/1073 L/1112
L/1126 L/1228 L/1229 L/1871 L/1963 L/2013
Fritillaria graeca Boiss. & Spruner var. graeca
Liliaceae
Fritillaria graeca Boiss. & Spruner var. unicolor Halácsy
Fritillaria guicciardii Heldr. & Sart var. zahnii Heldr.
FE 5:32 FGC 3:220 MFG 2:679
L/1073 L/1229 L/1963 L/2013
688
689
690
691
692
693
694
Taxon
Filago cretensis Gand. subsp. cycladum Wagenitz
Asteraceae
EFK :308 FCA :66 FCSII : FE 4:122
L/197 L/201 L/531 L/762 L/776 L/913 L/1049 L/1111 L/1627 L/1710 L/2089 L/2093 L/2181
Taxon
Qualifiers
687
Taxon
Number
.
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+
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+
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.
StE
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+
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Pe
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SPi
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IoI
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+
+
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.
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WAe
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+
.
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.
NPi
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EC
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NC
+
+
+
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NE
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NAe
+
+
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+
.
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+
Kik
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+
KK
.
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.
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+
EAe
Distribution
Qualifier
254 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Fritillaria graeca Boiss. & Spruner var. guicciardii (Heldr. & Sart.)
Boiss.
Liliaceae
Fritillaria graeca Boiss. & Spruner subsp. guicciardii (Heldr. & Sart.)
Zaharof
Fritillaria guicciardii Heldr. & Sart.
FE 5:32 FGC 3:220 MFG 2:679
L/160 L/512 L/1073 L/1229 L/1963 L/2013
Fritillaria messanensis Raf. subsp. sphaciotica (Gand.) Kamari &
Phitos
Liliaceae
Fritillaria graeca auct. Fl. Graec.
Fritillaria sphaciotica Gand.
FCSII : FE 5:32
L/1914
Fritillaria mutabilis Kamari
Liliaceae
Fritillaria graeca var. thessala auct., non (Boiss.) Rix
Fritillaria graeca var. thessala auct., non Boiss.
MFG 2:679
L/121 L/1073 L/1846 L/2013 L/2200
Fritillaria obliqua Ker-Gawl.
Liliaceae
FE 5:33
L/138 L/1073 L/1229
Fritillaria obliqua Ker-Gawl. subsp. obliqua
Liliaceae
FE 5:33 RDB :276
L/138 L/1071 L/1073 L/1229 L/1871
Fritillaria obliqua Ker-Gawl. subsp. tuntasia (Heldr. ex Halácsy)
Kamari
Liliaceae
Fritillaria tuntasia Heldr. ex Halácsy
L/138 L/177 L/1073 L/1229
Fritillaria pelinaea Kamari
Liliaceae
FT 11:244 RDB :278
L/494 L/1957
695
696
697
698
699
700
701
+
.
.
.
.
+
.
.
+
.
.
+
+
+
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
.
.
+
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+
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+
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+
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+
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+
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+
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.
.
.
.
.
.
+
.
+
.
.
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 255
Fritillaria sporadum Kamari
Liliaceae
RDB :286
L/443 L/1072
Fritillaria theophrasti Kamari & Phitos
Liliaceae
Fritillaria pontica Wahlenb. var. substipelata P.Candargy
Fritillaria pontica sensu auct. fl. graec. non Wahlenb.
L/1832f
Fritillaria thessala (Boiss.) Kamari
Liliaceae
Fritillaria graeca Boiss. & Spruner subsp. thessala (Boiss.) Rix
Fritillaria graeca Boiss. & Spruner var. thessala Boiss.
Fritillaria thessalica Spruner
FE 5:32 MFG 2:680
L/513 L/1073 L/1112 L/1229 L/1832d L/1846 L/2013 L/2200
!
!
705
706
707
708
709
Fritillaria spetsiotica Kamari
Liliaceae
RDB :284
L/1073
704
Fritillaria thessala (Boiss.) Kamari subsp. reiseri Kamari
Liliaceae
Fritillaria ionica Halácsy p.p.
FE 5:32 MFG 2:680 RDB :288
L/1073
Fritillaria thessala (Boiss.) Kamari subsp. ionica (Halácsy) Kamari
Liliaceae
Fritillaria graeca Wahlenb var. ionica (Halácsy) Zaharof
Fritillaria ionica Halácsy p.p.
Fritillaria pontica Wahlenb var. ionica (Halácsy) Turrill
MFG 2:682
L/1073 L/1832d
Fritillaria rhodocanakis Orph. ex Baker
Liliaceae
FE 5:32 RDB :282
L/138 L/1073 L/1126 L/1229 L/1963
703
Taxon
Fritillaria rhodia Hansen
Liliaceae
FT 8:295 RDB :280
L/197 L/494 L/1037 L/1073 L/1111
Taxon
Qualifiers
702
Taxon
Number
?
.
.
+
.
+
.
.
.
.
.
.
.
StE
+
+
.
Pe
.
.
.
.
+
.
.
.
WAe
.
+
+
.
.
.
.
.
IoI
.
+
+
.
.
.
.
.
SPi
.
+
+
.
.
.
.
.
NPi
.
.
+
.
.
.
.
.
EC
.
+
+
.
.
.
.
.
NC
.
.
.
.
.
.
.
.
NE
.
.
.
.
.
.
.
.
NAe
.
.
.
.
.
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.
.
Kik
.
.
.
.
.
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.
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KK
.
.
.
+
.
.
.
+
EAe
Distribution
Qualifier
256 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Galanthus ikariae Baker
Amaryllidaceae
Galanthus latifolius Rupr.
FE 5:78 FT 8:365 RDB :290
L/296 L/539 L/869 L/1069 L/1663 L/1807 L/2181 L/2181 L/2237
Galanthus reginae-olgae Orph. subsp. reginae-olgae
Amaryllidaceae
FE 5:77 FT 8:367 RDB :294
L/296 L/607 L/1069 L/1963 L/1964
Galium absurdum Krendl
Rubiaceae
MFG 2:318
L/790 L/2200
Galium advenum Krendl
Rubiaceae
MFG 2:319
L/790
Galium amorginum Halácsy
Rubiaceae
EFK :240 FCA :138 FCSI :79 FE 4:26 MFG 2:321
L/255 L/790 L/869 L/1111
Galium avascense Krendl
Rubiaceae
L/790
712
713
714
715
716
717
Fumana paphlagonica Bornm. & Janchen subsp. alpina (Janchen)
Greuter
Cistaceae
Fumana procumbens sensu Bald. 1916
Fumana vulgaris f. alpina Janchen
Helianthemum ericoides sensu Gandoger 1916
EFK :199 FAe :253 FCA :56 FCSI :25 FCSII : FT 1:520 MCL 1:317 MFG 1:644
L/255 L/607 L/1111
711
?
Fritillaria thessala (Boiss.) Kamari subsp. thessala
Liliaceae
MFG 2:681
L/121 L/1073 L/1244 L/2013
710
.
+
+
.
.
.
.
.
.
.
.
+
.
.
+
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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+
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+
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+
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+
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+
+
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+
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+
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+
+
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+
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+
.
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+
.
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+
.
.
.
.
.
.
+
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 257
Galium capreum Krendl
Rubiaceae
MFG 2:319
L/790
Galium circae Krendl
Rubiaceae
MFG 2:314
L/790 L/2013
Galium citraceum Boiss.
Rubiaceae
Galium aureum Vis. var. oblongifolium Boiss.
Galium firmum Tausch var. citraceum (Boiss.) Boiss. & Heldr.
Galium firmum Tausch var. euboeum Tocl & Rochl.
EFK :239 FCA :138 MFG 2:313
L/121 L/790 L/895 L/1111 L/1871 L/1963 L/2013 L/2200
Galium conforme Krendl
Rubiaceae
L/790 L/1957
Galium cyllenium Boiss. & Heldr.
Rubiaceae
Galium incanum Sm. subsp. cyllenium (Boiss. & Heldr.) Maire &
Petitm.
FE 4:29 MFG 2:323
L/160 L/395 L/512 L/607 L/1963
Galium decorum Krendl
Rubiaceae
MFG 2:320
L/790
Galium exaltatum Krendl
Rubiaceae
L/565 L/566 L/790 L/1975
Galium extensum Krendl
Rubiaceae
EFK :240 FCA :138 MFG 2:311
L/790 L/1111
719
720
721
722
723
724
725
726
Taxon
Galium capitatum Bory & Chaub.
Rubiaceae
FAe :577 FE 4:35 FGC 1:725
L/323b L/607 L/765 L/1052 L/1111 L/1846 L/1871 L/1878 L/1963 L/1973
Taxon
Qualifiers
718
Taxon
Number
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
+
.
.
+
WAe
+
+
.
+
StE
+
.
.
+
Pe
.
.
.
.
.
.
.
.
.
.
.
.
+
+
.
.
.
.
SPi
IoI
.
.
.
.
.
.
.
.
.
NPi
.
+
.
.
.
.
.
.
+
EC
.
+
.
.
.
.
.
.
.
NC
.
+
.
.
.
.
.
.
?
NE
.
.
+
.
.
.
.
+
.
NAe
.
.
.
.
+
.
.
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.
Kik
+
.
.
.
.
+
.
.
.
KK
.
.
.
.
+
.
.
.
.
EAe
Distribution
Qualifier
258 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Galium fruticosum Willd.
Rubiaceae
EFK :238 FCA :138 FE 4:20
L/1111
Galium graecum L. subsp. pseudocanum Ehrend.
Rubiaceae
EFK :239 FCA :139 FE 4:34
L/268 L/1111
Galium incanum Sm. subsp. creticum Ehrend.
Rubiaceae
EFK :239 FAe :575 FCA :139 FCSI :69 FE 4:29 FGC 1:717 MFG 2:322
L/1111
Galium incrassatum Halácsy
Rubiaceae
Galium capitatum Boiss. sensu auct. Kret.
Galium capitatum Boiss. var. hispidulum Raulin
EFK :238 FAe :577 FCA :139 FCSI :79 FCSII : FE 4:35
L/751 L/1111
Galium incurvum Sm.
Rubiaceae
Galium amorginum sensu Greuter 1973, non Halácsy 1901
EFK : FCA :139 FCSI :79 FE 4:26 MFG 2:321
L/255 L/1111
Galium insulare Krendl
Rubiaceae
FAe :572 MFG 2:312
L/790 L/1349 L/1973 L/1975
Galium malickyi Krendl
Rubiaceae
L/790
Galium melanantherum Boiss.
Rubiaceae
FE 4:26
L/530 L/790 L/1713 L/1871 L/1963 L/1981 L/2181
727
728
729
730
731
732
733
734
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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+
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+
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+
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+
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+
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+
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+
+
+
+
+
.
.
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PATTERNS OF PLANT ENDEMISM IN GREECE 259
Galium monasterium Krendl
Rubiaceae
MFG 2:316
L/790
Galium ophiolithicum Krendl
Rubiaceae
MFG 2:317
L/790
Galium pastorale Krendl
Rubiaceae
L/790
Galium peloponnesiacum Ehrend. & Krendl
Rubiaceae
EFK :240 FCA :139 FE 4:26 MFG 2:314
L/607 L/790 L/1111 L/1123 L/1963
Galium pisoderium Krendl
Rubiaceae
MFG 2:311
L/790 L/2201
Galium reiseri Halácsy
Rubiaceae
FE 4:24
L/196 L/443 L/790 L/915 L/928
Galium sacrorum Krendl
Rubiaceae
MFG 2:316
L/601 L/790
Galium samium Krendl
Rubiaceae
FAe :572 FT 10:216 FT 11:218
L/539 L/540 L/790 L/796
Galium samothracicum Rech.f.
Rubiaceae
FCA :139 FE 4:24 FT 10:216 MFG 2:313
L/321 L/790 L/796 L/895 L/1111 L/1349 L/1871 L/1871 L/1957 L/1973
736
737
738
739
740
741
742
743
744
Taxon
Galium monachinii Boiss. & Heldr.
Rubiaceae
EFK :237 FAe :577 FCA :139 FE 4:34 MFG 2:324
L/122 L/273 L/869 L/1111
Taxon
Qualifiers
735
Taxon
Number
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
StE
+
.
.
.
.
Pe
+
.
.
+
.
.
.
.
.
+
WAe
.
.
.
.
.
+
.
.
.
.
IoI
.
.
.
.
+
.
.
.
+
.
SPi
.
.
+
.
+
.
.
+
+
.
NPi
.
.
.
.
.
.
.
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.
.
EC
.
.
.
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+
.
.
.
.
.
NC
.
.
.
.
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.
.
NE
+
.
.
.
.
.
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.
.
.
NAe
.
.
.
.
.
.
.
.
.
+
Kik
+
.
.
.
.
+
.
.
.
+
KK
+
+
.
.
.
.
+
.
.
.
EAe
Distribution
Qualifier
260 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
754
Geocaryum bornmuelleri (Wolff) Engstr.
Apiaceae
Biasolettia bornmuelleri Wollf.
RDB :296
L/365 L/406 L/1267b
Genista sakellariadis Boiss. & Orph.
Fabaceae
FE 2:96 FGC 1:331 MCL 4:100 MFG 1:455
L/159
?
753
Genista halácsyi Heldr.
Fabaceae
FE 2:97 MCL 4:100 MFG 1:456
L/607 L/1963
750
Genista parnassica Halácsy
Fabaceae
Genista salzamanii subsp. parnassica (Halácsy) Ponert
Genista subsericans (Bornm.) Rech.f.
FE 2:97 FT 3:28 MCL 4:93 MFG 1:456
L/796 L/895 L/975 L/1252
Galium violaceum Krendl
Rubiaceae
MFG 2:319
L/790 L/1052 L/1111 L/1830 L/1963
749
752
Galium thymifolium Boiss. & Heldr.
Rubiaceae
FE 4:22 MFG 2:305
L/160 L/512 L/607 L/1871 L/1963 L/2013
748
Genista millii Heldr. ex Boiss.
Fabaceae
FE 2:97 MCL 4:100 MFG 1:456
L/121 L/391 L/1871 L/2013 L/2200
Galium thracicum Krendl
Rubiaceae
L/790
747
751
Galium thasium Stoj. & Kit.
Rubiaceae
L/1267b
746
?
Galium taygeteum Krendl
Rubiaceae
MFG 2:318
L/790 L/1963
745
.
.
+
.
.
+
.
.
.
.
+
.
.
.
+
+
+
.
.
+
.
.
.
+
.
.
+
.
.
.
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.
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+
.
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.
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.
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.
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+
.
.
+
.
+
.
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+
+
.
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+
.
.
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.
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PATTERNS OF PLANT ENDEMISM IN GREECE 261
Geocaryum parnassicum (Boiss. & Heldr.) Engstr.
Apiaceae
Butinia parnassica (Boiss. & Heldr.) Heldr. ex Nyman
Biasolettia parnassica (Boiss. & Heldr.) Nyman
Freyera parnassica Boiss. & Heldr.
Huetia cynapioides (Guss.) P.W. Ball subsp. macrocarpa (Boiss. &
Spruner) P.W. Ball p.p.
FAe :406 FE 2:330 MFG 1:674
L/159 L/395 L/406 L/512 L/530 L/587 L/1871 L/1963 L/1973
Geocaryum peloponnesiacum Engstr.
Apiaceae
MFG 1:677
L/395 L/406 L/1123 L/1963
Geocaryum pumilum (Sm.) Nyman
Apiaceae
Bunium pumilum Sm.
Freyera pumila (Sm.) Boiss.
Huetia pumila (Sm.) Boiss.
FE 2:330 MFG 1:676
L/406 L/512
758
759
760
Geocaryum divaricatum (Boiss. & Orph.) Engstr.
Apiaceae
Freyera divaricata Boiss. & Orph.
Geocaryum cynapioides (Guss.) P.W. Ball subsp. divaricatum (Boiss. &
Orph.) P.W. Ball
Huetia cynapioides (Guss.) P.W. Ball subsp. divaricata (Boiss. & Orph.)
P.W. Ball
FE 2:330 RDB :298
L/395 L/406 L/512 L/607 L/608 L/1176 L/1871 L/1963
Geocaryum creticum (Boiss. & Heldr.) Engstr.
Apiaceae
Butinia cretica Boiss. & Heldr.
Freyera cretica (Boiss. & Heldr.) Boiss. & Heldr.
Huetia cretica (Boiss. & Heldr.) P.W. Ball
FCA :151 FE 2:330 MFG 1:676
Taxon
Geocaryum euboicum (Rech.f.) Engstr.
Apiaceae
RDB :298
L/323a L/406 L/1176 L/1871
?
Taxon
Qualifiers
757
756
755
Taxon
Number
.
+
.
+
.
.
.
StE
+
+
.
+
.
Pe
.
.
+
+
.
.
WAe
.
.
.
.
.
+
.
.
.
SPi
.
.
.
IoI
.
.
.
.
.
.
NPi
.
.
.
.
.
.
EC
.
.
.
.
.
.
NC
.
.
.
.
.
.
NE
.
.
.
.
.
.
NAe
.
.
.
.
.
.
Kik
.
.
.
.
.
+
KK
.
.
.
.
.
.
EAe
Distribution
Qualifier
262 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Geranium thessalum Franzén
Geraniaceae
MCL 3:257 MFG 1:541
L/1063 L/1097
Globularia stygia Orph. ex Boiss.
Plantaginaceae
Globularia cordifolia L. subsp. stygia (Orph.) Wettst.
FE 3:283 MCL 3:262 MFG 2:259 RDB :300
L/138 L/160 L/395 L/512 L/607 L/1963
Gnaphalium roeseri Boiss. & Heldr. subsp. roeseri
Asteraceae
FE 4:127 MFG 2:413
L/607 L/1963
Goniolimon heldreichii Halácsy
Plumbaginaceae
Statice heldreichii Halácsy
FE 3:50 FGC 3:15 MCL 4:320
Goniolimon sartorii Boiss.
Plumbaginaceae
FAe :427 FE 3:51 FGC 3:14 MCL 4:320
L/776 L/1260 L/1871 L/1871
Gymnospermium altaicum (Pall.) Spach subsp. peloponnesiacum
Phitos
Berberidaceae
Gymnospermium altaicum (Pall.) Spach subsp. odessanum sensu auct.
Grec.
Leontice odessana (DC.) G. Don
AFE 9:22 FE 1:294 FHe 2:81 MCL 1:63 RDB :302
L/418 L/1133 L/1963
Gypsophila nana Bory & Chaub.
Caryophyllaceae
Gypsophila achaia Bornm. (included in taxon)
Gypsophila nana Bory & Chaub. var. glabrifolia Halácsy
AFE 7:122 EFK :90 FAe :150 FCA :47 FE 1:220 FHe 1:324 MCL 1:209 MFG 1:171
L/160 L/499 L/512 L/518 L/530 L/554 L/607 L/1963
Halimium voldii Kit Tan, Perdetzoglou & Raus
Cistaceae
L/1772 L/1963
761
762
763
764
765
766
767
768
.
.
.
.
.
.
+
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
+
+
.
.
+
+
.
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.
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+
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+
+
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.
.
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+
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.
.
.
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+
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 263
Helichrysum doerfleri Rech.f.
Asteraceae
EFK :308 FAe :613 FCA :67 FCSI :32 FE 4:129 MFG 2:415
L/518 L/522 L/554
Helichrysum heldreichii Boiss.
Asteraceae
EFK :309 FCA :67 FCSI :32 FE 4:130 RDB :306
L/518 L/519 L/1583
Helichrysum sibthorpii Rouy
Asteraceae
Helichrysum virgineum (Sm.) Griseb.
FE 4:129 MFG 2:415 RDB :308
L/404a L/1972 L/1973
Helichrysum taenari Rothm.
Asteraceae
FE 4:129 MFG 2:415 RDB :310
L/485 L/607 L/824 L/1963
Helictotrichon agropyroides (Boiss.) Henrard
Poaceae
Avenula agropyroides (Boiss.) Holub
Avena agropyroides Boiss.
Avena sibthorpii Nyman
Avenastrum cycladum Rech.f. & Scheffer
Avenula cycladum (Rech.f. & Scheffer) Greuter
Avenula peloponnesiaca J. Holub
Helictotrichon cycladum (Rechf. & Scheffer) Rech.f.
EFK :394 FAe :791 FCA :164 FCSI :99 FE 5:215 FGC 3:370 MFG 2:800
L/111 L/443 L/530 L/592 L/607 L/613 L/767 L/869
L/1830 L/1871
771
772
773
774
775
Helianthemum hymettium Boiss. & Heldr.
Cistaceae
EFK :198 FAe :251 FCA :56 FCSI :25 FE 2:290 MCL 1:322 MFG 1:643
L/518 L/530 L/543 L/607 L/707 L/856 L/1112 L/1844 L/1963 L/1981
Taxon
Helichrysum amorginum Boiss. & Orph.
Asteraceae
FE 4:129 MFG 2:415 RDB :304
L/169 L/206
Taxon
Qualifiers
770
769
Taxon
Number
+
+
+
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
SPi
.
.
.
.
.
.
.
.
IoI
WAe
.
.
+
StE
.
+
Pe
.
.
.
.
.
.
.
NPi
.
.
.
.
.
.
.
EC
.
.
.
.
.
.
.
NC
.
.
+
.
.
.
.
NE
.
.
.
.
.
.
.
NAe
+
.
.
.
.
+
+
Kik
+
.
.
+
+
.
+
KK
.
.
.
.
.
.
.
EAe
Distribution
Qualifier
264 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Heptaptera colladonioides Marg. & Reut.
Apiaceae
Colladonia colladonioides (Marg. & Reut.) Halácsy
Colladonia heptaptera Boiss.
FE 2:344 FGC 1:661
L/607 L/765 L/1123 L/1244 L/1963
Herniaria degenii (Hermann) Chaudhri
Caryophyllaceae
Herniaria nigrimontium Hermann var. degenii Hermann
FE 1:184 FE 6:152 FHe 1:232 MCL 1:211 MFG 1:133
Herniaria parnassica Boiss. subsp. cretica Chaudhri
Caryophyllaceae
Herniaria parnassica Boiss. var. hirtella Chaudhri
AFE 6:147 EFK :83 FAe :129 FCA :47 FCSI :19 FCSII : FE 1:183 FHe 1:229 MCL 1:213 MFG 1:132
L/518 L/522 L/554 L/1766 L/1963
Hesperis laciniata All. subsp. secundiflora (Boiss. & Spruner)
Breistr.
Brassicaceae
Hesperis glutinosa Vis. subsp. secundiflora (Boiss.) F. Dvořák
Hesperis glutinosa Vis. var. secundiflora (Boiss. & Spruner) Borbas
Hesperis secundiflora Boiss & Spruner
AFE 10:92 FE 1:336 FGC 1:72 FHe 2:153 MCL 3:119 MFG 1:248
L/310a L/323a L/530 L/532 L/1871 L/1963
Hesperis theophrasti Borbás subsp. rechingeri (F. Dvořák) Kit Tan
& J. Suda
Brassicaceae
Hesperis rechingeri F. Dvořák
AFE 10:98 FE 1:337 FHe 2:155 MCL 3:121 MFG 1:248
778
779
780
781
782
Helictotrichon convolutum (C. Presl) Henrard subsp. heldreichii
(Parl.) Gervais
Poaceae
FE 5:209 MFG 2:798
L/121 L/607 L/2013 L/2200 L/2237
Heliotropium halacsyi Riedl
Boraginaceae
Heliotropium bocconei sensu Hayek p.p., non Guss.
FE 3:85 MCL 1:90
L/121 L/798 L/971 L/1123 L/1156 L/1846 L/2200
?
777
776
.
.
.
.
.
+
.
+
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
.
+
.
.
+
+
.
+
.
+
+
+
.
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.
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.
.
+
+
.
.
.
.
.
.
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.
.
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.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
*
PATTERNS OF PLANT ENDEMISM IN GREECE 265
Hieracium dasycraspedum Buttler
Asteraceae
MFG 2:630
L/601
Hieracium epirense (Zahn) Buttler
Asteraceae
Hieracium wiesbaurianum Uechtr. ex Baenitz subsp. retroversidens
Zahn
Hieracium wiesbaurianum Uechtr. ex Baenitz subsp. kyllenense Zahn
Hieracium wiesbaurianum Uechtr. ex Baenitz subsp. epirense Zahn
MFG 2:618
L/121 L/1963 L/2200
Hieracium ferdinandi-coburgi J. Wagner & Zahn
Asteraceae
MFG 2:627
L/1019
Hieracium gaudryi Boiss.
Asteraceae
Hieracium pannosum Boiss. subsp. gaudryi (Boiss. & Orph.) Nyman
FE 4:390 MFG 2:621
L/1107
Hieracium gracilifurcum Zahn
Asteraceae
Hieracium sartorianum sensu Boiss. 1875; Halácsy 1902 p.p.
FE 4:391 MFG 2:636
Hieracium graecum Boiss. & Heldr.
Asteraceae
Hieracium graecum f. flocciceps Zahn
MFG 2:630
L/895 L/1963
786
787
788
789
790
791
!
Hieracium cylleneum Halácsy
Asteraceae
FGS 2:69 MFG 2:597 MFG 2:639
L/1963
785
Hieracium chalasinense Zahn
Asteraceae
FE 4:408 MFG 2:597
L/607
?
784
Taxon
Hieracium bonaquae Buttler & Lippert
Asteraceae
MFG 2:608
Taxon
Qualifiers
783
Taxon
Number
.
.
+
+
.
+
.
+
.
.
.
.
StE
.
+
.
+
+
.
Pe
.
.
.
.
.
.
.
.
.
.
.
.
.
.
IoI
.
.
.
.
WAe
.
.
.
+
+
.
.
.
.
SPi
+
.
.
+
+
+
.
.
.
NPi
.
.
.
.
.
.
.
.
.
EC
.
+
.
+
+
.
.
.
+
NC
.
.
.
.
.
.
.
.
.
NE
.
.
.
.
.
.
.
.
.
NAe
.
.
.
.
.
.
.
.
.
Kik
.
.
.
.
.
.
.
.
.
KK
.
.
.
.
.
.
.
.
.
EAe
Distribution
Qualifier
266 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Hieracium leucocomum Arvet-Touvet
Asteraceae
Hieracium sericophyllum Nejceff & Zahn subsp. eriocomum Zahn
Hieracium sericophyllum Nejceff & Zahn subsp. fiedleri Zahn
MFG 2:631
L/1963
Hieracium necopinum Buttler
Asteraceae
MFG 2:639
L/601
Hieracium pallidum Biv. subsp. creticum Zahn
Asteraceae
FCSII:
Hieracium psaridianum Zahn
Asteraceae
FE 4:408 MFG 2:597 PFPB 2:928
L/607 L/869
795
796
797
798
Hieracium sartorianum Boiss. & Heldr.
Asteraceae
Hieracium sartorianum Boiss. & Heldr. subsp. naegelianoides Zahn
FE 4:391 MFG 2:598 MFG 2:635
L/512 L/587 L/607 L/1107 L/1871 L/1871 L/1963
Hieracium leithneri (Heldr. & Sart. ex Boiss.) Zahn
Asteraceae
Hieracium leithneri (Heldr. & Sart. ex Boiss.) Zahn var. eurytanicum
Zahn
Hieracium sartorianum (Boiss. & Heldr.) var. leithneri Heldr. & Sart. ex
Boiss.
Hieracium sartorianum sensu auct. Rchb.f. (1860), Boiss (1875); Consp.
Fl. Graec; Zahn 1921 p.p.
Hieracium sericophyllum Nejceff & Zahn subsp. olenium Zahn
FCA :67 FE 4:390 FGC 2:242 MFG 2:633
L/121 L/2013 L/2200
794
799
Hieracium heldreichii Boiss.
Asteraceae
EFK :342 FCA :67 FCSI :32 FE 4:391 MFG 2:622
L/346 L/530 L/607 L/1963
793
?
Hieracium greuteri Gottschl.
Asteraceae
L/1923
792
+
+
.
.
+
+
+
+
+
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
+
.
.
.
.
.
.
+
.
.
.
.
.
+
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
+
.
.
.
.
.
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 267
.
+
.
Hymenonema laconicum Boiss. & Heldr.
Asteraceae
FE 4:305
L/607 L/1963
808
+
+
.
Hyacinthella atchleyi (A.K. Jackson & Turrill) Feinbrun
Asparagaceae
FE 5:46 RDB :314
L/530 L/1009 L/1116 L/1117 L/1173 L/1259a
.
.
807
.
.
Hordeum murinum L. subsp. setariurum H. Scholz & Raus
Poaceae
L/924 L/2237
805
Horstrissea dolinicola Greuter, Gerstberger & Egli
Apiaceae
EFK :211 FCA :151 FCSI :87 RDB :312
L/166 L/522
.
.
Hirtellina fruticosa (L.) Dittrich
Asteraceae
Centaurea fruticosa L.
Staehelina fruticosa (L.) L.
EFK :321 FCA :72 FCSI :32 FE 4:217 FT 5:595
L/196 L/1563
804
806
.
.
Himantoglossum × samariense C. Alibertis & A. Alibertis
Orchidaceae
EFK :428 FCA :187 FCSI :111 FT 8:508
L/255 L/526 L/557 L/611 L/791 L/808 L/1463 L/1816 L/2185
.
.
.
.
.
803
.
.
.
Hieracium trikalense Buttler
Asteraceae
MFG 2:626
.
.
.
WAe
802
+
StE
+
Pe
Hieracium szilyanum (J. Wagner & Zahn) Buttler
Asteraceae
Hieracium graecum Boiss. & Heldr. subsp. szilyanum J. Wagner &
Zahn
MFG 2:623
L/1019
Hieracium scapigerum Boiss
Asteraceae
FE 4:398 MFG 2:637
L/160 L/499 L/512 L/530 L/607 L/1963 L/2013
Taxon
801
?
!
800
!
Taxon
Qualifiers
Taxon
Number
.
.
.
.
.
.
.
.
.
IoI
.
.
.
.
.
.
+
.
.
SPi
.
.
.
.
.
.
+
.
.
NPi
.
.
.
+
.
.
.
.
.
EC
.
+
.
.
.
.
.
+
.
NC
.
.
.
.
.
.
.
.
.
NE
.
.
.
.
.
.
.
.
.
NAe
.
.
.
.
+
.
.
.
.
Kik
.
.
+
.
+
+
.
.
.
KK
.
.
.
.
+
.
.
.
.
EAe
Distribution
Qualifier
268 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Hypecoum procumbens L. subsp. fragrantissimum A. Dahl
Papaveraceae
AFE 9:60 FE 1:303 FHe 2:97 MCL 4:282
L/403
Hypericum aciferum (Greuter) N.K.B. Robson
Hypericaceae
Elodes acifera Greuter
EFK :194 FCA :93 FE 2:264 MCL 3:263 RDB :316
L/336 L/408 L/1159 L/1583
Hypericum amblycalyx Coust. & Gand.
Hypericaceae
EFK :193 FCA :93 FE 2:264 MCL 3:266
L/1583
Hypericum athoum Boiss. & Orph.
Hypericaceae
Hypericum montanum L. var. athoum Griseb.
Hypericum sanctum Degen
FE 2:265 MCL 3:264 MFG 1:602
L/895 L/1107 L/1267b L/1973
Hypericum cuisinii Barbey
Hypericaceae
FAe :264 FCA :93 FE 2:266 FT 2:387 MCL 3:265 MFG 1:602
L/539 L/540 L/796 L/1049 L/1957
Hypericum delphicum Boiss. & Heldr.
Hypericaceae
FAe :264 FE 2:265 MCL 3:265 MFG 1:602
L/323a L/1871 L/1973 L/2181
Hypericum elongatum Ledeb. subsp. tymphresteum (Boiss. &
Spruner) N.K.B. Robson
Hypericaceae
Hypericum hyssopifolium Chaix subsp. tymphresteum (Boiss. & Spruner)
Hayek
Hypericum tymphresteum Boiss. & Spruner
FE 2:265 FGC 1:281 MCL 3:268 MFG 1:596
L/121 L/1160 L/2200
809
810
811
812
813
814
815
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
?
.
+
.
.
.
.
.
.
+
.
.
.
.
+
.
.
.
.
+
.
.
+
.
+
+
.
.
.
+
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 269
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Hypericum fragile Heldr. & Sart. ex Boiss.
Hypericaceae
FAe :264 FE 2:265 MCL 3:266
L/323a L/1871
Hypericum jovis Greuter
Hypericaceae
EFK :193 FCA :94 MCL 3:266 RDB :318
L/253 L/1583
Hypericum kelleri Bald.
Hypericaceae
EFK :195 FAe :264 FCA :94 FCSI :50 FE 2:267 MCL 3:268 MFG 1:607 RDB :320
L/518 L/522 L/554 L/707 L/987 L/1766
Hypericum taygeteum Quézel & Contandr.
Hypericaceae
FE 2:265 MCL 3:273
L/607 L/1588 L/1963
Hypericum trichocaulon Boiss. & Heldr.
Hypericaceae
Hypericum repens Sm.
EFK :195 FCA :94 FE 2:267 MCL 3:274 MFG 1:607
L/336 L/1766
Hypochaeris tenuiflora (Boiss.) Boiss.
Asteraceae
Hypochoeris tenuiflora (Boiss.) Boiss.
Robertia tenuiflora Boiss.
EFK :335 FCA :68 FCSI :32 FE 4:309 MFG 2:525
L/1766
818
819
820
821
822
823
Hypericum empetrifolium Willd. subsp. oliganthum (Rech.f.)
I. Hagemann
Hypericaceae
Hypericum empetrifolium Willd. var. oliganthum Rech.f. (including
taxon)
EFK :193 FCA :94 FCSI :49 MFG 1:595
L/554 L/1036
Taxon
Hypericum empetrifolium Willd. subsp. tortuosum (Rech.f.)
I. Hagemann
Hypericaceae
Hypericum empetrifolium Willd. var. tortuosum Rech.f.
EFK :193 FCA :94 FCSI :49 FCSII : MFG 1:595
L/518 L/554 L/1036 L/1766
Taxon
Qualifiers
817
816
Taxon
Number
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
+
.
.
WAe
.
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StE
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IoI
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SPi
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NPi
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NC
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Kik
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EAe
Distribution
Qualifier
270 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Iberis epirota Halácsy
Brassicaceae
Iberis spathulata sensu Bald. 1899
FGC 1:106 FHe 2:267 MFG 1:333
Iberis runemarkii Greuter & Burdet
Brassicaceae
Iberis arbuscula Runemark
FHe 2:266 FT 1:310 FT 11:31 MCL 3:128 RDB :322
L/138 L/540 L/941
Inula candida (L.) Cass.
Asteraceae
Conyza candida L.
Inula candida (L.) Cass. subsp. candida
Inula candida (L.) Cass subsp. limoniifolia Hayek
Inula candida var. p.p. Halácsy Fl. Graec.
Inula limoniifolia (Sm.) Boiss. ex Raulin
Inula limoniifolia subsp. rotundifolia
EFK :310 FAe :617 FCA :68 FCSI :33 FE 4:135
B/3060c L/119 L/554 L/765 L/932 L/1052 L/1903 L/1917 L/1963
Inula oxylepis Hausskn.
Asteraceae
Inula candida subsp. limonella (Heldr.) Rech.f.
Inula sophiae Beauverd
Inula sophiae Beauverd var. sporadum Phitos
FE 4:135
L/196 L/323b L/443 L/608 L/1871 L/1871 L/2013
Inula pseudolimonella (Rech.f.) Rech.f.
Asteraceae
Inula candida (L.) Cass. subsp. decalvans (Halácsy) P.W. Ball
Inula limonifolia (Sm.) Boiss. var. decalvans Halácsy
Inula limonifolia (Sm.) Boiss. var. pseudolimonella Rech.f.
EFK :310 FCA :68 FCSI :33 FE 4:135
L/554
Inula rotundifolia (Halácsy) Greuter
Asteraceae
Inula candida var. rotundifolia Halácsy
FE 4:136
L/607 L/1963
824
825
826
827
828
829
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 271
Iris unguicularis Poiret subsp. cretensis (Janka) Davis & Jury
Iridaceae
Iris cretensis Janka
Iris cretica Herbert ex Baker nomen nudum
Iris humilis Sieber ex Baker
Iris stylosa Desf. var. angustifolia Boiss. & Heldr.
Siphonostylis cretensis (Janka) Schulze subsp. cretensis
EFK :365 FCA :178 FCSI :104 FE 5:88 FGC 3:189 FT 8:391 MFG 2:722
L/159 L/201 L/321 L/411 L/1188
Isatis grammotis Kit Tan
Brassicaceae
FHe 2:128
835
836
!
Iris unguicularis Poiret subsp. carica (W. Schulze) Davis & Jury var.
angustifolia (Boiss. & Heldr.) Davis & Jury
Iridaceae
Iris cretica Herbert ex Baker nomen nudum
Iris stylosa Mazziari
Siphonostylis Hybridocomplex Schulze
FGC 3:189
L/411 L/776 L/1188
Iris falcata Babalonas & Papan.
Iridaceae
L/1284 L/2237
833
834
Inula verbascifolia (Willd.) Hausskn. subsp. parnassica (Boiss. &
Heldr.) Tutin
Asteraceae
Inula parnassica Boiss. & Heldr.
FE 4:136
L/121 L/395 L/607 L/1830 L/1871 L/1963 L/2013 L/2200
832
?
Inula verbascifolia (Willd.) Hausskn. subsp. methanea (Hausskn.)
Tutin
Asteraceae
Inula candida subsp. methanea (Hausskn.) Hayek
Inula methanea Hausskn.
FCA :68 FE 4:136
L/122 L/439 L/516 L/530 L/607 L/1101 L/1963 L/1973
831
Taxon
Inula subfloccosa Rech.f.
Asteraceae
FE 4:136 RDB :324
L/323b L/1871
Taxon
Qualifiers
830
Taxon
Number
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+
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+
+
+
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StE
+
+
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Pe
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+
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+
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IoI
+
WAe
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+
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NPi
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SPi
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EC
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NC
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NE
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NAe
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Kik
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+
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KK
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EAe
Distribution
Qualifier
272 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Isatis vermia Papan.
Brassicaceae
AFE 10:46 FE 1:324 FHe 2:128 MCL 3:131 MFG 1:238
L/1290 L/2201
Isoetes heldreichii Wettst.
838
839
Jacobaea ambigua (Biv.) Pelser & Veldkamp subsp. taygetea (Boiss.
& Heldr.) B. Nord. & Greuter
Asteraceae
Senecio ambiguus (Biv.) DC. subsp. taygeteus (Boiss. & Heldr.) Greuter
Cineraria nebrodensis Guss. subsp. taygetea (Boiss.) Nyman
Senecio ambiguus subsp. ambiguus sec. Strid, A. (ed.) Mountain flora of
Greece 2 Edinburgh 1991
Senecio taygeteus Boiss. & Heldr.
FAe :637 FE 4:194 MFG 2:460
L/323b L/1904 L/1905
Jacobaea gnaphalioides (Spreng.) Veldkamp
Asteraceae
Conyza gnaphalodes (Sieber) Nyman
Conyza gnaphalodes Sieber
Senecio gnaphalioides Spreng.
Senecio gnaphalodes Sieber
EFK :317 FCA :72 FCSI :34 FCSII : FE 4:194
Jankaea heldreichii (Boiss.) Boiss.
Gesneriaceae
Haberlea heldreichii Boiss.
Ramonda heldrechii (Boiss.) Fritsch.
FE 3:285 MCL 3:261 MFG 2:262 RDB :326
L/138 L/183 L/404a L/485 L/966 L/1180
840
841
842
Isoetaceae
AFE 1:30 FE 1:6 FGC 3:481 MCL 1:4
L/536 L/1573 L/1760
Isatis tinctoria L. subsp. athoa (Boiss.) Papan.
Brassicaceae
Isatis tinctoria L. var. stenocarpa Griseb.
Isatis athoa Boiss.
AFE 10:44 FE 1:324 FHe 2:127 MCL 3:129 MFG 1:238
L/1973
837
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+
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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?
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PATTERNS OF PLANT ENDEMISM IN GREECE 273
Jurinea taygetea Halácsy
Asteraceae
FE 4:220 MFG 2:477
L/512 L/1963
Lactuca alpestris (Gand.) Rech.f.
Asteraceae
Scariola alpestris (Gand.) Greuter, Matthäs & Risse
Lactuca viminea (L.) J. Presl & C. Presl subsp. alpestris (Gand.)
Ferakova
Lactuca viminea (L.) J. Presl & C. Presl var. decumbens Halácsy
Phoenixopus alpestris Gand.
EFK :337 FCA :71 FCSI :34 FCSII : FE 4:329
L/253 L/1766
Lamium garganicum L. subsp. pictum (Boiss. & Heldr.) P.W. Ball
Lamiaceae
Lamium garganicum L. subsp. striatum (Sm.) Hayek var. pictum
(Boiss.) Hayek
Lamium pictum Boiss. & Orph.
Lamium striatum Sm. var. minus Boiss.
FE 3:147 MCL 3:288 MFG 2:94
L/261 L/512 L/587 L/607 L/895 L/1252 L/1963 L/2200
Lamyropsis carpini Greuter
Asteraceae
L/991
Laserpitium pseudomeum Orph., Heldr. & Sart. ex Boiss.
Apiaceae
FE 2:370 FGC 1:620 FGS 1:41 MFG 1:733
L/395 L/587 L/607 L/1963 L/2013
Lathyrus neurolobus Boiss. & Heldr.
Fabaceae
EFK :152 FAe :336 FCA :106 FCSI :58 FE 2:140 FGC 1:472 MCL 4:121
L/257 L/554
844
845
846
847
848
849
Taxon
Johrenia distans (Griseb.) Halácsy
Apiaceae
Caroselinum distans Gris.
Johrenia graeca Boiss. & Spruner
FAe :410 FE 2:358 FGC 1:634 FGS 1:42
L/395 L/516 L/530 L/607 L/736 L/781 L/1871 L/1963 L/1973 L/1981
Taxon
Qualifiers
843
Taxon
Number
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StE
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IoI
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WAe
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NPi
SPi
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EC
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NC
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NAe
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Kik
+
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KK
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EAe
Distribution
Qualifier
274 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Leontodon hellenicus Phitos
Asteraceae
MFG 2:531 RDB :330
L/330 L/1118 L/1694
Leptoplax emarginata (Boiss.) O.E. Schulz
Brassicaceae
Peltaria emarginata (Boiss.) Hausskn.
Alyssum emarginatum Rouy
Koniga emarginata (Boiss.) Nyman
Ptilotrichon emarginatum Boiss.
AFE 11:21 FE 1:358 FGC 1:116 FHe 2:231 MCL 3:135 MFG 1:275 PFPB 1:480
L/159 L/323a L/995 L/1871 L/1901
Ligusticum olympicum Novak
Apiaceae
MFG 1:704
L/404a
Ligusticum rhizomaticum Hartvig
Apiaceae
MFG 1:707
L/1041
Limodorum abortivum (L.) Swartz f. gracile (B. Willing & E. Willing)
P. Delforge
Orchidaceae
Limodorum abortivum (L.) Swartz var. gracile (B. Willing & E. Willing)
Kreutz
Limodorum abortivum (L.) Swartz subsp. gracile B. Willing &
E. Willing
L/526 L/611
Limonium albomarginatum Brullo
Plumbaginaceae
MCL 4:322
L/422 L/1963
851
852
853
854
855
856
A
Leontodon graecus Boiss. & Heldr.
Asteraceae
Leontodon crispus Vill. subsp. graecus (Boiss. & Heldr.) Hayek
FAe :681 FE 4:314 FGC 2:187 FT 5:677 MFG 2:530
L/121 L/321 L/323b L/361 L/443 L/516 L/530 L/869 L/895 L/928 L/1123 L/1349 L/1846 L/1871 L/1907 L/1957 L/1960 L/1963 L/2200
850
.
.
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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+
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+
PATTERNS OF PLANT ENDEMISM IN GREECE 275
U
S
U
862
863
864
U
U
861
865
A
860
S
A
858
859
S
?
Taxon
Qualifiers
857
Taxon
Number
Limonium corinthiacum (Boiss. & Heldr.) O. Kuntze
Plumbaginaceae
Statice corinthiaca Boiss. & Heldr.
FE 3:47 FGC 3:17 MCL 4:335
Limonium chrisianum Brullo & Guarino
Plumbaginaceae
FCSII :
L/2093
Limonium cephalonicum Artelari
Plumbaginaceae
Statice cancellata auct. (non Bernh.) var. laxa Boiss. p.p.
Statice cancellata auct. (non Bernh.) var. laxa Boiss. p.p.
MCL 4:328
L/747 L/903 L/1123
Limonium carpathum (Rech.f.) Rech.f.
Plumbaginaceae
Statice carpatha Rech.f.
EFK :226 FAe :429 FCA :125 FE 3:46 MCL 4:325
L/551 L/786 L/1049
Limonium calliopsium A. Mayer
Plumbaginaceae
EFK :227 FCSI :70
B/2246
Limonium brevipetiolatum Artelari & Erben
Plumbaginaceae
MCL 4:343
L/339 L/829 L/1963
Limonium arcuatum Artelari
Plumbaginaceae
Statice cancellata auct., non Bernh.
MCL 4:328
L/747 L/903
Limonium aphroditae Artelari & Georgiou
Plumbaginaceae
L/671 L/1584 L/1963
Limonium antipaxorum Artelari
Plumbaginaceae
MCL 4:328
L/747 L/903
Taxon
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+
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StE
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+
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Pe
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WAe
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+
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+
+
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+
IoI
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SPi
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NPi
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EC
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NC
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NAe
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Kik
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+
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KK
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EAe
Distribution
Qualifier
276 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
U
S
A
A
S
U
U
U
866
867
868
869
870
871
872
873
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Limonium frederici (Barbey) Rech.f.
Plumbaginaceae
Statice frederici Barbey
FAe :429 FCA :126 FE 3:45 MCL 4:330
L/111 L/196 L/255 L/268 L/554 L/932 L/1049
Limonium elaphonisicum A. Mayer
Plumbaginaceae
EFK :226 FCSI :71
B/2246 L/762
Limonium doerfleri (Halácsy) Rech.f.
Plumbaginaceae
Limonium ramosissimum (Poiret) Maire subsp. doerfleri (Halácsy)
Pignatti
Statice doerfleri Halácsy
EFK :228 FAe :429 FCA :126 FE 3:47 FT 7:470 MCL 4:328
L/169 L/201 L/707 L/776 L/861 L/1210a
Limonium damboldtianum Phitos & Artelari
Plumbaginaceae
MCL 4:328 RDB :338
L/747 L/903 L/1121 L/1123
Limonium cythereum Artelari & Georgiou
Plumbaginaceae
L/671 L/1584 L/1963
Limonium creticum Artelari
Plumbaginaceae
EFK :228 FCSI :71 RDB :336
B/2246 L/762 L/904
Limonium coronense Artelari
Plumbaginaceae
Limonium melium (Nyman) Pignatti p.p.
Statice corcyrense Guss. var. major Boiss. p.p.
Statice melia Nyman p.p.
MCL 4:328
L/607 L/747 L/903 L/1963
Limonium cornarianum Kypr. & Artelari
Plumbaginaceae
FCSII :
L/554 L/1572
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+
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+
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+
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+
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+
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+
+
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+
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+
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+
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PATTERNS OF PLANT ENDEMISM IN GREECE 277
A
A
A
S
S
U
S
A
875
876
877
878
879
880
881
Taxon
Qualifiers
874
Taxon
Number
Limonium ocymifolium (Poiret) O. Kuntze
Plumbaginaceae
Limonium pigadiense (Rech.f.) Rech.f.
Statice ocymifolia Poiret
Statice pigadiensis Rech.f.
EFK :227 FCA :126 FE 3:47 FT 7:470 MCL 4:335
L/111 L/119 L/197 L/421 L/446 L/671 L/765 L/796 L/913 L/932 L/1049 L/1288 L/1627 L/1713 L/2181
Limonium messeniacum Artelari & Kamari
Plumbaginaceae
L/1687 L/1963
Limonium melium (Nyman) Pignatti
Plumbaginaceae
Statice cosyrensis Guss. subsp. melia Nyman
FAe :427 FE 3:44 FGC 3:21 MCL 4:333
L/772
Limonium kardamylii Artelari & Kamari
Plumbaginaceae
L/1963
Limonium ithacense Artelari
Plumbaginaceae
MCL 4:328 RDB :342
L/747 L/903 L/1123
Limonium hyssopifolium (Girard) Rech.f.
Plumbaginaceae
Statice hyssopifolia Girard
EFK :226 FAe :427 FCA :126 FCSII : FE 3:46 FGC
3:20 FT 7:471 MCL 4:336
L/308e L/537 L/539 L/540 L/776 L/786 L/932 L/2093
Limonium hierapetrae Rech.f.
Plumbaginaceae
EFK :228 FCA :126 FCSI :71 MCL 4:335
B/2246 L/554
Limonium graecum (Poiret) Rech.f. subsp. ammophilon Papatsou &
Phitos
Plumbaginaceae
EFK :226 FCSI :71 FT 7:472 MCL 4:336
B/2246 L/445 L/671 L/2089
Taxon
+
+
+
+
.
+
.
.
Pe
.
.
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.
+
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.
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.
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+
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IoI
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WAe
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+
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StE
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SPi
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NPi
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EC
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NC
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NE
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NAe
+
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+
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.
Kik
+
.
.
.
.
+
+
+
KK
+
.
.
.
.
+
.
+
EAe
Distribution
Qualifier
278 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
A
S
S
U
A
S
U
882
883
884
885
886
887
888
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Limonium sitiacum Rech.f.
Plumbaginaceae
EFK :227 FCA :126 FCSI :72 MCL 4:336
B/2246 L/201 L/268 L/537 L/554 L/1710
Limonium saracinatum Artelari
Plumbaginaceae
Statice cancellata auct. (non Bernh.) var. laxa Boiss. p.p.
MCL 4:328
L/747 L/903 L/1123
Limonium runemarkii Rech.f.
Plumbaginaceae
Limonium pigadiense sensu Christod. et al. Gramvousa
Limonium ocymifolium sensu Greuter & Rechinger 1967;
Yannitsaros 1969 Kythira
FCSII 0: FE 3:46 MCL 4:336
L/119 L/323a L/671 L/765 L/932 L/1871
Limonium rigidum A. Mayer
Plumbaginaceae
EFK :227 FCSI :71
B/2246
Limonium pylium Artelari
Plumbaginaceae
Statice cancellata auct. (non Bernh.) var. glabra Boiss. p.p.
MCL 4:328
L/747 L/903 L/1963
Limonium phitosianum Artelari
Plumbaginaceae
MCL 4:328 RDB :344
L/722 L/747 L/903 L/1800
Limonium palmare (Sm.) Rech.f.
Plumbaginaceae
Statice graeca Poiret var. palmaris (Sm.) Hayek
Statice palmaris Sm.
Statice rorida var. palmaris Boiss.
FAe :430 FE 3:46 MCL 4:336
L/111 L/323a L/1871 L/2227
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PATTERNS OF PLANT ENDEMISM IN GREECE 279
S
890
Linum gyaricum Vierh. subsp. gyaricum
Linaceae
FT 2:434
L/329 L/423 L/767
Linum gyaricum Vierh. subsp. icaricum Christod.
Linaceae
FT 2:435
L/329 L/539 L/540
Linum hellenicum Iatroú
Linaceae
MCL 4:222 RDB :350
L/267 L/1963
Linum leucanthum Boiss. & Spruner
Linaceae
FAe :273 FE 2:208 FGC 1:258 MCL 4:223 MFG 1:559
L/99 L/323b L/443 L/530 L/767 L/1871 L/1973 L/1981
Linum olympicum Boiss. subsp. athoum Hartvig
Linaceae
Linum iberidifolium Boiss. (nom. inval.)
MCL 4:224 MFG 1:563
L/1973
893
894
895
896
897
Linum goulimyi Rech.f.
Linaceae
FE 2:208 MCL 4:220
L/323a L/1871
Linum gyaricum Vierh.
Linaceae
FE 2:208 FT 2:434 MCL 4:222
L/329 L/423 L/443 L/539 L/856
?
Limonium zacynthium Artelari
Plumbaginaceae
MCL 4:328 RDB :346
L/722 L/747 L/903 L/1800
Limonium tenium (Heldr.) Rech.f.
Plumbaginaceae
Statice virgata Willd. var. tenia Heldr.
FE 3:46 FGS 1:73 MCL 4:343
L/99 L/323a
Taxon
892
891
U
Taxon
Qualifiers
889
Taxon
Number
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WAe
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StE
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Pe
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IoI
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SPi
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NPi
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EC
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NC
+
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
280 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Linum phitosianum Christod. & Iatroú
Linaceae
RDB :352
L/976 L/1963
Linum punctatum C.Presl. subsp. pycnophyllum (Boiss. & Heldr.)
Gustavsson
Linaceae
Linum pycnophyllum Boiss. & Heldr.
FE 2:209 FT 2:445 MCL 4:219 MFG 1:561
L/715 L/1264 L/2013
Lithodora zahnii (Heldr. ex Halácsy) I.M.Johnston
Boraginaceae
Lithospermum zahnii Heldr. ex Halácsy
FE 3:88 MCL 1:93 RDB :354
L/97 L/607 L/970 L/1963
Lomelosia albocincta (Greuter) Greuter
Caprifoliaceae
Scabiosa albocincta Greuter
Trochocephalus albocinctus (Greuter) J.Holub
EFK :293 FCA :87 FCSI :44 FE 4:69 MCL 3:187
L/255 L/336 L/518 L/750 L/941 L/1247 L/1583
Lomelosia crenata (Cyr.) Greuter & Burdet subsp. breviscapa
(Boiss. & Heldr.) Greuter & Burdet
Caprifoliaceae
Scabiosa breviscapa Boiss. & Heldr.
Scabiosa crenata Cyr. subsp. breviscapa (Boiss. & Heldr.) Hayek
Scabiosa crenata Cyr. var. breviscapa (Boiss. & Heldr.) Boiss.
FE 4:70 MCL 3:166 MFG 2:362
L/476 L/512 L/607 L/941 L/1963
Lomelosia hymettia (Boiss. & Spruner) Greuter & Burdet
Caprifoliaceae
Scabiosa cretica L. var. heterophylla Hampe
Scabiosa cretica L. var. trifoliolata Bory & Chaub.
Scabiosa hymettia Boiss. & Spruner
Scabiosa trifoliolata (Bory & Chaub.) Halácsy
Trochocephalus hymettius (Boiss. & Spruner) J.Holub
FAe :594 FE 4:69 FGC 1:764 MCL 3:188 PFPB 2:510
L/98 L/267 L/443 L/530 L/607 L/736 L/928 L/941 L/1247 L/1262e L/1871 L/1963
898
899
900
901
902
903
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PATTERNS OF PLANT ENDEMISM IN GREECE 281
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Lomelosia minoana (P.H.Davis) Greuter & Burdet subsp. asterusica
(Greuter) Greuter & Burdet
Caprifoliaceae
Scabiosa cretica Scabiosa cretica sensu Bald. 1903, p.p.; Fl. Aeg. p.p.
Scabiosa minoana (P.H.Davis) Greuter subsp. asterusica Greuter
EFK :293 FCA :87 FE 4:69 MCL 3:188 RDB :356
L/941 L/1247 L/1583
Lomelosia minoana (P.H.Davis) Greuter & Burdet subsp. minoana
Caprifoliaceae
Scabiosa cretica (L.) Greuter & Burdet subsp. minoana P.H.Davis
Scabiosa cretica sensu auct. cret. Fl. Aeg. p.p. (non L. 1753 s.str.)
Scabiosa minoana (P.H.Davis) Greuter
Scabiosa minoana (P.H.Davis) Greuter subsp. minoana
Trochocephalus minoanus (P.H.Davis) J.Holub
EFK :293 FAe :594 FCA :87 FE 4:69 MCL 3:188
L/518 L/1247 L/1583
Lomelosia sphaciotica (Roemer & J.A.Schultes) Greuter
Caprifoliaceae
Scabiosa sphaciotica Roemer & J.A.Schultes
EFK :292 FCA :87 FCSI :45 FE 4:70 MCL 3:191 MFG 2:360
L/941
Lonicera alpigena L. subsp. hellenica (Boiss.) Kit Tan & Ziel.
Caprifoliaceae
Lonicera hellenica Orph. ex Boiss.
FE 4:47 MCL 1:158
L/607 L/1772 L/1963
Lotus aduncus (Griseb.) Nyman
Fabaceae
Tetragonolobus aduncus Griseb.
FE 2:175 MCL 4:131 MFG 1:521
L/533 L/1004 L/1267b L/1349
905
906
907
908
909
Taxon
Lomelosia minoana (P.H.Davis) Greuter & Burdet
Caprifoliaceae
Scabiosa cretica (L.) Greuter subsp. minoana P.H.Davis
Scabiosa minoana (P.H.Davis) Greuter
Trochocephalus minoanus (P.H.Davis) J.Holub
EFK :293 FAe :594 FCA :87 FCSI :45 FE 4:69 MCL 3:188
L/941 L/1247 L/1583
Taxon
Qualifiers
904
Taxon
Number
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WAe
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StE
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IoI
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SPi
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NPi
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EC
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NC
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NE
+
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NAe
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Kik
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KK
.
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EAe
Distribution
Qualifier
282 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Malcolmia graeca Boiss. & Spruner subsp. bicolor (Boiss. & Heldr.)
Stork
Brassicaceae
Malcolmia bicolor Boiss. & Heldr. var. veluchensis (Boiss. & Heldr.)
Boiss.
Malcolmia bicolor Boiss. & Heldr.
Malcolmia veluchensis Boiss. & Heldr.
Wilckia bicolor Boiss. & Heldr.
AFE 10:104 FE 1:338 FHe 2:159 MCL 3:137 MFG 1:251
L/512 L/530 L/895 L/930 L/1772 L/1844 L/1963 L/2200
Malcolmia graeca Boiss. & Spruner subsp. graeca
Brassicaceae
Malcolmia graeca Boiss. & Spruner var. tenuior Hausskn.
AFE 10:104 FE 1:338 FHe 2:159 MCL 3:137 MFG 1:251
L/1178 L/1871 L/1963
914
Malabaila psaridiana Heldr.
Apiaceae
FE 2:367
L/607
912
913
Lysimachia serpyllifolia Schreber
Primulaceae
Lysimachia anagalloides Sm. (from Kriti)
EFK :323 FAe :435 FCA :129 FE 3:26 MCL 4:378 MFG 1:746
L/512 L/518 L/607 L/1769 L/1871 L/1963 L/2013 L/2201
911
?
Lutzia cretica (L.) Greuter & Burdet
Brassicaceae
Lutzia fruticosa Gand.
Alyssum oocarpum Gand.
Scleroptychis cretica (L.) Degen
Alyssoides cretica (L.) Medicus
Alyssum creticum L.
AFE 11:23 EFK :117 FCA :84 FCSI :42 FCSII : FE 1:359 FHe 2:199 MCL 3:136
L/145 L/165 L/201 L/518 L/554 L/830
910
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 283
Marrubium cylleneum Boiss. & Heldr.
Lamiaceae
Marrubium velutinum Sm. subsp. cylleneum (Boiss. & Heldr.) Nyman
Marrubium velutinum Sm. var. cylleneum (Boiss. & Heldr.) Boiss.
FE 3:137 MCL 3:293 MFG 2:83
L/160 L/512 L/1830 L/1963
Marrubium velutinum Sm.
Lamiaceae
Marrubium haussknechtii Hausskn.
FE 3:138 MCL 3:294 MFG 2:83
L/121 L/159 L/160 L/512 L/587 L/953 L/1211 L/1830 L/1963 L/2200 L/2201
920
Malcolmia orsiniana (Ten.) Ten. subsp. angulifolia (Boiss. & Orph.)
A.Stork
Brassicaceae
Malcolmia angulifolia Boiss. & Orph.
Malcolmia cymbalaria Boiss.
Wilckia angulifolia (Boiss. & Orph.) Halácsy
AFE 10:107 FE 1:339 FHe 2:164 MCL 3:139 MFG 1:249
L/529 L/2013
918
919
Malcolmia macrocalyx (Halácsy) Rech.f. subsp. scyria (Rech.f.)
P.W.Ball
Brassicaceae
Malcolmia scyria Rech.f.
AFE 10:107 FAe :217 FE 1:339 FHe 2:161 MCL 3:138
L/196 L/323a L/915 L/1178 L/1871 L/2181
917
!
Malcolmia macrocalyx (Halácsy) Rech.f. subsp. macrocalyx
Brassicaceae
AFE 10:107 FAe :217 FE 1:339 FHe 2:161 MCL 3:138
L/122 L/423 L/443 L/447 L/1832d
916
Taxon
Malcolmia macrocalyx (Halácsy) Rech.f.
Brassicaceae
Malcolmia maritima (L.) R.Br. var. macrocalyx (Halácsy) Hayek
Wilckia maritima (L.) Halácsy var. macrocalyx Halácsy
AFE 10:107 FAe :217 FE 1:339 FHe 2:161 MCL 3:138
L/323a L/443 L/447 L/1178 L/1832d L/2181
Taxon
Qualifiers
915
Taxon
Number
.
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+
IoI
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WAe
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StE
Pe
+
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SPi
+
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NPi
.
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+
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EC
+
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+
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NC
.
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+
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?
?
NE
.
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NAe
.
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Kik
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KK
.
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EAe
Distribution
Qualifier
284 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Melica rectiflora Boiss. & Heldr.
Poaceae
EFK :384 FAe :774 FCA :172 FCSI :101 FE 5:178 MFG 2:784
L/159 L/257 L/512 L/518 L/551 L/554 L/1963
Melilotus graecus (Boiss. & Spruner) Lassen
Fabaceae
Pocockia graeca Boiss. & Spruner
EFK :156 FCA :109 FE 2:151 MCL 4:147
L/121 L/253 L/267 L/516 L/530 L/607 L/923 L/1800 L/1830 L/1846 L/1917 L/1963 L/1981 L/2200
927
Melica cretica Boiss. & Heldr.
Poaceae
EFK :384 FCA :172 FE 5:179 MFG 2:785
925
926
Melampyrum ciliatum Boiss. & Heldr.
Orobanchaceae
Melampyrum barbatum Waldst. & Kit. ex Willd. subsp. ciliatum
(Boiss. & Heldr.) Nyman
FE 3:254 FGC 2:444
924
?
Medicago strasseri Greuter, Matthäs & Risse
Fabaceae
EFK :158 FCA :109 FCSI :60 MCL 4:136
L/253 L/908 L/1583
923
Matricaria rosella (Boiss. & Orph.) Nyman
Asteraceae
Chamaemelum rosellum Boiss. & Orph.
Tripleurospermum rosellum (Boiss. & Orph.) Hayek
FAe :633 FE 4:166 FGC 3:67 FT 5:300
L/111 L/121 L/377 L/1588 L/2200
Matricaria tempskyana (Freyn & Sint.) Rauschert
Asteraceae
Chamaemelum tempskyanum Freyn & Sint.
Tripleurospermum tempskyanum (Freyn & Sint.) Hayek
FE 4:167 MFG 2:452
L/995
!
922
921
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PATTERNS OF PLANT ENDEMISM IN GREECE 285
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Mentha pulegium L. subsp. erinoides (Heldr.) Kokkini
Lamiaceae
Mentha erinoides Heldr.
Mentha gryparia Heldr. f. erinoides Heldr.
Mentha pulegium L. var. erinoides (Heldr.) Hayek
EFK :267 FCA :96 MCL 3:296
L/253 L/522 L/1082
Micromeria hispida Boiss. & Heldr. ex Benth.
Lamiaceae
Satureja candica Greuter & Burdet
Satureja hispida (Benth.) Nyman
EFK :264 FCA :99 FCSII : FE 3:168 MCL 3:335 MFG 2:135
L/940 L/1941
Micromeria sphaciotica Boiss. & Heldr. ex Benth.
Lamiaceae
Satureja sphaciotica (Boiss. & Heldr. ex Benth.) Greuter & Burdet
Micromeria carpatha Rech.f.
Micromeria microphylla sensu auct. karp., kret., Consp. Fl. Graec.;
Fl. Aeg.; Boiss. Fl. Or.
EFK :264 FAe :526 FCA :100 FCSI :54 FCSII : FE 3:168 MCL 3:36
L/273 L/518 L/707 L/940 L/1941
Micromeria taygetea P.H.Davis
Lamiaceae
Satureja taygetea (P.H.Davis) Greuter & Burdet
FE 3:168 MCL 3:340 MFG 2:132
L/138 L/607 L/940 L/1963
Minuartia athoa (Griseb.) Kamari
Caryophyllaceae
Alsine setacea (Thuill.) Mert. & W.D.J.Koch
Minuartia setacea (Thuill.) Hayek var. athoa (Griseb.) Mattf.
FHe 1:180
Minuartia athoa (Griseb.) Kamari subsp. athoa
Caryophyllaceae
FHe 1:180
929
930
931
932
933
934
Taxon
Mentha pulegium L. subsp. cephalonia (H.Braun) Kokkini
Lamiaceae
Mentha pulegium L. var. subtomentella f. cephalonia H.Braun
MCL 3:296
L/1082 L/1123 L/1846
Taxon
Qualifiers
928
Taxon
Number
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WAe
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StE
Pe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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+
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Kik
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+
+
+
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KK
.
.
.
.
.
.
.
EAe
Distribution
Qualifier
286 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Minuartia athoa (Griseb.) Kamari subsp. neoiraklitsa Kamari
Caryophyllaceae
FHe 1:180
Minuartia attica (Boiss. & Spruner) Vierh. subsp. idaea (Halácsy)
Kamari & Constantin.
Caryophyllaceae
Minuartia verna (L.) Hiern var. acutipetala Boiss.
Alsine verna (L.) Wahlenb var. idaea Halácsy
Minuartia idaea (Halácsy) Pawlsowski
Minuartia verna (L.) Hiern subsp. idaea (Halácsy) Hayek
FCA :48 FCSI :19 FE 1:158 FHe 1:186 FT 11:46 MCL 1:228 MFG 1:104
L/321 L/539 L/540 L/551 L/554 L/796 L/974 L/1075 L/1957
Minuartia confusa (Boiss.) Maire & Petitm.
Caryophyllaceae
Alsine confusa Boiss.
Alsine trichocalycina sensu Boissier
Cherleria sedoides sensu Sm. non L.
Minuartia trichocalycina sensu auct., non Arenaria trichocalycina Ten.
& Guss.
AFE 6:49 FE 1:156 FHe 1:179 MCL 1:216 MFG 1:95
L/160 L/607 L/1963 L/2013
Minuartia dirphya Trigas & Iatroú
Caryophyllaceae
L/1871 L/1898
Minuartia eurytanica (Boiss. & Heldr.) Hand.-Mazz
Caryophyllaceae
Alsine eurytanica Boiss. & Heldr.
Alsine recurva All. var. eurytanica (Boiss. & Heldr.) Halácsy
Minuartia hirsuta (M.Bieb.) Hand.-Mazz. subsp. eurytanica (Boiss. &
Heldr.) Strid
AFE 6:51 FAe :137 FE 1:156 FHe 1:189 FT 11:45 FT 2:47 MCL 1:219 MFG 1:97
L/513 L/2013 L/2200
Minuartia graminifolia (Ard.) Jav. subsp. brachypetala Kamari
Caryophyllaceae
FHe 1:190
935
936
937
938
939
940
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 287
Minuartia parnonia (Kamari) Iatroú, Trigas & Kit Tan
Caryophyllaceae
Minuartia wettsteinii Mattf. subsp. parnonia Kamari
FHe 1:182 RDB :366
L/1963
Minuartia pichleri (Boiss.) Maire & Petitm.
Caryophyllaceae
Minuartia favargeri Iatroú & T.Georgiadis
Alsine pichleri Boiss.
AFE 6:57 FE 1:158 FHe 1:184 MCL 1:223 MFG 1:103 RDB :364
L/512 L/607 L/756 L/1915 L/1963
Minuartia pseudosaxifraga (Mattf.) Greuter & Burdet
Caryophyllaceae
Alsine graminifolia sensu auct. from Tymfi, Cons. Fl. Graec.
Minuartia stellata (E.D.Clarke) Maire & Petitmegin subsp.
pseudosaxifraga Mattf.
AFE 6:53 FE 1:157 FGC 3:237 FHe 1:191 MCL 1:223 MFG 1:101
L/601
Minuartia setacea (Thuill.) Hayek subsp. olympica Kamari
Caryophyllaceae
FHe 1:179
Minuartia wettsteinii Mattf.
Caryophyllaceae
Minuartia wettsteinii Mattf. subsp. wettsteinii
EFK :78 FAe :138 FCA :48 FCSI :20 FCSII : FE 1:159 FHe 1:182 MCL 1:229 MFG 1:105 RDB :366
L/707
Muscari cycladicum P.H.Davis & Stuart
Asparagaceae
Leopoldia cycladica (Davis & Stuart) Bentzer
FCA :184 FE 5:47
L/430 L/730 L/776 L/783 L/869 L/948 L/1641 L/2093
942
943
944
945
946
947
Taxon
Minuartia greuteriana Kamari
Caryophyllaceae
Minuartia greuteriana Kamari var. pessana
FHe 1:188 RDB :362
L/342
Taxon
Qualifiers
941
Taxon
Number
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WAe
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StE
Pe
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IoI
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SPi
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NPi
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EC
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NC
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
288 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Muscari cycladicum P.H.Davis & Stuart subsp. cycladicum
Asparagaceae
Leopoldia cycladica (Davis & Stuart) Bentzer subsp. cycladica
L/730 L/776 L/948
Muscari cycladicum P.H.Davis & Stuart subsp. subsessilis (Bentzer)
Raus
Asparagaceae
Leopoldia cycladica (Davis & Stuart) Garbari subsp. subsessilis Bentzer
L/730 L/776 L/869 L/948
Muscari dionysicum Rech.f.
Asparagaceae
Leopoldia dionysica (Rech.f.) Greuter
EFK :358 FCA :184 FE 5:47 FT 8:251
L/268 L/423 L/554 L/730 L/767 L/948 L/1710
Muscari kerkis Karlén
Asparagaceae
FT 10:225 MFG 2:701 RDB :368
L/282 L/796
Muscari pulchellum Heldr. & Sart.
Asparagaceae
Botryanthus pulchellus (Heldr. & Sart.) Nyman
Botryanthus sartorii Todaro
Muscari racemosum subsp. pulchellum ( Heldr. & Sart.) Stojanov &
Stefanof
FAe :732 FE 5:48 MFG 2:701
L/282 L/530 L/607 L/685 L/776 L/869 L/1288 L/1711 L/1844 L/1871 L/1957 L/1963
Muscari pulchellum Heldr. & Sart. subsp. clepsydroides Karlén
Asparagaceae
Muscari pulchellum Heldr. & Sart. var. amoenum Heldr. ex Halácsy
FE 5:48 MFG 2:701
L/282 L/607 L/685 L/776 L/869 L/1288
Muscari pulchellum Heldr. & Sart. subsp. pulchellum
Asparagaceae
FE 5:48 MFG 2:701
L/282 L/607 L/1711 L/1963
948
949
950
951
952
953
954
.
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 289
Myosotis solange Greuter & Zaffran
Boraginaceae
EFK :251 FCA :41 FCSI :15 MCL 1:100 MFG 2:56 RDB :370
L/252 L/518 L/519 L/709 L/1766
Nepeta argolica Bory & Chaub. subsp. argolica
Lamiaceae
Nepeta sibthorpii Benth. subsp. sibthorpii
MFG 2:110
L/351 L/498 L/530 L/1830 L/1963 L/1963
Nepeta argolica Bory & Chaub. subsp. dirphya (Boiss.) Strid & Kit
Tan
Lamiaceae
Nepeta dirphya (Boiss.) Heldr. ex Halácsy
Nepeta sibthorpii Benth. subsp. dirphya (Boiss.) Baden
Nepeta sibthorpii Benth. var. dirphya Boiss.
FAe :508 FE 3:159 MCL 3:304 MFG 2:112
L/99 L/111 L/323b L/351 L/498 L/1871 L/1963
Nepeta argolica Bory & Chaub. subsp. malacotrichos (Baden) Strid
& Kit Tan
Lamiaceae
Nepeta sibthorpii Benth. subsp. malacotrichos Baden
MFG 2:112
L/351 L/1963 L/1963
Nepeta camphorata Boiss. & Heldr.
Lamiaceae
MFG 2:114
L/351 L/497 L/498 L/607 L/1963
Nepeta camphorata Boiss. & Heldr. var. camphorata
Lamiaceae
FE 3:159 MCL 3:303 MFG 2:114
L/351 L/497 L/498 L/607 L/1963
956
957
958
959
960
961
Taxon
Muscari spreitzenhoferi (Heldr.) Vierh.
Asparagaceae
Leopoldia spreitzenhoferi Heldr.
Muscari amoenocomum Rech.f. (included in taxon)
Muscari creticum Vierh.
Muscari cycladicum sensu auct. cret. [Bentzer 1973]?
Muscari maritimum sensu auct. cret.
Muscari weissii sensu auct. cret. [Rechinger 1944, Bentzer 1973]?
EFK :358 FAe :731 FCA :184 FCSI :109 FE 5:47
L/255 L/268 L/430 L/730 L/762 L/932 L/1766 L/2093
Taxon
Qualifiers
955
Taxon
Number
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StE
Pe
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WAe
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IoI
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
290 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Nepeta camphorata Boiss. & Heldr. var. heldreichii (Halácsy)
C.Baden
Lamiaceae
Nepeta heldreichii Halácsy
FE 3:159 MCL 3:303 MFG 2:116
L/497 L/498 L/1963
Nepeta melissifolia Lam.
Lamiaceae
Nepeta melissaefolia Lam. (sphalm.)
EFK :262 FAe :508 FCA :97 FCSI :52 FE 3:160 FGC 2:535 MCL 3:302
L/351 L/762
Nepeta orphanidea Boiss.
Lamiaceae
Nepeta laconia Orphanides, nom. nud.
Nepeta parnassica Heldr. & Sart. (b) orphanidea
FE 3:159 MCL 3:303 MFG 2:116
L/351 L/1963
Nepeta parnassica Heldr. & Sart. ex Boiss.
Lamiaceae
Nepeta orientalis Miller var. parnassica (Heldr. & Sart.) Bald.
Nepeta sibthorpii Benth. subsp. parnassica (Heldr. & Sart.) Maire &
Petitm.
FE 3:159 MCL 3:304 MFG 2:112
L/351 L/498 L/512 L/1963
Nepeta scordotis L.
Lamiaceae
EFK :262 FCA :97 FE 3:159 FGC 2:x MCL 3:303
L/366 L/765 L/1052 L/1963
Nepeta sphaciotica P.H.Davis
Lamiaceae
EFK :262 FCA :97 FE 3:159 MCL 3:304 MFG 2:116 RDB :372
L/138 L/255 L/351 L/485 L/498 L/518 L/519 L/981
962
963
964
965
966
967
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 291
Nigella carpatha Strid
Ranunculaceae
Nigella dejenii sensu Davis Fl. Turk p.p.; Tutin Fl. Eur.p.p.; non Vierh.
AFE 8:33 EFK :96 FCA :131 FE 1:252 FHe 2:9 MCL 4:412
L/316 L/429
Nigella degenii Vierh.
Ranunculaceae
Nigella aristata sensu Halácsy Consp. Fl. Graec. p.p.; Hayek Prod. Fl.
pen. Balk. p.p.; Rechinger Fl. Aeg. p.p.;. non Sibth. & Sm.
AFE 8:34 FE 1:252 FHe 2:9 MCL 4:412
L/205 L/316 L/429 L/783 L/856 L/869 L/2181
Nigella degenii Vierh. subsp. barbro Strid
Ranunculaceae
AFE 8:34 FHe 2:10 MCL 4:412
L/205 L/429 L/2181
970
971
972
Nigella arvensis L. subsp. aristata (Sm.) Nyman
Ranunculaceae
Nigella aristata Sm.
Nigella aristata subsp. rechingeri Tunin
Nigella arvensis L. subsp. involucrata (Boiss.) Terracc. p.p.
Nigella arvensis L. a typica Boiss. sensu Haussknecht, 1892; non sensu
orig. Boissier, Fl. Or.
Nigella arvensis L. subsp. foeniculacea (DC.) Bornm. sensu Bornueller,
1928; non sensu orig. De Candolle, 1818
Nigella arvensis L. subsp. rechingeri (Tutin) Tutin
Nigella arvensis L. var. glauca Boiss. sensu Hayek Prodr. Flor. pen.
Balc.; non sensu orig. Boissier Fl. Or.
Nigella arvensis L. var. involucrata Boiss. p.p.
Nigella huthii Brand. var. aristata (Sm.) Brand, illeg. comb.
AFE 8:32 FAe :178 FE 1:252 FHe 2:7 FT 1:102 MCL 4:411
L/205 L/316 L/429 L/447 L/516 L/530 L/607 L/1871 L/1963
Taxon
Nigella arvensis L. subsp. brevifolia Strid
Ranunculaceae
Nigella arvensis L. subsp. cretica (Miller) Ciferri & Ciacomini
Nigella arvensis L. var. glaucescens sensu Rechinger 1943
Nigella arvensis L. var. microcarpa Boiss. [sphalm.macrocarpa] p.p.
Nigella cretensis Steven
Nigella cretica sensu Halácsy Consp. Fl. Graec.; Hayek Prodr. Fl. pen.
Balk.; Rechinger Fl. Aeg.; non sensu orig. Miller
AFE 8:32 EFK :95 FAe :78 FCA :131 FE 1:252 FHe 2:8 FT 10:13 MCL 4:411
L/197 L/205 L/316 L/429 L/750 L/1963
!
Taxon
Qualifiers
969
968
Taxon
Number
.
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+
StE
+
+
Pe
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WAe
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IoI
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SPi
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+
NPi
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+
EC
.
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NC
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NAe
NE
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Kik
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+
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KK
.
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+
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EAe
Distribution
Qualifier
292 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Nigella degenii Vierh. subsp. degenii
Ranunculaceae
AFE 8:34 FHe 2:9 MCL 4:412
L/128 L/205 L/429 L/776
Nigella degenii Vierh. subsp. jenny Strid
Ranunculaceae
Nigella aristata sensu Halácsy Consp. Fl. Graec. p.p.; Hayek Prod. Fl.
pen. Balk. p.p.; Rechinger Fl. Aeg. p.p.;. non Sibth. & Sm.
Nigella divaricata Beaupre ex DC. sensu Gandoger, 1920; non sensu
orig. De Candolle
AFE 8:34 FHe 2:10 MCL 4:412
L/205 L/429
Nigella degenii Vierh. subsp. minor Strid
Ranunculaceae
AFE 8:34 FHe 2:10 MCL 4:412
L/205 L/429
Nigella doerfleri Vierh.
Ranunculaceae
Nigella cretensis sensu Greuter & Rechinger, 1967 p.p.; non sensu orig.
Steven
Nigella cretica auct. p.p. non Miller
Nigella degenii sensu Tutin & al. Fl. Eur. 1967 p.p.; Rechinger Fl. Aeg.
p.p., non Vierh.
AFE 8:35 EFK :96 FCA :131 FE 1:252 FHe 2:11 MCL 4:412
L/128 L/197 L/205 L/316 L/429 L/518 L/762 L/776
L/783 L/869 L/932 L/2093
Nigella icarica Strid
Ranunculaceae
Nigella arvensis var. involucrata Boiss. sensu Fors.-Major & Barbey,
1897; non sensu orig. Boissier Fl. Or.
AFE 8:32 FAe :179 FHe 2:10 FT 10:14 MCL 4:412
L/205 L/429 L/539 L/540
Nigella stricta Strid
Ranunculaceae
Nigella cretensis sensu Greuter & Rechinger, 1967 p.p.; non sensu orig.
Steven, 1856
AFE 8:35 EFK :96 FCA :132 FE 1:253 FHe 2:12 MCL 4:413
L/205 L/429 L/518 L/765 L/1052 L/1878 L/1963
973
974
975
976
977
978
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 293
Omphalodes runemarkii Strid & Kit Tan
Boraginaceae
L/1895
Onobrychis aliacmonia Rech.f.
Fabaceae
MCL 4:152
L/1025 L/1081 L/1100 L/1150 L/1963
Onobrychis ebenoides Boiss. & Spruner
Fabaceae
Onobrychis ebenoides Boiss. & Spruner var. minor Boiss.
Onobrychis carnea Boiss. & Heldr.
Onobrychis hybrida Halácsy
FAe :388 FE 2:188 FGC 1:458 FT 11:98 MCL 4:151
L/323a L/530 L/587 L/607 L/1713 L/1830 L/1844 L/1871 L/1963 L/1980 L/1981 L/2013
Onobrychis montana DC. subsp. macrocarpa Strid
Fabaceae
Onobrychis montana DC. subsp. cadmea (Boiss.) P.W.Ball
Onobrychis scardica var. brevicaulis Halácsy
MCL 4:156 MFG 1:537
L/121 L/395 L/1963 L/2200
Onobrychis peloponnesiaca (Iatroú & Kit Tan) Iatroú & Kit Tan
Fabaceae
Onobrychis aliacmonia Rech.f. subsp. peloponnesiaca Iatroú & Kit Tan
L/1081 L/1600 L/1963
Onobrychis sphaciotica Greuter
Fabaceae
EFK :173 FCA :110 FE 2:189 MCL 4:154 MFG 1:535 RDB :376
L/255 L/408 L/518 L/1583
980
981
982
983
984
985
Taxon
Odontites linkii Heldr. & Sart. ex Boiss. subsp. linkii
Orobanchaceae
Odontites bocconei (Guss.) Walpers subsp. linkii (Heldr. & Sart.) Maire
& Petitm.
Odontites cretica Boiss. (included in taxon)
Odontites frutescens Halácsy
Odontites linkii Heldr. & Sart. ex Boiss. subsp. creticus (Boiss.) Greuter
(included in taxon)
EFK :281 FCA :143 FE 3:268 MFG 2:241
L/255 L/529 L/584 L/607 L/1844 L/1871 L/1963 L/2013
Taxon
Qualifiers
979
Taxon
Number
.
.
.
.
.
+
.
.
.
.
+
+
.
.
+
.
+
.
.
.
.
.
SPi
IoI
+
+
.
.
+
WAe
.
.
.
+
StE
.
.
+
+
Pe
.
.
.
.
.
.
.
NPi
.
.
.
.
.
.
+
EC
.
.
.
.
+
.
+
NC
.
.
.
.
.
.
.
NE
.
.
.
.
.
.
.
NAe
.
.
.
.
.
.
.
Kik
+
.
.
.
.
.
+
KK
.
.
.
+
.
.
+
EAe
Distribution
Qualifier
294 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Onopordum laconicum Heldr. & Sart. ex Rouy
Asteraceae
Onopordum tauricum Willd. var. elatum auct. cyth.
FE 4:246
L/119 L/607 L/765 L/1963
Onopordum bracteatum Boiss. & Heldr. subsp. myriacanthum
(Boiss.) Franco
Asteraceae
Onopordum myriacanthum Boiss.
FCA :69 FE 4:247 FT 5:363
L/197 L/456d L/554 L/607 L/1963
990
992
Onopordum bracteatum Boiss. & Heldr. subsp. creticum Franco
Asteraceae
EFK :325 FCA :69 FCSI :33 FE 4:247
L/253 L/456d
989
Onopordum caulescens D’Urv. subsp. caulescens
Asteraceae
FAe :657 FE 4:246
L/776 L/1641
Onopordum acanthium L. subsp. parnassicum (Boiss. & Heldr.)
Nyman
Asteraceae
FE 4:246
988
991
Ononis verae Sirj.
Fabaceae
EFK :155 FAe :340 FCA :111 FCSI :61 FE 2:145 MCL 4:163
L/762 L/1766
987
!
Ononis spinosa L. subsp. diacantha (Rchb.) Greuter
Fabaceae
Ononis antiquorum L. subsp. diacantha (Rchb.) Greuter
Ononis diacantha Rchb.
Ononis repens L. subsp. diacantha (Rchb.) Greuter
Ononis spinosa L. var. diacantha (Rchb.) Sirj.
EFK :155 FAe :342 FCA :111 FE 2:147 MCL 4:166
L/119 L/355 L/765 L/786 L/1052
986
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 295
Onosma elegantissima Rech.f. & Goulimy
Boraginaceae
FE 3:93 MCL 1:107 MFG 2:35 RDB :378
L/614 L/951 L/995 L/1145
Onosma epirotica Teppner
Boraginaceae
MFG 2:39
L/558
Onosma erecta Sm.
Boraginaceae
EFK :248 FAe :456 FCA :41 FCSI :15 FE 3:94 FT 6:369 MCL 1:107 MFG 2:33
L/395 L/530 L/607 L/1262a L/1963 L/1973 L/2013 L/2201
Onosma erecta Sm. subsp. erecta
Boraginaceae
Onosma laconica Boiss. & Orph.
Onosma orphanidis Boiss.
EFK :248 FAe :456 FCA :41 FCSI :15 FT 6:369 MFG 2:33
L/395 L/518 L/530 L/554 L/607 L/1262a L/1871 L/1963 L/2013
996
997
998
999
Onosma erecta Sm. subsp. malickyi Teppner
Boraginaceae
MFG 2:34
L/530 L/607 L/1963
Onopordum rhodense Boiss. ex Rech.f.
Asteraceae
Onopordum tauricum Willd. var. mazzocchianum Pamp.
FAe :656 FT 5:361
L/197 L/707
995
1000
Onopordum messeniacum Halácsy
Asteraceae
FE 4:247 FGC 2:122
L/607 L/765 L/1963
994
Taxon
Onopordum majoris Beauverd
Asteraceae
Onopordon majorii Beauverd (sphalm.)
EFK :324 FAe :657 FCA :69 FE 4:248 FT 5:364
L/111 L/539 L/540 L/796
Taxon
Qualifiers
993
Taxon
Number
+
+
+
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+
.
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WAe
+
+
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+
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+
StE
Pe
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SPi
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IoI
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+
.
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NPi
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EC
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+
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+
.
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NC
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+
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NE
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NAe
.
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Kik
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+
+
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KK
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+
+
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+
.
+
EAe
Distribution
Qualifier
296 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Onosma euboica Rech.f.
Boraginaceae
FE 3:93 MCL 1:107 MFG 2:36
L/1262a L/1871 L/1901
Onosma kaheirei Teppner
Boraginaceae
Onosma erecta Sm. var. pubiflora Halácsy
FT 11:189 MFG 2:34
L/530 L/539 L/540 L/558 L/759I L/1769 L/1871 L/1963 L/2201
Onosma kittanae Strid
Boraginaceae
L/1894 L/1901
Onosma leptantha Heldr.
Boraginaceae
FE 3:94 MCL 1:108 MFG 2:32
L/512 L/607 L/1963
Onosma paradoxa Janka
Boraginaceae
Onosma aucheriana DC. subsp. pallida (Boiss.) Hayek var. paradoxa
(Janka) Hayek
FE 3:93 MCL 1:108 MFG 2:35
Onosma pygmaea Riedl
Boraginaceae
MCL 1:110 MFG 2:31
L/1157 L/1901
Onosma sangiasense Teppner & Iatroú
Boraginaceae
MFG 2:32
L/839 L/1963
Onosma spruneri Boiss.
Boraginaceae
FE 3:94 MCL 1:110 MFG 2:31
L/369 L/530 L/607 L/1963
Onosma stridii Teppner
Boraginaceae
MFG 2:35 RDB :380
L/558 L/995 L/1963
1001
1002
1003
1004
1005
1006
1007
1008
1009
.
.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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+
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+
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+
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+
.
PATTERNS OF PLANT ENDEMISM IN GREECE 297
Ophrys argolica H.Fleischm. subsp. aegaea (Kalteisen &
H.R.Reinhard) H.A.Pedersen & Faurh.
Orchidaceae
Ophrys aegaea Kalteisen & H.R.Reinhard
Ophrys argolica sensu auct. karp., non H.Fleischm.
EFK :435 EFK :436 FAe :813 FCA :188 FCA :188 FCSI :111 FE 5:347 FE 5:347 FT 11:295 FT 8:487 FT 8:487 RDB :382
L/271 L/430 L/526 L/567 L/611 L/1463 L/1797 L/1816 L/2185 L/2256 L/2257
Ophrys argolica H.Fleischm. subsp. argolica
Orchidaceae
Ophrys argolica H.Fleischm. subsp. argolica
Ophrys ferrum-equinum Desf. subsp. argolica (H.Fleischm.) Soó
EFK :436 FAe :813 FCA :188 FE 5:347 FT 11:295 FT 8:487 RDB :382
L/138 L/194 L/197 L/271 L/273 L/430 L/526 L/611 L/869 L/872 L/1463 L/1797 L/1963 L/2185 L/2256 L/2257
1015
Ophrys andria P.Delforge subsp. halkionis (G.Kretzschmar &
H.Kretzschmar) Kreutz
Orchidaceae
Ophrys thesei P.Delforge
Ophrys andria P.Delforge var. halkionis G.Kretzschmar &
H.Kretzschmar
L/1793 L/2185 L/2256 L/2257 L/2260
1013
1014
Ophrys andria P.Delforge subsp. andria
Orchidaceae
L/744 L/2185 L/2256 L/2257 L/2260
1012
?
Ophrys andria P.Delforge
Orchidaceae
Ophrys fuciflora (F.W.Schmidt) Moench subsp. andria (P.Delforge)
Faurh.
Ophrys holoserica (Burm.f.) Greuter subsp. andria (P.Delforge) Faurh.
L/744 L/1793 L/2185 L/2256 L/2257 L/2260
1011
Taxon
Ophrys aesculapii Renz
Orchidaceae
Ophrys aesculapii Renz subsp. pseudoaranifera Renz
Ophrys renzii (Soó) Soó
Ophrys sphegodes Mill. subsp. aesculapii (Renz) Soó
FE 5:346 FT 11:292
L/526 L/530 L/607 L/611 L/1347 L/1797 L/1871 L/1963 L/2185 L/2256 L/2257
Taxon
Qualifiers
1010
Taxon
Number
.
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+
.
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+
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+
StE
.
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+
Pe
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SPi
IoI
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+
WAe
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NPi
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?
EC
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NC
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NE
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NAe
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+
+
+
+
.
Kik
.
+
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KK
.
.
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.
?
EAe
Distribution
Qualifier
298 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
?
?
1018
1019
1020
Ophrys argolica H.Fleischm. subsp. icariensis (M.Hirth & H.Spaeth)
Kreutz
Orchidaceae
Ophrys icariensis M.Hirth & H.Spaeth
FT 11:292
L/539 L/540 L/611 L/811 L/815 L/1797 L/2185 L/2256 L/2257
?
1017
Ophrys cretica (Vierh.) E.Nelson
Orchidaceae
Ophrys ‘tuberculata-cretica’
Ophrys doerfleri auct., non H.Fleischm.?
EFK :437 FAe :821 FCA :189 FCSI :112 FCSII : FE 5:348 FT 8:489
L/194 L/197 L/526 L/611 L/776 L/791 L/807 L/819 L/869 L/913 L/1463 L/1797 L/1816 L/2093 L/2256 L/2257
Ophrys candica E.Nelson ex Soó) H.Baumann & Künkele subsp.
lacaena (P.Delforge) Kreutz
Orchidaceae
Ophrys lacaena P.Delforge
Ophrys minoa sensu P.Delforge p.p.
L/2185
Ophrys candica (E.Nelson ex Sóo) H.Baumann & Künkele subsp.
cytherea B.Baumann & H.Baumann
Orchidaceae
L/872 L/2185
Ophrys argolica H.Fleischm. subsp. icariencis (M.Hirth & H.Spaeth)
Kreutz
Orchidaceae
Ophrys icariencis M.Hirth & H.Spaeth
L/2256 L/2258
1016
+
+
.
.
.
.
?
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+
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+
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+
+
+
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.
+
.
.
+
+
PATTERNS OF PLANT ENDEMISM IN GREECE 299
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Ophrys ferrum-equinum Desf. subsp. gottfriediana (Renz) E.Nelson
Orchidaceae
Ophrys gottfriediana Renz
Ophrys spruneri Nyman subsp. gottfriediana (Renz) Soó
Ophrys ferrum-equinum Desf. var. gottfriediana (Renz) Biel
FAe :813 FCA :189 FE 5:347
L/119 L/526 L/607 L/611 L/803 L/1123 L/1312 L/1463 L/1797 L/2185 L/2256 L/2257
Ophrys fusca Link subsp. calocaerina (Devillers-Tersch. & Devillers)
Kreutz
Orchidaceae
Ophrys calocaerina Devillers-Tersch. & Devillers
L/1797 L/2185 L/2256 L/2257 L/2258
?
?
1023
1024
1025
Ophrys fusca Link subsp. creberrima (Renz) H.Kretzschmar
Orchidaceae
Ophrys ‘creberrima-fusca’ Paulus
Ophrys bilunulata auct. cret. non Risso
Ophrys creberrima Paulus
FCSII :
L/611 L/791 L/1113 L/1429 L/1463 L/1797 L/1816 L/2185 L/2256 L/2257
Ophrys cretica (Vierh.) E.Nelson subsp. cretica
Orchidaceae
Ophrys cretica (Vierh.) E.Nelson subsp. bicornuta H.Kretzschmar &
R.L.Jahn
Ophrys cretica (Vierh.) E.Nelson subsp. beloniae G.Kretzschmar &
H.Kretzschmar
Ophrys kotschyi H.Fleischm. & Soó subsp. cretica H.Sundermann.
Ophrys spruneri Nyman f. cretica Vierh.
Ophrys spruneri Nyman subsp. cretica (Vierh.) Renz
FCSII :
L/1463 L/1816 L/2256 L/2257
1022
Taxon
Ophrys cretica (Vierh.) E.Nelson subsp. ariadnae (Paulus)
H.Kretzschmar
Orchidaceae
Ophrys kotschyi H.Fleischm. & Soó subsp. ariadnae (Paulus) Faurh.
Ophrys ariadnae Paulus, nom. inval.
Ophrys ‘albifrons-cretica’
Ophrys cretica (Vierh) E.Nelson subsp. karpathensis E.Nelson
Ophrys cretica (Vierh) E.Nelson subsp. naxia E.Nelson?
Ophrys doerfleri H.Fleischm.
Ophrys karpathensis (E.Nelson) Paulus & Gack, nom. inval.
EFK :437 FAe :821 FCA :189 FCSI :112 FE 5:348 FT 8:489
L/194 L/567 L/611 L/791 L/807 L/819 L/872 L/1463 L/1797 L/1816 L/1816 L/2256 L/2257
Taxon
Qualifiers
1021
Taxon
Number
+
.
.
+
.
.
+
+
.
+
.
.
IoI
.
.
?
.
WAe
StE
+
+
+
+
Pe
.
.
.
.
+
.
.
NPi
+
.
.
SPi
.
.
.
.
.
EC
.
.
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.
.
NC
.
.
.
.
.
NE
.
.
.
.
.
NAe
?
.
.
+
+
Kik
+
.
.
+
+
KK
.
.
.
+
?
EAe
*
Distribution
Qualifier
300 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Ophrys fusca Link subsp. perpusilla (Devillers-Tersch. & .Devillers)
H.Kretzschmar
Orchidaceae
Ophrys perpusilla Devillers-Tersch. & Devillers
Ophrys funerea auct. non Viviani
Ophrys punctulata auct. non Renz
L/2185 L/2256 L/2257
Ophrys fusca Link subsp. parosica (P.Delforge) Kreutz
Orchidaceae
Ophrys parosica P.Delforge
L/744 L/1429 L/2185 L/2256 L/2257 L/2258
1031
?
1029
Ophrys fusca Link subsp. eptapigiensis (Paulus) Kreutz
Orchidaceae
Ophrys eptapigiensis Paulus
Ophrys fusca Link subsp. eptapigiensis (Paulus) Kreutz
L/1978 L/2185 L/2256 L/2257 L/2258
Ophrys fusca Link subsp. parvula (Paulus) Kreutz
Orchidaceae
Ophrys parvula Paulus
L/1978 L/2185 L/2256 L/2257
?
1028
Ophrys fusca Link subsp. creticola (Paulus) H.Kretzschmar
Orchidaceae
Ophrys ‘Jouchtas-fusca’
Ophrys ‘Kreta-fusca, gross, früh’ Paulus nom. prov.
Ophrys creticola Paulus
EFK :431 FCSI :114 FCSII :
L/611 L/791 L/1113 L/1429 L/1816 L/2185 L/2256 L/2257
Ophrys fusca Link subsp. cressa (Paulus) H.Kretzschmar
Orchidaceae
Ophrys cressa Paulus
Ophrys fusca Link subsp. cressa (Paulus) H.Kretzschmar
Ophrys ‘merula-fusca’ Paulus nom. prov.
Ophrys ‘Thripti-fusca spät’ Paulus nom. prov.
EFK :433 FCSI :112 FCSII :
L/823 L/1113 L/1429 L/1463 L/1816 L/2185 L/2256 L/2257
1030
?
1027
!
?
1026
.
.
.
.
.
.
+
.
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+
.
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+
.
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+
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+
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+
+
+
+
?
+
.
.
*
PATTERNS OF PLANT ENDEMISM IN GREECE 301
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Ophrys heldreichii Schltr. subsp. polyxo (J.Mast, M.-A.Garnier,
Devillers-Tersch. & Devillers) Kreutz
Orchidaceae
Ophrys polyxo J.Mast, M.-A.Garnier, Devillers-Tersch. & Devillers
L/2184 L/2256 L/2257 L/2258
?
?
1036
1037
Ophrys holoserica (Burm.f.) Greuter subsp. aeoli (P.Delforge) Kreutz
Orchidaceae
Ophrys aeoli P.Delforge
L/1793 L/2185 L/2256 L/2257 L/2258
Ophrys heldreichii Schltr. subsp. heldreichii
Orchidaceae
Ophrys cornuta Steven ex M.Bieb. subsp. heldreichii (Schltr.) Renz.
Ophrys oestrifera Bieb. subsp. heldreichii (Schltr.) E.Nelson
Ophrys scolopax Cav. subsp. heldreichii (Schltr.) E.Nelson
EFK :438 FAe :817 FCA :189 FCSI :112 FE 5:348 FT 8:496
L/122 L/194 L/197 L/439 L/526 L/611 L/743 L/765 L/776 L/791 L/796 L/809 L/819 L/869 L/872 L/928 L/1313 L/1463 L/1797 L/1816 L/1871 L/1917 L/2181 L/2185 L/2256 L/2257 L/2258
1035
Ophrys grammica (E.Willing & B.Willing) Devillers-Tersch. &
Devillers
Orchidaceae
Ophrys mammosa Desf. subsp. oodcheila (Renz) A.Riechelman
Ophrys sphegodes Mill. subsp. grammica (B.Willing & E.Willing) Kreutz
Ophrys mammosa Desf. subsp. grammica B.Willing & E.Willing
L/526 L/611 L/1797 L/2185 L/2256 L/2257
?
1034
!
Ophrys gortynia (H.Baumann & Künkele) Paulus
Orchidaceae
Ophrys sphegodes Mill. subsp. gortynia H.Baumann & Künkele
EFK :436 FCA :190 FCSI :113
L/611 L/743 L/744 L/791 L/819 L/959 L/1463 L/1797 L/1816 L/2185 L/2256 L/2257
1033
Taxon
Ophrys fusca Link subsp. thriptiensis (Paulus) H.Kretzschmar
Orchidaceae
Ophrys thriptiensis Paulus
Ophrys fusca Link subsp. thriptiensis (Paulus) H.Kretzschmar
Ophrys ‘bicolor-fusca’ Paulus nom. prov.
Ophrys ‘Thripti-fusca früh’ Paulus nom. prov.
EFK :433 FCSI :112 FCSII :
L/611 L/1429 L/1463 L/1816 L/2185 L/2256 L/2257
Taxon
Qualifiers
1032
Taxon
Number
.
.
.
.
.
.
.
.
.
.
.
?
StE
Pe
.
.
?
.
.
.
WAe
.
.
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+
.
.
IoI
.
.
.
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.
.
SPi
.
.
.
+
.
.
NPi
.
.
.
.
.
.
EC
.
.
.
+
.
.
NC
.
.
.
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.
.
NE
.
.
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.
.
.
NAe
+
.
+
.
+
.
Kik
.
.
+
.
+
+
KK
.
+
?
.
.
?
EAe
Distribution
Qualifier
302 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
?
?
!
!
!
1040
1041
1042
1043
1044
?
1039
?
?
?
?
1038
Ophrys oestrifera M.Bieb. subsp. leptomera (P.Delforge) Kreutz
Orchidaceae
Ophrys leptomera P.Delforge
L/1796 L/2185
Ophrys oestrifera M.Bieb. subsp. dodekanensis (H.Kretzschmar &
Kreutz) Kreutz
Orchidaceae
Ophrys dodekanensis H.Kretzschmar & Kreutz
L/1977 L/2185 L/2256 L/2257 L/2258
Ophrys mammosa Desf. subsp. serotina B.Willing & E.Willing
Orchidaceae
Ophrys hystera Kreutz & R.Peter
L/2185 L/2256 L/2257 L/2258
Ophrys macedonica (H.Fleischm. ex Soó) Devillers-Tersch. &
Devillers
Orchidaceae
Ophrys mammosa Desf. subsp. macedonica (H.Fleischm. ex Soó) Soó
nom. inval.
Ophrys sphegodes auct. p.p.
Ophrys aranifera Huds. var. macedonica H.Fleischm., nom. nud.
Ophrys aranifera Huds. subsp. x macedonica H.Fleischm. ex Soó
L/1797 L/2185 L/2256 L/2257
Ophrys iricolor Desf. subsp. astypaleica (P.Delforge) Kreutz
Orchidaceae
Ophrys astypalaeica P.Delforge
L/1793 L/1795 L/2185 L/2256 L/2257 L/2258
Ophrys holoserica (Burm.f.) Greuter subsp. helios (Kreutz) Kreutz
Orchidaceae
Ophrys ‘cressa-holoserica’ Paulus nom. nud.
Ophrys karpatica A.Alibertis nom. nud.
Ophrys helios Kreutz
FCSII :
L/2185 L/2256 L/2257 L/2258
Ophrys holoserica (Burm.f.) Greuter subsp. halia (Paulus) Kreutz
Orchidaceae
Ophrys halia Paulus
L/1814 L/2185 L/2256 L/2257 L/2258
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*
PATTERNS OF PLANT ENDEMISM IN GREECE 303
Ophrys omegaifera H.Fleischm. subsp. fleischmannii (Hayek)
Del Prete
Orchidaceae
Ophrys fleischmannii Hayek
Ophrys funerea Viviani subsp. fleischmannii Soó
Ophrys fusca Link subsp. fleischmannii (Soó) Soó
Ophrys omegaifera H.Fleischm. var. fleischmannii (Hayek) Soó
EFK :434 FCA :189 FE 5:346 FT 11:295 FT 8:482
L/430 L/526 L/611 L/738 L/776 L/791 L/819 L/869 L/913 L/1463 L/1816 L/2185 L/2185 L/2256 L/2257
Ophrys sphegodes Mill. subsp. cretensis H.Baumann & Künkele
Orchidaceae
Ophrys cretensis (H.Baumann & Künkele) Paulus
Ophrys araneola Rchb. subsp. cretensis (H.Baumann & Künkele) Kreutz
Ophrys araneola sensu auct. cret., non Rchb. p.p.
Ophrys sphegodes Mill. subsp. cretensis H.Baumann & Künkele
EFK :436 FCA :190 FCSI :113
L/526 L/611 L/743 L/791 L/819 L/959 L/1463 L/1797 L/1816 L/2185 L/2256 L/2257 L/2258
Ophrys sphegodes Mill. subsp. zeusii (M.Hirth) Kreutz
Orchidaceae
Ophrys zeusii M.Hirth
Ophrys sphegodes Mill. subsp. zeusii (M. Hirth) Kreutz
1047
1048
1049
Ophrys oestrifera M.Bieb. subsp. schlechteriana (Soó) Kreutz
Orchidaceae
Ophrys schlechteriana (Soó) Devillers-Tersch. & Devillers
Ophrys fuciflorra ( F.W. Schmidt ) Moench f. pseudoestrifera Renz
Ophrys heldreichii Schlechter var. schlechteriana (Soó) Soó
Ophrys heldreichii Schlechter subsp. schlechteriana (Soó) Kreutz
FT 11:287
L/1797 L/2184 L/2185 L/2256 L/2257 L/2258
Taxon
Ophrys omegaifera H.Fleischm. subsp. basilissa (C.Alibertis,
A.Alibertis & H.R.Reinhard) H.Kretzschmar
Orchidaceae
Ophrys basilissa C.Alibertis & A.Alibertis
Ophrys omegaifera H.Fleischm. subsp. omegaifera var. basilissa
(C.Alibertis, A.Alibertis & H.R.Reinhard) Faurh.
EFK :434 FCA :188 FCSI :111 FT 11:292
L/611 L/742 L/743 L/900 L/1463 L/1816 L/1957 L/2185 L/2256 L/2257
?
?
Taxon
Qualifiers
1046
1045
Taxon
Number
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StE
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WAe
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SPi
IoI
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NPi
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EC
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NC
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NAe
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KK
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EAe
Distribution
Qualifier
304 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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Ophrys × delphinensis O.Danesch & E.Danesch (pro hybr.)
Orchidaceae
L/526 L/607 L/611 L/1313 L/1797 L/1871 L/2185
Ophrys × diakoptensis Bayer
Orchidaceae
L/607
Ophrys × kiparissensis E.Willing & B.Willing
Orchidaceae
L/607
1053
1054
1055
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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Ophrys spruneri Nyman subsp. spruneri
Orchidaceae
Ophrys ferrum-equinum Desf. subsp. spruneri (Nyman) Camus
Ophrys hiulka Spruner ex Rchb.f.
Ophrys sphegodes Mill. subsp. spruneri (Nyman) E.Nelson nom. inval.
Ophrys spruneri Nyman subsp. spruneri
EFK :437 FAe :820 FCA :190 FCSI :114 FE 5:347 FT 8:486
L/321 L/539 L/540 L/607 L/611 L/791 L/811 L/819 L/869 L/1123 L/1312 L/1313 L/1463 L/1797 L/1800 L/1816 L/1871 L/1871 L/1957 L/1963 L/2185 L/2256 L/2257
+
+
1052
Ophrys spruneri Nyman subsp. grigoriana (G.Kretzschmar &
H.Kretzschmar) H.Kretzschmar
Orchidaceae
Ophrys grigoriana G.Kretzschmar & H.Kretzschmar
Ophrys sphaciotica H.Fleischm.
FCSII :
L/791 L/1463 L/1816 L/2256 L/2257
1051
?
Ophrys spruneri Nyman
Orchidaceae
EFK :437 FAe :820 FCA :190 FCSI :114 FCSII : FE 5:347 FT 8:486
L/321 L/539 L/540 L/607 L/611 L/791 L/791 L/811 L/819 L/869 L/1123 L/1312 L/1313 L/1463 L/1463 L/1797 L/1800 L/1816 L/1816 L/1871 L/1871 L/1957 L/1963 L/2185 L/2256 L/2257
1050
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PATTERNS OF PLANT ENDEMISM IN GREECE 305
Anacamptis papilionacea (L.) R.M.Bateman, Pridgeon & M.W.Chase
subsp. messenica (Renz) H.Kretzschmar, Eccarius & H.Dietr.,
[as Orchis papilionacea L. var. messenica Renz]
Orchidaceae
L/2185
Orchis prisca Hautz.
Orchidaceae
Orchis patens Desf. subsp. falcicalcarata Wildhaber
Orchis patens Desf. subsp. nitidifolia Teschner
Orchis patens Desf. subsp. orientalis sensu Rechinger 1943
Orchis spitzelii Sauter ex Koch subsp. nitidifolia (Teschner) Soó
Orchis spitzelii sensu Seitz 1978, Henke 1983
EFK :427 FAe :836 FCA :192 FCSI :116 FE 3:340 MFG 1:881 RDB :390
L/430 L/611 L/791 L/813 L/819 L/867 L/870 L/1043 L/1043 L/1185 L/1463 L/2185
Orchis sitiaca (Renz.) P.Delforge
Orchidaceae
Orchis anatolica Boiss. subsp. sitiaca Renz
Orchis anatolica Boiss. var. sitiaca (Renz) Soó
EFK :427 FCA :191 FCSI :116 FCSII : FE 5:341
L/611 L/791 L/819 L/1463 L/2185
Origanum calcaratum Juss.
Lamiaceae
Amaracus tournefortti (Aiton) Benth.
Origanum tournefortii Aiton
EFK :266 FCA :97 FE 3:172 FT 10:206 FT 7:300 MCL 3:305 MFG 2:138
L/145 L/197 L/255 L/279
1058
1059
1060
1061
Anacamptis boryi (Rchb.f.) R.M.Bateman, Pridgeon & M.W.Chase
[as Orchis boryi Rchb.f.]
Orchidaceae
Anacamptis boryi (Rchb.f.) R.M.Bateman, Pridgeon & M.W.Chase
Orchis morio L. subsp. boryi (Rchb.f.) Soó
EFK :424 FCA :192 FE 5:338
L/607 L/611 L/791 L/819 L/1313 L/1463 L/1871 L/1963 L/2181 L/2185
Taxon
Anacamptis papilionacea (L.) R.M.Bateman, Pridgeon &
M.W.Chase subsp. alibertis (G.Kretzschmar & H.Kretzschmar)
H.Kretzschmar, Eccarius & H.Dietr. [as Orchis papilionacea
L. var. alibertis (G.Kretzschmar & H.Kretzschmar) P.Delforge]
Orchidaceae
Orchis papilionacea L. var. alibertis G.Kretzschmar & H.Kretzschmar
FCSII :
L/1813
Taxon
Qualifiers
1057
1056
Taxon
Number
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StE
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Pe
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WAe
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SPi
IoI
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NPi
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NC
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NAe
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Kik
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KK
+
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EAe
Distribution
Qualifier
306 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Origanum lirium Halácsy
Lamiaceae
Amaracus lirius (Halácsy) Hayek
FAe :531 FE 3:172 FGC 2:554 MCL 3:306
L/99 L/323b L/608 L/1871 L/1871
Origanum microphyllum (Benth.) Boiss.
Lamiaceae
Majorana microphylla Benth.
EFK :266 FCA :97 FE 3:172 MCL 3:306 MFG 2:137
Origanum scabrum Boiss. & Heldr.
Lamiaceae
Origanum pulchrum Boiss. & Heldr.
Amaracus pulcher (Boiss. & Heldr.) Briq.
Amaracus scaber (Boiss. & Heldr.) Briq.
Origanum scabrum Boiss. & Heldr. subsp. pulchrum (Boiss. & Heldr.)
P.H.Davis
FAe :531 FCSI :52 FCSII : FE 3:172 FGC 2:553 MCL 3:307 MFG 2:138
L/323b L/607 L/1871 L/1963
Origanum symes A.Carlström
Lamiaceae
FT 10:206 MCL 3:307 RDB :396
L/197 L/279
Origanum vetteri Briq. & Barbey
Lamiaceae
Amaracus vetteri (Briq. & Barbey) Hayek
EFK :266 FCA :97 FE 3:172 MCL 3:307 RDB :398
L/111
Ornithogalum atticum Boiss. & Heldr.
Asparagaceae
Ornithogalum glaucophyllum Baker?
Ornithogalum montanum Cyr. subsp. atticum (Bois. & Heldr.) Nyman
FE 5:38
L/111 L/447 L/512 L/515 L/607 L/1223 L/1711 L/1832d L/1871 L/1963 L/2181
1063
1064
1065
1066
1067
1068
!
Origanum dictamnus L.
Lamiaceae
Amaracus dictamnus (L.) Benth.
EFK :266 FCA :97 FE 3:172 FT 7:312 MCL 3:305 MFG 2:138 RDB :394
L/1583
1062
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+
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 307
!
1069
Ornithogalum creticum Zahar.
Asparagaceae
Ornithogalum visianicum auct. non Tommasini, nec Rechinger 1944, &
Greuter, 1973
EFK :354 FCA :184 FE 5:37
L/255 L/268 L/430 L/495 L/537 L/1049 L/1220 L/2093
Ornithogalum dictaeum Landström
Asparagaceae
EFK :354 FCSII :
L/515 L/869 L/1583
Ornithogalum dictaeum Landström subsp. dictaeum
Asparagaceae
EFK :354 FCSII :
L/515 L/1583
Ornithogalum dictaeum Landström subsp. naxense Landström
Asparagaceae
L/515 L/869
Ornithogalum exaratum Zahar.
Asparagaceae
FE 5:40 FT 8:243
L/515 L/608 L/1220 L/1223 L/1871
Ornithogalum fimbriatum Willd. subsp. gracilipes (Zahar.)
Landström
Asparagaceae
Ornithogalum gracilipes Zahar.
Ornithogalum gracilipes Zahar.
MFG 2:689
L/121 L/515 L/515 L/530 L/1224 L/1963 L/2200
Ornithogalum prasinantherum Zahar.
Asparagaceae
FE 5:37 FT 8:233
L/530 L/607 L/1220 L/1223 L/1830 L/1957 L/1963
Ornithogalum pumilum Zahar.
Asparagaceae
FCA :185 FT 10:224 FT 8:563 MFG 2:690
L/1223 L/2237
1071
1072
1073
1074
1075
1076
1077
Ornithogalum brachystylum Zahar.
Asparagaceae
L/197 L/1222
Taxon
1070
?
Taxon
Qualifiers
Taxon
Number
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StE
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Pe
+
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IoI
+
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WAe
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SPi
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NPi
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EC
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NC
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NE
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NAe
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Kik
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KK
+
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EAe
Distribution
Qualifier
308 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Paeonia clusii Stern subsp. clusii
Paeoniaceae
Paeonia cretica Tausch (nom. inval.)
AFE 9:17 EFK :102 FAe :177 FCA :122 FE 1:293 FHe 2:78 MCL 4:267
L/111 L/468 L/518 L/522 L/527 L/554 L/856
Paeonia clusii Stern subsp. rhodia (Stearn) Tzanoud.
Paeoniaceae
Paeonia corallina sensu Rechinger
Paeonia officinalis sensu Rechinger 1943
Paeonia rhodia Stearn
AFE 9:17 FAe :177 FE 1:293 FHe 2:78 FT 1:205 FT 10:22 MCL 4:267 RDB :400
L/138 L/197 L/468 L/527
Paeonia mascula (L.) Miller subsp. hellenica Tzanoud.
Paeoniaceae
AFE 9:21 FE 1:294 FHe 2:80 MCL 4:268
B/3060b L/468 L/527 L/607 L/1278 L/1871 L/2181
1081
1082
1083
Paeonia mascula (L.) Miller subsp. icarica Tzanoud.
Paeoniaceae
Paeonia mascula (L.) Miller subsp. hellenica Tzanoud. var. icarica
Stearn
FHe 2:80 FT 10:22 MCL 4:268
B/3060b L/468 L/527 L/539 L/540
Paeonia clusii Stern
Paeoniaceae
Paeonia officinalis L. var. glabra (Boiss.) Hayek
Paeonia peregrina Miller var. cretica Huth
Paeonia peregrina Miller var. glabra Boiss.
AFE 9:17 FAe :177 FCA :122 FE 1:293 FHe 2:77 FT 1:205 FT 10:22 MCL 4:267 RDB :400
L/111 L/468 L/522 L/527 L/554 L/856
1080
1084
Orobanche baumanniorum Greuter
Orobanchaceae
MCL 4:257 MFG 2:272
L/530 L/923 L/1963
1079
!
Ornithogalum sphaerolobum Zahar.
Asparagaceae
FT 10:225 FT 8:563
L/192 L/515 L/1220 L/1223
1078
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+
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+
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+
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+
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+
.
+
.
+
PATTERNS OF PLANT ENDEMISM IN GREECE 309
Paronychia cephalotes (MB.) Besser subsp. thracica Chaudhri
Caryophyllaceae
AFE 6:143 FHe 1:226 MCL 1:233
Petromarula pinnata (L.) A.DC.
Caryophyllaceae
Petromarula oxyloba Gand.
Petromarula pinnataoxyloba (L.) A.DC. var. oxyloba (Gand.) Hayek
Phyteuma pinnatum L.
EFK :297 FAe :605 FCA :43 FCSI :16 FE 4:95 MCL 1:150
L/518 L/554 L/856 L/1766
Petrorhagia candica P.W.Ball & Heywood
Caryophyllaceae
Gypsophila cretica sensu Smith (1809)
Tunica cretica auct. p.p. [non (L.) Fisch. & C.A.Mey.]
Tunica taygetea auct. pro parte, non (Boiss.) P.H.Davis
AFE 7:142 EFK :91 FAe :151 FCA :49 FCSI :20 FE 1:225 FHe 1:337 MCL 1:238
L/255 L/1766
Petrorhagia dianthoides (Sm.) P.W.Ball & Heywood
Caryophyllaceae
Gypsophila dianthoides Sm.
Tunica dianthoides (Sm.) Fisch. & C.A.Mey
AFE 7:145 EFK :91 FAe :151 FCA :49 FCSI :20 FE 1:226 FHe 1:340 MCL 1:238
L/518 L/707 L/916
1088
1089
1090
1091
Paronychia albanica Chaudhri subsp. graeca Chaudhri
Caryophyllaceae
AFE 6:142 FE 1:181 FHe 1:225 MCL 1:231 MFG 1:129
L/530 L/1123 L/1846 L/1963 L/2013
Paeonia parnassica Tzanoud.
Paeoniaceae
Paeonia arietina auct. non G.Anderson
Paeonia peregrina Miller var. latifolia Boiss.
AFE 9:15 FE 1:293 FHe 2:77 MCL 4:269 RDB :402
L/138 L/468 L/527
Taxon
Paronychia bornmuelleri Chaudhri
Caryophyllaceae
AFE 6:143 FE 1:181 FHe 1:226 MCL 1:233 RDB :404
L/973 L/1349
!
Taxon
Qualifiers
1087
1086
1085
Taxon
Number
.
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+
+
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StE
Pe
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WAe
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+
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IoI
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+
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SPi
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NPi
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EC
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+
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+
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NC
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+
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NE
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+
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NAe
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Kik
+
+
+
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KK
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EAe
Distribution
Qualifier
310 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Petrorhagia fasciculata (Marg. & Reut.) P.W.Ball & Heywood
Caryophyllaceae
Gypsophila fasciculata Marg. & Reut.
Tunica fasciculata (Marg. & Reut.) Boiss.
AFE 7:143 EFK :91 FCSI :20 FE 1:226 FHe 1:338 MCL 1:238
L/346 L/798 L/943 L/1123 L/1800 L/2013 L/2200
Petrorhagia glumacea (Bory & Chaub.) P.W.Ball & Heywood
Caryophyllaceae
Dianthus glumaceus Bory & Chaub.
Kohlrauschia glumacea (Bory & Chaub.) Hayek
Tunica glumacea (Chaub. & Bory) Boiss.
AFE 7:148 FE 1:227 FHe 1:342 MCL 1:239
L/121 L/441 L/607 L/1218 L/1244 L/1246 L/1913 L/1963
Petrorhagia graminea (Sm.) P.W.Ball & Heywood
Caryophyllaceae
Gypsophila graminea Sm.
Tunica graminea (Sm.) Boiss.
AFE 7:145 FE 1:226 FHe 1:339 MCL 1:238
L/339 L/607 L/943 L/1123 L/1830 L/1963
Petrorhagia grandiflora Iatroú
Caryophyllaceae
AFE 7:145 FE 1:226 FHe 1:340 RDB :408
L/607 L/916 L/1963
Petrorhagia illyrica (L.) P.W.Ball & Heywood subsp. taygetea
P.W.Ball & Heywood
Caryophyllaceae
Tunica cretica sensu Hayek p.p., Bald. 1895, Rechinger 1944
Tunica illyrica (Ard.) Fisch. var. taygetea Boiss.
Tunica taygetea (Boiss.) P.H.Davis
AFE 7:141 EFK :91 FCA :49 FCSI :20 FE 1:225 FHe 1:335 MCL 1:239
L/518 L/554 L/607 L/707 L/1963
Petrorhagia ochroleuca (Sm.) P.W.Ball & Heywood
Caryophyllaceae
Gypsophila ochroleuca Sm.
Tunica ochroleuca Sm. Fisch. & C.A.Mey.
AFE 7:142 FE 1:225 FHe 1:336 MCL 1:239
L/607 L/607 L/607 L/943
1092
1093
1094
1095
1096
1097
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 311
Peucedanum achaicum Halácsy
Apiaceae
FE 2:362 FGS 1:42 RDB :410
L/607 L/683 L/1844 L/1938b L/1963
Peucedanum stridii Hartvig
Apiaceae
MFG 1:720
L/389
Peucedanum vourinense (Leute) Hartvig
Apiaceae
Peucedanum longifolium Waldst. & Kit. subsp. vourinense Leute
MFG 1:716
L/1901
Phagnalon pygmaeum (Sieber) Greuter
Asteraceae
Phagnalon pumilum (Sm.) DC.
EFK :309 FCA :69 FCSI :34 FE 4:133
Phitosia crocifolia (Boiss. & Heldr.) Kamari & Greuter
Asteraceae
Crepis crocifolia Boiss. & Heldr.
Hieracioides crocifolium (Boiss. & Heldr.) O.Kuntze
FE 4:352 MFG 2:591 RDB :210
L/512 L/607 L/1104 L/1685 L/1963
Phleum exaratum Hochst. ex Griseb. subsp. aegaeum (Vierh.) Dogan
Poaceae
Phleum arenarium auct. p.p., non L.
Phleum arenarium L. subsp. aegaeum Vierh.
Phleum graecum Boiss. & Heldr. subsp. aegaeum (Vierh.) Greuter
EFK :399 FAe :800 FCA :173 FE 5:241 FT 9:397
L/111 L/119 L/128 L/197 L/272 L/430 L/540 L/607 L/776 L/796 L/869 L/913 L/1235 L/1627 L/1917 L/2181
1099
1100
1101
1102
1103
1104
Taxon
Petrorhagia phthiotica (Boiss. & Heldr.) P.W.Ball & Heywood
Caryophyllaceae
Tunica ochroleuca (Sm.) Fisch & C.A.Mey. var. phtiotica (Boiss. &
Heldr.) Hayek
Tunica phtiotica Boiss. & Heldr.
AFE 7:143 FE 1:226 FHe 1:338 MCL 1:239 MFG 1:175
L/607 L/1467 L/1963 L/2013
Taxon
Qualifiers
1098
Taxon
Number
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Distribution
Qualifier
312 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Pimpinella rigidula (Boiss. & Orph.) H.Wolff
Apiaceae
Reutera rigidula Boiss. & Orph.
FE 2:331 FGC 1:685 FGS 2:46
L/607
Pimpinella tragium Vill. subsp. depressa (DC.) Tutin
Apiaceae
Pimpinella depressa DC.
Pimpinella tragium Vill var. depressa (DC.) Boiss.
EFK :212 FCA :152 FE 2:331 MFG 1:679
B/3060f L/607 L/1190 L/1766
Plantago atrata Hoppe subsp. graeca (Halácsy) Holub
Plantaginaceae
Plantago atrata Hoppe f. nana Pilger
Plantago graeca Halácsy
Plantago montana Lam. var. graeca (Halácsy) Halácsy
MCL 4:297 MFG 2:337
L/121 L/513 L/832 L/1846 L/1963 L/2200
1108
1109
1110
Poa cephalonica H.Scholz
Poaceae
MFG 2:771
L/304 L/889 L/1123
Pimpinella pretenderis (Heldr.) Orph. ex Halácsy
Apiaceae
Pimpinella tragium var. maritima Boiss.
EFK :212 FCA :152 FE 2:332 MFG 1:680 RDB :416
L/111 L/128 L/196 L/206 L/273 L/938
1107
1111
Phlomis pichleri Vierh.
Lamiaceae
Phlomis fruticosa sensu Fors.-Major & Barbey 1849, Rechinger 1943
EFK :259 FAe :511 FCA :98 FE 3:145 MCL 3:311
L/273 L/551
1106
!
Phlomis lanata Willd.
Lamiaceae
EFK :259 FCA :98 FE 3:145 MCL 3:311
L/856 L/1766
1105
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PATTERNS OF PLANT ENDEMISM IN GREECE 313
Poa trichophylla Heldr. & Sart.
Poaceae
Poa trichopoda Heldr. & Sart.
FE 5:165 MFG 2:770
L/269 L/304 L/512 L/513 L/2200
Podospermum roseum (Waldst. & Kit.) Gemeinholzer & Greuter
subsp. peristericum (Formánek) Gemeinholzer & Greuter
Asteraceae
Scorzonera rosea Waldst. & Kit. subsp. peristerica (Formánek) Greuter
Scorzonera purpurea Hausskn. subsp. peristerica Formánek
Scorzonera purpurea Hausskn. subsp. rhodantha (Hausskn.) Maire &
Petitm.
Scorzonera rhodantha Hausskn.
FE 4:339 MFG 2:535
L/121 L/512 L/513 L/1769 L/1902 L/1905 L/2013 L/2200 L/2201
Polygala crista-galli Chodat
Polygalaceae
FE 2:235 MCL 4:347
L/607 L/1963
Polygala helenae Greuter
Polygalaceae
MCL 4:347 RDB :418
L/119 L/1963
Polygala nicaeensis Risso ex Koch subsp. tomentella (Boiss.) Chodat
Polygalaceae
Polygala nicaeensis Risso ex Koch subsp. graeca Chodat
Polygala nicaeensis Risso ex Koch var. tomentella Boiss.
FE 2:234 FGC 1:147 MCL 4:349 MFG 1:580
L/441 L/530 L/607 L/1123 L/1871 L/1963 L/2201
1114
1115
1116
1117
1118
Poa ophiolithica H.Scholz
Poaceae
MFG 2:767
L/269 L/304
!
1113
Taxon
Poa glauca Vahl. subsp. frearitis (Orph. ex Halácsy) H.Scholz
Poaceae
Poa caesia Sm. var. frearitis Orph. ex Halácsy
MFG 2:769
L/304
Taxon
Qualifiers
1112
Taxon
Number
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SPi
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NPi
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Distribution
Qualifier
314 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Polygala rausiana U. Raabe, Kit Tan, Iatroú, Vold & Parolly
Polygalaceae
L/2229
Polygala subuniflora Boiss. & Heldr.
Polygalaceae
FE 2:233 MCL 4:350 MFG 1:578 RDB :420
L/1120 L/1963
Polygonum icaricum Rech.f.
Polygonaceae
AFE 4:10 FE 1:93 FHe 1:78 FT 2:275 MCL 4:359 MFG 1:60
L/533 L/539 L/540 L/575 L/895 L/1252 L/1349 L/1957
Polygonum idaeum Hayek
Polygonaceae
Polygonum aviculare L. var. alpinum Boiss.
Polygonum aviculare var. nanum sensu Halácsy
AFE 4:12 EFK :63 FAe :107 FCA :127 FCSI :72 FE 1:93 FHe 1:79 MCL 4:359 MFG 1:61 RDB :422
L/518 L/522 L/554 L/987
Polygonum papillosum Hartvig
Polygonaceae
FE 1:93 FHe 1:79 MCL 4:359 RDB :424
L/266
Potentilla arcadiensis Iatroú
Rosaceae
AFE 13:259
L/607 L/757 L/1915 L/1963 L/2095
Potentilla deorum Boiss. & Heldr.
Rosaceae
AFE 13:265 FE 2:46 MFG 1:414
L/953 L/1940
Potentilla kionaea Halácsy
Rosaceae
Potentilla apennina subsp. kionaea (Halácsy) Maire & Petitm.
AFE 13:264 FE 2:46 MFG 1:414 RDB :428
L/330 L/512
1119
1120
1121
1122
1123
1124
1125
1126
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PATTERNS OF PLANT ENDEMISM IN GREECE 315
Prunella cretensis Gand.
Lamiaceae
Prunella laciniata (L.) L. var. cretica (Boiss.) Hayek
EFK :263 FCA :98 FE 3:162 MCL 3:312 MFG 2:120
L/253
Pterocephalus perennis Coulter subsp. perennis
Caprifoliaceae
Pterocephalus parnassi Sprengel
Pterocephalus perennis Coulter subsp. parnassi (Sprengel) Vierh.
Scabiosa pterocephala L.
FE 4:68 FGC 1:762 MCL 3:192 MFG 2:358
L/529 L/530 L/607 L/843 L/1871 L/1963 L/1973 L/2013
Ptilostemon gnaphalodes (Cyr.) Sojak subsp. pseudofruticosus
(Pamp.) Greuter
Asteraceae
Chamaepeuce fruticosa auct. non Cnicus fruticosus Desf.
Cirsium fruticosum auct.
Ptilostemon gnaphaloides (Cyr.) Sojak subsp. pseudofruticosus (Pamp.)
Greuter
EFK :323 FCA :70 FE 4:243 FT 5:418
L/447 L/539 L/540 L/607 L/1021 L/1871
Pulmonaria cesatiana (Fenzl & Friedr.) Selvi, Bigazzi, Hilger &
Papini
Boraginaceae
Nonea auriculata Boiss.
Paraskevia cesatiana (Fenzl & Friedr.) W.Sauer & G.Sauer
Nonea cesatiana (Fenzl & Friedr.) Greuter & Burdet
Anchusa cesatiana Fenzl & Friedr.
Pulmonaria auriculata (Boiss.) Halácsy
MCL 1:102 RDB :374
L/607 L/709 L/1162 L/1963 L/2238
Quercus trojana Webb subsp. euboica (Papaioannou) K.I.Chr.
Fagaceae
Quercus euboica Papaioannou
FHe 1:45 RDB :434
L/391 L/1290 L/1871 L/1960
1128
1129
1130
1131
1132
Taxon
Prometheum tymphaeum (Quézel & Contandr.) ’t Hart
Crassulaceae
Sedum tymphaeum Quézel & Contandr.
Helladia tymphaea (Quézel & Contandr.) M.Kral
AFE 12:117 FE 1:335 FHe 2:335 MCL 3:329 MFG 1:354
L/275 L/601
Taxon
Qualifiers
1127
Taxon
Number
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WAe
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KK
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EAe
Distribution
Qualifier
316 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Ranunculus cupreus Boiss. & Heldr.
Ranunculaceae
AFE 8:155 EFK :81 FAe :191 FCA :132 FCSII : FE 1:278 FHe 2:53 MCL 4:429 MFG 1:218
L/758
Ranunculus millii Boiss. & Heldr.
Ranunculaceae
AFE 8:156 FE 1:279 FGC 1:19 FHe 2:54 MCL 4:434
L/607 L/1963
Ranunculus radinotrichus Greuter & Strid
Ranunculaceae
AFE 8:138 EFK :101 FCA :133 FE 1:276 FHe 2:46 MCL 4:438 MFG 1:216 RDB :438
L/253 L/336 L/518 L/721 L/1766
Ranunculus subhomophyllus (Halácsy) Vierh.
Ranunculaceae
Ranunculus subhomophyllus (Halácsy) Vierh. subsp. cylleneus Vierh.
Ranunculus subhomophyllus (Halácsy) Vierh. subsp. parviflorus (Boiss.)
Vierh.
Ranunculus cadmicus Boiss. var. parviflorus Boiss.
Ranunculus subhomophyllus (Halácsy) Vierh. var. grandiflorus Vierh.
Ranunculus cadmicus sensu Halácsy, non Boiss.
Ranunculus chaerophyllos L. var. subhomophyllus Halácsy
AFE 8:158 EFK :102 FAe :190 FCA :133 FCSII : FE 1:279 FHe 2:55 FT 1:181 MCL 4:428 MFG 1:220
L/539 L/540 L/758 L/796 L/1830 L/1871 L/1963
Ranunculus thasius Halácsy
Ranunculaceae
AFE 8:158 FAe :190 FE 1:279 FHe 2:55 FT 1:180 MCL 4:428 MFG 1:220
L/99 L/323b L/443 L/539 L/540 L/869 L/1267a L/1871 L/2181
Ranunculus veronicae N.Böhling
Ranunculaceae
FCSII : FHe 2:56
L/1770
1133
1134
1135
1136
1137
1138
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PATTERNS OF PLANT ENDEMISM IN GREECE 317
Rhamnus sibthorpiana Roemer & Schultes
Rhamnaceae
Oreoherzogia pubescens (Sm.) Vent
Rhamnus pubescens Sm.
FE 2:245 MCL 4:457 MFG 1:588
L/160 L/512 L/607 L/843 L/1830 L/1901 L/1963
Rhinanthus pubescens (Sterneck) Boiss. & Heldr. ex Soó
Orobanchaceae
Alectrolophus pubescens Sterneck
Rhinanthus intermedius Quézel & Contandr.
Rhinanthus major L. var. parnassicus Boiss.
MFG 2:254
L/121 L/395 L/512 L/587 L/1963 L/2013
Rhinanthus sintenisii (Sterneck) Soó
Orobanchaceae
Alectorolophus sintenisii Sterneck
FE 3:278 MFG 2:255
L/1112
Ricotia cretica Boiss. & Heldr.
Brassicaceae
AFE 11:17 EFK :116 FCA :85 FCSI :43 FE 1:358 FHe 2:195 MCL 3:151
L/165 L/518 L/554 L/561 L/780 L/856 L/1096
Ricotia isatoides (Barbey) B.L.Burtt
Brassicaceae
Peltaria isatoides Barbey
AFE 11:17 EFK :117 FAe :243 FCA :85 FCSI :43 FE 1:358 FHe 2:196 MCL 3:151 RDB :440
L/780
1141
1142
1143
1144
1145
Rhamnus guicciardi (Boiss.) Halácsy
Rhamnaceae
Oreoherzogia guicciardi (Boiss.) Halácsy
Rhamnus fallax Boiss. var. guicciardi Boiss.
Rhamnus sibthorpianus Roemer & Schultes (including taxon)
FE 2:245 MCL 4:457 MFG 1:589
L/512 L/587 L/2237
?
1140
Taxon
Reseda tymphaea Hausskn.
Resedaceae
Reseda tymphaea Hausskn. subsp. tymphaea
Reseda epirotica Formánek
Reseda tymphaea Hausskn. var. asperula Halácsy
AFE 12:26 FE 1:420 FGC 1:125 FHe 2:299 MCL 4:449
L/607 L/1963
Taxon
Qualifiers
1139
Taxon
Number
+
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StE
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IoI
WAe
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SPi
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NPi
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EAe
Distribution
Qualifier
318 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Rindera graeca (A.DC.) Boiss. & Heldr.
Boraginaceae
Cynoglossum graecum (A.DC.) Greuter & Burdet
Mattia graeca A.DC.
FE 3:122 MCL 1:77 MFG 2:65
L/160 L/310a L/330 L/512 L/530 L/587 L/607 L/709 L/1871 L/1963 L/1973 L/2013
Rorippa icarica Rech.f.
Brassicaceae
FAe :210 FHe 2:177 FT 1:432 MCL 3:152 RDB :442
L/539 L/540
Ruta chalepensis L. subsp. fumariifolia (Boiss. & Heldr.) Nyman
Rutaceae
Ruta chalepensis L. var. fumariifolia Boiss.
Ruta fumariifolia Boiss. & Heldr.
EFK :184 FAe :283 FCA :140 FCSI :80 FE 2:227
L/268 L/913 L/1049 L/1878 L/1917
Salsola aegaea Rech.f.
Amaranthaceae
Salsola brevifolia sensu d’Urv.
Salsola vermiculata auct. balcan., non L.
AFE 5:84 EFK :71 FAe :124 FCA :55 FE 1:128 FHe 1:135 FT 11:63 MCL 1:311
L/150 L/197 L/201 L/206 L/255 L/268 L/495 L/537 L/554 L/913 L/1049 L/1266 L/1627 L/1710 L/1917 L/2093 L/2181
Salsola carpatha P.H.Davis
Amaranthaceae
Salsola aegaea sensu auct. karp., Rechinger 1943
Salsola vermiculata auct. balcan., non L.
AFE 5:82 EFK :71 FAe :124 FCA :55 FE 1:127 FHe 1:135 MCL 1:309
L/255 L/268 L/273 L/551 L/554 L/1049 L/1710
Salvia eichleriana Heldr. ex Halácsy
Lamiaceae
FE 3:189 FGC 2:481 MCL 3:315
Salvia pomifera L. subsp. pomifera
Lamiaceae
FCA :99 FE 3:190 MCL 3:319
L/151 L/1052
1146
1147
1148
1149
1150
1151
1152
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 319
Satureja acropolitana (Halácsy) Greuter & Burdet
Lamiaceae
Micromeria acropolitana Halácsy
FE 3:168 FGS 1:87 MCL 3:323 RDB :448
L/940 L/2228
Satureja cretica (L.) Briq.
Lamiaceae
Calamintha cretica (L.) Lam.
Melissa cretica L.
EFK :264 FCA :99 FCSII 0: FE 3:167 MCL 3:329 MFG 2:131
L/1941
Satureja hellenica Heldr. ex Halácsy
Lamiaceae
Satureja montana L. var. hellenica (Heldr.) Halácsy
Satureja parnassica Heldr. & Sart. sensu Fl. Aeg. p.p.
FAe :524 FGC 2:551 MCL 3:336 MFG 2:125
L/530 L/1871
1157
1158
1159
Satureja horvatii Silic subsp. macrophylla (Halácsy) Baden
Lamiaceae
Satureja parnassica Heldr. & Sart. ex Boiss. var. macrophylla Halácsy
MFG 2:126
L/121 L/1027 L/2200
Saponaria jagelii Phitos & Greuter
Caryophyllaceae
FHe 1:332 RDB :446
L/313 L/1963
1156
1160
Saponaria aenesia Heldr.
Caryophyllaceae
Saponaria calabrica Guss. var. aenesia Heldr.
AFE 7:138 FE 1:224 FHe 1:331 MCL 1:245
L/1123
1155
?
Sanguisorba cretica Hayek
Rosaceae
Poterium sphacioticum Gand.
Sanguisorba cretica Hayek f. sphaciotica (Gand.) Hayek
AFE 13:135 EFK :136 FAe :304 FCA :137 FCSI :77 FE 2:34
L/518 L/519 L/554 L/1583 L/1766
1154
Taxon
Salvia teddii Turrill
Lamiaceae
FE 3:191 MCL 3:321 MFG 2:166
Taxon
Qualifiers
1153
Taxon
Number
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WAe
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IoI
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SPi
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NPi
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EC
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NC
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
320 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Scabiosa taygetea Boiss. & Heldr. subsp. taygetea
Caprifoliaceae
Scabiosa holocericea Bory
Scabiosa taygetea Boiss. & Heldr. var. pindicola Hausskn.
MFG 2:365
L/395 L/1963 L/2013 L/2200
1166
Scaligeria halophila (Rech.f.) Rech.f.
Apiaceae
Scaligeria cretica (Miller) Boiss. subsp. halophila Rech.f.
EFK :210 FAe :423 FCA :152 FE 2:328
L/268 L/1148 L/1963
Saxifraga sibthorpii Boiss.
Saxifragaceae
Saxifraga cymbalaria sensu Sibth. & Sm.
AFE 12:132 FE 1:443 FHe 2:338 FT 4:258 MFG 1:366
L/587 L/1871 L/1963
1165
1167
Satureja parnassica Heldr. & Sart. ex Boiss. subsp. parnassica
Satureja montana L. subsp. parnassica (Heldr. & Sart.) Maire & Petitm.
Satureja parnassica Heldr. & Sart var. stygia Orph.
FGC 2:552 MCL 3:337 MFG 2:125
L/530 L/607 L/1830 L/1963
1164
!
Satureja parnassica Heldr. & Sart. ex Boiss. subsp. athoa (K. Malý)
Baden
Lamiaceae
Satureja athoa K.Malý
FAe :524 FE 3:165 MCL 3:336 MFG 2:125
L/443 L/530 L/1973
1163
Satureja montana L. subsp. macedonica (Formánek) Baden
Lamiaceae
Satureja macedonica Formánek
Satureja montana L. var. pisidica (Wettst.) Halácsy
Satureja olympica Halácsy, nomen nud.
FGC 2:551 MCL 3:337 MFG 2:123 PFPB 2:315
L/895 L/902 L/1028 L/1246
Satureja nana (P.H.Davis & Doroszenco) R.Jahn
Lamiaceae
Acinos graveolens (M.Bieb.) Link var. integrifolia Raulin nom nud.
Acinos nanus P.H.Davis & Doroszenco
EFK :265 FCSI :53 FCSII : FT 10:208 FT 10:236
L/346 L/762
!
1162
1161
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PATTERNS OF PLANT ENDEMISM IN GREECE 321
?
?
?
?
1172
1173
1174
1175
1176
Scilla battagliae (Speta) Valdés
Asparagaceae
Prospero battagliae Speta
FCSII :
L/1930 L/1931 L/2237
?
1171
Scilla messeniaca Boiss.
Asparagaceae
Scilla albanica Turill?
Scilla amoena sensu Bory & Chaubard
FE 5:42
L/607 L/1170 L/1963
Scilla idaea (Speta) Valdés
Asparagaceae
Prospero idaeum Speta
FCSII :
L/1930 L/1931 L/2237
Scilla hierapytnense (Speta) Valdés
Asparagaceae
Prospero hierapytnense Speta
FCSII :
L/1930 L/1931 L/2237
Scilla depressa (Speta) Valdés
Asparagaceae
Prospero depressum Speta
FCSII :
L/1930 L/1931 L/2237
Scilla cydonia Speta
Asparagaceae
Scilla bifolia sensu auct. cret., non L.
FCSII :
L/2237 L/2239 L/2240
Scilla autumnalis L. subsp. latifolia Iatroú & Kit Tan
Asparagaceae
L/1963
1170
Scilla andria Speta
Asparagaceae
L/2181 L/2195
1169
Taxon
Scaligeria moreana Engstrand
Apiaceae
L/607 L/1123 L/1963
Taxon
Qualifiers
1168
Taxon
Number
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IoI
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SPi
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NPi
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KK
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EAe
Distribution
Qualifier
322 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Scilla rhadamanthi (Speta) Valdés
Asparagaceae
Prospero rhadamanthi Speta
FCSII :
L/1930 L/1931 L/2237
Scilla talosii Tzanoud. & Kypr.
Asparagaceae
Prospero talosii (Tzanoud. & Kypr.) Speta
FCSII :
L/1214 L/1930
Scorzonera cretica Willd.
Asteraceae
Scorzonera dependens Rech.f.?
Scorzonera lassitica Vierh.
Scorzonera mollis sensu auct., Rechinger Fl. Aeg.
EFK :335 FAe :686 FAe :687 FCA :71 FE 4:322 FGC 2:198 FT 5:646
B/3060c L/437 L/913 L/1878 L/1917 L/2093
Scorzonera crocifolia Sm.
Asteraceae
FAe :686 FE 4:320 FGC 2:196 FGS 2:62
L/122 L/128 L/267 L/323b L/395 L/450 L/516 L/530 L/607 L/765 L/776 L/781 L/928 L/1112 L/1123 L/1267b L/1830 L/1844 L/1871 L/1878 L/1963 L/2097
1179
1180
1181
1182
Scilla minima (Speta) Valdés
Asparagaceae
Prospero minimum Speta
FCSII :
L/1930 L/1931 L/2237
Scilla nana (Schult. & Schult. f) Speta
Asparagaceae
Scilla nana (Schult. & Schult.f) Speta subsp. albescens (Speta) Speta
Chinodoxa forbesii Baker var. idaea Baker
Chionodoxa cretica Boiss. & Heldr.
Chionodoxa nana Schult. & Schult.f.
Hyacinthus nanus Schult. & Schult.f.
Scilla albescens Speta
Scilla cretica (Boiss. & Heldr.) Speta
Scilla hyacinthoides sensu Seitz 1978
EFK :356 FCA :185 FCSI :110 FCSII : FE 5:44 MFG 2:696
L/430 L/731 L/856 L/1766 L/2237
?
1178
1177
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 323
Scrophularia heterophylla Willd. var. heterophylla
Scrophulariaceae
Scrophularia caesia Sm.
Scrophularia heterophylla Willd. var. caesia (Sm.) Weiss
Scrophularia heterophylla Willd. var. oliveriana (Wydler) Weiss
Scrophularia heterophylla Willd. var. pinnatisecta Boiss.
Scrophularia heterophylla Willd. var. tenuis (Hausskn.) Hayek
Scrophularia heterophylla Willd. var. obtisiloba (Marg. & Reut.) Bornm.
Scrophularia laciniata var. obtusiloba Marg. & Reut.
Scrophularia micrantha D’ Urv.
Scrophularia oliveriana Wydler
Scrophularia sibthorpiana Sprengel
Scrophularia tenuis Hausskn.
Scrophularia urvilleana Wydler
EFK :275 FAe :479 FCA :143 FE 3:219 FT 6:634 MFG 2:196
L/111 L/122 L/321 L/321 L/447 L/607 L/928 L/932 L/1267b L/1963 L/2013
1187
Scrophularia spinulescens Degen & Hausskn.
Scrophulariaceae
FE 3:220 MFG 2:196 RDB :452
L/895
Scorzonera serpentinica Rech.f.
Asteraceae
FE 4:320
L/99 L/323b L/530 L/781 L/1871 L/1960
1186
1188
Scorzonera scyria Gustafsson & Snogerup
Asteraceae
FE 4:321 RDB :450
L/163 L/915
1185
Scorzonera eximia Rech.f.
Asteraceae
Scorzonera araneosa Sibth & Sm. described from Kiklades?
FAe :686 FE 4:322
L/128 L/169 L/196 L/783 L/861 L/930 L/1641
Taxon
Scorzonera mollis M.Bieb. subsp. idaea (Gand.) Lack
Asteraceae
Podospermum idaeum Gand.
Scorzonera idaea (Gand.) Lipsic.
EFK :335 FAe :686 FCA :71 FE 4:319 MFG 2:534
!
Taxon
Qualifiers
1184
1183
Taxon
Number
+
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StE
Pe
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WAe
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IoI
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NPi
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SPi
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EC
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NC
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NE
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NAe
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Kik
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KK
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EAe
Distribution
Qualifier
324 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Scutellaria albida L. subsp. vacillans (Rech.f.) Bothmer
Lamiaceae
Scutellaria vacillans Rech.f.
FE 3:136 MCL 3:342 MFG 2:80
L/209 L/347 L/1972 L/1973
Scutellaria brevibracteata Stapf subsp. icarica (Rech.f.) Greuter
& Burdet
Lamiaceae
Scutellaria icarica Rech.f.
Scutellaria rubicunda Hornem. subsp. icarica (Rech.f.) Rech.f.
Scutellaria rubicunda Hornem. subsp. samia Rech.f.
Scutellaria sieberi
FAe :504 FT 7:85 MCL 3:344
L/111 L/347 L/539 L/540 L/796 L/940 L/963
Scutellaria goulimyi Rech.f.
Lamiaceae
Scutellaria velenovskyi Rech.f. subsp. goulimyi (Rech.f.) Rech.f.
FE 3:136 MCL 3:342
L/99 L/155 L/209 L/323b L/347 L/1871
Scutellaria hirta Sm.
Lamiaceae
EFK :258 FCA :100 FE 3:136 MCL 3:343 MFG 2:81
L/347 L/1766
Scutellaria rupestris Boiss. & Heldr. subsp. caroli-henrici Bothmer
Lamiaceae
RDB :456
L/347 L/963 L/1963
Scutellaria rupestris Boiss. & Heldr. subsp. cephalonica (Rech.f.)
Greuter & Burdet
Lamiaceae
Scutellaria rubicunda Hornem. subsp. cephalonica (Bornm.) Rech.f.
Scutellaria rupestris Boiss. & Heldr. var. cephalonica Bornm.
FE 3:136 MCL 3:345 MFG 2:80
L/347 L/940 L/963 L/1123 L/1846
1189
1190
1191
1192
1193
1194
+
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PATTERNS OF PLANT ENDEMISM IN GREECE 325
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Scutellaria rupestris Boiss. & Heldr. subsp. olympica Bothmer
Lamiaceae
RDB :456
L/347
Scutellaria rupestris Boiss. & Heldr. subsp. parnassica (Boiss. &
Heldr.) Greuter & Burdet
Lamiaceae
Scutellaria euboea Rech.f.
Scutellaria geraniana (Tuntas ex Halácsy) Rech.f.
Scutellaria peregrina L. var. geraniana (Halácsy) Hayek
Scutellaria peregrina L. var. parnassica Boiss.
Scutellaria rubicunda Hornem. subsp. geraniana (Halácsy)
I.B.K.Richardson
Scutellaria rubicunda Hornem. subsp. hellenica Rech.f.
Scutellaria rubicunda Hornem. subsp. parnassica (Boiss.) Rech.f.
Scutellaria rubicunda Hornem. subsp. rupestris (Boiss. & Heldr.)
I.B.K.Richardson p.p.
Scutellaria rupestris Boiss. & Heldr. subsp. geraniana (Halácsy) Greuter
Scutellaria sibthorpii var. geraniana Halácsy
FE 3:136 MCL 3:345 MFG 2:78
L/323b L/347 L/439 L/512 L/530 L/607 L/940 L/963 L/1830 L/1846 L/1871 L/1963
Scutellaria rupestris Boiss. & Heldr. subsp. rechingeri Bothmer
Lamiaceae
MFG 2:80 RDB :454
L/963
Scutellaria rupestris Boiss. & Heldr. subsp. rupestris
Lamiaceae
Scutellaria peregrina L. subsp. rupestris (Boiss. & Heldr.) Maire &
Petitm.
Scutellaria rubicunda Hornem. subsp. rupestris (Boiss. & Heldr.)
I.B.K.Richardson
FE 3:136 MCL 3:345 RDB :456
L/347 L/963 L/1846 L/1963
Scutellaria sieberi Benth.
Lamiaceae
Scutellaria hirta Sieber, non Sm.
FCA :100 FE 3:136 FGC 2:x MCL 3:346 MFG 2:81
L/209 L/347
1196
1197
1198
1199
1200
Taxon
Scutellaria rupestris Boiss. & Heldr. subsp. cytherea (Rech.f.)
Greuter & Burdet
Lamiaceae
Scutellaria rubicunda Hornem. subsp. cytherea Rech.f.
FE 3:136 MCL 3:345
L/119 L/347 L/765 L/940 L/963 L/1052 L/1915 L/1963
Taxon
Qualifiers
1195
Taxon
Number
+
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IoI
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WAe
StE
Pe
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SPi
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NPi
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NC
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NE
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NAe
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Kik
+
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KK
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EAe
Distribution
Qualifier
326 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Scutellaria sporadum Bothmer
Lamiaceae
Scutellaria geraniana (Tunt.) Rech.f.
MCL 3:342
L/209 L/347 L/439 L/443 L/766
Securigera globosa (Lam.) Lassen
Fabaceae
Coronilla globosa Lam.
EFK :171 FAe :384 FCA :112 FCSI :62 FE 2:183 MCL 4:174
L/554 L/932
Sedum apoleipon ’t Hart
Crassulaceae
Sedum sexangulare auct. non L.
AFE 12:89 FE 1:433 FHe 2:320 MCL 3:27 MFG 1:348
L/121 L/275 L/1039 L/2200
Sedum creticum C.Presl
Crassulaceae
Helladia cretica (C.Presl.) Kral.
AFE 12:120 EFK :131 FCA :76 FCSII : FE 1:435 FHe 2:332 MCL 3:22 MFG 1:353
L/275 L/392 L/432 L/534 L/551 L/1917
Sedum creticum C.Presl var. creticum
Crassulaceae
Oreosedum hierapetrae (Rech.f.) Grulich
Sedum creticum C.Presl var. hierapetrae (Rech.f.) ’t Hart & Hageman
Sedum hierapetrae Rech.f.
EFK :131 FCA :76 FCSII : FE 1:435 FHe 2:333 MCL 3:22 MFG 1:353
L/432 L/534
Sedum creticum C.Presl var. monocarpicum ’t Hart
Crassulaceae
Sedum cretense Maire
Sedum creticum Boiss. & Heldr.
Sedum creticum C.Presl var. creticum sensu Mount. Fl. Gr., 1986
AFE 12:120 EFK :131 FCA :76 FCSII 0: FHe 2:333
L/432 L/534 L/551
1201
1202
1203
1204
1205
1206
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PATTERNS OF PLANT ENDEMISM IN GREECE 327
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Sedum eriocarpum Sm. subsp. delicum (Vierh.) ’t Hart
Crassulaceae
Sedum delicum (Vierh.) A.Carlström
Sedum rubens L. subsp. delicum Vierh.
Sedum rubens L. var. delicum (Vierh.) Hayek
AFE 12:95 EFK :132 FAe :294 FCA :76 FCSI :38 FE 1:436 FHe 2:325 MCL 3:25
L/355 L/537 L/539 L/540 L/554 L/607 L/776 L/869 L/913 L/932 L/972 L/1078 L/1627
Sedum eriocarpum Sm. subsp. epiroticum (Bald.) ’t Hart
Crassulaceae
Sedum annuum L. subsp. epiroticum Bald.
AFE 12:95 FHe 2:325
Sedum eriocarpum Sm. subsp. eriocarpum
Crassulaceae
AFE 12:95 FHe 2:324
L/1963
Sedum eriocarpum Sm. subsp. spathulifolium ’t Hart
Crassulaceae
Sedum rubens subsp. delicum sensu auct. cret., non Vierh.
Sedum delicum sensu auct. cret., non (Vierh.) Carlström
AFE 12:95 FCA :76 FCSI :38 FCSII : FHe 2:325
Sedum laconicum Boiss. & Heldr. subsp. laconicum
Crassulaceae
Sedum laconicum Boiss. & Heldr. subsp. insulare (Rech.f.) Greuter &
Rech.f.
Sedum idaeum D.A.Webb.
Sedum laconicum Boiss. & Heldr. var. insulare Rech.f.
AFE 12:87 EFK :131 FCA :77 FCSI :37 FCSII : FE 1:433 FHe 2:317 MCL 3:21 MFG 1:347
L/119 L/1871 L/1996 L/2013 L/2200
Sedum litoreum Guss. var. creticum’t Hart
Crassulaceae
Sedum praesidis Runemark & Greuter
Sedum litoreum Guss. subsp. praesidis (Runemark & Greuter) Greuter
AFE 12:93 EFK :130 FCA :77 FCSI :37 FCSI :37 FE 1:436 FHe 2:323 MCL 3:23
L/253 L/551 L/921 L/1214
1208
1209
1210
1211
1212
1213
Taxon
Sedum eriocarpum Sm. subsp. apertiflorum ’t Hart
Crassulaceae
AFE 12:95 FHe 2:324
L/1871
Taxon
Qualifiers
1207
Taxon
Number
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NPi
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NC
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NAe
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KK
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EAe
Distribution
Qualifier
328 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Senecio eubaeus Boiss. & Heldr.
Asteraceae
FE 4:197 MFG 2:465
L/323b L/1871
Senecio fruticulosus Sm.
Asteraceae
EFK :318 FCA :71 FE 4:202 MFG 2:469
Serapias cordigera L. var. cretica (B.Baumann & H.Baumann)
P.Delforge
Orchidaceae
Serapias cordigera L. subsp. cretica B.Baumann & H.Baumann
FCSII :
L/1798 L/1816 L/2185
Serapias cycladum H.Baumann & Künkele
Orchidaceae
Serapias orientalis (Greuter) H.Baumann & Künkele subsp. cycladum
(H.Baumann & Künkele) Nelson
L/2181 L/2185
Serapias patmia Hirth & Spaeth
Orchidaceae
Serapias cordigera L. subsp. patmia (M.Hirth & H.Spaeth) Kreutz
FT 11:298
L/742
Seseli aroanicum Hartvig
Apiaceae
MFG 1:684
L/607 L/1041 L/1830 L/1938b L/1963
1216
1217
1218
1219
1220
1221
Sempervivum marmoreum Griseb. subsp. ballsii (Wale) Zonn.
Crassulaceae
Sempervivum ballsii Wale
AFE 12:62 FE 1:427 FGC 1: FHe 2:312 MCL 3:30 MFG 1:339
L/432 L/459
1215
!
Sedum tristriatum Boiss.
Crassulaceae
Oreosedum tristriatum (Boiss. & Heldr.) Grilich.
AFE 12:120 EFK :131 FCA :77 FE 1:434 FHe 2:333 MCL 3:28 MFG 1:354
L/275 L/432 L/518 L/607 L/1963
1214
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 329
Seseli parnassicum Boiss. & Heldr.
Apiaceae
FE 2:337 MFG 1:684
L/121 L/330 L/512 L/2200
Sesleria doerfleri Hayek
Poaceae
EFK :383 FCA :175 FE 5:175 RDB :460
L/430 L/750 L/1583
Sesleria taygetea Hayek
Poaceae
FE 5:176 MFG 2:779
L/269 L/530 L/607 L/736 L/1963
Sesleria vaginalis Boiss. & Orph.
Poaceae
Sesleria krajinae Deyl
FE 5:175 MFG 2:781
L/499 L/607 L/1871 L/1963 L/2200 L/2201
Sideritis clandestina (Bory & Chaub.) Hayek
Lamiaceae
Phlomis clandestina Bory & Chaub.
Sideritis cretica Sm. p.p., non L.
Sideritis syriaca sensu Bory & Chaub., non L.
Sideritis theezans Boiss. & Heldr.
FE 3:142 MCL 3:347 MFG 2:87
B/3060z L/512 L/1963
Sideritis clandestina (Bory & Chaub.) Hayek subsp. clandestina
Lamiaceae
MCL 3:347 MFG 2:88
L/607 L/1963
1223
1224
1225
1226
1227
1228
Taxon
Seseli gummiferum Pall. ex Sm. subsp. crithmifolium (DC.)
P.H.Davis
Apiaceae
Seseli crithmifolium (DC.) Boiss.
Seseli gummiferum Pall. ex Sm. subsp. aegaeum P.H.Davis (included in
taxon)
EFK :213 FAe :408 FCA :153 FE 2:335 FGC 1:649
L/196 L/197 L/206 L/707 L/869
Taxon
Qualifiers
1222
Taxon
Number
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WAe
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SPi
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NPi
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EAe
Distribution
Qualifier
330 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Sideritis clandestina (Bory & Chaub.) Hayek subsp. peloponnesiaca
(Boiss. & Heldr.) Baden
Lamiaceae
Sideritis clandestina (Bory & Chaub.) Hayek subsp. cyllenea (Hayek)
Papan. & Kokkini
Sideritis peloponnesiaca Boiss. & Heldr.
Sideritis theezans Boiss. & Heldr. subsp. peloponnesiaca (Boiss. &
Heldr.) Bornm.
Sideritis theezans Boiss. & Heldr. var. cyllenea Boiss.
FE 3:142 MCL 3:347 MFG 2:88
L/160 L/512 L/607 L/1963
Sideritis euboea Heldr.
Lamiaceae
Sideritis syriaca L. var. condensata Boiss. & Heldr.
FE 3:142 MCL 3:348 MFG 2:89
L/323b L/1871
Sideritis perfoliata L. subsp. athoa (Papan. & Kokkini) Baden
Lamiaceae
Sideritis athoa Papan. & Kokkini
Sideritis perfoliata L. var. lanata Turrill
FT 10:203 MCL 3:351 MFG 2:90
L/477 L/895 L/1973
Sideritis raeseri Boiss. & Heldr. subsp. attica (Heldr.) Papan. &
Kokkini
Lamiaceae
Sideritis attica Heldr.
Sideritis raeseri Boiss. & Heldr. var. attica (Heldr.) Halácsy
MCL 3:351 MFG 2:87
L/953 L/1109
Sideritis syriaca L. subsp. syriaca
Lamiaceae
EFK :259 FCA :100 FE 3:142 MCL 3:353
L/1766
Silene adelphiae Runemark
Caryophyllaceae
AFE 7:66 FE 1:206 FHe 1:278 MCL 1:254
L/254 L/358
1229
1230
1231
1232
1233
1234
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 331
Silene ammophila Boiss. & Heldr. subsp. ammophila
Caryophyllaceae
Conosilene conica (L.) A.Love & Kjellqvist subsp. ammophila
(Boiss. & Heldr.) A.Love & Kjellqvist
Pleconax conica (L.) Sourkova subsp. ammophila (Boiss. & Heldr.)
A.Love & Kjellqvist
AFE 7:118 EFK :86 FCA :50 FE 1:218 FHe 1:319 MCL 1:249 RDB :460
L/358 L/518 L/554 L/2093
Silene ammophila Boiss. & Heldr. subsp. carpathae Chowdhuri
Caryophyllaceae
Conosilene conica subsp. ammophila var. carpathae (Chowdhuri) A.Love
& Kjellqvist
Pleconax ammophila (Bois. & Heldr.) Šourková subsp. carpathae
(Chowdhuri) Šourková
Pleconax carpathae (Chowdhuri) Ikonn.
Pleconax conica subsp. ammophila var. carpathae (Chowdhuri) A.Love
& Kjellqvist
AFE 7:118 EFK :86 FCA :50 FE 1:218 FHe 1:319 MCL 1:249 RDB :462
L/201 L/358 L/551
Silene antri-jovis Greuter & Burdet
Caryophyllaceae
Silene fruticulosa Sieber ex Otth
Silene saxifraga L. subsp. fruticulosa Arcang.
Silene saxifraga sensu auct. Kret non L.
AFE 7:72 EFK :87 FAe :169 FCA :50 FCSI :20 FE 1:208 FHe 1:285 MCL 2:74
L/350 L/358 L/499 L/518 L/554 L/707 L/923
1237
1238
1239
Silene aegaea Oxelman
Caryophyllaceae
FHe 1:299 FT 11:52
L/345 L/358 L/539 L/540
Taxon
Silene ammophila Boiss. & Heldr.
Caryophyllaceae
Pleconax ammophila (Bois. & Heldr.) Šourková
AFE 7:118 EFK :86 FAe :165 FCA :50 FE 1:218 FHe 1:319 MCL 1:249 RDB :462
L/111 L/201 L/358 L/518 L/551 L/554 L/2093
Taxon
Qualifiers
1236
1235
Taxon
Number
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StE
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NPi
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Distribution
Qualifier
332 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Silene auriculata Sm.
Caryophyllaceae
Melandrium auriculatum (Sibth. & Sm.) Rohrb.
Silene auriculata subsp. lanuginosa (Bertol.) Arcangeli
Silene lanuginosa Bertol.
AFE 7:54 FE 1:203 FHe 1:270 MCL 1:250 MFG 1:146
L/160 L/358 L/512 L/529 L/587 L/1963
Silene barbeyana Heldr. ex Boiss.
Caryophyllaceae
AFE 7:79 FE 1:209 FHe 1:292 MCL 1:250 MFG 1:162
L/330 L/358 L/934
Silene cephallenia Heldr. subsp. cephallenia
Caryophyllaceae
Silene linifolia Sibth. & Sm. subsp. cephallenia (Heldr.) Maire & Petitm.
AFE 7:66 FE 1:206 FHe 1:280 MCL 1:252
L/358 L/603
Silene congesta Sm.
Caryophyllaceae
Silene congesta Sm. subsp. moreana Melzh.
Silene delphica Boiss. & Heldr.
AFE 7:66 FE 1:206 FHe 1:279 MCL 1:154
L/358 L/358 L/512 L/934 L/1588 L/1844 L/1963
Silene conglomeratica Melzh.
Caryophyllaceae
Silene saxifraga L. var. coenobitarum Hayek
AFE 7:74 FE 1:208 FHe 1:286 MCL 1:273 MFG 1:158
L/350 L/358 L/1092 L/1963
Silene corinthiaca Boiss. & Heldr.
Caryophyllaceae
Silene juncea auct. [non Roth]
Silene kotschyi Boiss. var. corinthiaca (Boiss. & Heldr.) F.N.Williams
Silene portensis auct. [non L.]
Silene portensis L. subsp. rigidula Greuter & Burdet
Silene rigidula Sm.
Silene vittata auct. [non Stapf]
AFE 7:96 FE 1:212 FGC 1:173 FHe 1:297 MCL 1:272
L/358 L/451 L/516 L/530 L/1588 L/1844 L/1871 L/1963
1240
1241
1242
1243
1244
1245
+
+
+
+
+
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PATTERNS OF PLANT ENDEMISM IN GREECE 333
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Silene dirphya Greuter & Burdet
Caryophyllaceae
Saponaria caespitosa Sm.
Saponaria saxifraga L. var. smithii (Ser.) Rohrb.
Saponaria smithii Ser.
Silene saxifraga L. subsp. smithii (Ser.) Nyman
Silene smithii (Ser.) Boiss. & Heldr.
AFE 7:74 FE 1:208 FHe 1:285 MCL 1:274 MFG 1:157
L/350 L/358 L/418 L/1871
Silene echinosperma Boiss. & Heldr.
Caryophyllaceae
AFE 7:96 FE 1:212 FHe 1:296 MCL 1:257
L/358 L/607 L/1963
Silene fabaria (L.) Sm. subsp. domokina Greuter
Caryophyllaceae
FHe 1:271
L/350 L/358 L/530 L/736 L/995
Silene flavescens Waldst. & Kit. subsp. dictaea (Rechf.) Greuter
Caryophyllaceae
Silene dictaea Rech.f.
Silene thessalonica Boiss. & Heldr. subsp. dictaea (Rech.f.) Melzh.
AFE 7:65 EFK :89 FCA :52 FCSI :22 FE 1:206 FHe 1:281 MCL 1:258 MFG 1:153 RDB :464
L/350 L/358 L/518 L/554 L/1267a L/1583
Silene gigantea (L.) L. subsp. hellenica Greuter
Caryophyllaceae
FHe 1:258
L/350 L/358 L/530 L/1871
Silene goulimyi Turrill
Caryophyllaceae
AFE 7:29 FE 1:199 FHe 1:257 MCL 1:259
L/358 L/607 L/771 L/785 L/934
1247
1248
1249
1250
1251
1252
Taxon
Silene cythnia (Halácsy) Walters
Caryophyllaceae
Silene italica (L.) Pers. var. cythnia Halácsy
AFE 7:29 FE 1:199 FGC 1:184 FHe 1:256 FT 11:50 MCL 1:256
L/128 L/169 L/358 L/442 L/539 L/540 L/759a L/776 L/783 L/869 L/934 L/1288
Taxon
Qualifiers
1246
Taxon
Number
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WAe
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NPi
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KK
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EAe
*
Distribution
Qualifier
334 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Silene grisebachii (Davidov) Pirker & Greuter
Caryophyllaceae
Pleconax subconica Sourkova subsp. grisebachii (Davidov) Sourkova
Pleconax subconica var. grisebachii (Davidov) Sourkova
Silene subconica Friv. subsp. grisebachii (Davidov) Jordanov & Panov
Silene subconica Friv. var. grisebachii Davidov
AFE 7:120 FHe 1:320 MCL 1:255
L/350 L/358 L/1349
Silene guicciardii Boiss. & Heldr.
Caryophyllaceae
Silene marschallii auct. non Mey.
AFE 7:34 FE 1:200 FHe 1:259 MCL 1:265
L/358
Silene haussknechtii Heldr. ex Hausskn.
Caryophyllaceae
Silene sedoides Poiret subsp. haussknechtii (Hausskn.) Maire & Petitm.
AFE 7:100 FE 1:213 FHe 1:299 MCL 1:259 MFG 1:169
L/345 L/358 L/513 L/931 L/995
Silene holzmannii Heldr. ex Boiss.
Caryophyllaceae
AFE 7:104 EFK :86 FAe :171 FCA :51 FE 1:214 FGC 1:177 FHe 1:306 FT 11:53 MCL 1:251 RDB :466
L/111 L/138 L/201 L/206 L/255 L/358 L/365 L/495 L/551 L/759a L/776 L/877 L/913 L/1023 L/1049 L/1710 L/1963
Silene insularis Barbey
Caryophyllaceae
AFE 7:99 EFK :88 FCA :51 FE 1:213 FHe 1:302 MCL 1:260
L/111 L/358
Silene integripetala Bory & Chaub.
Caryophyllaceae
AFE 7:99 EFK :89 FCSI :21 FCSII : FE 1:213 FHe 1:301 MCL 1:260 MFG 1:169
B/3060c L/345 L/358 L/607 L/1210d L/1917 L/1963
Silene integripetala Bory & Chaub. subsp. elaphonesiaca Oxelman
Caryophyllaceae
FHe 1:301
L/345 L/358
1253
1254
1255
1256
1257
1258
1259
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 335
Silene integripetala Bory & Chaub. subsp. lidenii Oxelman
Caryophyllaceae
FCSII : FHe 1:301
B/3060c L/345 L/358 L/1210d
Silene intonsa Greuter & Melzh.
Caryophyllaceae
Silene retzdorffiana sensu Quézel & Contandr.
AFE 7:87 FE 1:211 FHe 1:295 MCL 1:270
L/358 L/601 L/922
Silene ionica Halácsy
Caryophyllaceae
Oberna ionica (Halácsy) Ikonn.
AFE 7:63 FE 1:205 FHe 1:272 MCL 1:257
L/358 L/769 L/770 L/934 L/1123 L/1846
Silene italica (L.) Pers. subsp. peloponnesiaca Greuter
Caryophyllaceae
FHe 1:255 MFG 1:140
L/350 L/358 L/530 L/1963
Silene laconica Boiss. & Orph.
Caryophyllaceae
AFE 7:100 FE 1:214 FHe 1:300 MCL 1:262
L/345 L/358 L/607 L/1963
Silene linoides Otth
Caryophyllaceae
Silene linifolia Sm.
Silene linifolia Sm. var. ceccariniana Hausskn.
Silene linifolia Sm. var. gracilis Beauverd & Topali
AFE 7:71 FE 1:207 FGC 1:182 FHe 1:290 MCL 1:264
L/99 L/323a L/358 L/530 L/771 L/771 L/934 L/1963
1262
1263
1264
1265
1266
1267
Silene integripetala Bory & Chaub. subsp. greuteri (Phitos) Akeroyd
Caryophyllaceae
Silene greuteri Phitos
AFE 7:99 EFK :89 FCA :51 FCSI :21 FE 1:213 FHe 1:301 MCL 1:260 RDB :468
L/252 L/345 L/358 L/711 L/897
Taxon
Silene integripetala Bory & Chaub. subsp. integripetala
Caryophyllaceae
FCSI :21 FE 1:213 FHe 1:301 MCL 1:260 MFG 1:169
L/345 L/358 L/607 L/1963
Taxon
Qualifiers
1261
1260
Taxon
Number
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+
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EAe
Distribution
Qualifier
336 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Silene melzheimeri Greuter
Caryophyllaceae
Silene radicosa Boiss. & Heldr. subsp. rechingeri Melzh.
FE 1:201 FHe 1:262 MCL 1:272 MFG 1:144
L/350 L/358 L/774 L/2201
Silene multicaulis Guss. subsp. sporadum (Halácsy) Greuter
Caryophyllaceae
Silene genistifolia Halácsy
Silene linifolia Sm. subsp. sporadum (Halácsy) Phitos
Silene linifolia Sm. var. sporadum Halácsy
Silene multicaulis Guss. subsp. genistifolia (Halácsy) Melz.
Silene multicaulis Guss. subsp. stenocalycina (Rech.f.) Melz.
Silene stenocalycina Rech.f.
AFE 7:77 FE 1:208 FHe 1:290 MCL 1:266 MFG 1:159
L/358 L/388 L/443 L/767 L/771 L/774 L/895 L/928 L/934 L/1107 L/1871 L/1973 L/2202
Silene niederi Heldr. ex Boiss.
Caryophyllaceae
AFE 7:34 FE 1:200 FHe 1:259 MCL 1:267
L/358 L/441 L/603
Silene nutabunda Greuter
Caryophyllaceae
FHe 1:307
L/350 L/358 L/1963
Silene oligantha Boiss. & Heldr.
Caryophyllaceae
Silene radicosa Boiss. & Heldr. var. breviflora Boiss.
AFE 7:40 FE 1:201 FHe 1:262 MCL 1:272 MFG 1:144
L/350 L/358 L/775 L/934 L/1246 L/1871 L/1960
Silene oligantha Boiss. & Heldr. subsp. oligantha
Caryophyllaceae
FE 1:201 FHe 1:262 MCL 1:272 MFG 1:144
L/350 L/358
Silene oligantha Boiss. & Heldr. subsp. parnesia Greuter
Caryophyllaceae
FE 1:201 FHe 1:262 MCL 1:272 MFG 1:144
L/350 L/358
1268
1269
1270
1271
1272
1273
1274
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 337
Silene orphanidis Boiss.
Caryophyllaceae
AFE 7:79 FE 1:209 FHe 1:288 MCL 1:268 MFG 1:162 RDB :470
L/138 L/350 L/358 L/404a L/1972 L/1973
Silene parnassica Boiss. & Spruner subsp. dionysii Stoj. & Jordanov
Caryophyllaceae
Silene dionysii Stoj. & Jordanov
Silene dionysii Stoj. & Jordanov var. elata Stoj. & Jordanov
Silene dionysii Stoj. & Jordanov var. pumila Stoj. & Jordanov
AFE 7:79 FE 1:209 FHe 1:288 MCL 1:257 MFG 1:160
L/350 L/358 L/775
Silene parnassica Boiss. & Spruner subsp. vourinensis Greuter
Caryophyllaceae
AFE 7:75 FE 1:208 FHe 1:288 MCL 1:274 MFG 1:156
L/254 L/350 L/358
Silene pentelica Boiss.
Caryophyllaceae
Silene sedoides Poiret subsp. pentelica (Boiss.) Nyman
Silene sedoides Poiret var. pentelica (Boiss.) Rohrb.
AFE 7:100 FE 1:213 FGC 1:174 FHe 1:299 FT 11:52 MCL 1:269
L/282 L/310a L/345 L/358 L/361 L/539 L/540 L/1871 L/2181
Silene pinetorum Boiss. & Heldr.
Caryophyllaceae
AFE 7:96 EFK :86 FAe :170 FCA :51 FCSI :22 FE 1:212 FHe 1:296 MCL 1:269
L/350 L/358 L/518 L/554
1277
1278
1279
1280
1281
Silene oligantha Boiss. & Heldr. subsp. pseudoradicosa (Rech.f.)
Greuter
Caryophyllaceae
Silene radicosa Boiss. & Heldr. subsp. pseudoradicosa Rech.f.
FE 1:201 FHe 1:262 MCL 11:272 MFG 1:143
L/350 L/358 L/608 L/771 L/934 L/1871 L/1960
Taxon
Silene orbelica Greuter
Caryophyllaceae
FHe 1:282
L/350 L/358
Taxon
Qualifiers
1276
1275
Taxon
Number
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StE
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IoI
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SPi
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EAe
Distribution
Qualifier
338 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Silene pinetorum Boiss. & Heldr. subsp. pinetorum
Caryophyllaceae
FAe :170 FCSI :22 FHe 1:296
L/350 L/358 L/518 L/554
Silene pinetorum Boiss. & Heldr. subsp. sphaciotica Oxelman &
Greuter
Caryophyllaceae
FAe :170 FCSI :22 FHe 1:296
L/350 L/358 L/518 L/554
Silene pusilla Waldst. & Kit. subsp. chromodonta (Boiss. & Reuter)
Greuter
Caryophyllaceae
Heliosperma chromodontum (Boiss. & Reuter) Juratzka
Ixoca chromodonta (Boiss. & Reuter) Sojak
Ixoca phyllitica (Neumayer) Ikonn
Silene chromodonta Boiss. & Reut.
Silene quadridentata (Murray) Pers. subsp. chromodonta
(Boiss. & Reuter) H.Neumayer
Silene quadridentata subsp. phyllitica Neumayer
AFE 7:87 FE 1:211 FHe 1:294 MCL 1:270 MFG 1:106
L/350 L/358
Silene pusilla Waldst. & Kit. subsp. tymphaea Greuter
Caryophyllaceae
FHe 1:295
L/350 L/358 L/601 L/688
Silene salamandra Pamp.
Caryophyllaceae
Silene aegyptiaca auct. [non (L.) L.]
FAe :170 FHe 1:302 FT 10:80 MCL 1:249
L/197 L/288 L/358 L/995
Silene samothracica (Rech.f.) Greuter
Caryophyllaceae
Silene caesia sensu Rechinger 1943
Silene caesia Sm. subsp. samothracica (Rech.f.) Melzh.
Silene variegata (Desf.) Boiss. & Heldr. var. samothracica Rech.f.
AFE 7:63 FAe :165 FE 1:205 FHe 1:273 FT 11:51 MCL 1:251 MFG 1:150
L/321 L/350 L/358 L/796 L/895 L/974 L/1957
1282
1283
1284
1285
1286
1287
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PATTERNS OF PLANT ENDEMISM IN GREECE 339
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Silene sedoides Poiret subsp. runemarkii Oxelman
Caryophyllaceae
FHe 1:299
L/345 L/358 L/1963
Silene sieberi Fenzl
Caryophyllaceae
AFE 7:24 EFK :90 FAe :173 FCA :52 FE 1:197 FHe 1:255 MCL 1:262 MFG 1:141
L/321 L/358 L/518 L/554 L/1766
Silene spinescens Sm.
Caryophyllaceae
Silene spinescens Sm. var. argolica Boiss.
AFE 1:29 FE 1:199 FHe 1:256 MCL 1:276
L/358 L/516 L/530 L/607 L/759a L/771 L/934 L/1871 L/1963
Silene taygetea Halácsy ex Vierh.
Caryophyllaceae
Saxifraga fruticulosa subsp. taygetea (Halácsy) Hayek
Silene saxifraga L. subsp. taygetea (Halácsy ex Vierh.) Hayek
AFE 7:72 FE 1:208 FHe 1:285 MCL 1:274 MFG 1:155
L/358
Silene variegata (Desf.) Boiss. & Heldr.
Caryophyllaceae
Cucubalus cyclamineous Sieber
Lychnis variegata Desf.
Oberna variegata (Desf.) J.Holub
AFE 7:63 EFK :87 FAe :165 FCA :52 FE 1:205 FHe 1:273 MCL 1:280 MFG 1:151
L/358 L/518 L/554
1289
1290
1291
1292
1293
Taxon
Silene sartorii Boiss. & Heldr.
Caryophyllaceae
Conosilene conica var. sartorii (Boiss & Heldr.) A.Love & Kjellqvist
Pleconax conica var. sartorii (Boiss. & Heldr.) A.Love & Kjellqvist
Pleconax sartorii (Boiss. & Heldr.) Sourkova
Silene conica L. subsp. sartorii (Boiss. & Heldr.) Chater & Walters
AFE 7:120 FAe :166 FE 1:218 FHe 1:320 FT 2:240 MCL 1:254
L/111 L/128 L/153 L/197 L/358 L/759a L/776 L/783 L/869 L/1890 L/1963
Taxon
Qualifiers
1288
Taxon
Number
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+
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SPi
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IoI
WAe
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StE
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NPi
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EAe
Distribution
Qualifier
340 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Stachys candida Bory & Chaub.
Lamiaceae
FCA 3:156 FE 3:156 MCL 3:335
L/512 L/607 L/785 L/1963
Solenanthus pindicus Aldén
Boraginaceae
Cynoglossum pindicum (Aldén) Greuter & Burdet
MCL 2:79 MFG 2:61
L/477 L/709
1298
1300
Soldanella chrysosticta Kress subsp. pelia (Raus) Raus
Primulaceae
Soldanella pelia Raus
MCL 4:383 RDB :474
L/298 L/1896 L/1897
1297
Solenopsis minuta (L.) C.Presl subsp. annua Greuter, Matthäs &
Risse
Campanulaceae
EFK :297 FAe :606 FCA :43 FCSI :16 MCL 1:153
L/252 L/518 L/780 L/893
Silene vulgaris (Moench) Garcke subsp. vourinensis Greuter
Caryophyllaceae
FHe 1:278
L/350 L/358
1296
1299
Silene vulgaris (Moench) Garcke subsp. suffrutescens Greuter,
Matthäs & Risse
Caryophyllaceae
AFE 7:58 EFK :86 FCA :52 FCSII : FE 1:204 FHe 1:277 MCL 1:279
L/252 L/358
1295
?
Silene vulgaris (Moench) Garcke subsp. megalosperma (Sart. ex
Heldr.) Hayek
Caryophyllaceae
Silene cucubalus Wibel. subsp. megalosperma (Sart. ex Heldr.) Rech.f.
Silene inflata var. microphylla Boiss.
Silene megalosperma Sart. ex Heldr.
Silene venosa Ascherson var. megalosperma (Sart. ex Heldr.) Halácsy
AFE 7:59 FGC 1:260 FHe 1:277
L/160 L/512 L/530 L/759a L/1871 L/1963 L/2013
1294
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 341
Stachys macrotricha Rech.f. & Goulimy
Lamiaceae
FE 3:154 MCL 3:359
L/818
Stachys mucronata Sieber ex Sprengel
Lamiaceae
EFK :261 FCA :101 FE 3:155 MCL 3:361
L/273 L/551
1307
1308
Stachys graeca Boiss. & Heldr.
Lamiaceae
Stachys acutifolia sensu Halácsy (1902, 1908, 1912), Rechinger (1943),
Tutin & al. (1972)
FE 3:154 FGC 2:522 FGS 1:85 FGS 2:69 MCL 3:359
L/323b L/530 L/1871
1305
Stachys ionica Halácsy
Lamiaceae
Stachys candida auct., non Bory & Chaub.
Stachys ionica Halácsy var. cephalonica Halácsy
Stachys swansonii auct., non Benth.
Stachys swansonii Benth. var. ionica (Halácsy) Hayek f. cephalonica
(Halácsy) Hayek
FE 3:156 FGC 2:527 FGS 1:85 MCL 3:366
L/537 L/1123 L/1690 L/1800
Stachys goulimyi Rech.f.
Lamiaceae
FE 3:155 MFG 2:104
L/818 L/1901
1304
1306
Stachys euboica Rech.f.
Lamiaceae
FE 3:156 MCL 3:356 RDB :478
L/323b L/1115 L/1871 L/1871
1303
!
Stachys chrysantha Boiss. & Heldr.
Lamiaceae
Stachys candida Bory & Chaub. subsp. chrysantha Benth.
FE 3:156 MCL 3:355
L/119 L/512 L/607 L/1877 L/1963
1302
Taxon
Stachys canescens Bory & Chaub.
Lamiaceae
Stachys messeniaca Boiss.
FE 3:154 MCL 3:355
L/607 L/1963
Taxon
Qualifiers
1301
Taxon
Number
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StE
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SPi
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NPi
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EAe
Distribution
Qualifier
342 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Stachys pangaea Phitos
Lamiaceae
MCL 3:363 MFG 2:103 RDB :480
L/1118
Stachys parolinii Vis.
Lamiaceae
FE 3:155 FGC 2:524 MCL 3:364
L/607 L/1123 L/1244 L/1844 L/1963
Stachys recta L. subsp. olympica Stoj. & Jordanov
Lamiaceae
MFG 2:106
L/953
Stachys spinosa L.
Lamiaceae
EFK :261 FCA :101 FCSI :55 FE 3:156 FGC 2:x FGS 1:x MCL 3:365
L/196 L/268 L/273
Stachys spreitzenhoferi Heldr.
Lamiaceae
FE 3:156 MCL 3:355
L/119 L/607 L/765 L/1963
Stachys spreitzenhoferi Heldr. subsp. spreitzenhoferi
Lamiaceae
MCL 3:355
L/119 L/607 L/765 L/1963
Stachys spreitzenhoferi Heldr. subsp. virella D.Persson
Lamiaceae
MCL 3:355 RDB :482
L/607 L/1963
Stachys spruneri Boiss.
Lamiaceae
FE 3:155 FGC 2:x FGS 1:69 FGS 2:528 MCL 3:366
L/516 L/530
Stachys swainsonii Benth.
Lamiaceae
FE 3:156 FGC 2:x MCL 3:366
L/530 L/1122 L/1963
1309
1310
1311
1312
1313
1314
1315
1316
1317
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 343
Stachys swainsonii Benth. subsp. scyronica (Boiss.) Phitos &
Damboldt
Lamiaceae
Stachys scyronica Boiss.
FE 3:156 MCL 3:366 RDB :484
L/530 L/1115 L/1122
Stachys swainsonii Benth. subsp. swainsonii
Lamiaceae
FE 3:156 FGC 2:x MCL 3:366
Stachys tetragona Boiss. & Heldr.
Lamiaceae
FE 3:155 FGC 2:528 FGS 1:86 MCL 3:366
L/196 L/323b L/423 L/443 L/928 L/1871
Stachys virgata Bory & Chaub.
Lamiaceae
FE 3:155 FGC 2:525 MCL 3:366
L/607
Staehelina petiolata (L.) Hilliard & B.L.Burtt
Asteraceae
Staehelina arborea Schreb.
Staehelina arborescens L.
EFK :321 FCA :72 FE 4:217
L/1766
Sternbergia greuteriana Kamari & R.Artelari
Amaryllidaceae
Sternbergia sicula auct. plur. fl. cret., non Tineo ex Guss.
FCA :158
L/551 L/792 L/1049 L/1074 L/1807
1320
1321
1322
1323
1324
1325
Stachys swainsonii Benth. subsp. argolica (Boiss.) Phitos &
Damboldt
Lamiaceae
Stachys argolica Boiss.
Stachys swainsonii Benth. var. argolica (Boiss.) Boiss.
FE 3:156 FGC 2:x MCL 3:366
L/607 L/1122 L/1963
Taxon
Stachys swainsonii Benth. subsp. melangavica
Lamiaceae
RDB :484
L/530 L/1115
Taxon
Qualifiers
1319
1318
Taxon
Number
.
.
.
.
.
.
.
.
+
.
+
+
.
.
.
.
+
.
.
.
.
+
WAe
StE
.
+
Pe
.
.
.
.
.
.
.
.
IoI
.
.
.
.
.
.
.
.
SPi
.
.
.
.
.
.
.
.
NPi
.
.
.
.
.
.
.
.
EC
.
.
.
.
.
.
.
.
NC
.
.
.
.
.
.
.
.
NE
.
.
.
.
.
.
.
.
NAe
.
.
.
.
.
.
.
.
Kik
+
+
.
.
.
.
.
.
KK
.
.
.
.
.
.
.
.
EAe
Distribution
Qualifier
344 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
?
1330
Symphyandra cretica A.DC.
Campanulaceae
Campanula cretica (A.DC.) D.Dietr
Symphyandra cretica A.DC. subsp. cretica
Symphyandra cretica A.DC. subsp. eu-cretica Hayek
EFK :294 FAe :602 FCA :42 FCSI :15 FE 4:93 MCL 1:125
L/518 L/554 L/1111
Symphyandra samothracica (Degen) Halácsy
Campanulaceae
Campanula samothracica (Degen) Greuter & Burdet subsp. samothracica
Symphyandra cretica A.DC. subsp. samothracica (Degen) Hayek
FE 4:93 MCL 1:125 MFG 2:389
L/533 L/709 L/895 L/1252 L/1349
1331
1332
Stipa rechingeri Martinovský
Poaceae
FE 5:249 MFG 2:829
L/269 L/528 L/1211 L/2201 L/2237
Stipa monticola H.Scholz
Poaceae
L/283 L/2237
?
1329
Stipa isoldae H.Scholz
Poaceae
L/1163 L/1901 L/2237
Stipa lessingiana Tri. & Rupr. subsp. cyllenaea (Strid) Strid
Poaceae
Stipa cyllenea Strid
MFG 2:826
L/269 L/528 L/607 L/1181
?
1327
Stipa endotricha Martinovsky
Poaceae
FE 5:249 MFG 2:829
L/269 L/528 L/607 L/1217 L/1832d L/2237 L/2242
1328
?
1326
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
+
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
?
.
.
.
.
.
.
.
.
.
+
+
.
+
+
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 345
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Symphytum creticum (Willd.) Greuter & Rech.f.
Boraginaceae
Borago cretica Willd.
Procopiania cretica (Willd.) Gusuleac
Psilostemon creticus (Willd.) DC.
Trachystemon creticum (Willd.) D.Don ex G.Don
EFK :250 FAe :446 FCA :41 FE 3:105 MCL 1:115
B/3060c L/119 L/122 L/518 L/554 L/607 L/1917 L/1963
Symphytum cycladense Pawl.
Boraginaceae
Symphytum davisii Wickens subsp. cycladense (Pawl.) Stearn
FE 3:105 MCL 1:114
L/138 L/1177
Symphytum davisii Wickens
Boraginaceae
Symphytum davisii Wickens subsp. davisii
FE 3:105 MCL 1:114
L/1177
Symphytum icaricum Pawl.
Boraginaceae
Symphytum davisii sensu Wickens 1969 pp.
Symphytum davisii Wickens subsp. icarisum (Pawl.) Stearn
FT 6:382 MCL 1:114
L/539 L/540 L/1114 L/1177
Symphytum insulare (Pawl.) Greuter & Burdet
Boraginaceae
Procopiania insularis Pawl.
Prokopiania cretica sensu Fl. Aeg.
Trachystemon orientale auct.
EFK :250 FAe :446 FAe :447 FCA :41 FE 3:106 MCL 1:115
L/201 L/551 L/709 L/1161
1334
1335
1336
1337
1338
Taxon
Symphyandra sporadum Halácsy
Campanulaceae
Campanula samothracica (Degen) Greuter & Burdet subsp. sporadum
(Halácsy) Greuter & Burdet
Campanula sporadum s.l. sensu Gustafsson M. & Snogerup S. (1974)
from Skantzoura
Symphyandra cretica subsp. sporadum (Halácsy) Hayek
FAe :603 FE 4:93 FGC 2:248 MCL 1:125 MFG 2:389
L/163 L/196 L/423 L/443 L/709 L/767 L/928
Taxon
Qualifiers
1333
Taxon
Number
.
.
.
.
.
.
.
.
.
.
.
+
StE
Pe
.
.
.
.
+
+
WAe
.
.
.
.
+
.
IoI
.
.
.
.
.
.
SPi
.
.
.
.
.
.
NPi
.
.
.
.
.
.
EC
.
.
.
.
.
.
NC
.
.
.
.
.
.
NE
.
.
.
.
.
.
NAe
+
.
+
+
+
.
Kik
+
.
.
.
+
.
KK
.
+
.
.
+
.
EAe
Distribution
Qualifier
346 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
A
!
!
1341
1342
A
A
S
A
A
1343
1344
1345
1346
1347
A
Taraxacum acutiusculum Sonck
Asteraceae
MCL 2:746
U
1340
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Taraxacum decrepitum Kirschner & Štepánec
Asteraceae
MCL 2:758
L/1963
Taraxacum declivicola Kirschner, Sonck & Štepánec
Asteraceae
MCL 2:758
Taraxacum cylleneum Furnkranz
Asteraceae
Taraxacum apenninum sensu Hayek Prodr. Fl. Penins. Balc.
Taraxacum glaciale sensu Halácksy Fl. Graec.
FGC 2:223 MCL 2:762 MFG 2:545
L/607 L/1830 L/1938b L/1963 L/2013
Taraxacum copidophylloides A.J.Richards
Asteraceae
MCL 2:746 MFG 2:565
L/2200
Taraxacum amborum G.Hagl.
Asteraceae
MCL 2:744 MFG 2:571
Taraxacum albomarginatum A.J.Richards
Asteraceae
MCL 2:744 MFG 2:567
L/1211 L/2201
Taraxacum aganippeum Sonck
Asteraceae
MFG 2:551
Symphytum naxicola Pawl.
Boraginaceae
Symphytum anatolicum auct. from Naxos Fl. Aeg.
Symphytum davisii Wickens subsp. naxicola (Pawl.) Stearn
FAe :446 FE 3:104 FGC 2:330 MCL 1:114
L/857 L/869 L/1177
1339
.
.
.
.
.
.
+
.
+
.
+
.
.
.
.
.
.
+
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
+
.
.
.
.
.
+
.
.
.
.
.
.
.
.
+
.
+
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
*
*
PATTERNS OF PLANT ENDEMISM IN GREECE 347
A
A
A
A
A
A
1353
1354
1355
1356
1357
1358
!
U
1352
A
1351
!
U
U
1349
1350
A
Taxon
Qualifiers
1348
Taxon
Number
Taraxacum graecofontanum A.J.Richards & Sonck
Asteraceae
MCL 2:752 MFG 2:552
Taraxacum glabricaule Sonck
Asteraceae
MCL 2:758
L/2253
Taraxacum gionense A.J.Richards
Asteraceae
MCL 2:748 MFG 2:558
L/1830 L/2013 L/2200
Taraxacum fragosum Sonck
Asteraceae
MCL 2:748 MFG 2:563
L/405 L/2200
Taraxacum fibratum Sonck
Asteraceae
MCL 2:748 MFG 2:560
L/121 L/1211 L/2200 L/2201
Taraxacum extimum Kirschner, Sonck & Štepánec
Asteraceae
MCL 2:758
Taraxacum egnatiae Sonck
Asteraceae
MCL 2:747
Taraxacum edessicum Sonck
Asteraceae
MCL 2:747 MFG 2:564
Taraxacum divinum Sonck
Asteraceae
MCL 2:747
Taraxacum dialeptum Sonck
Asteraceae
MCL 2:744
L/1211 L/1880 L/2201
Taraxacum deorum A.J.Richards
Asteraceae
MCL 2:775 MFG 2:548
Taxon
?
+
.
.
+
.
+
+
.
+
.
.
.
.
.
.
StE
.
.
.
.
.
.
Pe
.
.
.
.
.
.
.
.
.
.
.
.
.
.
IoI
.
.
.
.
.
.
.
.
WAe
+
+
+
.
.
.
.
.
.
.
.
SPi
+
+
.
.
.
.
.
.
.
.
.
NPi
.
.
.
.
.
.
.
.
.
.
.
EC
+
.
.
+
+
.
.
+
.
+
+
NC
.
.
.
.
.
.
.
.
.
.
.
NE
.
.
.
.
.
.
.
.
.
.
.
NAe
.
.
.
.
.
.
.
.
.
.
.
Kik
.
.
.
.
.
.
.
.
.
.
.
KK
.
.
.
.
.
.
.
.
.
?
.
EAe
*
*
*
*
Distribution
Qualifier
348 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
A
U
U
U
U
A
U
A
S
U
1359
1360
1361
1362
1363
1364
1365
1366
1367
1368
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Taraxacum olympicola Sonck
Asteraceae
MCL 2:750 MFG 2:560
L/1211
Taraxacum nudum Soest
Asteraceae
MCL 2:775 MFG 2:546
L/1196
Taraxacum noterophilum Kirschner, Sonck & Štepánec
Asteraceae
MCL 2:759
L/2254
Taraxacum nanulum Sonck
Asteraceae
MCL 2:749
Taraxacum molybdocephalum Sonck
Asteraceae
MFG 2:561
L/394 L/2200
Taraxacum magnesicum Sonck
Asteraceae
MCL 2:749
Taraxacum lingulilobum Sonck
Asteraceae
MCL 2:749 MFG 2:560
Taraxacum kalambakae Sonck
Asteraceae
MCL 2:767
L/2254
Taraxacum herae Sonck
Asteraceae
MCL 2:748
Taraxacum graecum Dahlst.
Asteraceae
FE 4:334 FT 5:799 MCL 2:774
L/566 L/2200 L/2244
.
.
.
.
.
.
.
.
+
.
.
+
.
.
.
.
.
.
+
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
+
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
+
.
.
.
+
.
+
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
+
*
*
*
*
PATTERNS OF PLANT ENDEMISM IN GREECE 349
U
S
A
U
U
U
A
U
1372
1373
1374
1375
1376
1377
1378
A
1370
1371
U
Taxon
Qualifiers
1369
Taxon
Number
Taraxacum submicrocranum Sonck
Asteraceae
MCL 2:751
L/2254
Taraxacum sublimiforme Sonck
Asteraceae
MCL 2:772
L/2254
Taraxacum salonikiense Sonck
Asteraceae
MCL 2:751
Taraxacum rigidifolium Sonck
Asteraceae
MCL 2:771
L/2254
Taraxacum radinum Sonck
Asteraceae
MCL 2:751
Taraxacum protervum Sonck
Asteraceae
MCL 2:750 MFG 2:562
L/394 L/2200
Taraxacum pindicum Kirschner & Štěpánec
Asteraceae
MCL 2:762 MFG 2:545
L/1963 L/2245
Taraxacum phitosi Soest
Asteraceae
MCL 2:750
Taraxacum panhellenicum Sonck
Asteraceae
MCL 2:745 MFG 2:572
L/1963
Taraxacum olympophilum Sonck
Asteraceae
MCL 2:745
L/2253
Taxon
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
StE
.
.
.
+
.
Pe
.
.
.
.
.
.
.
.
.
.
WAe
.
.
.
.
.
.
.
.
.
.
IoI
.
.
.
.
.
.
.
.
+
.
SPi
+
+
.
+
.
.
+
.
+
.
NPi
.
.
.
.
.
.
.
.
+
.
EC
.
.
.
.
.
+
.
.
.
+
NC
.
.
.
.
.
.
.
.
.
.
NE
.
.
.
.
.
.
.
.
.
.
NAe
.
.
.
.
.
.
.
.
.
.
Kik
.
.
.
.
.
.
.
.
.
.
KK
.
.
.
.
.
.
.
.
.
.
EAe
*
*
*
Distribution
Qualifier
350 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
U
U
A
U
A
1382
1383
1384
1385
1386
1387
A
1381
A
U
!
1380
1379
Telephium imperati L. subsp. pauciflorum (Greuter) Greuter &
Burdet
Caryophyllaceae
Telephium imperati L. subsp. orientale (Boiss.) Nyman var. pauciflorum
Greuter
Telephium imperati L. var. pauciflorum Greuter
AFE 6:168 EFK :85 FAe :132 FCA :53 FCSI :23 FE 1:188 FHe 1:238 MCL 1:288 MFG 1:135
L/388 L/408 L/518 L/554
Taraxacum zagorae Sonck
Asteraceae
MCL 2:773
L/2254
Taraxacum xanthiense Soest
Asteraceae
FE 4:340 MCL 2:752 MFG 2:557
L/2245
Taraxacum viale Sonck
Asteraceae
MCL 2:752
L/1211 L/2201 L/2253
Taraxacum vexatum Sonck
Asteraceae
Taraxacum lacistophylloides Sonck, non Dahlst.
MCL 2:752 MFG 2:564
Taraxacum trigonense Sonck
Asteraceae
MCL 2:772
L/2254
Taraxacum thessalicum Soest
Asteraceae
MCL 2:775 MFG 2:549
Taraxacum terenodes Sonck
Asteraceae
MCL 2:752
L/1211 L/2201
Taraxacum subolivaceum Sonck
Asteraceae
MCL 2:760 MFG 2:551
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
+
.
.
.
.
.
.
.
.
.
.
.
.
+
.
+
.
.
.
.
.
.
.
.
.
.
+
.
.
+
+
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
+
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
*
PATTERNS OF PLANT ENDEMISM IN GREECE 351
Teucrium aroanium Orph. ex Boiss.
Lamiaceae
FE 3:131 MCL 3:367 MFG 2:70
L/395 L/607 L/1915 L/1963
Teucrium chamaedrys L. subsp. olympicum Rech.f.
Lamiaceae
MCL 3:368 MFG 2:72
Teucrium cuneifolium Sm.
Lamiaceae
EFK :258 FCA :101 FE 3:133 FGC 2:x MCL 3:369
L/268
Teucrium divaricatum Heldr. subsp. athoum (Hausskn.) Bornm.
Lamiaceae
Teucrium divaricatum Heldr. var. athoum Hausskn.
FE 3:132 MCL 3:369
L/367 L/447 L/1832d L/1973
Teucrium flavum L. subsp. gymnocalyx Rech.f.
Lamiaceae
FCA :101 FE 3:133 MCL 3:370
L/323b L/2013
Teucrium francisci-werneri Rech.f.
Lamiaceae
Teucrium werneri Rech.f.
FE 3:131 MCL 3:372 RDB :488
L/119 L/607 L/757 L/765 L/1877 L/1963
1390
1391
1392
1393
1394
1395
Tetragonolobus wiedemannii Boiss.
Fabaceae
Lotus wiedemannii (Boiss.) Hayek
FAe :373 FE 2:177 FGC 1:416 MCL 4:131
L/776
Taxon
Teucrium alpestre Sm.
Lamiaceae
Teucrium alpestre Sibth. & Smit subsp. alpestre
Teucrium gossypium Rech.f. (included in taxon)
EFK :258 FCA :101 FE 3:134 MCL 3:366 MFG 2:72
L/1766
?
Taxon
Qualifiers
1389
1388
Taxon
Number
.
.
+
+
+
.
.
.
.
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StE
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IoI
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SPi
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NPi
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NC
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EAe
Distribution
Qualifier
352 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Thamnosciadium junceum (Sm.) Hartvig
Apiaceae
Sclerochorton junceum (Sm.) Boiss.
Seseli junceum Sm.
Seseli stylare Boiss. & Heldr. (from Taygetos)
FE 2:333 MFG 1:709
L/607 L/1042 L/1694 L/1963
Thesium vlachorum Aldén
Santalaceae
FE 1:85 FHe 1:66 MFG 1:58 RDB :492
L/898
Thlaspi bulbosum Boiss.
Brassicaceae
Raparia bulbosa (Spruner) F.K. Mey.
AFE 11:153 FE 1:386 FGC 1:107 FHe 2:258 FT 1:334 MCL 3:162 PFPB 1:475
L/121 L/282 L/321 L/323a L/361 L/363 L/530 L/539 L/540 L/736 L/1871 L/1957 L/2200
1400
1401
Teucrium montbretii Benth. subsp. heliotropifolium (Barbey)
P.H.Davis
Lamiaceae
Teucrium heliotropifolium Barbey
EFK :257 FCA :102 FCSI :55 FE 3:131 FT 11:198 FT 7:70 MCL 3:373 RDB :490
L/196 L/197 L/789 L/981 L/1049
1398
1399
Teucrium halacsyanum Heldr.
Lamiaceae
Teucrium corcyanum Quézel & Contandr.
FE 3:131 FGC 2:471 MCL 3:373
L/537 L/607 L/1123 L/1690 L/1800 L/1963
1397
!
Teucrium gracile Barbey & Major
Lamiaceae
Teucrium alpestre Barbey & Major (sphalm.) subsp. gracilis
Teucrium alpestre Sibth. & Smit subsp. gracile (Barbey & Major)
D.Wood
EFK :258 FCA :101 FE 3:134 MCL 3:371 MFG 2:73
L/201 L/268 L/273 L/1049
1396
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PATTERNS OF PLANT ENDEMISM IN GREECE 353
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Thlaspi epirotum Halácsy
Brassicaceae
Noccaea epirota (Halácsy) F.K.Mey.
Thlaspi epiroticum Halácsy (sphalm.)
AFE 11:156 FE 1:387 FGC 1:109 FHe 2:258 MCL 3:165 MFG 1:324 PFPB 1:478
L/513
Thlaspi graecum Jordan
Brassicaceae
Thlaspi taygeteum Boiss.
Noccaea graeca (Jordan) F.K.Mey.
AFE 1:152 FE 1:386 FGC 1:107 FHe 2:256 MCL 3:169 MFG 1:322 PFPB 1:475
L/159 L/512 L/561 L/607 L/1851 L/1963
Thlaspi zaffranii (F.K.Mey.) Greuter & Burdet
Brassicaceae
Noccaea zaffranii F.K.Mey.
AFE 11:157 EFK :122 FCA :86 FCSI :44 FHe 2:258 MCL 3:171 RDB :494
L/346 L/518 L/942 L/1095 L/1766
Thymbra calostachya (Rech.f.) Rech.f.
Lamiaceae
Micromeria calostachya Rech.f.
EFK :265 FCA :102 FE 3:170 MCL 3:380 RDB :496
L/255 L/554 L/1583
Thymus dolopicus Formánek
Lamiaceae
Thymus euboeus Halácsy
Thymus serpyllum L. var. angustifolius sensu Boiss. Fl. Or. p.p.
FAe :540 FE 3:176 FGC 2:560 MCL 3:382 MFG 2:148
L/323a L/593 L/1871
1403
1404
1405
1406
1407
Taxon
Thlaspi creticum (Degen & Jav.) Greuter & Burdet
Brassicaceae
Noccaea cretica (Degen & Jav.) F.K.Mey.
Thlaspi graecum Jordan subsp. creticum (Degen & Jav.) Greuter
Thlaspi graecum sensu suct. cret., Fl. Eur.; Greuter 1974
Thlaspi microphyllum Boiss. & Orph. subsp. creticum Degen & Jav
AFE 11:152 EFK :122 FAe :242 FCA :86 FE 1:386 FGC 1:107 FHe 2:257 MCL 3:161 MFG 1:323 PFPB 1:474
L/518 L/522 L/561 L/830
Taxon
Qualifiers
1402
Taxon
Number
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EAe
Distribution
Qualifier
354 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Thymus hartvigi Morales
Lamiaceae
Thymus rechingeri Hartvig
Thymus teucrioides Boiss. & Spruner var. deorum Ronniger p.p.
Thymus teucrioides sensu Boiss. Fl. Or. p.p.
FGC 2:567 MFG 2:145 RDB :498
L/258 L/593 L/1871 L/1963 L/2013 L/2200
Thymus hartvigi Morales subsp. hartvigi
Lamiaceae
Thymus rechingeri Hartvig subsp. rechingeri
FGC 2:567 MFG 2:147 RDB :498
L/258 L/593 L/1963 L/2013
Thymus hartvigi Morales subsp. macrocalyx (Hartvig) Morales
Lamiaceae
Thymus rechingeri Hartvig subsp. macrocalyx Hartvig
Thymus rechingeri Hartvig subsp. rechingeri
MFG 2:147 RDB :498
L/121 L/258 L/593 L/1467 L/1871 L/1963 L/2200
Thymus holosericeus Celak.
Lamiaceae
Thymus leucadicus (Rchb.f.) Halácsy
Thymus striatus sensu auct., non Val.
FE 3:177 FGC 2:562 MCL 3:383 MFG 2:151
L/593 L/979 L/1123 L/1244
Thymus laconicus Jalas
Lamiaceae
Thymus pastoralis Turrill non Iljin
FE 3:177 MCL 3:383
B/3060a L/487 L/593 L/607 L/1915 L/1963
Thymus leucospermus Hartvig
Lamiaceae
Thymus teucrioides Boiss. & Spruner var. deorum Ronniger p.p.
Thymus teucrioides sensu Boiss. Fl. Or. p.p.
MFG 2:147
L/258 L/593
Thymus plasonii Adamović
Lamiaceae
Thymus zygioides var. plasonii (Adamović) H.Braun
FE 3:178 MCL 3:392
L/138 L/487 L/566 L/593 L/1729
1408
1409
1410
1411
1412
1413
1414
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+
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© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 355
Thymus teucrioides Boiss. & Spruner subsp. candilicus (Beauv.)
Hartvig
Lamiaceae
Thymus teucrioides Boiss. & Spruner var. candilicus Beauv.
MFG 2:145
L/258 L/530 L/736 L/781 L/995 L/1871 L/1901
Tragopogon brevirostris DC. subsp. longifolius (Heldr. & Sart. ex
Boiss.) I.B.K.Richardson
Asteraceae
Tragopogon longifolius Heldr. & Sart. ex Boiss.
FE 4:324 FGC 2:195
L/607 L/1871 L/1963 L/2013
Tragopogon lassithicus Rech.f.
Asteraceae
EFK :335 FCA :73 FCSI :35 FE 4:324 RDB :500
L/255 L/518 L/522 L/554
Trifolium andricum Lassen
Fabaceae
L/1758 L/2181
Trifolium aurantiacum Boiss. & Spruner
Fabaceae
Chrysaspis aurantiaca (Boiss. & Spruner) Hendrych
Trifolium agrarium L., nom. ambig.
EFK :162 FAe :359 FCA :112 FCSII : FE 2:166 FGC 1:405 MCL 4:181 MFG 1:510
L/159 L/561 L/607 L/1600 L/1830
Trifolium barbeyi Gibelli & Belli
Fabaceae
EFK :165 FCA :112 FCSI :62 FE 2:169 MCL 4:180 RDB :502
L/197 L/365 L/1067
Trifolium dolopium Heldr. & Hausskn. ex Gibelli & Belli
Fabaceae
Chrysaspis dolopia (Gibelli & Belli) Hendrych
FAe :359 FE 2:166 FGC 1:405 MCL 4:182
L/1973
1416
1417
1418
1419
1420
1421
1422
Taxon
Thymus samius Ronniger & Rech.f.
Lamiaceae
FAe :540 FT 7:363 MCL 3:392 MFG 2:154
L/593 L/796 L/975
Taxon
Qualifiers
1415
Taxon
Number
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StE
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WAe
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SPi
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NPi
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NC
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EAe
Distribution
Qualifier
356 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Trifolium praetermissum Greuter, Pleger & Raus
Fabaceae
EFK :162 FCA :114 FCSI :63 FCSII : MCL 4:189
L/253 L/273 L/522 L/551 L/1686
Trigonella rechingeri Sirj.
Fabaceae
EFK :157 FCA :115 FE 2:151 FT 11:96 MCL 4:195
L/196 L/197 L/201 L/206 L/268 L/423 L/495 L/537 L/551 L/776 L/1049 L/1710 L/1878
Trinia frigida (Boiss. & Heldr.) Drude
Apiaceae
Apinella frigida (Boiss. & Heldr.) Halácsy, Consp. Fl. Gr. p.p.
Apinella guicciardi (Boiss. & Heldr.) Halácsy, Consp. Fl. Gr. p.p.
Rumia frigida Boiss. & Heldr.
FE 2:351 FGC 1:674 MFG 1:694
L/121 L/160 L/512 L/607 L/1112 L/1123 L/1963 L/2013
Trinia guicciardii (Boiss. & Heldr.) Drude
Apiaceae
Apinella frigida (Boiss. & Heldr.) Halácsy, Consp. Fl. Gr. p.p.
Apinella guicciardi (Boiss. & Heldr.) Halácsy, Consp. Fl. Gr. p.p.
FE 2:351 FGC 1:675 MFG 1:695
L/512 L/530 L/607 L/1246
Trisetum laconicum Boiss. & Orph.
Poaceae
Trisetum rechingeri Chrtek
Avena laconica (Boiss. & Orph.) Nyman
FAe :791 FE 5:221 FGC 3:375
L/528 L/607 L/1588 L/1871 L/1963
1425
1426
1427
1428
1429
Trifolium parnassi Boiss. & Spruner
Fabaceae
FE 2:162 MCL 4:188 MFG 1:500
L/121 L/160 L/512 L/513 L/607 L/1112 L/1128 L/1246 L/2013 L/2200
Trifolium phitosianum Böhling, Greuter & Raus
Fabaceae
FCSII :
L/522 L/1686
!
1424
1423
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 357
Valeriana crinii Orph. ex Boiss. subsp. crinii
Caprifoliaceae
Valeriana montana L. subsp. crinii Orph. ex Boiss.
Valeriana montana L. var. crinii (Orph. ex Boiss.) Boiss
MFG 2:347 RDB :508
L/160 L/930 L/1064 L/1963
Umbilicus parviflorus (Desf.) DC.
Crassulaceae
Umbilicus sprunerianus Boiss.
Cotyledon parviflora Desf.
AFE 12:47 FCA :77 FE 1:424 FHe 2:307 FT 4:211 MCL 3:33
L/150 L/530 L/607 L/869 L/1957
1435
1437
Umbilicus albido-opacus A.Carlström
Crassulaceae
FHe 2:307 FT 10:140 MCL 3:33
L/197 L/279 L/355
1434
Valeriana asarifolia Dufr.
Caprifoliaceae
EFK :290 FCA :155 FCSII : FE 4:53
B/3060c L/551 L/856 L/1917
Tulipa goulimyi Sealy & Turrill
Liliaceae
EFK :352 FCA :186 FE 5:29 RDB :504
B/3060c L/119 L/267 L/450 L/607 L/765 L/785 L/810 L/905 L/964 L/1194 L/1210d L/1219 L/1917 L/1963
1433
1436
Tulipa doerfleri Gand.
Liliaceae
Tulipa hageri Heldr. var. doerfleri (Gand.) Hayek
EFK :353 FAe :722 FCA :186 FE 5:29 MFG 2:672
L/707
1432
!
Tulipa cretica Boiss. & Heldr.
Liliaceae
FCA :186 FE 5:30
1431
Taxon
Tulipa bakeri A.D.Hall
Liliaceae
EFK :353 FCA :186 FE 5:30 MFG 2:671
Taxon
Qualifiers
1430
Taxon
Number
.
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+
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WAe
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IoI
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SPi
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NPi
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NC
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NE
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NAe
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Kik
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+
+
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+
+
+
+
KK
.
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+
+
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EAe
Distribution
Qualifier
358 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Valeriana olenaea Boiss. & Heldr.
Caprifoliaceae
FE 4:54 MFG 2:345
L/512 L/607 L/1963
Verbascum acaule (Bory & Chaub.) O. Kuntze
Scrophulariaceae
Celsia acaulis Bory & Chaub.
FE 3:210 MFG 2:173
L/160 L/499 L/512 L/607 L/1963
Verbascum adeliae Heldr. ex Boiss.
Scrophulariaceae
FAe :468 FE 3:213 FGC 2:390
L/169 L/869
Verbascum aphentulium Heldr.
Scrophulariaceae
Verbascum pelium Halácsy
FE 3:212 FGC 2:382 MFG 2:181
L/122 L/458 L/890a
Verbascum arcturus L.
Scrophulariaceae
EFK :274 FCA :144 FE 3:209
L/1583 L/1766
Verbascum boissieri (Heldr. & Sart. ex Boiss.) O. Kuntze
Scrophulariaceae
Celsia boissieri Heldr. & Sart.
Celsia daenzeri Fauche & Chaub. var. lyrata Boiss. & Heldr.
FE 3:209 FGC 2:397
L/447 L/530 L/736 L/781 L/1832d L/1871
Verbascum botuliforme Murb.
Scrophulariaceae
FE 3:213
Verbascum cylindrocarpum Griseb.
Scrophulariaceae
FAe :467 FE 3:213
L/1267b L/1349
1438
1439
1440
1441
1442
1443
1444
1445
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+
.
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+
+
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
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PATTERNS OF PLANT ENDEMISM IN GREECE 359
Verbascum daenzeri (Fauche & Chaub.) O.Kuntze
Scrophulariaceae
Celsia daenzeri Fauche & Chaub.
FE 3:208
L/395 L/607 L/1963 L/2181
Verbascum delphicum Boiss. & Heldr.
Scrophulariaceae
FE 3:215 MFG 2:186
L/1871
Verbascum epixanthinum Boiss. & Heldr.
Scrophulariaceae
FE 3:212 MFG 2:179
L/121 L/607 L/1112 L/1467 L/1871 L/1963 L/2013 L/2200
Verbascum epixanthinum Boiss. & Heldr. var. epixanthinum
Scrophulariaceae
Verbascum epixanthinum Halácsy var. taygeteum
Verbascum epixanthinum Boiss. & Heldr. var. taygeteum (Halácsy)
Murb
MFG 2:180
L/1963 L/2013
Verbascum epixanthinum Boiss. & Heldr. var. parnassicum
(Halácsy) Murb.
Scrophulariaceae
Verbascum parnassicum Halácsy
MFG 2:180
L/1467 L/1963
Verbascum epixanthinum Boiss. & Heldr. var. pindicolum
(Freyn & Sint.) Murb.
Scrophulariaceae
Verbascum agrimonioides Degen & Borbas
Verbascum tymphaeum Freyn & Sint.
Verbascum adenotrichum Halácsy
Verbascum viscidulum Freyn & Sint.
Verbascum pindicolum Freyn & Sint.
MFG 2:180
L/1963
1447
1448
1449
1450
1451
1452
Taxon
Verbascum cylleneum (Boiss. & Heldr.) O.Kuntze
Scrophulariaceae
Celsia acaulis subsp. cyllena (Boiss. & Heldr.) O.Kuntze
Celsia cyllenea Boiss. & Heldr.
FE 3:210 MFG 2:173 RDB :510
L/395 L/607 L/1963
Taxon
Qualifiers
1446
Taxon
Number
+
+
+
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+
+
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IoI
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StE
+
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Pe
+
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SPi
+
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NPi
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EC
+
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NC
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.
.
NE
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.
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.
.
.
.
NAe
.
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.
.
+
.
Kik
.
.
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.
.
.
.
KK
.
.
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.
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EAe
Distribution
Qualifier
360 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Verbascum pangaeum Murb. & Rech.f.
Scrophulariaceae
FE 3:210 MFG 2:176
L/1107 L/1996
Verbascum pentelicum Murb.
Scrophulariaceae
FE 3:213
1461
Verbascum orphanideum Murb.
Scrophulariaceae
FE 3:213
L/607
1459
1460
Verbascum longifolium Ten. var. samaritanii (Hedr.) Murb.
Scrophulariaceae
Verbascum epirotum Halácsy
Verbascum epixanthinum Boiss. & Heldr. var. samaritanii Boiss.
Verbascum samaritanii Heldr. ex Boiss.
FE 3:211 FGC 2:379 MFG 2:178
1458
?
Verbascum ikaricum Murb.
Scrophulariaceae
FAe :468 FT 6:576 MFG 2:187
L/111 L/539 L/540 L/796
1457
Verbascum hypoleucum Boiss. & Heldr.
Scrophulariaceae
Verbascum nigrum subsp. hypoleucum (Boiss. & Heldr.) Nyman
FE 3:216 FGC 2:392
L/607
1456
?
Verbascum halacsyanum Sint. & Bornm. ex Halácsy
Scrophulariaceae
FE 3:213
1455
Verbascum friedrichsthalianum O.Kuntze
Scrophulariaceae
Celsia speciosa Fenzl
FE 3:208 FGC 3:396
L/607
1454
?
Verbascum euboicum Murb. & Rech.f.
Scrophulariaceae
FAe :466 FE 3:212
L/323a L/1871
1453
.
.
+
.
.
+
.
+
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+
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+
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+
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+
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 361
Verbascum spathulisepalum Greuter & Rech.f.
Scrophulariaceae
L/755
Verbascum speciosum Schrader subsp. megaphlomos
(Boiss. & Heldr.) Nyman
Scrophulariaceae
Verbascum megaphlomos (Boiss. & Heldr.) Halácsy
Verbascum megaphlomos (Boiss. & Heldr.) Halácsy var. macrophyllum
Halácsy
Verbascum speciosum Schrader var. megaphlomos Boiss. & Heldr.
FE 3:213 FGC 2:386 MFG 2:182
L/395 L/607 L/890a L/1963 L/2013
Verbascum spinosum L.
Scrophulariaceae
EFK :274 FCA :144 FCSII : FE 3:208 MFG 2:172
L/1132 L/1766
Verbascum zuccarinii (Boiss.) I.K.Ferguson
Scrophulariaceae
Celsia tomentosa Zucc.
Verbascum graecum Heldr. & Sart. var. zuccarinii Boiss.
Verbascum tomentosum (Zucc.) O.Kuntze
FE 3:210 FGC 2:393
L/323a L/1010 L/1871
Veronica chamaedrys L. subsp. chamaedryoides (Bory & Chaub.)
M.A.Fisch.
Plantaginaceae
Veronica chamaedryoides Bory & Chaub.
Veronica chamaedrys L. var. chamaedryoides (Bory & Chaub.) Nyman
Veronica chamaedrys L. var. pindica Hausskn.
Veronica chamaedrys L. var. rigida Turrill
Veronica rigida (Turrill) Turrill
MFG 2:222
L/121 L/395 L/530 L/1267b L/1846 L/1871 L/1963 L/2013 L/2200 L/2201
1464
1465
1466
1467
1468
!
Verbascum reiseri Halácsy
Scrophulariaceae
FE 3:215 MFG 2:186
L/330
1463
Taxon
Verbascum phlomoides L. var. cycladum Murb.
Scrophulariaceae
Verbascum sartorii Halácsy non Boiss. & Heldr.
FAe :466
L/367 L/776 L/2181
Taxon
Qualifiers
1462
Taxon
Number
.
.
+
.
.
+
+
.
+
.
+
.
.
.
StE
Pe
+
+
+
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.
IoI
.
.
+
WAe
+
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.
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SPi
+
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+
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NPi
+
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EC
+
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NC
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+
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NAe
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+
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.
NE
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.
+
Kik
.
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+
.
.
.
.
KK
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.
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.
.
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EAe
Distribution
Qualifier
362 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
1475
Veronica oetaea Gustavsson
Plantaginaceae
MFG 2:224 RDB :512
L/138 L/330 L/1961 L/2013
Veronica glauca Sm. subsp. peloponnesiaca (Boiss. & Orph.) Maire &
Petitm.
Plantaginaceae
Veronica peloponnesiaca Boiss. & Orph.
FAe :481 FE 3:249 FGC 2:433 MFG 2:226
L/530 L/1244 L/1846 L/1871 L/1963 L/1973 L/2013 L/2181
1474
!
Veronica glauca Sm. subsp. kavusica (Rech.f.) M.A.Fisch.
Plantaginaceae
Veronica kavusica Rech.f.
Veronica praecox sensu Rechinger, 1943
EFK :280 FCA :144 FE 3:248 MFG 2:228
L/253
1473
Veronica glauca Sm. subsp. glauca
Plantaginaceae
Veronica amoena sensu Heldr., non Bieb.
Veronica graeca Spruner, nomen nudum
EFK :280 FAe :481 FE 3:249 MFG 2:226
L/530 L/1871
1472
!
Veronica glauca Sm. subsp. chaubardii (Boiss. & Reuter) Maire &
Petitm.
Plantaginaceae
Veronica peduncularis Bory & Chaub., non Bieb.
FGC 2:433 MFG 2:228
L/1830
1471
Veronica contandriopouli Quézel
Plantaginaceae
FE 3:244 MFG 2:216
L/512 L/607 L/1144 L/2237
Veronica erinoides Boiss. & Spruner
Plantaginaceae
FE 3:244 MFG 2:216
L/607 L/947 L/1963 L/2200
?
1470
1469
+
+
.
.
.
+
+
.
.
+
.
+
.
.
+
.
.
.
+
+
+
.
+
+
.
+
.
.
.
+
.
.
.
.
.
.
+
.
.
.
.
.
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+
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.
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+
.
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+
.
.
.
.
.
.
.
+
+
.
.
.
.
.
.
.
.
.
.
*
PATTERNS OF PLANT ENDEMISM IN GREECE 363
!
1476
Vicia pinetorum Boiss. & Spruner
Fabaceae
EFK :150 FAe :328 FCA :116 FE 2:131 FGC 1:486 MCL 4:209
L/145 L/323a L/361 L/561 L/869 L/1871 L/1973
Vicia villosa Roth subsp. maniatissa Kit Tan & Lassen
Fabaceae
L/1963
1481
1482
Vicia cretica Boiss. & Heldr. subsp. aegaea (Halácsy) P.W.Ball
Fabaceae
Vicia aegaea (Halácsy) Rech.f.
Vicia cretica Boiss. & Heldr. var. aegaea Halácsy
FAe :330 FE 2:133 MCL 4:204
L/128 L/958
1479
Vicia davisii Greuter
Fabaceae
Vicia villosa subsp. semicretica Ponert
FT 10:110 FT 11:90 FT 3:293 MCL 4:204
L/192
Veronica thymifolia Sm.
Plantaginaceae
Veronica cretica Pall. ex Link
Veronica tymfrestea Boiss. & Spruner
EFK :278 FCA :145 FE 3:246 MFG 2:219
L/121 L/160 L/512 L/587 L/607 L/1963 L/2013 L/2200
1478
1480
Veronica sartoriana Boiss. & Heldr.
Plantaginaceae
FCA :145 FE 3:249 FT 6:713 MFG 2:231
L/530 L/1963
Veronica orsiniana Ten. subsp. teucrioides (Boiss. & Heldr.)
M.A.Fisch.
Plantaginaceae
Veronica austriaca L. subsp. dentata (F.W.Schmidt) Watzl var.
teucrioides (Boiss. & Heldr.) Waltzl
Veronica austriaca L. var. teucrioides (Boiss. & Heldr.) Halácsy quoad
basion. solo
Veronica prostrata var. kajmakcalanica Adamović?
Veronica teucrioides Boiss. & Heldr.
FGC 2:427 MFG 2:220
L/121 L/785 L/2200 L/2201
Taxon
1477
?
Taxon
Qualifiers
Taxon
Number
.
+
.
.
.
+
.
+
+
+
+
?
.
+
StE
Pe
.
+
.
.
.
.
.
.
.
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+
.
IoI
.
.
WAe
.
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+
.
.
SPi
.
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.
.
NPi
.
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.
EC
.
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.
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+
+
NC
.
+
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.
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NE
.
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+
.
NAe
.
+
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+
.
.
.
Kik
.
?
.
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+
+
.
KK
.
.
+
.
.
+
.
EAe
Distribution
Qualifier
364 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Vincetoxicum creticum Browicz
Apocynaceae
EFK :233 FCA :38 FCSI :13 FCSII : MCL 1:58 MFG 2:10
L/252 L/519 L/708 L/1860
Viola athois W.Becker
Violaceae
FAe :256 FE 2:259 MFG 1:625
L/1972 L/1973
Viola cephalonica Bornm.
Violaceae
FE 2:279 MFG 1:631
L/306 L/1123
Viola chelmea Boiss. & Heldr.
Violaceae
Viola chelmea Boiss. & Heldr. subsp. chelmea sensu Tutin & al.,
Fl. Eur.; Strid M. Fl. Gr.
FE 2:273 MFG 1:616
L/160 L/607 L/822 L/1871 L/1963
Viola cretica Boiss. & Heldr.
Violaceae
EFK :196 FCA :156 FCSI :90 FE 2:273 MFG 1:615
L/308b L/822
Viola cretica Boiss. & Heldr. subsp. cretica
Violaceae
EFK :196 FCA :156 FCSI :90 FE 2:273 MFG 1:615
L/822
Viola cretica Boiss. & Heldr. subsp. glabra Tiniakou
Violaceae
EFK :196 FCA :156 FCSI :90 FE 2:273 MFG 1:615
L/822
1483
1484
1485
1486
1487
1488
1489
.
.
.
.
.
.
+
.
.
+
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.
.
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.
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+
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+
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+
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+
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+
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+
+
+
.
.
.
+
.
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.
.
.
.
.
PATTERNS OF PLANT ENDEMISM IN GREECE 365
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Viola euboea (Halácsy) Halácsy
Violaceae
Viola gracilis Sm. var. euboea Halácsy
Viola heterophylla Bertol subsp. euboea (Halácsy) W.Becker
Viola heterophylla Bertol var. euboea (Halácsy) W.Becker
FE 2:279 MFG 1:628
L/323a L/1871
Viola fragrans Sieber
Violaceae
EFK :196 FCA :156 FCSI :90 FE 2:277 MFG 1:622
L/1766
Viola ganiatsasii Erben
Violaceae
L/565
Viola graeca (W.Becker) Halácsy
Violaceae
Viola gracilis sensu auct., Fl. Or., Consp. Fl. Graec.
Viola heterophylla Bertol. subsp. graeca (W.Becker) W.Becker
Viola heterophylla Bertol. var. graeca W.Becker
MFG 1:627
L/121 L/160 L/512 L/513 L/587 L/911 L/931 L/1716
L/1846 L/1963 L/2013 L/2200 L/2201
Viola mercurii Orph. ex Halácsy
Violaceae
Viola heldreichiana Boiss. f. mercurii (Orph.) W.Becker
Viola tricolor L. var. demetria Boiss.
FE 2:281 MFG 1:638
L/512 L/607 L/1088 L/1716 L/1963
1492
1493
1494
1495
1496
Viola epirota (Halácsy) Raus
Violaceae
Viola declinata Waldst. & Kit. var. epirota Halácsy
Viola euboea (Halácsy) Halácsy var. epirota Halácsy
Viola heterophylla Bertol. subsp. epirota (Halácsy) W.Becker
MFG 1:632
L/830
!
1491
Taxon
Viola dirphya Tiniakou
Violaceae
L/608 L/822 L/1871
Taxon
Qualifiers
1490
Taxon
Number
+
+
.
.
.
.
.
+
.
.
.
.
IoI
.
.
.
.
+
.
+
WAe
.
.
.
.
.
.
.
.
StE
.
Pe
.
.
.
.
.
+
.
SPi
.
.
.
.
.
+
.
NPi
.
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.
.
.
EC
.
+
.
.
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.
.
NC
.
.
+
.
.
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NE
.
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.
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.
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NAe
.
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.
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.
.
Kik
.
.
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+
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.
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KK
.
.
.
.
.
.
.
EAe
Distribution
Qualifier
366 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
Viola oligyrtia Tiniakou
Violaceae
RDB :516
L/822 L/1716 L/1963
Viola parnonia Kit Tan, Sfikas & Vold
Violaceae
MFG 1:628
L/909 L/911 L/1264 L/1716 L/1716 L/1963
Viola phitosiana Erben
Violaceae
L/121 L/530 L/1716 L/1871 L/1963 L/2200 L/2201
Viola poetica Boiss. & Spruner
Violaceae
FE 2:277 MFG 1:623
L/306 L/330 L/512 L/587 L/2013 L/2200
Viola pseudograeca
Violaceae
MFG 1:628
L/911
Viola rausii Erben
Violaceae
L/306
Viola samothracica (Degen) Raus
Violaceae
Viola gracilis var. samothracica (Degen) Rech.f.
Viola olympica var. samothracica Degen
FAe :256
L/895 L/1252 L/1911
Viola serresiana Erben
Violaceae
L/1688
1497
1498
1499
1500
1501
1502
1503
1504
.
.
.
.
.
.
.
+
+
.
.
.
.
+
+
+
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
.
.
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+
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+
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.
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.
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+
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+
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+
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+
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+
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.
+
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+
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+
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.
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PATTERNS OF PLANT ENDEMISM IN GREECE 367
Viola voliotisii Erben
Violaceae
L/306 L/496
Viola vourinensis Erben
Violaceae
MFG 1:628
L/911 L/1901
Viscum album L. subsp. creticum
Santalaceae
EFK :61 FCA :117
L/1342
Zelkova abelicea (Lam.) Boiss.
Ulmaceae
Abelicea cretica (Spach) O.Kuntze
Planera abelicea (Lam.) Schultes
Planera cretica (Spach) Kotschy
Quercus abelicea Lam.
Ulmus abelicea (Lam.) Sm.
Zelkova crenata var. cretica Spach
Zelkova cretica (Spach) Spach
AFE 3:83 EFK :59 FAe :97 FCA :147 FCY 2:1463 FE 1:77 FHe 1:52 RDB :526
L/138 L/204 L/518 L/519 L/522 L/554 L/569 L/685 L/686 L/687 L/750 L/983 L/1927
1507
1508
1509
1510
Viola sfikasiana M. Erben
Violaceae
L/1716 L/1963
Taxon
Viola striis-notata (J.Wagner) Merxm. & Lippert
Violaceae
Viola cenisia L. var. striis notata J.Wagner
Viola grisebachiana Vis. f. integrifolia Stoj. & Jordanov
MFG 1:619 RDB :520
L/1180
Taxon
Qualifiers
1506
1505
Taxon
Number
.
.
.
.
.
.
.
.
.
StE
.
.
+
Pe
.
.
.
.
.
.
WAe
.
.
.
.
.
.
IoI
.
.
.
.
.
.
SPi
.
.
.
.
.
.
NPi
.
.
.
.
.
.
EC
.
.
+
+
+
.
NC
.
.
.
.
.
.
NE
.
.
.
.
.
.
NAe
.
.
.
.
.
.
Kik
+
+
.
.
.
.
KK
.
.
.
.
.
.
EAe
Distribution
Qualifier
368 K. GEORGHIOU and P. DELIPETROU
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 369
Alphabetic synonym list
Synonym
Endemic List Number
Abelicea cretica (Spach) O. Kuntze
1510
Aberboa lancifolia (Sprengel) DC.
390
Achillea umbellata Sm. subsp. vulgaris Heimerl
nom. illeg.
14
Achillea umbellata Sm. subsp. vulgaris Heimerl
var. monocephala Heldr. ex Heimerl
14
Achillea umbellata Sm. subsp. vulgaris Heimerl
var. polycephala Heimerl
14
Achillea umbellata Sm. var. pauciloba (Heldr. ex
Heimerl) Heimerl
12
Abies apollinis Link
1
Abies cephalonica Loudon f. arcadiaca Henkel &
W. Hochst
1
Abies cephalonica Loudon f. parnassica Henkel &
W. Hochst
1
Abies cephalonica Loudon var. apollinis (Link)
Beissner
1
Acinos graveolens (M. Bieb.) Link var. integrifolia
Raulin nom nud.
1162
Abies cephalonica Loudon var. graeca (Fraas) Liu
1
Acinos nanus P.H. Davis & Doroszenco
1162
Abies panachaica Heldr.
1
Acosta chalcidicaea (Hayek) Holub
364
Abies reginae-amaliae Heldr.
1
Acosta pallidior (Halácsy) Holub
349
Acantholimon androsaceum sensu auct. Samos
2
Acosta subsericans (Halácsy) Holub
425
Acantholimon echinus (L.) Boiss. subsp. creticum
(Boiss.) Papan. & Kokkini
3
Adonis aestivalis L. subsp. cretica (Hunth.)
C. Steinb.
17
Acantholimon echinus L. var. creticum Boiss.
3
2
Adonis dentata Delile subsp. cretica (Hunth.)
Riedl
17
Acantholimon echinus sensu auct. Samos
Acantholimon ulicinum sensu auct. cret.
3
17
Acer amaliae Boiss. (nom. illeg.)
5
Adonis microcarpa DC. subsp. cretica (Huth.)
Vierh.
Acer hyrcanum Fischer & C.A. Mey. subsp.
reginae amaliae (Boiss.) E. Murray
5
Aegilops biuncialis Vis. var. archipelagica Eig
19
Aegilops comosa Sm. var. subventricosa Boiss.
20
Acer opalus Mill. subsp. hyrcanum (Fisch. & C.A.
Mey.) var. reginae amaliae (Orph. ex Boiss.) Alden
5
Aethionema creticum Boiss. & Heldr.
28
Aethionema graecum Boiss. & Spruner var.
creticum (Boiss. & Heldr.) Halácsy
28
Aethionema orbiculatum auct. non (Boiss.) Hayek
24
Aethionema ovalifolium (DC.) Boiss.
28
Aethionema polygaloides DC.
28
Aethionema saxatile (L.) R. Br. var. creticum
(Boiss. & Heldr.) Halácsy
28
Agrostemma gracile auct.
29
Agrostemma gracile L. var. thessalum Bornm.
29
Acer reginae-amaliae Orph. ex Boiss.
5
Achillea aegyptiaca L. subsp. taygetea (Boiss. &
Heldr.) Hayek
13
Achillea aegyptiaca L. var. taygetea (Boiss. &
Heldr.) Halácsy
13
Achillea clavennae L. var. integrifolia Halácsy
11
Achillea erba-rotta All. subsp. olympica (Heimerl)
I.B.K. Richardson
8
Achillea moschata Wulfen subsp. olympica
Heimerl
8
Agrostemma thessalum (Formanek) Hayek
29
31
Achillea olympica (Heimerl) Halácsy
8
Alchemilla acutiloba Steven subsp. aroanica
Buser
Achillea pauciloba (Heldr. ex Heimerl) Heimerl
12
Achillea pauciloba (Heldr. ex Heimerl) Heimerl
var. Halαcsyana Heimerl
11
Achillea pauciloba (Heldr. ex Heimerl) Heimerl
var. integrifolia (Halácsy) Heimerl
Achillea tournefortii DC. (nom. illeg.)
11
7
Achillea umbellata Sm. subsp. monocephala
(Heldr. ex Heimerl) Tzanoud. & Iatroú
14
Achillea umbellata Sm. subsp. pauciloba Heldr.
ex Heimerl
12
Alectorolophus sintenisii Sterneck
1143
Alectrolophus pubescens Sterneck
1142
Alkanna baeotica DC.
34
Alkanna stribrnyi var. pelia Halácsy
37
Allium achaium Boiss. & Orph. var. parnassicum
Boiss.
75
Allium arvense Guss. var. trachypus Boiss.
84
Allium boryanum Kunth
51
Allium bourgeaui subsp. bourgeaui sensu Bothmer
1974 p.p.
45
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
370 K. GEORGHIOU and P. DELIPETROU
Allium callimischon sensu Rechinger 1949
52
Alyssum gionae Quezel & Contadriopoulos
Allium circinnatum Sieber, Rechinger 1943; Tutin
& al. 1980; Stearn 1978
55
Alyssum idaeum var. curetum (Gand.) Hayek
Allium cruentatum Sieber nom. nud.
52
Allium dilatatum Zahar.
65
Allium erythraeum sensu auct. non Griseb. p.p.
86
Allium obtusiflorum DC. (including taxon)
74
Allium obtusiflorum DC. (including taxon)
59
Allium obtusiflorum sensu auct. non DC. p.p.
86
Allium paniculatum L. subsp. euboicum (Rech. f.)
Stearn
227
95
Alyssum lunarioides Willd.
683
Alyssum montanum L. subsp. epiroticum
(Baumg.) Hayek
101
Alyssum montanum L. subsp. graecum (Halácsy)
Hayek
98
Alyssum montanum L. subsp. montanum proles
epiroticum Baumg.
101
60
Alyssum montanum L. subsp. montanum proles
humile Baumg.
100
Allium paniculatum sensu auct. karp., non L.
1759
85
Alyssum montanum L. subsp. montanum proles
laxum Baumg.
100
Allium parnassicum auct. non Halácsy
61
98
Allium sphaerocephalon L. subsp.
sphaerocephalon var. trachypus Boiss.
84
Alyssum montanum L. subsp. montanum proles
thessalum (Halácsy) Baumg.
Alyssum montanum L. var. angustifolium Heldr.
99
Allium sphaerocephalon L. var. aegaeum Heldr. &
Halácsy
84
Alyssum montanum L. var. graecum Halácsy p.p.
minor
99
Allium stamineum Boiss. var. hymettium Boiss.
67
88
Allium stamineum Orphanides, non Boiss.
61
Alyssum murale Waldst. & Kit. subsp. stojanoffi
(Nyár.) Dudley in J. Arnold Arbor. 1964 non
sensu orig.
Allium staticiforme sensu auct. cret., non Sm?
85
84
Alyssum nebrodense Tineo var. fragillimum
(Baldacci) Halácsy
91
Allium trachypus Boiss. & Spruner
Alsine confusa Boiss.
937
Alyssum olympicum Halácsy
Alsine eurytanica Boiss. & Heldr.
939
Alyssum oocarpum Gand.
910
Alsine graminifolia sensu auct. from Tymfi, Cons.
Fl. Graec.
944
Alyssum orientale Ard. var. alpinum Halácsy
228
Alsine pichleri Boiss.
943
Alsine recurva All. var. eurytanica (Boiss. &
Heldr.) Halácsy
939
Alsine setacea (Thuill.) Mert. & W.D.J. Koch
933
Alsine trichocalycina sensu Boissier
937
Alsine verna (L.) Wahlenb var. idaea Halácsy
Alyssoides cretica (L.) Medicus
Alyssum praecox auct. graec. non Boiss.
98
89
Alyssum repens subsp. trichostachyum (Rupr.)
Hayek
101
Alyssum repens var. stenophyllum Halácsy
101
Alyssum repens Rupr.
101
936
Alyssum saxatile L. var. alpinum (Halácsy)
Halácsy
228
910
Alyssum spathulaefolium Formánek
101
Alyssum alspestre L. var. fragillimum Baldacci
91
Alyssum spruneri Jordan & Fourr.
99
Alyssum argenteum sensu Boiss. Fl. Orient.
87
Alyssum thessalum Halácsy
98
Alyssum atlanticum sensu Sm. Fl. Graec. Prodr.
105
Alyssum tymphaeum (Hausskn.) Halácsy
251
Alyssum condensatum Boiss. & Hausskn. subsp.
flexibile (Nyár) sensu T.R. Dudley 1964
103
Alyssum umbellatum Desv. sensu T.R. Dudley et
al. in Fl. Turkey
108
Alyssum corymbosum sensu Halácsy Consp. Fl.
Graec. p.p. minor
227
Alyssum virescens Halácsy
101
Alyssum creticum L.
910
Alyssum curetum Gand.
95
Alyssum doerfleri var. taygeteum (Heldr.) Bornm.
106
Alyssum emarginatum Rouy
852
Alyssum fallacinum sensu auct. cret. non
Hausskn.
Alyssum foliosum sensu Halácsy Consp. Fl. Graec.
‘Pentelikon’ p.p.
87
104
Alyssum vourinonense T.R. Dudley & Rech. f.
98
Amaracus dictamnus (L.) Benth.
1062
Amaracus lirius (Halácsy) Hayek
1063
Amaracus pulcher (Boiss. & Heldr.) Briq.
1065
Amaracus scaber (Boiss. & Heldr.) Briq.
1065
Amaracus tournefortti (Aiton) Benth.
1061
Amaracus vetteri (Briq. & Barbey) Hayek
1067
Amelanchier chelmea (Halácsy) Browicz
109
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 371
Amelanchier cretica var. chelmea Halácsy
109
Ammanthus ageratifolius auct. cret.
Ammanthus filicaulis Boiss. & Heldr.
145
122
Anthemis heracleotica sensu Rechinger Fl. Aeg.
p.p.
128
Anthemis montana L. var. incana (Boiss.) Hayek
127
Ammanthus glaberrimus Rech.f.
129
Anthemis montana L. var. linnaeana Hal.
125
Ammanthus intermedius Gand. (non Anthemis
intermedia Guss. )
122
Anthemis montana L. var. spruneri (Boiss. &
Heldr.) Halácsy
127
Ammanthus maritimus auct. p.p. [non Boiss. &
Heldr.]
123
127
Ammanthus maritimus Boiss. & Heldr. (non
Anthemis maritima L. Chrysanthemum
maritimum)
122
Anthemis orientalis (L.) Degen subsp. incana
(Boiss.) Hayek var. spruneri (Boiss. & Heldr.)
Hayek
Anthemis orientalis (L.) Degen subsp. sibthorpii
(Griseb.) Hayek
126
Ammanthus maritimus Fl. Aeg. p.p.; Halácsy
1912
142
Anthemis panachaica Halácsy
125
132
Ammanthus tomentellus p.p.
123
Anthemis peregrina L. subsp. guicciardii (Heldr.
ex Sart.) Zangeri
Ammanthus tomentellus sensu Gandoger 1915,
1916, 1920, p.p.; Rechinger 1943
142
Anthemis peregrina L. var. guicciardii (Heldr. ex
Sart.) Boiss.
132
Anacamptis boryi (Rchb.f.) R.M. Bateman,
Pridgeon & M.W. Chase
1056
Anthemis peregrina L. var. heracleotica Boiss. &
Heldr.
132
Anchusa cesatiana Fenzl & Friedr.
1131
Anthemis pusilla subsp. ammanthiformis sensu
Greuter & Rechinger, 1967 p.p.
140
Anthemis pusilla Greuter var. liguliflora (Halácsy)
Greuter
134
Anthemis sibthorpii Griseb.
126
Anthemis spruneri Boiss. & Heldr.
127
Anthemis tomentosa L. subsp. heracleotica (Boiss.
& Heldr.) R. Fernandes
132
Anchusa hybrida Ten. var. setigera Halácsy
115
Anchusa macrosyrinx Rech.f.
115
Anchusa ovata auct non Lehm., Mazziari
116
Anchusa perlata Lam. nom. illeg.
116
Anchusa sartorii Guşul.
115
Anchusa spruneri Boiss.
114
Anchusa stylosa M. Bieb. subsp. spruneri (Boiss.)
Nyman
114
Anthemis tomentosa L. subsp. scopulorum (Rech.
f.) Grierson
137
Anchusa stylosa M. Bieb. var. major DC.
114
Anthemis tomentosa sensu Rechinger 1960 p.p.
145
Anchusa undulata Sm. var. maritima Rech.f.
115
Anthyllis aegaea Turrill
Anchusa variegata (L.) Lehm.
116
1427
Anchusa variegata (L.) Lehm. subvar. cephalonica
(Bornm. & Gusul.) Gusul.
116
Apinella frigida (Boiss. & Heldr.) Halácsy, Consp.
Fl. Gr. p.p.
116
Apinella frigida (Boiss. & Heldr.) Halácsy, Consp.
Fl. Gr. p.p.
1428
Anchusa variegata (L.) Lehm. var. cephalonica
Bornm. & Gusul.
118
Apinella guicciardi (Boiss. & Heldr.) Halácsy,
Consp. Fl. Gr. p.p.
1428
Androcymbium gramineum (Cav.) J.F. Macbr.
var. puniceum (Maire) Maire
119
Apinella guicciardi (Boiss. & Heldr.) Halácsy,
Consp. Fl. Gr. p.p.
1427
Anemone stellata var. heldreichii Boiss.
Anthemis boissieri Stoj. & Acht.
120
Aquilegia amaliae var. taygetea (Orph.) Hayek
148
Anthemis cretica (L.) Nyman var. liguliflora
Halácsy
134
Aquilegia amaliae Heldr. ex Boiss. subsp.
meridionalis Quézel & Contandr.
147
Anthemis cretica auct., non L.
120
Aquilegia taygetea Orph.
148
Anthemis filicaulis (Boiss. & Heldr.) Greuter f.
discoidea Greuter
128
Arabis muralis Bertol var. cretica (Boiss. &
Heldr.) B.M.G. Jones
149
Anthemis flexicaulis Rech.f.
143
150
Anthemis guicciardii Heldr. ex Sart.
132
Arabis ochroleuca Boiss. & Heldr. non (Lam.)
Lam.
Anthemis guicciardii sensu Rechinger Fl. Aeg.
p.p.; Hansen 1971
145
Arabis serpillifolia Vill. subsp. cretica (Boiss. &
Heldr) B.M.G. Jones
149
Anthemis heracleotica sensu Rechinger Fl. Aeg.
p.p.
134
Arenaria litoralis Phitos
158
Arenaria oxypetala Sm. sensu Halácsy
153
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
146
372 K. GEORGHIOU and P. DELIPETROU
Arenaria pamphylica sensu Hayek
153
Arenaria serpyllifolia L. subsp. aegaea (Rech.f.)
Akeroyd
151
Arenaria teddii Turrill
Armeria majellensis Boiss. var. brachyphylla
Boiss.
Asperula tournefortii Sieber ex Sprengel var.
revoluta Rech.f.
200
Astracantha cretica (Lam.) Podlech
206
152
Astracantha dolinicola Lam.
209
165
Astracantha monachorum (Širj.) Podlech
218
163
Astracantha thracica (Griseb.) Podlech subsp.
cyllenea (Fisch.) Greuter
208
Armeria undulata sensu Rechinger 1960, eub.
Arum maculatum sensu Greuter 1973
166
218
Asperula aristata L. subsp. peristeraea Halácsy ex
Hayek
170
Astracantha thracica (Griseb.) Podlech subsp.
monachorum (Širj.) Greuter
Astragalus baldaccii Degen
215
208
Asperula athoa Boiss.
197
Astragalus calavrytensis Beauverd & Topali
Asperula boissieri Heldr. ex Boiss. var. transiens
Halácsy
183
Astragalus creticus Lam. f. glaberrimus Greuter &
Zaffran
206
Asperula coa Rech.f.
180
Astragalus gilvus sensu Agerer-Kirchhof (1976)
211
Asperula crassifolia sensu Gandoger 1916,
Rechinger 1943
175
Astragalus hirsutus Raul, non Vahl.
213
Astragalus insulae-carpathi Eig
204
Asperula gionae Quezel & Contandr.
172
Astragalus monachorum Širj.
218
Asperula idaea Halácsy var. fallax Rech.f.
179
208
Asperula idaea Halácsy var. lassitica Rech.f.
179
Astragalus parnassi Boiss. subsp. cylleneus
(Fisch.) Hayek
Asperula incana Sm.
191
204
Asperula kritensis Coust. & Gand.
193
Astragalus tauricolus auct. non Boiss. Hayek
Prodr. fl. Balc.; Rechinger fl. Aeg.
Asperula longiflora Waldst. & Kit var. thessala
(Boiss. & Heldr.) Halácsy
170
Astragalus tauricolus Boiss. subsp. austoaegaeus
(Rech.f.) Ponert
204
Asperula lutea subsp. oetaea (Boiss.) Nyman
188
Astragalus tauricolus Boiss. var. niveus Barbey
204
Asperula lutea var. abbreviata Halácsy
168
Astragalus thracicus Griseb. subsp. cylleneus
(Boiss. & Heldr.) Strid
208
Asperula lutea var. pinifolia Boiss.
190
180
Athamanta macedonica (L.) Sprengel subsp.
arachnoidea (Boiss. & Orph.) Tutin
220
Asperula lutea var. rigida sensu FAe rhodos
Asperula lutea Sm. subsp. mungieri (Boiss. &
Heldr.) Ehrend. & Krendl
184
Athamanta macedonica (L.) Sprengel var.
arachnoidea (Boiss. & Orph.) Halácsy
220
Asperula lutea Sm. subsp. rigidula (Halácsy)
Ehrend.
194
Atriplex tatarica auct.
221
Atriplex tatarica L. subsp. recurva (D’Urv.) Rech.f.
221
Asperula lutea Sm. var. oetaea Boiss.
188
Atriplex tatarica L. var. recurva Halácsy
221
Asperula lutea Sm. var. pulvinaris Boiss.
172
Atriplex tatarica L. var. sibthorpii Halácsy
221
Asperula lutea Sm. var. rigida Boiss.
194
223
Asperula lutea Sm. var. rigidula Halácsy
194
Aubrieta deltoidea (L.) DC. var. degeniana
Baldacci
Asperula majori Barbey (included in taxon)
200
226
Asperula nitida sensu auct. graec.
190
Aubrieta gracilis Spruner ex Boiss. subsp.
thessala (Boiss.) Maire & Petitmengin
Asperula nitida Sm. var. hirtella Boiss. p.p.
187
223
Asperula stricta auct. eur., non Boiss.
189
Aubrieta gracilis Spruner f. degeniana (Bald.)
Hayek
Asperula stricta auct. eur., non Boiss. subsp.
pruinosa Ehrend.
189
Aubrieta gracilis Spruner subsp. glabrescens
(Turrill) Akeroyd
223
189
Aubrieta gracilis Spruner subsp. scardica (Wettst.)
Phitos var. degeniana (Baldacci) Phitos
223
Asperula stricta auct. eur., non Boiss. subsp.
stricta
194
Aubrieta gracilis Spruner var. thessala (Boissieu)
Hayek
226
Asperula stricta Boiss. var. stricta Boiss.
Asperula stricta Boiss. var. tomentosa Boiss.
194
Aubrieta intermedia auct. non Heldr. & Orph.
226
Asperula thessala Boiss. & Heldr.
170
Avena agropyroides Boiss.
775
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 373
Avena laconica (Boiss. & Orph.) Nyman
1429
Brassica nivalis Boiss. & Heldr.
479
Avena sibthorpii Nyman
775
Brassica nivea Boiss. & Spruner
254
Avenastrum cycladum Rech.f. & Scheffer
775
775
Brassica nivea Boiss. & Spruner subsp. cretica
(Lam.) Onno
254
Avenula agropyroides (Boiss.) J. Holub
Avenula cycladum (Rech.f. & Scheffer) Greuter
775
Brassicella nivalis (Boiss. & Heldr.) O.E. Schultz
479
Avenula peloponnesiaca J. Holub
775
259
Bellevalia brevipedicellata sensu auct. cret.
orient., non Turrill
232
Bufonia brachyphylla Boiss & Heldr. var.
cecconiana Baldacci
Bufonia brachyphylla Boiss. & Heldr. nom. illeg.
258
Bellevalia spicata Boiss.
231
Bufonia brachyphylla sensu auct. p.p.
259
Berteroa mutabilis (Vemt.) DC. subsp. obliqua
(Sm.) Bald. sensu Bald. 1897
234
Bufonia stricta var. cecconiana sensu auct. p.p.
260
259
Berteroa obliqua (Sm.) DC. var. pindicola
(Halácsy) Quezél & Contadr.
234
Bufonia stricta var. typica sensu Halácsy Consp.
Fl. Graec. p.p.
234
Bufonia stricta (Sm.) Gurke var. cecconiana
(Baldacci) Gurke
259
Berteroa obliqua sensu auct., Halácsy 1894
Berteroa stricta Boiss. & Heldr. var. pindicola
Halácsy
234
Bufonia stricta (Sm.) Gurke var. hymettia Halácsy
260
Betonica coronariaarboreacretica J. Bauhin
587
Bufonia stricta (Sm.) Gurke var. typica Halácsy
p.p.
260
Biarum abbreviatum Schott
239
Buglossoides cretica Rvinus
116
Biarum fraasianum (Schott) N.E. Br. nom.
superfl.
237
Buglossum creticum minimum odoratum flore
varion eleganti Jonqcuet
116
Biarum spruneri Boiss.
238
Buglossum creticum verrucosum Strisser
116
Biarum tenuifolium (L.) Schott var. abbreviatum
(Schott) Engler
239
116
Biarum tenuifolium auct. kret., Rechinger 1949
240
Buglossum creticum culicolisb procumbentibus,
flore variegato odorato, echii foliis verrucosis
Hermann
Biarum zelebori cret. sensu Greuter 1973, p.p.
240
Bunium pumilum Sm.
760
Biasolettia bornmuelleri Wollf.
754
Bupleurum glumaceum Greuter & Rechinger 1967
266
Biasolettia parnassica (Boiss. & Heldr.) Nyman
758
Butinia cretica Boiss. & Heldr.
755
Bolanthus graecus var. thymifolius (Sm.)
Barkoudah
249
Butinia parnassica (Boiss. & Heldr.) Heldr. ex
Nyman
758
Bolanthus thessalus (Jaub. & Spach) Barkoudah
249
Bolanthus thymifolius sensu Phitos 1981
243
Calamintha alpina (L.) Lam. subsp.
nomismophylla Rech.f.
Borago cretica minima Contareni non descripta
Bellucci
116
Calamintha cretica (L.) Lam.
Borago cretica Willd.
1334
16
1158
Calamintha sylvatica Bromf. subsp. ascendens
(Jordan) P.W. Ball
268
Borago variegata cretica Wheler
116
Campanula acarnanica Damboldt
292
Borragine murale variegata con flore odorato, di
Candia Zanoni
116
Campanula aizoides Zaffran ex Greuter
270
Borrago verrucosa minor flore variegato Muntig
116
Campanula anchusiflora auct. non Sibth. & Sm.
284
Botryanthus pulchellus (Heldr. & Sart.) Nyman
952
Campanula andrewsii A.DC. var. pelia Halácsy
320
Botryanthus sartorii Todaro
952
Campanula andrewsii A.DC. subsp. breueri Tocl
& Rohlena
300
Brachypodium pinnatum sensu Zaffran 1976
252
254
Campanula andrewsii A.DC. var. lavrensis Tocl
& Rohlena
300
Brassica cretica Lam. subsp. nivea (Boiss. &
Spruner) Gustafsson. & Snogerup
254
Campanula asperuloides (Boiss. & Orph.) Engler
subsp. taygetea (Quezel & Contandr.) Greuter &
Burdet
278
Brassica cretica Lam. var. nivea (Boiss. &
Spruner) O.E. Schulz (included in taxon)
Brassica fruticulosa Cyr.
253
Campanula attica Boiss. & Heldr.
290
Brassica montana Pourret subsp. cretica (Lam.)
O.E. Schulz
254
Campanula calaminthifolia Lam. var. olivieri
(A.DC.) Halácsy
279
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
374 K. GEORGHIOU and P. DELIPETROU
Campanula calavrytana Beauverd & Top.
326
Campanula carpatha auct. cyth. non Halácsy
318
Campanula celsii sensu auct. Euboea, Rechinger
1960 p.p.
289
Campanula celsii sensu auct. Skop., Fl. Aeg. p.p.
321
Campanula cephallenica Feer
Campanula corymbosa Desf.
Campanula cretica (A.DC.) D. Dietr
Campanula rupestris Sibth. & Sm. subsp.
eu-rupestris Hayek var. virescens (Halácsy)
Hayek
326
Campanula rupestris Sibth. & Sm. var. virescens
Halácsy
326
293
Campanula rupestris Sm. var. andrewsii (A.DC.)
Hayek f. breueri (Tocl & Rohlena) Hayek
300
307
Campanula rupestris Sm. var. gracilis Heldr.
285
Campanula rupestris Sm. var. spathulifolia
Turrill.
285
1331
Campanula drabifolia Steud.
313
Campanula drabifolia subsp. creutzburgii
(Greuter) Fed.
288
Campanula samothracica (Degen) Greuter &
Burdet subsp. samothracica
1332
Campanula drabifolia subsp. pinatzii (Greuter &
Phitos) Fed.
308
1333
Campanula drabifolia Sm. var. major Boiss.
290
Campanula samothracica (Degen) Greuter &
Burdet subsp. sporadum (Halácsy) Greuter &
Burdet
Campanula eriantha Hampe
281
Campanula eriantha Vent.
281
Campanula eriantha Vent. var. typica Heldr.
Campanula saxatilis auct. non L. Boiss. Fl. Or.
Suppl.
280
281
Campanula saxatilis L. var. cythaerensis
Beauverd & Top., nom. nud.
318
Campanula erucifolia Feer
299
Campanula saxatilis L. var. simonelli Baldacci
317
Campanula glomerata L. var. stenosiphon Boiss.
323
275
Campanula sibthorpiana Halácsy var. filicaulis
Halácsy
322
Campanula laciniata Andr., non L.
Campanula lavrensis A.DC. var. breueri (Tocl &
Rohlena) Phitos
300
Campanula spatulata subsp. sibthorpiana var.
filicaulis (Halácsy) Hayek
322
Campanula leutweinii Heldr.
297
322
Campanula lyrata auct. non Lam.
281
Campanula spatulata Sm. subsp. spruneriana
sensu auct. Kret., Rechinger 1943 p.p.
Campanula mollis auct. non L.
317
Campanula olivieri A.DC.
279
Campanula parnassica Boiss. & Spruner
621
Campanula patenticalyx Bocquet?
324
Campanula pauciflora Desf.
322
Campanula pelia Hausskn.
320
Campanula pelviformis Lam. var. micrantha DC.
307
Campanula reiseri var. leonis Halácsy
312
Campanula reiseri var. pelagia Phitos
312
Campanula reiseri sensu auct. Skop., Fl. Aeg. p.p.
321
Campanula rotundifolia L. subsp. sancta Hayek
272
Campanula rupestris subsp. anchusiflora (Sm.)
Hayek
274
Campanula rupestris subsp. celsii (A.DC.) Hayek
281
Campanula rupestris sensu auct. Athos, Fl. Aeg.
p.p.
300
Campanula rupestris sensu auct. Euboea,
Rechinger 1960 p.p.
Campanula sporadum s.l. sensu Gustafsson M. &
Snogerup S. (1974) from Skantzoura
1333
Campanula subidaea Gand.
329
Campanula tomentosa var. bracteosa Heldr., p.p.
275
Campanula tomentosa sensu auct. non Vent.,
Boiss. Fl. Or., p.p.
275
Campanula tomentosa Vent. subsp. eu-rupestris
Hayek
314
Campanula tomentosa Vent. var. brachyantha
Boiss.
314
Campanula tomentosa Vent., non Lam. var.
calycina Heldr.
289
Campanula tubulosa auct. cyth. non Lam.
318
Campanula tubulosa auct. non Lam. Karp.
280
Campanula tubulosa Lam. var. carpatha
(Halácsy) Hayek
280
Campanula tubulosa Lam. var. oreophila Phitos
280
287
Cardamine barbaraeoides Halácsy
330
330
Campanula rupestris sensu auct. Euboea,
Rechinger 1960 p.p.
291
Cardamine raphanifolia Pourret subsp.
barbareoides (Halácsy) Strid
Carlina curetum Heldr. ex Halácsy
334
Campanula rupestris Sibth. & Sm. subsp.
eu-rupestris Hayek var. andrewsii (DC.) Hayek
275
Carlina curetum Heldr. ex Halácsy subsp.
curetum
334
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 375
Caroselinum distans Gris.
843
Carthamus gracilimus Rech.f.
339
Carthamus leucocaulos Sm. var. arachnoideus
Davis
337
Carum rigidulum (Viv.) W.D.J. Koch ex DC.
subsp. palmatum Hartvig
340
Caryophyllus arborescenscreticus C. Bauhin
587
Caryophyllus creticus, arboreus Tournefort
585
Caryophyllus sylvestrisarboreus Alpini
587
Centaurea alba L. subsp. princeps (Boiss. &
Heldr.) Gugler
409
Centaurea argentea L. subsp. macrothysana
(Rech.f.) Turland & L. Chilton var. chionantha
Turland & L. Chilton
353
Centaurea argentea L. var. brachythysana Rech.f.
351
Centaurea argentea L. var. chionantha Turland &
L. Chilton
353
Centaurea argentea L. var. macrothysana Rech.f.
351
Centaurea argentea L. var. macrothysana Rech.f.
354
Centaurea asperula Halácsy
391
Centaurea attica Nyman subsp. drakiensis (Freyn
& Sint.) Dostál
428
Centaurea attica Nyman subsp. asperula
(Halácsy) Dostal
391
Centaurea attica Nyman subsp. ossaea (Halácsy)
Dosral
401
Celsia acaulis subsp. cyllena (Boiss. & Heldr.)
O. Kuntze
1446
Celsia acaulis Bory & Chaub.
1439
Celsia boissieri Heldr. & Sart.
1443
Celsia cyllenea Boiss. & Heldr.
1446
Celsia daenzeri Fauche & Chaub.
1447
Celsia daenzeri Fauche & Chaub. var. lyrata
Boiss. & Heldr.
1443
Centaurea attica Nyman var. pateraea Halácsy
360
Celsia speciosa Fenzl
1454
Centaurea baldaccii Degen ex Halácsy
542
Celsia tomentosa Zucc.
1467
Centaurea boissieri Walp., non DC.
358
Centaurea brevispina Hausskn.
433
Centaurea chalkensis Fors.-Maj. & Barbey
389
Centaurea cineraria Sieb.
351
Centaurea confusa Halácsy
375
Centaurea cytherea Rech.f.
419
Centaurea achaia Boiss. & Heldr. subsp.
corinthiaca (Boiss. & Heldr.) Phitos &
Georgiadis
368
Centaurea achaia Boiss. & Heldr. var. corinthiaca
(Boiss. & Heldr.) Boiss.
368
Centaurea achaia Boiss. & Heldr. var. ferox Boiss.
& Heldr.
345
Centaurea deusta Baldacci
350
Centaurea achaia Boiss. & Heldr. var. intermedia
Boiss. & Heldr.
345
Centaurea deusta Ten. var. brunnea Halácsy
362
Centaurea achaia Boiss. & Heldr. var. submutica
Boiss. & Heldr.
345
Centaurea deustiformia Adamović subsp.
pseudocadmea (Wagenitz) Dostál
410
Centaurea affinis Friv. subsp. chalcidicaea
(Hayek) T. Georgiadis
364
Centaurea deustiformia Adamović subsp.
ptarmicifolia (Halácsy & Hayek) Dostál
412
Centaurea affinis Friv. subsp. lacerata (Hausskn.)
Maire & Petitmengin
384
Centaurea dissecta var. alpina Heldr., pl. exs. A
1884, non Heldr in Boiss. Fl. Or.
401
348
Centaurea dissecta Boiss. subsp. affinis var.
parlatoris
348
Centaurea affinis Friv. subsp. peloponnesiaca
(Halácsy) Dostál
Centaurea dissecta Boiss. var. peloponnesiaca
348
Centaurea affinis Friv. var. microcaphala Halácsy
375
Centaurea drakiensis Freyn & Sint.
428
Centaurea affinis Friv. var. pallidior (Halácsy)
Halácsy
349
Centaurea epirotica (Baldacci) Halácsy
412
Centaurea affinis Friv. var. peloponnesiaca
Halácsy
348
Centaurea eriopoda Rech.f.
414
386
Centaurea alba var. princeps (Boiss. & Heldr.)
Boiss.
409
Centaurea eryngioides Lam. var. subarachnoidea
Boiss. & Heldr.
Centaurea erythracantha Halácsy
421
Centaurea alba Halácsy subsp. brunnea (Halácsy)
Dostál
362
Centaurea finazzeri sensu auct. from Taygetos.
355
Centaurea fruticosa L.
804
Centaurea alba L. subsp. albanica (Halácsy)
Dostál
350
Centaurea graeca sensu auct., Boiss. & Spruner;
Fl. Aeg., non Griseb.
358
Centaurea alba L. subsp. heldreichii (Halácsy)
Dostál
377
Centaurea grisebachii (Nyman) Heldr. subsp.
paucijuga (Halácsy) Dostál
349
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
376 K. GEORGHIOU and P. DELIPETROU
Centaurea grisebachii (Nyman) Formanek subsp.
transiens (Halácsy) T. Georgiadis
430
Centaurea parlatoris Heldr var. boissieri Heldr. &
Sart.
358
Centaurea grisebachii (Nyman) Formanek var.
confusa (Halácsy) Hayek
375
Centaurea parnonia Halácsy
355
Centaurea paucijuga Halácsy
349
Centaurea guicciardi Boiss. subsp. minoa Heldr.
ex Boiss.
423
Centaurea pelia DC. f. hamata (Hausskn.) Hayek
405
Centaurea guicciardi Boiss. var. lineariloba
Halácsy & Dörfler
387
Centaurea pelia DC. var. refracta Hausskn.
405
Centaurea pentelica Hausskn.
361
Centaurea halacsyi Dörfler
414
Centaurea raquisina Sibth. & Sm.
351
Centaurea hellenica Boiss. & Spruner
414
373
Centaurea hellenica Boiss. & Spruner f.
intermedia Heldr.
414
Centaurea rupestris L. subsp. kozanii Routsi &
T. Georgiadis
355
Centaurea iberica Trev. ex Sprengel var.
holzmanniana Boiss.
379
Centaurea rupestris L. subsp. parnonia (Halácsy)
Routsi & T. Georgiadis
Centaurea rupestris L. var. minor Boiss.
355
364
Centaurea ipsaria Stoj. & Kitanov
426
Centaurea sancta-annae J. Wagner
Centaurea lactucifolia Boiss. subsp. chalkensis
(Fors.-Maj. & Barbey) Wagenitz
389
Centaurea saxatilis (K. Koch) B.D. Jacks.
416
Centaurea scyria Runemark
417
Centaurea laureotica Heldr. ex Halácsy
391
345
Centaurea macedonica Boiss. subsp. parnonia
(Halácsy) Dostál
355
Centaurea sibthorpii (Halácsy) Boiss. & Heldr.
p.p.
Centaurea sibthorpii Halácsy p.p.
370
Centaurea macedonica Formanek
375
380
Centaurea macedonica Heldr., non Griseb.,
non C. diffusa var. brevispina Boiss.
433
Centaurea solstialis subsp. idaea (Boiss. & Heldr.)
Gugler
359
Centaurea solstialis var. idaea (Boiss. & Heldr.)
Baldacci
380
Centaurea megarensis Halácsy & Hayek
Centaurea minoa Heldr. ex Boiss.
423
387
Centaurea mixta DC.
414
Centaurea spruneri Boiss. & Heldr. subsp.
lineariloba (Halácsy & Dörfl.) Dostál
Centaurea mixta Sm. subsp. eu-mixta Gugler
414
387
Centaurea mixta Sm. var. eriopoda Rech.f.
414
Centaurea spruneri Boiss. & Heldr. var.
lineariloba (Halácsy & Dörfl.) Hay.
Centaurea mixta Sm. var. eu-mixta (Gugler)
Rech.f.
414
Centaurea spruneri Boiss. & Heldr. var. minoa
(Boiss.) Hayek
423
Centaurea mixta Sm. var. halacsyi (Dörfler)
Rech.f.
414
Centaurea spruneri Boiss. & Heldr. var. minoa
(Boiss.) Rech.f.
423
Centaurea mixta Sm. var. myconia (Boiss. &
Sart.) Hayek
414
387
Centaurea mixta Sm. var. nigrotriangulata
Rech.f.
414
Centaurea spruneri Boiss. subsp. guicciardi
(Boiss.) Hayek var. lineariloba (Halácsy &
Dörfler) Hayek
Centaurea subarachnoidea Boiss. & Heldr.
386
Centaurea myconia Boiss. & Sart.
414
Centaurea subciliaris subsp. acarnanica Matthäs
344
Centaurea nigrotriangulata Rech.f.
414
Centaurea thessala Hausskn.
429
Centaurea oliveriana DC.
356
Centaurea tragacanthoides Rech.f.
422
Centaurea oliveriana DC. var. amorgina Boiss.
356
433
Centaurea orphanidea Heldr. & Sart. var.
thessala Hausskn.
429
Centaurea tymphaea Hausskn. var. brevispina
(Hausskn.) Hayek
Centaurea ustulata Hal.
350
Centaurea ovina Formanek
388
Centaurea xanthina Boiss. & Heldr. non Spreng.
405
Centaurea pallidior Halácsy
349
Centaurium limoniiforme Greuter
437
Centaurea pallidior Halácsy var. pallidissima
Halácsy
349
Centaurium saxatile K. Koch
416
438
Centaurea paniculata sensu auct., Sibth. & Sm.
Fl. Graec. Prodr. subsp. auct. sensu Sibth. & Sm.
Fl. Graec. Prodr.
358
Centranthus longiflorus Steven subsp. junceus
(Boiss. & Heldr) I.B.K. Richardson
Centranthus nevadensis Boiss. subsp. sieberi
(Heldr.) I.B.K. Richardson
439
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 377
Cerastium tomentosum auct. non L.
449
Cerastium uniflorum var. mitkaense Wagner
462
Cerastium uniflorum auct. non Clairv.
462
443
Chaenorhinum minus L. subsp. idaeum (Rech.f.)
R. Fernandes
464
Cephalaria balearica sensu Barbey
443
Chamaecytisus subidaeus (Gand.) Rothm.
468
Cephalaria centauroides Schur var. cretacea sensu
Raulin 1878
443
Chamaemelum rosellum Boiss. & Orph.
921
Cephalaria cretacea sensu Rchb. et Schultes 1827
443
Chamaemelum tempskyanum Freyn & Sint.
922
Cephalaria setulifera Boiss. & Heldr.
442
Chamaepeuce fruticosa auct. non Cnicus
fruticosus Desf.
Cerastium alpinum var. glanduliferum sensu Stoj.
& Stefanov
462
Chartolepis lancifolia (Sprengel) Fenzl
390
Cerastium alpinum auct. non L.
462
Cheiranthus cheiri auct non L.
654
Cerastium alpinum L. var. glutinosum Halácsy
454
Cheiranthus cheiri auct. p.p., non L.
640
Cerastium atheniense Lonsing
445
Cheiranthus cheiri L. var. corinthius (Boiss.)
Hausskn.
640
Cerastium brachiatum subsp. prolixum Lonsing
451
451
Cheiranthus cheiri L. var. corinthius (Boiss.)
Hausskn.
638
Cerastium brachiatum Lonsing
Cerastium brachiatum Lonsing subsp. decrescens
Lonsing
451
Cheiranthus cheiri L., p.p.
636
Cheiranthus corinthius Boiss.
640
Cerastium brachypetalum sensu Halácsy non
Pers. s. stricto
445
Cheiranthus senoneri Helsr. & Sart.
654
Cheirantus parnassi Boiss. & Heldr.
649
Cerastium corcyrense Moschl
446
Cherleria sedoides sensu Sm. non L.
937
Cerastium coronense Runemark
456
Chinodoxa forbesii Baker var. idaea Baker
1178
Cerastium crinitum Lonsing
452
Chionodoxa cretica Boiss. & Heldr.
1178
Cerastium doerfleri Halácsy ex Hayek
447
Chionodoxa nana Schult. & Schult.f.
1178
Cerastium illyricum Ard. subsp. decrescens
(Lonsing) P.D. Sell & Whitehead
451
Chrysaspis aurantiaca (Boiss. & Spruner)
Hendrych
1420
Cerastium illyricum Ard. subsp. prolixum
(Lonsing) P.D. Sell & Whitehead
451
Chrysaspis dolopia (Gibelli & Belli) Hendrych
1422
Cerastium illyricum Ard. var. macropetalum
Boiss.
451
Chrysocyathus cylleneus (Boiss., Heldr. & Orph.)
Chrtek & Slavikova
Cerastium illyricum Bory non Ard.
451
Cineraria nebrodensis Guss. subsp. taygetea
(Boiss.) Nyman
840
Cerastium laxum Boiss. & Heldr.
455
Cirsium cylleneum Halácsy
475
Cerastium litigiosum sensu Halácsy
446
Cirsium decussatum Boiss.
474
Cerastium litigiosum sensu Halácsy
448
Cirsium dolopicum Form.
474
Cerastium moesiacum Friv. f. halacsyi Borza
454
Cirsium fruticosum auct.
1130
Cerastium moesiacum Friv. f. thasium Hayek
454
Cirsium hypopsilum auct., non Boiss. & Heldr.
477
Cerastium pedunculare sensu Halácsy [non Bory
& Chaub.]
453
Cirsium lobelii Ten. var. hypopsilum (Boiss. &
Heldr.) Halácsy
475
Cerastium pedunculare sensu Hayek p.p. [non
Bory & Chaub.]
453
Colchicum amabile auct. non Heldr.
493
484
Cerastium pedunculare sensu Hayek p.p., non
Bory & Chaub.
451
Colchicum bivonae Guss. subsp. euboeum (Orph.)
Nyman
453
Colchicum boissieri Orph. var. gracile Orph., in
sched., nom. Nud.
490
Cerastium pelligerum Bornm. & Hayek
Cerastium pilosum Sibth & Sm., nomen
453
Colchicum creticum Davis 1949; Goulimis 1956
483
Cerastium pilosum Sibth. & Sm. nom. illeg.
450
Colchicum euboeum Orph.
484
Cerastium pindigenum Lonsing
448
Colchicum kochii auct. non Parl.
493
Centranthus ruber (L.) DC. subsp. junceus (Boiss.
& Heldr) Maire & Petitmengin
438
Cephalanthera rubra sensu Teschner 1975, non
(L.) Rich.?
440
Cephalaria ?ambrosioides sensu Consp. Fl. Graec.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
1130
18
378 K. GEORGHIOU and P. DELIPETROU
Corydalis rutifolia (Sm.) DC. subsp. uniflora
(Sieber) Cullen & P.H. Davis
506
Corydalis rutifolia (Sm.) DC. var. subuniflora
Boiss. & Heldr.
506
487
Corydalis solida (L.) Clairv. subsp. longicarpa
Lidén
505
Colchicum parnassicum auct. non Sartori & al.
493
484
Corydalis tenella Ledeb. ex Nordm. var. thasia
Stoj. & Kit.
505
Colchicum parnassicum auct. non Sartori et al.
Colchicum parnassicum auct., non Sartori et al.
(1859)
485
Cotula abrotanifolia Willd.
120
Cotyledon parviflora Desf.
1435
Colchicum polymorphum Orph., nom. nud.
493
Crenularia orbiculata Boiss.
Colchicum psaridis Heldr. ex Nyman, nom. nud.
490
Crepis crocifolia Boiss. & Heldr.
Colchicum pusillum sensu Gandoger 1916;
Rechinger 1935
483
Crepis divaricata Boiss. & Heldr. subsp.
sibthorpiana (Boiss. & Heldr.) Hayek
522
Colchicum rhenium Heldr., Epet. Syllog.
Parnassou 1901: 252 (1901)
487
Crepis flexiscapa Rech.f.
523
Colchicum taygeteum Heldr. in sched. nom. nud.
485
Crepis foetida L. subsp. maritima var. tybakensis
(Vierh.) Hayek
523
Colchicum zahnii Heldr.
490
514
Colladonia colladonioides (Margot & Reuter)
Halácsy
778
Crepis fraasii Sch. Bip. subsp. mungieri (Boiss. &
Heldr.) P.D. Sell
778
Crepis fuliginosa auct. plur., non Crepis
fuliginosa Sm.
517
Colladonia heptaptera Boiss.
Colutea arborescens sensu Rechinger 1943
494
Crepis fuliginosa Sm. var. graeca (Vierh.) Greuter
517
Colymbada acicularis (Sm.) Holub.
346
Crepis mungieri Boiss. & Heldr.
514
Colymbada ebenoides (Heldr.) Holub
369
517
Colymbada guicciardii (Boiss.) Holub.
423
Crepis neglecta f. fuliginosa auct. plur., non
Crepis fuliginosa Sm.
Colymbada laconica (Boiss.) Holub
386
517
Colymbada mixta DC.
414
Crepis neglecta L. var. fuliginosa auct. plur., non
Crepis fuliginosa Sm.
Colymbada oliveriana (DC.) Holub.
356
Crepis neglecta L. var. fuliginosa sensu auct.,
Rechinger Fl. Aeg.
513
Colymbada parnonia (Halácsy) Holub
355
411
Crepis neglecta L. var. graeca sensu auct. Kret.,
Rechinger Fl. Aeg.; 1943
513
Colymbada psilacantha (Boiss. & Heldr.) Holub
Colymbada raphanina (Sm.) Holub
413
513
Colymbada sibthorpii (Halácsy) Holub
345
Crepis neglecta L. var. stricta sensu auct.,
Rechinger Fl. Aeg.
Colymbada spruneri (Boiss. & Heldr.) Holub.
423
Crepis neglecta L. var. typica sensu auct.,
Rechinger Fl. Aeg.
513
Crepis neglecta L. f. cretica (Boiss.) Vierh.
513
Crepis neglecta L. f. graeca Vierh.
517
Crepis neglecta L. subsp. cretica (Boiss.) Hay.
513
Crepis neglecta L. subsp. graeca (Vierh.) Rech.f.
517
Crepis neglecta L. var. graeca (Vierh.) Hayek
517
Crepis neglecta sensu Barb. karp.; Halácsy
Conspect. Fl. Graec. p.p.
513
Crepis raulinii Boiss.
512
Colchicum latifolium Sm. var. euboeum Boiss.
484
Colchicum lingulatum Boiss. & Spruner var.
parnassicum Sartori, Orph. & Heldr.) Stef. nom.
illegit.
488
Colchicum neapolitanum (Ten.) Ten. subsp.
parlatoris (Orph.) K. Richt
25
1103
Conium maculatum L. var. divaricatum Boiss.
495
Conosilene conica var. sartorii (Boiss & Heldr.)
A Love & Kjellqvist
1288
Conosilene conica (L.) A. Love & Kjellqvist subsp.
ammophila (Boiss. & Heldr.) A. Love & Kjellqvist
1237
Conosilene conica subsp. ammophila var.
carpathae (Chowdhuri) A. Love & Kjellqvist
1238
Consolida tomentosa sensu Rechinger 1943,
partim?
496
Conyza candida L.
826
841
Crepis sieberi. var. mungieri (Boiss. & Heldr.)
Boiss.
514
Conyza gnaphalodes (Sieber) Nyman
Conyza gnaphalodes Sieber
841
Crepis taygetea Babcock
516
Crocus atticus (Boiss. & Orph.) Orph.
535
Crocus boryanus var. graius Herb.
527
Coronilla globosa Lam.
1202
Corydalis parnassica Orph. & Heldr.
504
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 379
Crocus boryi Bory & Chaub. var. laevigatus (Bory
& Chaub.) Baker
531
Cyclamen aeginiticum Hildebr.
544
538
Cyclamen creticum Hildebr. var. pallide-roseum
Grey-Wilson. nom. inval.
543
Crocus boryi Gay subsp. tournefortii (Gay)
Greuter, Matthäs & Risse
Cyclamen gaidurowryssii Glasau
544
Crocus boryi Gay var. orphanidis (Hook.f.) Baker
538
527
Cyclamen gaydurowryssii (orthographical variant)
Glasau
544
Crocus cretensis Körnicke
Crocus fontenayi Nyman non Reuter
538
545
Crocus fontenayi Reuter
531
Cyclamen graecum Link subsp. candicum Letsw.
nom. illeg.
Crocus goulimyi Turrill subsp. leucanthus
(B. Mathew) B. Mathew
529
Cyclamen hederifolium Aiton subsp. creticum
(Hildebr.) O Schwarz
543
Crocus graecus Chapp.
528
Cyclamen miliarakesii Heldr. ex Halácsy
544
Crocus graecus Chapp. var. cartwrightianus
(Herb.) Hayek
528
Cyclamen miliarakesii Heldr. ex Hildebr.
544
Cyclamen mindleri Heldr.
545
548
Crocus hadriaticus Herb. subsp. parnassicus
(B. Mathew) B. Mathew
530
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan
547
Crocus hadriaticus Herb. subsp. parnonicus B.
Mathew
530
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan
f. albiflorum B. Mathew
530
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan
subsp. rhodense (Meikle) Kit-Tan
549
Crocus hadriaticus Herb. var. chrysovelonicus
Herb.
530
Cyclamen peloponnesiacum (Grey-Wilson) Kit Tan
subsp. vividum (Grey-Wilson) Kit Tan
550
Crocus hadriaticus Herb. var. saundersianus
Herb.
Cyclamen penteliki Hildebr.
544
Crocus laevigatus Bory & Chaub. var. fontenayi
(Reuter) Halácsy
531
Cyclamen persicum Miller subsp. mindleri
(Heldr.) Knuth
545
Crocus marathonicus Heldr.
527
Cyclamen persicum sensu Sibth. & Sm.
544
Crocus melantherus Boiss. & Orph.
525
Cyclamen pseudograecum Hildebr.
545
Crocus nivalis Bory & Chaub.
536
Cyclamen repandum Sm. f. vividum Grey-Wilson
550
Crocus orphanidis Hook.f.
538
528
Cyclamen repandum Sm. subsp. creticum (Dörfler)
Debussche & Thompson
543
Crocus pallasii Spruner ex Boiss., non Goldb.
Crocus peloponnesiacus L.
530
548
Crocus pholegandrius Orph.
538
Cyclamen repandum Sm. subsp. peloponnesiacum
Grey-Wilson
Crocus sativus L. var. cartwrightianus (Herb.)
Maw
528
Cyclamen repandum Sm. subsp. peloponnesiacum
Grey-Wilson f. albiflorum B. Mathew
547
Crocus sibthorpianus Herbert var. latifolius
Herbert
537
Cyclamen repandum Sm. subsp. rhodense (Meikle)
Grey-Wilson
549
Crocus sieberi f. nivalis (Bory & Chaub.) Heldr.
536
Cyclamen repandum Sm. var. creticum Dörfler
543
Crocus sieberi var. heterochromus Halácsy
537
Cyclamen repandum Sm. var. rhodense Meikle
549
Crocus sieberi var. versicolor Boiss. & Heldr.
537
550
Crocus sieberi Gay var. atticus Boiss. & Orph.
535
Cyclamen repandum Sm. var. vividum (GreyWilson) Kit-Tan
Crocus sieberianus Herb.
537
Cyclamen velutinum Jord.
544
Crocus stridii Papan. & Zacharof
526
Cyclaminos graeca (Link) Heldr.
544
Cyclaminos miliakiaresii Heldr.
544
Crocus veneris Maw, non Tappeiner ex Poech
531
Cyclaminus graeca (Link) Asch.
544
Crucianella pubescens Willd.
191
Cyllenium spruneri (Boiss.) Schott
238
Cruciata coronata (Sm.) Ehrend. subsp. euboea
(Ehrend.) Ehrend.
539
Cymbalaria acutiloba (Boiss. & Heldr.) Speta
subsp. dodekanesi (Greuter) Speta
552
Cucubalus cyclamineus Sieber
1293
246
Cymbalaria acutiloba (Boiss. & Heldr.) Speta
subsp. microcalyx (Greuter) Speta
552
Cucubalus polygonoides Willd.
Cuscuta epithymum var. micrantha sensu Davis
1953
540
Cymbalaria microcalyx (Boiss.) Wettst. subsp.
microcalyx var. alba Voliotis
551
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
380 K. GEORGHIOU and P. DELIPETROU
Cymbalaria microcalyx (Boiss.) Wettst. var.
heterosepala (Cuf.) Cuf.
553
Cynoglossum graecum (A.DC.) Greuter & Burdet
1146
Cynoglossum pindicum (Alden) Greuter & Burdet
1298
Dianthus juniperinus Sm. var. sieberi sensu
Rechinger (Phyton, 1943) p.p.
591
Dianthus juniperinus sensu auct. Raulin 1869
p.p., non Sm.
588
Dianthus juniperinus Sm. var. sieberi sensu auct.
non Boiss.
588
Cytisus creticus Boiss. & Heldr.
468
Cytisus dorycnioides Davidoff
469
Cytisus subidaeus Gand.
468
Dianthus juniperinus Sm. var. aciphyllus (Sieber
ex Ser.) Halácsy
586
Dactylis rigida Boiss. & Heldr.
556
Dianthus juniperinus Sm. var. sieberi Boiss.
586
Dactylorhiza cordigera (Fries) Soó subsp. graeca
(H. Baumann) Kreutz
557
Dianthus kajmaktzalanicus Micevski
594
Dactylorhiza cordigera (Fries) Soó var. graeca (H.
Baumann) H. Presser
557
Dianthus lilacinus Boiss. & Heldr. var.
androsaceus Boiss. & Heldr.
564
Dactylorhiza cordigera (Fries) Soó var. rhodopeia
H. Presser
557
Dianthus myrtinervius Griseb. var. oxylepis Boiss.
594
Dianthus oxylepis (Boiss.) Kummerle & Jav.
594
Dactylorhiza kalopissii E. Nelson subsp.
pythagorae (Gölz & Reinchard) E. Nelson
558
Dianthus pruinosus Boiss. & Orph.
583
Dianthus pubescens Sm.
571
Delphinium junceum auct. non DC.
562
Dianthus pubescens Sm. var. glabratus Boiss.
571
Delphinium longipes sensu Hayek, non Moris
562
Dianthus pulviniformis Greuter
592
Delphinium peregrinum sensu Halácsy, non L.
562
Dianthus rhodius Rech.f.
579
Delphinium tenuissimum Sm.
498
Dianthus samaritani Heldr.
565
Delphinium tuntasianum Halácsy
499
596
Dianthus aciphyllus var. bauhinorum Greuter
587
Dianthus serratifolius Sm. var. abbreviatus
Heldr. & Halácsy
Dianthus aciphyllus Ser. var. aciphyllus
586
Dianthus sibthorpii Vierh.
584
Dianthus alpinus Sibth. sensu Smith (1809)
584
Dianthus taygeteus Quezel & Contandr.
596
Dianthus arboreus auct. plur.
572
Dianthus ventricosus Heldr. ex Halácsy
584
Dianthus arboreus L., nomen ambig.
586
Dianthus viscidus Bory & Chaub. var.
tymphresteus Boiss. & Spruner
601
Dianthus arboreus L., non Sm. et auct. plur.,
nomen ambiguum
587
Dianthus xanthianus Davidov
581
Dianthus arboreus sensu auct. p.p., non s. str.
585
Digitalis lanata Ehrh. subsp. leucophaea (Sibth.
& Sm.) H. Werner
606
Dianthus biflorus Sm. subsp. samaritani (Heldr.)
Maire & Pet.
565
Diosphaera asperuloides (Boiss. & Orph.) Buser
278
Dianthus chalcidicus Halácsy
569
Diosphaera jacquinii (Sieber) Buser
298
Dianthus cinnabarinus Spruner.
565
Draba aizoides L. subsp. lacaitae (Boiss.) Maire &
Petitmengin
608
Dianthus cinnabarinus Spruner. var. alpinus
Boiss.
565
Draba aizoides L. var. brevistyla Boiss. nom. illeg.
608
Dianthus creticus Tausch
575
Draba boissieri Baldacci
610
Dianthus cylleneus Boiss. & Heldr.
571
Draba cretica Boiss. & Heldr. var. lassitica
Halácsy
607
Dianthus diffusus Sm. var. cylleneus (Boiss. &
Heldr.) Halácsy
571
Draba cretica Boiss. & Heldr. var. sphaciotica
Halácsy
607
Dianthus glandulosopubescens Halácsy
571
Draba erostra Halácsy
608
610
Dianthus glumaceus Bory & Chaub.
1093
Dianthus haematocalyx Boiss. & Heldr. subsp.
sibthorpii (Vieh.) Hayek
584
Draba hispanica Boiss. subsp. parnassica (Boiss.
& Heldr.) Nyman
584
Echinops ritro L. var. sartorianus (Boiss. &
Heldr.) Boiss.
619
Dianthus haematocalyx Boiss. & Heldr. var.
alpinus Boiss.
Echinops sartorianus Boiss. & Heldr.
619
Dianthus haematocalyx Boiss. & Heldr. var.
pruinosus (Boiss. & Orph.) Boiss.
583
Echinops sphaerocephalus L. var. taygeteus
(Boiss. & Heldr.) Boiss.
620
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 381
661
810
Euphorbia myrsinites sensu auct. kret. Rechinger
1943
Epipactis atrorubens (Hoffm.) Besser subsp.
subclausa (Robatsch) Kreutz
626
Euphorbia myrsinites sensu Rechinger 1944
662
Evax cretensis (Gand.) Gand.
686
Epipactis atrorubens auct. non (Hoffm. ex Bernh.)
Besser
626
Farsetia lunaroides (Willd.) R.Br.
683
Epipactis cucullata (Boiss. & Heldr. ex Rchb.f.)
Wettstein
440
Ferulago serpentinica Rech.f.
664
Festuca callieri sensu Krause 1956
677
Epipactis graeca Halácsy nom nud.
624
670
Epipactis helleborine sensu auct. cret. non (L.)
Crantz, Künkele 1979?
623
Festuca hirtovaginata (Acht.) Markgr.-Dann.
(included in taxon)
676
Epipactis purpurata auct. non J.E. Smith
624
Festuca polita (Halácsy) Tzvelev var. cretica
Markgr.-Dann.
Epipactis thessala B. & H. Baumann
626
Festuca thracica Markgr.-Dann. (included in
taxon)
670
Eriolepis morinaefolia Gandoger
477
Fibigia lunaroides (Willd.) Sm. (sphalm.)
683
Erodium absinthoides Willd. subsp. chrysanthum
(L’Herit. ex DC.) Maire & Pettimengin
628
Filago spathulata C.Presl var. syria Hal.
685
Erodium vetteri W. Barbey & Major
630
Foxia spicata (Boiss.) Parl.
231
Erysimum aciphyllum sensu Boiss Fl. Orient.;
Meikle 1954 Chios
643
Freyera cretica (Boiss. & Heldr.) Boiss. & Heldr.
755
Freyera divaricata Boiss. & Orph.
756
640
Freyera parnassica Boiss. & Heldr.
758
Erysimum boryanum Boiss. & Spruner subsp.
parnassi (Boiss. & Heldr.) Nyman
649
Freyera pumila (Sm.) Boiss.
760
640
Fritillaria erhartii Boiss. & Orph. var. octavii
Gand.
691
Erysimum cheiri (L.) Crantz var. corinthium
(Boiss.) Polatschek
634
Fritillaria graeca var. thessala auct., non (Boiss.)
Rix
697
Erysimum parnassi Bory & Chaub var. atticum
(Heldr. & Sart. ex Boiss.) Hayek
Fritillaria graeca var. thessala auct., non Boiss.
697
Erysimum pusillum Bory & Chaub subsp. atticum
(Heldr. & Sart. ex Boiss.) P.W. Ball
634
Fritillaria graeca auct. Fl. Graec.
696
Erysimum pusillum Bory & Chaub var. atticum
(Heldr. & Sart. ex Boiss.) Hayek
634
Fritillaria graeca Boiss. & Spruner subsp. graeca
var. graeca
693
Erysimum pusillum Bory & Chaub. subsp.
cephalonicum (Polatschek) P.W. Ball
639
Fritillaria graeca Boiss. & Spruner subsp.
guicciardii (Heldr. & Sart.) Zaharof
695
Erysimum pusillum Bory & Chaub. subsp.
hayekii Jav. & Rech.f.
643
Fritillaria graeca Boiss. & Spruner subsp.
thessala (Boiss.) Rix
707
Erysimum pusillum Bory & Chaub. subsp.
microstylum (Hausskn.) Hayek
645
Fritillaria graeca Boiss. & Spruner var. thessala
Boiss.
707
Erysimum pusillum Bory & Chaub. subsp.
parnassi (Boiss. & Heldr.) Hayek
649
Fritillaria graeca Boiss. & Spruner var. unicolor
Halácsy
694
Erysimum pusillum Bory & Chaub. subsp.
rechingeri
653
Fritillaria graeca Wahlenb var. ionica (Halácsy)
Zaharof var.
708
Erysimum pusillum Bory & Chaub. subsp.
trichophyllum (Halácsy) Hayek
651
Fritillaria guicciardii Heldr. & Sart var. zahnii
Heldr.
694
Erysimum smyrnaeum sensu auct. Samothraki,
non Boiss. & Balansa
644
Fritillaria guicciardii Heldr. & Sart.
695
Fritillaria ionica Halácsy p.p.
708
Erysimum trichophyllum Halácsy
651
Fritillaria ionica Halácsy p.p.
709
Echinops taygeteus Boiss. & Heldr.
620
Elodes acifera Greuter
Erysimum bisaccatum Form.
Fritillaria macrandra Baker
691
Euphorbia bivonae sensu Turland 1989, non
Steud.
662
Fritillaria pontica sensu auct. fl. graec. non
Wahlenb.
706
Euphorbia myrsinites L. subsp. rechingeri
(Greuter) Alden
661
Fritillaria pontica Wahlenb var. ionica (Halácsy)
Turrill
708
Eunomia orbiculata Griseb.
25
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
382 K. GEORGHIOU and P. DELIPETROU
Fritillaria pontica Wahlenb. var. substipelata P.C.
Candargy
706
Gypsophila nana Bory & Chaub. var. grabrifolia
Halácsy
Fritillaria regis georgii Heldr. & Holzm. ex
Nyman
691
Gypsophila ocellata Sm.
Fritillaria rixii Zaharof
692
Fritillaria sibthorpiana auct. balkan., non (Sm.)
Baker
692
Fritillaria sphaciotica Gand.
696
Fritillaria thessalica Spruner
707
Fritillaria tuntasia Heldr. ex Halácsy
700
Fumana procumbens sensu Baldacci 1916
711
Fumana vulgaris f. alpina Janchen
711
Fumaria uniflora Sieber
506
Galanthus latifolius Rupr.
712
Galium amorginum sensu Greuter 1973, non
Halácsy 1901
731
Gypsophila ochroleuca Sm.
767
246
1097
Gypsophila polygonoides subsp. ocellata (Sm.)
Hayek
246
Gypsophila polygonoides (Willd.) Halácsy
246
Gypsophila polygonoides (Willd.) Halácsy subsp.
thessala (Jaub. & Spach) Hayek
249
Gypsophila polygonoides (Willd.) Halácsy subsp.
thymifolia (Sm.) Hayek
249
Gypsophila polygonoides (Willd.) Halácsy var.
thymifolia (Sm.) Hayek
249
Gypsophila thessala (Jaub. & Spach) Halácsy
249
Gypsophila thymifolia Sm.
249
Haberlea heldreichii Boiss.
842
Halαcsyella parnassica (Boiss. & Spruner)
Janchen
621
Helianthemum ericoides sensu Gandoger 1916
711
Helichrysum virgineum (Sm.) Griseb.
773
Helictotrichon cycladum (Rechf. & Scheffer)
Rech.f.
775
Galium aureum Vis. var. oblongifolium Boiss.
721
Galium boryanum Walp.
173
Galium capitatum Boiss. sensu auct. Kret.
730
Galium capitatum Boiss. var. hispidulum Raulin
730
Galium coronatum Sm. var. euboeum Ehrend.
539
Galium firmum Tausch var. citraceum (Boiss.)
Boiss. & Heldr.
721
Heliosperma chromodontum (Boiss. & Reuter)
Juratzka
1284
Galium firmum Tausch var. euboeum Tocl &
Rochl.
721
Heliotropium bocconei sensu Hayek p.p., non
Guss.
777
Galium incanum Sm. subsp. cyllenium (Boiss. &
Heldr.) Maire & Petitmengin
723
Helladia cretica (C.Presl.) Kral.
1204
Galium suberosum sensu auct. cret., non Sm.
193
Helladia tymphaea (Quézel & Contandr.) M. Kral
1127
Genista salzamanii subsp. parnassica (Halácsy)
Ponert
752
Herniaria nigrimontium Hermann var. degenii
Hermann
779
Genista subsericans (Bornm.) Rech.f.
752
Herniaria parnassica Boiss. var. hirtella
Chaudhri
780
Geocaryum cynapioides (Guss.) P.W. Ball subsp.
divaricatum (Boiss. & Orph.) P.W. Ball
756
Hesperis glutinosa Vis. subsp. secundiflora
(Boiss.) F. Dvořák
781
Githago thessala Formanek
29
762
Hesperis glutinosa Vis. var. secundiflora (Boiss. &
Spruner) Borbas
781
Globularia cordifolia subsp. stygia (Orph.) Wettst.
Gymnospermium altaicum (Pall.) Spach subsp.
odessanum sensu auct. Grec.
766
Hesperis rechingeri F. Dvořák
782
Hesperis secundiflora Boiss & Spruner
781
Gypsophila achaia Bornm. (included in taxon)
767
Hieracioides athoum (Boiss.) O. Kuntze
511
512
Gypsophila cretica sensu Smith (1809)
1090
Hieracioides auriculaefolium (Sieber.) O. Kuntze
Gypsophila dianthoides Sm.
1091
Gypsophila fasciculata Margot & Reuter
1092
Hieracioides crocifolium (Boiss. & Heldr.) O.
Kuntze
Gypsophila fasciculata Margot & Reuter var.
laconica Boiss.
248
Gypsophila fruticulosa (Bory & Chaub.) Boiss.
245
Gypsophila graeca (Schreber) Britten
246
Gypsophila graminea Sm.
Gypsophila laconica (Boiss.) Boiss.
1094
248
1103
Hieracioides heldreichianum O. Kuntze
516
Hieracioides incanum (Sm.) O. Kuntze
520
Hieracioides sibthorpianum (Boiss. & Heldr.) O.
Kuntze
522
Hieracium foetidum Sm.
522
Hieracium graecum f. flocciceps Zahn
791
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 383
Hieracium graecum Boiss. & Heldr. subsp.
szilyanum J. Wagner & Zahn
801
Hieracium leithneri (Heldr. & Sart. ex Boiss.)
Zahn var. eurytanicum Zahn
794
Hieracium pannosum Boiss. subsp. gaudryi
(Boiss. & Orph.) Nyman
789
Hieracium sartorianum (Boiss. & Heldr.) var.
leithneri Heldr. & Sart. ex Boiss.
794
Hieracium sartorianum Boiss. & Heldr. subsp.
naegelianoides Zahn
Iberis spathulata sensu Bald. 1899
824
Inula candida subsp. limonella (Heldr.) Rech.f.
827
Inula candida subsp. methanea (Hausskn.) Hayek
831
Inula candida var. p.p. Halácsy Fl. Graec.
826
Inula candida var. rotundifolia Halácsy
829
Inula candida (L.) Cass subsp. limoniifolia Hayek
826
Inula candida (L.) Cass. subsp. candida
826
799
Inula candida (L.) Cass. subsp. decalvans
(Halácsy) P.W. Ball
828
Hieracium sartorianum sensu auct. Rchb.f.
(1860), Boiss. (1875); Consp. Fl.. Graec; Zahn
1921 p.p.
794
Inula limonifolia (Sm.) Boiss. var. decalvans
Halácsy
828
790
Inula limonifolia (Sm.) Boiss. var.
pseudolimonella Rech.f.
828
Hieracium sartorianum sensu Boiss. 1875;
Halácsy 1902 p.p.
Inula limoniifolia subsp. rotundifolia
826
Hieracium sericophyllum Nejceff & Zahn subsp.
eriocomum Zahn
795
Inula limoniifolia (Sm.) Boiss. ex Raulin
826
Hieracium sericophyllum Nejceff & Zahn subsp.
fiedleri Zahn
795
Inula methanea Hausskn.
831
Inula parnassica Boiss. & Heldr.
832
Hieracium sericophyllum Nejceff & Zahn subsp.
olenium Zahn
794
Inula sophiae Beauverd
827
Inula sophiae Beauverd var. sporadum Phitos
827
Hieracium wiesbaurianum Uechtr. ex Baenitz
subsp. epirense Zahn
787
Iris cretensis Janka
835
Iris cretica Herbert ex Baker nomen nudum
835
Hieracium wiesbaurianum Uechtr. ex Baenitz
subsp. kyllenense Zahn
787
Iris cretica Herbert ex Baker nomen nudum
834
Hieracium wiesbaurianum Uechtr. ex Baenitz
subsp. retroversidens Zahn
787
Iris humilis Sieber ex Baker
835
Iris stylosa Desf. var. angustifolia Boiss. & Heldr.
835
834
Huetia cretica (Boiss. & Heldr.) P.W. Ball
755
Iris stylosa Mazziari
Huetia cynapioides (Guss.) P.W. Ball subsp.
divaricata (Boiss. & Orph.) P.W. Ball
756
Isatis athoa Boiss.
837
Isatis tinctoria L. var. stenocarpa Griseb.
837
Huetia cynapioides (Guss.) P.W. Ball subsp.
macrocarpa (Boiss. & Spruner) P.W. Ball p.p.
758
Ischarum fraasianum Schott
237
Ischarum spruneri (Boiss.) Schott
238
Huetia pumila (Sm.) Boiss.
760
Ixoca chromodonta (Boiss. & Reuter) Sojak
1284
Hutera nivalis (Boiss. & Heldr.) Gomez-Campo
479
Ixoca phyllitica (Neumayer) Ikonn
1284
1178
Johrenia graeca Boiss. & Spruner
843
Hyacinthus nanus Schult. & Schult.f.
Hyacinthus spicatus Sm.
231
Kentrophyllum leucocaulon DC.
Hyalea oliveriana Jaub. & Spach
356
Kohlrauschia glumacea (Bory & Chaub.) Hayek
Hypericum empetrifolium Willd. var. oliganthum
Rech. f.(including taxon)
816
Koniga emarginata (Boiss.) Nyman
852
Hypericum empetrifolium Willd. var. tortuosum
Rech. f.
817
Lactuca viminea J. Presl & C. Presl subsp.
alpestris (Gand.) Ferakova
845
Hypericum hyssopifolium Chaix subsp.
tymphresteum (Boiss. & Spruner) Hayek
815
Lactuca viminea J. Presl & C. Presl var.
decumbens Halácsy
845
Hypericum montanum L. var. athoum Griseb.
812
Lamium garganicum L. subsp. striatum (Sm.)
Hayek var. pictum (Boiss.) Hayek
846
Hypericum repens Sm.
822
Lamium pictum Boiss. & Orph.
846
Hypericum sanctum Degen
812
Lamium striatum Sm. var. minus Boiss.
846
Hypericum tymphresteum Boiss. & Spruner
815
Leontice odessana (DC.) G. Don
766
Hypochoeris tenuiflora (Boiss.) Boiss.
823
825
Leontodon crispus Vill. subsp. graecus (Boiss. &
Heldr) Hayek
850
Iberis arbuscula Runemark
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
337
1093
384 K. GEORGHIOU and P. DELIPETROU
Leopoldia cycladica (Davis & Stuart) Bentzer
947
Malcolmia cymbalaria Boiss.
918
Leopoldia cycladica (Davis & Stuart) Bentzer
subsp. cycladica
948
Malcolmia graeca Boiss. & Spruner var. tenuior
Hausskn.
914
Leopoldia cycladica (Davis & Stuart) Garbari
subsp. subsessilis Bentzer
949
Malcolmia maritima (L.) R. Br. var. macrocalyx
(Halácsy) Hayek
915
Leopoldia dionysica (Rech.f.) Greuter
950
Malcolmia scyria Rech.f.
917
Leopoldia spreitzenhoferi Heldr.
955
Malcolmia veluchensis Boiss. & Heldr.
913
Limodorum abortivum (L.) Swartz subsp. gracile
B. Willing & E. Willing
855
Marrubium haussknechtii Hausskn.
920
Limodorum abortivum (L.) Swartz var. gracile
(B. Willing & E. Willing) Kreutz
855
Marrubium velutinum Sm. subsp. cylleneum
(Boiss. & Heldr.) Nyman
919
Limonium melium (Nyman) Pignatti p.p.
867
Marrubium velutinum Sm. var. cylleneum (Boiss.
& Heldr.) Boiss.
919
Limonium ocymifolium sensu Greuter &
Rechinger 1967; Yannitsaros 1969 Kythira
886
Mattia graeca A. DC.
Limonium pigadiense (Rech.f.) Rech.f.
881
Melampyrum barbatum Waldst. & Kit. ex Willd.
subsp. ciliatum (Boiss. & Heldr.) Nyman
924
Limonium pigadiense sensu Christod. et al.
Gramvousa
886
Melandrium auriculatum (Sibth. & Sm.) Rohrb.
1240
Limonium ramosissimum (Poiret) Maire subsp.
doerfleri (Halácsy) Pignatti
871
Melissa cretica L.
1158
Linaria microcalyx (Boiss.) Wettst. var.
heterosepala Cuf.
553
Linaria microcalyx (Boiss.) Wettst. var. minor
Cuf.
1146
Mentha erinoides Heldr.
929
Mentha gryparia Heldr. f. erinoides Heldr.
929
Mentha pulegium L. var. erinoides (Heldr.) Hayek
929
554
Mentha pulegium L. var. subtomentella f.
cephalonia H. Braun
928
Linaria microcalyx (Boiss.) Wettst. var.
orphanidiana Cuf.
553
Micromeria acropolitana Halácsy
1157
Micromeria calostachya Rech.f.
1406
Linum iberidifolium Boiss. (nom. inval.)
897
Micromeria carpatha Rech.f.
931
Linum pycnophyllum Boiss. & Heldr.
899
21
Micromeria microphylla sensu auct. karp., kret.,
Consp. Fl. Graec.; Fl. Aeg.; Boiss. Fl. Or.
931
Lithospermum goulandriorum Rech.f.
Lithospermum goulandriorum Rech.f. subsp.
goulandriorum
22
Microrrhinum idaeum (Rech.f.) Speta
464
Minuartia favargeri Iatroú & T. Georgiadis
943
Lithospermum goulandriorum Rech.f. subsp.
thessalicum Aldén
23
Minuartia greuteriana Kamari var. pessana
941
900
Minuartia hirsuta (M. Bieb.) Hand.-Mazz. subsp.
eurytanica (Boiss. & Heldr.) Strid
939
Lonicera hellenica Orph. ex Boiss.
908
Minuartia idaea (Halácsy) Pawlsowski
936
Lotus wiedemannii (Boiss.) Hayek
1388
Minuartia setacea (Thuill.) Hayek var. athoa
(Griseb.) Mattf.
933
Minuartia stellata (E. D. Clarke) Maire &
Petitmegin subsp. pseudosaxifraga Mattf.
944
Minuartia trichocalycina sensu auct., non
Arenaria trichocalycina Ten. & Guss.
937
Lithospermum zahnii Heldr. ex Halácsy
Lutzia fruticosa Gand.
910
Lychnis variegata Desf.
1293
Lycopsis aegyptiaca auct non L., Gand
116
Lycopsis foliis drepano-dentatis L.
116
Lycopsis variegata L.
116
Lyrolepis diae Rech f.
335
Minuartia verna (L.) Hiern subsp. idaea (Halácsy)
Hayek
936
Lyrolepis piae Norderstam
335
Minuartia verna (L.) Hiern var. acutipetala Boiss.
936
Lysimachia anagalloides Sm. (from Kriti)
911
Minuartia wettsteinii Mattf. subsp. parnonia
Kamari
942
1064
Malcolmia angulifolia Boiss. & Orph.
918
Minuartia wettsteinii Mattf. subsp. wettsteinii
946
Malcolmia bicolor Boiss. & Heldr.
913
Moehringia stricta Sm.
258
Malcolmia bicolor Boiss. & Heldr. var.
veluchensis (Boiss. & Heldr.) Boiss.
913
Muscari amoenocomum Rech.f. (included in
taxon)
955
Majorana microphylla Benth.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 385
968
955
Nigella arvensis L. var. glauca Boiss. sensu Hayek
Prodr. Flor. pen. Balc.; non sensu orig. Boissier
Fl. Or.
Muscari macrocarpum sensu auct. Siteia, non
Sweet
232
Nigella arvensis L. var. glaucescens sensu
Rechinger 1943
969
Muscari maritimum sensu auct. cret.
955
Nigella arvensis L. var. involucrata Boiss. p.p.
968
Muscari pulchellum Heldr. & Sart. var. amoenum
Heldr. ex Halácsy
953
Nigella arvensis L. var. microcarpa Boiss.
[sphalm.macrocarpa] p.p.
969
Muscari racemosum subsp. pulchellum (Heldr. &
Sart.) Stojanov & Stefanof
952
Nigella cretensis sensu Greuter & Rechinger, 1967
p.p.; non sensu orig. Steven
976
Muscari weissii sensu auct. cret. [Rechinger 1944,
Bentzer 1973]?
955
Nigella cretensis sensu Greuter & Rechinger, 1967
p.p.; non sensu orig. Steven, 1856
978
Nepeta dirphya (Boiss.) Heldr. ex Halácsy
958
Nigella cretensis Steven
969
Nepeta heldreichii Halácsy
962
Nigella cretica auct. p.p. non Miller
976
Nepeta laconia Orphanides, nom. nud.
964
969
Nepeta melissaefolia Lam. (sphalm.)
963
Nigella cretica sensu Halácsy Consp. Fl. Graec.;
Hayek Prodr. Fl. pen. Balk.; Rechinger Fl. Aeg.;
non sensu orig. Miller
Nepeta orientalis Miller var. parnassica (Heldr. &
Sart) Baldacci
965
Nigella degenii sensu Tutin & al. Fl. Eur. 1967
p.p.; Rechinger Fl. Aeg. p.p., non Vierh.
976
Nepeta parnassica Heldr. & Sart. (β) orphanidea
964
970
Nepeta sibthorpii Benth. subsp. dirphya (Boiss.)
Baden
958
Nigella dejenii sensu Davis Fl. Turk p.p.; Tutin
Fl. Eur.p.p.; non Vierh.
974
Nepeta sibthorpii Benth. subsp. malacotrichos
Baden
959
Nigella divaricata Beaurpre ex DC. sensu
Gandoger, 1920; non sensu orig. De Candolle
968
Nepeta sibthorpii Benth. subsp. parnassica
(Heldr. & Sart.) Maire & Petitmengin
965
Nigella huthii Brand. var. aristata (Sm.) Brand,
illeg. comb.
Nepeta sibthorpii Benth. subsp. sibthorpii
Nepeta sibthorpii Benth. var. dirphya Boiss.
Muscari creticum Vierh.
955
Muscari cycladicum sensu auct. cret. [Bentzer
1973]?
Noccaea cretica (Degen & Javorka) F.K. Mey.
1402
957
Noccaea epirota (Halácsy) F.K. Mey.
1403
958
Noccaea graeca (A. Jordan) F.K. Mey
1404
Nigella aristata sensu Halácsy Consp. Fl. Graec.
p.p.; Hayek Prod. Fl. pen. Balk. p.p.; Rechinger
Fl. Aeg. p.p.;. non Sibth. & Sm.
974
Noccaea zaffranii F.K. Mey.
1405
Nonea auriculata Boiss.
1131
1131
Nigella aristata sensu Halácsy Consp. Fl. Graec.
p.p.; Hayek Prod. Fl. pen. Balk. p.p.; Rechinger
Fl. Aeg. p.p.;. non Sibth. & Sm.
971
Nonea cesatiana (Fenzl & Friedr.) Greuter &
Burdet
Oberna ionica (Halácsy) Ikonn.
1264
Nigella aristata Sm.
968
Oberna variegata (Desf.) J. Holub
1293
Nigella aristata subsp. rechingeri Tunin
968
Nigella arvensis var. involucrata Boiss. sensu
Forsyth Major & Barbey, 1897; non sensu orig.
Boissier Fl. Or.
977
Nigella arvensis L. a typica Boiss. sensu
Haussknecht, 1892; non sensu orig. Boissier, Fl.
Or.
968
Nigella arvensis L. subsp. cretica (Miller) Ciferri
& Ciacomini
Odontarrhena lesbiaca P. Candargy
97
Odontites bocconei (Guss.) Walpers subsp. linkii
(Heldr. & Sart.) Maire & Petitmengin
979
Odontites cretica Boiss. (included in taxon)
979
Odontites frutescens Halácsy
979
Odontites linkii Heldr. & Sart. ex Boiss. subsp.
creticus (Boiss.) Greuter (included in taxon)
979
969
Onobrychis aliacmonia Rech.f. subsp.
peloponnesiaca Iatroú & Kit Tan
984
Nigella arvensis L. subsp. foeniculacea (DC.)
Bornm. sensu Bornueller, 1928; non sensu orig.
De Candolle, 1818
968
Onobrychis carnea Boiss. & Heldr.
982
Onobrychis ebenoides Boiss. & Spruner var.
minor Boiss.
982
Nigella arvensis L. subsp. involucrata (Boiss.)
Terracc. p.p.
968
Onobrychis hybrida Halácsy
982
968
Onobrychis montana DC. subsp. cadmea (Boiss.)
P.W. Ball
983
Nigella arvensis L. subsp. rechingeri (Tutin) Tutin
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
386 K. GEORGHIOU and P. DELIPETROU
Onobrychis scardica var. brevicaulis Halácsy
983
Ophrys bilunulata auct. cret. non Risso
1025
Ononis antiquorum L. subsp. diacantha (Rchb.)
Greuter
986
Ophrys calocaerina Devilliers-Tersch. & Devilliers
1024
986
Ophrys cornuta Steven ex M. Bieb. subsp.
heldreichii (Schltr.) Renz.
1035
Ononis diacantha Rchb.
Ononis repens L. subsp. diacantha (Rchb.)
Greuter
986
Ophrys creberrima Paulus
1025
Ophrys cressa Paulus
1026
Ononis spinosa L. var. diacantha (Rchb.) Sirj.
986
Ophrys ‘cressa-holoserica’ Paulus nom. nud.
1039
Onopordon majorii Beauverd (sphalm.)
993
Ophrys cretensis (H. Baumann & Künkele) Paulus
1048
Onopordum myriacanthum Boiss.
990
1021
Onopordum tauricum Willd. var. elatum auct.
cyth.
992
Ophrys cretica (Vierh) E. Nelson subsp.
karpathensis E. Nelson
1021
Onopordum tauricum Willd. var. mazzocchianum
Pamp.
995
Ophrys cretica (Vierh) E. Nelson subsp. naxia
E. Nelson?
Ophrys cretica (Vierh.) E. Nelson subsp. beloniae
G. Kretzschmar & H. Kretzschmar
1022
Ophrys cretica (Vierh.) E. Nelson subsp. bicornuta
H. Kretzschmar & R.L. Jahn
1022
Ophrys creticola Paulus
1027
Ophrys dodekanensis H. Kretzschmar & Kreutz
1043
Ophrys doerfleri auct., non H. Fleischm.?
1020
Ophrys doerfleri H. Fleischm.
1021
Ophrys eptapigiensis Paulus
1028
Ophrys ferrum-equinum Desf. subsp. argolica
(H. Fleischm.) Soó
1015
Onosma aucheriana DC. subsp. pallida (Boiss.)
Hayek var. paradoxa (Janka) Hayek
1005
Onosma erecta Sm. var. pubiflora Halácsy
1002
Onosma laconica Boiss. & Orph.
999
Onosma orphanidis Boiss.
999
Ophrys ‘albifrons-cretica’
1021
Ophrys ‘bicolor-fusca’ Paulus nom. prov.
1032
Ophrys ‘creberrima-fusca’ Paulus
1025
Ophrys ‘Jouchtas-fusca’
1027
Ophrys ‘Kreta-fusca, gross, früh’ Paulus nom.
prov.
1027
Ophrys ferrum-equinum Desf. subsp. spruneri
(Nyman) Camus
1052
Ophrys ‘merula-fusca’ Paulus nom. prov.
1026
1032
Ophrys ferrum-equinum Desf. var. gottfriediana
(Renz) Biel
1023
Ophrys ‘Thripti-fusca früh’ Paulus nom. prov.
Ophrys ‘Thripti-fusca spät’ Paulus nom. prov.
1026
Ophrys fleischmannii Hayek
1047
Ophrys ‘tuberculata-cretica’
1020
1011
Ophrys aegaea Kalteisen & H.R. Reinhard
1014
Ophrys fuciflora (F.W. Schmidt) Moench subsp.
andria (P. Delforge) Faurh.
Ophrys aeoli P. Delforge
1037
1045
Ophrys aesculapii Renz subsp. pseudoaranifera
Renz
1010
Ophrys fuciflorra (F.W. Schmidt) Moench f.
pseudoestrifera Renz
Ophrys funerea auct. non Viviani
1031
Ophrys andria P. Delforge var. halkionis G.
Kretzschmar & H. Kretzschmar
1013
Ophrys funerea Viviani subsp. fleischmannii Soó
1047
1026
Ophrys araneola Rchb. subsp. cretensis
(H. Baumann & Künkele) Kreutz
1048
Ophrys fusca Link subsp. cressa (Paulus) H.
Kretzschmar
1028
Ophrys araneola sensu auct. cret., non Rchb. p.p.
1048
Ophrys fusca Link subsp. eptapigiensis (Paulus)
Kreutz
Ophrys aranifera Huds. subsp. x macedonica
H. Fleischmann ex Soó
1041
Ophrys fusca Link subsp. fleischmannii (Soó) Soó
1047
1032
Ophrys aranifera Huds. var. macedonica H.
Fleischmann, nom. nud.
1041
Ophrys fusca Link subsp. thriptiensis (Paulus)
H. Kretzschmar
Ophrys gottfriediana Renz
1023
Ophrys argolica H. Fleischm. subsp. argolica
1015
1051
Ophrys argolica sensu auct. karp., non H.
Fleischm
1014
Ophrys grigoriana G. Kretzschmar & H.
Kretzschmar
Ophrys halia Paulus
1038
Ophrys ariadnae Paulus, nom. inval.
1021
Ophrys heldreichii Schlechter subsp.
schlechteriana (Soó) Kreutz
1045
Ophrys astypalaeica P. Delforge
1040
1046
Ophrys heldreichii Schlechter var. schlechteriana
(Soó) Soó
1045
Ophrys basilissa C. Alibertis & A. Alibertis
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 387
Ophrys sphegodes Mill. subsp. grammica (B.
Willing & E. Willing) Kreutz
1034
1011
Ophrys sphegodes Mill. subsp. spruneri (Nyman)
E. Nelson nom. inval.
1052
Ophrys hystera Kreutz & R. Peter
1042
1049
Ophrys icariencis M. Hirth & H. Spaeth
1016
Ophrys sphegodes Mill. subsp. zeusii (M. Hirth)
Kreutz
Ophrys icariensis M. Hirth & H. Spaeth
1017
Ophrys spruneri Nyman f. cretica Vierh.
1022
Ophrys karpathensis (E. Nelson) Paulus & Gack,
nom. inval.
1021
Ophrys spruneri Nyman subsp. cretica (Vierh.)
Renz
1022
Ophrys karpatica A. Alibertis nom. nud.
1039
Ophrys spruneri Nyman subsp. gottfriediana
(Renz) Soó
1023
Ophrys kotschyi H. Fleischm. & Soó subsp.
ariadnae (Paulus) Faurh.
1021
Ophrys spruneri Nyman subsp. spruneri
1052
Ophrys kotschyi H. Fleischm. & Soó subsp. cretica
H. Sundermann.
1022
Ophrys thesei P. Delforge
1013
Ophrys thriptiensis Paulus
1032
1049
Ophrys helios Kreutz
1039
Ophrys hiulka Spruner ex Rchb.f.
1052
Ophrys holoserica (Burm.f.) Greuter subsp. andria
(P. Delforge) Faurh.
Ophrys lacaena P. Delforge
1019
Ophrys zeusii M. Hirth
Ophrys leptomera P. Delforge
1044
Orchis anatolica Boiss. subsp. sitiaca Renz
1060
Ophrys mammosa Desf. subsp. grammica B.
Willing & E. Willing
1034
Orchis anatolica Boiss. var. sitiaca (Renz) Soó
1060
Orchis morio L. subsp. boryi (Rchb.f.) Soó
1056
Ophrys mammosa Desf. subsp. macedonica
(H. Fleischmann ex Soó) Soó nom. inval.
1041
Orchis papilionacea L. var. alibertis G.
Kretzschmar & H. Kretzschmar
1057
Ophrys mammosa Desf. subsp. oodcheila (Renz)
A. Riechelman
1034
Orchis patens Desf. subsp. falcicalcarata
Wildhaber
1059
Ophrys minoa sensu P. Delforge p.p.
1019
Orchis patens Desf. subsp. nitidifolia Teschner
1059
Ophrys oestrifera Bieb. subsp. heldreichii (Schltr.)
E. Nelson
1035
Orchis patens Desf. subsp. orientalis sensu
Rechinger 1943
1059
Ophrys omegaifera H. Fleischm. subsp.
omegaifera var. basilissa (C. Alibertis, A. Alibertis
& H.R. Reinhard) Faurh.
1046
Orchis spitzelii Sauter ex Koch subsp. nitidifolia
(Teschner) Soó
1059
Orchis spitzelii sensu Seitz 1978, Henke 1983
1059
Ophrys omegaifera H. Fleischm. var.
fleischmannii (Hayek) Soó
1047
Oreoherzogia guicciardi (Boiss.) Halácsy
1140
Ophrys parosica P. Delforge
1029
Oreoherzogia pubescens (Sm.) Vent
1141
Ophrys parvula Paulus
1030
Oreosedum hierapetrae (Rech.f.) Grulich
1205
Ophrys perpusilla Devilliers-Tersch. & Devilliers
1031
Oreosedum tristriatum (Boiss. & Heldr.) Grilich.
1214
Ophrys polyxo J. Mast, M.-A. Garnier, DevillersTersch. & Devillers
1036
Origanum pulchrum Boiss. & Heldr.
1065
1065
Ophrys punctulata auct. non Renz
1031
Origanum scabrum Boiss. & Heldr. subsp.
pulchrum (Boiss. & Heldr.) P.H. Davis
Ophrys renzii (Soó) Soó
1010
Origanum tournefortii Aiton
1061
Ophrys schlechteriana (Soó) Devilliers-Tersch. &
Devilliers
1045
Ornithogalum glaucophyllum Baker?
1068
Ornithogalum gracilipes Zahar.
1075
Ophrys scolopax Cav. subsp. heldreichii (Schltr.)
E. Nelson
1035
Ornithogalum gracilipes Zahar.
1075
1068
Ophrys sphaciotica H. Fleischm.
1051
Ornithogalum montanum Cyr. subsp. atticum
(Bois. & Heldr.) Nyman
Ophrys sphegodes auct. p.p.
1041
1070
Ophrys sphegodes Mill. subsp. aesculapii (Renz)
Soó
1010
Ornithogalum visianicum auct. non Tommasini,
nec Rechinger 1944, & Greuter, 1973
Paeonia arietina auct. non G. Anderson
1085
Paeonia corallina sensu Rechinger
1082
Paeonia cretica Tausch (nom. inval.)
1081
Paeonia mascula (L.) Miller subsp. hellenica
Tzanoud. var. icarica Stearn
1084
Ophrys sphegodes Mill. subsp. cretensis H.
Baumann & Künkele
1048
Ophrys sphegodes Mill. subsp. gortynia H.
Baumann & Künkele
1033
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
388 K. GEORGHIOU and P. DELIPETROU
Pleconax conica (L.) Sourkova subsp. ammophila
(Boiss. & Heldr.) A. Love & Kjellqvist subsp.
ammophila (Boiss. & Heldr.) A. Love & Kjellqvist
1237
1080
Pleconax conica subsp. ammophila var. carpathae
(Chowdhuri) A. Love & Kjellqvist
1238
Paeonia peregrina Miller var. latifolia Boiss.
1085
Pleconax sartorii (Boiss. & Heldr.) Sourkova
1288
Paeonia rhodia Stearn
1082
1131
Pleconax subconica var. grisebachii (Davidov)
Sourkova
1253
Paraskevia cesatiana (Fenzl & Friedr.) W. & G.
Sauer
Pleconax subconica Sourkova subsp. grisebachii
(Davidov) Sourkova
1253
1145
Poa caesia Sm. var. frearitis Orph. ex Halácsy
1112
Petromarula oxyloba Gand.
1089
Poa trichopoda Heldr. & Sart.
1114
Petromarula pinnataoxyloba (L.) A.DC. var.
oxyloba (Gand.) Hayek
1089
Pocockia graeca Boiss. & Spruner
Paeonia officinalis L. var. glabra (Boiss.) Hayek
1080
Paeonia officinalis sensu Rechinger 1943
1082
Paeonia peregrina Miller var. cretica Huth
1080
Paeonia peregrina Miller var. glabra Boiss.
Peltaria emarginata (Boiss.) Hausskn.
Peltaria isatoides W. Barbey
Petrorhagia suffruticosa Rech.f. & Phitos
Peucedanum longifolium Waldst. & Kit. subsp.
vourinense Leute
Pevalekia lunarioides (Willd.) Trinajstic
Phaeopappus saxatilis (K. Koch) Boiss.
852
245
1101
683
416
Phagnalon pumilum (Sm.) DC.
1102
Phleum arenarium auct. p.p., non L.
1104
Phleum arenarium L. subsp. aegaeum Vierh.
1104
Phleum graecum Boiss. & Heldr. subsp. aegaeum
(Vierh.) Greuter
1104
Phlomis clandestina Bory & Chaub.
1227
Phlomis fruticosa sensu Forsyth Major & Barbey
1849, Rechinger 1943
1106
927
Podanthum giganteum Boiss.
219
Podospermum idaeum Gand.
1184
Polygala nicaeensis Risso ex Koch subsp. graeca
Chodat
1118
Polygala nicaeensis Risso ex Koch var. tomentella
Boiss.
1118
Polygonum aviculare var. nanum sensu Halácsy
1122
Polygonum aviculare L. var. alpinum Boiss.
1122
Potentilla apennina subsp. kionaea (Halácsy)
Maire & Petitmengin
1126
Potentilla geoides Bieb. subsp. geoides sensu Strid
p.p.
614
Potentilla geoides Bieb. subsp. longisepala Strid
613
Potentilla geoides Bieb. subsp. halacsyana (Degen)
Strid
612
Phoenixopus alpestris Gand.
845
Potentilla halacsyana Degen
612
Phyteuma jacquinii Sieber
298
Potentilla longisepala (Strid) Soják
613
Potentilla longisepala (Strid) Soják subsp.
epirotica Soják
614
Potentilla rupestris L. var. baeniczkyi (Friv.)
T. Wolf subvar. halacsyana (Degen) Hayek
612
Potentilla rupestris L. var. halacsyana (Degen)
T. Wolf
612
Phyteuma pinnatum L.
1089
Pimpinella depressa DC.
1109
Pimpinella tragium var. maritima Boiss.
1107
Pimpinella tragium Vill var. depressa (DC.) Boiss.
1109
Pistolochia uniflora (Sieber) Sojak
506
Planera abelicea (Lam.) Schultes
1510
Planera cretica (Spach) Kotschy
1510
Plantago atrata Hoppe f. nana Pilger
1110
Plantago graeca Halácsy
1110
Plantago montana Lam. var. graeca (Halácsy)
Halácsy
1110
Pleconax ammophila (Bois. & Heldr.) Šourková
1236
Pleconax ammophila (Bois. & Heldr.) Šourková
subsp. carpathae (Chowdhuri) Šourková
1238
Pleconax carpathae (Chowdhuri) Ikonn.
Pleconax conica var. sartorii (Boiss. & Heldr.) A
Love & Kjellqvist
Poterium sphacioticum Gand.
1154
Procopiania cretica (Willd.) Gusuleac
1334
Procopiania insularis Pawl.
1338
Prokopiania cretica sensu Fl. Aeg.
1338
Prospero battagliae Speta
1171
Prospero depressum Speta
1173
Prospero hierapytnense Speta
1174
Prospero idaeum Speta
1175
1238
Prospero minimum Speta
1177
1288
Prospero rhadamanthi Speta
1179
Prospero talosii (Tzanoud. & Kypr.) Speta
1180
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 389
Prunella laciniata (L.) L. var. cretica (Boiss.)
Hayek
1128
Psilostemon creticus (Willd.) DC.
1334
Ptarmica barbeyana (Heldr. & Heimerl) Nyman
Ptarmica umbellata (Sm.) DC.
Ptarmica vulgaris DC.
9
14
9
Salsola vermiculata auct. balcan., non L.
1149
Salsola vermiculata auct. balcan., non L.
1150
Sanguisorba cretica Hayek f. sphaciotica (Gand.)
Hayek
1154
Santolina montana Sm.
126
Saponaria caespitosa Sm.
1247
Pterocephalus parnassi Sprengel
1129
Saponaria calabrica Guss. var. aenesia Heldr.
1155
Pterocephalus perennis Coulter subsp. parnassi
(Sprengel) Vierh.
1129
Saponaria fruticulosa Bory & Chaub.
245
Ptilostemon gnaphaloides (Cyr.) Sojak subsp.
pseudofruticosus (Pamp.) W. Greuter
1130
Saponaria graeca Schreber
246
Saponaria polygonoides (Willd.) Jaub & Spach
246
Ptilotrichon emarginatum Boiss.
852
Ptilotrichon tymphaeum (Hausskn.) Halácsy
251
Pulmonaria auriculata (Boiss.) Halácsy
Puschkinia dubia Kunth
1131
231
Quercus abelicea Lam.
1510
Quercus euboica Papaioannou
1132
Ramonda heldrechii (Boiss.) Fritsch.
842
Ranunculus cadmicus Boiss. var. parviflorus
Boiss.
1136
Ranunculus cadmicus sensu Halácsy, non Boiss.
1136
Ranunculus chaerophyllos L. var. subhomophyllus
Halácsy
Saponaria saxifraga L. var. smithii (Ser.) Rohrb.
1247
Saponaria smithii Ser.
1247
Saponaria thymifolia var. ocellata (Sm.) Boiss.
246
Saponaria thymifolia (Sm.) Boiss.
249
Satureja athoa K. Malύ
1163
Satureja calamintha (L.) Scheele var. hirta Briq.
268
Satureja candica Greuter & Burdet
930
Satureja hispida (Benth.) Nyman
930
Satureja macedonica Form.
1161
1164
1136
Satureja montana L. subsp. parnassica (Heldr. &
Sart.) Maire & Petitmengin
1159
Ranunculus subhomophyllus (Halácsy) Vierh.
subsp. cylleneus Vierh.
1136
Satureja montana L. var. hellenica (Heldr.)
Halácsy
1136
Satureja montana L. var. pisidica (Wettst.)
Halácsy
1161
Ranunculus subhomophyllus (Halácsy) Vierh.
subsp. parviflorus (Boiss.) Vierh.
Satureja olympica Halácsy, nomen nud.
1161
Ranunculus subhomophyllus (Halácsy) Vierh. var.
grandiflorus Vierh.
1136
Satureja parnassica Heldr. & Sart var. stygia
Orph.
1164
Raparia bulbosa (Boiss.) F.K. Mayer
1401
1160
Reseda epirotica Form.
1139
Satureja parnassica Heldr. & Sart. ex Boiss. var.
macrophylla Halácsy
Reseda tymphaea Hausskn. subsp. tymphaea
1139
1159
Reseda tymphaea Hausskn. var. asperula Halácsy
1139
Satureja parnassica Heldr. & Sart. sensu Fl. Aeg.
p.p.
Reutera rigidula Boiss. & Orph.
1108
Rhamnus fallax Boiss. var. guicciardi Boiss.
1140
Rhamnus pubescens Sm.
1141
Rhamnus sibthorpianus Roemer & Schultes
(including taxon)
1140
Rhinanthus intermedius Quezel & Contandr.
1142
Rhinanthus major L. var. parnassicus Boiss.
1142
Rhynchosinapis nivalis (Boiss. & Heldr.) Heywood
479
Robertia tenuiflora Boiss.
823
Satureja sancta Greuter & Burdet
268
Satureja sphaciotica (Boiss. & Heldr. ex Benth.)
Greuter & Burdet
931
Satureja taygetea (P.H. Davis) Greuter & Burdet
932
Saxifraga cymbalaria sensu Sinth. & Sm.
1165
Saxifraga fruticulosa subsp. taygetea (Halácsy)
Hayek
1292
Scabiosa albocincta Greuter
901
Scabiosa breviscapa Boiss. & Heldr.
902
Scabiosa centauroides sensu Sieber et auct. cret.
443
Scabiosa crenata Cyr. subsp. breviscapa (Boiss. &
Heldr.) Hayek
902
Rumia frigida Boiss. & Heldr.
1427
Ruta chalepensis L. var. fumariifolia Boiss.
1148
Ruta fumariifolia Boiss. & Heldr.
1148
1150
Scabiosa crenata Cyr. var. breviscapa (Boiss. &
Heldr.) Boiss.
902
Salsola aegaea sensu auct. karp., Rechinger 1943
Salsola brevifolia sensu d’Urv.
1149
Scabiosa cretacea auct.
443
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
390 K. GEORGHIOU and P. DELIPETROU
Scabiosa cretica (L.) Greuter & Burdet subsp.
minoana P.H. Davis
906
Scorzonera purpurea Hausskn. subsp. rhodantha
(Hausskn.) Maire & Petitmengin
1115
Scabiosa cretica (L.) Greuter subsp. minoana P.H.
Davis
904
Scorzonera rhodantha Hausskn.
1115
1115
Scabiosa cretica L. var. heterophylla Hampe
903
Scorzonera rosea Waldst. & Kit. subsp. peristerica
(Formánek) Greuter
Scabiosa cretica L. var. trifoliolata Bory & Chaub.
903
Scrophularia caesia Sm.
1187
Scabiosa cretica Scabiosa cretica sensu Baldacci
1903, p.p.; Fl. Aeg. p.p.
905
Scrophularia heterophylla Willd. var. caesia (Sm.)
Weiss
1187
Scabiosa cretica sensu auct. cret. Fl. Aeg. p.p.
(non L. 1753 s.str.)
906
Scrophularia heterophylla Willd. var. obtisiloba
(Margot & Reuter) Bornm.
1187
1166
Scrophularia heterophylla Willd. var. oliveriana
(Wydler) Weiss
1187
Scrophularia heterophylla Willd. var. pinnatisecta
Boiss.
1187
Scabiosa holocericea Bory
Scabiosa hymettia Boiss. & Spruner
903
Scabiosa minoana (P.H. Davis) Greuter
904
Scabiosa minoana (P.H. Davis) Greuter
906
905
Scrophularia heterophylla Willd. var. tenuis
(Hausskn.) Hayek
1187
Scabiosa minoana (P.H. Davis) Greuter subsp.
asterusica Greuter
906
Scrophularia laciniata var. obtusiloba Margot &
Reuter
1187
Scabiosa minoana (P.H. Davis) Greuter subsp.
minoana
Scrophularia micrantha D’ Urv.
1187
Scrophularia oliveriana Wydler
1187
Scrophularia sibthorpiana Sprengel
1187
Scrophularia tenuis Hausskn.
1187
Scrophularia urvilleana Wydler
1187
Scutellaria euboea Rech.f.
1197
Scutellaria geraniana (Tunt.) Rech.f.
1201
Scutellaria geraniana (Tuntas ex Halácsy) Rech.f.
1197
Scutellaria hirta Sieber, non Sm.
1200
Scutellaria icarica Rech.f.
1190
Scutellaria peregrina L. subsp. rupestris (Boiss. &
Heldr.) Maire & Petitmengin
1199
Scutellaria peregrina L. var. geraniana (Halácsy)
Hayek
1197
Scutellaria peregrina L. var. parnassica Boiss.
1197
Scutellaria rubicunda Hornem. subsp. cephalonica
(Bornm.) Rech.f.
1194
Scabiosa pterocephala L.
1129
Scabiosa sphaciotica Roemer & J.A. Schultes
Scabiosa taygetea Boiss. & Heldr. var. pindicola
Hausskn.
907
1166
Scabiosa trifoliolata (Bory & Chaub.) Halácsy
903
Scabiosa uralensis auct. var. cretacea sensu Rchb.
1826
443
Scaligeria cretica (Miller) Boiss. subsp. halophila
Rech.f.
1167
Scariola alpestris (Gand.) Greuter, Matthäs &
Risse
845
Scilla albanica Turill?
1176
Scilla albescens Speta
1178
Scilla amoena sensu Chaub.ard & Bory
1176
Scilla bifolia sensu auct. cret., non L.
1172
Scilla cretica (Boiss. & Heldr.) Speta
1178
Scilla hyacinthoides sensu Seitz 1978
1178
1178
Scutellaria rubicunda Hornem. subsp. cytherea
Rech.f.
1195
Scilla nana (Schult. & Schult. f) Speta subsp.
albescens (Speta) Speta
1399
Scutellaria rubicunda Hornem. subsp. geraniana
(Halácsy) I.B.K. Richardson
1197
Sclerochorton junceum (Sm.) Boiss.
Scutellaria rubicunda Hornem. subsp. hellenica
Rech.f.
1197
Scutellaria rubicunda Hornem. subsp. icarica
(Rech.f.) Rech.f.
1190
Scutellaria rubicunda Hornem. subsp. parnassica
(Boiss.) Rech.f.
1197
Scutellaria rubicunda Hornem. subsp. rupestris
(Boiss. & Heldr.) I.B.K. Richardson
1199
Scutellaria rubicunda Hornem. subsp. rupestris
(Boiss. & Heldr.) I.B.K. Richardson p.p.
1197
Scleroptychis cretica (L.) Degen
910
Scorzonera araneosa Sibth & Sm. described from
Kiklades?
1183
Scorzonera dependens Rech.f.?
1181
Scorzonera idaea (Gandoger) Lipsic.
1184
Scorzonera lassitica Vierh.
1181
Scorzonera mollis sensu auct., Rechinger Fl. Aeg.
1181
Scorzonera purpurea Hausskn. subsp. peristerica
Formánek
1115
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 391
Serapias orientalis (Greuter) H. Baumann &
Künkele subsp. cycladum (H. Baumann &
Künkele) Nelson
1219
Seseli crithmifolium (DC.) Boiss.
1222
Seseli gummiferum Pall. ex Sm. subsp. aegaeum
P.H. Davis (included in taxon)
1222
1197
Seseli junceum Sm.
1399
Scutellaria sieberi
1190
Seseli stylare Boiss. & Heldr. (from Taygetos)
1399
Scutellaria vacillans Rech.f.
1189
Sesleria krajinae Deyl
1226
Scutellaria velenovskyi Rech.f. subsp. goulimyi
(Rech.f.) Rech.f.
1191
Sherardia pusilla Chaub. & Bory
173
Sideritis athoa Papan. & Kokkini
1231
Sedum annuum L. subsp. epiroticum Baldacii
1209
Sideritis attica Heldr.
1232
Sedum cretense Maire
1206
1229
Sedum creticum Boiss. & Heldr.
1206
Sideritis clandestina (Bory & Chaub.) Hayek
subsp. cyllenea (Hayek) Papan. & Kokkini
Sedum creticum C.Presl var. creticum sensu
Mount. Fl. Gr., 1986
1206
Sideritis cretica Sm. p.p., non L.
1227
Sideritis peloponnesiaca Boiss. & Heldr.
1229
Sedum creticum C.Presl var. hierapetrae (Rech.f.)
‘T Hart & Hageman
1205
Sideritis perfoliata L. var. lanata Turrill
1231
Sedum delicum (Vierh.) A. Carlström
1208
Sideritis raeseri Boiss. & Heldr. var. attica
(Heldr.) Halácsy
1232
Sedum delicum sensu auct. cret., non (Vierh.)
Carlström
1211
Sideritis syriaca L. var. condensata Boiss. &
Heldr.
1230
Sedum hierapetrae Rech.f.
1205
Sideritis syriaca sensu Bory & Chaub., non L.
1227
Sedum idaeum D.A. Webb.
1212
Sideritis theezans Boiss. & Heldr.
1227
Sedum laconicum Boiss. & Heldr. subsp. insulare
(Rech.f.) Greuter & Rech.f.
1212
Sideritis theezans Boiss. & Heldr. subsp.
peloponnesiaca (Boiss. & Heldr.) Bornm.
1229
Sedum laconicum Boiss. & Heldr. var. insulare
Rech.f.
1212
Sideritis theezans Boiss. & Heldr. var. cyllenea
Boiss.
1229
Sedum litoreum Guss. subsp. praesidis
(Runemark & Greuter) Greuter
1213
Silene aegyptiaca auct. [non (L.) L.]
1286
Sedum praesidis Runemark & Greuter
1213
Silene auriculata subsp. lanuginosa (Bertol.)
Arcangeli
1240
Sedum rubens subsp. delicum sensu auct. cret.,
non Vierh.
1211
Silene caesia sensu Rechinger 1943
1287
Sedum rubens L. subsp. delicum Vierh.
1208
Silene caesia Sm. subsp. samothracica (Rech.f.)
Melzh.
1287
Sedum rubens L. var. delicum (Vierh.) Hayek
1208
Silene chromodonta Boiss. & Reuter
1284
Sedum sexangulare auct. non L.
1203
Silene congesta Sm. subsp. moreana Melzh.
1243
Sedum tymphaeum Quézel & Contandr.
1127
1288
Sempervivum ballsii Wale
1215
Silene conica L. subsp. sartorii (Boiss. & Heldr.)
Chater & Walters
Scutellaria rubicunda Hornem. subsp. samia
Rech.f.
1190
Scutellaria rupestris Boiss. & Heldr. subsp.
geraniana (Halácsy) Greuter
1197
Scutellaria rupestris Boiss. & Heldr. var.
cephalonica Bornm.
1194
Scutellaria sibthorpii var. geraniana Halácsy
Senecio ambiguus subsp. ambiguus sec. Strid, A.
(ed.) – Mountain flora of Greece 2 Edinburgh 1991
840
Silene cucubalus Wibel. subsp. megalosperma
(Sart. ex Heldr.) Rech.f.
1294
Senecio ambiguus (Biv.) DC. subsp. taygeteus
(Boiss. & Heldr.) Greuter
840
Silene delphica Boiss. & Heldr.
1243
Silene dictaea Rech.f.
1250
Senecio gnaphalioides Spreng.
841
Silene dionysii Stoj. & Jordanov
1278
Senecio gnaphalodes Sieber
841
Silene dionysii Stoj. & Jordanov var. elata Stoj.
& Jordanov
1278
Senecio taygeteus Boiss. & Heldr.
840
Serapias cordigera L. subsp. cretica B. Baumann
& H. Baumann
1218
Silene dionysii Stoj. & Jordanov var. pumila Stoj.
& Jordanov
1278
Serapias cordigera L. subsp. patmia (M. Hirth &
H. Spaeth) Kreutz
1220
Silene fruticulosa Sieber ex Otth
1239
Silene genistifolia Halácsy
1269
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
392 K. GEORGHIOU and P. DELIPETROU
Silene greuteri Phitos
1260
Silene smithii (Ser.) Boiss. & Heldr.
1247
Silene inflata var. microphylla Boiss.
1294
Silene spinescens Sm. var. argolica Boiss.
1291
Silene italica (L.) Pers. var. cythnia Halácsy
1246
Silene stenocalycina Rech.f.
1269
Silene juncea auct. [non Roth]
1245
1245
Silene subconica Friv. subsp. grisebachii
(Davidov) Jordanov & Panov
1253
Silene kotschyi Boiss. var. corinthiaca (Boiss. &
Heldr.) F.N. Williams
Silene subconica Friv. var. grisebachii Davidov
1253
Silene lanuginosa Bertol.
1240
1242
Silene thessalonica Boiss. & Heldr. subsp. dictaea
(Rech.f.) Melzh.
1250
Silene linifolia Sibth. & Sm. subsp. cephallenia
(Heldr.) Maire & Petitmengin
1267
Silene variegata (Desf.) Boiss. & Heldr. var.
samothracica Rech.f.
1287
Silene linifolia Sm.
Silene linifolia Sm. subsp. sporadum (Halácsy)
Phitos
1269
Silene venosa Ascherson var. megalosperma
(Sart. ex Heldr.) Halácsy
1294
Silene linifolia Sm. var. ceccariniana Hausskn.
1267
Silene vittata auct. [non Stapf]
1245
Silene linifolia Sm. var. gracilis Beauverd &
Topali
1267
Sinapis radicata S. & Sm.
253
Silene linifolia Sm. var. sporadum Halácsy
1269
Siphonostylis cretensis (Janka) Schulze subsp.
cretensis
835
Silene marschallii auct. non Mey.
1254
Siphonostylis Hybridocomplex Schulze
834
Silene megalosperma Sart. Ex Heldr.
1294
Silene multicaulis Guss. subsp. genistifolia
(Halácsy) Melz.
1269
Silene multicaulis Guss. subsp. stenocalycina
(Rech. fil) Melz.
1269
Silene polygonoides (Willd.) Poiret
246
Silene portensis auct. [non L.]
1245
Silene portensis L. subsp. rigidula Greuter &
Burdet
1245
Soldanella pelia Raus
1297
Stachys acutifolia sensu Halácsy (1902, 1908,
1912), Rechinger (1943), Tutin & al. (1972)
1305
Stachys argolica Boiss.
1318
Stachys candida auct., non Bory & Chaub.
1306
Stachys candida Bory & Chaub. subsp.
chrysantha Benth.
1302
Stachys ionica Halácsy var. cephalonica Halácsy
1306
1301
Silene quadridentata subsp. phyllitica Neumayer
1284
Stachys messeniaca Boiss.
Silene quadridentata (Murray) Pers. subsp.
chromodonta (Boiss. & Reuter) H. Neumayer
1284
Stachys scyronica Boiss.
1320
1275
Stachys swainsonii Benth. var. argolica (Boiss.)
Boiss.
1318
Silene radicosa Boiss. & Heldr. subsp.
pseudoradicosa Rech.f.
Stachys swansonii auct., non Benth.
1306
Silene radicosa Boiss. & Heldr. subsp. rechingeri
Melzh.
1268
Stachys swansonii Benth. var. ionica (Halácsy)
Hayek f. cephalonica (Halácsy) Hayek
1306
Silene radicosa Boiss. & Heldr. var. breviflora
Boiss.
1272
Staehelina arborea Schreb.
1324
Silene retzdorffiana sensu – Quezel & Contandr.
1263
Staehelina arborescens L.
1324
Silene rigidula Sm.
1245
Silene saxifraga L. subsp. fruticulosa Arcang.
1239
Silene saxifraga L. subsp. smithii (Ser.) Nyman
1247
Silene saxifraga L. subsp. taygetea (Halácsy ex
Vierh.) Hayek
1292
Silene saxifraga L. var. coenobitarum Hayek
1244
Silene saxifraga sensu auct. Kret non L.
1239
Silene sedoides Poiret subsp. haussknechtii
(Hausskn.) Maire & Petitm.
1255
Silene sedoides Poiret subsp. pentelica (Boiss.)
Nyman
1280
Silene sedoides Poiret var. pentelica (Boiss.)
Rohrb.
1280
Staehelina fruticosa (L.) L.
Statice androsacea Jaub. & Spach
804
3
Statice cancellata auct. (non Bernh.) var. glabra
Boiss. p.p.
884
Statice cancellata auct. (non Bernh.) var. laxa
Boiss. p.p.
863
Statice cancellata auct. (non Bernh.) var. laxa
Boiss. p.p.
887
Statice cancellata auct. (non Bernh.) var. laxa
Boiss. p.p.
863
Statice cancellata auct., non Bernh.
859
Statice carpatha Rech.f.
862
Statice corcyrense Guss. var. major Boiss. p.p.
867
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 393
Statice corinthiaca Boiss. & Heldr.
865
Teucrium corcyanum Quezel & Contandr.
1397
Statice cosyrensis Guss. subsp. melia Nyman
879
1393
Statice doerfleri Halácsy
871
Teucrium divaricatum Heldr. var. athoum
Hausskn.
Statice frederici Barbey
873
Teucrium gossypium Rech.f. (included in taxon)
1389
Statice graeca Poiret var. palmaris (Sm.) Hayek
882
Teucrium heliotropifolium Barbey
1398
Statice heldreichii Halácsy
764
Teucrium werneri Rech.f.
1395
Statice hyssopifolia Girard
876
Thlaspi epiroticum Halácsy (sphalm.)
1403
Statice melia Nyman p.p.
867
1402
Statice ocymifolia Poiret
881
Thlaspi graecum Jordan subsp. creticum (Degen
& Jav.) Greuter
Statice palmaris Sm.
882
Thlaspi graecum sensu suct. cret., Fl. Eur.;
Greuter 1974
1402
Statice pigadiensis Rech.f.
881
1402
Statice rorida var. palmaris Boiss.
882
Thlaspi microphyllum Boiss. & Orph. subsp.
creticum Degen & Jav
Statice virgata Willd.var. tenia Heldr.
889
Thlaspi taygeteum Boiss.
1404
Sternbergia sicula auct. plur. fl. cret., non Tineo
ex Guss.
1325
Thymus euboeus Halácsy
1407
Thymus leucadicus (Rchb.f.) Halácsy
1411
Stipa cyllenea Strid
1328
Strangweia hyacinthoides Sm.
Strangweia spicata (Sm.) Boiss.
Thymus pastoralis Turrill non Iljin
1412
231
Thymus rechingeri Hartvig
1408
231
Thymus rechingeri Hartvig subsp. macrocalyx
Hartvig
1410
Thymus rechingeri Hartvig subsp. rechingeri
1409
Symphyandra cretica subsp. sporadum (Halácsy)
Hayek
1333
Symphyandra cretica A. DC. subsp. samothracica
(Degen) Hayek
1332
Thymus rechingeri Hartvig subsp. rechingeri
1410
1407
Symphyandra cretica A. DC. subsp. cretica
1331
Thymus serpyllum L. var. angustifolius sensu
Boiss. Fl. Or. p.p.
Symphyandra cretica A. DC. subsp. eu-cretica
Hayek
1331
Thymus striatus sensu auct., non Val.
1411
1416
Symphytum anatolicum auct. from Naxos Fl. Aeg.
1339
Thymus teucrioides Boiss. & Spruner var.
candilicus Beauv.
Symphytum davisii sensu Wickens 1969 pp.
1337
1335
Thymus teucrioides Boiss. & Spruner var. deorum
Ronniger p.p.
1408
Symphytum davisii Wickens subsp. cycladense
(Pawl.) Stearn
1413
Symphytum davisii Wickens subsp. davisii
1336
Thymus teucrioides Boiss. & Spruner var. deorum
Ronniger p.p.
Symphytum davisii Wickens subsp. icarisum
(Pawl.) Stearn
1337
Thymus teucrioides sensu Boiss. Fl. Or. p.p.
1408
Thymus teucrioides sensu Boiss. Fl. Or. p.p.
1413
1414
Symphytum davisii Wickens subsp. naxicola
(Pawl.) Stearn
1339
Thymus zygioides var. plasonii (Adamović) H.
Braun
Taraxacum apenninum sensu Hayek Prodr. Fl.
Penins. Balc.
1345
Trachelium asperuloides Boiss. & Orph.
278
Trachelium jacquinii (Sieber) Boiss. p.p.
298
Taraxacum glaciale sensu Halácksy Fl. Graec.
1345
1383
Trachelium jacquinii (Sieber) Boiss. subsp.
jacquinii
298
Taraxacum lacistophylloides Sonck, non Dahlst.
Telephium imperati L. subsp. orientale (Boiss.)
Nyman var. pauciflorum Greuter
1387
Trachystemon creticus (Willd.) G. Donf.
1334
Trachystemon orientalis auct.
1338
Telephium imperati L. var. pauciflorum Greuter
1387
Tragopogon longifolius Heldr. & Sart. ex Boiss
1417
Trifolium agrarium L., nom. ambig.
1420
Tetragonolobus aduncus Griseb.
909
Teucrium alpestre Barbey & Major (sphalm.)
subsp. gracilis
1396
Tripleurospermum rosellum (Boiss. & Orph.)
Hayek
921
Teucrium alpestre Sibth. & Smit subsp. alpestre
1389
922
Teucrium alpestre Sibth. & Smit subsp. gracile
(Barbey & Major) D. Wood
1396
Tripleurospermum tempskyanum (Freyn & Sint.)
Hayek
Trisetum rechingeri Chrtek
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
1429
394 K. GEORGHIOU and P. DELIPETROU
Triticum comosum Sm. K. Richt. subsp.
heldreichii (Boiss.) Greuter
20
Triticum heldreichii (Boiss.) K. Richt.
20
Verbascum parnassicum Halácsy
1451
Verbascum pelium Halácsy
1441
Verbascum pindicolum Freyn & Sint.
1452
Trochocephalus albocinctus (Greuter) j. Holug
901
Verbascum samaritanii Heldr. ex Boiss.
1458
Trochocephalus hymettius (Boiss. & Spruner)
J. Holug
903
Verbascum sartorii Halácsy non Boiss. & Heldr.
1462
Trochocephalus minoanus (P.H. Davis) Holug
904
Verbascum speciosum Schrader var. megaphlomos
Boiss. & Heldr.
1465
Trochocephalus minoanus (P.H. Davis) J. Holub
906
Verbascum tomentosum (Zucc.) O. Kuntze
1467
Verbascum tymphaeum Freyn & Sint.
1452
Verbascum viscidulum Freyn & Sint.
1452
Veronica amoena sensu Heldr., non Bieb.
1472
Veronica austriaca L. subsp. dentata (F.W.
Schmidt) Watzl var. teucrioides (Boiss. & Heldr.)
Waltzl
1476
Tulipa hageri Heldr. var. doerfleri (Gandoger)
Hayek
1432
Tunica cretica auct. p.p. [non (L.) Fischer &
C.A. Mey.]
1090
Tunica cretica sensu Hayek p.p., Baldacci 1895,
Rechinger 1944
1096
Tunica dianthoides (Sm.) Fischer & C.A. Mayer
1091
Tunica fasciculata (Margot & Reuter) Boiss.
1092
Veronica austriaca L. var. teucrioides (Boiss. &
Heldr.) Halácsy quoad basion. solo
1476
Tunica glumacea (Chaub. & Bory) Boiss.
1093
Veronica chamaedryoides Bory & Chaub.
1468
Tunica graminea (Sm.) Boiss.
1094
1468
Tunica illyrica (Ard.) Fischer var. taygetea Boiss.
1096
Veronica chamaedrys L. var. chamaedryoides
(Bory & Chaub.) Nyman
Tunica ochroleuca (Sm.) Fischer & C.A. Mey. var.
phtiotica (Boiss. & Heldr.) Hayek
1098
Veronica chamaedrys L. var. pindica Hausskn.
1468
Veronica chamaedrys L. var. rigida Turrill
1468
Tunica ochroleuca Sm. Fischer & C.A. Mey.
1097
Veronica cretica Pall. ex Link
1478
Tunica phtiotica Boiss. & Heldr.
1098
Veronica graeca Spruner, nomen nudum
1472
Tunica taygetea (Boiss.) P.H. Davis
1096
Veronica kavusica Rech.f.
1473
Tunica taygetea auct. pro parte, non (Boiss.)
P.H. Davis
1090
Veronica peduncularis Bory & Chaub., non Bieb.
1471
Ulmus abelicea (Lam.) Sm.
1510
Veronica peloponnesiaca Boiss. & Orph.
1474
Umbilicus sprunerianus Boiss.
1435
Veronica praecox sensu Rechinger, 1943
1473
Valeriana montana L. subsp. crinii Orph. ex
Boiss.
1437
Veronica prostrata var. kajmakcalanica
Adamović?
1476
Valeriana montana L. var. crinii (Orph. ex Boiss.)
Boiss
1437
Veronica rigida (Turrill) Turrill
1468
Veronica teucrioides Boiss. & Heldr.
1476
Verbascum adenotrichum Halácsy
1452
Veronica tymfrestea Boiss. & Spruner
1478
Verbascum agrimonioides Degen & Borbas
1452
Verbascum epirotum Halácsy
1458
Verbascum epixanthinum Boiss. & Heldr. var.
samaritanii Boiss.
1458
Verbascum epixanthinum Boiss. & Heldr. var.
taygeteum (Halácsy) Murb
Vesicaria tymphaea Hausskn.
251
Vicia aegaea (Halácsy) Rech.f.
1479
Vicia cretica Boiss. & Heldr. var. aegaea Halácsy
1479
Vicia villosa subsp. semicretica Ponert
1480
1450
Viola cenisia L. var. striis notata J. Wagner
1506
1486
Verbascum epixanthinum Halácsy var. taygeteum
1450
Viola chelmea Boiss. & Heldr. subsp. chelmea
sensu Tutin et al., Fl. Eur.; Strid M. Fl. Gr.
Verbascum graecum Heldr. & Sart. var. zuccarinii
Boiss.
1467
Viola declinata Waldst. & Kit. var. epirota
Halácsy
1491
Verbascum megaphlomos (Boiss. & Heldr.)
Halácsy
1465
Viola euboea (Halácsy) Halácsy var. epirota
Halácsy
1491
Verbascum megaphlomos (Boiss. & Heldr.)
Halácsy var. macrophyllum Halácsy
1465
Viola gracilis var. samothracica (Degen) Rech.f.
1503
1495
Verbascum nigrum subsp. hypoleucum (Boiss. &
Heldr.) Nyman
1456
Viola gracilis sensu auct., Fl. Or., Consp. Fl.
Graec.
Viola gracilis Sm. var. euboea Halácsy
1492
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 395
Viola grisebachiana Vis. f. integrifolia Stoj. &
Jordanov
1506
Viola heldreichiana Boiss. f. mercurii (Orph.)
W. Becker
1496
Viola heterophylla Bertol subsp. euboea (Halácsy)
W. Becker
1492
Viola heterophylla Bertol var. euboea (Halácsy)
W. Becker
1492
Viola heterophylla Bertol. subsp. epirota (Halácsy)
W. Becker
1491
Viola heterophylla Bertol. subsp. graeca
(W. Becker) W. Becker
1495
Viola heterophylla Bertol. var. graeca W. Becker
1495
Viola olympica var. samothracica Degen
1503
Viola tricolor L. var. demetria Boiss.
1496
Wagenitzia lancifolia (Sieber ex Sprengel) Dostál
390
Wilckia angulifolia (Boiss. & Orph.) Halácsy
918
Wilckia bicolor Boiss. & Heldr.
913
Wilckia maritima (L.) Halácsy var. macrocalyx
Halácsy
915
Zelkova crenata var. cretica Spach
1510
Zelkova cretica (Spach) Spach
1510
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AFE
Jalas J. & Suominen J. (eds). 1972–1997. Atlas Florae Europaeae. Distribution of vascular plants in
Europe. Vols 1–11. Helsinki University Printing House, Helsinki.
EFK
Jahn R. & Schönfelder P. 1995. Exkursionsflora für Kreta. Verlang Eugen Ulmer, Stuttgart (Hohenheim).
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FAe
Rechinger K.H. 1943. Flora Aegaea. Flora der Inseln und Halbinseln des ägäischen Meeres. Acad. Wiss.
Wien, Math.-Naturwiss. Kl., Denkschr. 105(1): xx + 924p. +25 plates and 3 maps. Reprinted by Otto Koeltz
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FCA
Turland, N. J., Chilton, L. & Press, J. R. 1993. Flora of the Cretan Area. Annotated Check List and Atlas.
The Natural History Museum, HMSO, London. 439p.
FCSI
Chilton L., Turland N.J. 1997. Flora of Crete. A supplement. Marengo Publications, UK, 125p. ISBN
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FCSII
Turland N.J., Chilton L. 2008. Flora of Crete: Supplement II, Additions 1997–2007. Availalable at http://
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Tutin, T.G. et al. (eds). 1968–1980. Flora Europaea. Vols 2–5. Cambridge University Press, Cambridge, UK.
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FGC
Halacsy E. De. 1901; 1902; 1904. Conspectus Florae Graecae 1; 2; 3. G. Engelmann, Lipsiae.
FGS
Halacsy E. De. 1908. Supplementum Conspectus Florae Graecae. G. Engelmann. Lipsiae.
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FHe
Strid A. & Tan K. (eds) 1997–2002. Flora Hellenica. Vols. 1–2. Koeltz Scientific Books, Köningstein,
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FT
Davis P.H. (ed.) 1965–1985. Flora of Turkey and the East Aegean Islands. Vols 1–9. Edinburgh University
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MCL
Greuter, W., Burdet, H.M. & Long, G. 1984–1989. Med-Checklist. A critical inventory of vascular plants of
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MFG
Strid, A. (ed). 1986. Mountain Flora of Greece. Vol 1. Cambridge University Press, Cambridge.
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D. Phitos, A. Strid, S. Snogerup, W. Greuter (eds). The Red Data Book of rare and threatened plants of
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Taxa currently considered synonyms or non-existent or mistaken names cited as greek endemics
Taxon
Detail
Alyssum chlorocarpum Hausskn.
Synonym of Alyssum chalcidicum Janca.
Alyssum obtusifolium Steven ex DC. subsp.
helioscopioides E.I. Nyár.
Synonym of Alyssum sibiricum Willd.
Anchusa macrosyrinx Rech.f.
Included in Anchusa undulata subsp. sartorii.
Anchusa rechingeri Riedl
Synonym of Nonea obtusifolia (Willd.) DC
Anchusa stridii Selvi & Bigazzi
A name (which was apparently never published) for A.
leucantha Selvi & Bigazzi.
Arabis longistyla Rech.f.
Synonym of Diplotaxis viminea (L.) DC.
Arabis olympica Stoj. & Jordanov
Synonym of Arabis collina Ten.
Arenaria filicaulis Fenzl subsp. euboica McNeill
Synonym of Arenaria filicaulis Fenzl. subsp. graeca (Boiss.)
McNeil
Aristolochia tournefortii Jaub. & Spach
Synonym of Aristolochia parvifolia Sm.
Asperula purpurea (L.) Ehrend. subsp. spadicum Boyce
Mistaken name cited in IUCN-WCMC database
Astragalus hellenicus Boiss.
Synonym of Astragalus exscapus L. subsp. exscapus.
Astragalus sempervirens Lam. subsp. cephalonicus (C.
Presl.) Ascherson & Graebner
Synonym of Astragalus sempervirens.
Astragalus taygeteus Jim.Perss. & Strid
Synonym of Astragalus angustifolius Lam.
Barbarea conferta Boiss. & Heldr.
According to the known collections Barbarea conferta falls
into the range of variation of Barbarea vulgaris R. Br. and
is included in it; further collections are needed to establish
whether it should be retained as a separate taxon.
Bunium capillare
mistaken name cited in IUCN-WCMC database
Carduus macrocephalus Desf. subsp. sporadum
(Halácsy) Franco
Synonym of C. nutans subsp. leiophyllus (Petrovic) Stoj. &
Stef.
Carduus taygeteus Boiss. & Heldr. subsp. insularis
Franco
Synonym of C. nutans L. subsp. taygeteus (Boiss. & Heldr.)
Hayek.
Carduus taygeteus Boiss. & Heldr. subsp. taygeteus
Synonym of C. nutans L. subsp. taygeteus (Boiss. & Heldr.)
Hayek.
Carduus thessalus Boiss. & Heldr.
Synonym of Carduus acanthoides L.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
PATTERNS OF PLANT ENDEMISM IN GREECE 421
Centaurea nicopolitana Bornm.
Not a Greek species: it is only known from the type specimen
whose origin is not specified and whose collector had never
visited Greece; probably it is a Bulgarian plant.
Colchicum amabile Heldr.
Synonym of Colchicum bivonae Guss.
Colchicum andrium Rech.f. & P. H. Davis
Synonym of Colchicum pusillum Sieber
Colchicum cousturieri Greuter
Included in Colchicum cupanii.
Colchicum pinatziorum Rech.f.
Synonym of Colchicum boissieri Orph.
Crataegus monogyna Jacq. subsp. aegeica (Pojark.)
Franco
Synonym of Crataegus monogyna Jacq. subsp. monoyna
Crepis pawlowskii Strid
Synonym of Crepis reuteriana Boiss.
Cynoglottis barrelieri (All.) Vural & Kit Tan subsp.
phocidica (Gustavsson) Greuter & Burdet
Synonym of Cynoglottis barrelieri (All.) Vural & Kit Tan
subsp. serpentinicola (Rech.f.) Vural & Kit Tan
Dianthus athous Rech.f.
Synonym of Dianthus gracilis Sm
Dianthus epirotus Halácsy
Synonym of Dianthus armeria L. subsp. armeria
Elymus diae nomen nudum
Synonym of Elytrigia juncea (L.) Nevski subsp. juncea
Elymus hispidus (Opiz) Melderis subsp. graecus
Melderis
Synonym of Elytria intermedia (Host) Nevski subsp.
intermedia
Epipactis degenii Szentpeteri & Monus
Apparently a synonym of Epipactis helleborine.
Epipactis heraclea P. Delforge & Kreutz
Apparently a synonym of Epipactis helleborine.
Fumana pinatsii Rech.f.
Synonym of Fumana arabica.
Fumaria amarysia Boiss. & Heldr.
It is a small seeded extreme of Fumaria judaica Boiss. subsp.
judaica.
Genista melia Boiss.
Collected only once, apparently a mythical taxon or never
existed.
Geranium humbertii Beauverd
Synonym of Geranium subcaulescens L’Her. ex DC.
Hesperis verroiana F. Dvorák
Synonym of Hesperis theophrasti Borbás subsp. theophrasti.
It is noted that further collections are necessary in order to
determine whether the plants from Vermio named H.
verroiana are worthy of a separate taxonomic status.
Inula serpentinica Rech.f. & Goulimy
Synonym of Inula ensifolia L.
Lamium bifidum Cyr. subsp. albimontanum Rech.f.
Synonym of Lamium purpureum L.
Linaria hellenica Turrill
Synonym of Linaria tenuis L.
Linum caespitosum Sm.
Synonym of Linum arboreum L.
Linum doerfleri Rech.f.
Synonym of Linum arboreum L.
Linum euboeum Bornm.
Synonym of Linum elegans Spruner ex Boiss.
Micromeria tapeinantha Rech.f.
An ill known taxon, possibly of hybrid origin, related with
Satureja nana (P.H. Davis & Doroszenco) R. Jahn
Muscari pharmacusanum (Heldr.) Boiss.
Synonym of Muscari comosum (L.) Miller.
Myosotis refracta Boiss. subsp. aegagrophila Greuter &
Grau
Synonym of Myosotis refracta Boiss. subsp. refracta
Onopordum dirphyum Halácsy
Synonym of Onopordum bracteatum Boiss. & Heldr.
Onosma psammophila Rech.f. & Riedl
Synonym of Onosma heterophylla Griseb.
Onosma taygetea Boiss. & Heldr.
Synonym of Onosma visianii Clementi
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422
422 K. GEORGHIOU and P. DELIPETROU
Ophrys crassicornis (Renz) Devilliers-Tersch. &
Devilliers
A synonym of Ophrys oestrifera M.Bieb subsp. bicornis
(Sadler) Kreutz.
Ophrys dictynnae P. Delforge
A synonym of Ophrys tenthredinifera Willd.
Ophrys ulyssea P. Delforge
A synonym of Ophrys tenthredinifera Willd.
Ornithogalum costatum Zahar.
Synonym of Ornithogalum gussonei Ten.
Pinus heldreichii Christ var. heldreichii
Subspecific division is not accepted in Flora Hellenica 1:7
Pisum graecum Quezel & Contandr.
A dubious species, perhaps a synonym of Lathyrus
grandiflorus.
Platanus orientalis L. var. cretica Dode.
Not considered a valid taxon in FHe or in any other recent
flora.
Ranunculus brevifolius Ten. subsp. pindicus (Hausskn.)
E. Mayer
The characters that separate this taxon from the typical
Ranunculus brevifolius Ten. are not constant.
Ranunculus lavrentiadis Papan. & Kokkini
Probably a dwarf form of Ranunculus muricatus or
Ranunculus arvensis.
Ranunculus miliarakesii Halácsy
A slender form of Ranunculus pedatus Waldst. & Kit.
Ranunculus rhodensis Boiss.
An aberrant Rhodian population of Ranunculus paludosus
Poiret
Saxifraga biflora All. subsp. epirotica D.A. Webb
A taxon doubtfully or incorrectly reported from Greece since
the specimens examined appear to represent a lax form of S.
oppositifolia L.
Saxifraga taygetea (Boiss. & Heldr.) Maire &
Petitmengin subsp. chelmea Voliotis
Not considered a valid taxon in FHe or in any other recent
flora.
Scilla pneumonanthe Speta
Included in the range of variation of Scilla bifolia.
Scilla reuteri Speta
Included in the range of variation of Scilla bifolia.
Scutellaria naxensis Bothmer
Synonym of Scutellaria albida L. subsp. velenovskyi (Rech.
fil.) Greuter
Senecio bicolor (Willd.) Tod. subsp. bicolor var. plattii
(Meikle) Matthews
Synonym of Jacobaea maritima subsp. bicolor.
Silene multicaulis Guss. subsp. cretica Melzh.
Synonym of Silene multicaulis Guss. subsp. multicaulis
Silene tempskyana Freyn & Sint
Synonym of Silene conica L. subsp. subconica
Stellaria glochidisperma (Murb.) Freyn
Synonym of Stellaria nemorum L. subsp. montana (Pierrat)
Berher
Thesium coarctiflorum Hendrych
Synonym of Thesium bergeri Zucc.
Trifolium ottonis Spruner ex Boiss.
A synonym of Trifolium noricum Wulfen.
© 2010 The Linnean Society of London, Botanical Journal of the Linnean Society, 2010, 162, 130–422