9/27/2011 Population Growth Not so stable populations Limits to Growth • We are going to work now on adding some realism to growth • Reasonably, growth will be influenced by intrinsic (or finite) rates – Figure 9.1 1 9/27/2011 Limits to Growth • We are going to work now on adding some realism to growth • Reasonably, growth will be influenced by intrinsic (or finite) rates – Figure 9.1 • But, also reasonably, growth can be influenced by conditions such as population size Z = N – Neq (equilib pop size where R0 = 1) (‐0.02 is slope = B) 2 9/27/2011 3 9/27/2011 Conclusion • The behavior of the population will depend on the interaction of the slope (sensitivity) of the response to population size, and the equilibrium population size – If L (= B*Neq) is between 0 and 1, then things go pretty smoothly – Between 1 and 2, less smoothly – 2 and 2.57, indefinite but regular oscillations – Over 2.57, mayhem • So? – So the interplay between B and Neq can cause a variety of effects – In real populations you could see all of these interactions Switching to Overlapping Generations • So far we have been looking at discrete generations – What about when they are overlapping? • For overlapping generations we will need the instantaneous rate r dN/dt = rN = (b‐d)N Nt/N0 = ert Doubling Time • When Nt/N0 = ert = 2, t = 0.69315/r • See how r affects t: 4 9/27/2011 Adding “Carrying Capacity” K • K is the maximum sustainable population • To get an effect of population size on growth: dN/dt = rN(K‐N/K) K‐N/K has been called the “unutilized opportunity for population growth” dN = r(K‐N/K) Dt*N It can be positive or negative 5 9/27/2011 A Not So Good Fit 6 9/27/2011 K can Vary Seasonally or Between Years 7 9/27/2011 Don’t Worry About Theta Logistic 8
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