This is the published version: Velmala, William, Helle, Samuli, Ahola, Markus P., Klaassen, Marcel, Lehikoinen, Esa, Rainio, Kalle, Sirkiä, Paivi M. and Laaksonen, Toni 2015, Natural selection for earlier male arrival to breeding grounds through direct and indirect effects in a migratory songbird, Ecology and evolution, vol. 5, no. 6, pp. 1205-1213. Available from Deakin Research Online: http://hdl.handle.net/10536/DRO/DU:30072452 Reproduced with the kind permission of the copyright owner. Copyright : 2015, Wiley Open Access v Selection differential Direct selection Direct effects Indirect selection Indirect effects Specific (non-causal) indirect effects Specific (causal) indirect effects v Male arrival date Number of fledglings –0.06 (–0.11, –0.01) –0.04 (–0.11, 0.03) –0.07 (–0.11, –0.03) Female laying date 0.26 (0.21, 0.30) –0.01 (–0.04, 0.01) 0.96 (0.78, 1.15) 0.05 (0.03, 0.07) Clutch size 0.01 (0.01, 0.02)
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