350 Forest fire history of Desolation Peak, Washington Jnnps K. Acn,E. Ma.nx FINNEy.I AND Ror-RNooE GouvEruarN2 Collegeof Forest Resources,University of Woshington,Seattle, WA 98195,U.S.A. Received April 17, 1989 Accepted September 26, 1989 A c ; e e , J . K . , F I t t N t v , M . , a n d o n G o u v u N a t N , R . 1 9 9 0 .F o r e s t f i r e h i s t o r y o f D e s o l a t i o nP e a k , W a s h i n g t o n .C a n . J . F o r . R e s .2 0 : 3 5 0 - 3 5 6 . Forestsin the vicinity of DesolationPeak, Washington,are of specialecologicalinterestbecauseof their transitional nature betweencoastaland interior forest types. The area is west of the CascadeMountain crestbut in the rainshadow ofmountains farther to the west. Fire return intervalswere hypothesizedto be shorter than typical for coastal forest types, such as those dominated by western hemlock and Pacific silver fir, and longer than typical for interior forest types,such as ponderosapine, owing to the closejuxtaposition of thesetypesat DesolationPeak. Sevenforest comntunity typeswere defined, and a 400-yearl'ire history was developedfor this 3500-haarea. The averagenatural fire rotation was 100 years;this varied by a factor of two by century and by topographic aspect.Foresttypes typical of coastal regions,suchas Douglas-fir,- westernhemlock and mountain hemlock- Pacific silverfir, had mean fire return intervals (108-137 years)much lower than in other westernWashington areas. The most interior forest type, ponderosapine - Douglas-fir, had a higher mean fire return interval (52 years)than reported for similar forest typeseastof the Cascades. Historically, fire has createdstructural and landscapediversity on DesolationPeak and may be an important process in the maintenanceof such diversitv into the future. A c E E , J . K . , F t t ' t N t v ,M . , e t D E C - i o u v e r n t N R , . 1 9 9 0 .F o r e s tf i r e h i s t o r y o f D e s o l a t i o nP e a k , W a s h i n g t o n .C a n . J . F o r . R e s .2 0 : 3 5 0 - 3 5 6 . L,esior€ts de la r6gion de Desolation Peak, Washington, sont d'un inter€t ecologiqueparticulier de par leur nature de transition entre les types forestiersde la cOteet ceux de I'int6rieur. Ce territoire se situe d I'ouest de la cr€te des Cascadesmais est une rdgion sous le vent abritee de la pluie des montagnesplus a I'oucst. A causede I'etroite juxtaposition dcs types forestiersd Desolation Peak, on a 6mis I'hypothdseque les cyclesdes incendiesforestiersy sont plus courts que la normale pour les types forestierstypiquesde la c6te comme ceux dominds par la Pruche de I'ouest et le Sapin gracieux,et plus longs que la normale pour les types forestiersde I'int€rieur tels lespinddesd Pin ponderosa. Sept communaut6sforestidresont 6tc identifieeset un historique des incendiesforestiersdes 400 dernidresanndesa 6te dressepour ce tcrritoire de 1500 ha. La pdriodemoyennede rotation des incendie-s forestiersnaturelsest dc 100 ans, v a l e u rv a r i a n t d ' u n f a c t L ' u d r e d e u x s e l o nl c s i d c l ee t I ' e x p o s i t i o nd e l a p e n t e .L e s t y p e s f o r e s t i e r st y p i q u e sd e l a c 6 t e , comme les associationsSapin de Douglas - Pruchc de I'ouest et Pruche subalpine - Sapin gracieux, ont des cycles desincendiesforesticrsbeaucoupplus courts(108-137ans)que ceuxdesautresregionsde I'ouestdu Washington.Le type forestier le plus d I'int6rieur, I'associationPin ponderosa- Sapin de Douglas a un cycle des incendiesforestiersplus long (52 ans) que celui rapporte pour des types forestierssemblablesd I'est des Cascades.Dans le pass6,les incendies ont crde une diversitdde la structureet du paysageforestiersde DesolationPeak. lls pourraient constituerun el6ment i m p o r t a n t d u m a i n t i e n d ' u n e t e l l e d i v e r s i t 6d a n s I ' a v e n i r . [Traduitpar la revue] Inlroduclion The fire history of lorestson the westside ol the Cascade M o u n t a i n s i n W a s h i n g t o n i s u s u a l l y c h a r a c t e r i z e db y infrequent (200- to 500-yearinterval) fires that kill all the trees in the stand (Morris 1934; Hemstrom and Franklin 1982). Fire has been an important disturbance process in this maritime climate area for millennia (Sugita and T s u k a d a 1 9 8 2 ;C w y n a r 1 9 8 7 ) .T r e e r e g e n e r a t i o na f t e r f i r e is often prompt, but may extend in some instancesover d e c a d e s( F r a n k l i n a n d H e m s t r o m 1 9 8l ) . A t h i g h e l e v a t i o n (> 1500m), the tree re-establishmentprocessmay take a century or more (Agee and Smith 1984). In the eastern Cascadesof Washington,with an interior climaticinfluence, the subalpine fire regime is similar to that of west side forests,but the low-elevation( < 1000 m) fire regimeis one of more frequent (10- to 50-yearinterval) fires (Agee 198l), which may resultin forestswith multiple age-classes (Larson l98l). lPresent address:Department of Forestry and RangeManageme-nt,University of California, Berkeley, CA, U.S.A. 'Present address:Bureau of Land Management,Palm Springs, CA, U.S.A. Pnnred in aanadi lnrpnrnrl au ( anada T h e N o r t h C a s c a d e isn W a s h i n g t o n( F i g . I ) i n c l u d el o w Iand forestson the moist westsideof the Cascades, subalpine forestsalong the crest, and interior forestson the dry east side. A typical zonal forest-typesequenceup the coastalwest side of the Cascadesis westernhemlock (Tsuga heterophylla (Raf.) Sarg.), Pacific silver fir (Abies amabilis (Dougl.) Forbes),and mountain hemlock (Tsuga mertensians(Bong.) Carr). A typical zonal forest-typesequenccup the eastside of the Cascadesis ponderosapine (PinusponderosaDougl.), Douglas-fir (Pseudotsugamenziesii(Mirb.) Franco), often rvith grand fir (Abies grandis(Dougl.) Forbes)or lodgepole pine (Pinus contorta Dougl. var. latifolia Engelm.), and subalpine fir (Abies lasioc'arpa(Hook.) Nutt.l (Agee and Kertis 1987). F o r e s t si n t h e v i c i n i t y o f t h e u p p e r S k a g i t R i v e r , R o s s Lake area,are of a specialbotanicalinterestbecauseof therr transitional nature betweenmoist coastal and dry interior forests(Franklin and Dyrness1973).The area is westof thc Cascadecrestbut is in the rainshadowof mountainsfarther t o t h e w e s t , i n c l u d i n gM o u n t B a k e r , M o u n t S h u k s a n ,a n d the Picket Range.The uniquenessof the arealiesin the close juxtaposition of coastal and interior forest communities rather than the presenceof unique community types (Agee AGEE ET AL 351 for determinationof age; selectionwas basedon adequaterepresentation of speciesand sizecohorts in each stand. Wedge samples were cut and removed from 50 trees in all locations where single or multiple fire scarswere present(Arno and Sneck 1977),except along trails. Multiple sampleswere taken in the same area only when there was an obvious differencein the number of scarson adjacent samples. +-'). \, ul 'or,' Vegetation communit-v analysis Two-way speciesindicatory analysis(TwtNSpAN)was used to classifythe vegetationdata (Hilt 1979;Gauch 1982).Speciesimportance valuesusedas inputs to the programswere cover classesfor shrubs and herbs and an averageof relative basal area and relative cover (expressed on the same 1-6 scale)for trees(Agee and Kertis 1987). Fire history analysis Incrementcoreswere mounted on boards, sanded,and counted under a binocular microscope.Total tree age on a sarnplewas determined by adding to the ageof each core an estimatedage lor core height. Correction factors were basedon unpublisheddata from WASHINGTON similar sitesthroughout the region (J.K. Agee, unpublisheddata). At 30 cm height, for example, the correction factor was as follows: Douglas-fir and grand fir, 3 years (mesic site) to 5 years Ftr;. L Locationof DesolationPeak study area (shownin (dry site); lodgepolepine, 3 years(low elevation)to 5 years(high l e c r e a t i oAnr e a( N R A ) i n t h e b l a c k )w i t h i nR o s sL a k eN a t i o n aR elevation). Pacific silver fir and subalpine fir correction factors N o r t h C a s c a d eosf W a s h i n e t o nB. r o k e nl i n e i s t h e c r e s to f t h e were basedon height and the .spacingof the innernrosl rings on CascadeMountains. the sample:at 30 cm height, l0 rings/cm, the correctionfactor was 5 years,while at 30 cm height, 50 rings/cm, the correction factor and Kertis 1987). Commonly, the west side zonal forest was 20 years. Fire dates were recorded from cores (Barrett and Arno 1988) sequenceoccurs on mesic north and east aspects,whereas and wedges.USDA Forest Serviceand the United StatesDepartthe east side zonal forest sequenceis found on drier south ment of the Interior National Park Servicerecordswere reviewed and westslopes. for additional 20th century fire occurrences.Age-classinformaT h e o b j e c t i v eo f t h i s s t u d y w a s t o d e f i n e t h e f o r e s t f i r e tion rvas also used to determinedisturbancedates. Distinct agej u s l h i s t o r yi n t h e v i c i n i t y o l ' D e s o l a t i o nP e a k e a s to f R o s s clas.sseparationsbetweengroups of treeson a plot were determined Lake. It was hypothesizedthat the fire frequenciesfor east from clustersof'tree agesfrom each plot. Each defined age-class sideand westsideforestcommunitieswould reflectthe close was representedby the oldest tree age in the cluster for plotting proximity of thesequite different fire regimes:on the low on the site map. Recruitment periodsafter disturbancein multiaged end of the range for forest types common on the west side, standswere usually lessthan 30 years. Age-classcoverageacross and on the high end of the range for types common to the the study area was mapped using I : 12 000 aerial photography and the mapped age groups in the plots. east side of the Cascades.To achievethis objective it was Although fires weredated to specificyears(e.g., ca. 1648),there necessaryto define the forest community types on Desolawas a lack of correspondence betweendatesamong wedgesamples t i o n P e a k a n d r e c o n s t r u ctth e d i s t u r b a n c eh i s t o r y o v e r t h e (Arno patterns ring owing to unusual annual in the vicinity of .scars p u r p o s e s study area.For thc of this study,a low severity and Sneck 1977);otdesttreesin cohortsof age-class werealso data fire is defined as one that kills primarily understory trees. used to establishdatesof events.Accuracy of the pre-I 800 dates A moderate severity fire kills most understory trees and includesan approximate5-yearrange,while I 800 1900dateshave leavesat leasta patchy overstory, with residualtreesoften about a 2-year range and > 1900dates are accurateto the year. scarred.A high severityfire kills most trees in the stand. In ecosystemswith fire return intervals >20-30 years, dendrochronologicaltechniquesare lesscritical than in ecosystemswith Methods high fire frequency, where individual fire eventsare more easily The Desolation Peakstudyareais definedby RossLakeon the confused (Madany et al. 1982). In areas with longer mean fire west,LightningCreekon thesouthandeast,anda valleyand low return intervals,where fires may often be of high severity,frequent saddleto the north. Ross[,akeis a reservoircreatedin the 1940s fires may be uncommon, but if they do occur, their presenceis in theSkagitRivervalley;manyfiresbeforethenburnedin areas likely to be missed if dendrochronologicaltechniquesare not now inundated.A total of 97 plotsweresubjectively established employed. Furthermore, fire event dates determined without overthe 3500-ha area,representing dendrochronology, while suitable for calculating fire frequencies the differentforesttypesand apparentstandagesbasedon the sizeclassesrepresented in medium to long fire return interval regimes,cannot be usedto in the stand.Fixed100-m'plotsweremostcommonlyused,because in associatefire with climate or other characteristicsof specificyears. mostcasesplotsof this sizeencompassed the rangeof treesizes llistorical fire boundaries were mapped using the age-classmap present in the stand..lnwidelyspaced stands225-or 400-m'plots and fire agesfrom wedgeand core information. Criteria for mapping were similar to those of Hemstrom and Franklin (1982): wereoccasionally employed.Aspect,slope,and elevationwere measuredby compass,clinometer,and altimeter,respectively. (l) Uniformity. The nature of vegetationresponsesto fire has not Diameterat breastheight, total tree height, and basalarea by changed during the period of study. (2) Aee continuity. Trees in stands with similar age cohorts, species wererecordedfor eachtree in the plot. Coverageof subwas determlnedby coverclass(1 : 0-590, ordinatevegetatlon separatecl by younger standsbut not significantvegetationor 2 - 6-25V0, 3 : 26-50V0,4 : 5l-75V0, 5 : 76-95V0,and topographic barriers, probably originated after the same fire 6 : 96-l00Vo).Roughly l0 treesper plot were incrementcored episode. 3s2 CAN. J. FOR. RES. VOL. 20, I99O TABLI l. Average basal area (BA; m2 ha r), cover (CV; 9o), and constancy (CON; 9o) of tree speciesin Desolation Peak forest communlty [ypes Pipo-Psnre BA Abam Abgr Abla Chno Pien Pial Pico Pipo Psme Thpl Tshe Tsnre CV Pico-Psme C]ON BA CV CON Psme-Abgr BA CV CON Psme-Tshe BA CV CON tr tr ll ll Itr12trtr Psme-Abam BA CV Tsme-Abam CON BA CV CON 40 20 20 100 3tr tr tr 3tr tr 4 14 tr 5 25 l4 '70 100 A l 27 5 tr 35 26 1 96 40 l0 50 3tr40 31 50 100 Itr4035 tr tr 16 29 35 13 l0 100 5-s 83 l5 t) 80 604 Pico-Abla BA CV 18 15 tr 6 tr 25 250 tr CON '75 25 7250 6n50 l o r a n a l y s i sw c r e A c m a , b i g l e a l n a p l c a n d P i m o , w c s t c r n w h i l c p i n e . (3) T<tpographicc'onsistency.Disturbances other than fire, such operateon steepgulliedslopes, as avalanches, characteristically whereasmudflows and floods affect standsat valley bottoms. Typically, thesedisturbances do not havesurficialcharcoaland can be distinguishedfrom fires. (4) Conservativelimils. Reconstructedburns are often minimurn reconstructions of past areasburncd and representconservative fire events. Two other assumptionsby Hemstrom and Franklin (1982)were modified for this study: (5) The rcgenerationspanassumptionwith early seraltree cohorts spanning 75 years or more was not used in this study, even though regenerationcan be very slow at elevations> 1500rn has beenmore rapid in most on DesolationPeak. Regeneration locations becauseof the presenceof moderate severity fires evidencedby substantialsurvivalof residualsafter many fires. (6) Unlessspecilicfire evidencewas apparent, fires wereassumed to burn uphill only until they reach a topographic barrier or subalpineforest,sinceother studies(Schmidt 1960;J.K. Agee, unpublisheddata) suggesta dampening of fire spreadand intensity when fires burn upslope into more moist fuels at higher elevation. Fire events,defined as either single fires or a seriesof simultaneous or closelyspacedfires, were reconstructedusing aerial photo and topographic data along with, in priority order, (l) fire scars from wedgesamples,(li ) coresspecificallyextractedto date a fire scar, (lii ) coresexhibiting aberrant ring patterns,generallyassociated with datesdeterminedfrom i or ii, and (i v) age-classdata, particularly where severefires had removed previous evidence. Boundariesof past fires were estimatedby drawing lines equidistant betweenpoints where fire scarsor age-classdata indicated a disarea continuity in past disturbance.This techniqueunderestimates coverageof fires becauselow severityfires may leavelittle or no trace that is recognizabledecadesor centurieslater. Area coverage of each fire was calculatedusing a dot-grid overlay on I : 24 000 scale maps, with 15.5 dots/cm. Natural fire rotations (NFR) (Heinselman 1973)were calculated using the mapped fire areasfor various time periods and geographical units. A selectedtime interval divided by the proportion of total area that is burned within that intervalyieldsthe NFR. Point estimatesof fire frequency, using individual or clustered fire scar samples, are more applicable in very frequent fire regimes where trees are often scarred but rarely killed (Kilgore and Taylor 1979; Dieterich 1980). Fire return interval methods that rely solely on age-class distribution and assumptionsabout randomnessof ignition (Johnsonand Van Wagner 1985)work best in areasof uniform lires. A major disadvantage terrainand stand-replacing of the NFR method is the needlo reconstructpast fire events,which can result in conservativefire return intervals (Van Wagner 1978). Mean fire rcturn intervalswere calculatedfor community types defined by rwtNsnnN by dividing the averagenumber ol fires per plot for a community type into the time period covered.A second method of analyzingfire return intervalsfor vegetationwas used to comparestandspeciescompositionand structurewith mean fire return intervals(MFRls). On eachplot, the two speciesrepresenting the highest (primary dominant) and secondhighest (secondary dominant) tree densityin the combineddominant and codominant tree stratawere identified.The MFRI for a primary dominant specieswas calculatedby pooling all plots where that specieswas a primary dominant and dividing the mean number of fires per plot into the time period covered. The same techniquewas used for secortdarydominartt species. Results Foresl communities Vegetation community types were identified through rwrNSPANon the basis of floristic similarity. Sevenforest community types were identified; vegetationcharacteristics of the sitesare summarizedin Tables I and 2 and Fig. 2. Comnrunitieswere named for two common conifer species; the secondconifer in the name is the likely potential overstory dominant in the long-term absenceof disturbance (plant series sensu Daubenmire 1968), whereas the first named conifer is an associatedseral dominant. Foresl fire events Fires were detectedover a 400-yearperiod on Desolation Peak. Major fires are discernibleover the entire period, but the chanceof reconstructingsmall fire eventsdiminishesas those eventsbecome older. A total of 29 fire eventswere mapped (Figs. 3A-3L). The oldestfire (ca. 1573)was determined from fire wedge information and covered a southwest aspectthat has burned repeatedlysincethat time. The ca. 1648fire was identified from widely scatteredage-classes over the entire study area and probably burned a significant area adjacent to Desolation Peak. The ca. 1733 and ca. 1782fires burned much of the same area covered by the ca. 1573 fire. The ca. 178l fire burned south to southeast aspects;a portion of the ca. 1782 fire was reburned.A fire that probably started in an area that is now under RossLake A G E E F , TA L . 353 T , A , s t - E2 . R e l a t i v e c o n s t a n c y ( 9 0 ) o f m a j o r s h r u b s i n D e s o l a t i o n P e a k f o r e s t c o m m u n i t y types Community type PipoPsnre Acer circinalurn Amelonchier alnifolia A rclostophylos Uva-ursi Berberis nervosu Ceanothus velulinus Corylus cornuta Holodiscus discolor Lonicera hispidula Pachistimo myrsiniles R h odode nd ro n aI b ifl o ru m Rosa gymnocarpo Rubus parviflorus Sallr spp. Spiroea betuli.folia Vaccinium spyt. PicoPsme +l 70 60 90 100 4l 22 59 60 52 26 52 PsnteAbgr PsmeTshe PsrneAbam TsmeAbam PicoAbla )b 80 '76 80 10 88 40 100 72 24 24 36 48 l5 21 96 78 30 20 50 50 60 40 20 z) 50 60 100 100 N ( ) i l r : R e l a t i v ec o n s t a n c yi s t h e p e r c c n l a g co l p l o t s w i t h i n a c o r n n r u n i l yt y p e t h a r c o n l a i n s t h e s p c c i c s .S e eT a b l c I I o r i l r c r n l : i l r J l i r , Io l \ n c ( i c \ u b h r e r i : r t i r r r r i . C o mm u n i t y T y p e s P t P o P 1 ; 1 yRnle P)n)r) Psrne Tsnrt: Ptco Psrne Psnte A b g r 1 5 5 s A b a r n A L x i r | A f ; l : r tr h O ; E'p9lei"ol lltl I II I lP'"i h'"d _5 ++l Pll a J,_r_ O | LttH | lpicol E ,9 4ttt lPieol I -Y l=c| |cI I ll*-d F.j l I a ctt A r-------1 | -T- II | --r- .\r]-.' t|l e E I ll I ce 1i l*.4 1"".{I -= r I ll u/ r- a) | tt I l_ l l___i within eachcommunity Ftc. 2. Percentdensityof treespecies type by forestlayer:0-3 m height,3-10 m height,and >10 m lessthan l09o of relativedensityare height.Species constituting category(shadedblack). lumpedin a miscellaneous burned up two sides of a major west-facing drainage in ca. 1806.An area on the eastside of Desolation Peak, which had not been burned by a major fire for 165 years,burned i n c a . 1 8 1 5 .I n c a . l 8 l 9 a n d c a . 1 8 3 2 ,s m a l l f i r e s b u r n e d lower portions of the southwest-facingslopes. The fire of ca. l85l coveredmost of the mountain. Some areasshown as unburned islandswere not mapped as burned owing to lack of evidence for reconstruction of the fire, although it is possiblethey did burn. A similar area was mapped by Ayres (1899) as an old burn that had not restocked.In ca. 1865. another small fire burned on the w n7 @ W' !) %--) Br---r-\ \/ )\ ffitstz ffiroae 're S tzes l)tez ,C:N vlA rT T- !rasz ffitast fraosffirszzffiiaeo ffiros ffiiaea IN <;7 Y<-,/ \-/ $rars lrsrs '(-i) t.\ /i\ \\ lluffitaoa ffiraoo J,o---4 )I (.. \) <) $% \ \-_./ ltwffiiaea ffitass[rooa Irsrs ffires+firos ffieosltst+ \2/ 'n lil km -F---l o 2 4 6 ffiisza Irgqs eo 810 Irrc. 3. Individualfire eventssinceca. 15"73 at DesolationPeak. Multiplefireyearson a singlemaparedenotedby differentshading. lower elevationsouth slopes.In ca. 1868and ca. 1872,the northeastern portion of the mountain was burned. The ca. 1872 fire, which showed an age-classslightly younger than the lower elevationca. 1868fire, might have beeninterpreted as a singleevent, but fire scar data indicated a separa- CAN. J. FOR. RES. VOL.20, I99O 354 Te,sr-E 3. Natural fire rotations for Desolation Peak Time period Mean fire return interval (years) Natural fire rotation (years) r600- l 699 t'700-1799 r 800-l 899 I 900-I 985 100 208 60 103 I 5 7 3 *r 9 8 5 100 TABr-Fr4. Natural fire rotation by a s p e c to v e r t h e 1 5 7 3 - 1 9 8 5p e r i o d Natural fire rotation (years) North East South Southwcst West TaeLE 5. Mean fire return intervals for forest communitv tvoes on Desolation Peak Ponderosapine - Douglas-iir Lodgeple pine - Douglas-fir Douglas-fir - grand fir Douglas-fir - western hemlock Douglas-fir - Pacific silver fir Mountain hemlock - Pacific silver fir Lodgepole pine - subalpine fir Tneln 6. Mean fire return intervalsfor major species where they are either primary or r e c o n d a r yd o n r i n a n t s Mean fire return interval (years) t82 107 9'7 65 t20 t i o n o f 4 y e a r s .I n c a . 1 8 8 0 ,m u c h o f t h e w e s t s l o p eo f t h e mountain burned, and over the next 4 decadessmall fires occurredin scattcredlocationsaround the mountain. Other small fires may haveoccurredearlierin the record,but could not be identified on the landscape. T h e a r e ao f t h e l 9 l 9 f i r e w a s r e b u r n e db y t h e l a r g e r f i r e o f 1 9 2 6 .T h e 1 9 1 9f i r e w a s r o m a n t i c i z e di n J a c k K e r o u a c ' s (1958, 1960) novels The Dharma Bums and Desolation Angels, although he assumedthe burned area was a singlc large fire in 1919.The first fire apparentlyburned primarily in subalpineforests(Fig. 3K), whereasthe secondwas part of the 20 000-ha Big Beaver fire that started near what is t o d a y t h e s o u t he n d o f R o s sL a k e , j u m p e d t h e S k a g i tR i v e r , and endednorth of DesolationPeak (Fig. 3L). Sincethe big 1926fire, only scatteredareashave burned in a period whcn a fire suppressionpolicy has been in effect. Patterns of fire frequency Natural fire rotations were calculatedfor the entire area o v e r t h e t i m e p e r i o d 1 5 7 3 - 1 9 8 5a n d f o r e a c h c e n t u r y (Table 3). The NFRs were also calculated by aspect (Table 4). The average NFR for the area was 100 years, interpretedto mean that in the courseof a century an area equal in sizeto the study areawould burn; someareaswould burn twice and others not at all. Wide temporal and spatial variations are evident during the time period covered by the study. The post-1900period has a NFR equal to the averageover the whole study period, whereasthe period 1800-1899has a low NFR. The l'100-1799period has a high NFR, whereasthe large fire in ca. 1648resultsin an average NFR (100 years)for the 1600-1699period. lnfrequent fire eventsof wide extent markedly affect the calculations, and the probable occurrenceof additional but unidentified fires in the 1600-1700period may make this NFR conservative. Southwestaspectsburned most frequently whereas north aspectsburned least frequently. Mean fire return intervals for community types (Table 5) showed a range of 52-137 years. The ponderosa pine Douglas-fir type has the shortest MFRI; it is most common 52 76 93 l3'1 108 t3'7 109 Ponderosapine Lodgepolc pine Douglas-fir Weslern red cedar W e s t e r nh e r n l o c k Pacific silver fir S u b a l p i n ef i r Mountain hemlock Primary dominant Secondary dominant 44 69 94 > 4l I 169 192 154 137 56 82 84 204 t26 r08 89 I z-t on the southwesa t s p e c t st h a t h a v e b u r n e d r e p e a t e d l yw i t h low severity fires. The lodgepole pine - Douglas-fir comm u n i t y , w i t h t h e n e x t l o w e s tM F R I , i s f o u n d o n m o s t o t h e r aspectsat low elevation.The Douglas-fir - grand fir community is found in protected lorv-elevationdraws, which o c c a s i o n a l l yh a v e n o t b u r n e d , s o i t s M F R I i s h i g h e r . T h e Douglas-fir - westernhemlock community has burned the leastof the low-elevationcommunities.It is found primarily on north aspectsthat are floristicallyand climaticallysimilar to moist lowland forestsfarther to the west. The remaining community typestend to be subalpineforest typesand have m o r e s i m i l a r M F R I s t h a n t h e m o n t a n ef o r e s t s .T h i s i s d u e to the widespreadnature of past high-elevationfires and limited geographicseparationbetweenthe subalpinetypes. Fire frequency variation was also assessedfor species wherethey were primary or secondarydominants(Table 6). Ponderosa pine and lodgepole pine were the only species to have shorterMFRIs wherethey wereprimary dominants. These are the only two major speciesat Desolation Peak that are restrictedto early seralstatus;all of the other species or potentialvegetationdominants(sensu are late successional Daubenmire 1968)in some locations. As their dominance dependson seral community maintenancethrough disturbance, the early seral speciesshould be more dominant where disturbanceis more frequent and the MFRI is lower. Speciesthat are later successionalor relatively fire sensitive, including subalpine fir, mountain hemlock, and Pacific silver fir, have longer MFRIs where they are primary dominants and tend to be secondarydominants wheredisturbance by fire is more frequent. Douglas-fir is both a major seral speciesin middle- to high-elevation communities and AGEE ET AL a major potential vegetation dominant in the drier lowland communities. Over the study area, it has roughly equal MFRIs whether it is a primary or secondarydominant. Discussion The spatial arrangementof the forest communities on DesolationPeak is influencedby temperatureand moisture gradients,which were also found to be important in other Pacific Northwest forests(Zobel et al. 1976;del Moral and Watson 1978)and in the North Cascades(Agee and Kertis l 987). Variable fire environmentson DesolationPeak are illustrated by the spectrumof fire return intervals in the forest community types. The communitiesin drier, warmer environments have the shortestMFRIs, whereasthe communities in cooler, wetter environmentshave the longestMFRIs. Thesefire fiequenciesalso reflect the landscapeheterogeneity within which they occur. At DesolationPeak, the interspersionof west side and east side types has a significant influence on fire frequencywithin a forestcommunity. The ponderosapine - Douglas-fircommunity has an MFRI of 52 years, far higher than fire intervals for the type elsewherein the Pacific Northwest where environmental gradients are more gradual and a homogeneous contiguousforestis present(Wischnofskeand A n d e r s o n1 9 8 3 ;F i n c h 1 9 8 4 ;J . K . A g e e ,u n p u b l i s h e dd a t a ) . The lodgepolepine - Douglas-fir community has an average MFRI higherthan typically found in similar forest types of westernMontana (Arno 1980).The Douglas-fir- western hemlockcommunity fire intervalsare below thosecalculated for either Douglas-fir (230 years) or western hemlock (598 years)for westernWashington (Fahnestockand Agee 1983),but are closeto thoseof 100 yearsfrom Means(1982) and Morrison and Swanson (1990) for dry Douglas-fir forests of the Oregon Cascades.The Douglas-fir - grand f i r c o m m u n i t yh a sa n M F R I w i t h i n t h e w i d e r a n g eo f A r n o (1980)from the northernRocky Mountainsand Wischnolske and Anderson (1983) from the WenatcheeValley of W a s h i n g t o n .T h c m o s t c o o l a n d m o i s t f o r e s t e o m m u n i t y , t h e m o u n t a i n h e m l o c k- P a c i f i c s i l v e rf i r c o m m u n i t y , h a s a n M F R I f a r b e l o w t h e 4 3 4 y e a r sf o r s i m i l a r f o r e s t t y p e s a t M o u n t R a i n i e r( H e m s t r o ma n d F r a n k l i n 1 9 8 2 ) .T h c f i r e return intervals of Desolation Peak forest communities appearto reflect the juxtaposition of forest types not commonly found together,lengtheningfire return intervals for communities with typically frequent fire regimes and shorteningthem for communities with typically infequent fire regimes. Although forest environmentand interspersionof forest typesare important influenceson fire return intervals,patterns of lightning and aboriginal ignitions are also important. Indiansinhabitedthe Skagit River valley in the vicinity of DesolationPeak, and a preliminary analysisof fire scars on stumps now underwater most of the year about 5 km north of the study area suggestsa l0- to l5-year fire return interval on the vallby floor (Taylor 1977).Such closelyplaced fires are generally absent from the record on Desolation Peak, suggestingmore frequent burning in the vicinity of the Indian village (e.g., Barrett and Arno 1982).Although mining era fires were common in the Cascades(Mack 1988) during the last half of the l9th century, the settler influence found elsewhere in the North Cascades (Ayres 1899; J)) Hemstrom and Franklin 1982) is not locally apparent at Desolation Peak. The short NFR for the lTth century and the long NFR for the l8th century would not be notable if Desolation Peak were the only area in the Pacific Northwest to show such a record. However, a similar pattern is reported for the forests at Mount Rainier (Hemstrom and Franklin 1982), the southern Olympic Mountains (Henderson and Peter l98l), and the west-centralCascadesof Oregon (Morrison and Swanson 1990).This apparent regional pattern is not explained by any of the separatestudies, but may reflect climatic changessuch as the Little Ice Age (Hendersonand Brubaker 1986)and associatedchangesin biomass flammability, dry summer winds, or lightning storm frequency. The forests of Desolation Peak have been significantly influencedby fire over the past half millennia and no doubt for millennia before. The preservationof the structuraland landscapediversity of thesenatural ecosystemswill require maintenanceof thesedisturbancepatterns into the future. Over the last 4 centuriesa fire has burned on Desolation Peak about every 15 years.Although it is unlikely the major seral dominants ponderosapine and lodgepolepine would disappearwith completefire exclusion,the spatialextentof thesecommunitieswould shrink over time. Forest communities dominated by these speciesare unusual west of the Cascadecrest,and future activemanagementof disturbance by fire will help maintain this unique landscapediversity. Acknowledgment This researchwas supported by cooperalive agreement CA-9000-3-0004, subagreements5 and 8, between the N a t i o n a l P a r k S e r v i c ea n d t h e U n i v e r s i t vo f W a s h i n e t o n . Ac;ut.., J.K. 1981.Firecffectson PacificNorthwestforests:I'lora, fuels, and fauna. NorthwestFire Council 198I Conference , R. P r o c e e d i n gN s ,o r t h w e sFt i r eC o u n c i l ,P o r t l a n dO A ( ; n E ,J . K . , a n d K t n l ' l s , J . 1 9 8 7 .F o r e s tt y p e so f t h e N o r t h Cascades National Park ServiceComplex.Can. J. Bot. 65: l 520-I 530. . u b a l p i nter e er e c s t a b l i s h m e n t A r ; l l , J . K . , a n d S u l ' t ' t t L, . 1 9 8 4S Ecology,65: after fire in theOtympicMountains,Washington. 810-819. .1.For. AnNo,S. 1980.I"orestfire historyin thenorthernRockies. 78: 460-465. A n N o , S . , a n d S N n c x ,K . 1 9 7 ' 7A. m e t h o df o r d e t e r m i n i nfgi r e West.USDA forestsof the Intermountain historyin coniferous For. Serv.Gen. Tech. Rep. INT-42. Aynrs, H.B. 1899.WashingtonForestReserve./r? Nineteenth Surveyannualreport.Part 5. ForestReserves. U.S. Ceological C o v e r n m e nPt r i n t i n gO f f i c e ,W a s h i n g t o nD, C . p p . 2 8 3 - 3 1 3 . Bannerr, S., and AnNo, S. 1982.Indian fires as an ecological factor in the northernRockies.J. For. 60: 647-651. 1988.Increment-borer methodsfor determiningfire history in coniferousforests.USDA For. Serv.Gen.Tech.Rep. tNT-244. CwyNRn, L. 1987.Fire and the forest history of the North CascadeRange.Ecology,68: 791-802. DeusENt'4rne, R. 1968.Plant communities.Harper and Row, New York. DELMoRAL,R., and WarsoN,A.F. 1978.Gradientstructureof forestvegetation in the centralWashingtonCascades. Vegetatio, 3E: 29-48. DTETERTCH, J. 1980.The compositefire interval-a tool for more accurateinterpretation of fire history.In Fire HistoryWorkshop. 356 CAN. J. F'OR. RES. VOL. 20, I99O Edited by M.A. Stokesand J. Dieterich.USDA For. Serv. Gen. Tech. Rep. RM-81. pp. 8-14. FAHNESTocK, G.R., and AGEE,J.K. 1983.Biomassconsumption and smoke production by prehistoric and modern forest fires i n w e s t e r nW a s h i n g t o n .J . F o r . E l : 6 5 3 - 6 5 7 . FtNCH,R.B. 1984.Fire history of selectedsiteson the Okanogan National Forest. USDA Forest Service, Pacific Northwest Region, Okanogan National Forest, Okanogan, WA. F n a N x r - r N ,J . , a n d D v n N u s s , C . 1 9 7 3 . N a t u r a l v e g e t a t i o no f Oregon and Washington. USDA For. Serv. Cen. Tech. Rep. PNW-8. F R A N K L I NJ, . , a n d H e v s r n o v , M . 1 9 8 1 .A s p e c t so f s u c c e s s i o n in the coniferousforestsof the Pacific Northwest. 1n Forestsuccession: concepts and application. Chap. 14. Edited by D . C . W e s t , H . H . S h u g a r t ,a n d D . B . B o t k i n . S p r i n g e r - V e r l a g , New York. GAUcH, H.G. 1982.Multivariate analysisin community ecology. Cambridge University Press, New York. H L , T N S E L M AMN. , 1 9 7 3 .F i r e i n t h e v i r g i n f o r e s t so f t h e B o u n d a r y Waters Canoe Area, Minnesota. Quat. Res. 3: 329-382. H E M S T R o MM, . , a n d F n , q N x t - t NJ,. F . 1 9 8 2 .F i r e a n d o t h e r d i s t u r bancesof the forestsin Mount Rainier National Park. Quat. Res. l8:32-51. H n N o r n s o n , J . , a n d B n u g a x n n , L . 1 9 8 6 .R e s p o n s eosf D o u g l a s fir to long-term variations in precipitation and temperature in westernWashington.1n Douglas-fir:stand managementfor the f u t u r e . C h a p . 2 3 . U n i v e r s i t yo f W a s h i n g t o n ,S e a t t l e .I n s t i t u t e ol Forest ResourcesContribution No. 55. H E N t ) E R S O JN.,, a n d P n r n n , D . 1 9 8 1 .P r e l i m i n a r yp l a n t a s s o c i a tions and habitat typesof the Shelton Ranger District, Olympic National Forest. USDA Forest Service, Pacific Northwest R e g i o n ,P o r t l a n d ,O R . Hrlr-, M.O. 1979. r'wtNspnN-a FoRIRANprogram for arranging multivariate data in an ordered two-way table by classification of the individuals and attributes. Cornell University, Ithaca, New York. J o t n r s o N ,8 . A . , a n d V e r W n c ; N E n ,C . E . 1 9 8 5 .T h e t h e o r y a n d use of two fire history models. Can. .1.For. Res. 15: 214-220. K n p o u n c , J . 1 9 5 8 .T h e D h a r m a b u m s . V i k i n g P r e s s ,N e w Y o r k . 1960. Desolation angels. Viking Press, New York. KIt-(ioRE,B., and Tavlon, D. 1979. Fire history of a sequoiamixed conifer forest. Ecology, 60: 129-142. LARSoN,B.C. 1981.Developmentof even-agedand uneven-aged mixed conifer standsin easternWashington. 1n Forest Succession and Stand Development Research in the Northwest: Proceedingsof the Symposium,26 Mar. 1981,OregonStateUniversity, Corvallis. Forest ResearchLaboratory, Oregon State U n i v e r s i t y ,C o r v a l l i s .p p . l l 3 - l 1 8 . MRcr, R.N. 1988. First comprehensivebotanical survey of the Columbia Plateau, Washington: the Sandberg and Leiberg . ci.62: ll8-128. e x p e d i t i o no f 1 8 9 3 .N o r t h w e s t S M a o r r N y , M . H . , S w r r N A M , T . W . , a n d W n s r , N . E . 1 9 8 2 .C o m parison of two approachesfor determiningfire datesfrom tree s c a r s .F o r . S c i . 2 E : 8 5 6 - 8 6 1 . M r a N s , J . E . 1 9 8 2 . D e v e l o p m e n t a lh i s t o r y o f d r y c o n i f e r o u s forestsin the westernOregon Cascades.1n Forest Successionand Stand DevelopmentResearchin the Northwest: Proceedingsof the Symposium, 26 Mar. 1981, Oregon State University, Corvallis. Forest ResearchLaboratory, Oregon State University, Corvallisp . p. 142-158. Monnts, W.G. 1934. Forest fires in Oregon and Washington. Oreg. Hist. Q. 35: 313-339. M o R R t s o N ,P . , a n d S w n N s o N ,F . 1 9 9 0 .F i r e h i s t o r yi n t w o f o r e s t ecosystemsof the central westernCascadesof Oregon. USDA For. Serv.Cen. Tech. Rep. PNW. In press. Sc'uvtur', R.L. 1960. Factorscontrolling the distributionof Douglas-firin coastalBritish Columbia. Q. J. For. 54: 156-160. S u c r r a , S . , a n d T s u r n o n , M . 1 9 8 2 .T h e v e g e t a t i o nh i s t o r y i n westernNorth America. L Hall and Mineral lakes.Jpn. .f. Bot. (Tokyo),95:410-418. T n v r . o n , D . 1 9 7 7 .S o m e p r e l i m i n a r yo b s e r v a t i o n o s f the history of natural fircs in the Skagit District. [Report on file.] North CascadesNational Park ServiceComplex, SedroWoolley, WA. V A N W A G N E RC , . E . 1 9 7 8 .A g e - c l a s sd i s t r i b u t i o na n d t h e f o r e s t fire cycle. Can. J. For. Res. E: 220-22'7. W r s c H N o r . ' s r pM , . G . , a n d A N o E n s o N ,D . W . 1 9 8 3 .T h e n a t u r a l role oI fire in WenatchccValley. USDA ForestService,Pacil'ic Northwest Region, Wenatchee National Forest, Washington State. Z o t s E L ,D , . B . , M c K e e , A . , H a w x , G . M . , a n d D y R N F t s sC, . T . 1 9 7 6 .R e l a t i o n s h i pos i e n v i r o n m e n lt o c o m p o s i t i o n ,s l r u c t u r e , and diversity of forest communities of the central western C a s c a d e so f O r e g o n . E c o l . M o n o g r . 4 6 : 1 3 5 - 1 5 6 .
© Copyright 2024 Paperzz